<documentid="F1076492EA18C5F7BB07455A5F9BE50A"ID-DOI="10.12651/JSR.2022.11.1.029"ID-ISSN="2713-8615"ID-Zenodo-Dep="13140154"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1722399903568"checkinUser="felipe"docAuthor="Podenas, Sigitas, Park, Sun-Jae & Aukštikalnienė, Hye-Woo Byun and Rasa"docDate="2022"docId="038E434AFFA1665BD7A4FB4EFB0A293D"docLanguage="en"docName="JSpeciesRes.11.1.29-37.pdf"docOrigin="Journal of Species Research 11 (1)"docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article"docStyleId="4D398BFF45FF8CA9E08F546BB06AE372"docStyleName="JSpeciesRes.2012-.journal_article"docStyleVersion="2"docTitle="Prionolabis clavaria"docType="treatment"docVersion="2"lastPageNumber="34"masterDocId="FFB73B32FFA5665ED70AFFABFF832863"masterDocTitle="Prionolabis crane flies (Diptera: Limoniidae) of Korea"masterLastPageNumber="37"masterPageNumber="29"pageNumber="33"updateTime="1722400374751"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:affiliationid="769479D095CAC9B4A833A3041D759D2D">Nature Research Centre, Akademijos str. 2, LT- 08412 Vilnius, Lithuania & Life Sciences Center of Vilnius University, Sauletekio str. 7, LT- 10257 Vilnius, Lithuania</mods:affiliation>
<mods:affiliationid="2F3265DD098AF227BFC807D9B318724F">Animal Resources Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
<bibRefCitationid="EFB68FADFFA1665AD692FB4EFDD12C9E"author="Alexander, C. P."box="[408,594,1253,1277]"pageId="4"pageNumber="33"pagination="418 - 436"refId="ref6490"refString="Alexander, C. P. 1950. Undescribed species of Japanese crane-flies (Diptera: Tipulidae). Part VII. Annals of the Entomological Society of America 43: 418 - 436."type="journal article"year="1950">Alexander, 1950</bibRefCitation>
<treatmentCitationid="0A86D44DFFA1665AD7A4FAAEFC8B2D7E"author="Alexander, C. P."box="[174,776,1285,1309]"page="429"pageId="4"pageNumber="33"year="1950">
<bibRefCitationid="EFB68FADFFA1665AD528FAAEFC8B2D7E"author="Alexander, C. P."box="[546,776,1285,1309]"pageId="4"pageNumber="33"pagination="418 - 436"refId="ref6490"refString="Alexander, C. P. 1950. Undescribed species of Japanese crane-flies (Diptera: Tipulidae). Part VII. Annals of the Entomological Society of America 43: 418 - 436."type="journal article"year="1950">Alexander, 1950: 429</bibRefCitation>
<treatmentCitationid="0A86D44DFFA1665AD7A4FA8EFF602D3D"author="Savchenko, E. N. & G. O. Krivolutskaya"page="70"pageId="4"pageNumber="33"year="1976">
<taxonomicNameid="4C2789DFFFA1665AD7A4FA8EFF602D3D"ID-CoL="77WQY"authority="Savchenko & Krivolutskaya, 1976: 70"authorityName="Savchenko & Krivolutskaya"authorityPageNumber="70"authorityYear="1976"class="Insecta"family="Limoniidae"genus="Prionolabis"kingdom="Animalia"order="Diptera"pageId="4"pageNumber="33"phylum="Arthropoda"rank="species"species="clavaria">
<bibRefCitationid="EFB68FADFFA1665AD698FA8EFF602D3D"author="Savchenko, E. N. & G. O. Krivolutskaya"pageId="4"pageNumber="33"pagination="1 - 160"refId="ref7188"refString="Savchenko, E. N. and G. O. Krivolutskaya. 1976. Limoniidae of the south Kuril Islands and south Sakhalin. Akadimiya Nauk Ukrainian SSR, Kiev: 1 - 160 (in Russian)."type="book chapter"year="1976">Savchenko & Krivolutskaya, 1976: 70</bibRefCitation>
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;
<treatmentCitationid="0A86D44DFFA1665AD7E7FAEDFE5E2D3D"author="Savchenko, E. N."box="[237,477,1350,1374]"page="113"pageId="4"pageNumber="33"year="1989">
<bibRefCitationid="EFB68FADFFA1665AD7E7FAEDFE5E2D3D"author="Savchenko, E. N."box="[237,477,1350,1374]"pageId="4"pageNumber="33"pagination="1 - 377"refId="ref7125"refString="Savchenko, E. N. 1989. Komary-limoniidy fauny SSSR [Limoniidae fauna of the USSR]. Determination tables of superspecies taxa with catalogue survey of species. Akadimiya Nauk Ukrainian SSR, I. I. Schmalhausen Institute of Zoology of Academy of Sciences of Ukraine, Naukova Dumka, Kiev: 1 - 377 (in Russian)."type="book chapter"year="1989">Savchenko, 1989: 113</bibRefCitation>
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;
<bibRefCitationid="EFB68FADFFA1665AD6E2FAEDFD312D3D"author="Oosterbroek, P."box="[488,690,1350,1374]"pageId="4"pageNumber="33"refId="ref6850"refString="Oosterbroek, P. 2021. Catalogue of the Craneflies of the World (CCW). Available from https: // ccw. naturalis. nl /"type="url"year="2021">Oosterbroek, 2021</bibRefCitation>
Head: Dark brown, sparsely dusted with brownish, covered with blackish to yellowish setae, longer laterally, shorter dorsally. Distance between eyes in male approximately equals to the length of scape. Male antenna (
<figureCitationid="131CEED9FFA1665AD7F2F991FED32E31"box="[248,336,1594,1618]"captionStart="Fig"captionStartId="4.[839,872,1035,1057]"captionTargetBox="[841,1440,269,984]"captionTargetId="figure-654@4.[829,1448,249,1072]"captionTargetPageId="4"captionText="Fig. 3. Prionolabis clavaria (Alexander, 1950), male. A. antenna. B. wing. C. male genitalia in glycerol, dorsal view. D. male geni- talia slide-mounted, dorsal view, holotype. Scale bars: 0.5 mm (A, C, D), 1.0 mm (B)."figureDoi="http://doi.org/10.5281/zenodo.13140160"httpUri="https://zenodo.org/record/13140160/files/figure.png"pageId="4"pageNumber="33">Fig. 3A</figureCitation>
long, reaching frontal margin of prescutum if bent backwards. Scape elongate, nearly cylindrical, 1.6 times as long as pedicel, dark brown at base, black at apex, dusted with brownish gray and bearing few black semi-adjacent setae. Pedicel widening towards distal end, black, covered with black setae. Flagellum 14-segmented, entirely black. Basal flagel- lomere slightly elongate, less than twice as long as wide, 2-5 flagellomeres oval, distal flagellomeres slightly elongate, apical segment 1.3 times as long as preceding. Longest verticils up to 1.7 times as long as respective segment. Rostrum, palpus and mouth parts black, dusted
Thorax: Cervical sclerites, pronotum and mesonotal prescutum black, covered with brownish pruinosity and erect blackish to yellowish setae. Longitudinal stripes on mesonotum missing. Tubercular pits missing, pseudosutural fovea indistinct, wide and shallow. Scutal lobe black, pruinosity scarcer than on prescutum. Area between scutal lobes black and densely dusted. Scutellum black with dark reddish brown posterior apex. Mediotergite dark brown to black, sparsely pruinose. Pleuron black, more heavily dusted ventrally, sparsely variegated with semi-polished areas where the bloom has been denuded, katepisternum setoseless. Male wing (
<figureCitationid="131CEED9FFA1665AD445F9F0FC262E10"box="[847,933,1627,1651]"captionStart="Fig"captionStartId="4.[839,872,1035,1057]"captionTargetBox="[841,1440,269,984]"captionTargetId="figure-654@4.[829,1448,249,1072]"captionTargetPageId="4"captionText="Fig. 3. Prionolabis clavaria (Alexander, 1950), male. A. antenna. B. wing. C. male genitalia in glycerol, dorsal view. D. male geni- talia slide-mounted, dorsal view, holotype. Scale bars: 0.5 mm (A, C, D), 1.0 mm (B)."figureDoi="http://doi.org/10.5281/zenodo.13140160"httpUri="https://zenodo.org/record/13140160/files/figure.png"pageId="4"pageNumber="33">Fig. 3B</figureCitation>
) slightly iridescent, translucent, with brownish tinge, yellowish at base and costal area, without any darker spots except brown oval stigma. Veins brown, yellowish at wing base and in costal area. Venation:
. Stem of halter pale, widely rusty brown at base, knob brownish with pale base. Coxae black, densely dusted brownish gray, covered with long blackish to yellowish setae. Meron small, thus second and third pairs of coxae close to each other. Trochanters black, just posterior narrowly rusty brown ventrally. Femur dark brown, narrowly pale at base, black at apex. Tibia dark brown with black apex. Tarsus dark brown turning black towards distal end. Male femur I: 4.7-5.0 mm long, II: 5.0 mm, III: 5.5-6.0 mm, tibia I:
Abdomen: Very dark brown, nearly black, dusted with brown pollen. Erect setae covering abdominal sclerites blackish at base, pale at distal end. Paired transverse sutures on tergites black but narrow. Male terminalia (
<figureCitationid="131CEED9FFA0665BD792FC40FEA12C60"box="[152,290,1003,1027]"captionStart="Fig"captionStartId="4.[839,872,1035,1057]"captionTargetBox="[841,1440,269,984]"captionTargetId="figure-654@4.[829,1448,249,1072]"captionTargetPageId="4"captionText="Fig. 3. Prionolabis clavaria (Alexander, 1950), male. A. antenna. B. wing. C. male genitalia in glycerol, dorsal view. D. male geni- talia slide-mounted, dorsal view, holotype. Scale bars: 0.5 mm (A, C, D), 1.0 mm (B)."figureDoi="http://doi.org/10.5281/zenodo.13140160"httpUri="https://zenodo.org/record/13140160/files/figure.png"pageId="5"pageNumber="34">Fig. 3C, 3D</figureCitation>
) concolorous with the rest of abdomen. Ninth tergite wider than longer, posterior margin with U-shaped medial notch and blunt apexed lobules on both sides of it. Gonocoxite slightly elongate, 1.7 times as long as width at base, simple, without additional lobes. Outer gonostylus sclerotized, narrows distally, apex curved and hook-shaped, additional fleshy and setose lobe starting at base of dorsal surface and reaching to about two-thirds of gonostylus. Inner gonostylus elongate, sclerotized with bifid tip, outer branch longer with setose apex, inner branch rounded. Paramere long and narrow with needle-like apex that is turned downwards at a straight angle; very apex usually not visible in exactly dorsal view, thus in slide-mounted specimens only preapical part, that is rounded or with small apical notch, is visible. Aedeagus very long, narrows towards distal end, that is curved downwards.
<bibRefCitationid="EFB68FADFFA0665BD68FF97AFDC12E8A"author="Savchenko, E. N."box="[389,578,1745,1769]"pageId="5"pageNumber="34"pagination="1 - 156"refId="ref7039"refString="Savchenko, E. N. 1983. Limoniidae of South Primorye. Aka- demiy Nauk Ukrainskoy SSR, I. I. Schmalhausen Institute of Zoology of Academy of Sciences of Ukraine, Naukova Dumka, Kiev: 1 - 156 (in Russian)."type="book chapter"year="1983">Savchenko, 1983</bibRefCitation>
<materialsCitationid="3B4FF801FFA0665BD7ACF8F9FA02297E"collectingDate="1939-06-22"collectionCode="USNM"collectorName="A. M. Yankovsky"country="North Korea"county="Toorisani"elevation="1829"location="Examined"municipality="Puksu Pyaksan"pageId="5"pageNumber="34"specimenCount="2"specimenCount-male="2"typeStatus="holotype">
<figureCitationid="131CEED9FFA0665BD674F8F9FE572F09"box="[382,468,1874,1898]"captionStart="Fig"captionStartId="7.[258,291,728,750]"captionTargetBox="[145,1413,262,693]"captionTargetId="figure-521@7.[144,1414,262,694]"captionTargetPageId="7"captionText="Fig. 6. Distribution maps of Korean Prionolabis. A. P. acanthophora. B. P. clavaria. C. P. dis. D. P. yankovskyana."figureDoi="http://doi.org/10.5281/zenodo.13140166"httpUri="https://zenodo.org/record/13140166/files/figure.png"pageId="5"pageNumber="34">Fig. 6B</figureCitation>
<collectorNameid="26D2978AFFA0665BD49CFEAEFBD5297E"box="[918,1110,261,285]"pageId="5"pageNumber="34">A. M. Yankovsky</collectorName>
(
<collectionCodeid="ED366A99FFA0665BD369FEAEFB35297E"box="[1123,1206,261,285]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871"lsid="urn:lsid:biocol.org:col:34871"name="Smithsonian Institution, National Museum of Natural History"pageId="5"pageNumber="34"type="Museum">USNM</collectionCode>
<materialsCitationid="3B4FF801FFA0665BD423FE8EFB07293D"collectingDate="1938-07-10"collectionCode="USNM"collectorName="A. M. Yankovsky"country="North Korea"elevation="1158"location="Seren Mts."pageId="5"pageNumber="34"specimenCount="1"typeStatus="holotype">
<collectorNameid="26D2978AFFA0665BD462FEEDFBA3293E"box="[872,1056,325,350]"pageId="5"pageNumber="34">A. M. Yankovsky</collectorName>
(
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