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<document id="939F7B1E4D88956C0BBA02D1F80D62FB" ID-CLB-Dataset="5898" ID-DOI="10.11646/zootaxa.3768.4.5" ID-GBIF-Dataset="985f8c4c-c9ad-482b-8e0a-c25f6580b9bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="4909658" ID-ZooBank="DBDE3685-85AC-4739-95DB-F8BA16117AAB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623114156328" checkinUser="felipe" docAuthor="Sihvonen, Pasi, Skou, Peder, Flamigni, Claudio, Fiumi, Gabriele &amp; Hausmann, Axel" docDate="2014" docId="C37B8D572E2EC93AFF72B7B35CAA753A" docLanguage="en" docName="zt03768p486.pdf" docOrigin="Zootaxa 3768 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hylaea pinicolaria" docType="treatment" docVersion="9" lastPageNumber="482" masterDocId="3F42F52F2E22C937FFE5B3585D257667" masterDocTitle="Revision of the Hylaea fasciaria (Linnaeus, 1758) species group in the western Palaearctic (Lepidoptera: Geometridae, Ennominae)" masterLastPageNumber="486" masterPageNumber="469" pageNumber="481" updateTime="1699002952716" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="905649DD339C4DC2118C718F7F4CE583">Revision of the Hylaea fasciaria (Linnaeus, 1758) species group in the western Palaearctic (Lepidoptera: Geometridae, Ennominae)</mods:title>
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<mods:namePart id="1B83CC2BE23ACA04FBF9FF69A25EECF3">Sihvonen, Pasi</mods:namePart>
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<mods:namePart id="370D3077F62F125CF90F13DE4B3424B9">Skou, Peder</mods:namePart>
<mods:affiliation id="130B98B156A72BBB942639480DBF0BBB">Aamosen 1, DK 5762 Vester Skerninge, Denmark. E-mail: info @ apollobooks. dk</mods:affiliation>
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<mods:namePart id="33EFE754E07CF8B6AC64669FA050B563">Flamigni, Claudio</mods:namePart>
<mods:affiliation id="82FC663D587F96AA276CD75D375202B1">Via delle Belle Arti 21, I- 40126 Bologna, Italy. E-mail: claudio. flamigni @ alice. it</mods:affiliation>
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<mods:namePart id="D5037D93107393C482B6990DBE87CC48">Fiumi, Gabriele</mods:namePart>
<mods:affiliation id="9A2AADB7E96BD52302972663214D2CA1">Via Decio Raggi 167, I- 47121 Forlí, Italy. E-mail: gabfium @ tiscali. it</mods:affiliation>
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<mods:namePart id="0E8072666DC2A0853D4EB91FB863AFDD">Hausmann, Axel</mods:namePart>
<mods:affiliation id="32324ABC796E396899CDBFD0A087DB23">Bavarian State Collection of Zoology (ZSM), Münchhausenstr. 21, 81247 Munich, Germany - Bavarian Natural History Collections (SNSB), Menzinger Str. 71, 80638 Munich, Germany. E-mail: axel. hausmann @ zsm. mwn. de</mods:affiliation>
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<mods:number id="271C51E003CA82CDF2B97952BA676E3F">3768</mods:number>
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<treatment id="C37B8D572E2EC93AFF72B7B35CAA753A" ID-DOI="http://doi.org/10.5281/zenodo.5047213" ID-GBIF-Taxon="183228139" ID-Zenodo-Dep="5047213" LSID="urn:lsid:plazi:treatment:C37B8D572E2EC93AFF72B7B35CAA753A" httpUri="http://treatment.plazi.org/id/C37B8D572E2EC93AFF72B7B35CAA753A" lastPageId="13" lastPageNumber="482" pageId="12" pageNumber="481">
<subSubSection id="03C86FCA2E2EC93BFF72B7B35F097362" box="[151,556,1259,1285]" pageId="12" pageNumber="481" type="nomenclature">
<paragraph id="4B6D3C412E2EC93BFF72B7B35F097362" blockId="12.[151,556,1259,1285]" box="[151,556,1259,1285]" pageId="12" pageNumber="481">
<heading id="10258B2D2E2EC93BFF72B7B35F097362" bold="true" box="[151,556,1259,1285]" fontSize="11" level="1" pageId="12" pageNumber="481" reason="1">
<taxonomicName id="8CD247C22E2EC93BFF72B7B35F097362" ID-CoL="3NDGG" authority="(Bellier, 1861)" baseAuthorityName="Bellier" baseAuthorityYear="1861" box="[151,556,1259,1285]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="pinicolaria">
<emphasis id="79A6E0532E2EC93BFF72B7B35F097362" bold="true" box="[151,556,1259,1285]" pageId="12" pageNumber="481">
<emphasis id="79A6E0532E2EC93BFF72B7B35C5D7362" bold="true" box="[151,376,1259,1285]" italics="true" pageId="12" pageNumber="481">Hylaea pinicolaria</emphasis>
(
<bibRefCitation id="2F4341B02E2EC93BFE62B7B35F017362" author="Bellier, C." box="[391,548,1259,1285]" pageId="12" pageNumber="481" pagination="29 - 30" refId="ref10615" refString="Bellier, C. (1861) Description de trois lepidopteres nouveaux de l'ile de Corse. Annales de la Societ entomologique de France, 1, 29 - 30." type="journal article" year="1861">Bellier, 1861</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="03C86FCA2E2EC93BFF72B66A5EB273E2" pageId="12" pageNumber="481" type="reference_group">
<paragraph id="4B6D3C412E2EC93BFF72B66A5EB273E2" blockId="12.[151,1436,1330,1413]" pageId="12" pageNumber="481">
<taxonomicName id="8CD247C22E2EC93BFF72B66A5CB6732F" ID-CoL="3NDGG" authority="Bellier, 1861" authorityName="Bellier" authorityYear="1861" box="[151,403,1330,1352]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="pinicolaria">
<emphasis id="79A6E0532E2EC93BFF72B66A5C2D732F" box="[151,264,1330,1352]" italics="true" pageId="12" pageNumber="481">pinicolaria</emphasis>
<materialsCitation id="FBBA361C2E2EC93BFEEBB66A5CB6732F" ID-GBIF-Occurrence="3312254304" box="[270,403,1330,1352]" collectorName="C. Oberthur" country="United Kingdom" location="UK" pageId="12" pageNumber="481" specimenCount="1" typeStatus="syntype">
<bibRefCitation id="2F4341B02E2EC93BFEEBB66A5CB6732F" author="Bellier, C." box="[270,403,1330,1352]" pageId="12" pageNumber="481" pagination="29 - 30" refId="ref10615" refString="Bellier, C. (1861) Description de trois lepidopteres nouveaux de l'ile de Corse. Annales de la Societ entomologique de France, 1, 29 - 30." type="journal article" year="1861">Bellier, 1861</bibRefCitation>
</materialsCitation>
</taxonomicName>
, Annls Soc. ent. Fr. (4) 1 (1): 29, pl. 2, fig. 12, (
<taxonomicName id="8CD247C22E2EC93BFC9CB66A5EE1732F" box="[889,964,1330,1352]" class="Insecta" family="Geometridae" genus="Ellopia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="79A6E0532E2EC93BFC9CB66A5EE1732F" box="[889,964,1330,1352]" italics="true" pageId="12" pageNumber="481">Ellopia</emphasis>
</taxonomicName>
).
<typeStatus id="946982E32E2EC93BFC3DB66A5914732F" box="[984,1073,1330,1352]" pageId="12" pageNumber="481" type="syntype">Syntype</typeStatus>
(s), Corsica (mountains of). Bellier's collection was apparently included in the
<collectorName id="E62759972E2EC93BFD68B6095E2E7300" box="[653,779,1361,1383]" pageId="12" pageNumber="481">C. Oberthür</collectorName>
collection, which is currently housed in the Natural History Museum, London,
<collectingCountry id="33C57CD12E2EC93BFE62B6285C8A73E2" box="[391,431,1392,1413]" name="United Kingdom" pageId="12" pageNumber="481">UK</collectingCountry>
. Despite searches, we have not located the type.
</paragraph>
</subSubSection>
<subSubSection id="03C86FCA2E2EC93AFF72B6EA5FA476B2" lastPageId="13" lastPageNumber="482" pageId="12" pageNumber="481" type="description">
<paragraph id="4B6D3C412E2EC93BFF72B6EA5F53708C" blockId="12.[151,1438,1458,2023]" pageId="12" pageNumber="481">
<emphasis id="79A6E0532E2EC93BFF72B6EA5C0C73AC" bold="true" box="[151,297,1458,1483]" inLineHeading="true" pageId="12" pageNumber="481" reason="1">Description.</emphasis>
<emphasis id="79A6E0532E2EC93BFED3B6EA5E6C73AC" box="[310,841,1458,1483]" italics="true" pageId="12" pageNumber="481">External characters and pregenital abdomen</emphasis>
(
<figureCitation id="D3E920C42E2EC93BFCBBB6EA5EF473AC" box="[862,977,1458,1483]" captionStart="FIGURES 15" captionStartId="4.[151,264,1398,1420]" captionTargetBox="[183,1386,187,1369]" captionTargetId="figure-357@4.[172,1404,187,1386]" captionTargetPageId="4" captionText="FIGURES 15. Adults of the Palaearctic Hylaea fasciaria species group. 1a. Hylaea fasciaria (L.) male, 13.vi.1987, Volsk, Saratov Region, South East Russia (coll. Skou). 1b. H. fasciaria male, 6.vii.1972, Skanør, Falsterbo, Skåne, Sweden (coll Skou). 1c. H. fasciaria male, 1.viii.1972, Byerum, Öland, Sweden (coll. Skou). 1d. H. fasciaria male, 18.vii.1970, Dueodde, Bornholm, Denmark (coll. Skou). 1e. H. fasciaria flavella (Wehrli), type, original illustration from Wehrli (19391954, plate 24: g), Armenia, without date (coll. unknown). 1f. H. fasciaria female, 6.vii.1972, Skanør, Falsterbo, Skåne, Sweden (coll. Skou). 1g. H. fasciaria female, 29.vii.2000, Terre Rouge, Briancon, Dept. Hautes Alpes, France (coll. Skou). 1h. H. fasciaria cedricola (Wehrli) male, ex ovo, Bey Dağlari, Antalya, Turkey (coll. Müller). 1i. H. fasciaria cedricola female, ex ovo, Bey Dağlari, Antalya, Turkey (coll. Müller). 1j. H. fasciaria male, 2728.vii.1983, Vorarlberg, 1500 m, Austria (coll. Skou). 1k. H. fasciaria male, 10.vi.2010, Mount Parnassos, 1650 m, Greece (coll. Skou). 1l. H. fasciaria female, 2728.vii.1983, Vorarlberg, 1500 m, Austria (coll. Skou). 1m. H. fasciaria female, 28.vi.1981, Karpenision, Aetolia, Greece (coll. Skou). 2a. Hylaea mediterranea new species male, 910.x.2010, Pizzo della Rondine, Sicily, Italy (coll. Skou), holotype. 2b H. mediterranea male, 910.x.2010, Pizzo della Rondine, Sicily, Italy (coll. Skou), paratype. 2c. H. mediterranea female, 910.x.2010, Pizzo della Rondine, Sicily, Italy (coll. Skou), paratype. 2d. H. mediterranea female, 5.vi.2005, Castelbuono, Sicily, Italy (coll. Skou), paratype. 3a. Hylaea pinicolaria (Bellier) male, 2021.vii.2004, Camping Monte Cinto, Haut-Asco, Corsica, France (coll. Skou). 3b. H. pinicolaria male, 2021.vii.2004, Camping Monte Cinto, Haut-Asco, Corsica, France (coll. Skou). 3c. H. pinicolaria male, 6.vii.1951, La Foce, Vizzavona, Corsica, France (coll. Herbulot, coll. ZSM). 3d. H. pinicolaria female, 23.vi.1994, Haut Asco, Corsica, France (coll. Skou). 4a. Hylaea compararia (Staud.) male, 23.vi.1911, Blida, Glacieres, Algeria (coll. BMNH). 4b. Hylaea compararia female, 13.ix.1911, Glacieres, Blida, Algeria (coll. BMNH). 4c. Hylaea compararia female, 15.ix.1911, Glacieres, Blida, Algeria (coll. BMNH). 5a. Hemithea squalidaria O. G. Costa, original illustration from O. G. Costa, [1848]. Illustration is reproduced under a licence from the Picture Library of the Natural History Museum, London, UK." figureDoi="http://doi.org/10.5281/zenodo.4909660" httpUri="https://zenodo.org/record/4909660/files/figure.png" pageId="12" pageNumber="481">Figures 3</figureCitation>
,
<figureCitation id="D3E920C42E2EC93BFC01B6EA5ED773AC" box="[996,1010,1458,1483]" captionStart="FIGURES 69" captionStartId="5.[151,264,1100,1122]" captionTargetBox="[152,1435,201,1054]" captionTargetId="figure-391@5.[151,1435,201,1061]" captionTargetPageId="5" captionText="FIGURES 69. An overview of diagnostic characters in the Palaearctic Hylaea fasciaria species group. Figure 6. H. fasciaria (Linnaeus) (Denmark, 27.vi.1977). Figure 7. H. mediterranea, new species (Italy, Sicily, 5.vi.2005, paratype). Figure 8. H. pinicolaria (Bellier) (France, Corsica, 20.-21.vii.2004). Figure 9. H. compararia (Staudinger) (Algeria, 23.vi.1911)." figureDoi="http://doi.org/10.5281/zenodo.4909662" httpUri="https://zenodo.org/record/4909662/files/figure.png" pageId="12" pageNumber="481">8</figureCitation>
) (diagnostic characters underlined): Wingspan male
<quantity id="8C2A91A42E2EC93BFEAAB68E5CE97389" box="[335,460,1494,1519]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="12" pageNumber="481" unit="mm" value="33.0" valueMax="34.0" valueMin="32.0">3234 mm</quantity>
, female
<quantity id="8C2A91A42E2EC93BFDCBB68E5F897389" box="[558,684,1494,1519]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="12" pageNumber="481" unit="mm" value="40.5" valueMax="42.0" valueMin="39.0">3942 mm</quantity>
. Wings light green, medial lines white. Medial line curved before costa, basal part parallel with costa. Postmedial line rather straight, only weakly curved, angled before it reaches costa well before apex and evenly curved outwards on inner margin. Medial area concolorous with rest of wing. Terminal line and fringes concolorous with wings, forewing apex dark red. Hindwing postmedial line distinct, curved. Discal spots absent. Wings below as above, but paler. Frons red-brown, thorax and abdomen concolorus with wings. Area between antennae (vertex) white. Antennae white dorsally, male antennae bipectinate, female antennae fasciculate. Hindleg tibia of both sexes with 2+2 spurs. Tympanal organs medium-sized. Sternites and tergites 38 of both sexes undifferentiated.
</paragraph>
<paragraph id="4B6D3C412E2EC93BFF22B5AE5E4E711C" blockId="12.[151,1438,1458,2023]" pageId="12" pageNumber="481">
<emphasis id="79A6E0532E2EC93BFF22B5AE5C557168" box="[199,368,1782,1807]" italics="true" pageId="12" pageNumber="481">Male genitalia</emphasis>
(
<figureCitation id="D3E920C42E2EC93BFE67B5AE5CDC7168" box="[386,505,1782,1807]" captionStart="FIGURES 1013" captionStartId="6.[151,264,1641,1663]" captionTargetBox="[212,1348,195,1603]" captionTargetId="figure-233@6.[212,1349,195,1603]" captionTargetPageId="6" captionText="FIGURES 1013. Male genitalia of the Palaearctic Hylaea fasciaria species group. Hylaea fasciaria (Linnaeus). 10a. genitalia, Finland: Enonkoski, 1618.vii.1996, slide PS1344 (coll. P. Sihvonen). 10b. aedeagus, Germany: Bad Harzburg, 2.vii.1988, slide PS1741 (coll. P. Sihvonen). 10c. vesica, Finland: Enonkoski, 1618.vii.1996, slide PS1344 (coll. P. Sihvonen). 10d. base of vesica, Finland: Enonkoski, 1618.vii.1996, slide PS1344 (coll. P. Sihvonen). Hylaea mediterranea new species 11a. genitalia, Italy: Sicily, 1000 m, 910.x.2010, slide PS1738, holotype (coll Skou). 11b. aedeagus, Italy: Sicily, 1000 m, 9 10.x.2010, slide PS1738, holotype (coll Skou). 11c. vesica, Italy: Sicily, 1000 m, 910.x.2010, slide PS1738, holotype (coll Skou). 11d. base of vesica, Italy: Sicily, 1000 m, 910.x.2010, slide PS1738, holotype (coll Skou). Hylaea pinicolaria (Bellier) 12a. genitalia, France: Corsica, 1100 m, 2021.vii.2004, slide PS1542 (coll. Skou). 12b. aedeagus, France: Corsica, 1100 m, 2021.vii.2004, slide PS1742 (coll. Skou). 12c. vesica, France: Corsica, 1100 m, 2021.vii.2004, slide PS1542 (coll. Skou). 12d. base of vesica, France: Corsica, 1100 m, 2021.vii.2004, slide PS1542 (coll. Skou). Hylea compararia (Staudinger) 13a. genitalia, Algeria: Blida, 23.vi.1911, slide PS1866/BMNH GEO 24840 (coll. BMNH). 13b. aedeagus, Algeria: Blida, 23.vi.1911, slide PS1866/BMNH GEO 24840 (coll. BMNH). 13c. vesica, Algeria: Blida, 5.vi.1908, slide PS1870/BMNH GEO 24844 (coll. BMNH). 13d. base of vesica, Algeria: Blida, 5.vi.1908, slide PS1870/BMNH GEO 24844 (coll. BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4909664" httpUri="https://zenodo.org/record/4909664/files/figure.png" pageId="12" pageNumber="481">Figure 12</figureCitation>
): Generally as in
<taxonomicName id="8CD247C22E2EC93BFD34B5AF5EC57168" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[721,992,1782,1807]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="fasciaria">
<emphasis id="79A6E0532E2EC93BFD34B5AF5E7E7168" box="[721,859,1782,1807]" italics="true" pageId="12" pageNumber="481">H. fasciaria</emphasis>
(Linnaeus)
</taxonomicName>
and
<taxonomicName id="8CD247C22E2EC93BFBC5B5AF59F87168" authorityName="Sihvonen &amp; Skou &amp; Flamigni &amp; Fiumi &amp; Hausmann" authorityYear="2014" box="[1056,1245,1782,1807]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="mediterranea">
<emphasis id="79A6E0532E2EC93BFBC5B5AF59F87168" box="[1056,1245,1782,1807]" italics="true" pageId="12" pageNumber="481">H. mediterranea</emphasis>
</taxonomicName>
. Aedeagus with weakly curved additional arm, apex not expanded in
<taxonomicName id="8CD247C22E2EC93BFD10B4435EB37154" authorityName="Bellier" authorityYear="1861" box="[757,918,1818,1843]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="pinicolaria">
<emphasis id="79A6E0532E2EC93BFD10B4435EB37154" box="[757,918,1818,1843]" italics="true" pageId="12" pageNumber="481">H. pinicolaria</emphasis>
</taxonomicName>
(additional arm straight in
<taxonomicName id="8CD247C22E2EC93BFB35B443587D7154" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1232,1368,1818,1843]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="fasciaria">
<emphasis id="79A6E0532E2EC93BFB35B443587D7154" box="[1232,1368,1818,1843]" italics="true" pageId="12" pageNumber="481">H. fasciaria</emphasis>
</taxonomicName>
, apex expanded in
<taxonomicName id="8CD247C22E2EC93BFEC0B4675CEF7131" baseAuthorityName="Staudinger" baseAuthorityYear="1894" box="[293,458,1855,1878]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="compararia">
<emphasis id="79A6E0532E2EC93BFEC0B4675CEF7131" box="[293,458,1855,1878]" italics="true" pageId="12" pageNumber="481">H. compararia</emphasis>
</taxonomicName>
). Base of vesica with straight row of microcornuti in
<taxonomicName id="8CD247C22E2EC93BFBFBB46759987130" authorityName="Bellier" authorityYear="1861" box="[1054,1213,1854,1879]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="pinicolaria">
<emphasis id="79A6E0532E2EC93BFBFBB46759987130" box="[1054,1213,1854,1879]" italics="true" pageId="12" pageNumber="481">H. pinicolaria</emphasis>
</taxonomicName>
(vesica with angled row of microcornuti, reaching aedeagus apex in
<taxonomicName id="8CD247C22E2EC93BFD51B43B5E7F711D" baseAuthorityName="Staudinger" baseAuthorityYear="1894" box="[692,858,1891,1914]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="compararia">
<emphasis id="79A6E0532E2EC93BFD51B43B5E7F711D" box="[692,858,1891,1914]" italics="true" pageId="12" pageNumber="481">H. compararia</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="4B6D3C412E2EC93AFF22B4DE5FA476B2" blockId="12.[151,1438,1458,2023]" lastBlockId="13.[151,1437,151,861]" lastPageId="13" lastPageNumber="482" pageId="12" pageNumber="481">
<emphasis id="79A6E0532E2EC93BFF22B4DE5CAA71F8" box="[199,399,1926,1951]" italics="true" pageId="12" pageNumber="481">Female genitalia</emphasis>
(
<figureCitation id="D3E920C42E2EC93BFE43B4DE5F0471F8" box="[422,545,1926,1951]" captionStart="FIGURES 1417" captionStartId="7.[151,264,1592,1614]" captionTargetBox="[160,1436,193,1559]" captionTargetId="figure-213@7.[160,1445,193,1559]" captionTargetPageId="7" captionText="FIGURES 1417. Female genitalia of the Palaearctic Hylaea fasciaria species group. Circle indicates the point of origin of the ductus seminalis. Hylaea fasciaria (Linnaeus) 14a. genitalia, Finland: Föglö, 28.vii.1995, slide PS1448 (coll. P. Sihvonen). 14b. reduced signum, Finland: Föglö, 28.vii.1995, slide PS1448 (coll. P. Sihvonen). 14c. ostium bursae and adjacent structures, Finland: Föglö, 28.vii.1995, slide PS1448 (coll. P. Sihvonen). Hylaea mediterranea new species 15a. genitalia, Italy: Sicily, 1000 m, 910.x.2010, slide PS1739, paratype (coll. Skou). 15b. signum, Italy: Sicily, 1000 m, 910.x.2010, slide PS1739, paratype (coll. Skou). 15c. ostium bursae and adjacent structures, Italy: Sicily, 1000 m, 910.x.2010, slide PS1739, paratype (coll. Skou). Hylaea pinicolaria (Bellier) 16a. genitalia, France: Corsica, 1100 m, 2021.vii.2004, slide PS1543 (coll. Skou). 16b. reduced signum, France: Corsica, 1100 m, 2021.vii.2004, slide PS1543 (coll. Skou). 16c. ostium bursae and adjacent structures, France: Corsica, 1100 m, 2021.vii.2004, slide PS1543 (coll. Skou). Hylaea compararia (Staudinger) 17a. genitalia, Algeria: Blida, 13.ix.1911, slide PS1868/BMNH GEO 24842, margins of the corpus bursae highlighted (coll. BMNH). 17b. posterior part of corpus bursae (signum absent), Algeria: Blida, 13.ix.1911, slide PS1868/BMNH GEO 24842 (coll. BMNH). 17c. ostium bursae and adjacent structures, Algeria: Blida, 15.ix.1911, slide PS1871/BMNH GEO 24845 (coll. BMNH)." figureDoi="http://doi.org/10.5281/zenodo.4909666" httpUri="https://zenodo.org/record/4909666/files/figure.png" pageId="12" pageNumber="481">Figure 16</figureCitation>
): Generally as in
<taxonomicName id="8CD247C22E2EC93BFCECB4DF5EBD71F8" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[777,920,1926,1951]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="fasciaria">
<emphasis id="79A6E0532E2EC93BFCECB4DF5EBD71F8" box="[777,920,1926,1951]" italics="true" pageId="12" pageNumber="481">H. fasciaria</emphasis>
</taxonomicName>
and
<taxonomicName id="8CD247C22E2EC93BFC05B4DF598671F8" authorityName="Sihvonen &amp; Skou &amp; Flamigni &amp; Fiumi &amp; Hausmann" authorityYear="2014" box="[992,1187,1926,1951]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="mediterranea">
<emphasis id="79A6E0532E2EC93BFC05B4DF598671F8" box="[992,1187,1926,1951]" italics="true" pageId="12" pageNumber="481">H. mediterranea</emphasis>
</taxonomicName>
, but with following quantitative difference: signum large in
<taxonomicName id="8CD247C22E2EC93BFDB9B4F35E3371A4" authorityName="Sihvonen &amp; Skou &amp; Flamigni &amp; Fiumi &amp; Hausmann" authorityYear="2014" box="[604,790,1962,1987]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="mediterranea">
<emphasis id="79A6E0532E2EC93BFDB9B4F35E3371A4" box="[604,790,1962,1987]" italics="true" pageId="12" pageNumber="481">H. mediterranea</emphasis>
</taxonomicName>
(signum absent or minute in
<taxonomicName id="8CD247C22E2EC93BFB83B4F359C871A4" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1126,1261,1962,1987]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="fasciaria">
<emphasis id="79A6E0532E2EC93BFB83B4F359C871A4" box="[1126,1261,1962,1987]" italics="true" pageId="12" pageNumber="481">H. fasciaria</emphasis>
</taxonomicName>
,
<taxonomicName id="8CD247C22E2EC93BFB19B4F358B971A4" authorityName="Bellier" authorityYear="1861" box="[1276,1436,1962,1987]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="pinicolaria">
<emphasis id="79A6E0532E2EC93BFB19B4F358B971A4" box="[1276,1436,1962,1987]" italics="true" pageId="12" pageNumber="481">H. pinicolaria</emphasis>
</taxonomicName>
and in
<taxonomicName id="8CD247C22E2EC93BFF0CB4975CB67181" baseAuthorityName="Staudinger" baseAuthorityYear="1894" box="[233,403,1999,2022]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="compararia">
<emphasis id="79A6E0532E2EC93BFF0CB4975CB67181" box="[233,403,1999,2022]" italics="true" pageId="12" pageNumber="481">H. compararia</emphasis>
</taxonomicName>
). Genitalia are large in
<taxonomicName id="8CD247C22E2EC93BFD49B4975E6A7180" authorityName="Bellier" authorityYear="1861" box="[684,847,1998,2023]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="pinicolaria">
<emphasis id="79A6E0532E2EC93BFD49B4975E6A7180" box="[684,847,1998,2023]" italics="true" pageId="12" pageNumber="481">H. pinicolaria</emphasis>
</taxonomicName>
(genitalia considerably smaller in
<taxonomicName id="8CD247C22E2EC93BFB07B49758AE7181" baseAuthorityName="Staudinger" baseAuthorityYear="1894" box="[1250,1419,1999,2022]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="481" phylum="Arthropoda" rank="species" species="compararia">
<emphasis id="79A6E0532E2EC93BFB07B49758AE7181" box="[1250,1419,1999,2022]" italics="true" pageId="12" pageNumber="481">H. compararia</emphasis>
</taxonomicName>
). Shape and size of the lamella antevaginalis, and width and length of the posterior part of the corpus bursae are variable and should be treated with caution.
</paragraph>
</subSubSection>
<subSubSection id="03C86FCA2E2FC93AFF22B3875F557702" pageId="13" pageNumber="482" type="distribution">
<paragraph id="4B6D3C412E2FC93AFF22B3875F94769F" blockId="13.[151,1437,151,861]" box="[199,689,223,248]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B3875C45769F" bold="true" box="[199,352,223,248]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">Distribution.</emphasis>
Endemic to
<collectingCountry id="33C57CD12E2FC93AFE0BB3875F66769F" box="[494,579,223,248]" name="France" pageId="13" pageNumber="482">Corsica</collectingCountry>
(
<collectingCountry id="33C57CD12E2FC93AFDB6B3875F83769F" box="[595,678,223,248]" name="France" pageId="13" pageNumber="482">France</collectingCountry>
).
</paragraph>
<paragraph id="4B6D3C412E2FC93AFF22B25C5F557702" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B25C5C6C777A" bold="true" box="[199,329,260,285]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">Phenology.</emphasis>
According to
<bibRefCitation id="2F4341B02E2FC93AFE17B25C5F97777A" author="Robineau, R." box="[498,690,260,285]" pageId="13" pageNumber="482" refId="ref11855" refString="Robineau, R. (2007) Guide des papillons nocturnes de France. Delachaux et Niestl, Paris, 288 pp." type="book" year="2007">Robineau (2007)</bibRefCitation>
it is univoltine, but this is true probably only at higher altitudes (where it flies from June to early August), while on the coast the species has been collected from May to June and from October to November (
<bibRefCitation id="2F4341B02E2FC93AFE32B2145F467702" author="Rungs, C." box="[471,611,332,357]" pageId="13" pageNumber="482" pagination="221 - 229" refId="ref11878" refString="Rungs, C. (1982) Notes de lepidopterologie corse (II). Alexanor, XII (5), 221 - 229; (6), 265 - 271." type="journal article" year="1982">Rungs 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03C86FCA2E2FC93AFF22B2375ED67572" pageId="13" pageNumber="482" type="biology_ecology">
<paragraph id="4B6D3C412E2FC93AFF22B2375CEC7792" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B2375C0277EF" bold="true" box="[199,295,367,392]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">Biology.</emphasis>
Larva has been recorded to feed on needles of
<taxonomicName id="8CD247C22E2FC93AFCD9B2285EEB77EF" box="[828,974,367,392]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="482" phylum="Tracheophyta" rank="species" species="laricio">
<emphasis id="79A6E0532E2FC93AFCD9B2285EEB77EF" box="[828,974,367,392]" italics="true" pageId="13" pageNumber="482">Pinus laricio</emphasis>
</taxonomicName>
(
<bibRefCitation id="2F4341B02E2FC93AFC39B237594977EF" author="Bellier, C." box="[988,1132,367,392]" pageId="13" pageNumber="482" pagination="29 - 30" refId="ref10615" refString="Bellier, C. (1861) Description de trois lepidopteres nouveaux de l'ile de Corse. Annales de la Societ entomologique de France, 1, 29 - 30." type="journal article" year="1861">Bellier 1861</bibRefCitation>
,
<bibRefCitation id="2F4341B02E2FC93AFB9DB237580077EF" author="Robineau, R." box="[1144,1317,367,392]" pageId="13" pageNumber="482" refId="ref11855" refString="Robineau, R. (2007) Guide des papillons nocturnes de France. Delachaux et Niestl, Paris, 288 pp." type="book" year="2007">Robineau 2007</bibRefCitation>
) and on
<taxonomicName id="8CD247C22E2FC93AFA62B2295DD677CB" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="482" phylum="Tracheophyta" rank="species" species="pinaster">
<emphasis id="79A6E0532E2FC93AFA62B2295DD677CB" italics="true" pageId="13" pageNumber="482">P. pinaster</emphasis>
</taxonomicName>
(=
<emphasis id="79A6E0532E2FC93AFEFFB2CD5CA477CB" box="[282,385,405,428]" italics="true" pageId="13" pageNumber="482">maritima</emphasis>
) (
<bibRefCitation id="2F4341B02E2FC93AFE7CB2CC5F1077CA" author="Mabille, P." box="[409,565,404,429]" pageId="13" pageNumber="482" pagination="545 - 564" refId="ref11568" refString="Mabille, P. (1867) Lepidopteres de la Corse. I. re notice. Annales de la Societ entomologique de France, 4 ser., VI (1866), 4, 545 - 564." type="journal article" year="1867">Mabille 1867</bibRefCitation>
). In captivity it has been reared on
<taxonomicName id="8CD247C22E2FC93AFC5AB2CD580777CA" authority="(Reisser, 1929)" baseAuthorityName="Reisser" baseAuthorityYear="1929" box="[959,1314,404,429]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="13" pageNumber="482" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="79A6E0532E2FC93AFC5AB2CD594B77CA" box="[959,1134,404,429]" italics="true" pageId="13" pageNumber="482">Pinus sylvestris</emphasis>
(
<bibRefCitation id="2F4341B02E2FC93AFB98B2CC583F77CA" author="Reisser, H." box="[1149,1306,404,429]" pageId="13" pageNumber="482" pagination="79 - 81" refId="ref11813" refString="Reisser, H. (1929) Beschreibung der unbekannten ersten Stande einiger corsischer Geometriden. Zeitschrift des Osterreichischen Entomologischen Vereines, Wien, 14, 3 - 6, 31 - 33, 42 - 44, 79 - 81." type="journal article" year="1929">Reisser, 1929</bibRefCitation>
)
</taxonomicName>
. Larva is a twig or needle-mimic, green, with transverse yellowish stripes or reddish-brown with diamond-shaped patterns dorsally (
<bibRefCitation id="2F4341B02E2FC93AFF1AB2845C987792" author="Lepiforum" box="[255,445,476,501]" pageId="13" pageNumber="482" refId="ref11459" refString="Lepiforum, e. V. (2013) Bestimmung von Schmetterlingen (Lepidoptera) und ihren Praimaginalstadien. Available from: http: // www. lepiforum. de / (accessed 20 February 2013)" type="url" year="2013">Lepiforum 2013</bibRefCitation>
).
</paragraph>
<paragraph id="4B6D3C412E2FC93AFF22B2A75E10745A" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B2A75C0F747F" bold="true" box="[199,298,511,536]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">Habitat.</emphasis>
In pine forests and places with more scattered pine trees. It is found mostly in the mountains from 500 to 1500 metres, but it occurs also at sea level (
<bibRefCitation id="2F4341B02E2FC93AFD79B17C5E0C745A" author="Rungs, C." box="[668,809,548,573]" pageId="13" pageNumber="482" pagination="221 - 229" refId="ref11878" refString="Rungs, C. (1982) Notes de lepidopterologie corse (II). Alexanor, XII (5), 221 - 229; (6), 265 - 271." type="journal article" year="1982">Rungs 1982</bibRefCitation>
).
</paragraph>
<paragraph id="4B6D3C412E2FC93AFF22B11F5E5B74AA" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B11F5CA67407" bold="true" box="[199,387,583,608]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">Similar species.</emphasis>
All four species in Palaearctic
<taxonomicName id="8CD247C22E2FC93AFD11B11F5E977407" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[756,946,583,608]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="482" phylum="Arthropoda" rank="species" species="fasciaria">
<emphasis id="79A6E0532E2FC93AFD11B11F5E977407" box="[756,946,583,608]" italics="true" pageId="13" pageNumber="482">Hylaea fasciaria</emphasis>
</taxonomicName>
species group are similar. The diagnostic, external characters shown in
<figureCitation id="D3E920C42E2FC93AFE38B1345F4774E2" box="[477,610,620,645]" captionStart="FIGURES 69" captionStartId="5.[151,264,1100,1122]" captionTargetBox="[152,1435,201,1054]" captionTargetId="figure-391@5.[151,1435,201,1061]" captionTargetPageId="5" captionText="FIGURES 69. An overview of diagnostic characters in the Palaearctic Hylaea fasciaria species group. Figure 6. H. fasciaria (Linnaeus) (Denmark, 27.vi.1977). Figure 7. H. mediterranea, new species (Italy, Sicily, 5.vi.2005, paratype). Figure 8. H. pinicolaria (Bellier) (France, Corsica, 20.-21.vii.2004). Figure 9. H. compararia (Staudinger) (Algeria, 23.vi.1911)." figureDoi="http://doi.org/10.5281/zenodo.4909662" httpUri="https://zenodo.org/record/4909662/files/figure.png" pageId="13" pageNumber="482">Figures 69</figureCitation>
are somewhat tentative and should not be used in isolation, but should be combined with information on biology, collecting locality, male and female genitalia and DNA barcodes. An overview of diagnostic morphological features is given in
<tableCitation id="065009FA2E2FC93AFCC1B1EC5E5D74AA" box="[804,888,692,717]" captionStart="TABLE 1" captionStartId="9.[151,241,152,175]" captionTargetPageId="9" captionText="TABLE 1. An overview of diagnostic morphological features in the Palaearctic Hylaea fasciaria species group. See Figures 19 for the external characters, 1013 for the male genitalia and 1417 for the female genitalia. The shape and size of the lamella antevaginalis in the female genitalia is variable, and should be treated with caution." httpUri="http://table.plazi.org/id/1FAD6CC92E2BC93EFF72B3C059FC7688" pageId="13" pageNumber="482" tableUuid="1FAD6CC92E2BC93EFF72B3C059FC7688">Table 1</tableCitation>
.
</paragraph>
<paragraph id="4B6D3C412E2FC93AFF22B18F5ED67572" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B18F5C4F7497" bold="true" box="[199,362,727,752]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">Genetic data.</emphasis>
Genetically homogeneous (n=4), mean and maximum intraspecific variation 0.0%. Nearest species:
<taxonomicName id="8CD247C22E2FC93AFF13B1A45C977572" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[246,434,764,789]" class="Insecta" family="Geometridae" genus="Hylaea" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="482" phylum="Arthropoda" rank="species" species="fasciaria">
<emphasis id="79A6E0532E2FC93AFF13B1A45C977572" box="[246,434,764,789]" italics="true" pageId="13" pageNumber="482">Hylaea fasciaria</emphasis>
</taxonomicName>
(minimum pairwise distance 3.9%). See
<figureCitation id="D3E920C42E2FC93AFC65B1A45ECA7572" box="[896,1007,764,789]" captionStart="FIGURE 26" captionStartId="15.[151,249,1960,1982]" captionTargetBox="[194,1379,200,1915]" captionTargetId="figure-41@15.[187,1379,193,1919]" captionTargetPageId="15" captionText="FIGURE 26. Neighbour joining tree (Kimura 2-parameter distance model for COI-5P marker) for European Hylaea specimens. Terminals with specimen ID-number and species name from BOLD." figureDoi="http://doi.org/10.5281/zenodo.4909678" httpUri="https://zenodo.org/record/4909678/files/figure.png" pageId="13" pageNumber="482">Figure 26</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="03C86FCA2E2FC93AFF22B0475CAA753A" pageId="13" pageNumber="482" type="description">
<paragraph id="4B6D3C412E2FC93AFF22B0475CAA753A" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="482">
<emphasis id="79A6E0532E2FC93AFF22B0475C91755F" box="[199,436,799,824]" inLineHeading="true" pageId="13" pageNumber="482" reason="1">
<emphasis id="79A6E0532E2FC93AFF22B0475C1D755F" bold="true" box="[199,312,799,824]" pageId="13" pageNumber="482">Variation</emphasis>
(
<figureCitation id="D3E920C42E2FC93AFEADB0475C8D755F" box="[328,424,799,824]" captionStart="FIGURES 15" captionStartId="4.[151,264,1398,1420]" captionTargetBox="[183,1386,187,1369]" captionTargetId="figure-357@4.[172,1404,187,1386]" captionTargetPageId="4" captionText="FIGURES 15. Adults of the Palaearctic Hylaea fasciaria species group. 1a. Hylaea fasciaria (L.) male, 13.vi.1987, Volsk, Saratov Region, South East Russia (coll. Skou). 1b. H. fasciaria male, 6.vii.1972, Skanør, Falsterbo, Skåne, Sweden (coll Skou). 1c. H. fasciaria male, 1.viii.1972, Byerum, Öland, Sweden (coll. Skou). 1d. H. fasciaria male, 18.vii.1970, Dueodde, Bornholm, Denmark (coll. Skou). 1e. H. fasciaria flavella (Wehrli), type, original illustration from Wehrli (19391954, plate 24: g), Armenia, without date (coll. unknown). 1f. H. fasciaria female, 6.vii.1972, Skanør, Falsterbo, Skåne, Sweden (coll. Skou). 1g. H. fasciaria female, 29.vii.2000, Terre Rouge, Briancon, Dept. Hautes Alpes, France (coll. Skou). 1h. H. fasciaria cedricola (Wehrli) male, ex ovo, Bey Dağlari, Antalya, Turkey (coll. Müller). 1i. H. fasciaria cedricola female, ex ovo, Bey Dağlari, Antalya, Turkey (coll. Müller). 1j. H. fasciaria male, 2728.vii.1983, Vorarlberg, 1500 m, Austria (coll. Skou). 1k. H. fasciaria male, 10.vi.2010, Mount Parnassos, 1650 m, Greece (coll. Skou). 1l. H. fasciaria female, 2728.vii.1983, Vorarlberg, 1500 m, Austria (coll. Skou). 1m. H. fasciaria female, 28.vi.1981, Karpenision, Aetolia, Greece (coll. Skou). 2a. Hylaea mediterranea new species male, 910.x.2010, Pizzo della Rondine, Sicily, Italy (coll. Skou), holotype. 2b H. mediterranea male, 910.x.2010, Pizzo della Rondine, Sicily, Italy (coll. Skou), paratype. 2c. H. mediterranea female, 910.x.2010, Pizzo della Rondine, Sicily, Italy (coll. Skou), paratype. 2d. H. mediterranea female, 5.vi.2005, Castelbuono, Sicily, Italy (coll. Skou), paratype. 3a. Hylaea pinicolaria (Bellier) male, 2021.vii.2004, Camping Monte Cinto, Haut-Asco, Corsica, France (coll. Skou). 3b. H. pinicolaria male, 2021.vii.2004, Camping Monte Cinto, Haut-Asco, Corsica, France (coll. Skou). 3c. H. pinicolaria male, 6.vii.1951, La Foce, Vizzavona, Corsica, France (coll. Herbulot, coll. ZSM). 3d. H. pinicolaria female, 23.vi.1994, Haut Asco, Corsica, France (coll. Skou). 4a. Hylaea compararia (Staud.) male, 23.vi.1911, Blida, Glacieres, Algeria (coll. BMNH). 4b. Hylaea compararia female, 13.ix.1911, Glacieres, Blida, Algeria (coll. BMNH). 4c. Hylaea compararia female, 15.ix.1911, Glacieres, Blida, Algeria (coll. BMNH). 5a. Hemithea squalidaria O. G. Costa, original illustration from O. G. Costa, [1848]. Illustration is reproduced under a licence from the Picture Library of the Natural History Museum, London, UK." figureDoi="http://doi.org/10.5281/zenodo.4909660" httpUri="https://zenodo.org/record/4909660/files/figure.png" pageId="13" pageNumber="482">Figure 3</figureCitation>
).
</emphasis>
Very little. Ground colour varies from light green to light yellowish-green. Only green specimens are known.
</paragraph>
</subSubSection>
</treatment>
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