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<mods:title id="5C28163EA1EB8102C2A95464D79C0C80">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="57B06EEC9C3E5A9D5451FEE2DEB1811A">Pacaud, Jean-Michel</mods:namePart>
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<mods:affiliation id="2EFBB6F0FD56A673218DB15A6528C020">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:date id="DDC1432F87CCCA2CF56D1A6DE855ED50">2024</mods:date>
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Genus 
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<emphasis id="B953EADEFFCFE42FFBADFF0EFBF4FF42" box="[1045,1141,216,241]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
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, 
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(
<figureCitation id="131C2A49FFCFE42FFB88FF21FBFDFEA2" box="[1072,1148,247,273]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann &amp; Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet &amp; Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="41" pageNumber="534">Fig. 22</figureCitation>
E-H)
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urn:lsid:zoobank.org:act: 
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. — 
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<emphasis id="B953EADEFFCFE42FFC61FEA8FBDFFE26" box="[985,1118,382,405]" italics="true" pageId="41" pageNumber="534">Murex pereger</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFBDCFEABFB70FE26" author="BEYRICH E." box="[1124,1265,381,405]" pageId="41" pageNumber="534" pagination="408 - 500" refId="ref42899" refString="BEYRICH E. 1854. - Die Conchylien des norddeutschen Tertiargebirges. Zeitschrift der Deutschen Geologischen Gesellschaft 6: 408 - 500, 726 - 781. https: // www. digitale-sammlungen. de / de / view / bsb 10283013" type="journal article" year="1854">Beyrich, 1854</bibRefCitation>
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, late Eocene to late Oligocene: Paris Basin, 
<collectingCountry id="F330765CFFCFE42FFBA9FE4EFBE7FE03" box="[1041,1126,408,432]" name="Belgium" pageId="41" pageNumber="534">Belgium</collectingCountry>
, Mainz Basin and north 
<collectingCountry id="F330765CFFCFE42FFAECFE4EFA31FE03" box="[1364,1456,408,432]" name="Germany" pageId="41" pageNumber="534">Germany</collectingCountry>
(
<figureCitation id="131C2A49FFCFE42FFC8DFE64FC31FE79" box="[821,944,434,458]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann &amp; Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet &amp; Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="41" pageNumber="534">Fig. 22G, H</figureCitation>
).
</paragraph>
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. — Name based on the 
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species 
<taxonomicName id="4C274D4FFFCFE42FFA90FE34FC36FDA7" authority="Beyrich, 1854" authorityName="Beyrich" authorityYear="1854" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="pereger">
<emphasis id="B953EADEFFCFE42FFA90FE34FA30FE4A" box="[1320,1457,482,505]" italics="true" pageId="41" pageNumber="534">Murex pereger</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFC95FE2AFC36FDA7" author="BEYRICH E." box="[813,951,508,532]" pageId="41" pageNumber="534" pagination="408 - 500" refId="ref42899" refString="BEYRICH E. 1854. - Die Conchylien des norddeutschen Tertiargebirges. Zeitschrift der Deutschen Geologischen Gesellschaft 6: 408 - 500, 726 - 781. https: // www. digitale-sammlungen. de / de / view / bsb 10283013" type="journal article" year="1854">Beyrich, 1854</bibRefCitation>
</taxonomicName>
and using Bey (from Beyrich), the letters reg (from 
<taxonomicName id="4C274D4FFFCFE42FFC95FDC1FCECFD9D" authorityName="Beyrich" authorityYear="1854" box="[813,877,535,558]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="pereger">
<emphasis id="B953EADEFFCFE42FFC95FDC1FCECFD9D" box="[813,877,535,558]" italics="true" pageId="41" pageNumber="534">pereger</emphasis>
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) and rex (from 
<taxonomicName id="4C274D4FFFCFE42FFBB0FDC1FBC4FD9D" authorityName="Linnaeus" authorityYear="1758" box="[1032,1093,535,558]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFBB0FDC1FBC4FD9D" box="[1032,1093,535,558]" italics="true" pageId="41" pageNumber="534">Murex</emphasis>
</taxonomicName>
).
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. — 
<taxonomicName id="4C274D4FFFCFE42FFBA9FD91FAC1FDED" authority="Carez, 1879" authorityName="Carez" authorityYear="1879" box="[1041,1344,582,606]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="sarroniensis">
<emphasis id="B953EADEFFCFE42FFBA9FD91FB40FDED" box="[1041,1217,583,606]" italics="true" pageId="41" pageNumber="534">Murex sarroniensis</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFB70FD90FAC1FDED" author="CAREZ L." box="[1224,1344,582,606]" pageId="41" pageNumber="534" pagination="637 - 641" refId="ref43315" refString="CAREZ L. 1879. - Description d'especes nouvelles du bassin de Paris. Bulletin de la Societe geologique de France serie 3, 7: 637 - 641." type="journal article" year="1879">Carez, 1879</bibRefCitation>
</taxonomicName>
[synonym: 
<taxonomicName id="4C274D4FFFCFE42FFC96FDB7FBCDFDCA" authority="Wrigley, 1930" authorityName="Wrigley" authorityYear="1930" box="[814,1100,609,633]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="abbatiae">
<emphasis id="B953EADEFFCFE42FFC96FDB7FC41FDCA" box="[814,960,609,633]" italics="true" pageId="41" pageNumber="534">Murex abbatiae</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFC7AFDB7FBCDFDCA" author="WRIGLEY A." box="[962,1100,609,633]" pageId="41" pageNumber="534" pagination="91 - 115" refId="ref49157" refString="WRIGLEY A. 1930. - Notes on English Eocene and Oligocene Mollusca with descriptions of new species. IV - The Muricidae. Proceedings of the Malacological Society of London 19: 91 - 115." type="journal article" year="1930">Wrigley, 1930</bibRefCitation>
</taxonomicName>
], London and Paris Basins, Paleocene (Thanetian) to Early Eocene (Ypresian) (
<figureCitation id="131C2A49FFCFE42FFB0EFDADFAACFD20" box="[1206,1325,635,659]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann &amp; Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet &amp; Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="41" pageNumber="534">Fig. 22C, D</figureCitation>
); 
<taxonomicName id="4C274D4FFFCFE42FFA87FDAAFC52FD1E" authority="Edwards" authorityName="Edwards" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="hantoniensis">
<emphasis id="B953EADEFFCFE42FFA87FDAAFCFAFD1E" italics="true" pageId="41" pageNumber="534">Murex hantoniensis</emphasis>
Edwards
</taxonomicName>
<emphasis id="B953EADEFFCFE42FFC6EFD40FC6BFD1E" box="[982,1002,662,685]" italics="true" pageId="41" pageNumber="534">in</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFC57FD40FB1FFD1E" author="LOWRY J. W. &amp; ETHERIDGE R. &amp; EDWARDS F. E." box="[1007,1182,661,686]" pageId="41" pageNumber="534" refId="ref46148" refString="LOWRY J. W., ETHERIDGE R. &amp; EDWARDS F. E. 1866. - Chart of the Characteristic British Tertiary fossils, stratigraphically arranged. Tennant &amp; Stanford, London, 4 pls. https: // www. biodiversitylibrary. org / page / 51308319" type="journal volume" year="1866">
Lowry 
<emphasis id="B953EADEFFCFE42FFB89FD40FBDFFD1E" box="[1073,1118,661,685]" italics="true" pageId="41" pageNumber="534">et al.</emphasis>
, 1866
</bibRefCitation>
, 
<collectingRegion id="49E3F82EFFCFE42FFB1FFD43FA95FD1E" box="[1191,1300,661,685]" country="United Kingdom" name="Hampshire" pageId="41" pageNumber="534">Hampshire</collectingRegion>
Basin, 
<collectingCountry id="F330765CFFCFE42FFAE0FD43FA2DFD1E" box="[1368,1452,661,685]" name="United Kingdom" pageId="41" pageNumber="534">England</collectingCountry>
, late Eocene (Priabonian) (
<figureCitation id="131C2A49FFCFE42FFB88FD66FB22FD7B" box="[1072,1187,688,712]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann &amp; Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet &amp; Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="41" pageNumber="534">Fig. 22E, F</figureCitation>
); 
<taxonomicName id="4C274D4FFFCFE42FFB01FD67FA2EFD7B" authority="Beyrich, 1854" authorityName="Beyrich" authorityYear="1854" box="[1209,1455,688,712]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="pereger">
<emphasis id="B953EADEFFCFE42FFB01FD67FA9CFD7B" box="[1209,1309,689,712]" italics="true" pageId="41" pageNumber="534">M. pereger</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFA9CFD66FA2EFD7B" author="BEYRICH E." box="[1316,1455,688,712]" pageId="41" pageNumber="534" pagination="408 - 500" refId="ref42899" refString="BEYRICH E. 1854. - Die Conchylien des norddeutschen Tertiargebirges. Zeitschrift der Deutschen Geologischen Gesellschaft 6: 408 - 500, 726 - 781. https: // www. digitale-sammlungen. de / de / view / bsb 10283013" type="journal article" year="1854">Beyrich, 1854</bibRefCitation>
</taxonomicName>
[synonym: 
<taxonomicName id="4C274D4FFFCFE42FFC20FD1DFB57FD51" authority="Sandberger, 1861" authorityName="Sandberger" authorityYear="1861" box="[920,1238,714,738]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="areolifer">
<emphasis id="B953EADEFFCFE42FFC20FD1DFBA4FD51" box="[920,1061,714,738]" italics="true" pageId="41" pageNumber="534">Murex areolifer</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFB92FD1CFB57FD51" author="SANDBERGER C. L. F." box="[1066,1238,714,738]" pageId="41" pageNumber="534" pagination="153 - 232" refId="ref48585" refString="SANDBERGER C. L. F. 1861. - Die conchylien des Mainzer Tertiar-beckens. Heft 5 - 6, C. W. Kreidel, Wiesbaden, 153 - 232." type="book chapter" year="1861">Sandberger, 1861</bibRefCitation>
</taxonomicName>
], Paris Basin, 
<collectingCountry id="F330765CFFCFE42FFAEFFD1CFA2AFD51" box="[1367,1451,714,738]" name="Belgium" pageId="41" pageNumber="534">Belgium</collectingCountry>
, Mainz Basin and north 
<collectingCountry id="F330765CFFCFE42FFBACFD33FBF1FD4E" box="[1044,1136,741,765]" name="Germany" pageId="41" pageNumber="534">Germany</collectingCountry>
, late Eocene (Priabonian) to late Oligocene (Chattian) (
<figureCitation id="131C2A49FFCFE42FFBBDFD29FBFBFCAB" box="[1029,1146,767,792]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann &amp; Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet &amp; Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="41" pageNumber="534">Fig. 22G, H</figureCitation>
); 
<taxonomicName id="4C274D4FFFCFE42FFB31FD29FCE3FC81" authority="Janssen, 1978" authorityName="Janssen" authorityYear="1978" class="Gastropoda" family="Muricidae" genus="Hexaplex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="brevaculeatus">
<emphasis id="B953EADEFFCFE42FFB31FD29FADCFCA4" box="[1161,1373,767,791]" italics="true" pageId="41" pageNumber="534">Hexaplex brevaculeatus</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFAD9FCD6FCE3FC81" author="JANSSEN R." pageId="41" pageNumber="534" pagination="3 - 195" refId="ref45311" refString="JANSSEN R. 1978. - Die Scaphopoden und Gastropoden des Kasseler Meeressandes von Glimmerode (Niederhessen). Geologisches Jahrbuch (A) 41: 3 - 195." type="journal article" year="1978">Janssen, 1978</bibRefCitation>
</taxonomicName>
, Paris basin, early Oligocene (Rupelian), Mainz basins, early Oliogocene and North 
<collectingCountry id="F330765CFFCFE42FFBB6FCE2FBEBFCFF" box="[1038,1130,820,844]" name="Germany" pageId="41" pageNumber="534">Germany</collectingCountry>
, late Oligocene; 
<taxonomicName id="4C274D4FFFCFE42FFAB1FCE3FB07FCD4" authority="Cossmann &amp; Peyrot, 1924" authorityName="Cossmann &amp; Peyrot" authorityYear="1924" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="cedillatus" subGenus="Poirieria">
<emphasis id="B953EADEFFCFE42FFAB1FCE3FAC7FCFF" box="[1289,1350,821,844]" italics="true" pageId="41" pageNumber="534">Murex</emphasis>
(
<emphasis id="B953EADEFFCFE42FFAEDFCE3FA26FCFF" box="[1365,1447,821,844]" italics="true" pageId="41" pageNumber="534">Poirieria</emphasis>
) 
<emphasis id="B953EADEFFCFE42FFC95FC99FC03FCD4" box="[813,898,847,871]" italics="true" pageId="41" pageNumber="534">cedillatus</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFC3FFC99FB07FCD4" author="COSSMANN M. &amp; PEYROT A." box="[903,1158,847,871]" pageId="41" pageNumber="534" pagination="71 - 144" refId="ref43770" refString="COSSMANN M. &amp; PEYROT A. 1924. - Conchologie neogenique de l'Aquitaine (Species n ° 707 - 762). Actes de la Societe linneenne de Bordeaux 75 (2): 71 - 144." type="journal article" year="1924">Cossmann &amp; Peyrot, 1924</bibRefCitation>
</taxonomicName>
, 
<collectingRegion id="49E3F82EFFCFE42FFB37FC99FB6CFCD4" box="[1167,1261,847,871]" country="France" name="Aquitaine" pageId="41" pageNumber="534">Aquitaine</collectingRegion>
Basin, 
<collectingCountry id="F330765CFFCFE42FFA8AFC99FAF4FCD5" box="[1330,1397,847,870]" name="France" pageId="41" pageNumber="534">France</collectingCountry>
, Early Miocene (Burdigalian).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFCFE42FFC95FC4FFA2FFB86" pageId="41" pageNumber="534" type="diagnosis">
<paragraph id="8B9836CCFFCFE42FFC95FC4FFA2FFB86" blockId="41.[811,1456,921,1077]" pageId="41" pageNumber="534">
<smallCapsWord id="8D7EA010FFCFE42FFC95FC4FFC16FC1C" baselines="938,939" box="[813,919,921,944]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="41" pageNumber="534">DIAGNOSIS</smallCapsWord>
. — Biconic teleoconch with subangular whorls. D1 well developed. P6 not atrophied. IP weak, ADP, MP present. Varices formed by 1-3 lamellae abaxially turned on the early whorls, occasionally reinforced by several lamellae on the succeeding whorls. P1 spine short. Shoulder sinus widely open, short, moderately deep. Posterior sinus on sutural ramp shallow.Columellar lip usually smooth.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFCFE42FFC95FB82FAA2F85F" pageId="41" pageNumber="534" type="discussion">
<paragraph id="8B9836CCFFCFE42FFC95FB82FC2DFBDE" blockId="41.[811,1457,1108,2028]" box="[813,940,1108,1134]" pageId="41" pageNumber="534">
<smallCapsWord id="8D7EA010FFCFE42FFC95FB82FC2DFBDE" baselines="1127,1128" box="[813,940,1108,1134]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="41" pageNumber="534">COMMENTS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFCFE42FFC95FBA2FAA2F85F" blockId="41.[811,1457,1108,2028]" pageId="41" pageNumber="534">
Members of 
<taxonomicName id="4C274D4FFFCFE42FFC07FBA3FBECFB3D" authority="Merle" authorityName="Merle" authorityYear="2024" box="[959,1133,1140,1166]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFC07FBA3FBA3FB3D" box="[959,1058,1141,1166]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFBC3FBA3FB48FB3C" box="[1147,1225,1141,1167]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
differ from those of 
<taxonomicName id="4C274D4FFFCFE42FFC96FB42FC14FB1D" authorityName="Merle" authorityYear="2005" box="[814,917,1172,1198]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFCFE42FFC96FB42FC14FB1D" box="[814,917,1172,1198]" italics="true" pageId="41" pageNumber="534">Jsowerbya</emphasis>
</taxonomicName>
in having a deeper shoulder sinus, more lamellose varices, and a smooth columellar lip. 
<taxonomicName id="4C274D4FFFCFE42FFB5AFB62FCF0FB5D" authority="Merle" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFB5AFB62FA30FB7D" box="[1250,1457,1204,1230]" italics="true" pageId="41" pageNumber="534">Pseudotrophonopsis</emphasis>
Merle
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFCC4FB02FC48FB5D" box="[892,969,1236,1262]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
is also distinguishing from 
<taxonomicName id="4C274D4FFFCFE42FFB44FB02FA2AFB5D" authority="Merle" authorityName="Merle" authorityYear="2024" box="[1276,1451,1236,1262]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFB44FB02FADEFB5E" box="[1276,1375,1236,1261]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFC95FB22FCFBFABD" box="[813,890,1268,1294]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
in having rounded whorls, strong secondary cords of equal strength to the primary cords, and numerous split internal denticles within the outer lip (D1 to D5). Represented by five species, 
<taxonomicName id="4C274D4FFFCFE42FFBC4FA82FAB5FADE" authority="Merle" authorityName="Merle" authorityYear="2024" box="[1148,1332,1363,1389]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFBC4FA82FB62FADE" box="[1148,1251,1364,1389]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFAFAFA82FA15FADD" box="[1346,1428,1364,1390]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
is the most diverse genus of the group. It extends from the Anglo-Parisian late Paleocene/early Eocene with 
<taxonomicName id="4C274D4FFFCFE42FFADCFA42FBB8FA7E" authority="(Carez, 1879)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Carez" baseAuthorityYear="1879" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="sarroniensis" status="n. comb.">
<emphasis id="B953EADEFFCFE42FFADCFA42FC0DFA7E" italics="true" pageId="41" pageNumber="534">B. sarroniensis</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFC25FA65FBAEFA7E" author="CAREZ L." box="[925,1071,1459,1485]" pageId="41" pageNumber="534" pagination="637 - 641" refId="ref43315" refString="CAREZ L. 1879. - Description d'especes nouvelles du bassin de Paris. Bulletin de la Societe geologique de France serie 3, 7: 637 - 641." type="journal article" year="1879">Carez, 1879</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFBFFFA65FB2EFA7E" box="[1095,1199,1459,1485]" pageId="41" pageNumber="534" rank="species">n. comb.</taxonomicNameLabel>
to the Early Miocene (Burdigalian) from the Aquitaine Basin with 
<taxonomicName id="4C274D4FFFCFE42FFA95FA02FBE3F9BE" authority="(Cossmann &amp; Peyrot, 1924)" baseAuthorityName="Cossmann &amp; Peyrot" baseAuthorityYear="1924" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="cedillatus">
<emphasis id="B953EADEFFCFE42FFA95FA02FA31FA5E" box="[1325,1456,1491,1517]" italics="true" pageId="41" pageNumber="534">B. cedillatus</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFC8CFA25FBDBF9BE" author="COSSMANN M. &amp; PEYROT A." box="[820,1114,1523,1549]" pageId="41" pageNumber="534" pagination="71 - 144" refId="ref43770" refString="COSSMANN M. &amp; PEYROT A. 1924. - Conchologie neogenique de l'Aquitaine (Species n ° 707 - 762). Actes de la Societe linneenne de Bordeaux 75 (2): 71 - 144." type="journal article" year="1924">Cossmann &amp; Peyrot, 1924</bibRefCitation>
)
</taxonomicName>
. 
<taxonomicName id="4C274D4FFFCFE42FFBD2FA25FAC8F9BF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Carez" baseAuthorityYear="1879" box="[1130,1353,1523,1548]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="sarroniensis" status="comb. nov.">
<emphasis id="B953EADEFFCFE42FFBD2FA25FAC8F9BF" box="[1130,1353,1523,1548]" italics="true" pageId="41" pageNumber="534">Beyregrex sarroniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFCFE42FFAE8FA25FA31F9BE" box="[1360,1456,1523,1549]" pageId="41" pageNumber="534" rank="species">n. comb.</taxonomicNameLabel>
displays several primitive characters, such as missing IP, MP and ABP. During the middle Eocene, no 
<taxonomicName id="4C274D4FFFCFE42FFB44F9E5FA2AF9FF" authority="Merle" authorityName="Merle" authorityYear="2024" box="[1276,1451,1586,1612]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B953EADEFFCFE42FFB44F9E5FADEF9FF" box="[1276,1375,1587,1612]" italics="true" pageId="41" pageNumber="534">Beyregrex</emphasis>
Merle
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFC95F985FCFBF9DE" box="[813,890,1619,1645]" pageId="41" pageNumber="534" rank="genus">n. gen.</taxonomicNameLabel>
species are known, but the genus re-emerges in the late Eocene of Hampshire with 
<taxonomicName id="4C274D4FFFCFE42FFB22F9A5FAC1F93F" box="[1178,1344,1650,1676]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="hantoniensis">
<emphasis id="B953EADEFFCFE42FFB22F9A5FAC1F93F" box="[1178,1344,1650,1676]" italics="true" pageId="41" pageNumber="534">B. hantoniensis</emphasis>
</taxonomicName>
(Edwards 
<emphasis id="B953EADEFFCFE42FFC97F945FCC7F91F" box="[815,838,1683,1708]" italics="true" pageId="41" pageNumber="534">in</emphasis>
<bibRefCitation id="EFB64B3DFFCFE42FFCF5F945FBA7F91F" author="LOWRY J. W. &amp; ETHERIDGE R. &amp; EDWARDS F. E." box="[845,1062,1682,1709]" pageId="41" pageNumber="534" refId="ref46148" refString="LOWRY J. W., ETHERIDGE R. &amp; EDWARDS F. E. 1866. - Chart of the Characteristic British Tertiary fossils, stratigraphically arranged. Tennant &amp; Stanford, London, 4 pls. https: // www. biodiversitylibrary. org / page / 51308319" type="journal volume" year="1866">
Lowry 
<emphasis id="B953EADEFFCFE42FFC19F945FC59F91F" box="[929,984,1682,1708]" italics="true" pageId="41" pageNumber="534">et al.</emphasis>
, 1866
</bibRefCitation>
), n. comb.. That species is closely related to 
<taxonomicName id="4C274D4FFFCFE42FFC1BF965FB49F97E" authority="(Beyrich, 1854)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Beyrich" baseAuthorityYear="1854" box="[931,1224,1714,1741]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="pereger" status="n. comb.">
<emphasis id="B953EADEFFCFE42FFC1BF965FB8CF97F" box="[931,1037,1715,1740]" italics="true" pageId="41" pageNumber="534">B. pereger</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFBA4F964FB41F97E" author="BEYRICH E." box="[1052,1216,1714,1741]" pageId="41" pageNumber="534" pagination="408 - 500" refId="ref42899" refString="BEYRICH E. 1854. - Die Conchylien des norddeutschen Tertiargebirges. Zeitschrift der Deutschen Geologischen Gesellschaft 6: 408 - 500, 726 - 781. https: // www. digitale-sammlungen. de / de / view / bsb 10283013" type="journal article" year="1854">Beyrich, 1854</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="A26057A5FFCFE42FFB6FF965FABDF97F" box="[1239,1340,1714,1741]" pageId="41" pageNumber="534" rank="species">n. comb.</taxonomicNameLabel>
occurring in the Paris Basin and in 
<collectingCountry id="F330765CFFCFE42FFB85F904FB24F95F" box="[1085,1189,1746,1772]" name="Germany" pageId="41" pageNumber="534">Germany</collectingCountry>
(Mainz Basin and north 
<collectingCountry id="F330765CFFCFE42FFC93F924FC19F8BF" box="[811,920,1778,1804]" name="Germany" pageId="41" pageNumber="534">Germany</collectingCountry>
) during the Oligocene. According to 
<bibRefCitation id="EFB64B3DFFCFE42FFAE3F924FCF9F89F" author="JANSSEN R." pageId="41" pageNumber="534" pagination="3 - 195" refId="ref45311" refString="JANSSEN R. 1978. - Die Scaphopoden und Gastropoden des Kasseler Meeressandes von Glimmerode (Niederhessen). Geologisches Jahrbuch (A) 41: 3 - 195." type="journal article" year="1978">Janssen (1978)</bibRefCitation>
, 
<taxonomicName id="4C274D4FFFCFE42FFC3EF8C4FB65F89F" authority="(Janssen, 1978)" baseAuthorityName="Janssen" baseAuthorityYear="1978" box="[902,1252,1810,1836]" class="Gastropoda" family="Muricidae" genus="Hexaplex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="brevaculeatus">
<emphasis id="B953EADEFFCFE42FFC3EF8C4FBB3F89F" box="[902,1074,1810,1836]" italics="true" pageId="41" pageNumber="534">B. brevaculeatus</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFBF8F8C4FB5AF89F" author="JANSSEN R." box="[1088,1243,1810,1836]" pageId="41" pageNumber="534" pagination="3 - 195" refId="ref45311" refString="JANSSEN R. 1978. - Die Scaphopoden und Gastropoden des Kasseler Meeressandes von Glimmerode (Niederhessen). Geologisches Jahrbuch (A) 41: 3 - 195." type="journal article" year="1978">Janssen, 1978</bibRefCitation>
)
</taxonomicName>
is another species. It was originally described from the Chattian of 
<collectingCountry id="F330765CFFCFE42FFAFEF8E4FA2FF8FF" box="[1350,1454,1842,1868]" name="Germany" pageId="41" pageNumber="534">Germany</collectingCountry>
(at Glimmerode) but is also recorded in the Rupelian of the Paris Basin (
<bibRefCitation id="EFB64B3DFFCFE42FFC67F8A7FBE1F838" author="MERLE D." box="[991,1120,1905,1931]" pageId="41" pageNumber="534" refId="ref46629" refString="MERLE D. 1999. - La radiation des Muricidae (Gastropoda: Neogastropoda) au Paleogene: approche phylogenetique et evolutive. Unpublished PhD thesis, Museum national d'Histoire naturelle, Paris, 499 p." type="book" year="1999">
<taxonomicName id="4C274D4FFFCFE42FFC67F8A7FBA1F838" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[991,1056,1905,1931]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
1999
</bibRefCitation>
; 
<bibRefCitation id="EFB64B3DFFCFE42FFBD7F8A4FA8BF838" author="LOZOUET P." box="[1135,1290,1905,1932]" pageId="41" pageNumber="534" refId="ref46076" refString="LOZOUET P. (coord.) 2012. - Stratotype Stampien. Museum national d'Histoire naturelle, Paris; Biotope, Meze, 464 p. (coll. Patrimoine geologique; 4)." type="book" year="2012">Lozouet 2012</bibRefCitation>
). For the Neogene, 
<taxonomicName id="4C274D4FFFCFE42FFCC8F844FAC2F818" authority="(Cossmann &amp; Peyrot, 1924)" baseAuthorityName="Cossmann &amp; Peyrot" baseAuthorityYear="1924" box="[880,1347,1937,1964]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="41" pageNumber="534" phylum="Mollusca" rank="species" species="cedillatus">
<emphasis id="B953EADEFFCFE42FFCC8F844FC77F818" box="[880,1014,1937,1963]" italics="true" pageId="41" pageNumber="534">B. cedillatus</emphasis>
(
<bibRefCitation id="EFB64B3DFFCFE42FFBBFF847FABAF818" author="COSSMANN M. &amp; PEYROT A." box="[1031,1339,1937,1964]" pageId="41" pageNumber="534" pagination="71 - 144" refId="ref43770" refString="COSSMANN M. &amp; PEYROT A. 1924. - Conchologie neogenique de l'Aquitaine (Species n ° 707 - 762). Actes de la Societe linneenne de Bordeaux 75 (2): 71 - 144." type="journal article" year="1924">Cossmann &amp; Peyrot, 1924</bibRefCitation>
)
</taxonomicName>
from the Burdigalian of the Aquitaine Basin demonstrates that the genus was not restricted to northern Europe.
</paragraph>
</subSubSection>
</treatment>
</document>