<documentID-CLB-Dataset="9619"ID-DOI="http://dx.doi.org/10.3897/zookeys.988.56188"ID-GBIF-Dataset="a38f6648-1c49-486b-a7ee-a877315d4e09"ID-PMC="PMC7666111"ID-Pensoft-Pub="1313-2970-988-63"ID-Pensoft-UUID="D5BC323AA5F05894AE8FBB10A3F0960D"ID-PubMed="33223891"ID-ZooBank="6E64D69BDD734A7EAE2B3CD21247A5E3"ModsDocID="1313-2970-988-63"checkinTime="1604753996846"checkinUser="pensoft"docAuthor="Li, Ya, Lin, Yucheng & Li, Shuqiang"docDate="2020"docId="C3F99341FBE1513EB9BB6C5D8548D745"docLanguage="en"docName="ZooKeys 988: 63-128"docOrigin="ZooKeys 988"docSource="http://dx.doi.org/10.3897/zookeys.988.56188"docTitle="Crassignatha nantou Y. Lin & S. Li 2020, sp. nov."docType="treatment"docUuid="B01CEA82-FFB1-41DD-919C-6AC0B8D0EF59"docUuidSource="ZooBank"docVersion="6"id="D5BC323AA5F05894AE8FBB10A3F0960D"lastPageNumber="63"masterDocId="D5BC323AA5F05894AE8FBB10A3F0960D"masterDocTitle="A review of Crassignatha (Araneae, Symphytognathidae)"masterLastPageNumber="128"masterPageNumber="63"pageNumber="63"updateTime="1732684104256"updateUser="ExternalLinkService">
<mods:affiliationid="E9643B305A319EE39F34ADF3F599443D">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China & The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<mods:affiliationid="7C05D42C9E404CC17499BFC17AD19A22">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China & The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<taxonomicNameid="56184430A5609BAA460095B6C98633AA"ID-CoL="98ZF6"LSID="C3F99341-FBE1-513E-B9BB-6C5D8548D745"authority="Y. Lin & S. Li"authorityName="Y. Lin & S. Li"authorityYear="2020"class="Arachnida"family="Symphytognathidae"genus="Crassignatha"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Crassignatha nantou"order="Araneae"pageId="0"pageNumber="63"phylum="Arthropoda"rank="species"species="nantou"status="sp. nov.">Crassignatha nantou Y. Lin & S. Li</taxonomicName>
<figureCitationid="22FF6C29153369CE9BAB903EDF2E7331"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Crassignatha nantou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)."figureDoi="10.3897/zookeys.988.56188.figure14"httpUri="https://binary.pensoft.net/fig/472599"pageId="0"pageNumber="63">Figs 14</figureCitation>
<figureCitationid="7F70EF188EB875805947E3B27275E9B4"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Crassignatha nantou sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure15"httpUri="https://binary.pensoft.net/fig/472600"pageId="0"pageNumber="63">, 15</figureCitation>
<figureCitationid="0FE9FEED77C2D1962DBE33CE3F834D99"captionStart="Figure 38"captionStartId="F38"captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi."figureDoi="10.3897/zookeys.988.56188.figure38"httpUri="https://binary.pensoft.net/fig/472623"pageId="0"pageNumber="63">, 38</figureCitation>
; 2757 m), 2.VII.2013, S. Li and G. Zheng leg.; 1♂ (NHMSU-HA055) and 1♀ (NHMSU-HA055) used for sequencing, GenBank: MT991996 and MT991995, same data as for preceding.
<taxonomicNameid="F2629E81521D2CED82A2590308677051"authorityName="Y. Lin & S. Li"authorityYear="2020"class="Arachnida"family="Symphytognathidae"genus="Crassignatha"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Crassignatha nantou"order="Araneae"pageId="0"pageNumber="63"phylum="Arthropoda"rank="species"species="nantou">
<figureCitationid="C30FFD699F22C5BB90289801F84C3D0D"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Crassignatha nantou sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure15"httpUri="https://binary.pensoft.net/fig/472600"pageId="0"pageNumber="63">15A, B, F, G</figureCitation>
sp. nov. by the shape of the male palp and vulva, but the male can be distinguished by having a blunt cymbial tooth and a hook on the median apophysis vs. a sharp cymbial tooth and lack of a hook in the latter (Figs
<figureCitationid="DA507F054C57447C27221CAC50E3D6CA"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Crassignatha nantou sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure15"httpUri="https://binary.pensoft.net/fig/472600"pageId="0"pageNumber="63">15A</figureCitation>
<figureCitationid="95CD11E640B155ADE0408BC1FF6280E0"captionStart="Figure 29"captionStartId="F29"captionText="Figure 29. Crassignatha shunani sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure29"httpUri="https://binary.pensoft.net/fig/472614"pageId="0"pageNumber="63">29A</figureCitation>
<figureCitationid="3E84A37573E48F68028CFCDF1209F598"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Crassignatha nantou sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure15"httpUri="https://binary.pensoft.net/fig/472600"pageId="0"pageNumber="63">15F, G</figureCitation>
<figureCitationid="45BD59244012B6274BA09011B883EC8E"captionStart="Figure 29"captionStartId="F29"captionText="Figure 29. Crassignatha shunani sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure29"httpUri="https://binary.pensoft.net/fig/472614"pageId="0"pageNumber="63">29F, G</figureCitation>
<taxonomicNameid="7474D0C0913BD579B529CF56D04CBA55"authorityName="Y. Lin & S. Li"authorityYear="2020"class="Arachnida"family="Symphytognathidae"genus="Crassignatha"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Crassignatha nantou"order="Araneae"pageId="0"pageNumber="63"phylum="Arthropoda"rank="species"species="nantou">
<figureCitationid="599C24C5267DAEFAAA284FC9F88A4649"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Crassignatha nantou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)."figureDoi="10.3897/zookeys.988.56188.figure14"httpUri="https://binary.pensoft.net/fig/472599"pageId="0"pageNumber="63">14A-C</figureCitation>
<figureCitationid="39532872DA4F8CAE97F548088EA806DF"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Crassignatha nantou sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure15"httpUri="https://binary.pensoft.net/fig/472600"pageId="0"pageNumber="63">15A-C</figureCitation>
): pale, weakly sclerotized. Cymbium large, with a few setae distally, cymbial tooth near distal margin. Tegulum large, smooth, and plump. Median apophysis disciform, with a hooked process on margin. Embolic membrane slender, laminar, translucent, arises near anterior part of median apophysis. Embolus long, flexible, spiraled, and wide basally, narrow distally.
<figureCitationid="C7C2C879EC27EC30F0F2501A7339C77E"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Crassignatha nantou sp. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)."figureDoi="10.3897/zookeys.988.56188.figure14"httpUri="https://binary.pensoft.net/fig/472599"pageId="0"pageNumber="63">14D-F</figureCitation>
: carapace nearly pear shaped, sculptured, and granular. Cephalic region lower than in male. PER slightly recurved. Chelicerae fused near base, covered with setae anteriorly. Labium subtriangular, fused to sternum. Sternum as in male.
<figureCitationid="E7D65BEA7BA724B9BE76CF78DE48D668"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Crassignatha nantou sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)."figureDoi="10.3897/zookeys.988.56188.figure15"httpUri="https://binary.pensoft.net/fig/472600"pageId="0"pageNumber="63">15D-G</figureCitation>
): epigynal area lightly sclerotized, with setae on lateral margins. Scape short, wide, copulatory opening located at its terminus, split into two labella. Internal structures more or less visible via translucent tegument. Paired spermathecae globose, separated by half their diameter. Fertilization ducts thin, starting at inside central margin of spermathecae, bent downward, twisted, and furcate at end. Copulatory ducts thick, connected to posterior margin of spermathecae, passing under the spermathecae, up into the center of vulva, deflexed to copulatory opening, their proximal base unfused.
<taxonomicNameid="E755292FBCF18025CC4E6339137EF685"authorityName="Y. Lin & S. Li"authorityYear="2020"class="Arachnida"family="Symphytognathidae"genus="Crassignatha"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Crassignatha nantou"order="Araneae"pageId="0"pageNumber="63"phylum="Arthropoda"rank="species"species="nantou">
<paragraphid="B64D9048CD95CB0F80A433467F256EEB"pageId="0"pageNumber="63">The specific name is derived from the type locality; noun in apposition.</paragraph>
<figureCitationid="4A79FD17229D7D4A42ACC6685D13FE84"captionStart="Figure 38"captionStartId="F38"captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi."figureDoi="10.3897/zookeys.988.56188.figure38"httpUri="https://binary.pensoft.net/fig/472623"pageId="0"pageNumber="63">38</figureCitation>