<documentid="1B8E4D1A45A387C27C369ABDAE8E63AD"ID-CLB-Dataset="298361"ID-DOI="10.11646/zootaxa.5458.3.2"ID-GBIF-Dataset="18d6f856-ced1-49b9-b460-5b7bdf073060"ID-ISSN="1175-5326"ID-Zenodo-Dep="11547469"ID-ZooBank="54609B08-07DE-4009-B15E-FEBE2C577AAE"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="diego"IM.materialsCitations_approvedBy="diego"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch,operationResults"IM.taxonomicNames_approvedBy="diego"IM.treatments_approvedBy="diego"checkinTime="1718021059398"checkinUser="plazi"docAuthor="Gower, David J., Sampaio, Filipa L., Vidanapathirana, Dulan Ranga & Mendis Wickramasinghe, L. J."docDate="2024"docId="157D87FC7677A15CFF02FB78794972D7"docLanguage="en"docName="zootaxa.5458.3.2.pdf"docOrigin="Zootaxa 5458 (3)"docSource="http://dx.doi.org/10.11646/zootaxa.5458.3.2"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Rhinophis martin Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe 2024, sp. nov."docType="treatment"docVersion="4"lastPageNumber="343"masterDocId="E944FF847676A157FF95FFC17D60750A"masterDocTitle="Two new species of the shieldtail snake genus Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae), from the Rakwana and the Knuckles Massifs of Sri Lanka"masterLastPageNumber="360"masterPageNumber="332"pageNumber="333"updateTime="1718118452382"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="C11DB4938FB116154F0380B8BECF0CAB">Two new species of the shieldtail snake genus Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae), from the Rakwana and the Knuckles Massifs of Sri Lanka</mods:title>
<mods:namePartid="066A3AE1524077E9BF3E2826D758330F">Gower, David J.</mods:namePart>
<mods:affiliationid="21804D3C6BEE32571E956568D8E12D98">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
<mods:affiliationid="91D72145D395885FDAAA8FBD7EA64487">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK & Department of Genetics, Evolution and Environment, University College London, London WC 1 E 6 BT, UK</mods:affiliation>
<mods:affiliationid="31338EAD839B7E76BDC1CAB95BB7223B">Herpetological Foundation of Sri Lanka, 31 / 5, Alwis Town, Hendala, Wattala, Sri Lanka</mods:affiliation>
<mods:namePartid="731C87410B7CC904F07CCFFF98AF99D2">Mendis Wickramasinghe, L. J.</mods:namePart>
<mods:affiliationid="AD694C5382A11E24A688D9520960382C">Herpetological Foundation of Sri Lanka, 31 / 5, Alwis Town, Hendala, Wattala, Sri Lanka</mods:affiliation>
<taxonomicNameid="5AD44D697677A156FF02FB787C0E71D9"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[151,366,1209,1236]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="1"pageNumber="333"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<figureCitationid="05EF2A6F7677A156FF02FB1C7C6271FD"box="[151,258,1245,1271]"captionStart-0="FIGURE 1"captionStart-1="FIGURE 2"captionStart-2="FIGURE 3"captionStart-3="FIGURE 4"captionStart-4="FIGURE 5"captionStart-5="FIGURE 6"captionStartId-0="2.[151,250,1943,1967]"captionStartId-1="3.[151,250,1911,1935]"captionStartId-2="4.[151,250,1543,1567]"captionStartId-3="9.[151,250,1537,1561]"captionStartId-4="10.[151,250,1448,1472]"captionStartId-5="10.[151,250,1997,2021]"captionTargetBox-0="[228,1358,179,1916]"captionTargetBox-1="[273,1315,185,1881]"captionTargetBox-2="[170,1416,181,1520]"captionTargetBox-3="[151,1435,237,1513]"captionTargetBox-4="[152,1398,533,1406]"captionTargetBox-5="[151,1435,1533,1974]"captionTargetId-0="figure-18@2.[228,1358,179,1916]"captionTargetId-1="figure-31@3.[266,1321,181,1887]"captionTargetId-2="figure-18@4.[170,1416,181,1520]"captionTargetId-3="figure-62@9.[151,1435,237,1513]"captionTargetId-4="figure-240@10.[151,1435,532,1412]"captionTargetId-5="figure-217@10.[151,1435,1533,1974]"captionTargetPageId-0="2"captionTargetPageId-1="3"captionTargetPageId-2="4"captionTargetPageId-3="9"captionTargetPageId-4="10"captionTargetPageId-5="10"captionText-0="FIGURE 1. Holotype of Rhinophis martin sp. nov., USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor."captionText-1="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)."captionText-2="FIGURE 3. Seven paratypes of Rhinophis martin sp. nov. Left column approximately dorsal view; right column approximately ventral view. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bars 20 mm. Photographs by H. Taylor."captionText-3="FIGURE 4. Head and tail ends of seven paratypes of Rhinophis martin sp. nov. in four views. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bar increments in millimetres. Photographs by H. Taylor."captionText-4="FIGURE 5. Graph showing relationship between number of ventral scales and number of subcaudal scales (mean value for left and right) in female (triangles) and male (circles) Rhinophis martin sp. nov."captionText-5="FIGURE 6. Rhinophis martin sp. nov., paratype DWC 2024.05.01 prior to fixation, in upper (left) and lower (right) views."figureDoi-0="http://doi.org/10.5281/zenodo.11547471"figureDoi-1="http://doi.org/10.5281/zenodo.11547474"figureDoi-2="http://doi.org/10.5281/zenodo.11547476"figureDoi-3="http://doi.org/10.5281/zenodo.11547478"figureDoi-4="http://doi.org/10.5281/zenodo.11547480"figureDoi-5="http://doi.org/10.5281/zenodo.11547482"httpUri-0="https://zenodo.org/record/11547471/files/figure.png"httpUri-1="https://zenodo.org/record/11547474/files/figure.png"httpUri-2="https://zenodo.org/record/11547476/files/figure.png"httpUri-3="https://zenodo.org/record/11547478/files/figure.png"httpUri-4="https://zenodo.org/record/11547480/files/figure.png"httpUri-5="https://zenodo.org/record/11547482/files/figure.png"pageId="1"pageNumber="333">Figs. 1–6</figureCitation>
<tableCitationid="D05603517677A156FE9BFB1C7CEA71FD"box="[270,394,1245,1271]"captionStart-0="TABLE 1"captionStart-1="TABLE 2"captionStart-2="TABLE 3"captionStartId-0="5.[151,239,151,175]"captionStartId-1="6.[151,239,692,716]"captionStartId-2="8.[151,239,368,392]"captionTargetPageId-0="5"captionTargetPageId-1="6"captionTargetPageId-2="8"captionText-0="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded."captionText-1="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded"captionText-2="TABLE 3. Uncorrected p-distances (%) for intra- and interspecific comparisons for Rhinophis martin sp. nov. and closest sampled relatives, for two nuclear (prlr, cmos) and four mitochondrial genes (12s, 16s, cytb, nd4) as analysed by Sampaio et al. (2023). Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison."httpUri-0="http://table.plazi.org/id/C9AB66627673A152FF02FF56793E7411"httpUri-1="http://table.plazi.org/id/C9AB66627670A151FF02FD757CD5761E"httpUri-2="http://table.plazi.org/id/C9AB6662767EA15FFF02FEB17ED474FE"pageId="1"pageNumber="333"tableUuid-0="C9AB66627673A152FF02FF56793E7411"tableUuid-1="C9AB66627670A151FF02FD757CD5761E"tableUuid-2="C9AB6662767EA15FFF02FEB17ED474FE">Tables 1–3</tableCitation>
<bibRefCitationid="F9454B1B7677A156FE56FA647FD670B5"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."box="[451,694,1445,1471]"pageId="1"pageNumber="333"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
<figureCitationid="05EF2A6F7677A156FE45FA297F747308"box="[464,532,1512,1538]"captionStart="FIGURE 1"captionStartId="2.[151,250,1943,1967]"captionTargetBox="[228,1358,179,1916]"captionTargetId="figure-18@2.[228,1358,179,1916]"captionTargetPageId="2"captionText="FIGURE 1. Holotype of Rhinophis martin sp. nov., USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor."figureDoi="http://doi.org/10.5281/zenodo.11547471"httpUri="https://zenodo.org/record/11547471/files/figure.png"pageId="1"pageNumber="333">Fig. 1</figureCitation>
<tableCitationid="D05603517677A156FDB6FA297F167308"box="[547,630,1512,1538]"captionStart="TABLE 1"captionStartId="5.[151,239,151,175]"captionTargetPageId="5"captionText="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded."httpUri="http://table.plazi.org/id/C9AB66627673A152FF02FF56793E7411"pageId="1"pageNumber="333"tableUuid="C9AB66627673A152FF02FF56793E7411">Table 1</tableCitation>
) is a tea estate and Gongala is a nearby mountain (peaking at 6.386° N, 80.654° E; 1,358 m elevation), <2 km to the north-northwest on the eastern edge of Sinharaja Forest Reserve, part of the Rakwana Massif (
<figureCitationid="05EF2A6F7677A156FEF7F9957CC77364"box="[354,423,1620,1646]"captionStart="FIGURE 2"captionStartId="3.[151,250,1911,1935]"captionTargetBox="[273,1315,185,1881]"captionTargetId="figure-31@3.[266,1321,181,1887]"captionTargetPageId="3"captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)."figureDoi="http://doi.org/10.5281/zenodo.11547474"httpUri="https://zenodo.org/record/11547474/files/figure.png"pageId="1"pageNumber="333">Fig. 2</figureCitation>
<collectingDateid="F92EE9C27677A156FCDCF99579687364"box="[841,1032,1620,1646]"pageId="1"pageNumber="333"value="1979-10-24">24 October 1979</collectingDate>
<tableCitationid="D05603517677A156FE33F9B97C987398"box="[422,504,1656,1682]"captionStart="TABLE 1"captionStartId="5.[151,239,151,175]"captionTargetPageId="5"captionText="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded."httpUri="http://table.plazi.org/id/C9AB66627673A152FF02FF56793E7411"pageId="1"pageNumber="333"tableUuid="C9AB66627673A152FF02FF56793E7411">Table 1</tableCitation>
<tableCitationid="D05603517677A156FD85F8917F027260"box="[528,610,1872,1898]"captionStart="TABLE 2"captionStartId="6.[151,239,692,716]"captionTargetPageId="6"captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded"httpUri="http://table.plazi.org/id/C9AB66627670A151FF02FD757CD5761E"pageId="1"pageNumber="333"tableUuid="C9AB66627670A151FF02FD757CD5761E">Table 2</tableCitation>
<tableCitationid="D05603517677A156FE23F8B57F6E7284"box="[438,526,1908,1934]"captionStart="TABLE 2"captionStartId="6.[151,239,692,716]"captionTargetPageId="6"captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded"httpUri="http://table.plazi.org/id/C9AB66627670A151FF02FD757CD5761E"pageId="1"pageNumber="333"tableUuid="C9AB66627670A151FF02FD757CD5761E">Table 2</tableCitation>
): USNM 548096–548099 and 548101–548110 collected by Lalith Jayawickrama from Hayes 14 July–18 August, 1977; USNM 548111–548116, 548124, 548125 collected by Lalith Jayawickrama from Hayes/Gongala 24–26 October, 1979. These are considered referred rather than
<taxonomicNameid="5AD44D697674A155FE00F8567F2572A5"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[405,581,1943,1967]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="2"pageNumber="334"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
, USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor.
Map showing places relevant to the distribution of
<taxonomicNameid="5AD44D697675A154FC81F8B67EA47285"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[788,964,1911,1935]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="3"pageNumber="335"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
, including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for
<taxonomicNameid="5AD44D697675A154FB7CF85D782872B9"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[1257,1352,1948,1971]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="3"pageNumber="335"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details).
<taxonomicNameid="5AD44D697672A153FE48F9C97FED7315"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[477,653,1544,1568]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="4"pageNumber="336"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
females), a large, domed tail shield with small, homogenous spines, and a distinctive colour pattern (at least in adults) with a dark belly and paler, orange-bronze dorsum (except for darker anterior sixth) with irregular, blackish transverse markings.
<taxonomicNameid="5AD44D697672A153FF52F8857CEC7254"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[199,396,1860,1886]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="4"pageNumber="336"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D697672A153FDD3F8A87E977289"authority="Aengals and Ganesh, 2013"authorityName="Aengals and Ganesh"authorityYear="2013"box="[582,1015,1896,1923]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="4"pageNumber="336"phylum="Chordata"rank="species"species="goweri">
<bibRefCitationid="F9454B1B7672A153FD2AF8A97E977289"author="Aengals, R. & Ganesh, S. R."box="[703,1015,1896,1923]"pageId="4"pageNumber="336"pagination="61 - 65"refId="ref15770"refString="Aengals, R. & Ganesh, S. R. (2013) Rhinophis goweri - a new species of shieldtail snake from the Southern Eastern Ghats, India. Russian Journal of Herpetology, 20 (1), 61 - 65."type="journal article"year="2013">Aengals and Ganesh, 2013</bibRefCitation>
<bibRefCitationid="F9454B1B7672A153FC10F8717DB272E5"author="Cyriac, V. P. & Narayanan, S. & Sampaio, F. L. & Umesh, P. & Gower, D. J."pageId="4"pageNumber="336"pagination="329 - 342"refId="ref16068"refString="Cyriac, V. P., Narayanan, S., Sampaio, F. L., Umesh, P. & Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the Wayanad region of peninsular India. Zootaxa, 4778 (2), 329 - 342. https: // doi. org / 10.11646 / zootaxa. 4778.2.5"type="journal article"year="2020">Cyriac, Narayanan, Sampaio, Umesh & Gower, 2020</bibRefCitation>
</taxonomicName>
, and
<taxonomicNameid="5AD44D697672A153FE98F814797372E5"authority="Sampaio, Narayanan, Cyriac, Venu & Gower, 2020"authorityName="Sampaio, Narayanan, Cyriac, Venu & Gower"authorityYear="2020"box="[269,1043,2004,2031]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="4"pageNumber="336"phylum="Chordata"rank="species"species="karinthandani">
<bibRefCitationid="F9454B1B7672A153FE44F815797372E5"author="Sampaio, F. L. & Narayanan, S. & Cyriac, V. P. & Venu, G. & Gower, D. J."box="[465,1043,2004,2031]"pageId="4"pageNumber="336"refId="ref16885"refString="Sampaio, F. L., Narayanan, S., Cyriac, V. P., Venu, G. & Gower, D. J. (2020) A new Indian species of Rhinophis Hemprich, 1820"type="book"year="2020">Sampaio, Narayanan, Cyriac, Venu & Gower, 2020</bibRefCitation>
Meristic and morphometric (in mm) data for holotype (*) and paratypes of
<taxonomicNameid="5AD44D697673A152FB8DFF5679B075A5"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[1048,1232,151,175]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded.
<taxonomicNameid="5AD44D697673A152FDB9FAF77F92705A"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[556,754,1334,1360]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
in not having a flat tail shield or midline contact between the opposite nasal shields, and by having dorsal scales in 17 rather than 19 rows at midbody. The new species differs from
<taxonomicNameid="5AD44D697673A152FE8CFA62791B70B6"authority="Wickramasinghe, Vidanapathirana, Rajeev & Gower, 2017"authorityName="Wickramasinghe, Vidanapathirana, Rajeev & Gower"authorityYear="2017"box="[281,1147,1442,1468]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="roshanpererai">
<bibRefCitationid="F9454B1B7673A152FE74FA63791B70B6"author="Wickramasinghe, L. M. & Vidanapathirana, D. R. & Rajeev, M. G. & Gower, D. J."box="[481,1147,1442,1468]"pageId="5"pageNumber="337"pagination="153 - 164"refId="ref17165"refString="Wickramasinghe, L. M., Vidanapathirana, D. R., Rajeev, M. G. & Gower, D. J. (2017) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the central hills of Sri Lanka. Zootaxa, 4263 (1), 153 - 164. https: // doi. org / 10.11646 / zootaxa. 4263.1.7"type="journal article"year="2017">
Wickramasinghe, Vidanapathirana, Rajeev & Gower,
in having homogenously sized spines on a regularly domed tail shield (versus a less prominent and more-irregularly shaped tail shield with heterogeneously sized spines).
<taxonomicNameid="5AD44D697673A152FD96FA2B7FAB730E"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[515,715,1514,1540]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D697673A152FB2CF9CE7C3D7346"authority="Gower & Maduwage, 2011"authorityName="Gower & Maduwage"authorityYear="2011"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="zigzag">
<bibRefCitationid="F9454B1B7673A152FABAF9CF7C3D7346"author="Gower, D. J. & Maduwage, K."pageId="5"pageNumber="337"pagination="51 - 68"refId="ref16404"refString="Gower, D. J. & Maduwage, K. (2011) Two new species of Rhinophis Hemprich (Serpentes: Uropeltidae) from Sri Lanka. Zootaxa, 2881 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2881.1.4"type="journal article"year="2011">Gower & Maduwage, 2011</bibRefCitation>
</taxonomicName>
by having fewer than 190 ventral scales (versus more than 205).
<taxonomicNameid="5AD44D697673A152FBF9F9F378547346"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[1132,1332,1586,1612]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D697673A152FE6AF9967DAF739E"authority="Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella, 2009"authorityName="Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella"authorityYear="2009"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="erangarviraji">
<bibRefCitationid="F9454B1B7673A152FD29F9977DAF739E"author="Wickramasinghe, L. J. M. & Vidanapathirana, D. R. & Wickramasinghe, N. & Ranwella, P. N."pageId="5"pageNumber="337"pagination="1 - 22"refId="ref17090"refString="Wickramasinghe, L. J. M., Vidanapathirana, D. R., Wickramasinghe, N. & Ranwella, P. N. (2009) A new species of Rhinophis Hemprich, 1820 (Reptilia: Serpentes: Uropeltidae) from Rakwana massif, Sri Lanka. Zootaxa, 2044 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2044.1.1"type="journal article"year="2009">
Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella,
in having more than 166 ventral scales (versus fewer than 164).
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(154–168 and 164–179, respectively), and additionally differs from the former in having a more protuberant, conical (versus less-prominently domed) tail shield and from the latter in having a paler orange-bronze dorsum (versus darker, brown-grey) and having (versus lacking) substantial yellow-orange markings on the underside of the tail.
<taxonomicNameid="5AD44D697673A152FE4BF8EF7FC47242"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[478,676,1838,1864]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D697673A152FECCF8927E737266"authority="Gower & Maduwage, 2011"authorityName="Gower & Maduwage"authorityYear="2011"box="[345,787,1874,1900]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="lineatus">
<bibRefCitationid="F9454B1B7673A152FE4EF8937E737266"author="Gower, D. J. & Maduwage, K."box="[475,787,1874,1900]"pageId="5"pageNumber="337"pagination="51 - 68"refId="ref16404"refString="Gower, D. J. & Maduwage, K. (2011) Two new species of Rhinophis Hemprich (Serpentes: Uropeltidae) from Sri Lanka. Zootaxa, 2881 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2881.1.4"type="journal article"year="2011">Gower & Maduwage, 2011</bibRefCitation>
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(183–190 and up to 195, respectively), and has a darker ventral than dorsal surface (versus vice versa).
<taxonomicNameid="5AD44D697673A152FD39F8B77E10729A"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[684,880,1910,1936]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="5"pageNumber="337"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<bibRefCitationid="F9454B1B7673A152FB72F8B77818729A"author="Gower, D. J."box="[1255,1400,1910,1936]"pageId="5"pageNumber="337"pagination="495 - 510"refId="ref16321"refString="Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from southwestern Sri Lanka. Zootaxa, 4810 (3), 495 - 510. https: // doi. org / 10.11646 / zootaxa. 4810.3.6"type="journal article"year="2020">Gower, 2020</bibRefCitation>
</taxonomicName>
by having a more rounded than ridged dorsal apex to the rostral shield (in transverse section), having more ventrals (ca.
females versus 159–177), males typically having more subcaudals (at least 6 on one side to 7 [very rarely 5], versus 4–5 on each side), and having a darker venter than dorsum (versus darker dorsum than venter). The new species differs from
in having fewer ventrals (<187 versus>191: see
<bibRefCitationid="F9454B1B7670A151FE9FFF1E7CFA75F3"author="Gower, D. J."box="[266,410,223,249]"pageId="6"pageNumber="338"pagination="495 - 510"refId="ref16321"refString="Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from southwestern Sri Lanka. Zootaxa, 4810 (3), 495 - 510. https: // doi. org / 10.11646 / zootaxa. 4810.3.6"type="journal article"year="2020">Gower 2020</bibRefCitation>
).
<taxonomicNameid="5AD44D697670A151FE38FF1E7F1075F2"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[429,624,223,249]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D697670A151FCF1FF1E7F727417"authority="Wickramasinghe, Vidanapathirana, Wickramasinghe & Gower, 2020"authorityName="Wickramasinghe, Vidanapathirana, Wickramasinghe & Gower"authorityYear="2020"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="gunasekarai">
<bibRefCitationid="F9454B1B7670A151FB81FF1E7F727417"author="Wickramasinghe, L. M. & Vidanapathirana, D. R. & Wickramasinghe, N. & Gower, D. J."pageId="6"pageNumber="338"pagination="65 - 80"refId="ref17239"refString="Wickramasinghe, L. M., Vidanapathirana, D. R., Wickramasinghe, N. & Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Reptilia: Uropeltidae), from cloud forest of the Knuckles massif of Sri Lanka. Zootaxa, 4810 (1), 65 - 80. https: // doi. org / 10.11646 / zootaxa. 4810.1.3"type="journal article"year="2020">
Wickramasinghe, Vidanapathirana, Wickramasinghe & Gower,
in having a much broader tail shield relative to tail width, in lacking narrow, continuous longitudinal yellow lines on the dorsum, and in having pale markings on the underside of the tail.
(although with slightly higher ranges), and both have a relatively broad tail shield; however the two species differ clearly in colour pattern (paler dorsum than venter in the new species versus the reverse in
), in the shape of the tail shield (more prominent and conical in
<taxonomicNameid="5AD44D697670A151FD3DFE1A7E7474FE"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[680,788,475,500]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
), and in DNA sequence data (with phylogenetic analysis of these data providing maximal support for
<taxonomicNameid="5AD44D697670A151FD78FE3E7E377712"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[749,855,511,536]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D697670A151FBAAFDE279897736"authorityName="Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella"authorityYear="2009"box="[1087,1257,547,572]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="erangaviraji">
<taxonomicNameid="5AD44D697670A151FB63FDE2780E7737"authorityName="Gower & Maduwage"authorityYear="2011"box="[1270,1390,547,573]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="lineatus">
<taxonomicNameid="5AD44D697670A151FF02FD867D9F776A"authorityName="Gower & Maduwage"authorityYear="2011"box="[151,255,583,608]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="zigzag">
<bibRefCitationid="F9454B1B7670A151FE9BFD867C96776B"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."box="[270,502,583,609]"pageId="6"pageNumber="338"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
<taxonomicNameid="5AD44D697670A151FD9CFD867FAD776A"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[521,717,583,609]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<tableCitationid="D05603517670A151FE1DFDAA7CBF778F"box="[392,479,619,645]"captionStart="TABLE 3"captionStartId="8.[151,239,368,392]"captionTargetPageId="8"captionText="TABLE 3. Uncorrected p-distances (%) for intra- and interspecific comparisons for Rhinophis martin sp. nov. and closest sampled relatives, for two nuclear (prlr, cmos) and four mitochondrial genes (12s, 16s, cytb, nd4) as analysed by Sampaio et al. (2023). Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison."httpUri="http://table.plazi.org/id/C9AB6662767EA15FFF02FEB17ED474FE"pageId="6"pageNumber="338"tableUuid="C9AB6662767EA15FFF02FEB17ED474FE">Table 3</tableCitation>
<bibRefCitationid="F9454B1B7670A151FE7EFDAA7FAF778F"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."box="[491,719,619,645]"pageId="6"pageNumber="338"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of
<taxonomicNameid="5AD44D697670A151FAA5FD757DBE77FA"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="6"pageNumber="338"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<tableCitationid="D05603517670A151FDB5F9207F1573F6"box="[544,629,1761,1788]"captionStart="TABLE 1"captionStartId="5.[151,239,151,175]"captionTargetPageId="5"captionText="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded."httpUri="http://table.plazi.org/id/C9AB66627673A152FF02FF56793E7411"pageId="6"pageNumber="338"tableUuid="C9AB66627673A152FF02FF56793E7411">Table 1</tableCitation>
<figureCitationid="05EF2A6F7670A151FBC5F92079F373F6"box="[1104,1171,1761,1788]"captionStart="FIGURE 1"captionStartId="2.[151,250,1943,1967]"captionTargetBox="[228,1358,179,1916]"captionTargetId="figure-18@2.[228,1358,179,1916]"captionTargetPageId="2"captionText="FIGURE 1. Holotype of Rhinophis martin sp. nov., USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor."figureDoi="http://doi.org/10.5281/zenodo.11547471"httpUri="https://zenodo.org/record/11547471/files/figure.png"pageId="6"pageNumber="338">Fig. 1</figureCitation>
for photographs. Good condition; few superficial scratches to some scales. Head small, snout pointed. Rostral pointed, longer than wide, without dorsal crest; widest at level of anterior upper corner of first supralabials. Rostral many (>8) times longer (in dorsal view) than rostral-frontal gap. Frontal irregularly hexagonal, longer than wide, lateral (ocular) margins slightly converging posteriorly, posterolateral margins straight to very slightly concave; lateral (ocular) margin shortest, posterolateral edges longest. Frontal much shorter, wider than rostral. Paired nasals separated from each other by posterior half of rostral. External naris small, subcircular, slightly countersunk within small depression, located in anteroventral corner of nasal. Nasal contacts supralabials 1 and 2. Prefrontals only briefly in contact with each other along midline (left overlapping right), separating frontal from rostral. Prefrontals wider than long, shorter than frontal. Supralabials four, first smallest, making the least contribution to margin of mouth; fourth much the largest. Ocular contacts supralabials 3 and 4. Eye distinct, diameter approximately 0.3 times length of ocular, located near anterior corner of ocular, bulging slightly from ocular surface, pupil circular. Paired parietals longer than wide, approximately as long as but much wider than frontal, posteriorly broadly rounded, angle between posteromedial and posterolateral edges approximately 90°. Opposite parietals in brief midline contact (longer than contact between opposite prefrontals), left overlapping right. Each parietal contacts four scales other than head shields. No mental groove; mental approximately as wide as long, smaller than infralabials, contacting first infralabials but not first ventral; three pairs of infralabials, second largest, first smallest. First and second ventrals longer than wide, third approximately as long as wide, fourth and subsequent ventrals wider than long. Seven maxillary and eight mandibular teeth on each side. Teeth simple, pointed, distinctly retrorse, straight, evenly spaced.
Body cylindrical. Body scales generally evenly sized on dorsum and along body except for those involved in dorsal scale row reductions. Midline ventral scales between mental and anal of even size though anteriormost ones gradually narrow. At midbody, ventrals 1.24 times width of scales in first dorsal row. Ventrals 186. Dorsal scale rows 19 anteriorly, reducing to 17 by level with 41
<paragraphid="9D6B36EA7671A150FF52FC4A7E217007"blockId="7.[151,1437,907,2013]"pageId="7"pageNumber="339">Dorsal scale rows approximately 13 at base of tail. Head and body scales macroscopically smooth, lacking keels. Very faint, low, short keels towards posterior edges of small scales overlapped by anals and on lower dorsal scales of tail. Paired anal scales (right overlying left) considerably larger than posteriormost ventrals and subcaudals. Distal margin of each anal overlaps four (left) and three (right) other scales in addition to anteriormost subcaudals. Three subcaudals, last two undivided. Tail ‘shield’ broadly rounded cone, forming tip of tail, wider than long in dorsal view, many times longer than the frontal in dorsal view (a little shorter than distance between snout tip and back of frontal), visible from below and especially above, base surrounded by 11 scales (including last subcaudal). In posterior view shield circular to very slightly egg-shaped, wider ventrally than dorsally. Shield surface densely spinose, spines small and evenly sized, slightly posteriorly hooked (slightly dorsal upper margin, slightly concave posterior margin in lateral view) arranged in narrowly spaced rows approximately convergent on apex of shield. No notable ridges on scales of the tail and posterior end of body.</paragraph>
<emphasisid="AFA0EAF87671A150FF52FAD77CC1703B"bold="true"box="[199,417,1302,1329]"pageId="7"pageNumber="339">Colour in alcohol.</emphasis>
Tip of rostrum pale orange. Head shields otherwise generally brown with pale tan distal thin margins; a slightly larger, diffuse pale area across posterior of nasals and anterior of prefrontals. Lower edges of supralabials more irregularly and broadly pale. Three dorsally incomplete, narrow (one scale wide, distributed across two scales) whitish transverse bands behind head on dorsum, each band separated by length of four to five dorsal scales.
Ventral body colour and pattern more uniform than dorsal. Ventrally brown; paler (pale orange-tan) at head end. Slightly punctate, more so anteriorly, by virtue of each scale having a brown core and pale (mostly latero-) distal edges (the posterior distal edge typically almost transparent). Scale bases and centres mostly uniformly coloured, but flecked in the paler anterior region of the venter. Among ventrals, only 5
are entirely pale, as are a couple of adjacent lower dorsal scale row scales plus the mental and first infralabials. Further posteriorly, occasional ventrals with larger whitish patch; 7
dark brown, then abruptly and increasingly (coppery) orange-brown where scales are darkly flecked scales and have darker latero- and (especially) posterodistal edges. Midvertebral dorsal scale row with faintly indicated, narrow brown line each side. Darker, narrow, subparallel (4–5 scales apart) transverse dorsal bars (not dorsally complete; some asymmetrically arranged) taper upwards from dorsolateral apex of body, only one scale wide at dorsal end. Transition between darker more regular ventral colour and paler dorsal colour is somewhat irregular but typically lies on the 5
<paragraphid="9D6B36EA7671A15FFF52F85E7819744B"blockId="7.[151,1437,907,2013]"lastBlockId="8.[151,1437,151,321]"lastPageId="8"lastPageNumber="340"pageId="7"pageNumber="339">Subcaudals and first dorsal scale row mostly blackish with pale (yellowish) laterodistal and translucent posterior margins (more pale edges), together forming a posteriorly tapering, dark midventral tail stripe. Either side of this dark midventral region, the tail has a yellowish lateral tail stripe along its length from the vent to the tail shield, approximately 1.5 dorsal scale rows thick; anteriorly this stripe is much thicker, extending up onto the dorsum to form a transverse, middorsally incomplete pale band. Dark scales on upper and lower surface of tail darker than on body. Tail shield orange-yellow at apex and along ventral and ventrolateral outer edge, with almost symmetrical blackish blotches extending to dorsolateral outer edges; approximately half of shield surface orange-yellow, half blackish.</paragraph>
Uncorrected p-distances (%) for intra- and interspecific comparisons for
<taxonomicNameid="5AD44D69767EA15FFB9AFEB179A87482"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[1039,1224,368,392]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="8"pageNumber="340"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<bibRefCitationid="F9454B1B767EA15FFAAAFE557C7874DB"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."pageId="8"pageNumber="340"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
. Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison.
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<taxonomicNameid="5AD44D69767EA15FFF36FCAF7D837689"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[163,227,878,899]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="8"pageNumber="340"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D69767EA15FFF36FB917D83716F"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[163,227,1104,1125]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="8"pageNumber="340"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D69767EA15FFF36FB267D8371F6"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[163,227,1255,1276]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="8"pageNumber="340"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D69767EA15FFF36FA987D9D7064"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[163,253,1369,1390]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="8"pageNumber="340"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<tableCitationid="D0560351767EA15FFCA8FA037EF070D7"box="[829,912,1474,1501]"captionStart="TABLE 2"captionStartId="6.[151,239,692,716]"captionTargetPageId="6"captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded"httpUri="http://table.plazi.org/id/C9AB66627670A151FF02FD757CD5761E"pageId="8"pageNumber="340"tableUuid="C9AB66627670A151FF02FD757CD5761E">Table 2</tableCitation>
<figureCitationid="05EF2A6F767EA15FFF02FA277D86730B"box="[151,230,1510,1537]"captionStart="FIGURE 3"captionStartId="4.[151,250,1543,1567]"captionTargetBox="[170,1416,181,1520]"captionTargetId="figure-18@4.[170,1416,181,1520]"captionTargetPageId="4"captionText="FIGURE 3. Seven paratypes of Rhinophis martin sp. nov. Left column approximately dorsal view; right column approximately ventral view. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bars 20 mm. Photographs by H. Taylor."figureDoi="http://doi.org/10.5281/zenodo.11547476"httpUri="https://zenodo.org/record/11547476/files/figure.png"pageId="8"pageNumber="340">Figs. 3</figureCitation>
and
<figureCitationid="05EF2A6F767EA15FFEB4FA267C4F730B"box="[289,303,1511,1537]"captionStart="FIGURE 4"captionStartId="9.[151,250,1537,1561]"captionTargetBox="[151,1435,237,1513]"captionTargetId="figure-62@9.[151,1435,237,1513]"captionTargetPageId="9"captionText="FIGURE 4. Head and tail ends of seven paratypes of Rhinophis martin sp. nov. in four views. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bar increments in millimetres. Photographs by H. Taylor."figureDoi="http://doi.org/10.5281/zenodo.11547478"httpUri="https://zenodo.org/record/11547478/files/figure.png"pageId="8"pageNumber="340">4</figureCitation>
each have a small (temporal-like) scale separating the parietal from the fourth supralabial on each side, much smaller on the left in USNM 548100 (among uropeltids, temporals occur consistently only in
(USNM 548118, 548120) the first ventral does not contact the first infralabials. Scale row reductions simple (see Appendix 1), reducing from 19 to 17 dorsal scale rows by level with the 31
lack notable ridges on scales in the tail and posterior end of body; low ridges on upper posterior tail dorsal scale row scales in USNM 548117 and 548121, short low ridges on posterior ends of some tail dorsal scale row scales in 548122. Teeth not counted except in USNM 548122, which resembles the
<taxonomicNameid="5AD44D69767FA15EFD56F9C07E157313"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[707,885,1537,1561]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="9"pageNumber="341"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
lack pale patch across posterior of nasals and anterior of prefrontals; USNM 548100 with sinuous pale band across posterior of oculars and prefrontals and anterior of frontal. Whitish transverse bands on dorsum behind head typically shorter and/or more irregular and asymmetrical in
, forming spots or subtriangular patches rather than bands. One or two pale ventral scales and adjacent dorsal scale rows anteriorly (between ventral 3–5) in most
except USNM 548117, 548118 or 548122. Background body colour paler in USNM 548100, darker in 548123. Darker pair of paravertebral longitudinal lines particularly faint, diffuse and incomplete in USNM 548100, particularly clear in 548120. Dark dorsal cross bars similar to
dorsal scale row absent in USNM 548118 and 548122, forming notably larger, elongated pale patches with darker lower and upper rims in USNM 548123. First dorsal scale row on tail with blackish dots in USNM 548100, only blackish on a few scales in 548121, only blackish on anteriormost scales in 548120, not blackish in 548123. Pale lateral stripe on tail less regular and complete in
, absent in USNM 548122, thicker (two or more dorsal scale row scales thick) in 548120 and 548123, restricted to posterior end of tail in 548100 and 548118. Dorsolateral yellow cross bars on anterior of tail absent in USNM 548118, short and broadly incomplete in 548122, narrow and almost middorsally complete in 548117. Shield more orange-yellow in USNM 548121, mostly dark in 548118 and 548122 and especially in 548117 and
Graph showing relationship between number of ventral scales and number of subcaudal scales (mean value for left and right) in female (triangles) and male (circles)
<taxonomicNameid="5AD44D69767CA15DFD05FA0D7E2170EE"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[656,833,1484,1508]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="10"pageNumber="342"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D69767CA15DFE8CF80C7CAA72EF"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[281,458,1997,2021]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="10"pageNumber="342"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<tableCitationid="D0560351767DA15CFBE6FF5679AA75BB"box="[1139,1226,151,177]"captionStart="TABLE 2"captionStartId="6.[151,239,692,716]"captionTargetPageId="6"captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded"httpUri="http://table.plazi.org/id/C9AB66627670A151FF02FD757CD5761E"pageId="11"pageNumber="343"tableUuid="C9AB66627670A151FF02FD757CD5761E">Table 2</tableCitation>
(USNM 548106, 548113) the rostral contacts the frontal and prevents midline contact between opposite prefrontals. Left parietal overlaps right in all except USNM 548109; where prefrontals overlap, left above right in all except USNM 548096, 548097, 548103, 548107, and 548111. Temporal-like scale on each side in USNM 548096, on right in USNM 548101. Frontal partly fused with right prefrontal in USNM 548098. All have 3,3 infralabials except USNM 548101 (3,4). First and second ventrals in contact in all except USNM 548098 and 548111). Right anal shield overlies left in all specimens except female specimens USNM 548103, 548111, and 548114. Scales around shield 10–13. Ventrals in males 169–178, females 179–186; subcaudals in males 5–7 per side, females typically 3–4 per side (two on one side on USNM 548107; five on one side in USNM 548115). Most referred specimens lack notable ridges on scales in the tail and posterior end of body; if present then typically low and on upper posterior dorsal scale row scales on tail, also on posteriormost subcaudals in, for example, USNM 548097, 548098, and 548114.
Variation in the number of ventrals, subcaudals and in relative tail length is strongly bimodal across the sample as a whole (
<figureCitationid="05EF2A6F767DA15CFDD9FD867FF0776B"box="[588,656,582,609]"captionStart="FIGURE 5"captionStartId="10.[151,250,1448,1472]"captionTargetBox="[152,1398,533,1406]"captionTargetId="figure-240@10.[151,1435,532,1412]"captionTargetPageId="10"captionText="FIGURE 5. Graph showing relationship between number of ventral scales and number of subcaudal scales (mean value for left and right) in female (triangles) and male (circles) Rhinophis martin sp. nov."figureDoi="http://doi.org/10.5281/zenodo.11547480"httpUri="https://zenodo.org/record/11547480/files/figure.png"pageId="11"pageNumber="343">Fig. 5</figureCitation>
<emphasisid="AFA0EAF8767DA15CFF52FDAB7C0B778E"bold="true"box="[199,363,618,644]"pageId="11"pageNumber="343">Colour in life.</emphasis>
In life (
<figureCitationid="05EF2A6F767DA15CFE57FDAA7F65778F"box="[450,517,619,645]"captionStart="FIGURE 6"captionStartId="10.[151,250,1997,2021]"captionTargetBox="[151,1435,1533,1974]"captionTargetId="figure-217@10.[151,1435,1533,1974]"captionTargetPageId="10"captionText="FIGURE 6. Rhinophis martin sp. nov., paratype DWC 2024.05.01 prior to fixation, in upper (left) and lower (right) views."figureDoi="http://doi.org/10.5281/zenodo.11547482"httpUri="https://zenodo.org/record/11547482/files/figure.png"pageId="11"pageNumber="343">Fig. 6</figureCitation>
), the dorsal areas that are orange-brown in preserved specimens are golden yellow, and the brownish ventral colour seen in preservation is more blackish. The off-white markings on the lateral and ventral surfaces of preserved specimens are whitish in life. The pale yellowish white markings on the lateral surface of the tail of preserved specimens are similarly coloured in life. One uncollected, small specimen (<
Named in honour of the late Martin Wijesinghe (1939–2021), in recognition of his conservation and education actions and advocacy, especially with respect to forest biota and ecosystems of southwestern
<emphasisid="AFA0EAF8767DA15CFF52FC377E64711A"bold="true"box="[199,772,1014,1041]"pageId="11"pageNumber="343">Distribution, natural history, and conservation.</emphasis>
As far as is known,
<taxonomicNameid="5AD44D69767DA15CFC61FC367900711A"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[1012,1120,1015,1040]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<figureCitationid="05EF2A6F767DA15CFB46FBDA7878713F"box="[1235,1304,1051,1077]"captionStart="FIGURE 2"captionStartId="3.[151,250,1911,1935]"captionTargetBox="[273,1315,185,1881]"captionTargetId="figure-31@3.[266,1321,181,1887]"captionTargetPageId="3"captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)."figureDoi="http://doi.org/10.5281/zenodo.11547474"httpUri="https://zenodo.org/record/11547474/files/figure.png"pageId="11"pageNumber="343">Fig. 2</figureCitation>
). The upper elevational limit based on known specimens is not clear because of imprecise locality data, but is less than
and USNM referred specimens were collected more than 40 years ago. The available field collection data do not indicate whether these USNM specimens were found in forest or in agricultural habitats, both of which occur in the general area today. The more-recently collected
specimens from Deniyaya were found when unearthed by mechanical diggers clearing an area of tea plantation for a car park. The new species is likely to qualify for Data Deficient status under IUCN Red List criteria (
<bibRefCitationid="F9454B1B767DA15CFF0AFAD67F1B703B"author="IUCN Species Survival Commission & SC"box="[159,635,1303,1329]"pageId="11"pageNumber="343"refId="ref16624"refString="IUCN Species Survival Commission (SC) (2012) IUCN Red List categories and criteria. Version 3.1. 2 Edition. IUCN, Gland and Cambridge, 32 pp."type="book"year="2012">IUCN Species Survival Commission 2012</bibRefCitation>
), at least until new field surveys are undertaken to better clarify its range and any potential threats, and/or until additional specimens can be found in other collections.
<taxonomicNameid="5AD44D69767DA15CFE44FA427F5A7096"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[465,570,1411,1436]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<figureCitationid="05EF2A6F767DA15CFB76FA0A784670EF"box="[1251,1318,1483,1509]"captionStart="FIGURE 2"captionStartId="3.[151,250,1911,1935]"captionTargetBox="[273,1315,185,1881]"captionTargetId="figure-31@3.[266,1321,181,1887]"captionTargetPageId="3"captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)."figureDoi="http://doi.org/10.5281/zenodo.11547474"httpUri="https://zenodo.org/record/11547474/files/figure.png"pageId="11"pageNumber="343">Fig. 2</figureCitation>
. However, detailed comparisons have not yet been made, and the identities of those populations remain as yet unclear.
<bibRefCitationid="F9454B1B767DA15CFED1F9D27F5B7327"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."box="[324,571,1555,1581]"pageId="11"pageNumber="343"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
<taxonomicNameid="5AD44D69767DA15CFBE7F9D279BE7326"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[1138,1246,1555,1580]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
(Deniyaya)”. There are no morphological vouchers for those specimens; these sequences were obtained from tissues of parts of three different and partial animals that were found alongside the three complete
<taxonomicNameid="5AD44D69767DA15CFEF0F9627CAF73B6"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[357,463,1699,1724]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D69767DA15CFDC2F92A7FA1720E"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[599,705,1771,1796]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<bibRefCitationid="F9454B1B767DA15CFC82F8CE7ED77223"author="Gower, D. J."box="[791,951,1807,1833]"pageId="11"pageNumber="343"pagination="495 - 510"refId="ref16321"refString="Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from southwestern Sri Lanka. Zootaxa, 4810 (3), 495 - 510. https: // doi. org / 10.11646 / zootaxa. 4810.3.6"type="journal article"year="2020">Gower (2020)</bibRefCitation>
is from Rakwana (not far from the known localities of the new species:
<figureCitationid="05EF2A6F767DA15CFE4CF8F27F7A7247"box="[473,538,1843,1869]"captionStart="FIGURE 2"captionStartId="3.[151,250,1911,1935]"captionTargetBox="[273,1315,185,1881]"captionTargetId="figure-31@3.[266,1321,181,1887]"captionTargetPageId="3"captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)."figureDoi="http://doi.org/10.5281/zenodo.11547474"httpUri="https://zenodo.org/record/11547474/files/figure.png"pageId="11"pageNumber="343">Fig. 2</figureCitation>
) but differs from
<taxonomicNameid="5AD44D69767DA15CFD49F8F27E257246"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[732,837,1843,1868]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<taxonomicNameid="5AD44D69767DA15CFC75F896792A727A"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[992,1098,1879,1904]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
<bibRefCitationid="F9454B1B767DA15CFAAFF8967C40729F"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."pageId="11"pageNumber="343"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
<tableCitationid="D0560351767DA15CFAC8F8BA7DC772B3"captionStart="TABLE 3"captionStartId="8.[151,239,368,392]"captionTargetPageId="8"captionText="TABLE 3. Uncorrected p-distances (%) for intra- and interspecific comparisons for Rhinophis martin sp. nov. and closest sampled relatives, for two nuclear (prlr, cmos) and four mitochondrial genes (12s, 16s, cytb, nd4) as analysed by Sampaio et al. (2023). Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison."httpUri="http://table.plazi.org/id/C9AB6662767EA15FFF02FEB17ED474FE"pageId="11"pageNumber="343"tableUuid="C9AB6662767EA15FFF02FEB17ED474FE">Table 3</tableCitation>
<taxonomicNameid="5AD44D69767DA15CFE4CF85E7F2272B2"authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024"authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe"authorityYear="2024"box="[473,578,1951,1976]"class="Squamata"family="Uropeltidae"genus="Rhinophis"kingdom="Animalia"pageId="11"pageNumber="343"phylum="Chordata"rank="species"species="martin"status="sp. nov.">
were inferred to be distinct from all other sampled taxa at the species level in all of the species delimitation analyses reported by
<bibRefCitationid="F9454B1B767DA15CFCBAF802794572D7"author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J."box="[815,1061,1987,2013]"pageId="11"pageNumber="343"pagination="107651"refId="ref16965"refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651"type="journal article"year="2023">
