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<document id="7C25F8BAC4FF4BF575917C5A780CEA06" ID-DOI="10.11646/zootaxa.3630.2.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="222179" ID-ZooBank="70C11335-A5F6-4BCB-98E6-4D14EB1A4015" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1460584307806" checkinUser="plazi" docAuthor="Parker, Joseph &amp; Maruyama, Munetoshi" docDate="2013" docId="03FB8786FFE7FFCF498EFAC6B0DBFA83" docLanguage="en" docName="zt03630p378.pdf" docOrigin="Zootaxa 3630 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Jubogaster Parker &amp; Maruyama 2013, gen. nov." docType="treatment" docUuid="C4617491-F55F-4757-9189-A09F56E27AE2" docUuidSource="ZooBank" docVersion="2" lastPageNumber="372" masterDocId="FFC2FFFEFFE5FFCC4919FFE8B240FFFB" masterDocTitle="Jubogaster towai, a new Neotropical genus and species of Trogastrini (Coleoptera: Staphylinidae: Pselaphinae) exhibiting myrmecophily and extreme body enlargement" masterLastPageNumber="378" masterPageNumber="369" pageNumber="371" updateTime="1730809485467" updateUser="ExternalLinkService">
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<mods:title id="680FA9A8B06D22FDAEA5BBD120DD44D3">Jubogaster towai, a new Neotropical genus and species of Trogastrini (Coleoptera: Staphylinidae: Pselaphinae) exhibiting myrmecophily and extreme body enlargement</mods:title>
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<mods:namePart id="64B214466E52A94CD3DB5BAB37058EBF">Parker, Joseph</mods:namePart>
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<mods:namePart id="3B236B6E510A66B50AAEDD4CA50FBC5C">Maruyama, Munetoshi</mods:namePart>
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<mods:typeOfResource id="DD5D754FB376479BEEDB4CD37673FAA9">text</mods:typeOfResource>
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<mods:title id="200A845623AF8D03E45AE935F3E7D4E4">Zootaxa</mods:title>
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<mods:part id="B4EF0C898543A27D53167E8307798BBF">
<mods:date id="BB173061770AA8C5D44DA6BC3E7C5F5F">2013</mods:date>
<mods:detail id="4D3E7D2E5AD58B3E132B8733C8390575" type="pubDate">
<mods:number id="23E59DDEA809EC025246B48729CA4973">2013-03-20</mods:number>
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<mods:number id="E8EF333DA449E3C6D20654859F4EE278">3630</mods:number>
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<mods:classification id="EE490EC8DB68D658B9B1C1CF4914AC34">journal article</mods:classification>
<mods:identifier id="F42A4DFEBBCD82FB970B889B50481107" type="DOI">10.11646/zootaxa.3630.2.11</mods:identifier>
<mods:identifier id="A05A61D4F58B93EE030DF7B2E30970AE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FB8786FFE7FFCF498EFAC6B0DBFA83" ID-DOI="http://doi.org/10.5281/zenodo.14040089" ID-Zenodo-Dep="14040089" LSID="urn:lsid:zoobank.org:act:C4617491-F55F-4757-9189-A09F56E27AE2" httpUri="http://treatment.plazi.org/id/03FB8786FFE7FFCF498EFAC6B0DBFA83" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371">
<subSubSection id="C348651BFFE7FFCE498EFAC6B3C6FABC" box="[151,390,1326,1351]" pageId="2" pageNumber="371" type="nomenclature">
<paragraph id="8BED3690FFE7FFCE498EFAC6B3C6FABC" blockId="2.[151,390,1326,1351]" box="[151,390,1326,1351]" pageId="2" pageNumber="371">
<emphasis id="B926EA82FFE7FFCE498EFAC6B3C6FABC" bold="true" box="[151,390,1326,1351]" pageId="2" pageNumber="371">
<taxonomicName id="4C524D13FFE7FFCE498EFAC6B357FABC" authority="Parker &amp; Maruyama, 2013" authorityName="Parker &amp; Maruyama" authorityYear="2013" box="[151,279,1326,1351]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926EA82FFE7FFCE498EFAC6B357FABC" bold="true" box="[151,279,1326,1351]" italics="true" pageId="2" pageNumber="371">Jubogaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557F9FFE7FFCE4804FAC7B3C6FABC" box="[285,390,1327,1351]" pageId="2" pageNumber="371" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C348651BFFE7FFCE498EFA98B155FA7C" box="[151,789,1391,1416]" pageId="2" pageNumber="371" type="type_taxon">
<paragraph id="8BED3690FFE7FFCE498EFA98B155FA7C" blockId="2.[151,1436,1391,2028]" box="[151,789,1391,1416]" pageId="2" pageNumber="371">
<emphasis id="B926EA82FFE7FFCE498EFA98B372FA73" bold="true" box="[151,306,1391,1416]" pageId="2" pageNumber="371">
<typeStatus id="54E98832FFE7FFCE498EFA98B292FA73" box="[151,210,1392,1416]" pageId="2" pageNumber="371">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C524D13FFE7FFCE4820FA98B3BAFA73" box="[313,506,1392,1416]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai" status="sp. nov.">
<emphasis id="B926EA82FFE7FFCE4820FA98B3BAFA73" box="[313,506,1392,1416]" italics="true" pageId="2" pageNumber="371">Jubogaster towai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557F9FFE7FFCE4B18FA98B018FA7C" box="[513,600,1392,1415]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
here designated.
</paragraph>
</subSubSection>
<subSubSection id="C348651BFFE7FFCE498EFA7BB2B6F90B" pageId="2" pageNumber="371" type="diagnosis">
<paragraph id="8BED3690FFE7FFCE498EFA7BB2B6F90B" blockId="2.[151,1436,1391,2028]" pageId="2" pageNumber="371">
<emphasis id="B926EA82FFE7FFCE498EFA7BB351FA57" bold="true" box="[151,273,1427,1452]" pageId="2" pageNumber="371">Diagnosis.</emphasis>
Euplectite pselaphines with the following characters: transverse head articulating ventrally under pronotum when deflexed, typical of
<taxonomicName id="4C524D13FFE7FFCE4B21FA5FB0F7FA34" box="[568,695,1463,1487]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
. Head with vertex excavate, lacking vertexal fovea; ventral head without the typical dense “genal beard” of setae common to most
<taxonomicName id="4C524D13FFE7FFCE4AADFA34B673FA0F" box="[948,1075,1500,1524]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
. Eleven-segmented moderately geniculate antennae, received by large excavations anterior to eyes. Pronotum jubine-like: broad and obcordate, constricted at basal quarter, with weakly-evident medial longitudinal sulcus and distinct antebasal sulcus. Pronotum afoveate, with margins smooth, devoid of crenulations or small spines found in other
<taxonomicName id="4C524D13FFE7FFCE4DD7F9AFB70DF9A4" box="[1230,1357,1607,1631]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
. Elytra lacking any stereotypical fovea. Legs with contiguous, projecting coxae, broad femora, basally flattened tibia, and three tarsal segments; segment 1 very small, 2 and 3 elongate and subequal in length. Tarsi with two equally-sized claws, unlike other
<taxonomicName id="4C524D13FFE7FFCE486DF95CB3AFF937" box="[372,495,1716,1740]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
which have unequally-sized claws. Bifid anterior protarsal claw in the
<typeStatus id="54E98832FFE7FFCE4C0AF95DB703F936" box="[1299,1347,1717,1741]" pageId="2" pageNumber="371">type</typeStatus>
species
<taxonomicName id="4C524D13FFE7FFCE498EF931B2AFF90B" box="[151,239,1753,1776]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai">
<emphasis id="B926EA82FFE7FFCE498EF931B2AFF90B" box="[151,239,1753,1776]" italics="true" pageId="2" pageNumber="371">J. towai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C348651BFFE7FFCE49D0F914B1CEF85F" pageId="2" pageNumber="371" type="etymology">
<paragraph id="8BED3690FFE7FFCE49D0F914B1CEF85F" blockId="2.[151,1436,1391,2028]" pageId="2" pageNumber="371">
<emphasis id="B926EA82FFE7FFCE49D0F914B30FF8EF" bold="true" box="[201,335,1788,1812]" pageId="2" pageNumber="371">Etymology.</emphasis>
A combination of
<taxonomicName id="4C524D13FFE7FFCE4B38F914B09AF8EF" authority="Schaufuss" authorityName="Schaufuss" box="[545,730,1788,1812]" class="Insecta" family="Staphylinidae" genus="Jubus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B926EA82FFE7FFCE4B38F914B022F8EF" box="[545,610,1788,1812]" italics="true" pageId="2" pageNumber="371">Jubus</emphasis>
Schaufuss
</taxonomicName>
and
<taxonomicName id="4C524D13FFE7FFCE4A09F915B186F8EF" authority="Sharp" authorityName="Sharp" box="[784,966,1788,1812]" class="Insecta" family="Staphylinidae" genus="Trogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B926EA82FFE7FFCE4A09F915B13CF8EF" box="[784,892,1789,1812]" italics="true" pageId="2" pageNumber="371">Trogaster</emphasis>
Sharp
</taxonomicName>
, the
<typeStatus id="54E98832FFE7FFCE4AE3F915B66AF8EE" box="[1018,1066,1789,1813]" pageId="2" pageNumber="371">type</typeStatus>
genera of Jubini and
<taxonomicName id="4C524D13FFE7FFCE4C03F914B7D8F8EF" box="[1306,1432,1788,1812]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
, respectively. The name is chosen on account of the membership of the new genus in
<taxonomicName id="4C524D13FFE7FFCE4D43F8F7B698F8CC" box="[1114,1240,1823,1847]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="tribe" tribe="Trogastrini">Trogastrini</taxonomicName>
, despite apparent Jubini-like characters such as the shape of the pronotum and the presence of two equally-sized tarsal claws (a character state found in Jubini, but not other Euplectitae). The bifid anterior protarsal claw of the
<typeStatus id="54E98832FFE7FFCE4DEFF881B766F87A" box="[1270,1318,1897,1921]" pageId="2" pageNumber="371">type</typeStatus>
species
<taxonomicName id="4C524D13FFE7FFCE4C91F881B295F85F" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai">
<emphasis id="B926EA82FFE7FFCE4C91F881B295F85F" italics="true" pageId="2" pageNumber="371">J. towai</emphasis>
</taxonomicName>
is also shared with
<taxonomicName id="4C524D13FFE7FFCE48AAF864B15EF85F" authority="Silvestri" authorityName="Silvestri" box="[435,798,1932,1956]" class="Insecta" family="Staphylinidae" genus="Phamisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="myrmophilus">
<emphasis id="B926EA82FFE7FFCE48AAF864B0F8F85F" box="[435,696,1932,1956]" italics="true" pageId="2" pageNumber="371">Phamisus myrmophilus</emphasis>
Silvestri
</taxonomicName>
of Jubini.
</paragraph>
</subSubSection>
<subSubSection id="C348651BFFE7FFCF49D0F847B0DBFA83" lastPageId="3" lastPageNumber="372" pageId="2" pageNumber="371" type="description">
<paragraph id="8BED3690FFE7FFCE49D0F847B388F817" blockId="2.[151,1436,1391,2028]" pageId="2" pageNumber="371">
<emphasis id="B926EA82FFE7FFCE49D0F847B31AF833" bold="true" box="[201,346,1967,1992]" pageId="2" pageNumber="371">Description.</emphasis>
Body large (
<quantity id="4CAA9B75FFE7FFCE48F4F858B07EF833" box="[493,574,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="371" unit="mm" value="5.1">5.1mm</quantity>
for the
<typeStatus id="54E98832FFE7FFCE4B8EF859B087F832" box="[663,711,1969,1993]" pageId="2" pageNumber="371">type</typeStatus>
species
<taxonomicName id="4C524D13FFE7FFCE4A31F859B1C1F833" box="[808,897,1969,1992]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="towai">
<emphasis id="B926EA82FFE7FFCE4A31F859B1C1F833" box="[808,897,1969,1992]" italics="true" pageId="2" pageNumber="371">J. towai</emphasis>
</taxonomicName>
shown in
<figureCitation id="13692A15FFE7FFCE4AEEF858B615F833" box="[1015,1109,1968,1992]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="2" pageNumber="371">Figure 1</figureCitation>
). Body shape approximately ovoid and strongly convex.
</paragraph>
<paragraph id="8BED3690FFE6FFCF49D0FF7FB63FFE0F" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
<emphasis id="B926EA82FFE6FFCF49D0FF7FB350FF4B" bold="true" box="[201,272,151,176]" italics="true" pageId="3" pageNumber="372">Head:</emphasis>
Approximately triangular, strongly transverse (
<figureCitation id="13692A15FFE6FFCF4A30FF7FB129FF54" box="[809,873,151,175]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 4</figureCitation>
). Vertex afoveate, concave, with moderately deep excavation extending from basal quarter to anterior margin of frons; excavation narrowing apically along midline as it passes between raised frontolateral margins. Frontolateral margins carinate, extending roughly arcuately from posterior of antennal insertion cavity to clypeus (
<figureCitation id="13692A15FFE6FFCF4BD7FEECB14FFEE7" box="[718,783,260,284]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
). Frontolateral face accommodating large circular cavity anterior to eyes to receive antennae (
<figureCitation id="13692A15FFE6FFCF4B50FECFB0CBFEC4" box="[585,651,295,319]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
). Eye flanked directly anteriorly by additional large excavation for retracted maxillary palpus (
<figureCitation id="13692A15FFE6FFCF48CAFEA4B054FE9F" box="[467,532,332,356]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
). Ventral surface of head bisected by longitudinal gular carina situated within broader gular sulcus terminating in gular fovea. Gular sulcus widens basally, spanning the head-neck interface. Maxillary palpi apparently 4-segmented, with segment II long and curved at apex, III short and triangular, wider than II, IV oval, approximately
<date id="FFEC1050FFE6FFCF4B18FE5FB05DFE34" box="[513,541,439,463]" pageId="3" pageNumber="372">2x</date>
longer than wide (
<figureCitation id="13692A15FFE6FFCF4BECFE5FB174FE34" box="[757,820,439,463]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 5</figureCitation>
). Antennae (
<figureCitation id="13692A15FFE6FFCF4ADFFE5FB643FE34" box="[966,1027,439,463]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 3</figureCitation>
) with 11 antennomeres, moderately geniculate with enlarged segment I. Antennomeres IX
<emphasis id="B926EA82FFE6FFCF4BEDFE34B142FE0F" bold="true" box="[756,770,476,500]" pageId="3" pageNumber="372">–</emphasis>
XI strongly elongate (XI longest).
</paragraph>
<paragraph id="8BED3690FFE6FFCF49D0FE17B10CFCCC" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
<emphasis id="B926EA82FFE6FFCF49D0FE17B368FDE3" bold="true" box="[201,296,511,536]" italics="true" pageId="3" pageNumber="372">Thorax:</emphasis>
Pronotum (
<figureCitation id="13692A15FFE6FFCF48A8FE17B3B0FDEC" box="[433,496,511,535]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 4</figureCitation>
) obcordate,
<date id="FFEC1050FFE6FFCF4B98FE17B0DDFDEC" box="[641,669,511,535]" pageId="3" pageNumber="372">2x</date>
wider than head, almost as wide as elytra. Afoveate, with smooth margins. Anterior margin broadly arcuate, largely obscuring the deflexed head when viewed from above, and constricted sharply in basal quarter, so that pronotum is laterally expanded into broad, blunt spurs with curved edges. Medial longitudinal sulcus present on pronotal disc, weakly impressed, thin, terminating at deeper, bowshaped antebasal sulcus that extends between bases of mediolateral spurs. Prosternum afoveate, with lateral longitudinal carinae extending from sides of neck to base. Mesosternum with median mesosternal foveae (MMSF), lateral mesosternal fovea (LMSF) and lateral mesocoxal fovea (LMCF) present. Other typical pselaphine mesosternal fovea listed in
<bibRefCitation id="EFC34B61FFE6FFCF48CDFD14B0CFFCEF" author="Chandler, D. S." box="[468,655,764,788]" journalOrPublisher="Memoirs on Entomology International" pageId="3" pageNumber="372" pagination="1 - 560" part="15" refId="ref6070" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International, 15, 1 - 560." title="Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae)" type="journal article" year="2001">Chandler (2001)</bibRefCitation>
lacking. Metasternum strongly convex. Lateral metathoracic foveae (LMTF) present; median metathoracic fovea (MMTF) absent.
</paragraph>
<paragraph id="8BED3690FFE6FFCF49D0FCABB646FC17" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
<emphasis id="B926EA82FFE6FFCF49D0FCABB301FCA7" bold="true" box="[201,321,835,860]" italics="true" pageId="3" pageNumber="372">Abdomen:</emphasis>
Abdomen with tergites strongly convex. Visible segments 1
<emphasis id="B926EA82FFE6FFCF4AF3FCACB1B8FCA7" bold="true" box="[1002,1016,836,860]" pageId="3" pageNumber="372">–</emphasis>
3 (IV
<emphasis id="B926EA82FFE6FFCF4D2DFCACB602FCA7" bold="true" box="[1076,1090,836,860]" pageId="3" pageNumber="372">–</emphasis>
VI) subequal in length (
<figureCitation id="13692A15FFE6FFCF4C57FCACB7CAFCA7" box="[1358,1418,836,860]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
), segment 4 (VII) longest (not apparent in
<figureCitation id="13692A15FFE6FFCF4B8AFC8FB095FC84" box="[659,725,871,895]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
due to curvature of abdomen). Tergites 1
<emphasis id="B926EA82FFE6FFCF4DCFFC80B6A4FC7B" bold="true" box="[1238,1252,872,896]" pageId="3" pageNumber="372">–</emphasis>
3 (IV
<emphasis id="B926EA82FFE6FFCF4C3EFC80B775FC7B" bold="true" box="[1319,1333,872,896]" pageId="3" pageNumber="372">–</emphasis>
VI) with pronounced paratergites, 4 (VII) with small tapering paratergites. Tergite 6 (IX) internalised. Six sternites visible (
<figureCitation id="13692A15FFE6FFCF4986FC47B29EFC3C" box="[159,222,943,967]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
), sternite 7 (IX) internalised. First visible sternite (III) with median process projecting between metacoxae. Sternites 2 and 3 (IV and V) subequal in length, 4 (VI) slightly shorter (
<figureCitation id="13692A15FFE6FFCF4AA0FC3CB1B8FC17" box="[953,1016,980,1004]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
).
</paragraph>
<paragraph id="8BED3690FFE6FFCF49D0FC1FB723FB5B" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
<emphasis id="B926EA82FFE6FFCF49D0FC1FB35AFBEB" bold="true" box="[201,282,1015,1040]" italics="true" pageId="3" pageNumber="372">Elytra:</emphasis>
Strongly convex, slightly broader than pronotum (
<figureCitation id="13692A15FFE6FFCF4A70FC1FB1E7FBF4" box="[873,935,1015,1039]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 1</figureCitation>
), approximately 0.3x total body length. At their longest, equal in length to visible abdominal segments 2+3 (V+VI) combined. Elytral humeri broadly rounded to almost half elytral length. Sutural stria entire; lacking sutural fovea and other stereotypical antebasal fovea. Lacking subhumeral fovea and subhumeral stria. One weakly defined, somewhat discal depression is evident halfway between suture and humerus; its homology to canonical pselaphine antebasal elytral fovea is unclear.
</paragraph>
<paragraph id="8BED3690FFE6FFCF49D0FB44B0DBFA83" blockId="3.[151,1437,151,1401]" pageId="3" pageNumber="372">
<emphasis id="B926EA82FFE6FFCF49D0FB44B349FB38" bold="true" box="[201,265,1196,1219]" italics="true" pageId="3" pageNumber="372">Legs:</emphasis>
All coxae contiguous (
<figureCitation id="13692A15FFE6FFCF4B03FB44B01BFB3F" box="[538,603,1196,1220]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 2</figureCitation>
), procoxae conical, projecting ventrally, mesocoxae globose, metacoxae (
<figureCitation id="13692A15FFE6FFCF4987FB38B29BFB13" box="[158,219,1232,1256]" captionStart="FIGURES 1 – 7" captionStartId="4.[151,264,1908,1931]" captionTargetBox="[151,1435,211,1884]" captionTargetId="figure@4.[151,1436,195,1884]" captionTargetPageId="4" captionText="FIGURES 1 – 7. Jubogaster towai Parker and Maruyama. 1) dorsal habitus 2) ventral habitus 3) left antenna, exterior face is to the left 4) vertex of head and pronotum 5) frontal of view of head, showing excavations for antennae and maxillary palpi 6) right elytron apical angle showing excision 7) left metacoxa, metatrochanter with arrow pointing to rhomboid-shaped tubercle." httpUri="https://zenodo.org/record/222180/files/figure.png" pageId="3" pageNumber="372">Fig 7</figureCitation>
) transverse-conical, spanning from ventral midline to metasternal margin, and projecting posteriorly. Femora somewhat thickened, and excavated apically to receive retracted tibia. Tibia flattened basally. Femora and tibia lacking spines or processes. Tarsi elongate, 3-segmented, first segment extremely short, 0.2x the length of segment 2. Segment 2 and 3 subequal in length. All tarsi with 2 well-developed, equally-sized tarsal claws. Bifid anterior protarsal claw in
<typeStatus id="54E98832FFE6FFCF4841FA89B3C8FA82" box="[344,392,1377,1401]" pageId="3" pageNumber="372">type</typeStatus>
species
<taxonomicName id="4C524D13FFE6FFCF48FEFA89B000FA83" box="[487,576,1377,1400]" class="Insecta" family="Silphidae" genus="Jubogaster" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="towai">
<emphasis id="B926EA82FFE6FFCF48FEFA89B000FA83" box="[487,576,1377,1400]" italics="true" pageId="3" pageNumber="372">J. towai</emphasis>
</taxonomicName>
(
<figureCitation id="13692A15FFE6FFCF4B56FA88B0CDFA83" box="[591,653,1376,1400]" captionStart="FIGURES 8 – 10. 8" captionStartId="6.[151,264,1096,1119]" captionTargetBox="[163,1421,201,1057]" captionTargetId="figure@6.[157,1429,193,1075]" captionTargetPageId="6" captionText="FIGURES 8 – 10. 8) apical protarsus showing bifid anterior protarsal claw 9) aedeagus, ventral view 10) aedeagus, dorsal view." httpUri="https://zenodo.org/record/222181/files/figure.png" pageId="3" pageNumber="372">Fig 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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