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<document id="A845833ED80CDE161A4305D4618E6015" ID-DOI="10.5281/zenodo.176282" ID-GBIF-Dataset="fe89bad2-db11-4058-a77b-59dbc6714d89" ID-ISSN="1175-5326" ID-Zenodo-Dep="176282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459830473595" checkinUser="plazi" docAuthor="Mwabvu, Tarombera, Hamer, Michelle L. &amp; Slotow, Robert H." docDate="2007" docId="03B587B06D01FFFEFF54F8D660E6FEE7" docLanguage="en" docName="zt01452p023.pdf" docOrigin="Zootaxa 1452" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brevitibius nyathi Mwabvu, Hamer &amp; Slotow, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="16" masterDocId="FF8CFFC86D0CFFF1FFC3FFEE6305FFB5" masterDocTitle="A taxonomic review of the southern African millipede genus, Bicoxidens Attems, 1928 (Diplopoda: Spirostreptida: Spirostreptidae), with the description of three new species and a tentative phylogeny" masterLastPageNumber="23" masterPageNumber="1" pageNumber="14" updateTime="1698222893347" updateUser="plazi">
<mods:mods id="14FA6E64A847D52808DF78B43F38D4F6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0CA4CD16B68711D5963F1C0B8AA8665">
<mods:title id="10369C30181084C72333C221C3AC4F8D">A taxonomic review of the southern African millipede genus, Bicoxidens Attems, 1928 (Diplopoda: Spirostreptida: Spirostreptidae), with the description of three new species and a tentative phylogeny</mods:title>
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<mods:name id="B611200D80EC2F6AE918E06B647FE189" type="personal">
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<mods:namePart id="B2E4125239340C6E1FAC7B27319D85E2">Mwabvu, Tarombera</mods:namePart>
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<mods:namePart id="40C0C3636D65D813AC236F4082D40B12">Hamer, Michelle L.</mods:namePart>
</mods:name>
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<mods:role id="28D49B3C32DCE4000C8F0C9784434230">
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<mods:namePart id="F29F1D2CD0F185D63D9AD39AAB3CD6EF">Slotow, Robert H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9F678A54676B348A26D0EC65F82C4485">text</mods:typeOfResource>
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<mods:title id="DE629BF11344DDA053B16BE7B8960A9A">Zootaxa</mods:title>
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<mods:part id="79AFE0086020A92DD4471258EA80EFD9">
<mods:date id="2443DA13316502AD4F5751F996538A96">2007</mods:date>
<mods:detail id="5C4638B2D75BE9AD97A4EEF02EBDD2BB" type="volume">
<mods:number id="53CD2F4210535F4889252A535883FEC7">1452</mods:number>
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<mods:start id="6F9118E7ED4F1973D4D40E455D942CE2">1</mods:start>
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<mods:classification id="AE626FCA670789B26944B0712B8C9AFF">journal article</mods:classification>
<mods:identifier id="48F45519311818BB7645B087921FA0DB" type="DOI">10.5281/zenodo.176282</mods:identifier>
<mods:identifier id="3B45D62AC91C826338EF062628D15B49" type="GBIF-Dataset">fe89bad2-db11-4058-a77b-59dbc6714d89</mods:identifier>
<mods:identifier id="CEFED9B85D7BAE45D7EAE60F08B789D2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1C3E26238CA45AF58A6B3131653075E1" type="Zenodo-Dep">176282</mods:identifier>
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<treatment id="03B587B06D01FFFEFF54F8D660E6FEE7" ID-DOI="http://doi.org/10.5281/zenodo.6251147" ID-GBIF-Taxon="119357839" ID-Zenodo-Dep="6251147" LSID="urn:lsid:plazi:treatment:03B587B06D01FFFEFF54F8D660E6FEE7" httpUri="http://treatment.plazi.org/id/03B587B06D01FFFEFF54F8D660E6FEE7" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="C306652D6D01FFFCFF54F8D663E6F8C1" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BA336A66D01FFFCFF54F8D662D1F8E7" blockId="13.[151,468,1848,1908]" box="[151,468,1848,1874]" pageId="13" pageNumber="14">
<heading id="D0EB81CA6D01FFFCFF54F8D662D1F8E7" bold="true" box="[151,468,1848,1874]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B968EAB46D01FFFCFF54F8D662D1F8E7" bold="true" box="[151,468,1848,1874]" pageId="13" pageNumber="14">
<taxonomicName id="4C1C4D256D01FFFCFF54F8D66200F8E7" box="[151,261,1848,1874]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nyathi" status="sp. nov.">
<emphasis id="B968EAB46D01FFFCFF54F8D66200F8E7" bold="true" box="[151,261,1848,1874]" italics="true" pageId="13" pageNumber="14">B. nyathi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57CF6D01FFFCFECEF8D6624AF8E7" box="[269,335,1848,1874]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
Mwabvu
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336A66D01FFFCFF54F8B463E6F8C1" blockId="13.[151,468,1848,1908]" box="[151,227,1882,1908]" pageId="13" pageNumber="14">
<figureCitation id="13272A236D01FFFCFF54F8B463E6F8C1" box="[151,227,1882,1908]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C306652D6D01FFFFFF54F844674FFF6F" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BA336A66D01FFFCFF54F8446230F859" blockId="13.[151,1436,1962,2028]" pageId="13" pageNumber="14">
<typeStatus id="54A788046D01FFFCFF54F84463D4F871" box="[151,209,1962,1988]" pageId="13" pageNumber="14">Type</typeStatus>
material (examined):
<typeStatus id="54A788046D01FFFCFE21F8446151F871" box="[482,596,1962,1988]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F30B76366D01FFFCFDA0F844600DF871" box="[611,776,1962,1988]" name="Zimbabwe" pageId="13" pageNumber="14">ZIMBABWE</collectingCountry>
: 1 ɗ, Mudzwiru River [2030B4],
<date id="FFA210666D01FFFCFB6CF844661FF871" box="[1199,1306,1962,1988]" pageId="13" pageNumber="14" value="1994-02-04">4.ii.1994</date>
, F.
<taxonomicName id="4C1C4D256D01FFFCFA8FF8446699F871" box="[1356,1436,1962,1988]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="22" phylum="Arthropoda" rank="species" species="nyathi">Nyathi</taxonomicName>
(NMZ/D747)
</paragraph>
<paragraph id="8BA336A66D02FFFFFF05FF76674FFF6F" blockId="14.[151,1437,152,338]" pageId="14" pageNumber="15">
Etymology: Named after Mr Francis
<taxonomicName id="4C1C4D256D02FFFFFDB4FF7661C2FF07" box="[631,711,152,178]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="22" phylum="Arthropoda" rank="species" species="nyathi">Nyathi</taxonomicName>
(Natural History Museum, Bulawayo,
<collectingCountry id="F30B76366D02FFFFFB4CFF76660BFF07" box="[1167,1294,152,178]" name="Zimbabwe" pageId="14" pageNumber="15">Zimbabwe</collectingCountry>
) in recognition of his contribution to the collection and study of invertebrates in
<collectingCountry id="F30B76366D02FFFFFC07FF2E6743FF6F" box="[964,1094,192,218]" name="Zimbabwe" pageId="14" pageNumber="15">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C306652D6D02FFFFFF05FF0663C9F9E6" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BA336A66D02FFFFFF05FF0661C0FEE7" blockId="14.[151,1437,152,338]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05FF066238FEB4" box="[198,317,232,257]" italics="true" pageId="14" pageNumber="15">Diagnosis</emphasis>
: Apical telocoxite with a lateral and median process resulting in an oval shape (
<figureCitation id="13272A236D02FFFFFB20FF06662DFEB7" box="[1251,1320,232,258]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
a); subapically, the median leaf has a raised subapical process (
<figureCitation id="13272A236D02FFFFFCDDFEFE6072FE9F" box="[798,887,272,298]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig. 8b</figureCitation>
). A laterally deflected aboral lateral leaf lobe, more tapered than rounded, is present (
<figureCitation id="13272A236D02FFFFFDA3FED661B2FEE7" box="[608,695,312,338]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig. 8b</figureCitation>
).
</paragraph>
<caption id="DF63662E6D02FFFFFF54FAC56061FAD0" httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15" targetBox="[305,1289,386,1305]" targetPageId="14">
<paragraph id="8BA336A66D02FFFFFF54FAC56061FAD0" blockId="14.[151,1436,1323,1455]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF54FAC56219FAF6" bold="true" box="[151,284,1323,1347]" pageId="14" pageNumber="15">FIGURE 8.</emphasis>
<taxonomicName id="4C1C4D256D02FFFFFEE7FAC56282FAF7" box="[292,391,1323,1346]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nyathi" status="sp. nov.">
<emphasis id="B968EAB46D02FFFFFEE7FAC56282FAF7" box="[292,391,1323,1346]" italics="true" pageId="14" pageNumber="15">B. nyathi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57CF6D02FFFFFE4CFAC562E1FAF7" box="[399,484,1323,1346]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
(NMZ/D747).
<emphasis id="B968EAB46D02FFFFFD57FAC561ADFAF6" bold="true" box="[660,680,1323,1347]" pageId="14" pageNumber="15">a,</emphasis>
oral view of gonopods;
<emphasis id="B968EAB46D02FFFFFC7CFAC560D1FAF6" bold="true" box="[959,980,1323,1347]" pageId="14" pageNumber="15">b,</emphasis>
aboral view of gonopods;
<emphasis id="B968EAB46D02FFFFFB3FFAC5660DFAF6" bold="true" box="[1276,1288,1323,1347]" pageId="14" pageNumber="15">c</emphasis>
, oral view of prefemoral lobe of first pair of male legs;
<emphasis id="B968EAB46D02FFFFFD95FAA36169FAD0" bold="true" box="[598,620,1357,1381]" pageId="14" pageNumber="15">d,</emphasis>
lateral view of collum.
</paragraph>
</caption>
<paragraph id="8BA336A66D02FFFFFF54FA9E61EEFA1A" blockId="14.[151,1436,1323,1455]" pageId="14" pageNumber="15">Abbreviations: cf, collum fold; fl, femoral lobe; llb, lateral leaf lobe; lp, lateral process; ml, median leaf; mp, median process; pc, paracoxite; pfl, prefemoral lobe; tp, telopodite.</paragraph>
<paragraph id="8BA336A66D02FFFFFF05FA07625DF9B6" blockId="14.[151,1436,1513,2021]" box="[198,344,1513,1539]" pageId="14" pageNumber="15">Description:</paragraph>
<paragraph id="8BA336A66D02FFFFFF05F9FF63C9F9E6" blockId="14.[151,1436,1513,2021]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F9FF63F0F99F" box="[198,245,1553,1578]" italics="true" pageId="14" pageNumber="15">Size</emphasis>
: Body length
<quantity id="4CE49B436D02FFFFFE67F9FF610CF99E" box="[420,521,1553,1579]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="15" unit="mm" value="115.0">115 mm</quantity>
; minimum and maximum body width 4–5 and
<quantity id="4CE49B436D02FFFFFB80F9FF67AFF99E" box="[1091,1194,1553,1579]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="14" pageNumber="15" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
; antenna length
<quantity id="4CE49B436D02FFFFFAB3F9FF63CCF9E6" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="14" pageNumber="15" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C306652D6D02FFFEFF05F98F60E6FEE7" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="8BA336A66D02FFFFFF05F98F62F6F9CE" blockId="14.[151,1436,1513,2021]" box="[198,499,1633,1659]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F98F62C3F9CF" box="[198,454,1633,1658]" italics="true" pageId="14" pageNumber="15">Number of body rings</emphasis>
: 52
</paragraph>
<paragraph id="8BA336A66D02FFFFFF05F96761ECF916" blockId="14.[151,1436,1513,2021]" box="[198,745,1673,1699]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F967621CF917" box="[198,281,1673,1698]" italics="true" pageId="14" pageNumber="15">Colour</emphasis>
: Body black; legs and antennae brown.
</paragraph>
<paragraph id="8BA336A66D02FFFFFF05F95F60E0F97E" blockId="14.[151,1436,1513,2021]" box="[198,997,1713,1739]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F95F6219F97F" box="[198,284,1713,1738]" italics="true" pageId="14" pageNumber="15">Collum</emphasis>
: With three complete folds and an incomplete fold (
<figureCitation id="13272A236D02FFFFFC43F95F60C3F97E" box="[896,966,1713,1739]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
d).
</paragraph>
<paragraph id="8BA336A66D02FFFFFF05F93263C6F8A8" blockId="14.[151,1436,1513,2021]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F93261C8F940" box="[198,717,1752,1781]" italics="true" pageId="14" pageNumber="15">Pre-femoral process of 1st pair of male legs</emphasis>
: Subtriangular, proximally parallel, with a rounded tip (
<figureCitation id="13272A236D02FFFFFAAEF93563A3F8A8" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
c).
</paragraph>
<paragraph id="8BA336A66D02FFFFFF05F8C260E5F8F0" blockId="14.[151,1436,1513,2021]" box="[198,992,1835,1861]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F8C262B5F8F0" box="[198,432,1836,1861]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C1C4D256D02FFFFFF05F8C26216F8F0" box="[198,275,1836,1861]" class="Insecta" family="Mutillidae" genus="Sigilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Sigilla</taxonomicName>
on body ring
</emphasis>
: Three irregular rows, reduced to one ventrally.
</paragraph>
<paragraph id="8BA336A66D02FFFFFF05F8BA60A8F850" blockId="14.[151,1436,1513,2021]" pageId="14" pageNumber="15">
<emphasis id="B968EAB46D02FFFFFF05F8BA6236F8D8" box="[198,307,1876,1901]" italics="true" pageId="14" pageNumber="15">Gonopod</emphasis>
: Orally, apical telocoxite oval. A median process is present, directed distally, not overlapping the opposite one. Aborally, apex of median leaf slightly raised (
<figureCitation id="13272A236D02FFFFFCBEF89560CBF820" box="[893,974,1915,1941]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig.8b</figureCitation>
). An aboral, lateral leaf lobe extending from apical telocoxite (below the level of the apex of the median leaf) deflected laterally, more tapered than rounded. The telopodite forms an incomplete loop distally (
<figureCitation id="13272A236D02FFFFFC93F8256090F850" box="[848,917,1995,2021]" captionStart="FIGURE 8. B" captionStartId="14.[151,255,1323,1347]" captionTargetBox="[305,1289,386,1305]" captionTargetId="figure@14.[294,1303,376,1316]" captionTargetPageId="14" captionText="FIGURE 8. B. nyathi sp. nov. (NMZ / D 747). a, oral view of gonopods; b, aboral view of gonopods; c, oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum." httpUri="https://zenodo.org/record/176290/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
a)
</paragraph>
<paragraph id="8BA336A66D03FFFEFF05FF766245FF6F" blockId="15.[151,1437,152,338]" pageId="15" pageNumber="16">
Distribution: Known from Mudzwiru River, south of Gweru,
<collectingCountry id="F30B76366D03FFFEFC5BFF766710FF07" box="[920,1045,152,178]" name="Zimbabwe" pageId="15" pageNumber="16">Zimbabwe</collectingCountry>
(
<figureCitation id="13272A236D03FFFEFBE6FF766778FF07" box="[1061,1149,152,178]" captionStart="FIGURE 10" captionStartId="20.[151,255,1486,1510]" captionText="FIGURE 10: Distribution of Bicoxidens species" pageId="15" pageNumber="16">Fig. 10</figureCitation>
). Specimen found under a granite rock.
</paragraph>
<paragraph id="8BA336A66D03FFFEFF05FF0660E6FEE7" blockId="15.[151,1437,152,338]" pageId="15" pageNumber="16">
Remarks: The apical telocoxite is oval and simple, unlike other species. However, it resembles
<emphasis id="B968EAB46D03FFFEFADDFF06612DFE9C" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C1C4D256D03FFFEFADDFF0663BDFE9C" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="matopoensis">matopoensis</taxonomicName>
,
<taxonomicName id="4C1C4D256D03FFFEFF07FEFE624DFE9C" box="[196,328,272,297]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nigerrimus">nigerrimus</taxonomicName>
,
<taxonomicName id="4C1C4D256D03FFFEFE90FEFE62A1FE9C" box="[339,420,272,297]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="friendi">friendi</taxonomicName>
,
<taxonomicName id="4C1C4D256D03FFFEFE73FEFE612DFE9C" box="[432,552,272,297]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gokwensis">gokwensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C1C4D256D03FFFEFDA3FEFE61B6FE9C" box="[608,691,272,297]" class="Diplopoda" family="Spirostreptidae" genus="Bicoxidens" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="brincki">
<emphasis id="B968EAB46D03FFFEFDA3FEFE61B6FE9C" box="[608,691,272,297]" italics="true" pageId="15" pageNumber="16">brincki</emphasis>
</taxonomicName>
by having an aboral lobe on the lateral leaf. However, in
<taxonomicName id="4C1C4D256D03FFFEFA90FEFE6698FE9C" box="[1363,1437,272,297]" class="Diplopoda" family="Spirostreptidae" genus="Brevitibius" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nyathi">
<emphasis id="B968EAB46D03FFFEFA90FEFE6698FE9C" box="[1363,1437,272,297]" italics="true" pageId="15" pageNumber="16">nyathi</emphasis>
</taxonomicName>
the lobe is smaller, deflected laterally and is more tapered than rounded.
</paragraph>
</subSubSection>
</treatment>
</document>
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