A taxonomic review of the southern African millipede genus, Bicoxidens Attems, 1928 (Diplopoda: Spirostreptida: Spirostreptidae), with the description of three new species and a tentative phylogeny Author Mwabvu, Tarombera Author Hamer, Michelle L. Author Slotow, Robert H. text Zootaxa 2007 1452 1 23 journal article 10.5281/zenodo.176282 fe89bad2-db11-4058-a77b-59dbc6714d89 1175-5326 176282 B. nyathi sp. n. Mwabvu Fig. 8 Type material (examined): Holotype : ZIMBABWE : 1 ɗ, Mudzwiru River [2030B4], 4.ii.1994 , F. Nyathi (NMZ/D747) Etymology: Named after Mr Francis Nyathi (Natural History Museum, Bulawayo, Zimbabwe ) in recognition of his contribution to the collection and study of invertebrates in Zimbabwe . Diagnosis : Apical telocoxite with a lateral and median process resulting in an oval shape ( Fig. 8 a); subapically, the median leaf has a raised subapical process ( Fig. 8b ). A laterally deflected aboral lateral leaf lobe, more tapered than rounded, is present ( Fig. 8b ). FIGURE 8. B. nyathi sp. nov. (NMZ/D747). a, oral view of gonopods; b, aboral view of gonopods; c , oral view of prefemoral lobe of first pair of male legs; d, lateral view of collum. Abbreviations: cf, collum fold; fl, femoral lobe; llb, lateral leaf lobe; lp, lateral process; ml, median leaf; mp, median process; pc, paracoxite; pfl, prefemoral lobe; tp, telopodite. Description: Size : Body length 115 mm ; minimum and maximum body width 4–5 and 6–7 mm ; antenna length 5–6 mm . Number of body rings : 52 Colour : Body black; legs and antennae brown. Collum : With three complete folds and an incomplete fold ( Fig. 8 d). Pre-femoral process of 1st pair of male legs : Subtriangular, proximally parallel, with a rounded tip ( Fig. 8 c). Sigilla on body ring : Three irregular rows, reduced to one ventrally. Gonopod : Orally, apical telocoxite oval. A median process is present, directed distally, not overlapping the opposite one. Aborally, apex of median leaf slightly raised ( Fig.8b ). An aboral, lateral leaf lobe extending from apical telocoxite (below the level of the apex of the median leaf) deflected laterally, more tapered than rounded. The telopodite forms an incomplete loop distally ( Fig. 8 a) Distribution: Known from Mudzwiru River, south of Gweru, Zimbabwe ( Fig. 10 ). Specimen found under a granite rock. Remarks: The apical telocoxite is oval and simple, unlike other species. However, it resembles matopoensis , nigerrimus , friendi , gokwensis and brincki by having an aboral lobe on the lateral leaf. However, in nyathi the lobe is smaller, deflected laterally and is more tapered than rounded.