<documentid="466866F37DB17F687AD5957879133973"ID-CLB-Dataset="300057"ID-DOI="10.5252/geodiversitas2024v46a11"ID-GBIF-Dataset="a66a476e-10ee-4341-bafc-aa201e82e1dc"ID-ISSN="1638-9395"ID-Zenodo-Dep="12780247"ID-ZooBank="urn:lsid:zoobank.org:pub:558A3726-9C34-4C34-9FA2-C49B510BEDC7"IM.bibliography_approvedBy="guilherme"IM.illustrations_approvedBy="guilherme"IM.materialsCitations_approvedBy="guilherme"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="guilherme"IM.treatmentCitations_approvedBy="guilherme"IM.treatments_approvedBy="guilherme"checkinTime="1721351604131"checkinUser="plazi"docAuthor="Psarras, Christos, Merle, Didier, Moissette, Pierre & Koskeridou, Efterpi"docDate="2024"docId="03F43052FFA3A663FC7FF9403458F9E4"docLanguage="en"docName="geodiversitas.2024.46.11.pdf"docOrigin="Geodiversitas 46 (11)"docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a11.pdf"docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article"docStyleId="F830B10FF475E64C1F1601E3B32DDC00"docStyleName="Geodiversitas.2018-.journal_article"docStyleVersion="4"docTitle="Conus (Monteiroconus) tsampikaensis Psarras, Merle, Moissette & Koskeridou, 2024, n. sp."docType="treatment"docUuid="63A46EA6-1DD1-4454-8091-EA1866BD5D64"docUuidSource="ZooBank"docVersion="2"lastPageNumber="437"masterDocId="FFCD482AFFAEA673FFFEFFF23034FF88"masterDocTitle="The Pleistocene Conidae (Mollusca: Gastropoda) from the island of Rhodes (Greece) and their palaeoecological significance"masterLastPageNumber="444"masterPageNumber="423"pageNumber="434"updateTime="1721656402093"updateUser="ExternalLinkService"zenodo-license-document="CC0-1.0"zenodo-license-figures="CC0-1.0">
<mods:titleid="6C58EB681EEA8E0766EB9560E4AEF60B">The Pleistocene Conidae (Mollusca: Gastropoda) from the island of Rhodes (Greece) and their palaeoecological significance</mods:title>
<mods:affiliationid="B8F25B04CA07C7B121A044412964798E">National and Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Hist. Geology-Paleontology, University Campus 15784 Athens (Greece) cpsarras @ geol. uoa. gr (corresponding author)</mods:affiliation>
<mods:affiliationid="926C9E8729F3B16431FF3AF1A83834F9">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr</mods:affiliation>
<mods:affiliationid="5E261B57AB10F873BB15BE40546BFA82">National and Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Hist. Geology-Paleontology, University Campus 15784 Athens (Greece) pmoissette @ geol. uoa. gr</mods:affiliation>
<mods:affiliationid="CAACFFB9C6DB83EBBAD6A77BB9224C0C">National and Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Hist. Geology-Paleontology, University Campus 15784 Athens (Greece) ekosker @ geol. uoa. gr</mods:affiliation>
<taxonomicNameid="4C5DFAC7FFA3A67EFC7FF940352DF945"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"box="[897,1305,1714,1741]"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="13"pageNumber="434"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<treatmentCitationid="0AFCA755FFA3A67EFC01F8913530F8F3"author="CHIRLI C. & LINSE U."box="[1023,1284,1891,1915]"page="166"pageId="13"pageNumber="434"year="2011">
<bibRefCitationid="EFCCFCB5FFA3A67EFC01F8913530F8F3"author="CHIRLI C. & LINSE U."box="[1023,1284,1891,1915]"pageId="13"pageNumber="434"refId="ref13061"refString="CHIRLI C. & LINSE U. 2011. - The Pleistocene Marine Gastropods of Rhodes Island (Greece). Verlag Documenta Naturae, 447 p."type="book"year="2011">Chirli & Linse 2011: 166</bibRefCitation>
</treatmentCitation>
, pl. 57, figs 1a-d (non
<bibRefCitationid="EFCCFCB5FFA3A67EFC9AF88F33C6F81E"author="BROCCHI G."box="[868,1010,1917,1942]"pageId="13"pageNumber="434"pagination="1 - 2"refId="ref12435"refString="BROCCHI G. 1814. - Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Stamperia Reale, Milano, 1 - 2, 712 p. https: // doi. org / 10.5962 / bhl. title. 11569"type="book chapter"year="1814">Brocchi, 1814</bibRefCitation>
<taxonomicNameid="4C5DFAC7FFA0A67DFF2AF9F4323BF99F"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"box="[212,527,1542,1559]"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="14"pageNumber="435"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<figureCitationid="13669DC1FFA0A67DFD36F97332CDF911"box="[712,761,1665,1689]"captionStart="FIG"captionStartId="5.[132,143,1872,1889]"captionTargetBox="[132,1456,212,1836]"captionTargetId="figure-23@5.[698,1267,785,1347]"captionTargetPageId="5"captionText="FIG. 1. — Information about the studied section of Tsampika in Rhodes, Greece: A, general tectonic setting of the southern Aegean. B, geological map of Rhodes with the site of Tsampika as a blue star (*); C, satellite position of the studied section; D, geological map of Tsampika area, displaying the section (*), the cross section of Cornée et al. (2006) (red line a-b) and the location of the section of Cornée et al. (2018) (black arrow); E, cross section of the Ladiko-Tsampika Formation in Tsampika that was studied by Cornée et al. (2006), highlighting the area of study with a blue arrow; F, synthetic stratigraphic column of the section with the Conidae specimens recovered from TSS a. Notice that TSS d and TSS gravels (above arrow) were not visible on the figured section (Fig. 2), but were identified in nearby smaller sections. Maps A and B were modified from Cornée et al. 2018; D-E from Cornée et al. 2006. TSS beds belong to the TSS1 sequence of Cornée et al. (2006, 2018)."figureDoi="http://doi.org/10.5281/zenodo.12780249"httpUri="https://zenodo.org/record/12780249/files/figure.png"pageId="14"pageNumber="435">Fig.1</figureCitation>
<figureCitationid="13669DC1FFA0A67DFF2BF939313AF96B"box="[213,270,1739,1763]"captionStart="FIG"captionStartId="6.[132,143,1582,1599]"captionTargetBox="[132,1456,194,1557]"captionTargetId="figure-162@6.[131,795,782,1256]"captionTargetPageId="6"captionText="FIG. 2. — Small section of Tsampika along the new road on the south of Tsampika beach:A, the section is located in front of a palaeo-cliff of Jurassic limestones and has a small inclination; B, the section is divided into three units, from the base to the top: 1, sandy marl with cerithids and a rich malacofauna (= TSS a bed of Fig. 1F); 2, grey marl unit rich in Cladocora (= TSS b bed of Fig. 1F); 3, silty to sandy grey marl bed (= TSS c bed of Fig. 1F); numbers indicate the position of the associated faunas;C, detailed view of the palaeo-cliff perforated by the bivalve Lithophaga; D, in situ coral branches of Cladocora caespitosa; E, assemblage with Hexaplex (Trunculariopsis) trunculus (Linnaeus, 1758) (2), Cerithium vulgatum Bruguière, 1792 (1) and Ostrea edulis Linnaeus, 1758 (3) (hammer for scale); F, Assemblage with Hexaplex (Trunculariopsis) trunculus (Linnaeus, 1758) (2), Euthria cornea (Linnaeus, 1758) (4), and Bolma rugosa (Linnaeus, 1767) (5). Scale bar: F, c. 4 cm."figureDoi="http://doi.org/10.5281/zenodo.12780253"httpUri="https://zenodo.org/record/12780253/files/figure.png"pageId="14"pageNumber="435">Fig. 2</figureCitation>
), with low angled coeloconoid spire. Early spire whorls coeloconoid, slightly elevated with three to four spiral grooves on sutural ramp.Later spire whorls flatten with coeloconoid sutural ramps. Shoulder angulated, with maximum diameter below shoulder. Subsutural flexure moderately deep to shallow, strongly curved, moderately asymmetrical. Last whorl elongated, conical.
<taxonomicNameid="4C5DFAC7FFA1A67CFE87F9E9328EF9A4"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"box="[377,698,1563,1580]"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="15"pageNumber="436"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<paragraphid="8BE28144FFA1A67CFF7AF8C3342BF884"blockId="15.[131,777,1841,2027]"lastBlockId="15.[814,1457,1682,1804]"pageId="15"pageNumber="436">The colour pattern on the spire whorls consists of thin axial flammulae. The pattern on the last whorl consists of two levels of pigmentation. The first one consists of two broad spiral bands, separated by a wide non-fluorescent band along the anterior part of the last whorl. The second level of pigmentation consists of spirally and axially arranged fluorescent dots, dashes and small flakes interrupted by non-fluorescent bars or small blotches. The second level of pigmentation persists on the fluorescent bands and the non-fluorescent areas.</paragraph>
<bibRefCitationid="EFCCFCB5FFBEA663FF32FEA5319DFEF9"author="MONTEIRO A. & TENORIO M. J. & POPPE G. T."box="[204,425,343,369]"pageId="16"pageNumber="437"refId="ref16728"refString="MONTEIRO A., TENORIO M. J. & POPPE G. T. 2004. - A conchological Iconography. The family Conidae. The West African and Mediterranean species of Conus. Conchbooks, Hackenheim, 102 p."type="book"year="2004">
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) and thus differs from the specimens from Tsampika. In addition, the pattern of the last whorl of
is usually pale and consists of a purple-cream ground colour (
<bibRefCitationid="EFCCFCB5FFBEA663FE3CFE2432ACFE78"author="MONNIER E. & LIMPALAER L. & ROBIN A. & ROUX C."box="[450,664,470,496]"pageId="16"pageNumber="437"refId="ref16651"refString="MONNIER E., LIMPALAER L., ROBIN A. & ROUX C. 2018 a. - A Taxonomic Iconography of the Living Conidae. Vol. 1. ConchBooks, Hackenheim, 604 p."type="book"year="2018">
), although it has rarely been observed to have evenly distributed spiral rows and two slightly darker spiral bands in the middle and anterior part of the last whorl (Reeve 1844: pl. 54, fig. 244; see also
<bibRefCitationid="EFCCFCB5FFBEA663FF24FDA43189FDF8"author="MONNIER E. & LIMPALAER L. & ROBIN A. & ROUX C."box="[218,445,598,624]"pageId="16"pageNumber="437"refId="ref16651"refString="MONNIER E., LIMPALAER L., ROBIN A. & ROUX C. 2018 a. - A Taxonomic Iconography of the Living Conidae. Vol. 1. ConchBooks, Hackenheim, 604 p."type="book"year="2018">
<taxonomicNameid="4C5DFAC7FFBEA663FDE8FD843124FD38"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="16"pageNumber="437"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<taxonomicNameid="4C5DFAC7FFBEA663FF65FCC43227FCC7"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"box="[155,531,821,848]"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="16"pageNumber="437"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<taxonomicNameid="4C5DFAC7FFBEA663FD39FC8731EBFC47"authority="Pavia & Dulai & Festa & Gennari & Pavia & Carnevale, 2022"authorityName="Pavia & Dulai & Festa & Gennari & Pavia & Carnevale"authorityYear="2022"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="16"pageNumber="437"phylum="Mollusca"rank="species"species="villalvernensis"subGenus="Monteiroconus">
<bibRefCitationid="EFCCFCB5FFBEA663FE3AFC6731EBFC47"author="PAVIA G. & DULAI A. & FESTA A. & GENNARI R. & PAVIA M. & CARNEVALE G."pageId="16"pageNumber="437"pagination="129 - 210"refId="ref17085"refString="PAVIA G., DULAI A., FESTA A., GENNARI R., PAVIA M. & CARNEVALE G. 2022. - Palaeontology of the Upper Pliocene marine deposits of Rio Vaccaruzza, Villalvernia (Piedmont, NW Italy). Rivista Italiana Di Paleontologia e Stratigrafia 128 (1): 129 - 210. https: // doi. org / 10.54103 / 2039 - 4942 / 15178"type="journal article"year="2022">Pavia & Dulai & Festa & Gennari & Pavia & Carnevale, 2022</bibRefCitation>
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have a higher spire and elevated spire whorls, so we do not consider them conspecific to the Rhodes material.
<bibRefCitationid="EFCCFCB5FFBEA663FDC1FB86317FFB26"author="JUAREZ J. & MATAMALES-ANDREU R."pageId="16"pageNumber="437"pagination="39 - 67"refId="ref14825"refString="JUAREZ J. & MATAMALES-ANDREU R. 2016. - Taxons inedits o poc coneguts per al Pleistoce superior litoral de Mallorca (Illes Balears, Mediterrani occidental) i consideracions sobre alguns jaciments. Societat d'Historia Natural de Les Balears 59: 39 - 67. https: // raco. cat / index. php / BolletiSHNBalears / article / view / 336147"type="journal article"year="2016">Juárez & Matamales-Andreu (2016)</bibRefCitation>
, a species originally described from West Africa, from the Tyrrhenian (MIS 5e) of Mallorca. This species usually lacks a colour pattern or displays fine and irregular spiral stripes, or occasionally some subquadrate blotches on an overall white background. This pattern is very different from that observed on the Rhodes specimens.
<bibRefCitationid="EFCCFCB5FFBEA663FDD6FAA13112FA05"author="JUAREZ J. & MATAMALES-ANDREU R."pageId="16"pageNumber="437"pagination="39 - 67"refId="ref14825"refString="JUAREZ J. & MATAMALES-ANDREU R. 2016. - Taxons inedits o poc coneguts per al Pleistoce superior litoral de Mallorca (Illes Balears, Mediterrani occidental) i consideracions sobre alguns jaciments. Societat d'Historia Natural de Les Balears 59: 39 - 67. https: // raco. cat / index. php / BolletiSHNBalears / article / view / 336147"type="journal article"year="2016">Juárez & Matamales-Andreu (2016)</bibRefCitation>
did not describe a residual colour pattern on the Mallorcan material.
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. Temperature ranges (in Celsius degrees) of the four species. Map and temperatures modified from
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;
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<bibRefCitationid="EFCCFCB5FFBEA663FF7AF9813174F904"author="ZAZO C. & GOY J. L. & DABRIO C. J. & LARIO J. & GONZALEZ-DELGADO J. A. & BARDAJI T. & HILLAIRE-MARCEL C. & CABERO A. & GHALEB B. & BORJA F. & SILVA P. G. & ROQUERO E. & SOLER V."box="[132,320,1650,1677]"pageId="16"pageNumber="437"pagination="36 - 49"refId="ref19298"refString="ZAZO C., GOY J. L., DABRIO C. J., LARIO J., GONZALEZ-DELGADO J. A., BARDAJI T., HILLAIRE-MARCEL C., CABERO A., GHALEB B., BORJA F., SILVA P. G., ROQUERO E. & SOLER V. 2013. - Retracing the Quaternary history of sea-level changes in the Spanish Mediterranean-Atlantic coasts: Geomorphological and sedimentological approach, Geomorphology in Spain. Special issue in honour of Prof. Mateo Gutierrez. Geomorphology 196: 36 - 49. https: // doi. org / 10.1016 / j. geomorph. 2012.10.020"type="book chapter"year="2013">
<bibRefCitationid="EFCCFCB5FFBEA663FF17F9613153F924"author="BORN I. VON"box="[233,359,1682,1709]"pageId="16"pageNumber="437"refId="ref12376"refString="BORN I. VON 1778. - Index rerum naturalium Musei Caesarei Vindobonensis. Verzeichniss der Naturlichen Seltenheiten des K. K. Naturalien Kabinets zu Wien. Erster Theil, Schalthiere Pars 1 Testacea. Vindobonae: ex officina Krausiana, 596 p. https: // doi. org / 10.5962 / bhl. title. 11581"type="book"year="1778">
<bibRefCitationid="EFCCFCB5FFBEA663FEE1F94031EFF945"author="MUHS D. R. & SIMMONS K. R. & MECO, J. & PORAT N."box="[287,475,1714,1741]"pageId="16"pageNumber="437"pagination="408 - 424"refId="ref16830"refString="MUHS D. R., SIMMONS K. R., MECO, J. & PORAT N. 2015. - Uranium-series ages of fossil corals from Mallorca, Spain: The " Neotyrrhenian " high stand of the Mediterranean Sea revisited. Palaeogeography, Palaeoclimatology, Palaeoecology 438: 408 - 424. https: // doi. org / 10.1016 / j. palaeo. 2015.06.043"type="journal article"year="2015">
) is also questionable, as no one has yet shown any of the fossils under UV light.
<taxonomicNameid="4C5DFAC7FFBEA663FD39F92031FAF884"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="16"pageNumber="437"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<bibRefCitationid="EFCCFCB5FFBEA663FF65F8A331BCF8E4"author="CHIRLI C. & LINSE U."box="[155,392,1873,1900]"pageId="16"pageNumber="437"refId="ref13061"refString="CHIRLI C. & LINSE U. 2011. - The Pleistocene Marine Gastropods of Rhodes Island (Greece). Verlag Documenta Naturae, 447 p."type="book"year="2011">Chirli & Linse (2011)</bibRefCitation>
found specimens resembling
<taxonomicNameid="4C5DFAC7FFBEA663FD3AF8A031F0F803"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="16"pageNumber="437"phylum="Mollusca"rank="species"species="tsampikaensis"status="sp. nov."subGenus="Monteiroconus">
<taxonomicNameid="4C5DFAC7FFBEA663FF79F8233215F863"authorityName="Psarras & Merle & Moissette & Koskeridou"authorityYear="2024"box="[135,545,2001,2027]"class="Gastropoda"family="Conidae"genus="Conus"kingdom="Animalia"order="Neogastropoda"pageId="16"pageNumber="437"phylum="Mollusca"rank="species"species="tsampikaensis"subGenus="Monteiroconus">
). The presence of this species in Arkhangelos Fm, which is certainly older than Tsampika (
<bibRefCitationid="EFCCFCB5FFBEA663FCC8F9E13433F9A5"author="CORNEE J. - J. & QUILLEVERE F. & MOISSETTE P. & FIETZKE J. & LOPEZ-OTALVARO G. E. & MELINTE-DOBRINESCU M. & PHILIPPON M. & VAN HINSBERGEN D. J. J. & AGIADI K. & KOSKERIDOU E. & MUNCH P."box="[822,1031,1554,1581]"pageId="16"pageNumber="437"pagination="78 - 96"refId="ref13222"refString="CORNEE J. - J., QUILLEVERE F., MOISSETTE P., FIETZKE J., LOPEZ-OTALVARO G. E., MELINTE-DOBRINESCU M., PHILIPPON M., VAN HINSBERGEN D. J. J., AGIADI K., KOSKERIDOU E. & MUNCH P. 2018. - Tectonic motion in oblique subduction forearcs: insights from the revisited Middle and Upper Pleistocene deposits of Rhodes, Greece. Journal of the Geological Society 176 (1): 78 - 96. https: // doi. org / 10.1144 / jgs 2018 - 090"type="journal article"year="2018">
