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<mods:title id="F429D5EAA8FB2E005D309BA4379BDB6C">Two new species of the shieldtail snake genus Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae), from the Rakwana and the Knuckles Massifs of Sri Lanka</mods:title>
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<mods:namePart id="FDC67EFE1C002CD43884D05266ADFBAB">Gower, David J.</mods:namePart>
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<mods:affiliation id="1A4FB01B4C5EE9F4822936E8D2099FFE">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="4F34BA035F8ED92308A30CA9C7171739">Sampaio, Filipa L.</mods:namePart>
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<mods:affiliation id="7C44FE8F1DF73273C73981E52AB623A2">Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK & Department of Genetics, Evolution and Environment, University College London, London WC 1 E 6 BT, UK</mods:affiliation>
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<mods:namePart id="FBDF9049F6ED0A259CF4F43839C1FCC0">Vidanapathirana, Dulan Ranga</mods:namePart>
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<mods:affiliation id="B05B793B7A8F0AD15795D9AF75A26F1E">Herpetological Foundation of Sri Lanka, 31 / 5, Alwis Town, Hendala, Wattala, Sri Lanka</mods:affiliation>
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<mods:namePart id="A018E9D04DDEC289DE409D06F2F054B2">Mendis Wickramasinghe, L. J.</mods:namePart>
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<mods:affiliation id="30B7E1EC4A1815F13D1E9DDBBE27E6E2">Herpetological Foundation of Sri Lanka, 31 / 5, Alwis Town, Hendala, Wattala, Sri Lanka</mods:affiliation>
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<mods:title id="0F15636A13356431E275AF88F9F42553">Zootaxa</mods:title>
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<mods:part id="FEBF074D8B23A833A6374DB8ECA9A7AF">
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<mods:date id="09EEAFC1F2F455A479E8A7087A55AA8E">2024</mods:date>
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<mods:number id="98FCC46CF3F67243383FDC5EAC1AF6CB">2024-05-29</mods:number>
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<mods:number id="CA0C607F7B31597A348495208E80403D">5458</mods:number>
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<mods:number id="57154D2C08E2E9810034A2DF30D98C00">3</mods:number>
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<mods:url id="69F983535923F69FE70D91E90A7B82A2">http://dx.doi.org/10.11646/zootaxa.5458.3.2</mods:url>
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<mods:classification id="A8A67DC528B8743CE7C860B05AC542DC">journal article</mods:classification>
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<mods:identifier id="CA3B19DB98366A9F37A8163895D3BC60" type="DOI">10.11646/zootaxa.5458.3.2</mods:identifier>
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<mods:identifier id="ABAD48B007CB9ECDFB9952C164E65EF6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1D8F7AF568CE4A4788426EB92DACB2D2" type="Zenodo-Dep">11547469</mods:identifier>
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<mods:identifier id="89998D9DCB8B3021248DCDF575BE1DB8" type="ZooBank">54609B08-07DE-4009-B15E-FEBE2C577AAE</mods:identifier>
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<subSubSection id="D5CE65617677A156FF02FB787CB271D9" box="[151,466,1209,1236]" pageId="1" pageNumber="333" type="nomenclature">
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<paragraph id="9D6B36EA7677A156FF02FB787CB271D9" blockId="1.[151,536,1209,1271]" box="[151,466,1209,1236]" pageId="1" pageNumber="333">
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<heading id="C62381867677A156FF02FB787CB271D9" bold="true" box="[151,466,1209,1236]" fontSize="11" level="1" pageId="1" pageNumber="333" reason="1">
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<emphasis id="AFA0EAF87677A156FF02FB787CB271D9" bold="true" box="[151,466,1209,1236]" pageId="1" pageNumber="333">
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<taxonomicName id="5AD44D697677A156FF02FB787C0E71D9" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[151,366,1209,1236]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="1" pageNumber="333" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
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<emphasis id="AFA0EAF87677A156FF02FB787C0E71D9" bold="true" box="[151,366,1209,1236]" italics="true" pageId="1" pageNumber="333">Rhinophis martin</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B49357837677A156FEE0FB787CB271D9" box="[373,466,1209,1235]" pageId="1" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="D5CE65617677A156FF02FB1C7EEB703E" pageId="1" pageNumber="333" type="description">
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<paragraph id="9D6B36EA7677A156FF02FB1C7F7871FD" blockId="1.[151,536,1209,1271]" box="[151,536,1245,1271]" pageId="1" pageNumber="333">
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<figureCitation id="05EF2A6F7677A156FF02FB1C7C6271FD" box="[151,258,1245,1271]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1943,1967]" captionStartId-1="3.[151,250,1911,1935]" captionStartId-2="4.[151,250,1543,1567]" captionStartId-3="9.[151,250,1537,1561]" captionStartId-4="10.[151,250,1448,1472]" captionStartId-5="10.[151,250,1997,2021]" captionTargetBox-0="[228,1358,179,1916]" captionTargetBox-1="[273,1315,185,1881]" captionTargetBox-2="[170,1416,181,1520]" captionTargetBox-3="[151,1435,237,1513]" captionTargetBox-4="[152,1398,533,1406]" captionTargetBox-5="[151,1435,1533,1974]" captionTargetId-0="figure-18@2.[228,1358,179,1916]" captionTargetId-1="figure-31@3.[266,1321,181,1887]" captionTargetId-2="figure-18@4.[170,1416,181,1520]" captionTargetId-3="figure-62@9.[151,1435,237,1513]" captionTargetId-4="figure-240@10.[151,1435,532,1412]" captionTargetId-5="figure-217@10.[151,1435,1533,1974]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="10" captionText-0="FIGURE 1. Holotype of Rhinophis martin sp. nov., USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor." captionText-1="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)." captionText-2="FIGURE 3. Seven paratypes of Rhinophis martin sp. nov. Left column approximately dorsal view; right column approximately ventral view. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bars 20 mm. Photographs by H. Taylor." captionText-3="FIGURE 4. Head and tail ends of seven paratypes of Rhinophis martin sp. nov. in four views. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bar increments in millimetres. Photographs by H. Taylor." captionText-4="FIGURE 5. Graph showing relationship between number of ventral scales and number of subcaudal scales (mean value for left and right) in female (triangles) and male (circles) Rhinophis martin sp. nov." captionText-5="FIGURE 6. Rhinophis martin sp. nov., paratype DWC 2024.05.01 prior to fixation, in upper (left) and lower (right) views." figureDoi-0="http://doi.org/10.5281/zenodo.11547471" figureDoi-1="http://doi.org/10.5281/zenodo.11547474" figureDoi-2="http://doi.org/10.5281/zenodo.11547476" figureDoi-3="http://doi.org/10.5281/zenodo.11547478" figureDoi-4="http://doi.org/10.5281/zenodo.11547480" figureDoi-5="http://doi.org/10.5281/zenodo.11547482" httpUri-0="https://zenodo.org/record/11547471/files/figure.png" httpUri-1="https://zenodo.org/record/11547474/files/figure.png" httpUri-2="https://zenodo.org/record/11547476/files/figure.png" httpUri-3="https://zenodo.org/record/11547478/files/figure.png" httpUri-4="https://zenodo.org/record/11547480/files/figure.png" httpUri-5="https://zenodo.org/record/11547482/files/figure.png" pageId="1" pageNumber="333">Figs. 1–6</figureCitation>
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;
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<tableCitation id="D05603517677A156FE9BFB1C7CEA71FD" box="[270,394,1245,1271]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="5.[151,239,151,175]" captionStartId-1="6.[151,239,692,716]" captionStartId-2="8.[151,239,368,392]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded." captionText-1="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded" captionText-2="TABLE 3. Uncorrected p-distances (%) for intra- and interspecific comparisons for Rhinophis martin sp. nov. and closest sampled relatives, for two nuclear (prlr, cmos) and four mitochondrial genes (12s, 16s, cytb, nd4) as analysed by Sampaio et al. (2023). Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison." pageId="1" pageNumber="333">Tables 1–3</tableCitation>
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; Appendix 1
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</paragraph>
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<paragraph id="9D6B36EA7677A156FF02FADD7EEB703E" blockId="1.[151,907,1308,1332]" box="[151,907,1308,1332]" pageId="1" pageNumber="333">
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urn:lsid:zoobank.org:act:
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<uuid id="E9720C3F7677A156FE01FADD7EEB703E" box="[404,907,1308,1332]" pageId="1" pageNumber="333">6DA63C7E-C8D3-4318-83C4-18B1F5D7ACDC</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="D5CE65617677A156FF02FAA37FD670B5" box="[151,694,1378,1472]" pageId="1" pageNumber="333" type="reference_group">
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<paragraph id="9D6B36EA7677A156FF02FAA37C567076" blockId="1.[151,310,1378,1404]" box="[151,310,1378,1404]" pageId="1" pageNumber="333">
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<heading id="C62381867677A156FF02FAA37C567076" bold="true" box="[151,310,1378,1404]" fontSize="11" level="1" pageId="1" pageNumber="333" reason="1">
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<emphasis id="AFA0EAF87677A156FF02FAA37C567076" bold="true" box="[151,310,1378,1404]" pageId="1" pageNumber="333">Chresonymy</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9D6B36EA7677A156FF02FA647FD670B5" blockId="1.[151,694,1445,1472]" box="[151,694,1445,1472]" pageId="1" pageNumber="333">
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<taxonomicName id="5AD44D697677A156FF02FA647C5070CA" box="[151,304,1445,1472]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="1" pageNumber="333" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="AFA0EAF87677A156FF02FA647C6970B5" box="[151,265,1445,1471]" italics="true" pageId="1" pageNumber="333">Rhinophis</emphasis>
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sp.
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</taxonomicName>
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(Deniyaya):
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<bibRefCitation id="F9454B1B7677A156FE56FA647FD670B5" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." box="[451,694,1445,1471]" pageId="1" pageNumber="333" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
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Sampaio
|
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<emphasis id="AFA0EAF87677A156FDB9FA677F0570B5" box="[556,613,1445,1471]" italics="true" pageId="1" pageNumber="333">et al.</emphasis>
|
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(2023)
|
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</bibRefCitation>
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</paragraph>
|
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</subSubSection>
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<subSubSection id="D5CE65617677A156FF02FA297FDA72F0" pageId="1" pageNumber="333" type="materials_examined">
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<paragraph id="9D6B36EA7677A156FF02FA29796D7364" blockId="1.[151,1437,1512,2043]" pageId="1" pageNumber="333">
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<materialsCitation id="2DBC3CB77677A156FF02FA29796D7364" collectingDate="1979-10-24" collectionCode="USNM" collectorName="Lalith Jayawickrama" country="Sri Lanka" elevation="820" latitude="6.37" location="Hayes" longLatPrecision="783" longitude="80.65" municipality="Hayes" pageId="1" pageNumber="333" specimenCode="USNM 548119" specimenCount="1" specimenCount-female="1" stateProvince="Sabaragamuwa Province" typeStatus="holotype">
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<emphasis id="AFA0EAF87677A156FF02FA297C6B7308" bold="true" box="[151,267,1512,1538]" pageId="1" pageNumber="333">
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<typeStatus id="426F88487677A156FF02FA297C667308" box="[151,262,1512,1538]" pageId="1" pageNumber="333" type="holotype">Holotype</typeStatus>
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.
|
|||
|
|
</emphasis>
|
|||
|
|
<specimenCode id="CD729E917677A156FE86FA297CA07309" box="[275,448,1512,1539]" pageId="1" pageNumber="333">USNM 548119</specimenCode>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="05EF2A6F7677A156FE45FA297F747308" box="[464,532,1512,1538]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[228,1358,179,1916]" captionTargetId="figure-18@2.[228,1358,179,1916]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Rhinophis martin sp. nov., USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor." figureDoi="http://doi.org/10.5281/zenodo.11547471" httpUri="https://zenodo.org/record/11547471/files/figure.png" pageId="1" pageNumber="333">Fig. 1</figureCitation>
|
|||
|
|
,
|
|||
|
|
<tableCitation id="D05603517677A156FDB6FA297F167308" box="[547,630,1512,1538]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded." pageId="1" pageNumber="333">Table 1</tableCitation>
|
|||
|
|
), adult
|
|||
|
|
<specimenCount id="8BD2FD637677A156FD45FA297E7F7308" box="[720,799,1512,1538]" pageId="1" pageNumber="333" type="female">female</specimenCount>
|
|||
|
|
, “Hayes/Gongala”,
|
|||
|
|
<collectingRegion id="5F10F8087677A156FB95FA2978797308" box="[1024,1305,1512,1538]" country="Sri Lanka" name="Sabaragamuwa" pageId="1" pageNumber="333">Sabaragamuwa Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="E5C3767A7677A156FAB3FA2978F97308" box="[1318,1433,1512,1538]" name="Sri Lanka" pageId="1" pageNumber="333">Sri Lanka</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<collectingMunicipality id="7D0FAC907677A156FF02F9CC7DBD732D" box="[151,221,1549,1575]" pageId="1" pageNumber="333">Hayes</collectingMunicipality>
|
|||
|
|
(
|
|||
|
|
<geoCoordinate id="F8E0502D7677A156FF7EF9CD7C26732D" box="[235,326,1548,1575]" degrees="6.37" direction="north" orientation="latitude" pageId="1" pageNumber="333" precision="555" value="6.37">6.37° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D7677A156FEC5F9CD7CD4732D" box="[336,436,1548,1575]" degrees="80.65" direction="east" orientation="longitude" pageId="1" pageNumber="333" precision="555" value="80.65">80.65° E</geoCoordinate>
|
|||
|
|
; ca.
|
|||
|
|
<quantity id="5A2C9B0F7677A156FE71F9CD7F4B732C" box="[484,555,1548,1574]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="333" unit="m" value="820.0">
|
|||
|
|
<elevation id="16F9D1D97677A156FE71F9CD7F4B732C" box="[484,555,1548,1574]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="1" pageNumber="333" unit="m" value="820.0">820 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
) is a tea estate and Gongala is a nearby mountain (peaking at 6.386° N, 80.654° E; 1,358 m elevation), <2 km to the north-northwest on the eastern edge of Sinharaja Forest Reserve, part of the Rakwana Massif (
|
|||
|
|
<figureCitation id="05EF2A6F7677A156FEF7F9957CC77364" box="[354,423,1620,1646]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1935]" captionTargetBox="[273,1315,185,1881]" captionTargetId="figure-31@3.[266,1321,181,1887]" captionTargetPageId="3" captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)." figureDoi="http://doi.org/10.5281/zenodo.11547474" httpUri="https://zenodo.org/record/11547474/files/figure.png" pageId="1" pageNumber="333">Fig. 2</figureCitation>
|
|||
|
|
). Collected by
|
|||
|
|
<collectorName id="3021533C7677A156FDC4F9957E5E7364" box="[593,830,1620,1646]" pageId="1" pageNumber="333">Lalith Jayawickrama</collectorName>
|
|||
|
|
,
|
|||
|
|
<date id="E96A102A7677A156FCDCF99579687364" box="[841,1032,1620,1646]" pageId="1" pageNumber="333" value="1979-10-24">
|
|||
|
|
<collectingDate id="F92EE9C27677A156FCDCF99579687364" box="[841,1032,1620,1646]" pageId="1" pageNumber="333" value="1979-10-24">24 October 1979</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7677A156FF52F9B979CE724C" blockId="1.[151,1437,1512,2043]" pageId="1" pageNumber="333">
|
|||
|
|
<emphasis id="AFA0EAF87677A156FF52F9B97C5F7398" bold="true" box="[199,319,1656,1682]" pageId="1" pageNumber="333">
|
|||
|
|
<typeStatus id="426F88487677A156FF52F9B97C5F7398" box="[199,319,1656,1682]" pageId="1" pageNumber="333" type="paratype">Paratypes</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
(n = 10;
|
|||
|
|
<tableCitation id="D05603517677A156FE33F9B97C987398" box="[422,504,1656,1682]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded." pageId="1" pageNumber="333">Table 1</tableCitation>
|
|||
|
|
). USNM 548100 (female) collected “Hayes”,
|
|||
|
|
<date id="E96A102A7677A156FB97F9B979D77398" box="[1026,1207,1656,1682]" pageId="1" pageNumber="333" value="1977-08-10">10 August 1977</date>
|
|||
|
|
. USNM 548117 and 548118 (males) collected “Hayes/Gongala”,
|
|||
|
|
<date id="E96A102A7677A156FD1AF95D7E3073BC" box="[655,848,1692,1718]" pageId="1" pageNumber="333" value="1979-10-24">24 October 1979</date>
|
|||
|
|
. USNM 548120 (female), 548121 (female), 548122 (male), and 548123 (male) collected “Hayes/Gongala”,
|
|||
|
|
<date id="E96A102A7677A156FCB5F9017E8773D0" box="[800,999,1728,1754]" pageId="1" pageNumber="333" value="1979-10-26">26 October 1979</date>
|
|||
|
|
. All seven of these USNM
|
|||
|
|
<typeStatus id="426F88487677A156FAA4F90078FC73D1" box="[1329,1436,1729,1755]" pageId="1" pageNumber="333" type="paratype">paratypes</typeStatus>
|
|||
|
|
were collected by Lalith Jayawickrama. NMSL-NH
|
|||
|
|
<date id="E96A102A7677A156FD63F9257E1473F4" box="[758,884,1764,1790]" pageId="1" pageNumber="333" value="2024-01-01">2024.01.01</date>
|
|||
|
|
(female) and NMSL-NH
|
|||
|
|
<date id="E96A102A7677A156FB35F925787E73F4" box="[1184,1310,1764,1790]" pageId="1" pageNumber="333" value="2024-01-02">2024.01.02</date>
|
|||
|
|
and DWC
|
|||
|
|
<date id="E96A102A7677A156FF02F8C97C757229" box="[151,277,1800,1827]" pageId="1" pageNumber="333" value="2024-05-01">2024.05.01</date>
|
|||
|
|
(males) collected from Burusgala, near Deniyaya,
|
|||
|
|
<collectingRegion id="5F10F8087677A156FCDBF8C9797F7228" box="[846,1055,1800,1826]" country="Sri Lanka" name="Southern" pageId="1" pageNumber="333">Southern Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="E5C3767A7677A156FBBFF8C979F87228" box="[1066,1176,1800,1826]" name="Sri Lanka" pageId="1" pageNumber="333">Sri Lanka</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<geoCoordinate id="F8E0502D7677A156FB33F8C978617229" box="[1190,1281,1800,1827]" degrees="6.38" direction="north" orientation="latitude" pageId="1" pageNumber="333" precision="555" value="6.38">6.38° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D7677A156FA99F8C978117229" box="[1292,1393,1800,1827]" degrees="80.60" direction="east" orientation="longitude" pageId="1" pageNumber="333" precision="555" value="80.6">80.60° E</geoCoordinate>
|
|||
|
|
; ca.
|
|||
|
|
<quantity id="5A2C9B0F7677A156FF02F8ED7DBF724C" box="[151,223,1836,1863]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="333" unit="m" value="750.0">750 m</quantity>
|
|||
|
|
) by Dulan Ranga Vidanapathirana and L. J. Mendis Wickramasinghe,
|
|||
|
|
<date id="E96A102A7677A156FC63F8ED79CB724C" box="[1014,1195,1836,1862]" pageId="1" pageNumber="333" value="2018-08-22">22 August 2018</date>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7677A156FF52F8917FDA72F0" blockId="1.[151,1437,1512,2043]" pageId="1" pageNumber="333">
|
|||
|
|
<emphasis id="AFA0EAF87677A156FF52F8917CD07260" bold="true" box="[199,432,1872,1898]" pageId="1" pageNumber="333">Referred specimens</emphasis>
|
|||
|
|
(n = 22;
|
|||
|
|
<tableCitation id="D05603517677A156FD85F8917F027260" box="[528,610,1872,1898]" captionStart="TABLE 2" captionStartId="6.[151,239,692,716]" captionTargetPageId="6" captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded" pageId="1" pageNumber="333">Table 2</tableCitation>
|
|||
|
|
).
|
|||
|
|
<specimenCount id="8BD2FD637677A156FDE3F8917F9C7260" box="[630,764,1872,1898]" count="8" pageId="1" pageNumber="333" type="male">Eight males</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="8BD2FD637677A156FC93F8917EE07260" box="[774,896,1872,1898]" count="11" pageId="1" pageNumber="333" type="female">11 females</specimenCount>
|
|||
|
|
, and three incomplete specimens for which sex is less confidently inferred (
|
|||
|
|
<tableCitation id="D05603517677A156FE23F8B57F6E7284" box="[438,526,1908,1934]" captionStart="TABLE 2" captionStartId="6.[151,239,692,716]" captionTargetPageId="6" captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded" pageId="1" pageNumber="333">Table 2</tableCitation>
|
|||
|
|
): USNM 548096–548099 and 548101–548110 collected by Lalith Jayawickrama from Hayes 14 July–18 August, 1977; USNM 548111–548116, 548124, 548125 collected by Lalith Jayawickrama from Hayes/Gongala 24–26 October, 1979. These are considered referred rather than
|
|||
|
|
<typeStatus id="426F88487677A156FBF9F87C79FC72DD" box="[1132,1180,1981,2007]" pageId="1" pageNumber="333">type</typeStatus>
|
|||
|
|
material because they were examined in less detail than the
|
|||
|
|
<typeStatus id="426F88487677A156FDA9F8207F0C72F1" box="[572,620,2017,2043]" pageId="1" pageNumber="333">type</typeStatus>
|
|||
|
|
series.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="C9AB66627674A155FF02F85679FB72FD" ID-DOI="http://doi.org/10.5281/zenodo.11547471" ID-Zenodo-Dep="11547471" httpUri="https://zenodo.org/record/11547471/files/figure.png" pageId="2" pageNumber="334" startId="2.[151,250,1943,1967]" targetBox="[228,1358,179,1916]" targetPageId="2" targetType="figure">
|
|||
|
|
<paragraph id="9D6B36EA7674A155FF02F85679FB72FD" blockId="2.[151,1437,1943,2039]" pageId="2" pageNumber="334">
|
|||
|
|
<emphasis id="AFA0EAF87674A155FF02F8567C7272A5" bold="true" box="[151,274,1943,1967]" pageId="2" pageNumber="334">FIGURE 1.</emphasis>
|
|||
|
|
Holotype of
|
|||
|
|
<taxonomicName id="5AD44D697674A155FE00F8567F2572A5" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[405,581,1943,1967]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="2" pageNumber="334" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87674A155FE00F8567F2572A5" box="[405,581,1943,1967]" italics="true" pageId="2" pageNumber="334">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87674A155FDDFF8567FFA72A5" bold="true" box="[586,666,1943,1967]" pageId="2" pageNumber="334">
|
|||
|
|
<taxonomicNameLabel id="B49357837674A155FDDFF8567FFA72A5" box="[586,666,1943,1967]" pageId="2" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="C9AB66627675A154FF02F8B67E3E72F1" ID-DOI="http://doi.org/10.5281/zenodo.11547474" ID-Zenodo-Dep="11547474" httpUri="https://zenodo.org/record/11547474/files/figure.png" pageId="3" pageNumber="335" startId="3.[151,250,1911,1935]" targetBox="[273,1315,185,1881]" targetPageId="3" targetType="figure">
|
|||
|
|
<paragraph id="9D6B36EA7675A154FF02F8B67E3E72F1" blockId="3.[151,1436,1911,2043]" pageId="3" pageNumber="335">
|
|||
|
|
<emphasis id="AFA0EAF87675A154FF02F8B67C727285" bold="true" box="[151,274,1911,1935]" pageId="3" pageNumber="335">FIGURE 2.</emphasis>
|
|||
|
|
Map showing places relevant to the distribution of
|
|||
|
|
<taxonomicName id="5AD44D697675A154FC81F8B67EA47285" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[788,964,1911,1935]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87675A154FC81F8B67EA47285" box="[788,964,1911,1935]" italics="true" pageId="3" pageNumber="335">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87675A154FC5CF8B679787285" bold="true" box="[969,1048,1911,1935]" pageId="3" pageNumber="335">
|
|||
|
|
<taxonomicNameLabel id="B49357837675A154FC5CF8B679787285" box="[969,1048,1911,1935]" pageId="3" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for
|
|||
|
|
<taxonomicName id="5AD44D697675A154FB7CF85D782872B9" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[1257,1352,1948,1971]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="3" pageNumber="335" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87675A154FB7CF85D782872B9" box="[1257,1352,1948,1971]" italics="true" pageId="3" pageNumber="335">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87675A154FAD9F85A78FC72B9" bold="true" box="[1356,1436,1947,1971]" pageId="3" pageNumber="335">
|
|||
|
|
<taxonomicNameLabel id="B49357837675A154FAD9F85A78FC72B9" box="[1356,1436,1947,1971]" pageId="3" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="C9AB66627672A153FF02F9C67EE57362" ID-DOI="http://doi.org/10.5281/zenodo.11547476" ID-Zenodo-Dep="11547476" httpUri="https://zenodo.org/record/11547476/files/figure.png" pageId="4" pageNumber="336" startId="4.[151,250,1543,1567]" targetBox="[170,1416,181,1520]" targetPageId="4" targetType="figure">
|
|||
|
|
<paragraph id="9D6B36EA7672A153FF02F9C67EE57362" blockId="4.[151,1436,1543,1640]" pageId="4" pageNumber="336">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FF02F9C67C727315" bold="true" box="[151,274,1543,1567]" pageId="4" pageNumber="336">FIGURE 3.</emphasis>
|
|||
|
|
Seven paratypes of
|
|||
|
|
<taxonomicName id="5AD44D697672A153FE48F9C97FED7315" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[477,653,1544,1568]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FE48F9C97FED7315" box="[477,653,1544,1568]" italics="true" pageId="4" pageNumber="336">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87672A153FD07F9C97F82732A" bold="true" box="[658,738,1544,1568]" pageId="4" pageNumber="336">
|
|||
|
|
<taxonomicNameLabel id="B49357837672A153FD07F9C97F82732A" box="[658,738,1544,1568]" pageId="4" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Left column approximately dorsal view; right column approximately ventral view.
|
|||
|
|
<emphasis id="AFA0EAF87672A153FEB0F9EA7C5E7349" bold="true" box="[293,318,1579,1603]" pageId="4" pageNumber="336">A.</emphasis>
|
|||
|
|
USNM 548100,
|
|||
|
|
<emphasis id="AFA0EAF87672A153FE63F9ED7F6E734E" bold="true" box="[502,526,1580,1604]" pageId="4" pageNumber="336">B.</emphasis>
|
|||
|
|
USNM 548117,
|
|||
|
|
<emphasis id="AFA0EAF87672A153FD50F9EA7FBE7349" bold="true" box="[709,734,1579,1603]" pageId="4" pageNumber="336">C.</emphasis>
|
|||
|
|
USNM 548118,
|
|||
|
|
<emphasis id="AFA0EAF87672A153FC00F9ED7ECE734E" bold="true" box="[917,942,1580,1604]" pageId="4" pageNumber="336">D.</emphasis>
|
|||
|
|
USNM 548120,
|
|||
|
|
<emphasis id="AFA0EAF87672A153FBF3F9ED791E734E" bold="true" box="[1126,1150,1580,1604]" pageId="4" pageNumber="336">E.</emphasis>
|
|||
|
|
USNM 548121,
|
|||
|
|
<emphasis id="AFA0EAF87672A153FAA3F9ED7829734E" bold="true" box="[1334,1353,1580,1604]" pageId="4" pageNumber="336">F.</emphasis>
|
|||
|
|
USNM 548122,
|
|||
|
|
<emphasis id="AFA0EAF87672A153FF65F98E7C6A736D" bold="true" box="[240,266,1615,1639]" pageId="4" pageNumber="336">G.</emphasis>
|
|||
|
|
USNM 548123. Scale bars 20 mm. Photographs by H. Taylor.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="D5CE65617672A151FF52F95179A4778F" lastPageId="6" lastPageNumber="338" pageId="4" pageNumber="336" type="diagnosis">
|
|||
|
|
<paragraph id="9D6B36EA7672A153FF52F9517CE07230" blockId="4.[151,1437,1680,2031]" pageId="4" pageNumber="336">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FF52F9517C2273A0" bold="true" box="[199,322,1680,1706]" pageId="4" pageNumber="336">Diagnosis.</emphasis>
|
|||
|
|
A
|
|||
|
|
<taxonomicName id="5AD44D697672A153FEF7F9517CB473A0" authorityName="Hemprich" authorityYear="1820" box="[354,468,1680,1706]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FEF7F9517CB473A0" box="[354,468,1680,1706]" italics="true" pageId="4" pageNumber="336">Rhinophis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
restricted to the Rakwana Massif of
|
|||
|
|
<collectingCountry id="E5C3767A7672A153FCEDF9517E8773A0" box="[888,999,1680,1706]" name="Sri Lanka" pageId="4" pageNumber="336">Sri Lanka</collectingCountry>
|
|||
|
|
with 17 dorsal scale rows at midbody, more than 166 and fewer than 190 ventral scales (
|
|||
|
|
<quantity id="5A2C9B0F7672A153FD55F9757E2073C4" box="[704,832,1716,1742]" metricMagnitude="0" metricUnit="m" metricValue="4.4831" metricValueMax="4.7244" metricValueMin="4.2418" pageId="4" pageNumber="336" unit="in" value="176.5" valueMax="186.0" valueMin="167.0">167–186 in</quantity>
|
|||
|
|
the 33 known specimens;
|
|||
|
|
<quantity id="5A2C9B0F7672A153FBF2F975798573C4" box="[1127,1253,1716,1742]" metricMagnitude="0" metricUnit="m" metricValue="4.3815" metricValueMax="4.5212" metricValueMin="4.2418" pageId="4" pageNumber="336" unit="in" value="172.5" valueMax="178.0" valueMin="167.0">167–178 in</quantity>
|
|||
|
|
males,
|
|||
|
|
<quantity id="5A2C9B0F7672A153FAAFF9757DCD73F8" metricMagnitude="0" metricUnit="m" metricValue="4.5847" metricValueMax="4.7244" metricValueMin="4.445" pageId="4" pageNumber="336" unit="in" value="180.5" valueMax="186.0" valueMin="175.0">175–186 in</quantity>
|
|||
|
|
females), a large, domed tail shield with small, homogenous spines, and a distinctive colour pattern (at least in adults) with a dark belly and paler, orange-bronze dorsum (except for darker anterior sixth) with irregular, blackish transverse markings.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7672A153FF52F885797772E5" blockId="4.[151,1437,1680,2031]" pageId="4" pageNumber="336">
|
|||
|
|
<taxonomicName id="5AD44D697672A153FF52F8857CEC7254" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[199,396,1860,1886]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FF52F8857CEC7254" box="[199,396,1860,1886]" italics="true" pageId="4" pageNumber="336">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87672A153FE06F8847C8E7255" bold="true" box="[403,494,1861,1887]" pageId="4" pageNumber="336">
|
|||
|
|
<taxonomicNameLabel id="B49357837672A153FE06F8847C8E7255" box="[403,494,1861,1887]" pageId="4" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from all six Indian species of
|
|||
|
|
<taxonomicName id="5AD44D697672A153FC0CF885796B7254" authorityName="Hemprich" authorityYear="1820" box="[921,1035,1860,1886]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FC0CF885796B7254" box="[921,1035,1860,1886]" italics="true" pageId="4" pageNumber="336">Rhinophis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having a ventral count of> 165 and <190 (versus more than
|
|||
|
|
<quantity id="5A2C9B0F7672A153FE64F8A87F5C7288" box="[497,572,1896,1923]" metricMagnitude="0" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="336" unit="in" value="200.0">200 in</quantity>
|
|||
|
|
<taxonomicName id="5AD44D697672A153FDD3F8A87E977289" authority="Aengals and Ganesh, 2013" authorityName="Aengals and Ganesh" authorityYear="2013" box="[582,1015,1896,1923]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="goweri">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FDD3F8A87FD67288" box="[582,694,1897,1922]" italics="true" pageId="4" pageNumber="336">R. goweri</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7672A153FD2AF8A97E977289" author="Aengals, R. & Ganesh, S. R." box="[703,1015,1896,1923]" pageId="4" pageNumber="336" pagination="61 - 65" refId="ref15770" refString="Aengals, R. & Ganesh, S. R. (2013) Rhinophis goweri - a new species of shieldtail snake from the Southern Eastern Ghats, India. Russian Journal of Herpetology, 20 (1), 61 - 65." type="journal article" year="2013">Aengals and Ganesh, 2013</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, fewer than
|
|||
|
|
<quantity id="5A2C9B0F7672A153FB1FF8A979B47288" box="[1162,1236,1896,1923]" metricMagnitude="0" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="336" unit="in" value="150.0">150 in</quantity>
|
|||
|
|
<taxonomicName id="5AD44D697672A153FB4BF8A87C3772AD" authority="Boulenger, 1893" authorityName="Boulenger" authorityYear="1893" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="travancoricus">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FB4BF8A878FD7288" box="[1246,1437,1897,1922]" italics="true" pageId="4" pageNumber="336">R. travancoricus</emphasis>
|
|||
|
|
Boulenger, 1893
|
|||
|
|
</taxonomicName>
|
|||
|
|
); and in having 17 dorsal scale rows at or just behind midbody (versus
|
|||
|
|
<quantity id="5A2C9B0F7672A153FB0DF84D79B472AC" box="[1176,1236,1932,1959]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="336" unit="in" value="15.0">15 in</quantity>
|
|||
|
|
<taxonomicName id="5AD44D697672A153FB48F84C7C3472C0" authority="Boulenger, 1896" authorityName="Boulenger" authorityYear="1896" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="fergusonianus">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FB48F84C78FD72AD" box="[1245,1437,1933,1959]" italics="true" pageId="4" pageNumber="336">R. fergusonianus</emphasis>
|
|||
|
|
Boulenger, 1896
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697672A153FECAF8707FD972C0" authority="Beddome, 1863" authorityName="Beddome" authorityYear="1863" box="[351,697,1968,1994]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="sanguineus">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FECAF8707C9D72C0" box="[351,509,1969,1994]" italics="true" pageId="4" pageNumber="336">R. sanguineus</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7672A153FD91F8717FD972C0" author="Beddome" box="[516,697,1968,1994]" pageId="4" pageNumber="336" pagination="1 - 24" refId="ref16927" refString="closely related to R. sanguineus Beddome, 1863 (Serpentes: Uropeltidae). Zootaxa, 4881 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4881.1.1" type="journal article" year="1863">Beddome, 1863</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697672A153FD51F8707DB272E5" authority="Cyriac, Narayanan, Sampaio, Umesh & Gower, 2020" authorityName="Cyriac, Narayanan, Sampaio, Umesh & Gower" authorityYear="2020" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="melanoleucus">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FD51F8707E1E72C0" box="[708,894,1968,1994]" italics="true" pageId="4" pageNumber="336">R. melanoleucus</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7672A153FC10F8717DB272E5" author="Cyriac, V. P. & Narayanan, S. & Sampaio, F. L. & Umesh, P. & Gower, D. J." pageId="4" pageNumber="336" pagination="329 - 342" refId="ref16068" refString="Cyriac, V. P., Narayanan, S., Sampaio, F. L., Umesh, P. & Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the Wayanad region of peninsular India. Zootaxa, 4778 (2), 329 - 342. https: // doi. org / 10.11646 / zootaxa. 4778.2.5" type="journal article" year="2020">Cyriac, Narayanan, Sampaio, Umesh & Gower, 2020</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="5AD44D697672A153FE98F814797372E5" authority="Sampaio, Narayanan, Cyriac, Venu & Gower, 2020" authorityName="Sampaio, Narayanan, Cyriac, Venu & Gower" authorityYear="2020" box="[269,1043,2004,2031]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="4" pageNumber="336" phylum="Chordata" rank="species" species="karinthandani">
|
|||
|
|
<emphasis id="AFA0EAF87672A153FE98F8147CAB72E4" box="[269,459,2004,2030]" italics="true" pageId="4" pageNumber="336">R. karinthandani</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7672A153FE44F815797372E5" author="Sampaio, F. L. & Narayanan, S. & Cyriac, V. P. & Venu, G. & Gower, D. J." box="[465,1043,2004,2031]" pageId="4" pageNumber="336" refId="ref16885" refString="Sampaio, F. L., Narayanan, S., Cyriac, V. P., Venu, G. & Gower, D. J. (2020) A new Indian species of Rhinophis Hemprich, 1820" type="book" year="2020">Sampaio, Narayanan, Cyriac, Venu & Gower, 2020</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="C9AB66627673A152FF02FF56793E7411" pageId="5" pageNumber="337" startId="5.[151,239,151,175]" targetBox="[163,1416,301,1285]" targetIsTable="true" targetPageId="5" targetType="table">
|
|||
|
|
<paragraph id="9D6B36EA7673A152FF02FF56793E7411" blockId="5.[151,1437,151,284]" pageId="5" pageNumber="337">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FF02FF567C6875A5" bold="true" box="[151,264,151,175]" pageId="5" pageNumber="337">TABLE 1.</emphasis>
|
|||
|
|
Meristic and morphometric (in mm) data for holotype (*) and paratypes of
|
|||
|
|
<taxonomicName id="5AD44D697673A152FB8DFF5679B075A5" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[1048,1232,151,175]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FB8DFF5679B075A5" box="[1048,1232,151,175]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FB40FF59784975BA" bold="true" box="[1237,1321,152,176]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FB40FF59784975BA" box="[1237,1321,152,176]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="9D6B36EA7673A152FEC2FEEC7C6D700F" pageId="5" pageNumber="337">
|
|||
|
|
<table id="EFD4C44A76735EA8FF36FEEC78E8700F" box="[163,1416,301,1285]" gridcols="12" gridrows="27" pageId="5" pageNumber="337">
|
|||
|
|
<tr id="23E434A876735EA8FF36FEEC78E8744A" box="[163,1416,301,320]" gridrow="0" pageId="5" pageNumber="337" rowspan-0="1">
|
|||
|
|
<th id="60355DD476735EA8FEC4FEEC7CFB744A" box="[337,411,301,320]" gridcol="1" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FE20FEEC7C95744A" box="[437,501,301,320]" gridcol="2" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FD85FEEC7F2F744A" box="[528,591,301,320]" gridcol="3" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FDFFFEEC7FC9744A" box="[618,681,301,320]" gridcol="4" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FD50FEEC7E65744A" box="[709,773,301,320]" gridcol="5" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FC8AFEEC7E3F744A" box="[799,863,301,320]" gridcol="6" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FCEFFEEC7EDA744A" box="[890,954,301,320]" gridcol="7" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FC40FEEC7975744A" box="[981,1045,301,320]" gridcol="8" gridrow="0" pageId="5" pageNumber="337">USNM</th>
|
|||
|
|
<th id="60355DD476735EA8FBA7FEEC79F3744A" box="[1074,1171,301,320]" gridcol="9" gridrow="0" pageId="5" pageNumber="337">NMSL-NH</th>
|
|||
|
|
<th id="60355DD476735EA8FB3AFEEC7870744A" box="[1199,1296,301,320]" gridcol="10" gridrow="0" pageId="5" pageNumber="337">NMSL-NH</th>
|
|||
|
|
<th id="60355DD476735EA8FABDFEEC78E8744A" box="[1320,1416,301,320]" gridcol="11" gridrow="0" pageId="5" pageNumber="337">DWC</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FE9178E8746E" box="[163,1416,336,356]" gridrow="1" pageId="5" pageNumber="337" rowspan-0="1">
|
|||
|
|
<td id="60355DD476735EA8FEC4FE917CFB746E" box="[337,411,336,356]" gridcol="1" gridrow="1" pageId="5" pageNumber="337">548119*</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FE917C95746E" box="[437,501,336,356]" gridcol="2" gridrow="1" pageId="5" pageNumber="337">548100</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FE917F2F746E" box="[528,591,336,356]" gridcol="3" gridrow="1" pageId="5" pageNumber="337">548117</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFE917FC9746E" box="[618,681,336,356]" gridcol="4" gridrow="1" pageId="5" pageNumber="337">548118</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FE917E65746E" box="[709,773,336,356]" gridcol="5" gridrow="1" pageId="5" pageNumber="337">548120</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFE917E3F746E" box="[799,863,336,356]" gridcol="6" gridrow="1" pageId="5" pageNumber="337">548121</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFE917EDA746E" box="[890,954,336,356]" gridcol="7" gridrow="1" pageId="5" pageNumber="337">548122</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FE917975746E" box="[981,1045,336,356]" gridcol="8" gridrow="1" pageId="5" pageNumber="337">548123</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FE9179F3746E" box="[1074,1171,336,356]" gridcol="9" gridrow="1" pageId="5" pageNumber="337">2024.01.01</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFE917870746E" box="[1199,1296,336,356]" gridcol="10" gridrow="1" pageId="5" pageNumber="337">2024.01.02</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFE9178E8746E" box="[1320,1416,336,356]" gridcol="11" gridrow="1" pageId="5" pageNumber="337">2024.05.01</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FEB778E87480" box="[163,1416,374,394]" gridrow="2" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FEB77C5A7480" box="[163,314,374,394]" gridcol="0" gridrow="2" pageId="5" pageNumber="337">Sex</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FEB77CFB7480" box="[337,411,374,394]" gridcol="1" gridrow="2" pageId="5" pageNumber="337">F</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FEB77C957480" box="[437,501,374,394]" gridcol="2" gridrow="2" pageId="5" pageNumber="337">F</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FEB77F2F7480" box="[528,591,374,394]" gridcol="3" gridrow="2" pageId="5" pageNumber="337">M</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFEB77FC97480" box="[618,681,374,394]" gridcol="4" gridrow="2" pageId="5" pageNumber="337">M</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FEB77E657480" box="[709,773,374,394]" gridcol="5" gridrow="2" pageId="5" pageNumber="337">F</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFEB77E3F7480" box="[799,863,374,394]" gridcol="6" gridrow="2" pageId="5" pageNumber="337">F</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFEB77EDA7480" box="[890,954,374,394]" gridcol="7" gridrow="2" pageId="5" pageNumber="337">M</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FEB779757480" box="[981,1045,374,394]" gridcol="8" gridrow="2" pageId="5" pageNumber="337">M</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FEB779F37480" box="[1074,1171,374,394]" gridcol="9" gridrow="2" pageId="5" pageNumber="337">F</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFEB778707480" box="[1199,1296,374,394]" gridcol="10" gridrow="2" pageId="5" pageNumber="337">M</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFEB778E87480" box="[1320,1416,374,394]" gridcol="11" gridrow="2" pageId="5" pageNumber="337">M</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FE5A78E874A5" box="[163,1416,411,431]" gridrow="3" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FE5A7C5A74A5" box="[163,314,411,431]" gridcol="0" gridrow="3" pageId="5" pageNumber="337">Total length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FE5A7CFB74A5" box="[337,411,411,431]" gridcol="1" gridrow="3" pageId="5" pageNumber="337">243</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FE5A7C9574A5" box="[437,501,411,431]" gridcol="2" gridrow="3" pageId="5" pageNumber="337">226</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FE5A7F2F74A5" box="[528,591,411,431]" gridcol="3" gridrow="3" pageId="5" pageNumber="337">172</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFE5A7FC974A5" box="[618,681,411,431]" gridcol="4" gridrow="3" pageId="5" pageNumber="337">182</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FE5A7E6574A5" box="[709,773,411,431]" gridcol="5" gridrow="3" pageId="5" pageNumber="337">220</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFE5A7E3F74A5" box="[799,863,411,431]" gridcol="6" gridrow="3" pageId="5" pageNumber="337">232</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFE5A7EDA74A5" box="[890,954,411,431]" gridcol="7" gridrow="3" pageId="5" pageNumber="337">238</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FE5A797574A5" box="[981,1045,411,431]" gridcol="8" gridrow="3" pageId="5" pageNumber="337">241</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FE5A79F374A5" box="[1074,1171,411,431]" gridcol="9" gridrow="3" pageId="5" pageNumber="337">231</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFE5A787074A5" box="[1199,1296,411,431]" gridcol="10" gridrow="3" pageId="5" pageNumber="337">205</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFE5A78E874A5" box="[1320,1416,411,431]" gridcol="11" gridrow="3" pageId="5" pageNumber="337">186</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FE0078E874DE" box="[163,1416,449,468]" gridrow="4" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FE007C5A74DE" box="[163,314,449,468]" gridcol="0" gridrow="4" pageId="5" pageNumber="337">Tail length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FE007CFB74DE" box="[337,411,449,468]" gridcol="1" gridrow="4" pageId="5" pageNumber="337">5.7</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FE007C9574DE" box="[437,501,449,468]" gridcol="2" gridrow="4" pageId="5" pageNumber="337">5.2</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FE007F2F74DE" box="[528,591,449,468]" gridcol="3" gridrow="4" pageId="5" pageNumber="337">6.9</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFE007FC974DE" box="[618,681,449,468]" gridcol="4" gridrow="4" pageId="5" pageNumber="337">7.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FE007E6574DE" box="[709,773,449,468]" gridcol="5" gridrow="4" pageId="5" pageNumber="337">6.3</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFE007E3F74DE" box="[799,863,449,468]" gridcol="6" gridrow="4" pageId="5" pageNumber="337">6.3</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFE007EDA74DE" box="[890,954,449,468]" gridcol="7" gridrow="4" pageId="5" pageNumber="337">9.9</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FE00797574DE" box="[981,1045,449,468]" gridcol="8" gridrow="4" pageId="5" pageNumber="337">10.0</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FE0079F374DE" box="[1074,1171,449,468]" gridcol="9" gridrow="4" pageId="5" pageNumber="337">6</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFE00787074DE" box="[1199,1296,449,468]" gridcol="10" gridrow="4" pageId="5" pageNumber="337">8.5</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFE0078E874DE" box="[1320,1416,449,468]" gridcol="11" gridrow="4" pageId="5" pageNumber="337">7.5</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FE2778E874F0" box="[163,1416,486,506]" gridrow="5" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FE277C5A74F0" box="[163,314,486,506]" gridcol="0" gridrow="5" pageId="5" pageNumber="337">Midbody width</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FE277CFB74F0" box="[337,411,486,506]" gridcol="1" gridrow="5" pageId="5" pageNumber="337">8.6</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FE277C9574F0" box="[437,501,486,506]" gridcol="2" gridrow="5" pageId="5" pageNumber="337">7.2</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FE277F2F74F0" box="[528,591,486,506]" gridcol="3" gridrow="5" pageId="5" pageNumber="337">5.7</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFE277FC974F0" box="[618,681,486,506]" gridcol="4" gridrow="5" pageId="5" pageNumber="337">5.6</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FE277E6574F0" box="[709,773,486,506]" gridcol="5" gridrow="5" pageId="5" pageNumber="337">7.3</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFE277E3F74F0" box="[799,863,486,506]" gridcol="6" gridrow="5" pageId="5" pageNumber="337">7.9</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFE277EDA74F0" box="[890,954,486,506]" gridcol="7" gridrow="5" pageId="5" pageNumber="337">8.2</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FE27797574F0" box="[981,1045,486,506]" gridcol="8" gridrow="5" pageId="5" pageNumber="337">7.9</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FE2779F374F0" box="[1074,1171,486,506]" gridcol="9" gridrow="5" pageId="5" pageNumber="337">7.8</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFE27787074F0" box="[1199,1296,486,506]" gridcol="10" gridrow="5" pageId="5" pageNumber="337">7.3</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFE2778E874F0" box="[1320,1416,486,506]" gridcol="11" gridrow="5" pageId="5" pageNumber="337">6.8</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FDCD78E87715" box="[163,1416,524,543]" gridrow="6" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FDCD7C5A7715" box="[163,314,524,543]" gridcol="0" gridrow="6" pageId="5" pageNumber="337">Midbody</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FDCD7CFB7715" box="[337,411,524,543]" gridcol="1" gridrow="6" pageId="5" pageNumber="337">25</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FDCD7C957715" box="[437,501,524,543]" gridcol="2" gridrow="6" pageId="5" pageNumber="337">22</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FDCD7F2F7715" box="[528,591,524,543]" gridcol="3" gridrow="6" pageId="5" pageNumber="337">19</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFDCD7FC97715" box="[618,681,524,543]" gridcol="4" gridrow="6" pageId="5" pageNumber="337">19</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FDCD7E657715" box="[709,773,524,543]" gridcol="5" gridrow="6" pageId="5" pageNumber="337">24</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFDCD7E3F7715" box="[799,863,524,543]" gridcol="6" gridrow="6" pageId="5" pageNumber="337">24</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFDCD7EDA7715" box="[890,954,524,543]" gridcol="7" gridrow="6" pageId="5" pageNumber="337">26</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FDCD79757715" box="[981,1045,524,543]" gridcol="8" gridrow="6" pageId="5" pageNumber="337">26</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FDCD79F37715" box="[1074,1171,524,543]" gridcol="9" gridrow="6" pageId="5" pageNumber="337">25</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFDCD78707715" box="[1199,1296,524,543]" gridcol="10" gridrow="6" pageId="5" pageNumber="337">23</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFDCD78E87715" box="[1320,1416,524,543]" gridcol="11" gridrow="6" pageId="5" pageNumber="337">21</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FDF178E87749" box="[163,1416,560,579]" gridrow="7" pageId="5" pageNumber="337" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FDF17C5A7749" box="[163,314,560,579]" gridcol="0" gridrow="7" pageId="5" pageNumber="337">circumference</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FD9478E87763" box="[163,1416,597,617]" gridrow="8" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FD947C5A7763" box="[163,314,597,617]" gridcol="0" gridrow="8" pageId="5" pageNumber="337">Head length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FD947CFB7763" box="[337,411,597,617]" gridcol="1" gridrow="8" pageId="5" pageNumber="337">5.7</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FD947C957763" box="[437,501,597,617]" gridcol="2" gridrow="8" pageId="5" pageNumber="337">5.5</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FD947F2F7763" box="[528,591,597,617]" gridcol="3" gridrow="8" pageId="5" pageNumber="337">4.8</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFD947FC97763" box="[618,681,597,617]" gridcol="4" gridrow="8" pageId="5" pageNumber="337">5.0</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FD947E657763" box="[709,773,597,617]" gridcol="5" gridrow="8" pageId="5" pageNumber="337">5.4</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFD947E3F7763" box="[799,863,597,617]" gridcol="6" gridrow="8" pageId="5" pageNumber="337">5.3</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFD947EDA7763" box="[890,954,597,617]" gridcol="7" gridrow="8" pageId="5" pageNumber="337">6.1</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FD9479757763" box="[981,1045,597,617]" gridcol="8" gridrow="8" pageId="5" pageNumber="337">6.4</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FD9479F37763" box="[1074,1171,597,617]" gridcol="9" gridrow="8" pageId="5" pageNumber="337">5.7</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFD9478707763" box="[1199,1296,597,617]" gridcol="10" gridrow="8" pageId="5" pageNumber="337">5.7</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFD9478E87763" box="[1320,1416,597,617]" gridcol="11" gridrow="8" pageId="5" pageNumber="337">5.6</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FDBA78E87784" box="[163,1416,635,654]" gridrow="9" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FDBA7C5A7784" box="[163,314,635,654]" gridcol="0" gridrow="9" pageId="5" pageNumber="337">Head width</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FDBA7CFB7784" box="[337,411,635,654]" gridcol="1" gridrow="9" pageId="5" pageNumber="337">3.9</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FDBA7C957784" box="[437,501,635,654]" gridcol="2" gridrow="9" pageId="5" pageNumber="337">3.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FDBA7F2F7784" box="[528,591,635,654]" gridcol="3" gridrow="9" pageId="5" pageNumber="337">3.3</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFDBA7FC97784" box="[618,681,635,654]" gridcol="4" gridrow="9" pageId="5" pageNumber="337">3.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FDBA7E657784" box="[709,773,635,654]" gridcol="5" gridrow="9" pageId="5" pageNumber="337">3.6</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFDBA7E3F7784" box="[799,863,635,654]" gridcol="6" gridrow="9" pageId="5" pageNumber="337">4.0</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFDBA7EDA7784" box="[890,954,635,654]" gridcol="7" gridrow="9" pageId="5" pageNumber="337">4.4</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FDBA79757784" box="[981,1045,635,654]" gridcol="8" gridrow="9" pageId="5" pageNumber="337">4.0</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FDBA79F37784" box="[1074,1171,635,654]" gridcol="9" gridrow="9" pageId="5" pageNumber="337">4.4</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFDBA78707784" box="[1199,1296,635,654]" gridcol="10" gridrow="9" pageId="5" pageNumber="337">3.9</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFDBA78E87784" box="[1320,1416,635,654]" gridcol="11" gridrow="9" pageId="5" pageNumber="337">4.0</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FD6178E877BE" box="[163,1416,672,692]" gridrow="10" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FD617C5A77BE" box="[163,314,672,692]" gridcol="0" gridrow="10" pageId="5" pageNumber="337">Eye diameter</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FD617CFB77BE" box="[337,411,672,692]" gridcol="1" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FD617C9577BE" box="[437,501,672,692]" gridcol="2" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FD617F2F77BE" box="[528,591,672,692]" gridcol="3" gridrow="10" pageId="5" pageNumber="337">0.3</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFD617FC977BE" box="[618,681,672,692]" gridcol="4" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FD617E6577BE" box="[709,773,672,692]" gridcol="5" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFD617E3F77BE" box="[799,863,672,692]" gridcol="6" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFD617EDA77BE" box="[890,954,672,692]" gridcol="7" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FD61797577BE" box="[981,1045,672,692]" gridcol="8" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FD6179F377BE" box="[1074,1171,672,692]" gridcol="9" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFD61787077BE" box="[1199,1296,672,692]" gridcol="10" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFD6178E877BE" box="[1320,1416,672,692]" gridcol="11" gridrow="10" pageId="5" pageNumber="337">0.4</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FD0778E877D3" box="[163,1416,710,729]" gridrow="11" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FD077C5A77D3" box="[163,314,710,729]" gridcol="0" gridrow="11" pageId="5" pageNumber="337">Eye–snout tip</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FD077CFB77D3" box="[337,411,710,729]" gridcol="1" gridrow="11" pageId="5" pageNumber="337">3.0</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FD077C9577D3" box="[437,501,710,729]" gridcol="2" gridrow="11" pageId="5" pageNumber="337">3.0</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FD077F2F77D3" box="[528,591,710,729]" gridcol="3" gridrow="11" pageId="5" pageNumber="337">2.8</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFD077FC977D3" box="[618,681,710,729]" gridcol="4" gridrow="11" pageId="5" pageNumber="337">2.9</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FD077E6577D3" box="[709,773,710,729]" gridcol="5" gridrow="11" pageId="5" pageNumber="337">2.8</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFD077E3F77D3" box="[799,863,710,729]" gridcol="6" gridrow="11" pageId="5" pageNumber="337">2.8</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFD077EDA77D3" box="[890,954,710,729]" gridcol="7" gridrow="11" pageId="5" pageNumber="337">3.2</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FD07797577D3" box="[981,1045,710,729]" gridcol="8" gridrow="11" pageId="5" pageNumber="337">3.4</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FD0779F377D3" box="[1074,1171,710,729]" gridcol="9" gridrow="11" pageId="5" pageNumber="337">3.1</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFD07787077D3" box="[1199,1296,710,729]" gridcol="10" gridrow="11" pageId="5" pageNumber="337">2.9</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFD0778E877D3" box="[1320,1416,710,729]" gridcol="11" gridrow="11" pageId="5" pageNumber="337">2.9</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FD2A78E877F5" box="[163,1416,747,767]" gridrow="12" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FD2A7C5A77F5" box="[163,314,747,767]" gridcol="0" gridrow="12" pageId="5" pageNumber="337">Eye–eye</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FD2A7CFB77F5" box="[337,411,747,767]" gridcol="1" gridrow="12" pageId="5" pageNumber="337">2.6</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FD2A7C9577F5" box="[437,501,747,767]" gridcol="2" gridrow="12" pageId="5" pageNumber="337">2.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FD2A7F2F77F5" box="[528,591,747,767]" gridcol="3" gridrow="12" pageId="5" pageNumber="337">2.3</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFD2A7FC977F5" box="[618,681,747,767]" gridcol="4" gridrow="12" pageId="5" pageNumber="337">2.5</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FD2A7E6577F5" box="[709,773,747,767]" gridcol="5" gridrow="12" pageId="5" pageNumber="337">2.5</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFD2A7E3F77F5" box="[799,863,747,767]" gridcol="6" gridrow="12" pageId="5" pageNumber="337">2.6</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFD2A7EDA77F5" box="[890,954,747,767]" gridcol="7" gridrow="12" pageId="5" pageNumber="337">3.0</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FD2A797577F5" box="[981,1045,747,767]" gridcol="8" gridrow="12" pageId="5" pageNumber="337">2.7</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FD2A79F377F5" box="[1074,1171,747,767]" gridcol="9" gridrow="12" pageId="5" pageNumber="337">2.5</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFD2A787077F5" box="[1199,1296,747,767]" gridcol="10" gridrow="12" pageId="5" pageNumber="337">2.5</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFD2A78E877F5" box="[1320,1416,747,767]" gridcol="11" gridrow="12" pageId="5" pageNumber="337">2.6</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FCD078E8762E" box="[163,1416,785,804]" gridrow="13" pageId="5" pageNumber="337" rowspan-10="1" rowspan-11="1" rowspan-3="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FCD07C5A762E" box="[163,314,785,804]" gridcol="0" gridrow="13" pageId="5" pageNumber="337">Rostral length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FCD07CFB762E" box="[337,411,785,804]" gridcol="1" gridrow="13" pageId="5" pageNumber="337">2.6</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FCD07C95762E" box="[437,501,785,804]" gridcol="2" gridrow="13" pageId="5" pageNumber="337">2.2</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFCD07FC9762E" box="[618,681,785,804]" gridcol="4" gridrow="13" pageId="5" pageNumber="337">2.6</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FCD07E65762E" box="[709,773,785,804]" gridcol="5" gridrow="13" pageId="5" pageNumber="337">2.2</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFCD07E3F762E" box="[799,863,785,804]" gridcol="6" gridrow="13" pageId="5" pageNumber="337">2.2</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFCD07EDA762E" box="[890,954,785,804]" gridcol="7" gridrow="13" pageId="5" pageNumber="337">2.5</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FCD07975762E" box="[981,1045,785,804]" gridcol="8" gridrow="13" pageId="5" pageNumber="337">2.4</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FCD079F3762E" box="[1074,1171,785,804]" gridcol="9" gridrow="13" pageId="5" pageNumber="337">2.3</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FCF778E87643" box="[163,1416,822,841]" gridrow="14" pageId="5" pageNumber="337" rowspan-10="1" rowspan-11="1" rowspan-3="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FCF77C5A7643" box="[163,314,822,841]" gridcol="0" gridrow="14" pageId="5" pageNumber="337">Frontal length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FCF77CFB7643" box="[337,411,822,841]" gridcol="1" gridrow="14" pageId="5" pageNumber="337">1.9</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FCF77C957643" box="[437,501,822,841]" gridcol="2" gridrow="14" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFCF77FC97643" box="[618,681,822,841]" gridcol="4" gridrow="14" pageId="5" pageNumber="337">1.8</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FCF77E657643" box="[709,773,822,841]" gridcol="5" gridrow="14" pageId="5" pageNumber="337">1.6</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFCF77E3F7643" box="[799,863,822,841]" gridcol="6" gridrow="14" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFCF77EDA7643" box="[890,954,822,841]" gridcol="7" gridrow="14" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FCF779757643" box="[981,1045,822,841]" gridcol="8" gridrow="14" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FCF779F37643" box="[1074,1171,822,841]" gridcol="9" gridrow="14" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FC9D78E87665" box="[163,1416,860,879]" gridrow="15" pageId="5" pageNumber="337" rowspan-10="1" rowspan-11="1" rowspan-3="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FC9D7C5A7665" box="[163,314,860,879]" gridcol="0" gridrow="15" pageId="5" pageNumber="337">Frontal width</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FC9D7CFB7665" box="[337,411,860,879]" gridcol="1" gridrow="15" pageId="5" pageNumber="337">1.8</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FC9D7C957665" box="[437,501,860,879]" gridcol="2" gridrow="15" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFC9D7FC97665" box="[618,681,860,879]" gridcol="4" gridrow="15" pageId="5" pageNumber="337">1.6</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FC9D7E657665" box="[709,773,860,879]" gridcol="5" gridrow="15" pageId="5" pageNumber="337">1.6</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFC9D7E3F7665" box="[799,863,860,879]" gridcol="6" gridrow="15" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFC9D7EDA7665" box="[890,954,860,879]" gridcol="7" gridrow="15" pageId="5" pageNumber="337">1.7</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FC9D79757665" box="[981,1045,860,879]" gridcol="8" gridrow="15" pageId="5" pageNumber="337">1.6</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FC9D79F37665" box="[1074,1171,860,879]" gridcol="9" gridrow="15" pageId="5" pageNumber="337">1.6</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FC4078E8769E" box="[163,1416,897,916]" gridrow="16" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FC407C5A769E" box="[163,314,897,916]" gridcol="0" gridrow="16" pageId="5" pageNumber="337">Ventral scales</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FC407CFB769E" box="[337,411,897,916]" gridcol="1" gridrow="16" pageId="5" pageNumber="337">186</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FC407C95769E" box="[437,501,897,916]" gridcol="2" gridrow="16" pageId="5" pageNumber="337">180</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FC407F2F769E" box="[528,591,897,916]" gridcol="3" gridrow="16" pageId="5" pageNumber="337">172</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFC407FC9769E" box="[618,681,897,916]" gridcol="4" gridrow="16" pageId="5" pageNumber="337">173</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FC407E65769E" box="[709,773,897,916]" gridcol="5" gridrow="16" pageId="5" pageNumber="337">178</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFC407E3F769E" box="[799,863,897,916]" gridcol="6" gridrow="16" pageId="5" pageNumber="337">181</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFC407EDA769E" box="[890,954,897,916]" gridcol="7" gridrow="16" pageId="5" pageNumber="337">174</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FC407975769E" box="[981,1045,897,916]" gridcol="8" gridrow="16" pageId="5" pageNumber="337">172</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FC4079F3769E" box="[1074,1171,897,916]" gridcol="9" gridrow="16" pageId="5" pageNumber="337">175</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFC407870769E" box="[1199,1296,897,916]" gridcol="10" gridrow="16" pageId="5" pageNumber="337">173</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFC4078E8769E" box="[1320,1416,897,916]" gridcol="11" gridrow="16" pageId="5" pageNumber="337">167</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FC6678E876B0" box="[163,1416,935,954]" gridrow="17" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FC667C5A76B0" box="[163,314,935,954]" gridcol="0" gridrow="17" pageId="5" pageNumber="337">Subcaudal scales</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FC667CFB76B0" box="[337,411,935,954]" gridcol="1" gridrow="17" pageId="5" pageNumber="337">3,3 (2)</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FC667C9576B0" box="[437,501,935,954]" gridcol="2" gridrow="17" pageId="5" pageNumber="337">3,3 (1)</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FC667F2F76B0" box="[528,591,935,954]" gridcol="3" gridrow="17" pageId="5" pageNumber="337">6,6 (2)</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFC667FC976B0" box="[618,681,935,954]" gridcol="4" gridrow="17" pageId="5" pageNumber="337">6,6 (0)</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FC667E6576B0" box="[709,773,935,954]" gridcol="5" gridrow="17" pageId="5" pageNumber="337">4,4 (3)</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFC667E3F76B0" box="[799,863,935,954]" gridcol="6" gridrow="17" pageId="5" pageNumber="337">4,4 (2)</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFC667EDA76B0" box="[890,954,935,954]" gridcol="7" gridrow="17" pageId="5" pageNumber="337">6,6 (2)</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FC66797576B0" box="[981,1045,935,954]" gridcol="8" gridrow="17" pageId="5" pageNumber="337">7,6 (3)</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FC6679F376B0" box="[1074,1171,935,954]" gridcol="9" gridrow="17" pageId="5" pageNumber="337">4,4 (2)</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFC66787076B0" box="[1199,1296,935,954]" gridcol="10" gridrow="17" pageId="5" pageNumber="337">6,6 (0)</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFC6678E876B0" box="[1320,1416,935,954]" gridcol="11" gridrow="17" pageId="5" pageNumber="337">5,5 (4)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FC0D78E876D5" box="[163,1416,972,991]" gridrow="18" pageId="5" pageNumber="337" rowspan-9="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FC0D7C5A76D5" box="[163,314,972,991]" gridcol="0" gridrow="18" pageId="5" pageNumber="337">Scales around</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FC0D7CFB76D5" box="[337,411,972,991]" gridcol="1" gridrow="18" pageId="5" pageNumber="337">11</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FC0D7C9576D5" box="[437,501,972,991]" gridcol="2" gridrow="18" pageId="5" pageNumber="337">12</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FC0D7F2F76D5" box="[528,591,972,991]" gridcol="3" gridrow="18" pageId="5" pageNumber="337">13</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFC0D7FC976D5" box="[618,681,972,991]" gridcol="4" gridrow="18" pageId="5" pageNumber="337">11</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FC0D7E6576D5" box="[709,773,972,991]" gridcol="5" gridrow="18" pageId="5" pageNumber="337">11</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFC0D7E3F76D5" box="[799,863,972,991]" gridcol="6" gridrow="18" pageId="5" pageNumber="337">10</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFC0D7EDA76D5" box="[890,954,972,991]" gridcol="7" gridrow="18" pageId="5" pageNumber="337">12</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FC0D797576D5" box="[981,1045,972,991]" gridcol="8" gridrow="18" pageId="5" pageNumber="337">12</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFC0D787076D5" box="[1199,1296,972,991]" gridcol="10" gridrow="18" pageId="5" pageNumber="337">11</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFC0D78E876D5" box="[1320,1416,972,991]" gridcol="11" gridrow="18" pageId="5" pageNumber="337">12</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FC3178E87109" box="[163,1416,1008,1027]" gridrow="19" pageId="5" pageNumber="337" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FC317C5A7109" box="[163,314,1008,1027]" gridcol="0" gridrow="19" pageId="5" pageNumber="337">shield base</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FBD478E87123" box="[163,1416,1045,1065]" gridrow="20" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FBD47C5A7123" box="[163,314,1045,1065]" gridcol="0" gridrow="20" pageId="5" pageNumber="337">Shield base</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FBD47CFB7123" box="[337,411,1045,1065]" gridcol="1" gridrow="20" pageId="5" pageNumber="337">5.4</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FBD47C957123" box="[437,501,1045,1065]" gridcol="2" gridrow="20" pageId="5" pageNumber="337">5.2</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FBD47F2F7123" box="[528,591,1045,1065]" gridcol="3" gridrow="20" pageId="5" pageNumber="337">4.2</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFBD47FC97123" box="[618,681,1045,1065]" gridcol="4" gridrow="20" pageId="5" pageNumber="337">4.4</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FBD47E657123" box="[709,773,1045,1065]" gridcol="5" gridrow="20" pageId="5" pageNumber="337">5.2</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFBD47E3F7123" box="[799,863,1045,1065]" gridcol="6" gridrow="20" pageId="5" pageNumber="337">4.8</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFBD47EDA7123" box="[890,954,1045,1065]" gridcol="7" gridrow="20" pageId="5" pageNumber="337">5.8</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FBD479757123" box="[981,1045,1045,1065]" gridcol="8" gridrow="20" pageId="5" pageNumber="337">5.8</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FBD479F37123" box="[1074,1171,1045,1065]" gridcol="9" gridrow="20" pageId="5" pageNumber="337">5.8</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFBD478707123" box="[1199,1296,1045,1065]" gridcol="10" gridrow="20" pageId="5" pageNumber="337">5.5</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFBD478E87123" box="[1320,1416,1045,1065]" gridcol="11" gridrow="20" pageId="5" pageNumber="337">5.2</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FBFB78E87147" box="[163,1416,1082,1101]" gridrow="21" pageId="5" pageNumber="337" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FBFB7C5A7147" box="[163,314,1082,1101]" gridcol="0" gridrow="21" pageId="5" pageNumber="337">height</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FB9E78E87178" box="[163,1416,1119,1138]" gridrow="22" pageId="5" pageNumber="337">
|
|||
|
|
<th id="60355DD476735EA8FF36FB9E7C5A7178" box="[163,314,1119,1138]" gridcol="0" gridrow="22" pageId="5" pageNumber="337">Shield base width</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FB9E7CFB7178" box="[337,411,1119,1138]" gridcol="1" gridrow="22" pageId="5" pageNumber="337">4.9</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FB9E7C957178" box="[437,501,1119,1138]" gridcol="2" gridrow="22" pageId="5" pageNumber="337">4.3</td>
|
|||
|
|
<td id="60355DD476735EA8FD85FB9E7F2F7178" box="[528,591,1119,1138]" gridcol="3" gridrow="22" pageId="5" pageNumber="337">4.0</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFB9E7FC97178" box="[618,681,1119,1138]" gridcol="4" gridrow="22" pageId="5" pageNumber="337">3.9</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FB9E7E657178" box="[709,773,1119,1138]" gridcol="5" gridrow="22" pageId="5" pageNumber="337">4.8</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFB9E7E3F7178" box="[799,863,1119,1138]" gridcol="6" gridrow="22" pageId="5" pageNumber="337">4.5</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFB9E7EDA7178" box="[890,954,1119,1138]" gridcol="7" gridrow="22" pageId="5" pageNumber="337">4.9</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FB9E79757178" box="[981,1045,1119,1138]" gridcol="8" gridrow="22" pageId="5" pageNumber="337">4.4</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FB9E79F37178" box="[1074,1171,1119,1138]" gridcol="9" gridrow="22" pageId="5" pageNumber="337">4.9</td>
|
|||
|
|
<td id="60355DD476735EA8FB3AFB9E78707178" box="[1199,1296,1119,1138]" gridcol="10" gridrow="22" pageId="5" pageNumber="337">4.1</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFB9E78E87178" box="[1320,1416,1119,1138]" gridcol="11" gridrow="22" pageId="5" pageNumber="337">4.3</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FB4578E87192" box="[163,1416,1156,1176]" gridrow="23" pageId="5" pageNumber="337" rowspan-10="1" rowspan-3="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FB457C5A7192" box="[163,314,1156,1176]" gridcol="0" gridrow="23" pageId="5" pageNumber="337">Shield length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FB457CFB7192" box="[337,411,1156,1176]" gridcol="1" gridrow="23" pageId="5" pageNumber="337">4.8</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FB457C957192" box="[437,501,1156,1176]" gridcol="2" gridrow="23" pageId="5" pageNumber="337">3.5</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFB457FC97192" box="[618,681,1156,1176]" gridcol="4" gridrow="23" pageId="5" pageNumber="337">4.9</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FB457E657192" box="[709,773,1156,1176]" gridcol="5" gridrow="23" pageId="5" pageNumber="337">4.0</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFB457E3F7192" box="[799,863,1156,1176]" gridcol="6" gridrow="23" pageId="5" pageNumber="337">4.2</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFB457EDA7192" box="[890,954,1156,1176]" gridcol="7" gridrow="23" pageId="5" pageNumber="337">4.6</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FB4579757192" box="[981,1045,1156,1176]" gridcol="8" gridrow="23" pageId="5" pageNumber="337">4.5</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FB4579F37192" box="[1074,1171,1156,1176]" gridcol="9" gridrow="23" pageId="5" pageNumber="337">4.2</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFB4578E87192" box="[1320,1416,1156,1176]" gridcol="11" gridrow="23" pageId="5" pageNumber="337">3.4</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FB6978E871B1" box="[163,1416,1192,1211]" gridrow="24" pageId="5" pageNumber="337" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FB697C5A71B1" box="[163,314,1192,1211]" gridcol="0" gridrow="24" pageId="5" pageNumber="337">(middorsal)</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FB0F78E871EB" box="[163,1416,1230,1249]" gridrow="25" pageId="5" pageNumber="337" rowspan-10="1" rowspan-3="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FB0F7C5A71EB" box="[163,314,1230,1249]" gridcol="0" gridrow="25" pageId="5" pageNumber="337">Shield length</th>
|
|||
|
|
<td id="60355DD476735EA8FEC4FB0F7CFB71EB" box="[337,411,1230,1249]" gridcol="1" gridrow="25" pageId="5" pageNumber="337">1.9</td>
|
|||
|
|
<td id="60355DD476735EA8FE20FB0F7C9571EB" box="[437,501,1230,1249]" gridcol="2" gridrow="25" pageId="5" pageNumber="337">1.9</td>
|
|||
|
|
<td id="60355DD476735EA8FDFFFB0F7FC971EB" box="[618,681,1230,1249]" gridcol="4" gridrow="25" pageId="5" pageNumber="337">2.1</td>
|
|||
|
|
<td id="60355DD476735EA8FD50FB0F7E6571EB" box="[709,773,1230,1249]" gridcol="5" gridrow="25" pageId="5" pageNumber="337">2.0</td>
|
|||
|
|
<td id="60355DD476735EA8FC8AFB0F7E3F71EB" box="[799,863,1230,1249]" gridcol="6" gridrow="25" pageId="5" pageNumber="337">2.0</td>
|
|||
|
|
<td id="60355DD476735EA8FCEFFB0F7EDA71EB" box="[890,954,1230,1249]" gridcol="7" gridrow="25" pageId="5" pageNumber="337">2.2</td>
|
|||
|
|
<td id="60355DD476735EA8FC40FB0F797571EB" box="[981,1045,1230,1249]" gridcol="8" gridrow="25" pageId="5" pageNumber="337">2.3</td>
|
|||
|
|
<td id="60355DD476735EA8FBA7FB0F79F371EB" box="[1074,1171,1230,1249]" gridcol="9" gridrow="25" pageId="5" pageNumber="337">1.5</td>
|
|||
|
|
<td id="60355DD476735EA8FABDFB0F78E871EB" box="[1320,1416,1230,1249]" gridcol="11" gridrow="25" pageId="5" pageNumber="337">1.3</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876735EA8FF36FB3378E8700F" box="[163,1416,1266,1285]" gridrow="26" pageId="5" pageNumber="337" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
|||
|
|
<th id="60355DD476735EA8FF36FB337C5A700F" box="[163,314,1266,1285]" gridcol="0" gridrow="26" pageId="5" pageNumber="337">(midventral)</th>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7673A151FF52FAF7783C744B" blockId="5.[151,1437,1334,2009]" lastBlockId="6.[151,1437,150,645]" lastPageId="6" lastPageNumber="338" pageId="5" pageNumber="337">
|
|||
|
|
Among
|
|||
|
|
<collectingRegion id="5F10F8087673A152FEB1FAF77C25705A" box="[292,325,1334,1360]" country="United Kingdom" name="Surrey" pageId="5" pageNumber="337">Sri</collectingRegion>
|
|||
|
|
Lankan congeners,
|
|||
|
|
<taxonomicName id="5AD44D697673A152FDB9FAF77F92705A" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[556,754,1334,1360]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FDB9FAF77F92705A" box="[556,754,1334,1360]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FD6FFAF77E36705A" bold="true" box="[762,854,1334,1360]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FD6FFAF77E36705A" box="[762,854,1334,1360]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="5AD44D697673A152FC7AFAF67831705B" authority="(Kelaart, 1853)" baseAuthorityName="Kelaart" baseAuthorityYear="1853" box="[1007,1361,1334,1361]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="saffragamus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FC7AFAF679FA705A" box="[1007,1178,1334,1360]" italics="true" pageId="5" pageNumber="337">R. saffragamus</emphasis>
|
|||
|
|
(Kelaart, 1853)
|
|||
|
|
</taxonomicName>
|
|||
|
|
in not having a flat tail shield or midline contact between the opposite nasal shields, and by having dorsal scales in 17 rather than 19 rows at midbody. The new species differs from
|
|||
|
|
<taxonomicName id="5AD44D697673A152FCDAFABE79FF7093" authority="(Gray, 1858)" baseAuthorityName="Gray" baseAuthorityYear="1858" box="[847,1183,1406,1433]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="melanogaster">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FCDAFABE79677092" box="[847,1031,1406,1432]" italics="true" pageId="5" pageNumber="337">R. melanogaster</emphasis>
|
|||
|
|
(Gray, 1858)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697673A152FB3EFABE7DBB70B6" authority="(Nicholls, 1929)" baseAuthorityName="Nicholls" baseAuthorityYear="1929" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="phillipsi">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FB3EFABE78467092" box="[1195,1318,1406,1432]" italics="true" pageId="5" pageNumber="337">R. phillipsi</emphasis>
|
|||
|
|
(Nicholls, 1929)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="5AD44D697673A152FE8CFA62791B70B6" authority="Wickramasinghe, Vidanapathirana, Rajeev & Gower, 2017" authorityName="Wickramasinghe, Vidanapathirana, Rajeev & Gower" authorityYear="2017" box="[281,1147,1442,1468]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="roshanpererai">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FE8CFA627CB870B6" box="[281,472,1442,1468]" italics="true" pageId="5" pageNumber="337">R. roshanpererai</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7673A152FE74FA63791B70B6" author="Wickramasinghe, L. M. & Vidanapathirana, D. R. & Rajeev, M. G. & Gower, D. J." box="[481,1147,1442,1468]" pageId="5" pageNumber="337" pagination="153 - 164" refId="ref17165" refString="Wickramasinghe, L. M., Vidanapathirana, D. R., Rajeev, M. G. & Gower, D. J. (2017) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from the central hills of Sri Lanka. Zootaxa, 4263 (1), 153 - 164. https: // doi. org / 10.11646 / zootaxa. 4263.1.7" type="journal article" year="2017">
|
|||
|
|
Wickramasinghe, Vidanapathirana, Rajeev & Gower,
|
|||
|
|
<quantity id="5A2C9B0F7673A152FBD6FA63791B70B6" box="[1091,1147,1442,1468]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="5" pageNumber="337" unit="in" value="2017.0">2017</quantity>
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having homogenously sized spines on a regularly domed tail shield (versus a less prominent and more-irregularly shaped tail shield with heterogeneously sized spines).
|
|||
|
|
<taxonomicName id="5AD44D697673A152FD96FA2B7FAB730E" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[515,715,1514,1540]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FD96FA2B7FAB730E" box="[515,715,1514,1540]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FD42FA2B7E55730E" bold="true" box="[727,821,1514,1540]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FD42FA2B7E55730E" box="[727,821,1514,1540]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="5AD44D697673A152FC4DFA2A78F7730E" authority="Deraniyagala, 1941" authorityName="Deraniyagala" authorityYear="1941" box="[984,1431,1514,1540]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="dorsimaculatus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FC4DFA2A79C9730E" box="[984,1193,1514,1540]" italics="true" pageId="5" pageNumber="337">R. dorsimaculatus</emphasis>
|
|||
|
|
Deraniyagala, 1941
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697673A152FF02F9CE7F727322" authority="(Schneider, 1801)" baseAuthorityName="Schneider" baseAuthorityYear="1801" box="[151,530,1550,1576]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="oxyrynchus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FF02F9CE7C5A7322" box="[151,314,1550,1576]" italics="true" pageId="5" pageNumber="337">R. oxyrynchus</emphasis>
|
|||
|
|
(Schneider, 1801)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697673A152FDB7F9CE7E5B7322" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" box="[546,827,1550,1576]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="porrectus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FDB7F9CE7FD07322" box="[546,688,1551,1576]" italics="true" pageId="5" pageNumber="337">R. porrectus</emphasis>
|
|||
|
|
Wall, 1921
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697673A152FCDEF9CE791B7322" authority="Muller, 1832" authorityName="Muller" authorityYear="1832" box="[843,1147,1550,1576]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="punctatus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FCDEF9CE7EBB7322" box="[843,987,1551,1576]" italics="true" pageId="5" pageNumber="337">R. punctatus</emphasis>
|
|||
|
|
Müller, 1832
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="5AD44D697673A152FB2CF9CE7C3D7346" authority="Gower & Maduwage, 2011" authorityName="Gower & Maduwage" authorityYear="2011" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="zigzag">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FB2CF9CE78457322" box="[1209,1317,1551,1576]" italics="true" pageId="5" pageNumber="337">R. zigzag</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7673A152FABAF9CF7C3D7346" author="Gower, D. J. & Maduwage, K." pageId="5" pageNumber="337" pagination="51 - 68" refId="ref16404" refString="Gower, D. J. & Maduwage, K. (2011) Two new species of Rhinophis Hemprich (Serpentes: Uropeltidae) from Sri Lanka. Zootaxa, 2881 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2881.1.4" type="journal article" year="2011">Gower & Maduwage, 2011</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having fewer than 190 ventral scales (versus more than 205).
|
|||
|
|
<taxonomicName id="5AD44D697673A152FBF9F9F378547346" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[1132,1332,1586,1612]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FBF9F9F378547346" box="[1132,1332,1586,1612]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FAABF9F378FC7346" bold="true" box="[1342,1436,1586,1612]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FAABF9F378FC7346" box="[1342,1436,1586,1612]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="5AD44D697673A152FEB0F9967CA8737A" authorityName="Deraniyagala" authorityYear="1975" box="[293,456,1622,1648]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="tricoloratus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FEB0F9967CA8737A" box="[293,456,1622,1648]" italics="true" pageId="5" pageNumber="337">R. tricoloratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="5AD44D697673A152FE6AF9967DAF739E" authority="Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella, 2009" authorityName="Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella" authorityYear="2009" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="erangarviraji">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FE6AF9967FD5737A" box="[511,693,1623,1648]" italics="true" pageId="5" pageNumber="337">R. erangarviraji</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7673A152FD29F9977DAF739E" author="Wickramasinghe, L. J. M. & Vidanapathirana, D. R. & Wickramasinghe, N. & Ranwella, P. N." pageId="5" pageNumber="337" pagination="1 - 22" refId="ref17090" refString="Wickramasinghe, L. J. M., Vidanapathirana, D. R., Wickramasinghe, N. & Ranwella, P. N. (2009) A new species of Rhinophis Hemprich, 1820 (Reptilia: Serpentes: Uropeltidae) from Rakwana massif, Sri Lanka. Zootaxa, 2044 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2044.1.1" type="journal article" year="2009">
|
|||
|
|
Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella,
|
|||
|
|
<quantity id="5A2C9B0F7673A152FF02F9BB7DAF739E" box="[151,207,1658,1684]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="5" pageNumber="337" unit="in" value="2009.0">2009</quantity>
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having more than 166 ventral scales (versus fewer than 164).
|
|||
|
|
<taxonomicName id="5AD44D697673A152FC24F9BB7915739E" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[945,1141,1658,1684]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FC24F9BB7915739E" box="[945,1141,1658,1684]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FBE8F9BB79B8739E" bold="true" box="[1149,1240,1658,1684]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FBE8F9BB79B8739E" box="[1149,1240,1658,1684]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
has more ventral scales (ca.
|
|||
|
|
<quantity id="5A2C9B0F7673A152FE80F95F7CF673B2" box="[277,406,1694,1720]" metricMagnitude="0" metricUnit="m" metricValue="4.3815" metricValueMax="4.5212" metricValueMin="4.2418" pageId="5" pageNumber="337" unit="in" value="172.5" valueMax="178.0" valueMin="167.0">167–178 in</quantity>
|
|||
|
|
males,
|
|||
|
|
<quantity id="5A2C9B0F7673A152FE65F95F7F1173B2" box="[496,625,1694,1721]" metricMagnitude="0" metricUnit="m" metricValue="4.5847" metricValueMax="4.7244" metricValueMin="4.445" pageId="5" pageNumber="337" unit="in" value="180.5" valueMax="186.0" valueMin="175.0">175–186 in</quantity>
|
|||
|
|
females) than both
|
|||
|
|
<taxonomicName id="5AD44D697673A152FCC2F95E793C73B3" authority="Kelaart 1853" authorityName="Kelaart" authorityYear="1853" box="[855,1116,1694,1721]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="blythii">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FCC2F95E7EA073B2" box="[855,960,1694,1720]" italics="true" pageId="5" pageNumber="337">R. blythii</emphasis>
|
|||
|
|
Kelaart 1853
|
|||
|
|
</taxonomicName>
|
|||
|
|
(ca. 148–158 and 155–167, respectively) and
|
|||
|
|
<taxonomicName id="5AD44D697673A152FEF5F9027C9E73D6" baseAuthorityName="Cuvier" baseAuthorityYear="1829" box="[352,510,1730,1756]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="philippinus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FEF5F9027C9E73D6" box="[352,510,1730,1756]" italics="true" pageId="5" pageNumber="337">R. philippinus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(154–168 and 164–179, respectively), and additionally differs from the former in having a more protuberant, conical (versus less-prominently domed) tail shield and from the latter in having a paler orange-bronze dorsum (versus darker, brown-grey) and having (versus lacking) substantial yellow-orange markings on the underside of the tail.
|
|||
|
|
<taxonomicName id="5AD44D697673A152FE4BF8EF7FC47242" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[478,676,1838,1864]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FE4BF8EF7FC47242" box="[478,676,1838,1864]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FD38F8EF7E687242" bold="true" box="[685,776,1838,1864]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FD38F8EF7E687242" box="[685,776,1838,1864]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
has fewer ventral scales (ca.
|
|||
|
|
<quantity id="5A2C9B0F7673A152FBCBF8EF79807242" box="[1118,1248,1838,1864]" metricMagnitude="0" metricUnit="m" metricValue="4.4069" metricValueMax="4.5212" metricValueMin="4.2926" pageId="5" pageNumber="337" unit="in" value="173.5" valueMax="178.0" valueMin="169.0">169–178 in</quantity>
|
|||
|
|
males,
|
|||
|
|
<quantity id="5A2C9B0F7673A152FAAFF8EF7DCD7266" metricMagnitude="0" metricUnit="m" metricValue="4.5847" metricValueMax="4.7244" metricValueMin="4.445" pageId="5" pageNumber="337" unit="in" value="180.5" valueMax="186.0" valueMin="175.0">175–186 in</quantity>
|
|||
|
|
females) than
|
|||
|
|
<taxonomicName id="5AD44D697673A152FECCF8927E737266" authority="Gower & Maduwage, 2011" authorityName="Gower & Maduwage" authorityYear="2011" box="[345,787,1874,1900]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="lineatus">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FECCF8927C0A7266" box="[345,362,1875,1900]" italics="true" pageId="5" pageNumber="337">R</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="AFA0EAF87673A152FEEFF8937CB37266" box="[378,467,1874,1900]" italics="true" pageId="5" pageNumber="337">lineatus</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7673A152FE4EF8937E737266" author="Gower, D. J. & Maduwage, K." box="[475,787,1874,1900]" pageId="5" pageNumber="337" pagination="51 - 68" refId="ref16404" refString="Gower, D. J. & Maduwage, K. (2011) Two new species of Rhinophis Hemprich (Serpentes: Uropeltidae) from Sri Lanka. Zootaxa, 2881 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2881.1.4" type="journal article" year="2011">Gower & Maduwage, 2011</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(183–190 and up to 195, respectively), and has a darker ventral than dorsal surface (versus vice versa).
|
|||
|
|
<taxonomicName id="5AD44D697673A152FD39F8B77E10729A" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[684,880,1910,1936]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FD39F8B77E10729A" box="[684,880,1910,1936]" italics="true" pageId="5" pageNumber="337">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87673A152FCE2F8B77EB2729A" bold="true" box="[887,978,1910,1936]" pageId="5" pageNumber="337">
|
|||
|
|
<taxonomicNameLabel id="B49357837673A152FCE2F8B77EB2729A" box="[887,978,1910,1936]" pageId="5" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="5AD44D697673A152FBFDF8B67818729A" authority="Gower, 2020" authorityName="Gower" authorityYear="2020" box="[1128,1400,1910,1936]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="5" pageNumber="337" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF87673A152FBFDF8B679BF729A" box="[1128,1247,1910,1936]" italics="true" pageId="5" pageNumber="337">R. mendisi</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7673A152FB72F8B77818729A" author="Gower, D. J." box="[1255,1400,1910,1936]" pageId="5" pageNumber="337" pagination="495 - 510" refId="ref16321" refString="Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from southwestern Sri Lanka. Zootaxa, 4810 (3), 495 - 510. https: // doi. org / 10.11646 / zootaxa. 4810.3.6" type="journal article" year="2020">Gower, 2020</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having a more rounded than ridged dorsal apex to the rostral shield (in transverse section), having more ventrals (ca.
|
|||
|
|
<quantity id="5A2C9B0F7673A152FF02F87F7C7672D2" box="[151,278,1982,2008]" metricMagnitude="0" metricUnit="m" metricValue="4.3815" metricValueMax="4.5212" metricValueMin="4.2418" pageId="5" pageNumber="337" unit="in" value="172.5" valueMax="178.0" valueMin="167.0">167–178 in</quantity>
|
|||
|
|
males,
|
|||
|
|
<quantity id="5A2C9B0F7673A152FEF9F87F7C8B72D2" box="[364,491,1982,2009]" metricMagnitude="0" metricUnit="m" metricValue="4.5847" metricValueMax="4.7244" metricValueMin="4.445" pageId="5" pageNumber="337" unit="in" value="180.5" valueMax="186.0" valueMin="175.0">175–186 in</quantity>
|
|||
|
|
females versus 159–177), males typically having more subcaudals (at least 6 on one side to 7 [very rarely 5], versus 4–5 on each side), and having a darker venter than dorsum (versus darker dorsum than venter). The new species differs from
|
|||
|
|
<taxonomicName id="5AD44D697670A151FD14FF7A7E8675DF" authority="(Hemprich, 1820)" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[641,998,187,213]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FD14FF7A7E7475DF" box="[641,788,187,213]" italics="true" pageId="6" pageNumber="338">R. homolepis</emphasis>
|
|||
|
|
(Hemprich, 1820)
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having fewer ventrals (<187 versus>191: see
|
|||
|
|
<bibRefCitation id="F9454B1B7670A151FE9FFF1E7CFA75F3" author="Gower, D. J." box="[266,410,223,249]" pageId="6" pageNumber="338" pagination="495 - 510" refId="ref16321" refString="Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from southwestern Sri Lanka. Zootaxa, 4810 (3), 495 - 510. https: // doi. org / 10.11646 / zootaxa. 4810.3.6" type="journal article" year="2020">Gower 2020</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="5AD44D697670A151FE38FF1E7F1075F2" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[429,624,223,249]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FE38FF1E7F1075F2" box="[429,624,223,249]" italics="true" pageId="6" pageNumber="338">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87670A151FDE2FF1E7FB075F3" bold="true" box="[631,720,223,249]" pageId="6" pageNumber="338">
|
|||
|
|
<taxonomicNameLabel id="B49357837670A151FDE2FF1E7FB075F3" box="[631,720,223,249]" pageId="6" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="5AD44D697670A151FCF1FF1E7F727417" authority="Wickramasinghe, Vidanapathirana, Wickramasinghe & Gower, 2020" authorityName="Wickramasinghe, Vidanapathirana, Wickramasinghe & Gower" authorityYear="2020" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="gunasekarai">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FCF1FF1E796D75F3" box="[868,1037,223,249]" italics="true" pageId="6" pageNumber="338">R. gunasekarai</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B7670A151FB81FF1E7F727417" author="Wickramasinghe, L. M. & Vidanapathirana, D. R. & Wickramasinghe, N. & Gower, D. J." pageId="6" pageNumber="338" pagination="65 - 80" refId="ref17239" refString="Wickramasinghe, L. M., Vidanapathirana, D. R., Wickramasinghe, N. & Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Reptilia: Uropeltidae), from cloud forest of the Knuckles massif of Sri Lanka. Zootaxa, 4810 (1), 65 - 80. https: // doi. org / 10.11646 / zootaxa. 4810.1.3" type="journal article" year="2020">
|
|||
|
|
Wickramasinghe, Vidanapathirana, Wickramasinghe & Gower,
|
|||
|
|
<quantity id="5A2C9B0F7670A151FE4FFEC27F727417" box="[474,530,259,285]" metricMagnitude="1" metricUnit="m" metricValue="5.1308" pageId="6" pageNumber="338" unit="in" value="2020.0">2020</quantity>
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having a much broader tail shield relative to tail width, in lacking narrow, continuous longitudinal yellow lines on the dorsum, and in having pale markings on the underside of the tail.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7670A151FF52FE8A79A4778F" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="338">
|
|||
|
|
The ventral and subcaudal counts in the new species overlap substantially with those for
|
|||
|
|
<taxonomicName id="5AD44D697670A151FB00FE8A7DAF7483" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="drummondhayi">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FB00FE8A783D746F" box="[1173,1373,331,357]" italics="true" pageId="6" pageNumber="338">R. drummondhayi</emphasis>
|
|||
|
|
Wall, 1921
|
|||
|
|
</taxonomicName>
|
|||
|
|
(although with slightly higher ranges), and both have a relatively broad tail shield; however the two species differ clearly in colour pattern (paler dorsum than venter in the new species versus the reverse in
|
|||
|
|
<taxonomicName id="5AD44D697670A151FB51FE5278EC74A7" authorityName="Wall" authorityYear="1921" box="[1220,1420,403,429]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="drummondhayi">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FB51FE5278EC74A7" box="[1220,1420,403,429]" italics="true" pageId="6" pageNumber="338">R. drummondhayi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), in distribution (Rakwana Massif in the new species versus Central Highlands in
|
|||
|
|
<taxonomicName id="5AD44D697670A151FB8CFE76798274DB" authorityName="Wall" authorityYear="1921" box="[1049,1250,439,465]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="drummondhayi">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FB8CFE76798274DB" box="[1049,1250,439,465]" italics="true" pageId="6" pageNumber="338">R. drummondhayi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), in the shape of the tail shield (more prominent and conical in
|
|||
|
|
<taxonomicName id="5AD44D697670A151FD3DFE1A7E7474FE" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[680,788,475,500]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FD3DFE1A7E7474FE" box="[680,788,475,500]" italics="true" pageId="6" pageNumber="338">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87670A151FC89FE1A7E1774FF" bold="true" box="[796,887,475,501]" pageId="6" pageNumber="338">
|
|||
|
|
<taxonomicNameLabel id="B49357837670A151FC89FE1A7E1774FF" box="[796,887,475,501]" pageId="6" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
), and in DNA sequence data (with phylogenetic analysis of these data providing maximal support for
|
|||
|
|
<taxonomicName id="5AD44D697670A151FD78FE3E7E377712" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[749,855,511,536]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FD78FE3E7E377712" box="[749,855,511,536]" italics="true" pageId="6" pageNumber="338">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87670A151FCC8FE3E7ED77713" bold="true" box="[861,951,511,537]" pageId="6" pageNumber="338">
|
|||
|
|
<taxonomicNameLabel id="B49357837670A151FCC8FE3E7ED77713" box="[861,951,511,537]" pageId="6" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
being more closely related to
|
|||
|
|
<taxonomicName id="5AD44D697670A151FA9FFE3E78FC7713" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[1290,1436,511,537]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FA9FFE3E78FC7713" box="[1290,1436,511,537]" italics="true" pageId="6" pageNumber="338">R. homolepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="5AD44D697670A151FF53FDE27C087737" authorityName="Deraniyagala" authorityYear="1975" box="[198,360,547,573]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="tricoloratus">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FF53FDE27C087737" box="[198,360,547,573]" italics="true" pageId="6" pageNumber="338">R. tricoloratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
versus
|
|||
|
|
<taxonomicName id="5AD44D697670A151FE28FDE27FE57737" authorityName="Wall" authorityYear="1921" box="[445,645,547,573]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="drummondhayi">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FE28FDE27FE57737" box="[445,645,547,573]" italics="true" pageId="6" pageNumber="338">R. drummondhayi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
being more closely related to
|
|||
|
|
<taxonomicName id="5AD44D697670A151FC46FDE279527737" authorityName="Kelaart" authorityYear="1853" box="[979,1074,547,573]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="blythi">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FC46FDE279527737" box="[979,1074,547,573]" italics="true" pageId="6" pageNumber="338">R. blythi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697670A151FBAAFDE279897736" authorityName="Wickramasinghe, Vidanapathirana, Wickramasinghe & Ranwella" authorityYear="2009" box="[1087,1257,547,572]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="erangaviraji">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FBAAFDE279897736" box="[1087,1257,547,572]" italics="true" pageId="6" pageNumber="338">R. erangaviraji</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D697670A151FB63FDE2780E7737" authorityName="Gower & Maduwage" authorityYear="2011" box="[1270,1390,547,573]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="lineatus">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FB63FDE2780E7737" box="[1270,1390,547,573]" italics="true" pageId="6" pageNumber="338">R. lineatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="5AD44D697670A151FF02FD867D9F776A" authorityName="Gower & Maduwage" authorityYear="2011" box="[151,255,583,608]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="zigzag">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FF02FD867D9F776A" box="[151,255,583,608]" italics="true" pageId="6" pageNumber="338">R. zigzag</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
:
|
|||
|
|
<bibRefCitation id="F9454B1B7670A151FE9BFD867C96776B" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." box="[270,502,583,609]" pageId="6" pageNumber="338" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
|||
|
|
Sampaio
|
|||
|
|
<emphasis id="AFA0EAF87670A151FEEDFD867CD1776B" box="[376,433,583,609]" italics="true" pageId="6" pageNumber="338">et al.</emphasis>
|
|||
|
|
2023
|
|||
|
|
</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="5AD44D697670A151FD9CFD867FAD776A" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[521,717,583,609]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FD9CFD867FAD776A" box="[521,717,583,609]" italics="true" pageId="6" pageNumber="338">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87670A151FD41FD867E4F776B" bold="true" box="[724,815,583,609]" pageId="6" pageNumber="338">
|
|||
|
|
<taxonomicNameLabel id="B49357837670A151FD41FD867E4F776B" box="[724,815,583,609]" pageId="6" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
also differs notably from all other named congeners in DNA-sequence data (
|
|||
|
|
<tableCitation id="D05603517670A151FE1DFDAA7CBF778F" box="[392,479,619,645]" captionStart="TABLE 3" captionStartId="8.[151,239,368,392]" captionTargetPageId="8" captionText="TABLE 3. Uncorrected p-distances (%) for intra- and interspecific comparisons for Rhinophis martin sp. nov. and closest sampled relatives, for two nuclear (prlr, cmos) and four mitochondrial genes (12s, 16s, cytb, nd4) as analysed by Sampaio et al. (2023). Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison." pageId="6" pageNumber="338">Table 3</tableCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="F9454B1B7670A151FE7EFDAA7FAF778F" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." box="[491,719,619,645]" pageId="6" pageNumber="338" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
|||
|
|
Sampaio
|
|||
|
|
<emphasis id="AFA0EAF87670A151FDC1FDAA7FED778F" box="[596,653,619,645]" italics="true" pageId="6" pageNumber="338">et al.</emphasis>
|
|||
|
|
2023
|
|||
|
|
</bibRefCitation>
|
|||
|
|
), as discussed in the Remarks section below.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="C9AB66627670A151FF02FD757CD5761E" pageId="6" pageNumber="338" startId="6.[151,239,692,716]" targetBox="[163,1405,805,1714]" targetIsTable="true" targetPageId="6" targetType="table">
|
|||
|
|
<paragraph id="9D6B36EA7670A151FF02FD757CD5761E" blockId="6.[151,1437,692,788]" pageId="6" pageNumber="338">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FF02FD757C6C77C6" bold="true" box="[151,268,692,716]" pageId="6" pageNumber="338">TABLE 2.</emphasis>
|
|||
|
|
Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of
|
|||
|
|
<taxonomicName id="5AD44D697670A151FAA5FD757DBE77FA" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="6" pageNumber="338" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FAA5FD757DBE77FA" italics="true" pageId="6" pageNumber="338">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF87670A151FF70FD187C5A77FB" bold="true" box="[229,314,729,753]" pageId="6" pageNumber="338">
|
|||
|
|
<taxonomicNameLabel id="B49357837670A151FF70FD187C5A77FB" box="[229,314,729,753]" pageId="6" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="9D6B36EA7670A151FF36FCE4782973B8" pageId="6" pageNumber="338">
|
|||
|
|
<table id="EFD4C44A76705EA8FF36FCE4781D73B8" box="[163,1405,805,1714]" colsContinueIn="19.[163,1405,264,1978]" gridcols="6" gridrows="23" pageId="6" pageNumber="338">
|
|||
|
|
<tr id="23E434A876705EA8FF36FCE4781D7637" box="[163,1405,805,829]" gridrow="0" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FCE47C5E7637" box="[163,318,805,829]" gridcol="0" gridrow="0" pageId="6" pageNumber="338">Specimen</th>
|
|||
|
|
<th id="60355DD476705EA8FE5CFCE47C8F7637" box="[457,495,805,829]" gridcol="1" gridrow="0" pageId="6" pageNumber="338">Sex</th>
|
|||
|
|
<th id="60355DD476705EA8FD04FCE47FD77637" box="[657,695,805,829]" gridcol="2" gridrow="0" pageId="6" pageNumber="338">TL</th>
|
|||
|
|
<th id="60355DD476705EA8FCF7FCE47EE27637" box="[866,898,805,829]" gridcol="3" gridrow="0" pageId="6" pageNumber="338">tL</th>
|
|||
|
|
<th id="60355DD476705EA8FB95FCE479EB7637" box="[1024,1163,805,829]" gridcol="4" gridrow="0" pageId="6" pageNumber="338">Ventral scales</th>
|
|||
|
|
<th id="60355DD476705EA8FB44FCE4781D7637" box="[1233,1405,805,829]" gridcol="5" gridrow="0" pageId="6" pageNumber="338">Subcaudal scales</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FC8C781D766F" box="[163,1405,845,869]" gridrow="1" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FC8C7C5E766F" box="[163,318,845,869]" gridcol="0" gridrow="1" pageId="6" pageNumber="338">USNM 548096</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFC8C7C8F766F" box="[457,495,845,869]" gridcol="1" gridrow="1" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FC8C7FD7766F" box="[657,695,845,869]" gridcol="2" gridrow="1" pageId="6" pageNumber="338">229</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FC8C7EE2766F" box="[866,898,845,869]" gridcol="3" gridrow="1" pageId="6" pageNumber="338">9.4</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FC8C79EB766F" box="[1024,1163,845,869]" gridcol="4" gridrow="1" pageId="6" pageNumber="338">177</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FC8C781D766F" box="[1233,1405,845,869]" gridcol="5" gridrow="1" pageId="6" pageNumber="338">7,6 (0)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FCB4781D7687" box="[163,1405,885,909]" gridrow="2" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FCB47C5E7687" box="[163,318,885,909]" gridcol="0" gridrow="2" pageId="6" pageNumber="338">USNM 548097</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFCB47C8F7687" box="[457,495,885,909]" gridcol="1" gridrow="2" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FCB47FD77687" box="[657,695,885,909]" gridcol="2" gridrow="2" pageId="6" pageNumber="338">195</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FCB47EE27687" box="[866,898,885,909]" gridcol="3" gridrow="2" pageId="6" pageNumber="338">8.1</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FCB479EB7687" box="[1024,1163,885,909]" gridcol="4" gridrow="2" pageId="6" pageNumber="338">173</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FCB4781D7687" box="[1233,1405,885,909]" gridcol="5" gridrow="2" pageId="6" pageNumber="338">7,7 (5)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FC5C781D76BC" box="[163,1405,925,950]" gridrow="3" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FC5C7C5E76BC" box="[163,318,925,950]" gridcol="0" gridrow="3" pageId="6" pageNumber="338">USNM 548098</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFC5C7C8F76BC" box="[457,495,925,950]" gridcol="1" gridrow="3" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FC5C7FD776BC" box="[657,695,925,950]" gridcol="2" gridrow="3" pageId="6" pageNumber="338">213</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FC5C7EE276BC" box="[866,898,925,950]" gridcol="3" gridrow="3" pageId="6" pageNumber="338">7.8</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FC5C79EB76BC" box="[1024,1163,925,950]" gridcol="4" gridrow="3" pageId="6" pageNumber="338">174</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FC5C781D76BC" box="[1233,1405,925,950]" gridcol="5" gridrow="3" pageId="6" pageNumber="338">6,5 (1)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FC04781D76D4" box="[163,1405,965,990]" gridrow="4" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FC047C5E76D4" box="[163,318,965,990]" gridcol="0" gridrow="4" pageId="6" pageNumber="338">USNM 548099</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFC047C8F76D4" box="[457,495,965,990]" gridcol="1" gridrow="4" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FC047FD776D4" box="[657,695,965,990]" gridcol="2" gridrow="4" pageId="6" pageNumber="338">241</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FC047EE276D4" box="[866,898,965,990]" gridcol="3" gridrow="4" pageId="6" pageNumber="338">5.8</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FC0479EB76D4" box="[1024,1163,965,990]" gridcol="4" gridrow="4" pageId="6" pageNumber="338">182</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FC04781D76D4" box="[1233,1405,965,990]" gridcol="5" gridrow="4" pageId="6" pageNumber="338">4,4 (1)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FC2F781D710C" box="[163,1405,1006,1030]" gridrow="5" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FC2F7C5E710C" box="[163,318,1006,1030]" gridcol="0" gridrow="5" pageId="6" pageNumber="338">USNM 548101</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFC2F7C8F710C" box="[457,495,1006,1030]" gridcol="1" gridrow="5" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FC2F7FD7710C" box="[657,695,1006,1030]" gridcol="2" gridrow="5" pageId="6" pageNumber="338">173</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FC2F7EE2710C" box="[866,898,1006,1030]" gridcol="3" gridrow="5" pageId="6" pageNumber="338">6.0</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FC2F79EB710C" box="[1024,1163,1006,1030]" gridcol="4" gridrow="5" pageId="6" pageNumber="338">177</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FC2F781D710C" box="[1233,1405,1006,1030]" gridcol="5" gridrow="5" pageId="6" pageNumber="338">6,6 (2)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FBD7781D7124" box="[163,1405,1046,1070]" gridrow="6" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FBD77C5E7124" box="[163,318,1046,1070]" gridcol="0" gridrow="6" pageId="6" pageNumber="338">USNM 548102</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFBD77C8F7124" box="[457,495,1046,1070]" gridcol="1" gridrow="6" pageId="6" pageNumber="338">F?</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FBD77FD77124" box="[657,695,1046,1070]" gridcol="2" gridrow="6" pageId="6" pageNumber="338">254</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FBD77EE27124" box="[866,898,1046,1070]" gridcol="3" gridrow="6" pageId="6" pageNumber="338">6.0</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FBD779EB7124" box="[1024,1163,1046,1070]" gridcol="4" gridrow="6" pageId="6" pageNumber="338">–</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FBD7781D7124" box="[1233,1405,1046,1070]" gridcol="5" gridrow="6" pageId="6" pageNumber="338">–</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FBFF781D715C" box="[163,1405,1086,1110]" gridrow="7" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FBFF7C5E715C" box="[163,318,1086,1110]" gridcol="0" gridrow="7" pageId="6" pageNumber="338">USNM 548103</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFBFF7C8F715C" box="[457,495,1086,1110]" gridcol="1" gridrow="7" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FBFF7FD7715C" box="[657,695,1086,1110]" gridcol="2" gridrow="7" pageId="6" pageNumber="338">259</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FBFF7EE2715C" box="[866,898,1086,1110]" gridcol="3" gridrow="7" pageId="6" pageNumber="338">5.9</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FBFF79EB715C" box="[1024,1163,1086,1110]" gridcol="4" gridrow="7" pageId="6" pageNumber="338">186</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FBFF781D715C" box="[1233,1405,1086,1110]" gridcol="5" gridrow="7" pageId="6" pageNumber="338">4,4 (2)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FBA7781D7175" box="[163,1405,1126,1151]" gridrow="8" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FBA77C5E7175" box="[163,318,1126,1151]" gridcol="0" gridrow="8" pageId="6" pageNumber="338">USNM 548104</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFBA77C8F7175" box="[457,495,1126,1151]" gridcol="1" gridrow="8" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FBA77FD77175" box="[657,695,1126,1151]" gridcol="2" gridrow="8" pageId="6" pageNumber="338">242</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FBA77EE27175" box="[866,898,1126,1151]" gridcol="3" gridrow="8" pageId="6" pageNumber="338">9.8</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FBA779EB7175" box="[1024,1163,1126,1151]" gridcol="4" gridrow="8" pageId="6" pageNumber="338">178</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FBA7781D7175" box="[1233,1405,1126,1151]" gridcol="5" gridrow="8" pageId="6" pageNumber="338">6,5 (1)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FB4E781D71AD" box="[163,1405,1167,1191]" gridrow="9" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FB4E7C5E71AD" box="[163,318,1167,1191]" gridcol="0" gridrow="9" pageId="6" pageNumber="338">USNM 548105</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFB4E7C8F71AD" box="[457,495,1167,1191]" gridcol="1" gridrow="9" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FB4E7FD771AD" box="[657,695,1167,1191]" gridcol="2" gridrow="9" pageId="6" pageNumber="338">199</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FB4E7EE271AD" box="[866,898,1167,1191]" gridcol="3" gridrow="9" pageId="6" pageNumber="338">5.2</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FB4E79EB71AD" box="[1024,1163,1167,1191]" gridcol="4" gridrow="9" pageId="6" pageNumber="338">183</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FB4E781D71AD" box="[1233,1405,1167,1191]" gridcol="5" gridrow="9" pageId="6" pageNumber="338">4,3 (0)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FB76781D71C5" box="[163,1405,1207,1231]" gridrow="10" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FB767C5E71C5" box="[163,318,1207,1231]" gridcol="0" gridrow="10" pageId="6" pageNumber="338">USNM 548106</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFB767C8F71C5" box="[457,495,1207,1231]" gridcol="1" gridrow="10" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FB767FD771C5" box="[657,695,1207,1231]" gridcol="2" gridrow="10" pageId="6" pageNumber="338">228</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FB767EE271C5" box="[866,898,1207,1231]" gridcol="3" gridrow="10" pageId="6" pageNumber="338">8.1</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FB7679EB71C5" box="[1024,1163,1207,1231]" gridcol="4" gridrow="10" pageId="6" pageNumber="338">169</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FB76781D71C5" box="[1233,1405,1207,1231]" gridcol="5" gridrow="10" pageId="6" pageNumber="338">5,5 (3)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FB1E781D71FD" box="[163,1405,1247,1271]" gridrow="11" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FB1E7C5E71FD" box="[163,318,1247,1271]" gridcol="0" gridrow="11" pageId="6" pageNumber="338">USNM 548107</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFB1E7C8F71FD" box="[457,495,1247,1271]" gridcol="1" gridrow="11" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FB1E7FD771FD" box="[657,695,1247,1271]" gridcol="2" gridrow="11" pageId="6" pageNumber="338">192</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FB1E7EE271FD" box="[866,898,1247,1271]" gridcol="3" gridrow="11" pageId="6" pageNumber="338">4.7</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FB1E79EB71FD" box="[1024,1163,1247,1271]" gridcol="4" gridrow="11" pageId="6" pageNumber="338">185</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FB1E781D71FD" box="[1233,1405,1247,1271]" gridcol="5" gridrow="11" pageId="6" pageNumber="338">3,2 (0)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FAC6781D702A" box="[163,1405,1287,1312]" gridrow="12" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FAC67C5E702A" box="[163,318,1287,1312]" gridcol="0" gridrow="12" pageId="6" pageNumber="338">USNM 548108</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFAC67C8F702A" box="[457,495,1287,1312]" gridcol="1" gridrow="12" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FAC67FD7702A" box="[657,695,1287,1312]" gridcol="2" gridrow="12" pageId="6" pageNumber="338">175</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FAC67EE2702A" box="[866,898,1287,1312]" gridcol="3" gridrow="12" pageId="6" pageNumber="338">7.7</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FAC679EB702A" box="[1024,1163,1287,1312]" gridcol="4" gridrow="12" pageId="6" pageNumber="338">170</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FAC6781D702A" box="[1233,1405,1287,1312]" gridcol="5" gridrow="12" pageId="6" pageNumber="338">7,7 (6)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FAF1781D7042" box="[163,1405,1328,1352]" gridrow="13" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FAF17C5E7042" box="[163,318,1328,1352]" gridcol="0" gridrow="13" pageId="6" pageNumber="338">USNM 548109</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFAF17C8F7042" box="[457,495,1328,1352]" gridcol="1" gridrow="13" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FAF17FD77042" box="[657,695,1328,1352]" gridcol="2" gridrow="13" pageId="6" pageNumber="338">194</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FAF17EE27042" box="[866,898,1328,1352]" gridcol="3" gridrow="13" pageId="6" pageNumber="338">5.0</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FAF179EB7042" box="[1024,1163,1328,1352]" gridcol="4" gridrow="13" pageId="6" pageNumber="338">183</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FAF1781D7042" box="[1233,1405,1328,1352]" gridcol="5" gridrow="13" pageId="6" pageNumber="338">4,3 (2)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FA99781D707A" box="[163,1405,1368,1392]" gridrow="14" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FA997C5E707A" box="[163,318,1368,1392]" gridcol="0" gridrow="14" pageId="6" pageNumber="338">USNM 548110</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFA997C8F707A" box="[457,495,1368,1392]" gridcol="1" gridrow="14" pageId="6" pageNumber="338">M?</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FA997FD7707A" box="[657,695,1368,1392]" gridcol="2" gridrow="14" pageId="6" pageNumber="338">–</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FA997EE2707A" box="[866,898,1368,1392]" gridcol="3" gridrow="14" pageId="6" pageNumber="338">–</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FA9979EB707A" box="[1024,1163,1368,1392]" gridcol="4" gridrow="14" pageId="6" pageNumber="338">–</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FA99781D707A" box="[1233,1405,1368,1392]" gridcol="5" gridrow="14" pageId="6" pageNumber="338">6,6 (2)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FA41781D7092" box="[163,1405,1408,1432]" gridrow="15" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FA417C5E7092" box="[163,318,1408,1432]" gridcol="0" gridrow="15" pageId="6" pageNumber="338">USNM 548111</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFA417C8F7092" box="[457,495,1408,1432]" gridcol="1" gridrow="15" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FA417FD77092" box="[657,695,1408,1432]" gridcol="2" gridrow="15" pageId="6" pageNumber="338">258</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FA417EE27092" box="[866,898,1408,1432]" gridcol="3" gridrow="15" pageId="6" pageNumber="338">5.8</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FA4179EB7092" box="[1024,1163,1408,1432]" gridcol="4" gridrow="15" pageId="6" pageNumber="338">186</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FA41781D7092" box="[1233,1405,1408,1432]" gridcol="5" gridrow="15" pageId="6" pageNumber="338">4,4 (0)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FA69781D70CB" box="[163,1405,1448,1473]" gridrow="16" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FA697C5E70CB" box="[163,318,1448,1473]" gridcol="0" gridrow="16" pageId="6" pageNumber="338">USNM 548112</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFA697C8F70CB" box="[457,495,1448,1473]" gridcol="1" gridrow="16" pageId="6" pageNumber="338">M</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FA697FD770CB" box="[657,695,1448,1473]" gridcol="2" gridrow="16" pageId="6" pageNumber="338">245</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FA697EE270CB" box="[866,898,1448,1473]" gridcol="3" gridrow="16" pageId="6" pageNumber="338">9.1</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FA6979EB70CB" box="[1024,1163,1448,1473]" gridcol="4" gridrow="16" pageId="6" pageNumber="338">174</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FA69781D70CB" box="[1233,1405,1448,1473]" gridcol="5" gridrow="16" pageId="6" pageNumber="338">6,7 (1)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FA10781D70E3" box="[163,1405,1489,1513]" gridrow="17" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FA107C5E70E3" box="[163,318,1489,1513]" gridcol="0" gridrow="17" pageId="6" pageNumber="338">USNM 548113</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFA107C8F70E3" box="[457,495,1489,1513]" gridcol="1" gridrow="17" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FA107FD770E3" box="[657,695,1489,1513]" gridcol="2" gridrow="17" pageId="6" pageNumber="338">223</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FA107EE270E3" box="[866,898,1489,1513]" gridcol="3" gridrow="17" pageId="6" pageNumber="338">6.5</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FA1079EB70E3" box="[1024,1163,1489,1513]" gridcol="4" gridrow="17" pageId="6" pageNumber="338">181</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FA10781D70E3" box="[1233,1405,1489,1513]" gridcol="5" gridrow="17" pageId="6" pageNumber="338">4,4 (3)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36FA38781D731B" box="[163,1405,1529,1553]" gridrow="18" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36FA387C5E731B" box="[163,318,1529,1553]" gridcol="0" gridrow="18" pageId="6" pageNumber="338">USNM 548114</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CFA387C8F731B" box="[457,495,1529,1553]" gridcol="1" gridrow="18" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04FA387FD7731B" box="[657,695,1529,1553]" gridcol="2" gridrow="18" pageId="6" pageNumber="338">211</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7FA387EE2731B" box="[866,898,1529,1553]" gridcol="3" gridrow="18" pageId="6" pageNumber="338">5.7</td>
|
|||
|
|
<td id="60355DD476705EA8FB95FA3879EB731B" box="[1024,1163,1529,1553]" gridcol="4" gridrow="18" pageId="6" pageNumber="338">182</td>
|
|||
|
|
<td id="60355DD476705EA8FB44FA38781D731B" box="[1233,1405,1529,1553]" gridcol="5" gridrow="18" pageId="6" pageNumber="338">4,3 (0)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36F9E0781D7333" box="[163,1405,1569,1593]" gridrow="19" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36F9E07C5E7333" box="[163,318,1569,1593]" gridcol="0" gridrow="19" pageId="6" pageNumber="338">USNM 548115</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CF9E07C8F7333" box="[457,495,1569,1593]" gridcol="1" gridrow="19" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04F9E07FD77333" box="[657,695,1569,1593]" gridcol="2" gridrow="19" pageId="6" pageNumber="338">218</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7F9E07EE27333" box="[866,898,1569,1593]" gridcol="3" gridrow="19" pageId="6" pageNumber="338">6.2</td>
|
|||
|
|
<td id="60355DD476705EA8FB95F9E079EB7333" box="[1024,1163,1569,1593]" gridcol="4" gridrow="19" pageId="6" pageNumber="338">179</td>
|
|||
|
|
<td id="60355DD476705EA8FB44F9E0781D7333" box="[1233,1405,1569,1593]" gridcol="5" gridrow="19" pageId="6" pageNumber="338">4,5 (2)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36F988781D7368" box="[163,1405,1609,1634]" gridrow="20" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36F9887C5E7368" box="[163,318,1609,1634]" gridcol="0" gridrow="20" pageId="6" pageNumber="338">USNM 548116</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CF9887C8F7368" box="[457,495,1609,1634]" gridcol="1" gridrow="20" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04F9887FD77368" box="[657,695,1609,1634]" gridcol="2" gridrow="20" pageId="6" pageNumber="338">209</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7F9887EE27368" box="[866,898,1609,1634]" gridcol="3" gridrow="20" pageId="6" pageNumber="338">5.5</td>
|
|||
|
|
<td id="60355DD476705EA8FB95F98879EB7368" box="[1024,1163,1609,1634]" gridcol="4" gridrow="20" pageId="6" pageNumber="338">184</td>
|
|||
|
|
<td id="60355DD476705EA8FB44F988781D7368" box="[1233,1405,1609,1634]" gridcol="5" gridrow="20" pageId="6" pageNumber="338">4,4 (1)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36F9B3781D7380" box="[163,1405,1650,1674]" gridrow="21" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36F9B37C5E7380" box="[163,318,1650,1674]" gridcol="0" gridrow="21" pageId="6" pageNumber="338">USNM 548124</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CF9B37C8F7380" box="[457,495,1650,1674]" gridcol="1" gridrow="21" pageId="6" pageNumber="338">F</td>
|
|||
|
|
<td id="60355DD476705EA8FD04F9B37FD77380" box="[657,695,1650,1674]" gridcol="2" gridrow="21" pageId="6" pageNumber="338">221</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7F9B37EE27380" box="[866,898,1650,1674]" gridcol="3" gridrow="21" pageId="6" pageNumber="338">6.3</td>
|
|||
|
|
<td id="60355DD476705EA8FB95F9B379EB7380" box="[1024,1163,1650,1674]" gridcol="4" gridrow="21" pageId="6" pageNumber="338">181</td>
|
|||
|
|
<td id="60355DD476705EA8FB44F9B3781D7380" box="[1233,1405,1650,1674]" gridcol="5" gridrow="21" pageId="6" pageNumber="338">4,4 (3)</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A876705EA8FF36F95B781D73B8" box="[163,1405,1690,1714]" gridrow="22" pageId="6" pageNumber="338">
|
|||
|
|
<th id="60355DD476705EA8FF36F95B7C5E73B8" box="[163,318,1690,1714]" gridcol="0" gridrow="22" pageId="6" pageNumber="338">USNM 548125</th>
|
|||
|
|
<td id="60355DD476705EA8FE5CF95B7C8F73B8" box="[457,495,1690,1714]" gridcol="1" gridrow="22" pageId="6" pageNumber="338">F?</td>
|
|||
|
|
<td id="60355DD476705EA8FD04F95B7FD773B8" box="[657,695,1690,1714]" gridcol="2" gridrow="22" pageId="6" pageNumber="338">–</td>
|
|||
|
|
<td id="60355DD476705EA8FCF7F95B7EE273B8" box="[866,898,1690,1714]" gridcol="3" gridrow="22" pageId="6" pageNumber="338">10</td>
|
|||
|
|
<td id="60355DD476705EA8FB95F95B79EB73B8" box="[1024,1163,1690,1714]" gridcol="4" gridrow="22" pageId="6" pageNumber="338">–</td>
|
|||
|
|
<td id="60355DD476705EA8FB44F95B781D73B8" box="[1233,1405,1690,1714]" gridcol="5" gridrow="22" pageId="6" pageNumber="338">4,4 (1)</td>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<subSubSection id="D5CE65617670A15CFF52F9207DAF7633" lastPageId="11" lastPageNumber="343" pageId="6" pageNumber="338" type="description">
|
|||
|
|
<paragraph id="9D6B36EA7670A150FF52F92079267713" blockId="6.[151,1437,1761,2040]" lastBlockId="7.[151,1437,151,717]" lastPageId="7" lastPageNumber="339" pageId="6" pageNumber="338">
|
|||
|
|
<emphasis id="AFA0EAF87670A151FF52F9207C8873F6" bold="true" box="[199,488,1761,1788]" pageId="6" pageNumber="338">
|
|||
|
|
Description of
|
|||
|
|
<typeStatus id="426F88487670A151FEEFF9237C8373F6" box="[378,483,1762,1788]" pageId="6" pageNumber="338" type="holotype">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
See
|
|||
|
|
<tableCitation id="D05603517670A151FDB5F9207F1573F6" box="[544,629,1761,1788]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetPageId="5" captionText="TABLE 1. Meristic and morphometric (in mm) data for holotype (*) and paratypes of Rhinophis martin sp. nov. Subcaudal scales data are reported as counts for: left, right (with number of fused pairs in parentheses). Head length = distance between snout tip and posterior end of last supralabial. Head width = distance between posterior edges of last supralabials. “–“ indicates shortest distance between two points. Empty cells indicate data not recorded." pageId="6" pageNumber="338">Table 1</tableCitation>
|
|||
|
|
for morphometric and meristic data, and
|
|||
|
|
<figureCitation id="05EF2A6F7670A151FBC5F92079F373F6" box="[1104,1171,1761,1788]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[228,1358,179,1916]" captionTargetId="figure-18@2.[228,1358,179,1916]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Rhinophis martin sp. nov., USNM 548119. Upper row: head end in right lateral, dorsal, ventral and left lateral views; Central two rows whole specimen in ventral (upper) and dorsal (lower) views); Lower row: tail end in left lateral, ventral, dorsal and right lateral views. Scale bar increments in millimetres. Photographs by H. Taylor." figureDoi="http://doi.org/10.5281/zenodo.11547471" httpUri="https://zenodo.org/record/11547471/files/figure.png" pageId="6" pageNumber="338">Fig. 1</figureCitation>
|
|||
|
|
for photographs. Good condition; few superficial scratches to some scales. Head small, snout pointed. Rostral pointed, longer than wide, without dorsal crest; widest at level of anterior upper corner of first supralabials. Rostral many (>8) times longer (in dorsal view) than rostral-frontal gap. Frontal irregularly hexagonal, longer than wide, lateral (ocular) margins slightly converging posteriorly, posterolateral margins straight to very slightly concave; lateral (ocular) margin shortest, posterolateral edges longest. Frontal much shorter, wider than rostral. Paired nasals separated from each other by posterior half of rostral. External naris small, subcircular, slightly countersunk within small depression, located in anteroventral corner of nasal. Nasal contacts supralabials 1 and 2. Prefrontals only briefly in contact with each other along midline (left overlapping right), separating frontal from rostral. Prefrontals wider than long, shorter than frontal. Supralabials four, first smallest, making the least contribution to margin of mouth; fourth much the largest. Ocular contacts supralabials 3 and 4. Eye distinct, diameter approximately 0.3 times length of ocular, located near anterior corner of ocular, bulging slightly from ocular surface, pupil circular. Paired parietals longer than wide, approximately as long as but much wider than frontal, posteriorly broadly rounded, angle between posteromedial and posterolateral edges approximately 90°. Opposite parietals in brief midline contact (longer than contact between opposite prefrontals), left overlapping right. Each parietal contacts four scales other than head shields. No mental groove; mental approximately as wide as long, smaller than infralabials, contacting first infralabials but not first ventral; three pairs of infralabials, second largest, first smallest. First and second ventrals longer than wide, third approximately as long as wide, fourth and subsequent ventrals wider than long. Seven maxillary and eight mandibular teeth on each side. Teeth simple, pointed, distinctly retrorse, straight, evenly spaced.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FF52FDE27D9877C7" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="339">
|
|||
|
|
Body cylindrical. Body scales generally evenly sized on dorsum and along body except for those involved in dorsal scale row reductions. Midline ventral scales between mental and anal of even size though anteriormost ones gradually narrow. At midbody, ventrals 1.24 times width of scales in first dorsal row. Ventrals 186. Dorsal scale rows 19 anteriorly, reducing to 17 by level with 41
|
|||
|
|
<superScript id="6AA19BA27671A150FD5DFD4E7FB37797" attach="left" box="[712,723,655,669]" fontSize="6" pageId="7" pageNumber="339">st</superScript>
|
|||
|
|
ventral and maintained along most of body; scale row reduction formula:
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FEBAFD3A7CFF761F" blockId="7.[254,464,762,861]" box="[303,415,762,789]" pageId="7" pageNumber="339">4 + 5 (41)</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FF6BFCDE7CB07633" blockId="7.[254,464,762,861]" box="[254,464,799,825]" pageId="7" pageNumber="339">19 --------------- 17</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FEBAFC827CFE7657" blockId="7.[254,464,762,861]" box="[303,414,834,861]" pageId="7" pageNumber="339">4 + 5 (38)</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FF52FC4A7E217007" blockId="7.[151,1437,907,2013]" pageId="7" pageNumber="339">Dorsal scale rows approximately 13 at base of tail. Head and body scales macroscopically smooth, lacking keels. Very faint, low, short keels towards posterior edges of small scales overlapped by anals and on lower dorsal scales of tail. Paired anal scales (right overlying left) considerably larger than posteriormost ventrals and subcaudals. Distal margin of each anal overlaps four (left) and three (right) other scales in addition to anteriormost subcaudals. Three subcaudals, last two undivided. Tail ‘shield’ broadly rounded cone, forming tip of tail, wider than long in dorsal view, many times longer than the frontal in dorsal view (a little shorter than distance between snout tip and back of frontal), visible from below and especially above, base surrounded by 11 scales (including last subcaudal). In posterior view shield circular to very slightly egg-shaped, wider ventrally than dorsally. Shield surface densely spinose, spines small and evenly sized, slightly posteriorly hooked (slightly dorsal upper margin, slightly concave posterior margin in lateral view) arranged in narrowly spaced rows approximately convergent on apex of shield. No notable ridges on scales of the tail and posterior end of body.</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FF52FAD77C4D70CB" blockId="7.[151,1437,907,2013]" pageId="7" pageNumber="339">
|
|||
|
|
<emphasis id="AFA0EAF87671A150FF52FAD77CC1703B" bold="true" box="[199,417,1302,1329]" pageId="7" pageNumber="339">Colour in alcohol.</emphasis>
|
|||
|
|
Tip of rostrum pale orange. Head shields otherwise generally brown with pale tan distal thin margins; a slightly larger, diffuse pale area across posterior of nasals and anterior of prefrontals. Lower edges of supralabials more irregularly and broadly pale. Three dorsally incomplete, narrow (one scale wide, distributed across two scales) whitish transverse bands behind head on dorsum, each band separated by length of four to five dorsal scales.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FF52FA0A79087393" blockId="7.[151,1437,907,2013]" pageId="7" pageNumber="339">
|
|||
|
|
Ventral body colour and pattern more uniform than dorsal. Ventrally brown; paler (pale orange-tan) at head end. Slightly punctate, more so anteriorly, by virtue of each scale having a brown core and pale (mostly latero-) distal edges (the posterior distal edge typically almost transparent). Scale bases and centres mostly uniformly coloured, but flecked in the paler anterior region of the venter. Among ventrals, only 5
|
|||
|
|
<superScript id="6AA19BA27671A150FB9BF9F7797B734E" attach="left" box="[1038,1051,1590,1604]" fontSize="6" pageId="7" pageNumber="339">th</superScript>
|
|||
|
|
and 6
|
|||
|
|
<superScript id="6AA19BA27671A150FBF1F9F77911734E" attach="left" box="[1124,1137,1590,1604]" fontSize="6" pageId="7" pageNumber="339">th</superScript>
|
|||
|
|
are entirely pale, as are a couple of adjacent lower dorsal scale row scales plus the mental and first infralabials. Further posteriorly, occasional ventrals with larger whitish patch; 7
|
|||
|
|
<superScript id="6AA19BA27671A150FDBFF9BF7F577386" attach="left" box="[554,567,1662,1676]" fontSize="6" pageId="7" pageNumber="339">th</superScript>
|
|||
|
|
ventral anterior to anal shields is entirely whitish.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A150FF52F9627C2E729F" blockId="7.[151,1437,907,2013]" pageId="7" pageNumber="339">
|
|||
|
|
Dorsally, anterior ca.
|
|||
|
|
<quantity id="5A2C9B0F7671A150FE2FF9627F6273B7" box="[442,514,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="339" unit="mm" value="30.0">30mm</quantity>
|
|||
|
|
dark brown, then abruptly and increasingly (coppery) orange-brown where scales are darkly flecked scales and have darker latero- and (especially) posterodistal edges. Midvertebral dorsal scale row with faintly indicated, narrow brown line each side. Darker, narrow, subparallel (4–5 scales apart) transverse dorsal bars (not dorsally complete; some asymmetrically arranged) taper upwards from dorsolateral apex of body, only one scale wide at dorsal end. Transition between darker more regular ventral colour and paler dorsal colour is somewhat irregular but typically lies on the 5
|
|||
|
|
<superScript id="6AA19BA27671A150FD8EF8977F48726E" attach="left" box="[539,552,1878,1892]" fontSize="6" pageId="7" pageNumber="339">th</superScript>
|
|||
|
|
dorsal scale row. A few isolated scales along the 5
|
|||
|
|
<superScript id="6AA19BA27671A150FBCAF897790C726E" attach="left" box="[1119,1132,1878,1892]" fontSize="6" pageId="7" pageNumber="339">th</superScript>
|
|||
|
|
dorsal scale row largely or entirely whitish.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA7671A15FFF52F85E7819744B" blockId="7.[151,1437,907,2013]" lastBlockId="8.[151,1437,151,321]" lastPageId="8" lastPageNumber="340" pageId="7" pageNumber="339">Subcaudals and first dorsal scale row mostly blackish with pale (yellowish) laterodistal and translucent posterior margins (more pale edges), together forming a posteriorly tapering, dark midventral tail stripe. Either side of this dark midventral region, the tail has a yellowish lateral tail stripe along its length from the vent to the tail shield, approximately 1.5 dorsal scale rows thick; anteriorly this stripe is much thicker, extending up onto the dorsum to form a transverse, middorsally incomplete pale band. Dark scales on upper and lower surface of tail darker than on body. Tail shield orange-yellow at apex and along ventral and ventrolateral outer edge, with almost symmetrical blackish blotches extending to dorsolateral outer edges; approximately half of shield surface orange-yellow, half blackish.</paragraph>
|
|||
|
|
<caption id="C9AB6662767EA15FFF02FEB17ED474FE" pageId="8" pageNumber="340" startId="8.[151,239,368,392]" targetBox="[163,1396,517,1426]" targetIsTable="true" targetPageId="8" targetType="table">
|
|||
|
|
<paragraph id="9D6B36EA767EA15FFF02FEB17ED474FE" blockId="8.[151,1437,368,501]" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF02FEB17C697482" bold="true" box="[151,265,368,392]" pageId="8" pageNumber="340">TABLE 3.</emphasis>
|
|||
|
|
Uncorrected p-distances (%) for intra- and interspecific comparisons for
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFB9AFEB179A87482" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[1039,1224,368,392]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFB9AFEB179A87482" box="[1039,1224,368,392]" italics="true" pageId="8" pageNumber="340">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFB5BFEB078427483" bold="true" box="[1230,1314,369,393]" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicNameLabel id="B4935783767EA15FFB5BFEB078427483" box="[1230,1314,369,393]" pageId="8" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and closest sampled relatives, for two nuclear (
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFD99FE557F5574A6" box="[524,565,404,428]" italics="true" pageId="8" pageNumber="340">prlr</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFDD7FE547F1974A6" box="[578,633,405,428]" italics="true" pageId="8" pageNumber="340">cmos</emphasis>
|
|||
|
|
) and four mitochondrial genes (
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFC58FE547E9274A7" box="[973,1010,405,429]" italics="true" pageId="8" pageNumber="340">12s</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFC6BFE55794474A6" box="[1022,1060,404,428]" italics="true" pageId="8" pageNumber="340">16s</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFBA5FE55793C74A6" box="[1072,1116,404,428]" italics="true" pageId="8" pageNumber="340">cytb</emphasis>
|
|||
|
|
,
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFBFCFE5579F174A6" box="[1129,1169,404,428]" italics="true" pageId="8" pageNumber="340">nd4</emphasis>
|
|||
|
|
) as analysed by
|
|||
|
|
<bibRefCitation id="F9454B1B767EA15FFAAAFE557C7874DB" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." pageId="8" pageNumber="340" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
|||
|
|
Sampaio
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF02FE787DAC74DA" box="[151,204,440,464]" italics="true" pageId="8" pageNumber="340">et al.</emphasis>
|
|||
|
|
(2023)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
. Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="9D6B36EA767EA15FFE25FDC77C667098" pageId="8" pageNumber="340">
|
|||
|
|
<table id="EFD4C44A767E5EA8FF36FDC478147098" box="[163,1396,517,1426]" gridcols="7" gridrows="24" pageId="8" pageNumber="340">
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FDC478147713" box="[163,1396,517,537]" gridrow="0" pageId="8" pageNumber="340" rowspan-0="1">
|
|||
|
|
<th id="60355DD4767E5EA8FE08FDC47C817713" box="[413,481,517,537]" gridcol="1" gridrow="0" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFE25FDC77CAD7713" box="[432,461,518,537]" italics="true" pageId="8" pageNumber="340">12s</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<th id="60355DD4767E5EA8FE6FFDC47FE77713" box="[506,647,517,537]" gridcol="2" gridrow="0" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFDA7FDC47F2F7712" box="[562,591,517,536]" italics="true" pageId="8" pageNumber="340">16s</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<th id="60355DD4767E5EA8FD0AFDC47E2B7713" box="[671,843,517,537]" gridcol="3" gridrow="0" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFD76FDC47E677712" box="[739,775,517,536]" italics="true" pageId="8" pageNumber="340">cytb</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<th id="60355DD4767E5EA8FCE4FDC479637713" box="[881,1027,517,537]" gridcol="4" gridrow="0" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFC3FFDC47EAA7712" box="[938,970,517,536]" italics="true" pageId="8" pageNumber="340">nd4</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<th id="60355DD4767E5EA8FBA3FDC479A27713" box="[1078,1218,517,537]" gridcol="5" gridrow="0" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFBFEFDC479EC7712" box="[1131,1164,517,536]" italics="true" pageId="8" pageNumber="340">prlr</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<th id="60355DD4767E5EA8FB7DFDC478147713" box="[1256,1396,517,537]" gridcol="6" gridrow="0" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFA8DFDC778247712" box="[1304,1348,518,536]" italics="true" pageId="8" pageNumber="340">cmos</emphasis>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FD817814775C" box="[163,1396,576,598]" gridrow="1" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FD817C14775C" box="[163,372,576,598]" gridcol="0" gridrow="1" pageId="8" pageNumber="340">
|
|||
|
|
Within
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF7FFD807C25775C" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[234,325,577,598]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="martin">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF7FFD807C25775C" box="[234,325,577,598]" italics="true" pageId="8" pageNumber="340">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp.
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FD817C81775C" box="[413,481,576,598]" gridcol="1" gridrow="1" pageId="8" pageNumber="340">0 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFD817FE7775C" box="[506,647,576,598]" gridcol="2" gridrow="1" pageId="8" pageNumber="340">0 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFD817E2B775C" box="[671,843,576,598]" gridcol="3" gridrow="1" pageId="8" pageNumber="340">–</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FD817963775C" box="[881,1027,576,598]" gridcol="4" gridrow="1" pageId="8" pageNumber="340">0 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FD8179A2775C" box="[1078,1218,576,598]" gridcol="5" gridrow="1" pageId="8" pageNumber="340">0 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFD817814775C" box="[1256,1396,576,598]" gridcol="6" gridrow="1" pageId="8" pageNumber="340">0 [1]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FDA578147773" box="[163,1396,612,633]" gridrow="2" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FDA57C147773" box="[163,372,612,633]" gridcol="0" gridrow="2" pageId="8" pageNumber="340">nov.</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FD4A781477AB" box="[163,1396,651,673]" gridrow="3" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FD4A7C1477AB" box="[163,372,651,673]" gridcol="0" gridrow="3" pageId="8" pageNumber="340">
|
|||
|
|
Within
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF7FFD4D7C0877AA" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[234,360,651,673]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF7FFD4D7C0877AA" box="[234,360,651,673]" italics="true" pageId="8" pageNumber="340">R. homolepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FD4A7C8177AB" box="[413,481,651,673]" gridcol="1" gridrow="3" pageId="8" pageNumber="340">0 [21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFD4A7FE777AB" box="[506,647,651,673]" gridcol="2" gridrow="3" pageId="8" pageNumber="340">0.1 (0–0.1) [21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFD4A7E2B77AB" box="[671,843,651,673]" gridcol="3" gridrow="3" pageId="8" pageNumber="340">0.3 (0–1.2) [21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FD4A796377AB" box="[881,1027,651,673]" gridcol="4" gridrow="3" pageId="8" pageNumber="340">0 (0–0.6) [21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FD4A79A277AB" box="[1078,1218,651,673]" gridcol="5" gridrow="3" pageId="8" pageNumber="340">0.1 (0–0.4) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFD4A781477AB" box="[1256,1396,651,673]" gridcol="6" gridrow="3" pageId="8" pageNumber="340">0 [3]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FD72781477C3" box="[163,1396,691,713]" gridrow="4" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FD727C1477C3" box="[163,372,691,713]" gridcol="0" gridrow="4" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FD727C4877C2" authorityName="Gower" authorityYear="2020" box="[163,296,691,713]" class="Squamata" family="Uropeltidae" genus="Rhinophis" isUncertain="true" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FD727DD877C2" box="[163,184,691,712]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF49FD727C4877C2" box="[220,296,691,712]" italics="true" pageId="8" pageNumber="340">mendisi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEBDFD727C2877C2" box="[296,328,691,712]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
sp.
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FD727C8177C3" box="[413,481,691,713]" gridcol="1" gridrow="4" pageId="8" pageNumber="340">2.2 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFD727FE777C3" box="[506,647,691,713]" gridcol="2" gridrow="4" pageId="8" pageNumber="340">1.4 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFD727E2B77C3" box="[671,843,691,713]" gridcol="3" gridrow="4" pageId="8" pageNumber="340">11.3 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FD72796377C3" box="[881,1027,691,713]" gridcol="4" gridrow="4" pageId="8" pageNumber="340">7.2 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FD7279A277C3" box="[1078,1218,691,713]" gridcol="5" gridrow="4" pageId="8" pageNumber="340">0.9 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFD72781477C3" box="[1256,1396,691,713]" gridcol="6" gridrow="4" pageId="8" pageNumber="340">0.3 [1]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FD16781477E7" box="[163,1396,727,749]" gridrow="5" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FD167C1477E7" box="[163,372,727,749]" gridcol="0" gridrow="5" pageId="8" pageNumber="340">Palabaddala</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FD3F7814761E" box="[163,1396,766,788]" gridrow="6" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FD3F7C14761E" box="[163,372,766,788]" gridcol="0" gridrow="6" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FD3E7C487619" authorityName="Gower" authorityYear="2020" box="[163,296,766,788]" class="Squamata" family="Uropeltidae" genus="Rhinophis" isUncertain="true" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FD3E7DD8761E" box="[163,184,767,788]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF49FD3F7C487619" box="[220,296,766,787]" italics="true" pageId="8" pageNumber="340">mendisi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEBDFD3F7C287619" box="[296,328,766,787]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FD3F7C81761E" box="[413,481,766,788]" gridcol="1" gridrow="6" pageId="8" pageNumber="340">2.5 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFD3F7FE7761E" box="[506,647,766,788]" gridcol="2" gridrow="6" pageId="8" pageNumber="340">1.3 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFD3F7E2B761E" box="[671,843,766,788]" gridcol="3" gridrow="6" pageId="8" pageNumber="340">1.5 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FD3F7963761E" box="[881,1027,766,788]" gridcol="4" gridrow="6" pageId="8" pageNumber="340">5.7 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FD3F79A2761E" box="[1078,1218,766,788]" gridcol="5" gridrow="6" pageId="8" pageNumber="340">0.9 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFD3F7814761E" box="[1256,1396,766,788]" gridcol="6" gridrow="6" pageId="8" pageNumber="340">0.3 [1]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FCE37814763D" box="[163,1396,802,823]" gridrow="7" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FCE37C14763D" box="[163,372,802,823]" gridcol="0" gridrow="7" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FCE37C6B763D" authorityName="Deraniyagala" authorityYear="1975" box="[163,267,802,823]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="tricolorata">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FCE37C6B763D" box="[163,267,802,823]" italics="true" pageId="8" pageNumber="340">tricolorata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FC8878147655" box="[163,1396,841,863]" gridrow="8" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FC887C147655" box="[163,372,841,863]" gridcol="0" gridrow="8" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FC8B7C487654" authorityName="Gower" authorityYear="2020" box="[163,296,841,863]" class="Squamata" family="Uropeltidae" genus="Rhinophis" isUncertain="true" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FC8B7DD87655" box="[163,184,842,863]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF49FC887C487654" box="[220,296,841,862]" italics="true" pageId="8" pageNumber="340">mendisi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEBDFC887C287654" box="[296,328,841,862]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FC887C817655" box="[413,481,841,863]" gridcol="1" gridrow="8" pageId="8" pageNumber="340">2.2 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFC887FE77655" box="[506,647,841,863]" gridcol="2" gridrow="8" pageId="8" pageNumber="340">1.4 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFC887E2B7655" box="[671,843,841,863]" gridcol="3" gridrow="8" pageId="8" pageNumber="340">9.6 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FC8879637655" box="[881,1027,841,863]" gridcol="4" gridrow="8" pageId="8" pageNumber="340">7.5 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FC8879A27655" box="[1078,1218,841,863]" gridcol="5" gridrow="8" pageId="8" pageNumber="340">1.2 (1.1–1.3) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFC8878147655" box="[1256,1396,841,863]" gridcol="6" gridrow="8" pageId="8" pageNumber="340">0.8 [2]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FCAF78147689" box="[163,1396,878,899]" gridrow="9" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FCAF7C147689" box="[163,372,878,899]" gridcol="0" gridrow="9" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FCAF7D837689" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[163,227,878,899]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FCAF7D837689" box="[163,227,878,899]" italics="true" pageId="8" pageNumber="340">martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="B4935783767EA15FFF7CFCAF7C537689" box="[233,307,878,899]" pageId="8" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FC55781476A1" box="[163,1396,916,939]" gridrow="10" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FC557C1476A1" box="[163,372,916,939]" gridcol="0" gridrow="10" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FC547C4876A0" authorityName="Gower" authorityYear="2020" box="[163,296,917,939]" class="Squamata" family="Uropeltidae" genus="Rhinophis" isUncertain="true" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FC547DD876A0" box="[163,184,917,938]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF49FC547C4876A0" box="[220,296,917,938]" italics="true" pageId="8" pageNumber="340">mendisi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEBDFC547C2876A0" box="[296,328,917,938]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FC557C8176A1" box="[413,481,916,939]" gridcol="1" gridrow="10" pageId="8" pageNumber="340">2.2 [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFC557FE776A1" box="[506,647,916,939]" gridcol="2" gridrow="10" pageId="8" pageNumber="340">0.9 (0.8–0.9) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFC557E2B76A1" box="[671,843,916,939]" gridcol="3" gridrow="10" pageId="8" pageNumber="340">12.8 (12.4–14) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FC55796376A1" box="[881,1027,916,939]" gridcol="4" gridrow="10" pageId="8" pageNumber="340">5.8 (5.1–6.2) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FC5579A276A1" box="[1078,1218,916,939]" gridcol="5" gridrow="10" pageId="8" pageNumber="340">0.9 (0.7–0.9) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFC55781476A1" box="[1256,1396,916,939]" gridcol="6" gridrow="10" pageId="8" pageNumber="340">0.5 [3]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FC78781476C4" box="[163,1396,953,974]" gridrow="11" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FC787C1476C4" box="[163,372,953,974]" gridcol="0" gridrow="11" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FC787C6676C4" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[163,262,953,974]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FC787C6676C4" box="[163,262,953,974]" italics="true" pageId="8" pageNumber="340">homolepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FC21781476FC" box="[163,1396,992,1014]" gridrow="12" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FC217C1476FC" box="[163,372,992,1014]" gridcol="0" gridrow="12" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FC207DD876FC" box="[163,184,993,1014]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
sp. Palabaddala
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEC7FC207C1476FC" box="[338,372,993,1014]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FC217C8176FC" box="[413,481,992,1014]" gridcol="1" gridrow="12" pageId="8" pageNumber="340">1.9 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFC217FE776FC" box="[506,647,992,1014]" gridcol="2" gridrow="12" pageId="8" pageNumber="340">1.1 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFC217E2B76FC" box="[671,843,992,1014]" gridcol="3" gridrow="12" pageId="8" pageNumber="340">7 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FC21796376FC" box="[881,1027,992,1014]" gridcol="4" gridrow="12" pageId="8" pageNumber="340">5.3 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FC2179A276FC" box="[1078,1218,992,1014]" gridcol="5" gridrow="12" pageId="8" pageNumber="340">0 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFC21781476FC" box="[1256,1396,992,1014]" gridcol="6" gridrow="12" pageId="8" pageNumber="340">0.5 [1]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FBC578147113" box="[163,1396,1028,1049]" gridrow="13" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FBC57C147113" box="[163,372,1028,1049]" gridcol="0" gridrow="13" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FBC57C6B7113" authorityName="Deraniyagala" authorityYear="1975" box="[163,267,1028,1049]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="tricolorata">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FBC57C6B7113" box="[163,267,1028,1049]" italics="true" pageId="8" pageNumber="340">tricolorata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FBEA7814714B" box="[163,1396,1067,1089]" gridrow="14" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FBEA7C14714B" box="[163,372,1067,1089]" gridcol="0" gridrow="14" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FBED7DD8714B" box="[163,184,1068,1089]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
sp. Palabaddala
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEC7FBED7C14714B" box="[338,372,1068,1089]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FBEA7C81714B" box="[413,481,1067,1089]" gridcol="1" gridrow="14" pageId="8" pageNumber="340">2.7 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFBEA7FE7714B" box="[506,647,1067,1089]" gridcol="2" gridrow="14" pageId="8" pageNumber="340">1.3 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFBEA7E2B714B" box="[671,843,1067,1089]" gridcol="3" gridrow="14" pageId="8" pageNumber="340">6.2 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FBEA7963714B" box="[881,1027,1067,1089]" gridcol="4" gridrow="14" pageId="8" pageNumber="340">6.5 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FBEA79A2714B" box="[1078,1218,1067,1089]" gridcol="5" gridrow="14" pageId="8" pageNumber="340">0.3 (0.2–0.4) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFBEA7814714B" box="[1256,1396,1067,1089]" gridcol="6" gridrow="14" pageId="8" pageNumber="340">0.9 [2]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FB917814716F" box="[163,1396,1104,1125]" gridrow="15" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FB917C14716F" box="[163,372,1104,1125]" gridcol="0" gridrow="15" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FB917D83716F" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[163,227,1104,1125]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FB917D83716F" box="[163,227,1104,1125]" italics="true" pageId="8" pageNumber="340">martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="B4935783767EA15FFF7CFB917C53716F" box="[233,307,1104,1125]" pageId="8" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FBB778147187" box="[163,1396,1142,1165]" gridrow="16" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FBB77C147187" box="[163,372,1142,1165]" gridcol="0" gridrow="16" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FBB67DD87186" box="[163,184,1143,1164]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
sp. Palabaddala
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFEC7FBB67C147186" box="[338,372,1143,1164]" italics="true" pageId="8" pageNumber="340">–R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FBB77C817187" box="[413,481,1142,1165]" gridcol="1" gridrow="16" pageId="8" pageNumber="340">1.1 [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFBB77FE77187" box="[506,647,1142,1165]" gridcol="2" gridrow="16" pageId="8" pageNumber="340">1.2 (1.1–1.3) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFBB77E2B7187" box="[671,843,1142,1165]" gridcol="3" gridrow="16" pageId="8" pageNumber="340">11.7 (11.2–13.7) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FBB779637187" box="[881,1027,1142,1165]" gridcol="4" gridrow="16" pageId="8" pageNumber="340">6.6 (6.4–6.7) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FBB779A27187" box="[1078,1218,1142,1165]" gridcol="5" gridrow="16" pageId="8" pageNumber="340">0.3 (0.2–0.4) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFBB778147187" box="[1256,1396,1142,1165]" gridcol="6" gridrow="16" pageId="8" pageNumber="340">0.7 (0.5–0.8) [3]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FB5A781471BA" box="[163,1396,1179,1200]" gridrow="17" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FB5A7C1471BA" box="[163,372,1179,1200]" gridcol="0" gridrow="17" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FB5A7C6671BA" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[163,262,1179,1200]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FB5A7C6671BA" box="[163,262,1179,1200]" italics="true" pageId="8" pageNumber="340">homolepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FB03781471D2" box="[163,1396,1218,1240]" gridrow="18" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FB037C1471D2" box="[163,372,1218,1240]" gridcol="0" gridrow="18" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FB027C2771DD" box="[163,327,1218,1240]" italics="true" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FB027C4671DD" authorityName="Deraniyagala" authorityYear="1975" box="[163,294,1218,1240]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="tricolorata">R. tricolorata</taxonomicName>
|
|||
|
|
–R.
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FB037C8171D2" box="[413,481,1218,1240]" gridcol="1" gridrow="18" pageId="8" pageNumber="340">0.8 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFB037FE771D2" box="[506,647,1218,1240]" gridcol="2" gridrow="18" pageId="8" pageNumber="340">0.6 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFB037E2B71D2" box="[671,843,1218,1240]" gridcol="3" gridrow="18" pageId="8" pageNumber="340">2.8 [1]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FB03796371D2" box="[881,1027,1218,1240]" gridcol="4" gridrow="18" pageId="8" pageNumber="340">2 [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FB0379A271D2" box="[1078,1218,1218,1240]" gridcol="5" gridrow="18" pageId="8" pageNumber="340">0.3 (0.2–0.4) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFB03781471D2" box="[1256,1396,1218,1240]" gridcol="6" gridrow="18" pageId="8" pageNumber="340">0.5 [2]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FB26781471F6" box="[163,1396,1255,1276]" gridrow="19" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FB267C1471F6" box="[163,372,1255,1276]" gridcol="0" gridrow="19" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FB267D8371F6" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[163,227,1255,1276]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FB267D8371F6" box="[163,227,1255,1276]" italics="true" pageId="8" pageNumber="340">martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="B4935783767EA15FFF7CFB267C5371F6" box="[233,307,1255,1276]" pageId="8" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FACC78147029" box="[163,1396,1293,1315]" gridrow="20" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FACC7C147029" box="[163,372,1293,1315]" gridcol="0" gridrow="20" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FACF7C277028" box="[163,327,1293,1315]" italics="true" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FACF7C467028" authorityName="Deraniyagala" authorityYear="1975" box="[163,294,1293,1315]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="tricolorata">R. tricolorata</taxonomicName>
|
|||
|
|
–R.
|
|||
|
|
</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FACC7C817029" box="[413,481,1293,1315]" gridcol="1" gridrow="20" pageId="8" pageNumber="340">1.9 [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFACC7FE77029" box="[506,647,1293,1315]" gridcol="2" gridrow="20" pageId="8" pageNumber="340">0.7 (0.6–0.8) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFACC7E2B7029" box="[671,843,1293,1315]" gridcol="3" gridrow="20" pageId="8" pageNumber="340">14.5 (13.5–17.8) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FACC79637029" box="[881,1027,1293,1315]" gridcol="4" gridrow="20" pageId="8" pageNumber="340">5 (4.7–5.2) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FACC79A27029" box="[1078,1218,1293,1315]" gridcol="5" gridrow="20" pageId="8" pageNumber="340">0.1 (0–0.4) [3]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFACC78147029" box="[1256,1396,1293,1315]" gridcol="6" gridrow="20" pageId="8" pageNumber="340">0.8 [3]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FAF07814704C" box="[163,1396,1329,1350]" gridrow="21" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FAF07C14704C" box="[163,372,1329,1350]" gridcol="0" gridrow="21" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FAF07C66704C" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[163,262,1329,1350]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FAF07C66704C" box="[163,262,1329,1350]" italics="true" pageId="8" pageNumber="340">homolepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</th>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FA9878147064" box="[163,1396,1369,1390]" gridrow="22" pageId="8" pageNumber="340">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FA987C147064" box="[163,372,1369,1390]" gridcol="0" gridrow="22" pageId="8" pageNumber="340">
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF36FA987D9D7064" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[163,253,1369,1390]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF36FA987D9D7064" box="[163,253,1369,1390]" italics="true" pageId="8" pageNumber="340">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="B4935783767EA15FFE96FA987C2D7064" box="[259,333,1369,1390]" pageId="8" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
–
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFECCFA987C0E7064" box="[345,366,1369,1390]" italics="true" pageId="8" pageNumber="340">R.</emphasis>
|
|||
|
|
</th>
|
|||
|
|
<td id="60355DD4767E5EA8FE08FA987C817064" box="[413,481,1369,1390]" gridcol="1" gridrow="22" pageId="8" pageNumber="340">2.7 [21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FE6FFA987FE77064" box="[506,647,1369,1390]" gridcol="2" gridrow="22" pageId="8" pageNumber="340">1.1 (1–1.2) [21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FD0AFA987E2B7064" box="[671,843,1369,1390]" gridcol="3" gridrow="22" pageId="8" pageNumber="340">11.7 (10.7–14.2) [7]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FCE4FA9879637064" box="[881,1027,1369,1390]" gridcol="4" gridrow="22" pageId="8" pageNumber="340">5.6 (5.6–5.7)[21]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FBA3FA9879A27064" box="[1078,1218,1369,1390]" gridcol="5" gridrow="22" pageId="8" pageNumber="340">0.4 (0.2–0.7) [9]</td>
|
|||
|
|
<td id="60355DD4767E5EA8FB7DFA9878147064" box="[1256,1396,1369,1390]" gridcol="6" gridrow="22" pageId="8" pageNumber="340">1.3 (1.1–1.4) [6]</td>
|
|||
|
|
</tr>
|
|||
|
|
<tr id="23E434A8767E5EA8FF36FABC78147098" box="[163,1396,1405,1426]" gridrow="23" pageId="8" pageNumber="340" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
|
|||
|
|
<th id="60355DD4767E5EA8FF36FABC7C147098" box="[163,372,1405,1426]" gridcol="0" gridrow="23" pageId="8" pageNumber="340">homolepis</th>
|
|||
|
|
</tr>
|
|||
|
|
</table>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA767EA15EFF52FA037E9175DF" blockId="8.[151,1437,1474,2041]" lastBlockId="9.[151,1436,150,213]" lastPageId="9" lastPageNumber="341" pageId="8" pageNumber="340">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF52FA037CBA70D7" bold="true" box="[199,474,1474,1501]" pageId="8" pageNumber="340">
|
|||
|
|
Variation in
|
|||
|
|
<typeStatus id="426F8848767EA15FFEC8FA027CB570D7" box="[349,469,1475,1501]" pageId="8" pageNumber="340" type="paratype">paratypes</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Condition generally good. See
|
|||
|
|
<tableCitation id="D0560351767EA15FFCA8FA037EF070D7" box="[829,912,1474,1501]" captionStart="TABLE 2" captionStartId="6.[151,239,692,716]" captionTargetPageId="6" captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded" pageId="8" pageNumber="340">Table 2</tableCitation>
|
|||
|
|
for some meristic and morphometric data, and
|
|||
|
|
<figureCitation id="05EF2A6F767EA15FFF02FA277D86730B" box="[151,230,1510,1537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1543,1567]" captionTargetBox="[170,1416,181,1520]" captionTargetId="figure-18@4.[170,1416,181,1520]" captionTargetPageId="4" captionText="FIGURE 3. Seven paratypes of Rhinophis martin sp. nov. Left column approximately dorsal view; right column approximately ventral view. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bars 20 mm. Photographs by H. Taylor." figureDoi="http://doi.org/10.5281/zenodo.11547476" httpUri="https://zenodo.org/record/11547476/files/figure.png" pageId="8" pageNumber="340">Figs. 3</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="05EF2A6F767EA15FFEB4FA267C4F730B" box="[289,303,1511,1537]" captionStart="FIGURE 4" captionStartId="9.[151,250,1537,1561]" captionTargetBox="[151,1435,237,1513]" captionTargetId="figure-62@9.[151,1435,237,1513]" captionTargetPageId="9" captionText="FIGURE 4. Head and tail ends of seven paratypes of Rhinophis martin sp. nov. in four views. A. USNM 548100, B. USNM 548117, C. USNM 548118, D. USNM 548120, E. USNM 548121, F. USNM 548122, G. USNM 548123. Scale bar increments in millimetres. Photographs by H. Taylor." figureDoi="http://doi.org/10.5281/zenodo.11547478" httpUri="https://zenodo.org/record/11547478/files/figure.png" pageId="8" pageNumber="340">4</figureCitation>
|
|||
|
|
for photographs. In scalation all
|
|||
|
|
<typeStatus id="426F8848767EA15FFD3AFA267E7A730B" box="[687,794,1511,1537]" pageId="8" pageNumber="340" type="paratype">paratypes</typeStatus>
|
|||
|
|
resemble the
|
|||
|
|
<typeStatus id="426F8848767EA15FFC28FA27797F730A" box="[957,1055,1510,1536]" pageId="8" pageNumber="340" type="holotype">holotype</typeStatus>
|
|||
|
|
except where noted here. In
|
|||
|
|
<specimenCount id="8BD2FD63767EA15FFAE4FA277C62732F" count="5" pageId="8" pageNumber="340" type="generic" typeStatus="paratype">five paratypes</specimenCount>
|
|||
|
|
(USNM 548100, 548122; NMSL-NH
|
|||
|
|
<date id="E96A102A767EA15FFD21F9CA7E54732F" box="[692,820,1547,1573]" pageId="8" pageNumber="340" value="2024-01-01">2024.01.01</date>
|
|||
|
|
,
|
|||
|
|
<date id="E96A102A767EA15FFCAAF9CA7EA1732F" box="[831,961,1547,1573]" pageId="8" pageNumber="340" value="2024-01-02">2024.01.02</date>
|
|||
|
|
; DWC
|
|||
|
|
<date id="E96A102A767EA15FFB86F9CB79F3732F" box="[1043,1171,1546,1573]" pageId="8" pageNumber="340" value="2024-05-01">2024.05.01</date>
|
|||
|
|
) the rostral contacts the frontal and prevents midline contact between opposite prefrontals. USNM 548100 and NMSL-NH
|
|||
|
|
<date id="E96A102A767EA15FFB70F9EE78037343" box="[1253,1379,1583,1609]" pageId="8" pageNumber="340" value="2024-01-02">2024.01.02</date>
|
|||
|
|
each have a small (temporal-like) scale separating the parietal from the fourth supralabial on each side, much smaller on the left in USNM 548100 (among uropeltids, temporals occur consistently only in
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFBCAF9B7787A739A" authorityName="Wall" authorityYear="1921" box="[1119,1306,1654,1680]" class="Squamata" family="Uropeltidae" genus="Brachyophidium" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFBCAF9B7787A739A" box="[1119,1306,1654,1680]" italics="true" pageId="8" pageNumber="340">Brachyophidium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFABDF9B678FC739A" authorityName="Beddome" authorityYear="1886" box="[1320,1436,1655,1680]" class="Squamata" family="Uropeltidae" genus="Teretrurus" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFABDF9B678FC739A" box="[1320,1436,1655,1680]" italics="true" pageId="8" pageNumber="340">Teretrurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="5AD44D69767EA15FFF5DF95B7C1673BE" box="[200,374,1690,1716]" class="Squamata" family="Uropeltidae" genus="Platyplectrurus" kingdom="Animalia" pageId="8" pageNumber="340" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="AFA0EAF8767EA15FFF5DF95B7C1673BE" box="[200,374,1690,1716]" italics="true" pageId="8" pageNumber="340">Platyplectrurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
). Left parietal overlaps right in all except USNM 548118 and NMSL-NH
|
|||
|
|
<date id="E96A102A767EA15FFB5FF95A782C73BF" box="[1226,1356,1691,1717]" pageId="8" pageNumber="340" value="2024-01-02">2024.01.02</date>
|
|||
|
|
; where prefrontals overlap, left lies above right in all except USNM 548120 and 548121. Eye 0.2–0.3 times length of ocular. In
|
|||
|
|
<specimenCount id="8BD2FD63767EA15FFE9DF9227CC473F7" box="[264,420,1763,1789]" count="2" pageId="8" pageNumber="340" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
|||
|
|
(USNM 548118, 548120) the first ventral does not contact the first infralabials. Scale row reductions simple (see Appendix 1), reducing from 19 to 17 dorsal scale rows by level with the 31
|
|||
|
|
<superScript id="6AA19BA2767EA15FFB7EF8C77996721E" attach="left" box="[1259,1270,1798,1812]" fontSize="6" pageId="8" pageNumber="340">st</superScript>
|
|||
|
|
to 42
|
|||
|
|
<superScript id="6AA19BA2767EA15FFAA2F8C77827721E" attach="left" box="[1335,1351,1798,1812]" fontSize="6" pageId="8" pageNumber="340">nd</superScript>
|
|||
|
|
ventral without apparent sexual dimorphism. Ventrals in female
|
|||
|
|
<typeStatus id="426F8848767EA15FFC81F8EA7E1F724F" box="[788,895,1835,1861]" pageId="8" pageNumber="340" type="paratype">paratypes</typeStatus>
|
|||
|
|
175–181, males 167–174; subcaudals in female
|
|||
|
|
<typeStatus id="426F8848767EA15FFF02F88E7C627263" box="[151,258,1871,1897]" pageId="8" pageNumber="340" type="paratype">paratypes</typeStatus>
|
|||
|
|
3,3 or 4,4, in males 6,
|
|||
|
|
<quantity id="5A2C9B0F767EA15FFD99F88F7F5B7262" box="[524,571,1870,1896]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="340" unit="in" value="6.0">6 in</quantity>
|
|||
|
|
all except 5,
|
|||
|
|
<quantity id="5A2C9B0F767EA15FFD44F88F7F9F7262" box="[721,767,1870,1897]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="340" unit="in" value="5.0">5 in</quantity>
|
|||
|
|
DWC
|
|||
|
|
<date id="E96A102A767EA15FFCC1F88F7EB27263" box="[852,978,1870,1897]" pageId="8" pageNumber="340" value="2024-05-01">2024.05.01</date>
|
|||
|
|
and 7,
|
|||
|
|
<quantity id="5A2C9B0F767EA15FFB8DF88F79277262" box="[1048,1095,1870,1896]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="340" unit="in" value="6.0">6 in</quantity>
|
|||
|
|
USNM 548123. At midbody, ventrals 1.2–1.33 times width of scales in first dorsal row. At least one pair of subcaudals is ‘fused’ (undivided) in all
|
|||
|
|
<typeStatus id="426F8848767EA15FFF02F8567C6272BB" box="[151,258,1943,1969]" pageId="8" pageNumber="340" type="paratype">paratypes</typeStatus>
|
|||
|
|
except USNM 548118 and NMSL-NH
|
|||
|
|
<date id="E96A102A767EA15FFD52F8567E2872BB" box="[711,840,1943,1969]" pageId="8" pageNumber="340" value="2024-01-02">2024.01.02</date>
|
|||
|
|
. Scales around the tail shield 10–13. Most
|
|||
|
|
<typeStatus id="426F8848767EA15FFAA4F85678FC72BB" box="[1329,1436,1943,1969]" pageId="8" pageNumber="340" type="paratype">paratypes</typeStatus>
|
|||
|
|
lack notable ridges on scales in the tail and posterior end of body; low ridges on upper posterior tail dorsal scale row scales in USNM 548117 and 548121, short low ridges on posterior ends of some tail dorsal scale row scales in 548122. Teeth not counted except in USNM 548122, which resembles the
|
|||
|
|
<typeStatus id="426F8848767FA15EFC43FF56795875BB" box="[982,1080,151,177]" pageId="9" pageNumber="341" type="holotype">holotype</typeStatus>
|
|||
|
|
in having seven maxillary teeth per side, but differs in having seven rather than eight teeth in each mandible.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="C9AB6662767FA15EFF02F9C07F5C736B" ID-DOI="http://doi.org/10.5281/zenodo.11547478" ID-Zenodo-Dep="11547478" httpUri="https://zenodo.org/record/11547478/files/figure.png" pageId="9" pageNumber="341" startId="9.[151,250,1537,1561]" targetBox="[151,1435,237,1513]" targetPageId="9" targetType="figure">
|
|||
|
|
<paragraph id="9D6B36EA767FA15EFF02F9C07F5C736B" blockId="9.[151,1436,1537,1633]" pageId="9" pageNumber="341">
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFF02F9C07C747313" bold="true" box="[151,276,1537,1561]" pageId="9" pageNumber="341">FIGURE 4.</emphasis>
|
|||
|
|
Head and tail ends of seven paratypes of
|
|||
|
|
<taxonomicName id="5AD44D69767FA15EFD56F9C07E157313" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[707,885,1537,1561]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="9" pageNumber="341" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFD56F9C07E157313" box="[707,885,1537,1561]" italics="true" pageId="9" pageNumber="341">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFCE9F9C07EAE7313" bold="true" box="[892,974,1537,1561]" pageId="9" pageNumber="341">
|
|||
|
|
<taxonomicNameLabel id="B4935783767FA15EFCE9F9C07EAE7313" box="[892,974,1537,1561]" pageId="9" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
in four views.
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFBFEF9C079E37313" bold="true" box="[1131,1155,1537,1561]" pageId="9" pageNumber="341">A.</emphasis>
|
|||
|
|
USNM 548100,
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFAA0F9C0782C7313" bold="true" box="[1333,1356,1537,1561]" pageId="9" pageNumber="341">B.</emphasis>
|
|||
|
|
USNM 548117,
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFF7AF9E47C677337" bold="true" box="[239,263,1573,1597]" pageId="9" pageNumber="341">C.</emphasis>
|
|||
|
|
USNM 548118,
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFE21F9E47CAD7337" bold="true" box="[436,461,1573,1597]" pageId="9" pageNumber="341">D.</emphasis>
|
|||
|
|
USNM 548120,
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFDEEF9E47FF27337" bold="true" box="[635,658,1573,1597]" pageId="9" pageNumber="341">E.</emphasis>
|
|||
|
|
USNM 548121,
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFCD5F9E47E337337" bold="true" box="[832,851,1573,1597]" pageId="9" pageNumber="341">F.</emphasis>
|
|||
|
|
USNM 548122,
|
|||
|
|
<emphasis id="AFA0EAF8767FA15EFB97F9E4797C7337" bold="true" box="[1026,1052,1573,1597]" pageId="9" pageNumber="341">G.</emphasis>
|
|||
|
|
USNM 548123. Scale bar increments in millimetres. Photographs by H. Taylor.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="9D6B36EA767FA15DFF52F9517E6B74DB" blockId="9.[151,1437,1679,2030]" lastBlockId="10.[151,1437,150,465]" lastPageId="10" lastPageNumber="342" pageId="9" pageNumber="341">
|
|||
|
|
<typeStatus id="426F8848767FA15EFF52F9517C5173A0" box="[199,305,1680,1706]" pageId="9" pageNumber="341" type="paratype">Paratypes</typeStatus>
|
|||
|
|
similar in colour and pattern to
|
|||
|
|
<typeStatus id="426F8848767FA15EFD0BF94E7E6073A3" box="[670,768,1679,1705]" pageId="9" pageNumber="341" type="holotype">holotype</typeStatus>
|
|||
|
|
, with minor variations. All
|
|||
|
|
<typeStatus id="426F8848767FA15EFBA3F95179FF73A0" box="[1078,1183,1680,1706]" pageId="9" pageNumber="341" type="paratype">paratypes</typeStatus>
|
|||
|
|
lack pale patch across posterior of nasals and anterior of prefrontals; USNM 548100 with sinuous pale band across posterior of oculars and prefrontals and anterior of frontal. Whitish transverse bands on dorsum behind head typically shorter and/or more irregular and asymmetrical in
|
|||
|
|
<typeStatus id="426F8848767FA15EFE4EF93D7F24721C" box="[475,580,1788,1814]" pageId="9" pageNumber="341" type="paratype">paratypes</typeStatus>
|
|||
|
|
than in
|
|||
|
|
<typeStatus id="426F8848767FA15EFD0DF93A7F9A721F" box="[664,762,1787,1813]" pageId="9" pageNumber="341" type="holotype">holotype</typeStatus>
|
|||
|
|
, forming spots or subtriangular patches rather than bands. One or two pale ventral scales and adjacent dorsal scale rows anteriorly (between ventral 3–5) in most
|
|||
|
|
<typeStatus id="426F8848767FA15EFB41F8E178207230" box="[1236,1344,1824,1850]" pageId="9" pageNumber="341" type="paratype">paratypes</typeStatus>
|
|||
|
|
; USNM 548117 with mostly pale venter for ca.
|
|||
|
|
<quantity id="5A2C9B0F767FA15EFDCCF8827FC97254" box="[601,681,1859,1886]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="341" unit="mm" value="20.0">20 mm</quantity>
|
|||
|
|
behind third ventral; USNM 548122 without pale anterior patches ventrally. Final one or two ventrals pale in all
|
|||
|
|
<typeStatus id="426F8848767FA15EFD1AF8A97F987288" box="[655,760,1896,1922]" pageId="9" pageNumber="341" type="paratype">paratypes</typeStatus>
|
|||
|
|
except USNM 548117, 548118 or 548122. Background body colour paler in USNM 548100, darker in 548123. Darker pair of paravertebral longitudinal lines particularly faint, diffuse and incomplete in USNM 548100, particularly clear in 548120. Dark dorsal cross bars similar to
|
|||
|
|
<typeStatus id="426F8848767FA15EFA8AF86E781F72C3" box="[1311,1407,1967,1993]" pageId="9" pageNumber="341" type="holotype">holotype</typeStatus>
|
|||
|
|
in USNM 548117, 548121 and 548122 but less tall and more triangular or irregularly spot-like in other
|
|||
|
|
<typeStatus id="426F8848767FA15EFB4FF815782672E4" box="[1242,1350,2004,2030]" pageId="9" pageNumber="341" type="paratype">paratypes</typeStatus>
|
|||
|
|
. Border between darker dorsum and paler venter on 5
|
|||
|
|
<superScript id="6AA19BA2767CA15DFD19FF577FF975AE" attach="left" box="[652,665,150,164]" fontSize="6" pageId="10" pageNumber="342">th</superScript>
|
|||
|
|
or between 5
|
|||
|
|
<superScript id="6AA19BA2767CA15DFCA5FF577E5D75AE" attach="left" box="[816,829,150,164]" fontSize="6" pageId="10" pageNumber="342">th</superScript>
|
|||
|
|
and 6
|
|||
|
|
<superScript id="6AA19BA2767CA15DFC14FF577EEE75AE" attach="left" box="[897,910,150,164]" fontSize="6" pageId="10" pageNumber="342">th</superScript>
|
|||
|
|
dorsal scale row. Whitish spots along 5
|
|||
|
|
<superScript id="6AA19BA2767CA15DFAD0FF57783275AE" attach="left" box="[1349,1362,150,164]" fontSize="6" pageId="10" pageNumber="342">th</superScript>
|
|||
|
|
dorsal scale row absent in USNM 548118 and 548122, forming notably larger, elongated pale patches with darker lower and upper rims in USNM 548123. First dorsal scale row on tail with blackish dots in USNM 548100, only blackish on a few scales in 548121, only blackish on anteriormost scales in 548120, not blackish in 548123. Pale lateral stripe on tail less regular and complete in
|
|||
|
|
<typeStatus id="426F8848767CA15DFE58FEE67F58744B" box="[461,568,295,321]" pageId="10" pageNumber="342" type="paratype">paratypes</typeStatus>
|
|||
|
|
, absent in USNM 548122, thicker (two or more dorsal scale row scales thick) in 548120 and 548123, restricted to posterior end of tail in 548100 and 548118. Dorsolateral yellow cross bars on anterior of tail absent in USNM 548118, short and broadly incomplete in 548122, narrow and almost middorsally complete in 548117. Shield more orange-yellow in USNM 548121, mostly dark in 548118 and 548122 and especially in 548117 and
|
|||
|
|
<quantity id="5A2C9B0F767CA15DFF50FE777C5474DB" box="[197,308,438,465]" metricMagnitude="4" metricUnit="m" metricValue="1.3922324199999996" pageId="10" pageNumber="342" unit="in" value="548123.0">548123 in</quantity>
|
|||
|
|
which dark patches extend almost to apex.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="C9AB6662767CA15DFF02FA697EF870EF" ID-DOI="http://doi.org/10.5281/zenodo.11547480" ID-Zenodo-Dep="11547480" httpUri="https://zenodo.org/record/11547480/files/figure.png" pageId="10" pageNumber="342" startId="10.[151,250,1448,1472]" targetBox="[152,1398,533,1406]" targetPageId="10" targetType="figure">
|
|||
|
|
<paragraph id="9D6B36EA767CA15DFF02FA697EF870EF" blockId="10.[151,1436,1448,1509]" pageId="10" pageNumber="342">
|
|||
|
|
<emphasis id="AFA0EAF8767CA15DFF02FA697C7270CB" bold="true" box="[151,274,1448,1473]" pageId="10" pageNumber="342">FIGURE 5.</emphasis>
|
|||
|
|
Graph showing relationship between number of ventral scales and number of subcaudal scales (mean value for left and right) in female (triangles) and male (circles)
|
|||
|
|
<taxonomicName id="5AD44D69767CA15DFD05FA0D7E2170EE" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[656,833,1484,1508]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="10" pageNumber="342" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767CA15DFD05FA0D7E2170EE" box="[656,833,1484,1508]" italics="true" pageId="10" pageNumber="342">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="B4935783767CA15DFCD2FA0C7EF870EF" box="[839,920,1485,1509]" pageId="10" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="C9AB6662767CA15DFF02F80C781A72EF" ID-DOI="http://doi.org/10.5281/zenodo.11547482" ID-Zenodo-Dep="11547482" box="[151,1402,1997,2022]" httpUri="https://zenodo.org/record/11547482/files/figure.png" pageId="10" pageNumber="342" startId="10.[151,250,1997,2021]" targetBox="[151,1435,1533,1974]" targetPageId="10" targetType="figure">
|
|||
|
|
<paragraph id="9D6B36EA767CA15DFF02F80C781A72EF" blockId="10.[151,1402,1997,2022]" box="[151,1402,1997,2022]" pageId="10" pageNumber="342">
|
|||
|
|
<emphasis id="AFA0EAF8767CA15DFF02F80C7C7372EF" bold="true" box="[151,275,1997,2021]" pageId="10" pageNumber="342">FIGURE 6.</emphasis>
|
|||
|
|
<taxonomicName id="5AD44D69767CA15DFE8CF80C7CAA72EF" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[281,458,1997,2021]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="10" pageNumber="342" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767CA15DFE8CF80C7CAA72EF" box="[281,458,1997,2021]" italics="true" pageId="10" pageNumber="342">Rhinophis martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767CA15DFE45F80C7F4172EF" bold="true" box="[464,545,1997,2021]" pageId="10" pageNumber="342">
|
|||
|
|
<taxonomicNameLabel id="B4935783767CA15DFE45F80C7F4172EF" box="[464,545,1997,2021]" pageId="10" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, paratype DWC 2024.05.01 prior to fixation, in upper (left) and lower (right) views.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FF5778617713" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52FF577F4375BA" bold="true" box="[199,547,150,176]" pageId="11" pageNumber="343">Referred specimen variation.</emphasis>
|
|||
|
|
Some meristic and morphometric data presented in
|
|||
|
|
<tableCitation id="D0560351767DA15CFBE6FF5679AA75BB" box="[1139,1226,151,177]" captionStart="TABLE 2" captionStartId="6.[151,239,692,716]" captionTargetPageId="6" captionText="TABLE 2. Sex, total (TL) and tail (tL) length (in mm) and scalation data for the 22 referred specimens of Rhinophis martin sp. nov. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). “–“ indicates data not recorded" pageId="11" pageNumber="343">Table 2</tableCitation>
|
|||
|
|
. In
|
|||
|
|
<specimenCount id="8BD2FD63767DA15CFB62FF5678FD75BB" box="[1271,1437,151,177]" count="2" pageId="11" pageNumber="343" type="generic">two specimens</specimenCount>
|
|||
|
|
(USNM 548106, 548113) the rostral contacts the frontal and prevents midline contact between opposite prefrontals. Left parietal overlaps right in all except USNM 548109; where prefrontals overlap, left above right in all except USNM 548096, 548097, 548103, 548107, and 548111. Temporal-like scale on each side in USNM 548096, on right in USNM 548101. Frontal partly fused with right prefrontal in USNM 548098. All have 3,3 infralabials except USNM 548101 (3,4). First and second ventrals in contact in all except USNM 548098 and 548111). Right anal shield overlies left in all specimens except female specimens USNM 548103, 548111, and 548114. Scales around shield 10–13. Ventrals in males 169–178, females 179–186; subcaudals in males 5–7 per side, females typically 3–4 per side (two on one side on USNM 548107; five on one side in USNM 548115). Most referred specimens lack notable ridges on scales in the tail and posterior end of body; if present then typically low and on upper posterior dorsal scale row scales on tail, also on posteriormost subcaudals in, for example, USNM 548097, 548098, and 548114.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FDE37FFE776B" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52FDE37CDA7736" bold="true" box="[199,442,546,572]" pageId="11" pageNumber="343">Sexual dimorphism.</emphasis>
|
|||
|
|
Variation in the number of ventrals, subcaudals and in relative tail length is strongly bimodal across the sample as a whole (
|
|||
|
|
<figureCitation id="05EF2A6F767DA15CFDD9FD867FF0776B" box="[588,656,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1448,1472]" captionTargetBox="[152,1398,533,1406]" captionTargetId="figure-240@10.[151,1435,532,1412]" captionTargetPageId="10" captionText="FIGURE 5. Graph showing relationship between number of ventral scales and number of subcaudal scales (mean value for left and right) in female (triangles) and male (circles) Rhinophis martin sp. nov." figureDoi="http://doi.org/10.5281/zenodo.11547480" httpUri="https://zenodo.org/record/11547480/files/figure.png" pageId="11" pageNumber="343">Fig. 5</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FDAB7DAF7633" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52FDAB7C0B778E" bold="true" box="[199,363,618,644]" pageId="11" pageNumber="343">Colour in life.</emphasis>
|
|||
|
|
In life (
|
|||
|
|
<figureCitation id="05EF2A6F767DA15CFE57FDAA7F65778F" box="[450,517,619,645]" captionStart="FIGURE 6" captionStartId="10.[151,250,1997,2021]" captionTargetBox="[151,1435,1533,1974]" captionTargetId="figure-217@10.[151,1435,1533,1974]" captionTargetPageId="10" captionText="FIGURE 6. Rhinophis martin sp. nov., paratype DWC 2024.05.01 prior to fixation, in upper (left) and lower (right) views." figureDoi="http://doi.org/10.5281/zenodo.11547482" httpUri="https://zenodo.org/record/11547482/files/figure.png" pageId="11" pageNumber="343">Fig. 6</figureCitation>
|
|||
|
|
), the dorsal areas that are orange-brown in preserved specimens are golden yellow, and the brownish ventral colour seen in preservation is more blackish. The off-white markings on the lateral and ventral surfaces of preserved specimens are whitish in life. The pale yellowish white markings on the lateral surface of the tail of preserved specimens are similarly coloured in life. One uncollected, small specimen (<
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFA93FD17780377FB" box="[1286,1379,726,753]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="343" unit="mm" value="150.0">150 mm</quantity>
|
|||
|
|
total length) had a more whitish than black or brown venter but otherwise resembled the larger specimens documented here.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="D5CE6561767DA15CFF52FC827E9C76AF" pageId="11" pageNumber="343" type="etymology">
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FC827E9C76AF" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52FC827C2E7657" bold="true" box="[199,334,835,861]" pageId="11" pageNumber="343">Etymology.</emphasis>
|
|||
|
|
Named in honour of the late Martin Wijesinghe (1939–2021), in recognition of his conservation and education actions and advocacy, especially with respect to forest biota and ecosystems of southwestern
|
|||
|
|
<collectingCountry id="E5C3767A767DA15CFAEEFCA67D8176AF" name="Sri Lanka" pageId="11" pageNumber="343">Sri Lanka</collectingCountry>
|
|||
|
|
. For nomenclatural purposes, the species name is a noun in apposition.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="D5CE6561767DA15CFF52FC6F7F0A76E7" pageId="11" pageNumber="343" type="vernacular_names">
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FC6F7F0A76E7" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52FC6F7F4776C3" bold="true" box="[199,551,942,969]" pageId="11" pageNumber="343">Suggested vernacular name.</emphasis>
|
|||
|
|
Martinge thudulla (Sinhala); Maartinin nilakael pambu (Tamil); Martin’s rhinophis or Martin’s shieldtail (English).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="D5CE6561767DA15CFF52FC3778277303" pageId="11" pageNumber="343" type="distribution">
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FC3779C9705F" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52FC377E64711A" bold="true" box="[199,772,1014,1041]" pageId="11" pageNumber="343">Distribution, natural history, and conservation.</emphasis>
|
|||
|
|
As far as is known,
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFC61FC367900711A" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[1012,1120,1015,1040]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFC61FC367900711A" box="[1012,1120,1015,1040]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFBFCFC3679A5711B" bold="true" box="[1129,1221,1015,1041]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFBFCFC3679A5711B" box="[1129,1221,1015,1041]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is restricted to the southeastern part of the Rakwana Massif, where it occurs from
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFCC0FBDB7EFC713F" box="[853,924,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="343" unit="m" value="750.0">750 m</quantity>
|
|||
|
|
to at least
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFB87FBDA7938713F" box="[1042,1112,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="343" unit="m" value="820.0">820 m</quantity>
|
|||
|
|
elevation (
|
|||
|
|
<figureCitation id="05EF2A6F767DA15CFB46FBDA7878713F" box="[1235,1304,1051,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1935]" captionTargetBox="[273,1315,185,1881]" captionTargetId="figure-31@3.[266,1321,181,1887]" captionTargetPageId="3" captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)." figureDoi="http://doi.org/10.5281/zenodo.11547474" httpUri="https://zenodo.org/record/11547474/files/figure.png" pageId="11" pageNumber="343">Fig. 2</figureCitation>
|
|||
|
|
). The upper elevational limit based on known specimens is not clear because of imprecise locality data, but is less than
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFAAFFBFE78F77153" box="[1338,1431,1087,1113]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="11" pageNumber="343" unit="m" value="1360.0">1,360 m</quantity>
|
|||
|
|
, the peak of Gongala Mountain. The
|
|||
|
|
<typeStatus id="426F8848767DA15CFDA5FBA27FF27177" box="[560,658,1123,1149]" pageId="11" pageNumber="343" type="holotype">holotype</typeStatus>
|
|||
|
|
and USNM
|
|||
|
|
<typeStatus id="426F8848767DA15CFCB7FBA27EED7177" box="[802,909,1123,1149]" pageId="11" pageNumber="343" type="paratype">paratypes</typeStatus>
|
|||
|
|
and USNM referred specimens were collected more than 40 years ago. The available field collection data do not indicate whether these USNM specimens were found in forest or in agricultural habitats, both of which occur in the general area today. The more-recently collected
|
|||
|
|
<typeStatus id="426F8848767DA15CFF02FB0E7D9771E3" box="[151,247,1231,1257]" pageId="11" pageNumber="343" type="paratype">paratype</typeStatus>
|
|||
|
|
specimens from Deniyaya were found when unearthed by mechanical diggers clearing an area of tea plantation for a car park. The new species is likely to qualify for Data Deficient status under IUCN Red List criteria (
|
|||
|
|
<bibRefCitation id="F9454B1B767DA15CFF0AFAD67F1B703B" author="IUCN Species Survival Commission & SC" box="[159,635,1303,1329]" pageId="11" pageNumber="343" refId="ref16624" refString="IUCN Species Survival Commission (SC) (2012) IUCN Red List categories and criteria. Version 3.1. 2 Edition. IUCN, Gland and Cambridge, 32 pp." type="book" year="2012">IUCN Species Survival Commission 2012</bibRefCitation>
|
|||
|
|
), at least until new field surveys are undertaken to better clarify its range and any potential threats, and/or until additional specimens can be found in other collections.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52FA9E78277303" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
Superficially similar
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFE27FA9E7F447073" authorityName="Hemprich" authorityYear="1820" box="[434,548,1375,1401]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFE27FA9E7F447073" box="[434,548,1375,1401]" italics="true" pageId="11" pageNumber="343">Rhinophis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
specimens have been seen (LJMW, pers. obs.) but not collected ca.
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFA87FA9F78387073" box="[1298,1368,1374,1401]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="343" unit="km" value="15.0">15 km</quantity>
|
|||
|
|
to the South of the
|
|||
|
|
<typeStatus id="426F8848767DA15CFEB1FA427C347097" box="[292,340,1411,1437]" pageId="11" pageNumber="343">type</typeStatus>
|
|||
|
|
locality of
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFE44FA427F5A7096" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[465,570,1411,1436]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFE44FA427F5A7096" box="[465,570,1411,1436]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFDD5FA427FF87097" bold="true" box="[576,664,1411,1437]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFDD5FA427FF87097" box="[576,664,1411,1437]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, at Rammale (ca.
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFCF5FA437ED97097" box="[864,953,1410,1437]" degrees="6.25" direction="north" orientation="latitude" pageId="11" pageNumber="343" precision="555" value="6.25">6.25° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFC56FA4279477097" box="[963,1063,1411,1437]" degrees="80.64" direction="east" orientation="longitude" pageId="11" pageNumber="343" precision="555" value="80.64">80.64° E</geoCoordinate>
|
|||
|
|
; peak up to ca.
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFB45FA4278797097" box="[1232,1305,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="343" unit="m" value="800.0">800 m</quantity>
|
|||
|
|
), and ca.
|
|||
|
|
<quantity id="5A2C9B0F767DA15CFA15FA437DDB70CB" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="343" unit="km" value="25.0">25 km</quantity>
|
|||
|
|
to the West of the
|
|||
|
|
<typeStatus id="426F8848767DA15CFE1EFA667CDB70CB" box="[395,443,1447,1473]" pageId="11" pageNumber="343">type</typeStatus>
|
|||
|
|
locality, at Dorana Ella (
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFD47FA667E4D70CB" box="[722,813,1447,1473]" degrees="6.43" direction="north" orientation="latitude" pageId="11" pageNumber="343" precision="555" value="6.43">6.43° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFCADFA667EFD70CB" box="[824,925,1447,1473]" degrees="80.42" direction="east" orientation="longitude" pageId="11" pageNumber="343" precision="555" value="80.42">80.42° E</geoCoordinate>
|
|||
|
|
), Waddagala (
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFBAEFA6679F670CB" box="[1083,1174,1447,1473]" degrees="6.46" direction="north" orientation="latitude" pageId="11" pageNumber="343" precision="555" value="6.46">6.46° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFB34FA66786570CB" box="[1185,1285,1447,1473]" degrees="80.43" direction="east" orientation="longitude" pageId="11" pageNumber="343" precision="555" value="80.43">80.43° E</geoCoordinate>
|
|||
|
|
), Lankagama (
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFF0AFA0A7D9970EF" box="[159,249,1483,1509]" degrees="6.38" direction="north" orientation="latitude" pageId="11" pageNumber="343" precision="555" value="6.38">6.38° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFE96FA0B7C0770EF" box="[259,359,1482,1509]" degrees="80.45" direction="east" orientation="longitude" pageId="11" pageNumber="343" precision="555" value="80.45">80.45° E</geoCoordinate>
|
|||
|
|
), and Mederipitiya (
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFDD3FA0B7FC070EF" box="[582,672,1482,1509]" degrees="6.35" direction="north" orientation="latitude" pageId="11" pageNumber="343" precision="555" value="6.35">6.35° N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="F8E0502D767DA15CFD3FFA0A7E6F70EF" box="[682,783,1483,1509]" degrees="80.49" direction="east" orientation="longitude" pageId="11" pageNumber="343" precision="555" value="80.49">80.49° E</geoCoordinate>
|
|||
|
|
). These places are indicated on the map in
|
|||
|
|
<figureCitation id="05EF2A6F767DA15CFB76FA0A784670EF" box="[1251,1318,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1935]" captionTargetBox="[273,1315,185,1881]" captionTargetId="figure-31@3.[266,1321,181,1887]" captionTargetPageId="3" captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)." figureDoi="http://doi.org/10.5281/zenodo.11547474" httpUri="https://zenodo.org/record/11547474/files/figure.png" pageId="11" pageNumber="343">Fig. 2</figureCitation>
|
|||
|
|
. However, detailed comparisons have not yet been made, and the identities of those populations remain as yet unclear.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="D5CE6561767DA15CFF52F9D2794972D7" pageId="11" pageNumber="343" type="discussion">
|
|||
|
|
<paragraph id="9D6B36EA767DA15CFF52F9D2794972D7" blockId="11.[151,1437,150,2013]" pageId="11" pageNumber="343">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF52F9D27C5B7327" bold="true" box="[199,315,1555,1581]" pageId="11" pageNumber="343">Remarks.</emphasis>
|
|||
|
|
<bibRefCitation id="F9454B1B767DA15CFED1F9D27F5B7327" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." box="[324,571,1555,1581]" pageId="11" pageNumber="343" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
|||
|
|
Sampaio
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFE3BF9D27C887327" box="[430,488,1555,1581]" italics="true" pageId="11" pageNumber="343">et al.</emphasis>
|
|||
|
|
(2023)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
included DNA sequences for
|
|||
|
|
<specimenCount id="8BD2FD63767DA15CFC01F9D2792A7327" box="[916,1098,1555,1581]" count="3" pageId="11" pageNumber="343" type="generic">three specimens</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFBE7F9D279BE7326" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[1138,1246,1555,1580]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFBE7F9D279BE7326" box="[1138,1246,1555,1580]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFB72F9D278227327" bold="true" box="[1255,1346,1555,1581]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFB72F9D278227327" box="[1255,1346,1555,1581]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
in their phylogenetic analyses, labelled as “
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFDA0F9F67FB2735B" box="[565,722,1591,1617]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFDA0F9F67FC7735B" box="[565,679,1591,1617]" italics="true" pageId="11" pageNumber="343">Rhinophis</emphasis>
|
|||
|
|
sp.
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Deniyaya)”. There are no morphological vouchers for those specimens; these sequences were obtained from tissues of parts of three different and partial animals that were found alongside the three complete
|
|||
|
|
<typeStatus id="426F8848767DA15CFE4CF9BE7F597393" box="[473,569,1663,1689]" pageId="11" pageNumber="343" type="paratype">paratype</typeStatus>
|
|||
|
|
specimens collected in 2018. Sampaio
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFC7FF9BE79417393" box="[1002,1057,1663,1689]" italics="true" pageId="11" pageNumber="343">et al.</emphasis>
|
|||
|
|
’s (2023) analyses recovered (with maximal support)
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFEF0F9627CAF73B6" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[357,463,1699,1724]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFEF0F9627CAF73B6" box="[357,463,1699,1724]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFE40F9627F4F73B7" bold="true" box="[469,559,1699,1725]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFE40F9627F4F73B7" box="[469,559,1699,1725]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
as most-closely related to
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFCCFF9627E9D73B7" authorityName="Deraniyagala" authorityYear="1975" box="[858,1021,1699,1725]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="tricoloratus">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFCCFF9627E9D73B7" box="[858,1021,1699,1725]" italics="true" pageId="11" pageNumber="343">R. tricoloratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
among sampled taxa, with these two species in a maximally supported clade with
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFD0CF9067E5373EB" box="[665,819,1735,1761]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFD0CF9067E6B73EB" box="[665,779,1735,1761]" italics="true" pageId="11" pageNumber="343">Rhinophis</emphasis>
|
|||
|
|
sp.
|
|||
|
|
</taxonomicName>
|
|||
|
|
(from Palabaddala),
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFBB1F90679D873EB" baseAuthorityName="Hemprich" baseAuthorityYear="1820" box="[1060,1208,1735,1761]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="homolepis">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFBB1F90679D873EB" box="[1060,1208,1735,1761]" italics="true" pageId="11" pageNumber="343">R. homolepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFB6DF90678F573EB" authorityName="Gower" authorityYear="2020" box="[1272,1429,1735,1761]" class="Squamata" family="Uropeltidae" genus="Rhinophis" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFB6DF906787173EA" box="[1272,1297,1735,1760]" italics="true" pageId="11" pageNumber="343">R.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFAABF90678F573EB" box="[1342,1429,1735,1761]" italics="true" pageId="11" pageNumber="343">mendisi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The Palabaddala specimen differs from
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFDC2F92A7FA1720E" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[599,705,1771,1796]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFDC2F92A7FA1720E" box="[599,705,1771,1796]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFD5DF92A7E42720F" bold="true" box="[712,802,1771,1797]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFD5DF92A7E42720F" box="[712,802,1771,1797]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
in having a longer rostral shield, and a dark dorsum and pale venter. The material identified as
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFDD8F8CE7F8A7223" authorityName="Gower" authorityYear="2020" box="[589,746,1807,1833]" class="Squamata" family="Uropeltidae" genus="Rhinophis" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="mendisi">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFDD8F8CE7F067222" box="[589,614,1807,1832]" italics="true" pageId="11" pageNumber="343">R.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFD06F8CE7F8A7223" box="[659,746,1807,1833]" italics="true" pageId="11" pageNumber="343">mendisi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by
|
|||
|
|
<bibRefCitation id="F9454B1B767DA15CFC82F8CE7ED77223" author="Gower, D. J." box="[791,951,1807,1833]" pageId="11" pageNumber="343" pagination="495 - 510" refId="ref16321" refString="Gower, D. J. (2020) A new species of Rhinophis Hemprich, 1820 (Serpentes: Uropeltidae) from southwestern Sri Lanka. Zootaxa, 4810 (3), 495 - 510. https: // doi. org / 10.11646 / zootaxa. 4810.3.6" type="journal article" year="2020">Gower (2020)</bibRefCitation>
|
|||
|
|
is from Rakwana (not far from the known localities of the new species:
|
|||
|
|
<figureCitation id="05EF2A6F767DA15CFE4CF8F27F7A7247" box="[473,538,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1935]" captionTargetBox="[273,1315,185,1881]" captionTargetId="figure-31@3.[266,1321,181,1887]" captionTargetPageId="3" captionText="FIGURE 2. Map showing places relevant to the distribution of Rhinophis martin sp. nov., including the type locality of “Hayes/ Gongala” and paratype localities of Hayes, Hayes/Gongala and Burusgala. White circles indicate localities for R. martin sp. nov. verified by voucher specimens; black circles indicate localities at which animals superficially resembling the new species have been seen but not collected or examined in detail (see text for details)." figureDoi="http://doi.org/10.5281/zenodo.11547474" httpUri="https://zenodo.org/record/11547474/files/figure.png" pageId="11" pageNumber="343">Fig. 2</figureCitation>
|
|||
|
|
) but differs from
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFD49F8F27E257246" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[732,837,1843,1868]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFD49F8F27E257246" box="[732,837,1843,1868]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFCDFF8F27EC37247" bold="true" box="[842,931,1843,1869]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFCDFF8F27EC37247" box="[842,931,1843,1869]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
in that it has a darker dorsum than venter, and a more crest-like dorsal ridge on the rostral shield. The
|
|||
|
|
<specimenCount id="8BD2FD63767DA15CFC93F8967EDB727B" box="[774,955,1879,1905]" count="3" pageId="11" pageNumber="343" type="generic">three specimens</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFC75F896792A727A" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[992,1098,1879,1904]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFC75F896792A727A" box="[992,1098,1879,1904]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFBC4F89679CB727B" bold="true" box="[1105,1195,1879,1905]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFBC4F89679CB727B" box="[1105,1195,1879,1905]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
sampled by
|
|||
|
|
<bibRefCitation id="F9454B1B767DA15CFAAFF8967C40729F" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." pageId="11" pageNumber="343" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
|||
|
|
Sampaio
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFF02F8BA7DAF729F" box="[151,207,1915,1941]" italics="true" pageId="11" pageNumber="343">et al.</emphasis>
|
|||
|
|
(2023)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
have identical DNA sequences, which differ notably from those of their closest-known relatives (
|
|||
|
|
<tableCitation id="D0560351767DA15CFAC8F8BA7DC772B3" captionStart="TABLE 3" captionStartId="8.[151,239,368,392]" captionTargetPageId="8" captionText="TABLE 3. Uncorrected p-distances (%) for intra- and interspecific comparisons for Rhinophis martin sp. nov. and closest sampled relatives, for two nuclear (prlr, cmos) and four mitochondrial genes (12s, 16s, cytb, nd4) as analysed by Sampaio et al. (2023). Values are means, with ranges in parentheses (where there is variation), and number of pairwise comparisons in square brackets. “–“ indicates insufficient data for pairwise comparison." pageId="11" pageNumber="343">Table 3</tableCitation>
|
|||
|
|
). These
|
|||
|
|
<specimenCount id="8BD2FD63767DA15CFE91F85E7CD672B3" box="[260,438,1951,1977]" count="3" pageId="11" pageNumber="343" type="generic">three specimens</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="5AD44D69767DA15CFE4CF85E7F2272B2" authority="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe, 2024" authorityName="Gower & Sampaio & Vidanapathirana & Mendis Wickramasinghe" authorityYear="2024" box="[473,578,1951,1976]" class="Squamata" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" pageId="11" pageNumber="343" phylum="Chordata" rank="species" species="martin" status="sp. nov.">
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFE4CF85E7F2272B2" box="[473,578,1951,1976]" italics="true" pageId="11" pageNumber="343">R. martin</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFDD2F85E7FC072B3" bold="true" box="[583,672,1951,1977]" pageId="11" pageNumber="343">
|
|||
|
|
<taxonomicNameLabel id="B4935783767DA15CFDD2F85E7FC072B3" box="[583,672,1951,1977]" pageId="11" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
were inferred to be distinct from all other sampled taxa at the species level in all of the species delimitation analyses reported by
|
|||
|
|
<bibRefCitation id="F9454B1B767DA15CFCBAF802794572D7" author="Sampaio, F. L. & Day, J. J. & Wickramasinghe, L. J. M. & Cyriac, V. P. & Papadopoulou, A. & Brace, S. & Rajendran, A. & Simon-Nutbrown, C. & Flouris, T. & Kapli, P. & Vidanapathirana, D. R. & Kotharambath, R. & Kodandaramaiah, U. & Gower, D. J." box="[815,1061,1987,2013]" pageId="11" pageNumber="343" pagination="107651" refId="ref16965" refString="Sampaio, F. L., Day, J. J., Wickramasinghe, L. J. M., Cyriac, V. P., Papadopoulou, A., Brace, S., Rajendran, A., Simon-Nutbrown, C., Flouris, T., Kapli, P., Vidanapathirana, D. R., Kotharambath, R., Kodandaramaiah, U. & Gower, D. J. (2023) A nearcomplete species-level phylogeny of uropeltid snakes harnessing historical museum collections as a DNA source. Molecular Phylogenetics & Evolution, 178, 107651. https: // doi. org / 10.1016 / j. ympev. 2022.107651" type="journal article" year="2023">
|
|||
|
|
Sampaio
|
|||
|
|
<emphasis id="AFA0EAF8767DA15CFC0DF8027EB072D7" box="[920,976,1987,2013]" italics="true" pageId="11" pageNumber="343">et al.</emphasis>
|
|||
|
|
(2023)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|