Revision of the Hylaea fasciaria (Linnaeus, 1758) species group in the western Palaearctic (Lepidoptera: Geometridae, Ennominae)
Author
Sihvonen, Pasi
Author
Skou, Peder
Aamosen 1, DK 5762 Vester Skerninge, Denmark. E-mail: info @ apollobooks. dk
Author
Flamigni, Claudio
Via delle Belle Arti 21, I- 40126 Bologna, Italy. E-mail: claudio. flamigni @ alice. it
Author
Fiumi, Gabriele
Via Decio Raggi 167, I- 47121 Forlí, Italy. E-mail: gabfium @ tiscali. it
Author
Hausmann, Axel
Bavarian State Collection of Zoology (ZSM), Münchhausenstr. 21, 81247 Munich, Germany - Bavarian Natural History Collections (SNSB), Menzinger Str. 71, 80638 Munich, Germany. E-mail: axel. hausmann @ zsm. mwn. de
text
Zootaxa
2014
2014-02-27
3768
4
469
486
journal article
5898
10.11646/zootaxa.3768.4.5
985f8c4c-c9ad-482b-8e0a-c25f6580b9bd
1175-5326
4909658
DBDE3685-85AC-4739-95DB-F8BA16117AAB
Hylaea pinicolaria
(
Bellier, 1861
)
pinicolaria
Bellier, 1861
, Annls Soc. ent. Fr. (4) 1 (1): 29, pl. 2, fig. 12, (
Ellopia
).
Syntype
(s), Corsica (mountains of). Bellier's collection was apparently included in the
C. Oberthür
collection, which is currently housed in the Natural History Museum, London,
UK
. Despite searches, we have not located the type.
Description.
External characters and pregenital abdomen
(
Figures 3
,
8
) (diagnostic characters underlined): Wingspan male
32–34 mm
, female
39–42 mm
. Wings light green, medial lines white. Medial line curved before costa, basal part parallel with costa. Postmedial line rather straight, only weakly curved, angled before it reaches costa well before apex and evenly curved outwards on inner margin. Medial area concolorous with rest of wing. Terminal line and fringes concolorous with wings, forewing apex dark red. Hindwing postmedial line distinct, curved. Discal spots absent. Wings below as above, but paler. Frons red-brown, thorax and abdomen concolorus with wings. Area between antennae (vertex) white. Antennae white dorsally, male antennae bipectinate, female antennae fasciculate. Hindleg tibia of both sexes with 2+2 spurs. Tympanal organs medium-sized. Sternites and tergites 3–8 of both sexes undifferentiated.
Male genitalia
(
Figure 12
): Generally as in
H. fasciaria
(Linnaeus)
and
H. mediterranea
. Aedeagus with weakly curved additional arm, apex not expanded in
H. pinicolaria
(additional arm straight in
H. fasciaria
, apex expanded in
H. compararia
). Base of vesica with straight row of microcornuti in
H. pinicolaria
(vesica with angled row of microcornuti, reaching aedeagus apex in
H. compararia
).
Female genitalia
(
Figure 16
): Generally as in
H. fasciaria
and
H. mediterranea
, but with following quantitative difference: signum large in
H. mediterranea
(signum absent or minute in
H. fasciaria
,
H. pinicolaria
and in
H. compararia
). Genitalia are large in
H. pinicolaria
(genitalia considerably smaller in
H. compararia
). Shape and size of the lamella antevaginalis, and width and length of the posterior part of the corpus bursae are variable and should be treated with caution.
Distribution.
Endemic to
Corsica
(
France
).
Phenology.
According to
Robineau (2007)
it is univoltine, but this is true probably only at higher altitudes (where it flies from June to early August), while on the coast the species has been collected from May to June and from October to November (
Rungs 1982
).
Biology.
Larva has been recorded to feed on needles of
Pinus laricio
(
Bellier 1861
,
Robineau 2007
) and on
P. pinaster
(=
maritima
) (
Mabille 1867
). In captivity it has been reared on
Pinus sylvestris
(
Reisser, 1929
)
. Larva is a twig or needle-mimic, green, with transverse yellowish stripes or reddish-brown with diamond-shaped patterns dorsally (
Lepiforum 2013
).
Habitat.
In pine forests and places with more scattered pine trees. It is found mostly in the mountains from 500 to 1500 metres, but it occurs also at sea level (
Rungs 1982
).
Similar species.
All four species in Palaearctic
Hylaea fasciaria
species group are similar. The diagnostic, external characters shown in
Figures 6–9
are somewhat tentative and should not be used in isolation, but should be combined with information on biology, collecting locality, male and female genitalia and DNA barcodes. An overview of diagnostic morphological features is given in
Table 1
.
Genetic data.
Genetically homogeneous (n=4), mean and maximum intraspecific variation 0.0%. Nearest species:
Hylaea fasciaria
(minimum pairwise distance 3.9%). See
Figure 26
.
Variation
(
Figure 3
).
Very little. Ground colour varies from light green to light yellowish-green. Only green specimens are known.