<documentID-CLB-Dataset="3753"ID-DOI="http://dx.doi.org/10.3897/zookeys.1067.72369"ID-GBIF-Dataset="1165ef89-9b15-4c8a-a7a3-5a80ffae5903"ID-PMC="PMC8571248"ID-Pensoft-Pub="1313-2970-1067-19"ID-Pensoft-UUID="34D3B870A92C50819DE85FF5DDE19A1D"ID-PubMed="34759718"ID-ZooBank="AEAF44EE2CCD4E319CBCA4C5ACB0E2EB"ModsDocID="1313-2970-1067-19"checkinTime="1635566834375"checkinUser="pensoft"docAuthor="Srisonchai, Ruttapon, Sutcharit, Chirasak & Likhitrakarn, Natdanai"docDate="2021"docId="B86C57C06EFC5888AC19C847D7394223"docLanguage="en"docName="ZooKeys 1067: 19-56"docOrigin="ZooKeys 1067"docPubDate="2021-10-29"docSource="http://dx.doi.org/10.3897/zookeys.1067.72369"docTitle="Zephronia siamensis Hirst 1907"docType="treatment"docVersion="6"id="34D3B870A92C50819DE85FF5DDE19A1D"lastPageNumber="19"masterDocId="34D3B870A92C50819DE85FF5DDE19A1D"masterDocTitle="The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae)"masterLastPageNumber="56"masterPageNumber="19"pageNumber="19"updateTime="1732748198608"updateUser="ExternalLinkService">
<mods:titleid="72BC0BB1F936412D86E56D776C3C1D5B">The giant pill-millipede genus Zephronia Gray, 1832 from Thailand, with a redescription of Z. siamensis Hirst, 1907 and descriptions of three new species (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
<mods:affiliationid="334BFDFFD8431F23C3FF59DA0EA087C5">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
<mods:affiliationid="8999C2433BB10CF85290B4FDA92ADD6A">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand & Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
<figureCitationid="507E378ADEF8A24C2912E8E9FCEAF11C"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Photographs of living Zephronia spp. A-D Zephronia siamensis E, F Zephronia phrain. Photographs not to scale."figureDoi="10.3897/zookeys.1067.72369.figure1"httpUri="https://binary.pensoft.net/fig/604301"pageId="0"pageNumber="19">Figures 1A-D</figureCitation>
<figureCitationid="911C63B7F1CF1C207095209503C02D35"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">; 3</figureCitation>
<figureCitationid="C4786AB987397415CE1E7305FF3D4107"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Zephronia siamensis A-C left anterior telopod, anterior, posterior and sublateral views, respectively D, E Left and right posterior telopods, posterior view F Posterior telopod, anterior view. Abbreviations: cr-T = crenulated teeth, cx = coxa, ML = membranous lobe, sp = sclerotized process, syn-cx = syncoxite."figureDoi="10.3897/zookeys.1067.72369.figure4"httpUri="https://binary.pensoft.net/fig/604304"pageId="0"pageNumber="19">; 4</figureCitation>
<figureCitationid="4F79048A2295C43154889F06BDE853FA"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. SEM of endoterga of body ring 7, all from ventral views A, B Zephronia siamensis Hirst, 1907 C, D Zephronia enghoffi sp. nov. E, F Zephronia golovatchi sp. nov. G, H Zephronia panhai sp. nov. Abbreviations: cp = cuticular impression, ms = middle section, is = inner section, os = outer section, pm = posterior margin."figureDoi="10.3897/zookeys.1067.72369.figure13"httpUri="https://binary.pensoft.net/fig/604313"pageId="0"pageNumber="19">; 13A, B</figureCitation>
<figureCitationid="B67EB18B2D378166E3FDDCE424FC735D"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. SEM of endoterga in body ring 7, all from ventral views A Zephronia siamensis Hirst, 1907 B Zephronia enghoffi sp. nov. C Zephronia golovatchi sp. nov. D Zephronia panhai sp. nov. E inner area of endotergum in Zephronia panhai sp. nov."figureDoi="10.3897/zookeys.1067.72369.figure14"httpUri="https://binary.pensoft.net/fig/604314"pageId="0"pageNumber="19">; 14A</figureCitation>
<bibRefCitationid="79B2B21F581B6E3DA07767CACDDC74C0"author="Attems, C"journalOrPublisher="Memoirs of the Indian Museum"pageId="0"pageNumber="19"pagination="133 - 323"refId="B2"refString="Attems, C, 1936. Diplopoda of India. Memoirs of the Indian Museum 11: 133 - 323"title="Diplopoda of India."volume="11"year="1936">Attems 1936</bibRefCitation>
<bibRefCitationid="DA21A7B36EB056245539209A059822E5"DOI="https://doi.org/10.15298/arthsel.21.4.01"author="Golovatch, SI"journalOrPublisher="Arthropoda Selecta"pageId="0"pageNumber="19"pagination="273 - 294"refId="B9"refString="Golovatch, SI, Wesener, T, Mauries, J-P, Semenyuk, II, 2012. On the identities of Cryxus Leach, 1814 and Zephronia Gray, 1832, the oldest generic names in the millipede order Sphaerotheriida (Diplopoda). Arthropoda Selecta 21 (4): 273 - 294, DOI: https://doi.org/10.15298/arthsel.21.4.01"title="On the identities of Cryxus Leach, 1814 and Zephronia Gray, 1832, the oldest generic names in the millipede order Sphaerotheriida (Diplopoda)."url="https://doi.org/10.15298/arthsel.21.4.01"volume="21"year="2012">Golovatch et al. 2012</bibRefCitation>
<bibRefCitationid="A0ACA7A7B81C8DB3E74021C2AF0E7175"DOI="https://doi.org/10.11646/zootaxa.3220.1.2"author="Wongthamwanich, N"journalOrPublisher="Zootaxa"pageId="0"pageNumber="19"pagination="29 - 43"refId="B32"refString="Wongthamwanich, N, Panha, S, Sierwald, P, Wesener, T, Thirakhupt, K, 2012a. A new species of the giant pill-millipede genus Sphaerobelum Verhoeff, 1924 from northern Thailand, with an extensive description and molecular characters (Diplopoda: Sphaerotheriida: Zephroniidae). Zootaxa 3220: 29 - 43, DOI: https://doi.org/10.11646/zootaxa.3220.1.2"title="A new species of the giant pill-millipede genus Sphaerobelum Verhoeff, 1924 from northern Thailand, with an extensive description and molecular characters (Diplopoda: Sphaerotheriida: Zephroniidae)."url="https://doi.org/10.11646/zootaxa.3220.1.2"volume="3220"year="2012 a">Wongthamwanich et al. 2012a</bibRefCitation>
<bibRefCitationid="D184702B93C14588D343D010129EE72B"author="Wesener, T"journalOrPublisher="Bonn Zoological Bulletin Supplementum"pageId="0"pageNumber="19"pagination="1 - 104"refId="B28"refString="Wesener, T, 2016. The giant pill-millipedes, order Sphaerotheriida - An annotated species catalogue with morphological atlas and list of apomorphies (Arthropoda: Diplopoda). Bonn Zoological Bulletin Supplementum 63: 1 - 104"title="The giant pill-millipedes, order Sphaerotheriida - An annotated species catalogue with morphological atlas and list of apomorphies (Arthropoda: Diplopoda)."volume="63"year="2016">Wesener 2016</bibRefCitation>
<bibRefCitationid="A1A8696B3747BC7014C5044ADB6C3D53"author="Decker, P"journalOrPublisher="Schubartiana"pageId="0"pageNumber="19"pagination="23 - 34"publicationUrl="http://www.schubartiana.de/issues/pdf/vol4/Decker-2010-Myriapod_fauna_of_Thailand.pdf"refId="B6"refString="Decker, P, 2010. Contributions to the myriapod fauna of Thailand-New records of millipedes and centipedes from Thailand (Myriapoda: Diplopoda, Chilopoda). Schubartiana 4: 23 - 34, http://www.schubartiana.de/issues/pdf/vol4/Decker-2010-Myriapod_fauna_of_Thailand.pdf"title="Contributions to the myriapod fauna of Thailand-New records of millipedes and centipedes from Thailand (Myriapoda: Diplopoda, Chilopoda)."url="http://www.schubartiana.de/issues/pdf/vol4/Decker-2010-Myriapod_fauna_of_Thailand.pdf"volume="4"year="2010">Decker 2010</bibRefCitation>
<materialsCitationid="9467927F7C8603C06736DFE67412E451"collectingDate="2020-11-14"collectorName="R. Srisonchai, N. Likhitrakarn, P. Jirapatrasilp"country="Thailand"elevation="7"latitude="13.151055"location="Koh Sichang"longLatPrecision="15"longitude="100.81556"municipality="Sichang District"specimenCount="19"specimenCount-female="17"specimenCount-male="2"stateProvince="Chonburi Province">
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<materialsCitationid="DF8AA14A6B378E2B0DD8CEE8B3DD99F3"collectingDate="2016-07-04"collectorName="R. Srisonchai, P. Tongkerd"country="Thailand"elevation="128"latitude="13.208805"location="Province"longLatPrecision="1"longitude="101.65159"specimenCount="2"specimenCount-female="2"stateProvince="Chonburi">
<bibRefCitationid="6C8B9EF7ED2F7520692D95BD31E3E118"DOI="https://doi.org/10.1080/00222930709487327"author="Hirst, AS"journalOrPublisher="Arthropoda Selecta"pageId="0"pageNumber="19"refId="B10"refString="Hirst, AS, 1907. On four new pill-millipedes from the Malay Peninsula and Siam. Annals and Magazine of Natural History (series 7) 20: 215-219. https://doi.org/10.1080/00222930709487327"title="On four new pill-millipedes from the Malay Peninsula and Siam. Annals and Magazine of Natural History (series 7) 20: 215 - 219."url="https://doi.org/10.1080/00222930709487327"year="1907">Hirst 1907</bibRefCitation>
organ located next to the aberrant ommatidia, not inside the antennal groove. Adult body length relatively small, usually ca. 20 mm, <26.5 mm, tip of subanal plate concave, process of telopoditomere 2 of anterior telopod rather short and strongly curved distally, and process of telopoditomere 2 of anterior telopod shorter than the combination of telopoditomeres 3 and 4. Similar in these respects to
Attems, 1953. Differs from these two species by showing a body length> 16.7 mm, live specimens with unique dark green, tergites with two yellow-brown patches located in anterior half of tergites, surface of tergites with conspicuous setae, femur of leg extended with conspicuous teeth, and telopoditomere 4 of anterior telopods posteriorly with a row of conspicuous crenulated teeth (cr-T).
: Male: body length 15.0-26.5 mm. Width of thoracic shield 9.0-12.5 mm, of tergite 8 9.5-11.4 mm. Height of thoracic shield 5.2-6.4 mm, of tergite 7, 6.1-7.4 mm. Female: body length 15-23.0 mm. Width of thoracic shield ca. 12.1 mm, of tergite 8 ca. 12.8 mm. Height of thoracic shield ca. 7.3 mm, of tergite 7 ca. 8.2 mm.
<figureCitationid="23F0418C204854F43CD162DD01A7C1D7"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Photographs of living Zephronia spp. A-D Zephronia siamensis E, F Zephronia phrain. Photographs not to scale."figureDoi="10.3897/zookeys.1067.72369.figure1"httpUri="https://binary.pensoft.net/fig/604301"pageId="0"pageNumber="19">1A-D</figureCitation>
): Live specimens dark green; antennae dark brown; head, collum, thoracic shield, paratergites brown; legs bluish green. Tergites with two big patches, brown or yellowish brown, arranged in almost central part of anterior half; lateral part of tergites greenish dark, middle part of tergites brown. Anal shield with two colors contrasting each other, posterior half reddish brown, anterior half greenish dark brown. Color in alcohol after three months of preservation changed to greenish brown, head and collum dark greenish, tergites with a dark posterior margin, legs pale yellowish, distal podomeres rusty brown, antennae dark green.
<taxonomicNameid="C63B9B94015996448E7744FD281F9E4C"authorityName="Likhitrakarn & Golovatch"authorityYear="2021"class="Diplopoda"family="Zephroniidae"genus="Zephronia"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Zephronia phrain"order="Sphaerotheriida"pageId="0"pageNumber="19"phylum="Arthropoda"rank="species"species="phrain">
: Trapezoid, anterior part of head clothed with numerous long setae, posterior part sparsely punctate; anterior margin of labrum with a single tooth. Each eye with 75-92 ommatidia. Aberrant ocellus located inside antennal groove.
<figureCitationid="18532F1C1DD968CC0B33A58AA3E68C54"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">3A</figureCitation>
<figureCitationid="77D0A5BA9EC1BB7E3BC6D4FFE638501D"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">3C</figureCitation>
<figureCitationid="FE4D93FC0E871748B25E0CE559AB9063"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Photographs of living Zephronia spp. A-D Zephronia siamensis E, F Zephronia phrain. Photographs not to scale."figureDoi="10.3897/zookeys.1067.72369.figure1"httpUri="https://binary.pensoft.net/fig/604301"pageId="0"pageNumber="19">1A-C</figureCitation>
<figureCitationid="CFA3BC512FD1FD900E9C1D8E2FCF8F5B"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. SEM of endoterga of body ring 7, all from ventral views A, B Zephronia siamensis Hirst, 1907 C, D Zephronia enghoffi sp. nov. E, F Zephronia golovatchi sp. nov. G, H Zephronia panhai sp. nov. Abbreviations: cp = cuticular impression, ms = middle section, is = inner section, os = outer section, pm = posterior margin."figureDoi="10.3897/zookeys.1067.72369.figure13"httpUri="https://binary.pensoft.net/fig/604313"pageId="0"pageNumber="19">13A, B</figureCitation>
<figureCitationid="409EC9A3E11C41343B9B68308D545D4D"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. SEM of endoterga in body ring 7, all from ventral views A Zephronia siamensis Hirst, 1907 B Zephronia enghoffi sp. nov. C Zephronia golovatchi sp. nov. D Zephronia panhai sp. nov. E inner area of endotergum in Zephronia panhai sp. nov."figureDoi="10.3897/zookeys.1067.72369.figure14"httpUri="https://binary.pensoft.net/fig/604314"pageId="0"pageNumber="19">14A</figureCitation>
margin. Inner section (inner area) smooth, with a few setae. Middle section (middle area) with a single row of conspicuous, elliptical cuticular impressions; distance between impressions as wide as individual diameter. Bristles arranged in one row, tip of the longest bristles not extended beyond posterior margin or not reaching to posterior margin.
: Sexually dimorphic, in female large and well-rounded, in male slightly more rectangular, in both sexes glabrous. Surface similar to that of tergites. Inner surface (underside) with a single, long, black locking carina half as long as width of last laterotergite.
<figureCitationid="875451AB19B83B559172979C5D4AF77A"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">3B</figureCitation>
): Leg-pair 1 with one ventral spine, leg-pair 2 with two or three, leg-pair 3 with 4-6 ventral spines. Leg-pairs 4-18 with eight or nine ventral spines and two or three apical ones; thereafter slightly reduced into 5-8 ventral spines. In leg 9, femur ca. 1.7
longer than wide. Length of tarsus> femur> prefemur> coxa> tibia ≥ postfemur. All podomeres densely setose. Coxa large, with dentate ridge marginally (coxal process). Coxal process absent in leg-pairs 1 and 2. Prefemur without teeth. Femur large and stout, extended mesally, with 7-11 conspicuous teeth.
<figureCitationid="599D62A96F6B06E8A877CF3C11831D80"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">3F</figureCitation>
): Large and wide, semicircular, divided by a conspicuous mesal constriction; central margin (tip) concave, wide; lateral margin slightly convex. Densely setose.
<figureCitationid="D2081BFC769011F16F6B584601E69581"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">3D</figureCitation>
): First telopoditomere rectangular, slightly longer than wide. Telopoditomere 2 large, as long as telopoditomeres 3 and 4 combined. Process of telopoditomere 2 located posteriorly, but partly visible laterally in anterior view. Process of telopoditomere 2 wide, broader than telopoditomeres 3 and 4. Process of telopoditomere 2 conspicuously unciform, protruding as high as basal part of telopoditomere 4, apically with a well-rounded tip. Margin towards telopoditomere 3 with a membranous area carrying a rather small and sclerotized process (sp), apically with a rounded tip. Telopoditomere 3 slender, 1.4X longer than wide, 1.5X longer than telopoditomere 4. Telopoditomere 4 posteriorly with a row of 7 small and crenulated teeth (cr-T) with two prominent spines. All podomeres covered with long sparse setae, except for central part of telopoditomere 1 and posterior surfaces of 2-4.
): Inner horns with sharply edged tips, slightly curved caudad. Telopodite consisting of four podomeres. First podomere stout and narrow, nearly twice as wide as long. Immovable finger (process of telopoditomere 2) shorter than movable finger (consisting of telopoditomeres 3 and 4). Immovable finger stout and narrow, 1.6X longer than wide, not curved, glabrous distally. Margin towards movable finger with two massive, triangular, membranous lobes (ML). Telopoditomere 3 elongated, slightly curved, twice as long as telopoditomere 4; with a large, swollen, membranous ledge; postero-apically slightly enlarged, with a row of 11 or 12 crenulated teeth (cr-T). Telopoditomere 4 slender, twice as long as wide, slightly tapering apically; with a large, swollen, membranous ledge; with two long and sclerotized spines. Telopoditomeres 1 and 2 on both sides covered with few setae. Telopoditomere 3 at base of inner margin with a few setae, remaining parts of telopoditomeres 3 and 4 almost glabrous.
<figureCitationid="F8445FB1983B31AB2433495E22764728"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Zephronia siamensis A-C male D, E female (CUMZ-Zeph 0013) A right antenna, ventral view B the ninth left leg, posterior view C first coxae with stigmatic plates, posterior view D coxae of second legs with gonopores, posterior view E coxae and prefemur of second legs with vulvae, posterior view F subanal plate, ventral view. Abbreviations: cx = coxa, o = operculum, pre = prefemur, syn-cx = syncoxite, St-Pl = stigmatic plate."figureDoi="10.3897/zookeys.1067.72369.figure3"httpUri="https://binary.pensoft.net/fig/604303"pageId="0"pageNumber="19">3E</figureCitation>
): Vulva large, covering ca. 2/3 coxa, located at mesal side, extending mesally to basal third of prefemur. Operculum regularly rounded, margin straight, mesal margin not protruding.
<figureCitationid="BEB58BA1CA06E560788FC142DC4D4B40"captionStart="Figure 15"captionStartId="F15"captionText="Figure 15. Limestone habitats of some Zephronia spp. A type locality of Zephronia siamensis (Koh Srichang, Chonburi Province) B, C type locality of Zephronia enghoffi sp. nov. (Tham Phaya Nakharat, Khon Kaen Province) D Habitat of Zephronia phrain at Phawor shrine, Tak Province."figureDoi="10.3897/zookeys.1067.72369.figure15"httpUri="https://binary.pensoft.net/fig/604315"pageId="0"pageNumber="19">15A</figureCitation>
trees in a limestone area near a beach, while the other material from the Chonburi and Rayong provinces were likewise taken from limestone habitats. Currently, this species is known to occur only in eastern Thailand.
<paragraphid="10E64B435633F87B4664ADEB220F86A2"pageId="0"pageNumber="19">The live coloration of adults is generally dark green with two yellowish brown patches in the anterior half of tergites, this being quite unique for this species.</paragraph>
<bibRefCitationid="7C8C68352C1FDB6AC7BE7B91FDE499BB"DOI="https://doi.org/10.1080/00222930709487327"author="Hirst, AS"journalOrPublisher="Arthropoda Selecta"pageId="0"pageNumber="19"refId="B10"refString="Hirst, AS, 1907. On four new pill-millipedes from the Malay Peninsula and Siam. Annals and Magazine of Natural History (series 7) 20: 215-219. https://doi.org/10.1080/00222930709487327"title="On four new pill-millipedes from the Malay Peninsula and Siam. Annals and Magazine of Natural History (series 7) 20: 215 - 219."url="https://doi.org/10.1080/00222930709487327"year="1907">Hirst (1907)</bibRefCitation>
compared to the topotypes: the type specimen was ca. 26.5 mm in length, whereas the new material we examined were within the size range of 16.7-23.5 mm.
