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161 lines
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<document ID-DOI="http://doi.org/10.5281/zenodo.6103250" ID-GBIF-Dataset="f0d443f2-b8b9-46fa-a1aa-2d0f0d3c8748" ID-GBIF-Taxon="114923360" ID-ISSN="1175-5326" ID-Zenodo-Dep="6103250" ID-ZooBank="CEAB4AA5-C4F8-437E-A2F4-D82F383579C1" checkinTime="1447288886152" checkinUser="donat" docAuthor="Taylor, Robert W." docDate="2015" docId="03B3878D9B0FFFFBFF534EA69A1EFF92" docLanguage="en" docName="29627.pdf.imf" docOrigin="Zootaxa 3947 (2)" docTitle="Heteroponera trachypyx Taylor, 2015, sp. n." docType="treatment" docVersion="5" lastPageNumber="174" masterDocId="FF8AFFF59B18FFECFFC44D279F4AF867" masterDocTitle="Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus" masterLastPageNumber="180" masterPageNumber="151" pageId="23" pageNumber="174" updateTime="1645000973839" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Taylor, Robert W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>3947</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>151</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3947.2.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="ZooBank">CEAB4AA5-C4F8-437E-A2F4-D82F383579C1</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6103250" ID-GBIF-Taxon="114923360" ID-Zenodo-Dep="6103250" LSID="urn:lsid:plazi:treatment:03B3878D9B0FFFFBFF534EA69A1EFF92" httpUri="http://treatment.plazi.org/id/03B3878D9B0FFFFBFF534EA69A1EFF92" lastPageNumber="174" pageId="23" pageNumber="174">
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<subSubSection pageId="23" pageNumber="174" type="nomenclature">
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<paragraph box="[151,547,897,922]" pageId="23" pageNumber="174">
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<emphasis bold="true" box="[151,547,897,922]" pageId="23" pageNumber="174">
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<heading bold="true" box="[151,547,897,922]" fontSize="11" level="1" pageId="23" pageNumber="174" reason="1">
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8.
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<emphasis bold="true" box="[180,474,897,922]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName box="[180,474,897,922]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="trachypyx" status="sp. n.">Heteroponera trachypyx</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[481,547,897,922]" pageId="23" pageNumber="174" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</emphasis>
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</paragraph>
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<paragraph box="[158,309,932,956]" pageId="23" pageNumber="174">
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(
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<figureCitation box="[166,297,932,956]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,263,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." pageId="23" pageNumber="174">Figs 42–45</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="174" type="materials_examined">
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<paragraph box="[151,1134,1003,1029]" pageId="23" pageNumber="174">
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<emphasis bold="true" box="[151,481,1004,1029]" pageId="23" pageNumber="174">Type locality. AUSTRALIA</emphasis>
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: QUEENSLAND:
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<emphasis box="[699,895,1004,1029]" italics="true" pageId="23" pageNumber="174">McDowall Range</emphasis>
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(16°06'S, 145°20'E).
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</paragraph>
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<paragraph box="[199,885,1041,1066]" pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,394,1041,1066]" pageId="23" pageNumber="174">Type deposition.</emphasis>
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Holotype ANIC. Paratypes ANIC, QMBA.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="174" type="distribution">
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<paragraph pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,588,1077,1101]" pageId="23" pageNumber="174">Distribution, material examined.</emphasis>
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Known only from the following accessions (grid cells -15/145, -16/145 and -17/145). Unless otherwise indicated each a single worker. All samples collected (unless noted) by Berlese funnel sampling of rainforest leafmould or litter (usually “sieved”).
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<emphasis bold="true" box="[834,1072,1149,1173]" pageId="23" pageNumber="174">EA1 (Mt Finnigan):</emphasis>
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<emphasis box="[1079,1230,1149,1173]" italics="true" pageId="23" pageNumber="174">Shiptons Flat</emphasis>
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, -15 48, 145 14, 2 workers (Davies & Raven, 16–21.xi.1975).
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<emphasis box="[654,841,1186,1210]" italics="true" pageId="23" pageNumber="174">Mt Finnigan SF,</emphasis>
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-15 48, 145 17, moist litter pocket, 400 m (S. & J. Peck, 3.vii.1982).
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<emphasis bold="true" box="[365,643,1221,1245]" pageId="23" pageNumber="174">EA2 (Thornton Peak):</emphasis>
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<emphasis box="[654,966,1221,1245]" italics="true" pageId="23" pageNumber="174">12 Mile Scrub, Gap Creek</emphasis>
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, -15 50, 145 20 (Davies & Raven, 22– 28.xi.1975).
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<emphasis box="[296,639,1257,1282]" italics="true" pageId="23" pageNumber="174">3.5 km W of Cape Tribulation</emphasis>
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, Site 7, -16 0 5, 145 27, QMBA berlesate 440, 680m (GBM, DKY & GIT, 2.x.1982).
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<emphasis box="[346,550,1293,1317]" italics="true" pageId="23" pageNumber="174">McDowall Range</emphasis>
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, -16 0 6, 145 20, QMBA berlesate 684, 520 m (GBM, 27.xi.1985)(Type Locality).
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<emphasis box="[269,388,1330,1353]" italics="true" pageId="23" pageNumber="174">Mt Sorrow</emphasis>
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, summit, -16 0 8, 145 26, 2 workers, QMBA berlesates 256, 262, 800 m (GBM, 15.x.1980).
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<emphasis box="[151,340,1365,1389]" italics="true" pageId="23" pageNumber="174">Table Mountain,</emphasis>
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-16 0 9, 145 26, 4 workers, QMBA berlesates 540, 542, 320m (GBM & DWC, 24.iv.1983).
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<emphasis box="[151,339,1401,1426]" italics="true" pageId="23" pageNumber="174">Thornton Range</emphasis>
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, -16 14, 145 26, 100 m (RWT & JDF, 24.vi.1971)—2 workers, 150–180 m (RWT & JDF, 23.vi.1971).
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<emphasis box="[294,374,1437,1461]" italics="true" pageId="23" pageNumber="174">Rex Ck</emphasis>
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, site 3, -16.28 4, 145 19 4, rainforest berlesate (C Burwell, 14 ii 2005).
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</paragraph>
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<paragraph pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,420,1473,1498]" pageId="23" pageNumber="174">Worker diagnosis.</emphasis>
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General characteristics, conformation, coloration and major sculpturation as illustrated. Diagnostic features as in relevant key couplets above.
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<taxonomicName box="[823,1105,1509,1533]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="trachypyx">
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<emphasis box="[823,1105,1509,1533]" italics="true" pageId="23" pageNumber="174">Heteroponera trachypyx</emphasis>
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</taxonomicName>
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is less darkly colored and sculptured than the other
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<emphasis box="[444,597,1546,1569]" italics="true" pageId="23" pageNumber="174">H. ecarinata-</emphasis>
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group species. Head and mesosoma more-or-less uniformly medium-dark brown, the gaster, mandibles, antennae and legs a shade lighter. Coloration overall similar to the sympatric species
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<emphasis box="[151,320,1617,1642]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName box="[151,320,1617,1642]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="rhodopygea">H. rhodopygea</taxonomicName>
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</emphasis>
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and
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<emphasis box="[378,522,1618,1641]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName box="[378,522,1618,1641]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="viviennae">H. viviennae</taxonomicName>
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</emphasis>
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. Propodeal declivity shining, microsculpture distinct to vestigial, with very fine radial microstriae. Dimensions: (holotype; smallest available specimen (12–mile Scrub), largest available specimen (Mt Sorrow summit)): TL (ca): 5.1, 4.4, 5.1; HW: 1.18, 1.11, 1.19; HL: 1.14, 1.06, 114; CI: 103, 104, 104; EL: 0.17, 0.16, 0.18; SL: 0.68, 0.61, 0.71; SI: 57, 57, 59; PW: 0.95, 0.84, 0.87; WL: 1.40, 1.40, 1.48; petH: 0.76, -, -; petW: 0.48, 0.52, 0.50; GW 1.04, 1.03, 1.04.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="23" pageNumber="174" type="description">
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<paragraph box="[199,1025,1797,1821]" pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,356,1798,1821]" pageId="23" pageNumber="174">Related taxa.</emphasis>
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<emphasis box="[363,504,1798,1821]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName box="[363,504,1798,1821]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="ecarinata">H. ecarinata</taxonomicName>
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</emphasis>
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and
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<taxonomicName box="[559,735,1797,1821]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="pendergrasti">
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<emphasis box="[559,735,1797,1821]" italics="true" pageId="23" pageNumber="174">H. pendergrasti</emphasis>
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</taxonomicName>
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at species-complex level.
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</paragraph>
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<paragraph pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,592,1833,1858]" pageId="23" pageNumber="174">In-group sympatric associations.</emphasis>
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Sympatric with
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<taxonomicName box="[784,954,1833,1858]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="rhodopygea">
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<emphasis box="[784,954,1833,1858]" italics="true" pageId="23" pageNumber="174">H. rhodopygea</emphasis>
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</taxonomicName>
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at Mt Finnigan, Mt Sorrow and Thornton Range, and
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<taxonomicName box="[284,451,1869,1893]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="rhodopygea">
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<emphasis box="[284,451,1869,1893]" italics="true" pageId="23" pageNumber="174">H. rhodopygea</emphasis>
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</taxonomicName>
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+
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<taxonomicName box="[481,622,1870,1893]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="viviennae">
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<emphasis box="[481,622,1870,1893]" italics="true" pageId="23" pageNumber="174">H. viviennae</emphasis>
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</taxonomicName>
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in the Cape tribulation area and Mc Dowall Range.
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</paragraph>
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<paragraph pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,371,1905,1930]" pageId="23" pageNumber="174">Biogeography.</emphasis>
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Known only from the northern EAs 1 & 2, distantly allopatric from the other known
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<emphasis box="[1319,1426,1906,1929]" italics="true" pageId="23" pageNumber="174">
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<taxonomicName box="[1319,1426,1906,1929]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="ecarinata">ecarinata</taxonomicName>
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</emphasis>
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- group species.
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,1268,1977,2002]" pageId="23" pageNumber="174" type="discussion">
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<paragraph box="[199,1268,1977,2002]" pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,315,1978,2001]" pageId="23" pageNumber="174">Remarks.</emphasis>
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Variation in coloration suggests a prolonged callow to sub-callow period in workers.
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,1364,2013,2037]" pageId="23" pageNumber="174" type="etymology">
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<paragraph box="[199,1364,2013,2037]" pageId="23" pageNumber="174">
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<emphasis bold="true" box="[199,333,2014,2037]" pageId="23" pageNumber="174">Etymology.</emphasis>
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The name, a noun in apposition, refers to the coarse sculpturation of the first gastral tergite.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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