treatments-xml/data/D3/72/33/D3723344842638A6D42382F7CD46EBAD.xml

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<mods:title id="D76135575137B8BF3094ACFBF0067C9D">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart id="C6F920E8D6607FCD160D8405C29F3A63">Salata, Sebastian</mods:namePart>
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<mods:namePart id="64463D5471542FF442FC1A61FB290FE5">Fisher, Brian L.</mods:namePart>
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<treatment id="D3723344842638A6D42382F7CD46EBAD" ID-GBIF-Taxon="161777035" LSID="urn:lsid:zoobank.org:act:88804AF2-4F5F-4A67-B2BD-9012E4F43AE9" httpUri="http://treatment.plazi.org/id/D3723344842638A6D42382F7CD46EBAD" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="E41A6D3D0B86E5B22F5B407E97DE13B1" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="8011F53769B6494175ED5691B9FD137E" pageId="0" pageNumber="1">
<taxonomicName id="3FB735F9E41575EDF41F55BDF2837DA5" ID-CoL="CMLF3" LSID="http://zoobank.org/88804AF2-4F5F-4A67-B2BD-9012E4F43AE9" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longipilosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longipilosa" status="sp. nov.">Pheidole longipilosa</taxonomicName>
<taxonomicNameLabel id="8794B79F6CAAED517314568111EB563D" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="99B90BD15A19A65242B23A64AD8A0EEE" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">Figs 39A-F</figureCitation>
<figureCitation id="7AFBEEFD3333C8255312770692901907" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84X</figureCitation>
<figureCitation id="A986DA22B56B026147EE87BB86690E87" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E517DC104EE07A05EB9283F2A24B4B27" pageId="0" pageNumber="1" type="type material">
<paragraph id="56E05ED43A71F5EF802063532A91E0FC" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="BCED85C5ECB7E301751C72A7F02F9145" pageId="0" pageNumber="1">
<emphasis id="05998A580DCEBB24E4DE585714398E30" bold="true" pageId="0" pageNumber="1">
<emphasis id="07E35931160CC7B441EB568E43C5CBB1" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Toliara; 
<normalizedToken id="124BAEF859FA933E0664250ECABB410A" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="F2D31A9AF8C88A8B06559EED0E30D5C0" originalValue="Classée">Classee</normalizedToken>
<normalizedToken id="B86F541B837EF53BCEE8837B4998DF87" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
, 29.2 km 343°NNW Mahaboboka; -22.675, 44.19; alt. 1100 m; 18 Feb 2003; B.L. Fisher et al. leg.; CASENT0498298 (CASC). 
<emphasis id="B570484D9CB669F2DBF1B5AC19C5BBF6" bold="true" pageId="0" pageNumber="1">
<emphasis id="48557DE95FF79BCBD6F9441F6C161E2D" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Madagascar. •9w.; same data as for holotype; CASENT0235043, CASENT0498295-CASENT0498297, CASENT0872154-CASENT0872158 (CASC).
</paragraph>
</subSubSection>
<subSubSection id="038F3C1AC7A9E400F3E873F22374824B" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="F9F9868ADBAB824FA5E8472CE8D30A81" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="24F89F3AA10013736A5E6A3899A68FD6" pageId="0" pageNumber="1">
Madagascar. - 
<emphasis id="F2EB4D97C3CC298398E6FF7E3A61406E" bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
: •1w.; Parc National 
<normalizedToken id="3828BA7865AA652E18EED8F35A2B4828" originalValue="d’Isalo">d'Isalo</normalizedToken>
, Sahanafa River, 29.2 km 351°N Ranohira; 
<geoCoordinate id="D3475C88E4B4043A5113214D743E3FDA" degrees="22.31333" direction="south" orientation="latitude" precision="1" value="-22.31333">-22.31333</geoCoordinate>
, 
<geoCoordinate id="2CE86AD7BEFC0D8335033F8FC64870A1" degrees="45.29167" direction="east" orientation="longitude" precision="1" value="45.29167">45.29167</geoCoordinate>
; alt. 500 m; 10 Feb 2003; Fisher et al. leg.; CASENT0031767 (CASC). - 
<emphasis id="2DC83900DC9DEF3E589F4EADD83EE104" bold="true" italics="true" pageId="0" pageNumber="1">Toliara</emphasis>
: •7w., 4s., 3q.; 
<normalizedToken id="4210075314B9B5053EB48D88660F2473" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="DC74E87AF23D18B0C9D61FFC192E8ECA" originalValue="Classée">Classee</normalizedToken>
<normalizedToken id="046257DE4580BCC4EACEE9EEEF344D07" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
, 29.2 km 343°NNW Mahaboboka; -22.675, 44.19; alt. 1100 m; 18 Feb 2003; Fisher et al. leg.; CASENT0031011, CASENT0031099, CASENT0049501, CASENT0073507, CASENT0235043, CASENT0496936, CASENT0496937, CASENT0496938 (CASC). •1w., 5s.; 
<normalizedToken id="016140602B96821EB331D6F9F9EBECE8" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="8253A12D5F42D88757BCEAE279768B49" originalValue="Classée">Classee</normalizedToken>
<normalizedToken id="A9F82C85C45C985573BE385F0A38EBD8" originalValue="d’Analavelona">d'Analavelona</normalizedToken>
, 29.4 km 343°NNW Mahaboboka; 
<geoCoordinate id="CD1DCF1F32D4440329B8A75649B7FB94" degrees="22.675" direction="south" orientation="latitude" precision="55" value="-22.675">-22.675</geoCoordinate>
, 
<geoCoordinate id="00963321A539EC87A6A0CAD512EADCFC" degrees="44.18667" direction="east" orientation="longitude" precision="1" value="44.18667">44.18667</geoCoordinate>
; alt. 1050 m; 21 Feb 2003; Fisher et al. leg.; CASENT0023935, CASENT0023938, CASENT0023943, CASENT0023946, CASENT0023947, CASENT0024032 (CASC).
</paragraph>
</subSubSection>
<subSubSection id="3F7C83100AB2BC87071257797B11A0DB" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="E3B2F4B0663867F9DCD4AD2D59FD736F" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="02EC153C6674B057B2A71D31FC2FCBDA" pageId="0" pageNumber="1">
<emphasis id="AB61AFFB9982F2CFA8D319E637F304E6" bold="true" pageId="0" pageNumber="1">
<emphasis id="33A200022A5CE53511E6FDE16CBC12C7" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Head in full-face view sub-oval; sides of the head with dense, long, erect pilosity; occipital lobes shiny, with thick, irregular rugae, interspaces with distinct, irregular rugulae not fading posteriorly; inner hypostomal teeth distinct, high, closely spaced, triangular, with rounded apex directed inward, and wide base; outer hypostomal teeth taller and wider than inner hypostomal teeth, lobe-like, directed outward; gaster smooth. 
<emphasis id="0000A83E7B20D9A13FFA3EEBB3123054" bold="true" pageId="0" pageNumber="1">
<emphasis id="B9B88286455B05723D3219E272823BE0" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Head shiny, foveolate, with additional short, longitudinal, thick rugae on frons, genae with fading sculpture and smooth notch; scape, when laid back, surpassing the posterior head margin by one-fifth of its length; promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; mesosoma foveolate, katepisternum smooth; propodeal spines short, triangular.
</paragraph>
</subSubSection>
<subSubSection id="62049B4575DB927328C07DD24713FC0C" pageId="0" pageNumber="1" type="description">
<paragraph id="1B46FABD83999E1A25D1BBC80DFB7626" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="BB085002939D10244622E830679E58F6" pageId="0" pageNumber="1">
<emphasis id="377D2D1656025EF99E1EE9BCFB174011" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis id="CBAA46D82A84ED9FBCC2E92C4DC33D0A" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 1.05-1.11 (1.09); HW: 0.98-1.04 (1.0); SL: 0.46-0.52 (0.48); EL: 0.13-0.16 (0.13); WL: 0.84-0.97 (0.88); PSL: 0.14-0.17 (0.15); MTL: 0.47-0.5 (0.49); PNW: 0.52-0.58 (0.54); PTW: 0.12-0.14 (0.13); PPW: 0.35-0.4 (0.37); CI: 90.0-93.9 (91.7); SI: 46.5-51.8 (48.1); PSLI: 12.5-15.4 (13.7); PPI: 31.9-39.0 (34.9); PNI: 53.1-55.5 (54.3); MTI: 45.8 -50.9 (48.8). 
<emphasis id="3D8716A3C526AC3415BE347C0FA5A51C" bold="true" pageId="0" pageNumber="1">
<emphasis id="09A44ED83DE5091F8877AF3A53919B26" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view longer than wide, anterior of eyes slightly convex, posterior of eyes convex (Fig. 
<figureCitation id="5829CEAC308D0E7F12E3991D48662F1A" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39B</figureCitation>
). In lateral view sub-oval; ventral and dorsal faces convex; dorsal face not depressed posteriorly; inner hypostomal teeth visible. Sides of the head with dense, very long, erect pilosity; whole head with moderately dense, long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, with thick, longitudinal, and long rugae; interspaces distinctly rugulose. Occipital lobes shiny, with thick, irregular rugae, interspaces with distinct, irregular rugulae not fading posteriorly; frons with moderately dense, thick, longitudinal rugae, interspaces smooth, with indistinct rugulae weakening anteriorly; genae shiny, with dense and thin, irregular rugulae, central part with smooth notch; malar area with thin, longitudinal, dense rugae, interspaces with distinct rugulae. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugulae; median notch present, narrow, and moderately deep; median longitudinal carina present, indistinct; lateral longitudinal carinae present. Scape, when laid back, reaching the midlength of head; pilosity erect (Fig. 
<figureCitation id="873B24F05421BC8BEB74CB17AEDE352F" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39B, D</figureCitation>
). Inner hypostomal teeth distinct, high, closely spaced, triangular, with rounded apex directed inward, and wide base; outer hypostomal teeth taller and wider than inner hypostomal teeth, lobe-like, directed outward (Fig. 
<figureCitation id="C7283CBBBB634EDDB1818A7B75B52871" captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84X</figureCitation>
). 
<emphasis id="BCDD8214C2711FE51FE1B8F29C238593" bold="true" pageId="0" pageNumber="1">
<emphasis id="5F78A81E87904AF9B19D775BB8E90C15" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum relatively high and arched, posterior mesonotum relatively steep, with tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines small to moderately long, triangular, with sharp apex and wide base; humeral area laterally weakly produced (Fig. 
<figureCitation id="4E173D55AF8802C6F50748C94A893D14" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D</figureCitation>
). Surface shiny, promesonotum foveolate with additional indistinct, sparse rugoreticulations, sometimes sculpture fading on dorsal surface and lower parts of lateral sides; katepisternum rugoreticulate, with smooth notch; anepisternum and lateral sides of propodeum with distinct rugoreticulation; dorsoventral side of propodeum smooth or with indistinct sculpture. Pilosity moderately dense, very long, and erect (Fig. 
<figureCitation id="AB2565D82032CF0FDD8A624443464069" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
). 
<emphasis id="22BF66EC4E2089FC40F71AFD1A1511EA" bold="true" pageId="0" pageNumber="1">
<emphasis id="22082A09B4770271D13C600E07A079E6" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny; peduncle short, finely foveolate, with indistinct horizontal lobes on its basal part; node with fading sculpture, relatively low, triangular, with rounded apex, in rear view node slightly convex; pilosity long and erect (Fig. 
<figureCitation id="6D6C814110679E2C4FB6A615FCF5DD3C" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
). 
<emphasis id="343FC1C803DE33976B94121D4F99629F" bold="true" pageId="0" pageNumber="1">
<emphasis id="782D3CDDD5559896B8F27ABD0CF843EA" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny, finely shagreened, with dorsum at least partially smooth; in dorsal view sides with moderately short, acute, and triangular projections; pilosity long, long and erect (Fig. 
<figureCitation id="1F663A741CE110C672B4715B658B6395" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
). 
<emphasis id="50855C5250E2DAE24535BAF2A62167AC" bold="true" pageId="0" pageNumber="1">
<emphasis id="C794BB784EF89A5034597EDCC412ACDF" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and smooth, only basal part of first gastral tergite finely shagreened; pilosity dense, long, and erect (Fig. 
<figureCitation id="ABAEA51C301F3F0ACB9FF96497695D84" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
). 
<emphasis id="0D229780F10A5B612B33AF3D7C37D9A8" bold="true" pageId="0" pageNumber="1">
<emphasis id="9C1C026DBA378C513125F60D3A9B6685" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Reddish brown to bright brown (Fig. 
<figureCitation id="582E5A24DD3DB662F24525B2C342CCFE" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39D, F</figureCitation>
).
</paragraph>
<caption id="52E5D70CC32951F30F4E13CFD936A47A" doi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1" start="Figure 39" startId="F39">
<paragraph id="F1453EFAEF545E51F24FECF9845927EF" pageId="0" pageNumber="1">
<emphasis id="5AAFFAAD66612B1DE0836C495D435F9E" bold="true" pageId="0" pageNumber="1">Figure 39.</emphasis>
<taxonomicName id="A73FC1556AB9557AFBCAE661D548D1C9" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole longipilosa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="longipilosa">
<emphasis id="0B957F7DB174F3755D27D2F9B8E2DDB7" italics="true" pageId="0" pageNumber="1">Pheidole longipilosa</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis id="8F6282D5C253BDF106E185F6D2D7B883" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis id="0C0410F88821572B0D42846E998FE228" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis id="7EEFD4ED0F4D7C78B3F164D274B85228" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0235043) and full-face view (
<emphasis id="D8718A7D1B8E246E180B552099C82B15" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis id="9FCFA945E0617032791C01AA23353C74" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis id="44EFD410BB910417044593E48E9BC0BF" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0498298).
</paragraph>
</caption>
<paragraph id="399188C2C8194FB66D789CB6C825681C" pageId="0" pageNumber="1">
<emphasis id="2646087E78E2FE389A0B728646CF7A1A" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis id="3F4EFADF34D764C913FC8A99EB410A0E" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 0.49-0.55 (0.51); HW: 0.44-0.47 (0.45); SL: 0.46-0.47 (0.46); EL: 0.09-0.11 (0.1); WL: 0.57-0.6 (0.58); PSL: 0.09-0.11 (0.1); MTL: 0.33-0.36 (0.34); PNW: 0.29-0.32 (0.31); PTW: 0.06-0.08 (0.07); PPW: 0.1-0.12 (0.11); CI: 85.7-90.4 (88.6); SI: 100.2-105.6 (103.0); PSLI: 18.1-21.8 (20.0); PPI: 56.5-64.5 (60.8); PNI: 65.9-70.1 (68.2); MTI: 73.5-80.1 (76.1). 
<emphasis id="3617028DC05DCA78B9ACFC85093352EC" bold="true" pageId="0" pageNumber="1">
<emphasis id="841454701A29D1F32200ADC541BA50B8" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina absent (Fig. 
<figureCitation id="D407665653ED22DB790D9AB89AE396DF" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39A</figureCitation>
). Pilosity moderately sparse, moderately long, suberect to erect. Head shiny, foveolate, with additional short, longitudinal, thick rugae on frons; genae with fading sculpture and smooth notch. Clypeus smooth, with basal part finely rugulose; median longitudinal carina present; two lateral longitudinal carinae indistinct. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity suberect to erect (Fig. 
<figureCitation id="1F14F4D1D981ADAF2DCD51948F713421" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39A, C</figureCitation>
). 
<emphasis id="AB2CF3918B528A8CA592FB1BA7B82DDD" bold="true" pageId="0" pageNumber="1">
<emphasis id="789776920B5680264E1E3C148AD8BDD5" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, convex, short, with posterior declivity smoothly declining towards propodeum; promesonotal groove absent; metanotal groove absent; propodeal spines short, triangular, with acute apex (Fig. 
<figureCitation id="D4C845D79DD3B5D3D392D1D95DD906CF" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C</figureCitation>
). Sculpture foveolate; katepisternum smooth; anepisternum and mesonotum with fading sculpture and sometimes smooth. Pilosity moderately sparse, long, and erect (Fig. 
<figureCitation id="B8B1989009A2F086F3B90B85B47165A6" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
). 
<emphasis id="513D67D9F5D1000A644228844D6ADDA3" bold="true" pageId="0" pageNumber="1">
<emphasis id="90BDE4835CC5AEE785FC349352F66040" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle short and thin with ventral face slightly convex; with few short, erect setae (Fig. 
<figureCitation id="F04DDE84CF3C9CB42B2B3082DD68D9A9" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
). 
<emphasis id="989D58046BD4E7CB8B7936360522FC94" bold="true" pageId="0" pageNumber="1">
<emphasis id="01951126A1F370A09C29CDB6E6A6C14D" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and convex; with few short, erect setae (Fig. 
<figureCitation id="B8BA95041141CE2A678ED8579A216D13" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
). 
<emphasis id="6658337918FDC025AAE24E4D45E7C016" bold="true" pageId="0" pageNumber="1">
<emphasis id="59CE5EB3F4ED0918F6068A1C219E99BE" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With moderately sparse, erect pilosity (Fig. 
<figureCitation id="DD8516A997AF1F6DC4CB04D07629FC05" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
). 
<emphasis id="536A67E6CCA011FB8DBF34EA647FD80E" bold="true" pageId="0" pageNumber="1">
<emphasis id="A30086538DB451D98EAFB95E840B7745" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Yellow (Fig. 
<figureCitation id="D249B7E7E3DED059334E1923373AA5AB" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Pheidole longipilosa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0235043) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0498298)." figureDoi="10.3897/zookeys.905.39592.figure39" httpUri="https://binary.pensoft.net/fig/373086" pageId="0" pageNumber="1">39C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BB39166B1AF505784EB42720EB56FB28" pageId="0" pageNumber="1" type="etymology">
<paragraph id="AF602AB98C27003504AA898C56D4C2F7" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="83FC8721C46D978C2BECFAF02FB10986" pageId="0" pageNumber="1">Latin for long pilosity, in reference to the very long pilosity on sides of head in major workers.</paragraph>
</subSubSection>
<subSubSection id="F0B3D1DA5697300626DF9A74C1864A71" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="638C23B63AD64F2CAC66082F4172115F" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="6CF63A6E37E9633DBEC21984BBFFA1EA" pageId="0" pageNumber="1">The species was collected between 500-1100 m in elevation, in montane and gallery forests. Nests were located in rotten logs.</paragraph>
</subSubSection>
</treatment>
</document>