treatments-xml/data/CF/28/93/CF28935B6719FF90FF39FCC8FDEAFAD4.xml

<document id="6C5D9F78E0564DF596B68741B24895DD" ID-CLB-Dataset="298635" ID-DOI="10.11646/zootaxa.5468.1.8" ID-GBIF-Dataset="fbbf012f-ea10-47e4-8a26-d501081767eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="11615151" ID-ZooBank="FFC4DF39-964D-4967-8F7F-EA83CB4A827D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1718185085663" checkinUser="plazi" docAuthor="Salvador, Jewel Anne G., Nuñeza, Olga Macas, Robillard, Tony &amp; Tan, Ming Kai" docDate="2024" docId="CF28935B6719FF90FF39FCC8FDEAFAD4" docLanguage="en" docName="zootaxa.5468.1.8.pdf" docOrigin="Zootaxa 5468 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5468.1.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ducetia laniae Salvador &amp; Nuñeza &amp; Robillard &amp; Tan 2024, sp. nov." docType="treatment" docVersion="6" lastPageNumber="185" masterDocId="3311EB23671BFF96FFAEFFD7FFFFFFB5" masterDocTitle="A new species of Ducetia (Tettigoniidae, Phaneropterinae) from Mindanao, Philippines based on bioacoustics, stridulatory file morphology and genetic data" masterLastPageNumber="190" masterPageNumber="179" pageNumber="181" updateTime="1718732787391" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4EF8364AB43EFB86D759756D294C83B1">A new species of Ducetia (Tettigoniidae, Phaneropterinae) from Mindanao, Philippines based on bioacoustics, stridulatory file morphology and genetic data</mods:title>
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<mods:namePart id="E35531710E42116DEEB533DAB9903AB6">Salvador, Jewel Anne G.</mods:namePart>
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<mods:namePart id="878903B855A89102CC8E52A18F7491D7">Nuñeza, Olga Macas</mods:namePart>
<mods:affiliation id="09652E5D2C6E2D3A411B0176E37871FF">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines.</mods:affiliation>
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<mods:namePart id="B5F90634BEEFE9C3B073417F269A1075">Robillard, Tony</mods:namePart>
<mods:affiliation id="4637DB6E9EA2C2A9694457080EEB5823">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<emphasis id="75F5FE5F6719FF94FF39FCC8FEB7FC8C" bold="true" box="[151,328,798,825]" italics="true" pageId="2" pageNumber="181">Ducetia laniae</emphasis>
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(
<figureCitation id="DFBA3EC86719FF94FF31FC94FEFDFCE8" box="[159,258,835,861]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1060,1084]" captionStartId-1="3.[151,250,2011,2035]" captionStartId-2="4.[151,250,1041,1065]" captionStartId-3="5.[151,250,1736,1760]" captionStartId-4="6.[151,250,705,729]" captionStartId-5="7.[151,250,1943,1967]" captionStartId-6="8.[151,250,1541,1565]" captionTargetBox-0="[151,1436,181,1036]" captionTargetBox-1="[158,1430,1140,1987]" captionTargetBox-2="[151,1436,181,1017]" captionTargetBox-3="[151,1436,181,1712]" captionTargetBox-4="[161,1426,191,671]" captionTargetBox-5="[177,1410,181,1919]" captionTargetBox-6="[156,1429,191,1507]" captionTargetId-0="figure-18@3.[151,1436,181,1036]" captionTargetId-1="figure-36@3.[158,1430,1140,1987]" captionTargetId-2="figure-21@4.[151,1436,181,1017]" captionTargetId-3="figure-18@5.[151,1436,181,1712]" captionTargetId-4="figure-21@6.[151,1436,181,681]" captionTargetId-5="figure-18@7.[177,1410,181,1919]" captionTargetId-6="figure-21@8.[151,1436,181,1517]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURE 1. Ducetia laniae sp. nov. ♂ in its natural environment in Agusan del Sur." captionText-1="FIGURE 2. Ducetia laniae sp. nov. ♂ in its natural environment in Agusan del Sur." captionText-2="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." captionText-3="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." captionText-4="FIGURE 5. Tooth distribution on the stridulatory file of Ducetia laniae sp. nov. ♂ based on tooth length (A) and inter-tooth distance (B)." captionText-5="FIGURE 6. Ducetia laniae sp. nov. ♀: habitus in lateral (A) and dorsal (B) views; head, pronotum and anterior part of tegmina in dorsal view (C); abdominal apex in lateral (D) and ventral (E) views. Scale bars: 5 mm (A, B); 2 mm (C–E)." captionText-6="FIGURE 7. Ducetia laniae sp. nov. ♂ calling song: oscillogram (A) and spectrogram (B) showing two echeme-sequences; oscillograms showing four echemes (C) and one echeme (D); power spectra of an echeme (E) where the gray line represents the power [dB] (axis name and scale in gray) and the black solid peak represents the relative amplitude (axis name in black)." figureDoi-0="http://doi.org/10.5281/zenodo.11615155" figureDoi-1="http://doi.org/10.5281/zenodo.11615159" figureDoi-2="http://doi.org/10.5281/zenodo.11615163" figureDoi-3="http://doi.org/10.5281/zenodo.11615165" figureDoi-4="http://doi.org/10.5281/zenodo.11615169" figureDoi-5="http://doi.org/10.5281/zenodo.11615173" figureDoi-6="http://doi.org/10.5281/zenodo.11615177" httpUri-0="https://zenodo.org/record/11615155/files/figure.png" httpUri-1="https://zenodo.org/record/11615159/files/figure.png" httpUri-2="https://zenodo.org/record/11615163/files/figure.png" httpUri-3="https://zenodo.org/record/11615165/files/figure.png" httpUri-4="https://zenodo.org/record/11615169/files/figure.png" httpUri-5="https://zenodo.org/record/11615173/files/figure.png" httpUri-6="https://zenodo.org/record/11615177/files/figure.png" pageId="2" pageNumber="181">Figs 1–7</figureCitation>
)
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:
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• 
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, 19h15; coll. 
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&amp; 
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[molecular TRORT142-23 BC_JAS_047, GenBank Accession number 
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</date>
, 22h22; coll. 
<collectorName id="EA74479B6719FF94FB95FBCCFB39FB80" box="[1083,1222,1051,1077]" pageId="2" pageNumber="181">J. Amarante</collectorName>
, J.A. 
<collectorName id="EA74479B6719FF94FAA4FBCCFA67FB80" box="[1290,1432,1051,1077]" pageId="2" pageNumber="181">Salvador, M</collectorName>
. 
<collectorName id="EA74479B6719FF94FF39FBE8FEE0FBEC" box="[151,287,1087,1113]" pageId="2" pageNumber="181">Salvador, B</collectorName>
. 
<collectorName id="EA74479B6719FF94FE84FBE8FE68FBEC" box="[298,407,1087,1113]" pageId="2" pageNumber="181">Kiseo, M</collectorName>
. 
<collectorName id="EA74479B6719FF94FE0DFBE8FDF8FBEC" box="[419,519,1087,1113]" pageId="2" pageNumber="181">Kiseo, E</collectorName>
. 
<collectorName id="EA74479B6719FF94FDBDFBE8FD54FBEC" box="[531,683,1087,1113]" pageId="2" pageNumber="181">Bancairen, D</collectorName>
. 
<collectorName id="EA74479B6719FF94FD19FBE8FCD9FBEC" box="[695,806,1087,1113]" pageId="2" pageNumber="181">Lastua, A</collectorName>
. 
<collectorName id="EA74479B6719FF94FC9FFBE8FC20FBEC" box="[817,991,1087,1113]" pageId="2" pageNumber="181">Anobong, M.A</collectorName>
. 
<collectorName id="EA74479B6719FF94FC44FBE8FB76FBEC" box="[1002,1161,1087,1113]" pageId="2" pageNumber="181">Anobong, A.J</collectorName>
. 
<collectorName id="EA74479B6719FF94FB3BFBE8FAEEFBEC" box="[1173,1297,1087,1113]" pageId="2" pageNumber="181">Zamora, R</collectorName>
. 
<collectorName id="EA74479B6719FF94FAB2FBE8FA67FBEC" box="[1308,1432,1087,1113]" pageId="2" pageNumber="181">Zamora, A</collectorName>
. 
<collectorName id="EA74479B6719FF94FF39FBB4FE9AFBC8" box="[151,357,1123,1149]" pageId="2" pageNumber="181">Ruiz, M.A. Kiseo</collectorName>
; 64JAS1211 [
<specimenCode id="17278A366719FF94FDADFBB4FD4FFBC8" box="[515,688,1123,1149]" collectionCode="TRORT" pageId="2" pageNumber="181">TRORT146-23</specimenCode>
BC_JAS_051, GenBank Accession number 
<accessionNumber id="58D2BFAE6719FF94FB00FBB5FADAFBC8" box="[1198,1317,1122,1149]" pageId="2" pageNumber="181">PP625720</accessionNumber>
] (
<specimenCode id="17278A366719FF94FA95FBB4FEA4FB14" pageId="2" pageNumber="181">MNHN-EO-ENSIF11337</specimenCode>
) •
</materialsCitation>
<materialsCitation id="F7E928106719FF94FED5FB50FBFBFB70" ID-GBIF-Occurrence="4891618303" collectingDate="2022-06-01" collectingDateMax="2022-08-16" collectingDateMin="2022-06-01" collectionCode="R" collectorName="J. A. Salvador &amp; M. Salvador &amp; J. Noval" country="Philippines" elevation="20" latitude="8.712075" location="Bayugan City, Brgy" longLatPrecision="1" longitude="125.77753" municipality="Mindanao Island" pageId="2" pageNumber="181" specimenCode="MNHN-EO-ENSIF11338" specimenCount="51" specimenCount-female="1" specimenCount-male="1" stateProvince="Agusan del Sur province" typeStatus="paratype">
<specimenCount id="5187E9C46719FF94FED5FB50FE60FB14" box="[379,415,1159,1185]" count="1" pageId="2" pageNumber="181" type="female">1♀</specimenCount>
; 
<location id="425E74966719FF94FE00FB50FD70FB14" LSID="urn:lsid:plazi:treatment:CF28935B6719FF90FF39FCC8FDEAFAD4:425E74966719FF94FE00FB50FD70FB14" box="[430,655,1159,1185]" country="Philippines" latitude="8.712075" longLatPrecision="1" longitude="125.77753" municipality="Mindanao Island" name="Bayugan City, Brgy" pageId="2" pageNumber="181" stateProvince="Agusan del Sur province">Bayugan City, Brgy</location>
. Fili; 
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, 
<geoCoordinate id="22B5448A6719FF94FCF3FB51FC09FB14" box="[861,1014,1158,1185]" degrees="125.777530" direction="east" orientation="longitude" pageId="2" pageNumber="181" precision="1" value="125.77753">E125.777530</geoCoordinate>
, 
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<quantity id="80798FA86719FF94FBAFFB51FBBDFB14" box="[1025,1090,1158,1185]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="181" unit="m" value="50.0">50 m</quantity>
.
</specimenCount>
a.s.l; 
<date id="333F048D6719FF94FBD0FB50FAF5FB14" box="[1150,1290,1159,1185]" pageId="2" pageNumber="181" value="2022-06-01">
<collectingDate id="237BFD656719FF94FBD0FB50FAF5FB14" box="[1150,1290,1159,1185]" pageId="2" pageNumber="181" value="2022-06-01">1 June 2022</collectingDate>
</date>
, 20h37; coll. 
<collectorName id="EA74479B6719FF94FF39FB7CFECFFB70" box="[151,304,1195,1221]" pageId="2" pageNumber="181">J.A. Salvador</collectorName>
, 
<collectorName id="EA74479B6719FF94FE92FB7CFE38FB70" box="[316,455,1195,1221]" pageId="2" pageNumber="181">M. Salvador</collectorName>
&amp; 
<collectorName id="EA74479B6719FF94FE44FB7CFDB4FB70" box="[490,587,1195,1221]" pageId="2" pageNumber="181">J. Noval</collectorName>
; 71JAS460 (
<specimenCode id="17278A366719FF94FD77FB7CFC02FB70" box="[729,1021,1195,1221]" pageId="2" pageNumber="181">MNHN-EO-ENSIF11338</specimenCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C66719FF94FF69FB18FB5FFB5C" box="[199,1184,1231,1257]" pageId="2" pageNumber="181" type="etymology">
<paragraph id="473E224D6719FF94FF69FB18FB5FFB5C" blockId="2.[151,1437,906,1977]" box="[199,1184,1231,1257]" pageId="2" pageNumber="181">
<emphasis id="75F5FE5F6719FF94FF69FB18FEB1FB5C" bold="true" box="[199,334,1231,1257]" pageId="2" pageNumber="181">Etymology.</emphasis>
This species is named after Mellany “lani” 
<collectingCountry id="3F9662DD6719FF94FC97FB18FC61FB5C" box="[825,926,1231,1257]" name="El Salvador" pageId="2" pageNumber="181">Salvador</collectingCountry>
, mother of first author.
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C66719FF94FF69FB25FC35F9BC" pageId="2" pageNumber="181" type="diagnosis">
<paragraph id="473E224D6719FF94FF69FB25FAD0FA74" blockId="2.[151,1437,906,1977]" pageId="2" pageNumber="181">
<emphasis id="75F5FE5F6719FF94FF69FB25FEBDFAB9" bold="true" box="[199,322,1266,1292]" pageId="2" pageNumber="181">Diagnosis.</emphasis>
This new species is typical of the 
<taxonomicName id="808159CE6719FF94FD7AFB24FC67FAB9" baseAuthorityName="Thunberg" baseAuthorityYear="1815" box="[724,920,1267,1292]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="75F5FE5F6719FF94FD7AFB24FC67FAB9" box="[724,920,1267,1292]" italics="true" pageId="2" pageNumber="181">Ducetia japonica</emphasis>
</taxonomicName>
species group by the male cerci having long interno-ventral ridge and the tegmen having 4–6 closely parallel posterior branches that do not bifurcate. 
<taxonomicName id="808159CE6719FF94FAEAFAC0FF22FAE0" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F6719FF94FAEAFAC0FF22FAE0" italics="true" pageId="2" pageNumber="181">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F6719FF94FF49FAECFEBDFAE0" bold="true" box="[231,322,1339,1365]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="6EC643246719FF94FF49FAECFEBDFAE0" box="[231,322,1339,1365]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other species by the absence of distinctly larger teeth in the anal part (similar to 
<taxonomicName id="808159CE6719FF94FA2FFAECFF03FACC" authorityName="Brunner von Wattenwyl" authorityYear="1878" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="75F5FE5F6719FF94FA2FFAECFF03FACC" italics="true" pageId="2" pageNumber="181">D. adspersa</emphasis>
</taxonomicName>
and 
<taxonomicName id="808159CE6719FF94FE98FA88FC8AFACC" authority="(Brunner von Wattenwyl, 1891))" authorityName="" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1891" box="[310,885,1375,1401]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="75F5FE5F6719FF94FE98FA88FE08FACD" box="[310,503,1375,1400]" italics="true" pageId="2" pageNumber="181">Ducetia javanica</emphasis>
(Brunner von Wattenwyl, 1891))
</taxonomicName>
, and having distinctly more teeth, with a row of closely packed 97 teeth (see 
<bibRefCitation id="23105FBC6719FF94FE73FA54FD50FA28" author="Heller, K. - G. &amp; Ingrisch, S. &amp; Liu, C. &amp; Shi, F. - M. &amp; Hemp, C. &amp; Warchalowska-Sliwa, E. &amp; Rentz, D. C. F." box="[477,687,1411,1437]" pageId="2" pageNumber="181" pagination="286 - 307" refId="ref4945" refString="Heller, K. - G., Ingrisch, S., Liu, C., Shi, F. - M., Hemp, C., Warchalowska-Sliwa, E. &amp; Rentz, D. C. F. (2017) Complex songs and cryptic ethospecies: the case of the Ducetia japonica group (Orthoptera: Tettigonioidea: Phaneropteridae: Phaneropterinae). Zoological Journal of the Linnean Society, 181 (2), 286 - 307. https: // doi. org / 10.1093 / zoolinnean / zlw 019" type="journal article" year="2017">
Heller 
<emphasis id="75F5FE5F6719FF94FD85FA54FD9BFA28" box="[555,612,1411,1437]" italics="true" pageId="2" pageNumber="181">et al.</emphasis>
[2017
</bibRefCitation>
, Table 4]). Likewise in 
<taxonomicName id="808159CE6719FF94FC13FA54FBBAFA28" authorityName="Brunner von Wattenwyl" authorityYear="1878" box="[957,1093,1411,1437]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="adspersa">
<emphasis id="75F5FE5F6719FF94FC13FA54FBBAFA28" box="[957,1093,1411,1437]" italics="true" pageId="2" pageNumber="181">D. adspersa</emphasis>
</taxonomicName>
from Luzon (
<collectingCountry id="3F9662DD6719FF94FB4CFA54FA99FA28" box="[1250,1382,1411,1437]" name="Philippines" pageId="2" pageNumber="181">Philippines</collectingCountry>
), the number of teeth on the stridulatory file is notably lower, with fewer than 30 teeth, than 
<taxonomicName id="808159CE6719FF94FBC9FA70FB30FA74" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[1127,1231,1447,1473]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F6719FF94FBC9FA70FB30FA74" box="[1127,1231,1447,1473]" italics="true" pageId="2" pageNumber="181">D. laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F6719FF94FB78FA70FAD0FA74" bold="true" box="[1238,1327,1447,1473]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="6EC643246719FF94FB78FA70FAD0FA74" box="[1238,1327,1447,1473]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="473E224D6719FF94FF69FA1CFC35F9BC" blockId="2.[151,1437,906,1977]" pageId="2" pageNumber="181">
<taxonomicName id="808159CE6719FF94FF69FA1CFE91FA50" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[199,366,1483,1509]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F6719FF94FF69FA1CFE91FA50" box="[199,366,1483,1509]" italics="true" pageId="2" pageNumber="181">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F6719FF94FED9FA1CFE2DFA50" bold="true" box="[375,466,1483,1509]" pageId="2" pageNumber="181">
<taxonomicNameLabel id="6EC643246719FF94FED9FA1CFE2DFA50" box="[375,466,1483,1509]" pageId="2" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs by the calling song, which is relatively simple, consisting of an echeme-sequence made up of a closely-spaced repetition of echemes (see below).
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C66719FF94FF69F9C4FE29F9E4" pageId="2" pageNumber="181" type="discussion">
<paragraph id="473E224D6719FF94FF69F9C4FE29F9E4" blockId="2.[151,1437,906,1977]" pageId="2" pageNumber="181">
<emphasis id="75F5FE5F6719FF94FF69F9C4FEC4F998" bold="true" box="[199,315,1555,1581]" pageId="2" pageNumber="181">Remarks.</emphasis>
This new species represents the second species of 
<taxonomicName id="808159CE6719FF94FC2CF9C4FC24F999" authorityName="Stal" authorityYear="1874" box="[898,987,1555,1580]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="75F5FE5F6719FF94FC2CF9C4FC24F999" box="[898,987,1555,1580]" italics="true" pageId="2" pageNumber="181">Ducetia</emphasis>
</taxonomicName>
known to occur in the 
<collectingCountry id="3F9662DD6719FF94FB48F9C4FA98F998" box="[1254,1383,1555,1581]" name="Philippines" pageId="2" pageNumber="181">Philippines</collectingCountry>
, and the first in Mindanao Island.
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C66719FF90FF69F98DFD30FCDC" lastPageId="6" lastPageNumber="185" pageId="2" pageNumber="181" type="description">
<paragraph id="473E224D6719FF92FF69F98DFB0AFA6C" blockId="2.[151,1437,906,1977]" lastBlockId="4.[151,1437,1147,1966]" lastPageId="4" lastPageNumber="183" pageId="2" pageNumber="181">
<emphasis id="75F5FE5F6719FF94FF69F98DFEA6F9C1" bold="true" box="[199,345,1626,1652]" pageId="2" pageNumber="181">Description.</emphasis>
Habitus typical of 
<taxonomicName id="808159CE6719FF94FD9EF98CFD0EF9C1" baseAuthorityName="Thunberg" baseAuthorityYear="1815" box="[560,753,1627,1652]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="75F5FE5F6719FF94FD9EF98CFD0EF9C1" box="[560,753,1627,1652]" italics="true" pageId="2" pageNumber="181">Ducetia japonica</emphasis>
</taxonomicName>
species group; generally pale green with head and pronotum on dorsum yellow green, tegmina with stridulatory area and anal margins and legs (sometimes) brown when alive (
<figureCitation id="DFBA3EC86719FF94FF70F974FEBBF908" box="[222,324,1699,1725]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1060,1084]" captionStartId-1="3.[151,250,2011,2035]" captionStartId-2="4.[151,250,1041,1065]" captionTargetBox-0="[151,1436,181,1036]" captionTargetBox-1="[158,1430,1140,1987]" captionTargetBox-2="[151,1436,181,1017]" captionTargetId-0="figure-18@3.[151,1436,181,1036]" captionTargetId-1="figure-36@3.[158,1430,1140,1987]" captionTargetId-2="figure-21@4.[151,1436,181,1017]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Ducetia laniae sp. nov. ♂ in its natural environment in Agusan del Sur." captionText-1="FIGURE 2. Ducetia laniae sp. nov. ♂ in its natural environment in Agusan del Sur." captionText-2="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11615155" figureDoi-1="http://doi.org/10.5281/zenodo.11615159" figureDoi-2="http://doi.org/10.5281/zenodo.11615163" httpUri-0="https://zenodo.org/record/11615155/files/figure.png" httpUri-1="https://zenodo.org/record/11615159/files/figure.png" httpUri-2="https://zenodo.org/record/11615163/files/figure.png" pageId="2" pageNumber="181">Figs 1–3</figureCitation>
). Frontal rostrum conical, stout, with subacute apex; upper tubercle having median sulcus distinct (
<figureCitation id="DFBA3EC86719FF94FF31F910FF02F954" box="[159,253,1735,1761]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4A</figureCitation>
). Eyes oval and globular; in anterior view taller than wide (
<figureCitation id="DFBA3EC86719FF94FC1DF910FBF1F954" box="[947,1038,1735,1761]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4B</figureCitation>
); in dorsal view faintly protruding anteriorly (
<figureCitation id="DFBA3EC86719FF94FEBBF93CFE8DF8B0" box="[277,370,1771,1797]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4A</figureCitation>
); in lateral view obliquely elongated (
<figureCitation id="DFBA3EC86719FF94FC89F93CFC7DF8B0" box="[807,898,1771,1797]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4C</figureCitation>
). Mouthparts generally light colored (
<figureCitation id="DFBA3EC86719FF94FA9BF93CFA70F8B0" box="[1333,1423,1771,1797]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4B</figureCitation>
). Apical segment of maxillary palpi longest, slender, slightly curved, apex obtuse; fourth distinctly shorter than third segment in length (
<figureCitation id="DFBA3EC86719FF94FE17F8E4FDE9F8F8" box="[441,534,1843,1869]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4C</figureCitation>
). Pronotal disc not covering stridulatory area; 1.3–1.8 times as long as wide; anterior margin straight; slightly widening apically; median sulcus not distinct; lateral keel distinct near anterior and posterior margins; posterior margin broadly rounded 1.4–1.8 times as long as anterior margin (
<figureCitation id="DFBA3EC86719FF94FA9EF8ACFA6FF820" box="[1328,1424,1915,1941]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="2" pageNumber="181">Fig. 4A</figureCitation>
). Pronotal lateral lobe rectangular, 1.3–1.5 times as long as tall; with ventral margin fairly elongated and straight; anterior margin slightly concave; posterior margin broadly rounded (
<figureCitation id="DFBA3EC8671FFF92FC3EFBACFC17FB23" box="[912,1000,1147,1174]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4C</figureCitation>
). Thoracic auditory spiracle (= thoracic foramen) large and bean-shaped; partly or fully covered by pronotal lateral lobe (
<figureCitation id="DFBA3EC8671FFF92FBB2FB48FB8BFB0F" box="[1052,1140,1183,1210]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4C</figureCitation>
). Tegmen fully developed, well surpassing hind knees; infumated; rather narrow, with oblique apex; venation typical of 
<taxonomicName id="808159CE671FFF92FB26FB13FAB8FB68" baseAuthorityName="Thunberg" baseAuthorityYear="1815" box="[1160,1351,1220,1245]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="75F5FE5F671FFF92FB26FB13FAB8FB68" box="[1160,1351,1220,1245]" italics="true" pageId="4" pageNumber="183">Ducetia japonica</emphasis>
</taxonomicName>
species group: with 4–6 closely parallel posterior branches (
<figureCitation id="DFBA3EC8671FFF92FD45FB30FCB9FAB7" box="[747,838,1255,1282]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[151,1436,181,1017]" captionTargetId="figure-21@4.[151,1436,181,1017]" captionTargetPageId="4" captionText="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11615163" httpUri="https://zenodo.org/record/11615163/files/figure.png" pageId="4" pageNumber="183">Fig. 3A</figureCitation>
). Hind wing surpassing tegminal apices, reaching or surpassing middle of hind tibia; exposed part infumated (
<figureCitation id="DFBA3EC8671FFF92FCBAFADCFCA7FA93" box="[788,856,1291,1318]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[151,1436,181,1017]" captionTargetId="figure-21@4.[151,1436,181,1017]" captionTargetPageId="4" captionText="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11615163" httpUri="https://zenodo.org/record/11615163/files/figure.png" pageId="4" pageNumber="183">Fig. 3</figureCitation>
). Legs generally long and slender (
<figureCitation id="DFBA3EC8671FFF92FB4FFADCFADAFA93" box="[1249,1317,1291,1318]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[151,1436,181,1017]" captionTargetId="figure-21@4.[151,1436,181,1017]" captionTargetPageId="4" captionText="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11615163" httpUri="https://zenodo.org/record/11615163/files/figure.png" pageId="4" pageNumber="183">Fig. 3</figureCitation>
). Fore and middle legs and hind tibia sometimes brown (
<figureCitation id="DFBA3EC8671FFF92FD38FAF8FD08FAFF" box="[662,759,1327,1354]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1060,1084]" captionStartId-1="3.[151,250,2011,2035]" captionStartId-2="4.[151,250,1041,1065]" captionTargetBox-0="[151,1436,181,1036]" captionTargetBox-1="[158,1430,1140,1987]" captionTargetBox-2="[151,1436,181,1017]" captionTargetId-0="figure-18@3.[151,1436,181,1036]" captionTargetId-1="figure-36@3.[158,1430,1140,1987]" captionTargetId-2="figure-21@4.[151,1436,181,1017]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Ducetia laniae sp. nov. ♂ in its natural environment in Agusan del Sur." captionText-1="FIGURE 2. Ducetia laniae sp. nov. ♂ in its natural environment in Agusan del Sur." captionText-2="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11615155" figureDoi-1="http://doi.org/10.5281/zenodo.11615159" figureDoi-2="http://doi.org/10.5281/zenodo.11615163" httpUri-0="https://zenodo.org/record/11615155/files/figure.png" httpUri-1="https://zenodo.org/record/11615159/files/figure.png" httpUri-2="https://zenodo.org/record/11615163/files/figure.png" pageId="4" pageNumber="183">Fig. 1–3</figureCitation>
). Fore femur with both internal and external ventral margins having two to three very small spines, usually at apical half; middle femur with ventral margin having seven or eight small spines; hind femur with ventral margin having circa four spines widely spaced apart. Inner and outer knees of hind femora with rounded lobes each having a small apical spine. Fore tibia with both tympanum oval and open (
<figureCitation id="DFBA3EC8671FFF92FF31FA68FF06FA6F" box="[159,249,1471,1498]" captionStart="FIGURE 3" captionStartId="4.[151,250,1041,1065]" captionTargetBox="[151,1436,181,1017]" captionTargetId="figure-21@4.[151,1436,181,1017]" captionTargetPageId="4" captionText="FIGURE 3. Ducetia laniae sp. nov. ♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11615163" httpUri="https://zenodo.org/record/11615163/files/figure.png" pageId="4" pageNumber="183">Fig. 3A</figureCitation>
). Tibia ventrally and dorsally typically with numerous outer and inner dark-colored spines.
</paragraph>
<caption id="13FE72C56718FF95FF39FBF3FC13FB89" ID-DOI="http://doi.org/10.5281/zenodo.11615155" ID-Zenodo-Dep="11615155" box="[151,1004,1060,1084]" httpUri="https://zenodo.org/record/11615155/files/figure.png" pageId="3" pageNumber="182" startId="3.[151,250,1060,1084]" targetBox="[151,1436,181,1036]" targetPageId="3" targetType="figure">
<paragraph id="473E224D6718FF95FF39FBF3FC13FB89" blockId="3.[151,1004,1060,1084]" box="[151,1004,1060,1084]" pageId="3" pageNumber="182">
<emphasis id="75F5FE5F6718FF95FF39FBF3FEECFB89" bold="true" box="[151,275,1060,1084]" pageId="3" pageNumber="182">FIGURE 1.</emphasis>
<taxonomicName id="808159CE6718FF95FEB7FBF2FE50FB89" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[281,431,1060,1084]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F6718FF95FEB7FBF2FE50FB89" box="[281,431,1060,1084]" italics="true" pageId="3" pageNumber="182">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F6718FF95FE18FBF3FDF9FB89" bold="true" box="[438,518,1060,1084]" pageId="3" pageNumber="182">
<taxonomicNameLabel id="6EC643246718FF95FE18FBF3FDF9FB89" box="[438,518,1060,1084]" pageId="3" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ in its natural environment in Agusan del Sur.
</paragraph>
</caption>
<caption id="13FE72C56718FF95FF39F80CFC13F841" ID-DOI="http://doi.org/10.5281/zenodo.11615159" ID-Zenodo-Dep="11615159" box="[151,1004,2011,2036]" httpUri="https://zenodo.org/record/11615159/files/figure.png" pageId="3" pageNumber="182" startId="3.[151,250,2011,2035]" targetBox="[158,1430,1140,1987]" targetPageId="3" targetType="figure">
<paragraph id="473E224D6718FF95FF39F80CFC13F841" blockId="3.[151,1004,2011,2036]" box="[151,1004,2011,2036]" pageId="3" pageNumber="182">
<emphasis id="75F5FE5F6718FF95FF39F80CFEECF846" bold="true" box="[151,275,2011,2035]" pageId="3" pageNumber="182">FIGURE 2.</emphasis>
<taxonomicName id="808159CE6718FF95FEB7F80BFE50F846" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[281,431,2011,2035]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F6718FF95FEB7F80BFE50F846" box="[281,431,2011,2035]" italics="true" pageId="3" pageNumber="182">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F6718FF95FE18F80BFDF9F841" bold="true" box="[438,518,2012,2036]" pageId="3" pageNumber="182">
<taxonomicNameLabel id="6EC643246718FF95FE18F80BFDF9F841" box="[438,518,2012,2036]" pageId="3" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ in its natural environment in Agusan del Sur.
</paragraph>
</caption>
<caption id="13FE72C5671FFF92FF39FBC6FB74FB9C" ID-DOI="http://doi.org/10.5281/zenodo.11615163" ID-Zenodo-Dep="11615163" box="[151,1163,1041,1065]" httpUri="https://zenodo.org/record/11615163/files/figure.png" pageId="4" pageNumber="183" startId="4.[151,250,1041,1065]" targetBox="[151,1436,181,1017]" targetPageId="4" targetType="figure">
<paragraph id="473E224D671FFF92FF39FBC6FB74FB9C" blockId="4.[151,1163,1041,1065]" box="[151,1163,1041,1065]" pageId="4" pageNumber="183">
<emphasis id="75F5FE5F671FFF92FF39FBC6FEECFB9C" bold="true" box="[151,275,1041,1065]" pageId="4" pageNumber="183">FIGURE 3.</emphasis>
<taxonomicName id="808159CE671FFF92FEB7FBC5FE50FB9C" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[281,431,1041,1065]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F671FFF92FEB7FBC5FE50FB9C" box="[281,431,1041,1065]" italics="true" pageId="4" pageNumber="183">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F671FFF92FE18FBC6FDF9FB9C" bold="true" box="[438,518,1041,1065]" pageId="4" pageNumber="183">
<taxonomicNameLabel id="6EC64324671FFF92FE18FBC6FDF9FB9C" box="[438,518,1041,1065]" pageId="4" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ habitus in lateral (A) and dorsal (B) views. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="473E224D671FFF92FF69FA33FE15F8AB" blockId="4.[151,1437,1147,1966]" pageId="4" pageNumber="183">
<emphasis id="75F5FE5F671FFF92FF69FA33FEF4FA4B" bold="true" box="[199,267,1508,1534]" pageId="4" pageNumber="183">Male.</emphasis>
1A vein on left tegmen straight and swollen (
<figureCitation id="DFBA3EC8671FFF92FCB3FA34FC85FA4B" box="[797,890,1507,1534]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4A</figureCitation>
). Stridulatory file on 1A vein of left tegmen, in ventral view, about 
<quantity id="80798FA8671FFF92FEDAF9DFFE35F997" box="[372,458,1544,1570]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="183" unit="mm" value="1.7">1.7 mm</quantity>
long, mostly straight but slightly curved at basal end, with 97 stridulatory teeth; at anal end having teeth particularly densely packed, at basal end having teeth more widely spaced apart; tooth length generally increases from anal end to maximum of 
<quantity id="80798FA8671FFF92FD75F987FCBEF9DF" box="[731,833,1616,1642]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="183" unit="mm" value="0.12">0.12 mm</quantity>
before decreasing drastically at basal end (
<figureCitation id="DFBA3EC8671FFF92FA9DF998FA6FF9DF" box="[1331,1424,1615,1642]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4D</figureCitation>
). Average tooth length at anal third 
<quantity id="80798FA8671FFF92FDB0F9A3FD7AF93B" box="[542,645,1652,1678]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="4" pageNumber="183" unit="mm" value="0.07">0.07 mm</quantity>
, at middle 
<quantity id="80798FA8671FFF92FCABF9A3FC92F93B" box="[773,877,1652,1678]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="183" unit="mm" value="0.08">0.08 mm</quantity>
, at basal third 
<quantity id="80798FA8671FFF92FBB8F9A3FB84F93B" box="[1046,1147,1652,1678]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="183" unit="mm" value="0.1">0.10 mm</quantity>
(
<figureCitation id="DFBA3EC8671FFF92FB22F9A4FB17F93B" box="[1164,1256,1651,1678]" captionStart="FIGURE 5" captionStartId="6.[151,250,705,729]" captionTargetBox="[161,1426,191,671]" captionTargetId="figure-21@6.[151,1436,181,681]" captionTargetPageId="6" captionText="FIGURE 5. Tooth distribution on the stridulatory file of Ducetia laniae sp. nov. ♂ based on tooth length (A) and inter-tooth distance (B)." figureDoi="http://doi.org/10.5281/zenodo.11615169" httpUri="https://zenodo.org/record/11615169/files/figure.png" pageId="4" pageNumber="183">Fig. 5A</figureCitation>
). Average inter-tooth distance at anal end and middle region 
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, indistinctly smaller than at basal end of 
<quantity id="80798FA8671FFF92FB44F94FFAA1F907" box="[1258,1374,1688,1714]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="183" unit="mm" value="0.03">0.030 mm</quantity>
(
<figureCitation id="DFBA3EC8671FFF92FAC1F940FF44F963" captionStart="FIGURE 5" captionStartId="6.[151,250,705,729]" captionTargetBox="[161,1426,191,671]" captionTargetId="figure-21@6.[151,1436,181,681]" captionTargetPageId="6" captionText="FIGURE 5. Tooth distribution on the stridulatory file of Ducetia laniae sp. nov. ♂ based on tooth length (A) and inter-tooth distance (B)." figureDoi="http://doi.org/10.5281/zenodo.11615169" httpUri="https://zenodo.org/record/11615169/files/figure.png" pageId="4" pageNumber="183">Fig. 5B</figureCitation>
). Mirror area of left tegmen rectangular with anterior and posterior margins parallel, anal and basal margins slightly converging posteriorly; anterior margin 1.2–1.3 times as long as posterior margin, 1.8–1.9 times as long as the length of mirror (
<figureCitation id="DFBA3EC8671FFF92FE2DF8D4FE22F8AB" box="[387,477,1795,1822]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="473E224D671FFF92FF69F8F0FC57F81B" blockId="4.[151,1437,1147,1966]" pageId="4" pageNumber="183">
Abdominal apex typical of 
<taxonomicName id="808159CE671FFF92FE51F8FFFD3DF8F4" baseAuthorityName="Thunberg" baseAuthorityYear="1815" box="[511,706,1832,1857]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="75F5FE5F671FFF92FE51F8FFFD3DF8F4" box="[511,706,1832,1857]" italics="true" pageId="4" pageNumber="183">Ducetia japonica</emphasis>
</taxonomicName>
species group. Cercus in lateral view with basal half cylindrical and straight; with apical half curved dorsad; with apical third having characteristic interno-ventral ridge (
<figureCitation id="DFBA3EC8671FFF92FAC1F89CFF44F83F" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4E</figureCitation>
). Subgenital plate elongated; apical half deeply and narrowly emarginated at middle; with inner margins of emargination nearly touching each other except at basal end (
<figureCitation id="DFBA3EC8671FFF92FCE8F844FC63F81B" box="[838,924,1939,1966]" captionStart="FIGURE 4" captionStartId="5.[151,250,1736,1760]" captionTargetBox="[151,1436,181,1712]" captionTargetId="figure-18@5.[151,1436,181,1712]" captionTargetPageId="5" captionText="FIGURE 4. Ducetia laniae sp. nov. ♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D)." figureDoi="http://doi.org/10.5281/zenodo.11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="4" pageNumber="183">Fig. 4F</figureCitation>
).
</paragraph>
<caption id="13FE72C5671EFF93FF39F91FFB93F89D" ID-DOI="http://doi.org/10.5281/zenodo.11615165" ID-Zenodo-Dep="11615165" httpUri="https://zenodo.org/record/11615165/files/figure.png" pageId="5" pageNumber="184" startId="5.[151,250,1736,1760]" targetBox="[151,1436,181,1712]" targetPageId="5" targetType="figure">
<paragraph id="473E224D671EFF93FF39F91FFB93F89D" blockId="5.[151,1436,1736,1832]" pageId="5" pageNumber="184">
<emphasis id="75F5FE5F671EFF93FF39F91FFEEBF955" bold="true" box="[151,276,1736,1760]" pageId="5" pageNumber="184">FIGURE 4.</emphasis>
<taxonomicName id="808159CE671EFF93FEB5F91EFE4EF955" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[283,433,1736,1760]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F671EFF93FEB5F91EFE4EF955" box="[283,433,1736,1760]" italics="true" pageId="5" pageNumber="184">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F671EFF93FE17F91FFDF5F955" bold="true" box="[441,522,1736,1760]" pageId="5" pageNumber="184">
<taxonomicNameLabel id="6EC64324671EFF93FE17F91FFDF5F955" box="[441,522,1736,1760]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂: head, pronotum and anterior part of tegmina in dorsal view (A); face in anterior view (B); head, pronotum and anterior part of tegmina in lateral view (C); stridulatory file of the left tegmen in ventral view (D); abdominal apex in lateral (E) and ventral (F) views. Scale bars: 2 mm (A, C, E, F); 1 mm (B, D).
</paragraph>
</caption>
<paragraph id="473E224D671EFF93FF69F8ACFBA3F869" blockId="5.[151,1436,1914,2013]" pageId="5" pageNumber="184">
<emphasis id="75F5FE5F671EFF93FF69F8ACFE75F821" bold="true" box="[199,394,1914,1941]" pageId="5" pageNumber="184">
Female (
<figureCitation id="DFBA3EC8671EFF93FE9FF8ADFE85F821" box="[305,378,1914,1940]" captionStart="FIGURE 6" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[177,1410,181,1919]" captionTargetId="figure-18@7.[177,1410,181,1919]" captionTargetPageId="7" captionText="FIGURE 6. Ducetia laniae sp. nov. ♀: habitus in lateral (A) and dorsal (B) views; head, pronotum and anterior part of tegmina in dorsal view (C); abdominal apex in lateral (D) and ventral (E) views. Scale bars: 5 mm (A, B); 2 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.11615173" httpUri="https://zenodo.org/record/11615173/files/figure.png" pageId="5" pageNumber="184">Fig. 6</figureCitation>
).
</emphasis>
Habitus similar to male (
<figureCitation id="DFBA3EC8671EFF93FD14F8ADFCC2F821" box="[698,829,1914,1940]" captionStart="FIGURE 6" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[177,1410,181,1919]" captionTargetId="figure-18@7.[177,1410,181,1919]" captionTargetPageId="7" captionText="FIGURE 6. Ducetia laniae sp. nov. ♀: habitus in lateral (A) and dorsal (B) views; head, pronotum and anterior part of tegmina in dorsal view (C); abdominal apex in lateral (D) and ventral (E) views. Scale bars: 5 mm (A, B); 2 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.11615173" httpUri="https://zenodo.org/record/11615173/files/figure.png" pageId="5" pageNumber="184">Figs 6A–C</figureCitation>
). Ovipositor falcate with dorsal valve having dorsal margin from basal third dentated; with ventral valve having ventral margin dentated only at the apex (
<figureCitation id="DFBA3EC8671EFF93FA9DF849FA6FF80D" box="[1331,1424,1950,1976]" captionStart="FIGURE 6" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[177,1410,181,1919]" captionTargetId="figure-18@7.[177,1410,181,1919]" captionTargetPageId="7" captionText="FIGURE 6. Ducetia laniae sp. nov. ♀: habitus in lateral (A) and dorsal (B) views; head, pronotum and anterior part of tegmina in dorsal view (C); abdominal apex in lateral (D) and ventral (E) views. Scale bars: 5 mm (A, B); 2 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.11615173" httpUri="https://zenodo.org/record/11615173/files/figure.png" pageId="5" pageNumber="184">Fig. 6D</figureCitation>
). Subgenital plate small; in lateral and ventral view, triangular; apex subacute (
<figureCitation id="DFBA3EC8671EFF93FC56F815FBB0F869" box="[1016,1103,1986,2012]" captionStart="FIGURE 6" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[177,1410,181,1919]" captionTargetId="figure-18@7.[177,1410,181,1919]" captionTargetPageId="7" captionText="FIGURE 6. Ducetia laniae sp. nov. ♀: habitus in lateral (A) and dorsal (B) views; head, pronotum and anterior part of tegmina in dorsal view (C); abdominal apex in lateral (D) and ventral (E) views. Scale bars: 5 mm (A, B); 2 mm (C–E)." figureDoi="http://doi.org/10.5281/zenodo.11615173" httpUri="https://zenodo.org/record/11615173/files/figure.png" pageId="5" pageNumber="184">Fig. 6E</figureCitation>
).
</paragraph>
<caption id="13FE72C5671DFF90FF39FD16FEE6FD48" ID-DOI="http://doi.org/10.5281/zenodo.11615169" ID-Zenodo-Dep="11615169" httpUri="https://zenodo.org/record/11615169/files/figure.png" pageId="6" pageNumber="185" startId="6.[151,250,705,729]" targetBox="[161,1426,191,671]" targetPageId="6" targetType="figure">
<paragraph id="473E224D671DFF90FF39FD16FEE6FD48" blockId="6.[151,1436,705,765]" pageId="6" pageNumber="185">
<emphasis id="75F5FE5F671DFF90FF39FD16FEEBFD6C" bold="true" box="[151,276,705,729]" pageId="6" pageNumber="185">FIGURE 5.</emphasis>
Tooth distribution on the stridulatory file of 
<taxonomicName id="808159CE671DFF90FD4AFD15FC84FD6C" authority="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan, 2024" authorityName="Salvador &amp; Nuñeza &amp; Robillard &amp; Tan" authorityYear="2024" box="[740,891,705,729]" class="Insecta" family="Tettigoniidae" genus="Ducetia" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="185" phylum="Arthropoda" rank="species" species="laniae" status="sp. nov.">
<emphasis id="75F5FE5F671DFF90FD4AFD15FC84FD6C" box="[740,891,705,729]" italics="true" pageId="6" pageNumber="185">Ducetia laniae</emphasis>
</taxonomicName>
<emphasis id="75F5FE5F671DFF90FC2CFD16FC2BFD6C" bold="true" box="[898,980,705,729]" pageId="6" pageNumber="185">
<taxonomicNameLabel id="6EC64324671DFF90FC2CFD16FC2BFD6C" box="[898,980,705,729]" pageId="6" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ based on tooth length (A) and inter-tooth distance (B).
</paragraph>
</caption>
<paragraph id="473E224D671DFF90FF69FC98FD30FCDC" blockId="6.[151,1437,847,1378]" box="[199,719,847,877]" pageId="6" pageNumber="185">
<emphasis id="75F5FE5F671DFF90FF69FC98FDC0FCDC" bold="true" box="[199,575,847,877]" pageId="6" pageNumber="185">
Measurements (
<specimenCount id="5187E9C4671DFF90FE29FC86FE55FCD8" box="[391,426,849,877]" count="2" pageId="6" pageNumber="185" type="male">2♂</specimenCount>
, 
<specimenCount id="5187E9C4671DFF90FE16FC86FE24FCD8" box="[440,475,849,877]" count="1" pageId="6" pageNumber="185" type="female">1♀</specimenCount>
in mm).
</emphasis>
See 
<tableCitation id="0A0317F6671DFF90FDDBFC98FD36FCDC" box="[629,713,847,873]" captionStart="TABLE 1" captionStartId="6.[151,239,1425,1449]" captionTargetPageId="6" captionText="TABLE 1. Measurements of Ducetia laniae sp. nov. (2♂, 1♀ in mm)." httpUri="http://table.plazi.org/id/13FE72C5671DFF90FF39FA46FC7AFA1C" pageId="6" pageNumber="185" tableUuid="13FE72C5671DFF90FF39FA46FC7AFA1C">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C6671DFF90FF69FCA4FE57FC04" pageId="6" pageNumber="185" type="biology_ecology">
<paragraph id="473E224D671DFF90FF69FCA4FE57FC04" blockId="6.[151,1437,847,1378]" pageId="6" pageNumber="185">
<emphasis id="75F5FE5F671DFF90FF69FCA4FED3FC38" bold="true" box="[199,300,883,909]" pageId="6" pageNumber="185">Ecology.</emphasis>
This species is common in 
<collectingRegion id="8545ECAF671DFF90FDCBFCA3FCEAFC38" box="[613,789,883,910]" country="Philippines" name="Agusan del Sur" pageId="6" pageNumber="185">Agusan del Sur</collectingRegion>
, and can be mostly found in urban communities dwelling in dense bushes (
<figureCitation id="DFBA3EC8671DFF90FEF8FC40FE64FC04" box="[342,411,919,945]" captionStart="FIGURE 8" captionStartId="9.[151,250,1982,2006]" captionTargetBox="[151,1436,489,1959]" captionTargetId="figure-152@9.[151,1436,489,1959]" captionTargetPageId="9" captionText="FIGURE 8. Habitats in the municipalities of Sibagat (A) and Bunawan (B), Agusan del Sur where Ducetia laniae sp. nov. was found." figureDoi="http://doi.org/10.5281/zenodo.11615180" httpUri="https://zenodo.org/record/11615180/files/figure.png" pageId="6" pageNumber="185">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C6671DFF90FF69FC6CFC68FC4C" pageId="6" pageNumber="185" type="distribution">
<paragraph id="473E224D671DFF90FF69FC6CFD8EFC60" blockId="6.[151,1437,847,1378]" box="[199,625,955,981]" pageId="6" pageNumber="185">
<emphasis id="75F5FE5F671DFF90FF69FC6CFE9EFC60" bold="true" box="[199,353,955,981]" pageId="6" pageNumber="185">Distribution.</emphasis>
<collectingCountry id="3F9662DD671DFF90FEC9FC6CFE1AFC60" box="[359,485,955,981]" name="Philippines" pageId="6" pageNumber="185">Philippines</collectingCountry>
(Mindanao)
</paragraph>
<paragraph id="473E224D671DFF90FF69FC08FC68FC4C" blockId="6.[151,1437,847,1378]" box="[199,919,991,1018]" pageId="6" pageNumber="185">
<emphasis id="75F5FE5F671DFF90FF69FC08FE99FC4C" bold="true" box="[199,358,991,1017]" pageId="6" pageNumber="185">Type locality.</emphasis>
<collectingCountry id="3F9662DD671DFF90FEC2FC08FDEAFC4C" box="[364,533,991,1017]" name="Philippines" pageId="6" pageNumber="185">PHILIPPINES</collectingCountry>
: Mindanao Island: 
<collectingRegion id="8545ECAF671DFF90FD44FC37FC68FC4C" box="[746,919,991,1018]" country="Philippines" name="Agusan del Sur" pageId="6" pageNumber="185">Agusan del Sur</collectingRegion>
</paragraph>
</subSubSection>
<subSubSection id="0F9B71C6671DFF90FF69FBD4FDEAFAD4" pageId="6" pageNumber="185" type="biology_ecology">
<paragraph id="473E224D671DFF90FF69FBD4FDEAFAD4" blockId="6.[151,1437,847,1378]" pageId="6" pageNumber="185">
<emphasis id="75F5FE5F671DFF90FF69FBD4FD9EFBA8" bold="true" box="[199,609,1027,1057]" pageId="6" pageNumber="185">
Calling song (
<specimenCount id="5187E9C4671DFF90FEC1FBD2FE6DFB94" box="[367,402,1029,1057]" count="1" pageId="6" pageNumber="185" type="male">1♂</specimenCount>
, 28.6°C) (
<figureCitation id="DFBA3EC8671DFF90FDA5FBD4FDADFBA8" box="[523,594,1027,1053]" captionStart="FIGURE 7" captionStartId="8.[151,250,1541,1565]" captionTargetBox="[156,1429,191,1507]" captionTargetId="figure-21@8.[151,1436,181,1517]" captionTargetPageId="8" captionText="FIGURE 7. Ducetia laniae sp. nov. ♂ calling song: oscillogram (A) and spectrogram (B) showing two echeme-sequences; oscillograms showing four echemes (C) and one echeme (D); power spectra of an echeme (E) where the gray line represents the power [dB] (axis name and scale in gray) and the black solid peak represents the relative amplitude (axis name in black)." figureDoi="http://doi.org/10.5281/zenodo.11615177" httpUri="https://zenodo.org/record/11615177/files/figure.png" pageId="6" pageNumber="185">Fig. 7</figureCitation>
).
</emphasis>
The calling song is an echeme-sequence composed of complex echemes (n = 2), and has a duration of 8.28– 
<geoCoordinate id="22B5448A671DFF90FD98FBF0FD75FBF7" box="[566,650,1063,1090]" degrees="10.67" direction="south" orientation="latitude" pageId="6" pageNumber="185" precision="555" value="-10.67">10.67 s</geoCoordinate>
. Each echeme-sequence is made up of 46–56 echemes. The average echeme duration is 0.18± 
<geoCoordinate id="22B5448A671DFF90FE18FB9CFE04FBD3" box="[438,507,1099,1126]" degrees="0.02" direction="south" orientation="latitude" pageId="6" pageNumber="185" precision="555" value="-0.02">0.02 s</geoCoordinate>
(0.15– 
<geoCoordinate id="22B5448A671DFF90FDE5FB9CFD6EFBD0" box="[587,657,1099,1126]" degrees="0.23" direction="south" orientation="latitude" pageId="6" pageNumber="185" precision="555" value="-0.23">0.23 s</geoCoordinate>
), and the average interval between consecutive echemes is 20.8±2.4 ms (16.0–23.7 ms). At the start of each echeme, the first syllable is particularly long, having a duration of 12.8±2.5 ms (8.4–18.3 ms), and has the highest amplitude. The average interval between the first and second syllables is 9.0±3.8 ms (4.9–16.5 ms). The remaining part of the echeme consists of 16±3 (13–21) syllables and has an average duration of 0.16± 
<geoCoordinate id="22B5448A671DFF90FEF6FB0CFE64FB43" box="[344,411,1243,1270]" degrees="0.02" direction="south" orientation="latitude" pageId="6" pageNumber="185" precision="555" value="-0.02">0.02 s</geoCoordinate>
(0.14– 
<geoCoordinate id="22B5448A671DFF90FE47FB0CFDD2FB40" box="[489,557,1243,1269]" degrees="0.20" direction="south" orientation="latitude" pageId="6" pageNumber="185" precision="555" value="-0.2">0.20 s</geoCoordinate>
). These syllables show up as possibly rapid-decay pulses. The average syllable duration is distinctly shorter than the first syllable, i.e., 5.5±1.0 ms (3.4–7.1 ms), and the average interval between these consecutive syllables is 5.4±1.5 ms (2.3–7.5 ms). The call spectrum is broad-band, and has a peak frequency of 17.2±1.1 kHz (15.8–20.8 kHz).
</paragraph>
</subSubSection>
</treatment>
</document>