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<document ID-DOI="http://doi.org/10.5281/zenodo.3788771" ID-GBIF-Dataset="6dfdec5c-2e9d-4e1f-be4c-a120edc779e0" ID-GBIF-Taxon="163747514" ID-ISSN="1313–2970" ID-Zenodo-Dep="3788771" ID-ZooBank="6D59834F-82C0-4DCD-8F65-202AE8F03965" LSID="urn:lsid:zoobank.org:act:F24788C7-BB3D-40B9-BFC7-6BA5D4370601" approvalRequired="61" approvalRequired_for_textStreams="61" checkinTime="1587954401351" checkinUser="plazi" docAuthor="Hawks, David" docDate="2010" docId="777587FF090CFFA6FF4356B3B2CEFAEF" docLanguage="en" docName="ZK_article_2166.pdf" docOrigin="ZooKeys 39 (39)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@7e86ff3b" docStyleName="zookeys.2008.journal_article" docTitle="Catocala chelidonia subsp. occidentalis Hawks 2010, ssp. n." docType="treatment" docUuid="F24788C7-BB3D-40B9-BFC7-6BA5D4370601" docUuidSource="ZooBank" docVersion="5" lastPageNumber="31" masterDocId="8B4CFF87091EFFB4FFCF5615B045FFA5" masterDocTitle="Review of the Catocala delilah species complex (Lepidoptera, Erebidae)" masterLastPageNumber="35" masterPageNumber="13" pageId="18" pageNumber="31" updateTime="1643557595621" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Catocala delilah species complex (Lepidoptera, Erebidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hawks, David</mods:namePart>
<mods:affiliation>University of California, Riverside ,, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-03-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>39</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>39</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>13</mods:start>
<mods:end>35</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.39.439</mods:identifier>
<mods:identifier type="GBIF-Dataset">6dfdec5c-2e9d-4e1f-be4c-a120edc779e0</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576631</mods:identifier>
<mods:identifier type="ZooBank">6D59834F-82C0-4DCD-8F65-202AE8F03965</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788771" ID-GBIF-Taxon="163747514" ID-Zenodo-Dep="3788771" LSID="urn:lsid:zoobank.org:act:F24788C7-BB3D-40B9-BFC7-6BA5D4370601" httpUri="http://treatment.plazi.org/id/777587FF090CFFA6FF4356B3B2CEFAEF" lastPageNumber="31" pageId="18" pageNumber="31">
<subSubSection pageId="18" pageNumber="31" type="nomenclature">
<paragraph blockId="18.[140,977,166,263]" box="[140,703,166,193]" pageId="18" pageNumber="31">
<heading box="[140,703,166,193]" fontSize="11" level="2" pageId="18" pageNumber="31" reason="3">
<emphasis bold="true" box="[140,703,166,193]" pageId="18" pageNumber="31">
<taxonomicName authority="Hawks" authorityName="Hawks" authorityYear="2010" box="[140,615,166,193]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="chelidonia" status="ssp. n." subSpecies="occidentalis">
<emphasis bold="true" box="[140,523,166,193]" italics="true" pageId="18" pageNumber="31">
<emphasis box="[140,246,166,193]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Catocala</emphasis>
chelidonia occidentalis
</emphasis>
Hawks
</taxonomicName>
,
<taxonomicNameLabel box="[627,703,167,193]" pageId="18" pageNumber="31" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[140,977,166,263]" box="[140,977,202,229]" pageId="18" pageNumber="31">
<uri box="[140,977,202,229]" pageId="18" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid box="[421,977,202,229]" pageId="18" pageNumber="31">F24788C7-BB3D-40B9-BFC7-6BA5D4370601</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,265,237,263]" pageId="18" pageNumber="31" type="description">
<paragraph blockId="18.[140,977,166,263]" box="[140,265,237,263]" pageId="18" pageNumber="31">Figs 23, 24</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="31" type="materials_examined">
<paragraph blockId="18.[140,1108,307,1355]" pageId="18" pageNumber="31">
<emphasis bold="true" box="[140,313,307,333]" pageId="18" pageNumber="31">
<emphasis box="[140,198,307,333]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608811621" collectingDate="1987-06-15" collectingDateMax="1987-06-22" collectingDateMin="1987-06-15" collectionCode="PMNH" collectorName="D. C. Hawks" county="Riverside County" location="Santa Rosa Mountains" municipality="Pinyon Flat" pageId="18" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
<emphasis bold="true" box="[322,435,307,333]" pageId="18" pageNumber="31">
<typeStatus box="[322,435,307,333]" pageId="18" pageNumber="31">Holotype</typeStatus>
</emphasis>
♁ (Fig. 23,
<collectionCode box="[579,668,308,334]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="18" pageNumber="31" type="Museum">PMNH</collectionCode>
specimen #ENT 721051; FW length
<quantity box="[140,227,343,369]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="31" unit="mm" value="20.0">20 mm</quantity>
),
<emphasis bold="true" box="[247,343,343,369]" pageId="18" pageNumber="31">
<typeStatus box="[247,343,343,369]" pageId="18" pageNumber="31">allotype</typeStatus>
</emphasis>
<specimenCount box="[350,365,343,369]" pageId="18" pageNumber="31" type="female">♀</specimenCount>
(Fig. 24,
<collectionCode box="[476,565,343,369]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="18" pageNumber="31" type="Museum">PMNH</collectionCode>
specimen #ENT 721052; FW length
<quantity box="[1001,1088,343,369]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="31" unit="mm" value="21.0">21 mm</quantity>
):
<collectingMunicipality box="[140,275,378,404]" pageId="18" pageNumber="31">Pinyon Flat</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA6FF4356B3B2CEFAEF:FA036032090CFFA6FEEF576FB266FE31" box="[288,547,378,404]" county="Riverside County" municipality="Pinyon Flat" name="Santa Rosa Mountains" pageId="18" pageNumber="31" stateProvince="California">Santa Rosa Mountains</location>
,
<collectingCounty box="[559,759,378,404]" pageId="18" pageNumber="31">Riverside County</collectingCounty>
,
<collectingRegion box="[771,889,378,404]" country="United States of America" name="California" pageId="18" pageNumber="31">California</collectingRegion>
, leg.
<collectorName box="[948,1103,378,404]" pageId="18" pageNumber="31">D. C. Hawks</collectorName>
;
<typeStatus box="[140,239,413,439]" pageId="18" pageNumber="31">holotype</typeStatus>
<date box="[248,407,413,440]" pageId="18" pageNumber="31" value="1987-06-15">
<collectingDate box="[248,407,413,440]" pageId="18" pageNumber="31" value="1987-06-15">15 June 1987</collectingDate>
</date>
,
<typeStatus box="[420,509,413,439]" pageId="18" pageNumber="31">allotype</typeStatus>
<date box="[518,678,413,440]" pageId="18" pageNumber="31" value="1987-06-22">
<collectingDate box="[518,678,413,440]" pageId="18" pageNumber="31" value="1987-06-22">22 June 1987</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2608811619" collectorName="Remaining" county="Kern County" location="10 km NW of Frazier Park" pageId="18" pageNumber="31" specimenCount="100" specimenCount-female="100" stateProvince="California" typeStatus="paratype">
<emphasis bold="true" box="[691,806,413,439]" pageId="18" pageNumber="31">
<typeStatus box="[691,806,413,439]" pageId="18" pageNumber="31">Paratypes</typeStatus>
</emphasis>
(25 ♁
<specimenCount box="[889,942,413,440]" pageId="18" pageNumber="31" type="female">53 ♀</specimenCount>
): 23 ♁
<specimenCount box="[1033,1086,413,440]" pageId="18" pageNumber="31" type="female">46 ♀</specimenCount>
from the type locality, various dates and collectors.
<collectorName box="[717,841,448,474]" pageId="18" pageNumber="31">Remaining</collectorName>
<typeStatus box="[849,959,449,475]" pageId="18" pageNumber="31">paratypes</typeStatus>
:
<collectingCounty pageId="18" pageNumber="31">Kern County</collectingCounty>
:
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA6FF4356B3B2CEFAEF:FA036032090CFFA6FF7D57F1B1A9FE58" box="[178,492,483,510]" county="Kern County" name="10 km NW of Frazier Park" pageId="18" pageNumber="31" stateProvince="California">
<quantity box="[178,254,483,510]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="31" unit="km" value="10.0">10 km</quantity>
NW of Frazier Park
</location>
(
<specimenCount box="[513,550,484,510]" pageId="18" pageNumber="31" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811632" county="Los Angeles County" location="San Gabriel Mountains" pageId="18" pageNumber="31" specimenCount="4" specimenCount-female="4" stateProvince="California" typeStatus="paratype">
<collectingCounty box="[576,812,483,510]" pageId="18" pageNumber="31">Los Angeles County</collectingCounty>
:
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA6FF4356B3B2CEFAEF:FA036032090CFFA6FCF657F1B40DFE58" box="[825,1096,483,510]" county="Los Angeles County" name="San Gabriel Mountains" pageId="18" pageNumber="31" stateProvince="California">San Gabriel Mountains</location>
,
<locationDeviation box="[140,412,519,545]" location="Valyermo" pageId="18" pageNumber="31">
<quantity box="[140,200,519,545]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="31" unit="km" value="4.0">4 km</quantity>
SSW of Valyermo
</locationDeviation>
(1 ♁
<specimenCount box="[480,518,519,545]" pageId="18" pageNumber="31" type="female">4 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811625" box="[543,1034,519,545]" location="Juniper Hills" pageId="18" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA6FF4356B3B2CEFAEF:FA036032090CFFA6FDD05412B2F1FD84" box="[543,692,519,545]" name="Juniper Hills" pageId="18" pageNumber="31" stateProvince="California">Juniper Hills</location>
,
<locationDeviation box="[705,967,519,545]" location="Pearblossom" pageId="18" pageNumber="31">
<quantity box="[705,765,519,545]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="31" unit="km" value="3.0">3 km</quantity>
S of Pearblossom
</locationDeviation>
(
<specimenCount box="[987,1023,519,545]" pageId="18" pageNumber="31" type="female">1 ♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811637" collectorName="Halfway House" location="San Gabriel Mountains" municipality="Ridge Route" pageId="18" pageNumber="31" specimenCount="1" stateProvince="California" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA6FF4356B3B2CEFAEF:FA036032090CFFA6FBD75412B123FDE1" municipality="Ridge Route" name="San Gabriel Mountains" pageId="18" pageNumber="31" stateProvince="California">San Gabriel Mountains</location>
,
<collectingMunicipality box="[369,513,554,580]" pageId="18" pageNumber="31">Ridge Route</collectingMunicipality>
,
<collectorName box="[524,699,554,580]" pageId="18" pageNumber="31">Halfway House</collectorName>
(l ♁)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608811626" county="San Diego County" location="6 km S of Oak Grove" pageId="18" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
<collectingCounty box="[771,985,554,580]" pageId="18" pageNumber="31">San Diego County</collectingCounty>
:
<location LSID="urn:lsid:plazi:treatment:777587FF090CFFA6FF4356B3B2CEFAEF:FA036032090CFFA6FC2C543FB14CFDC2" county="San Diego County" name="6 km S of Oak Grove" pageId="18" pageNumber="31" stateProvince="California">
<quantity box="[995,1053,554,580]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="31" unit="km" value="6.0">6 km</quantity>
S of Oak Grove
</location>
(
<specimenCount box="[284,319,589,616]" pageId="18" pageNumber="31" type="female">1 ♀</specimenCount>
)
</materialsCitation>
.
<typeStatus box="[342,449,589,615]" pageId="18" pageNumber="31">Holotype</typeStatus>
and
<typeStatus box="[506,595,589,615]" pageId="18" pageNumber="31">allotype</typeStatus>
deposited at
<collectionCode box="[746,838,590,616]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="18" pageNumber="31" type="Museum">PMNH</collectionCode>
,
<typeStatus box="[850,956,590,616]" pageId="18" pageNumber="31">paratypes</typeStatus>
deposited at
<collectionCode box="[140,222,624,650]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="18" pageNumber="31" type="Museum">LACM</collectionCode>
and other
<collectingCountry box="[345,400,624,650]" name="United States of America" pageId="18" pageNumber="31">USA</collectingCountry>
museums as well as in private collections.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="31" type="diagnosis">
<paragraph blockId="18.[140,1108,307,1355]" pageId="18" pageNumber="31">
<emphasis bold="true" box="[185,312,659,685]" pageId="18" pageNumber="31">
<emphasis box="[185,307,659,685]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Diagnosis</emphasis>
.
</emphasis>
The forewings of
<emphasis box="[516,535,660,686]" italics="true" pageId="18" pageNumber="31">C</emphasis>
.
<emphasis box="[549,559,660,685]" italics="true" pageId="18" pageNumber="31">c</emphasis>
.
<emphasis box="[574,697,659,685]" italics="true" pageId="18" pageNumber="31">occidentalis</emphasis>
are similar to those of other subspecies. Distinguishing hindwing characters include: dorsal surface has many black scales at the base; wider inner band; narrower postmedial band; yellowish color of the dorsal surface consistently darker. About 5 percent of the specimens of
<emphasis box="[854,873,766,792]" italics="true" pageId="18" pageNumber="31">C</emphasis>
.
<emphasis box="[886,896,766,791]" italics="true" pageId="18" pageNumber="31">c</emphasis>
.
<emphasis box="[910,1033,765,791]" italics="true" pageId="18" pageNumber="31">occidentalis</emphasis>
examined could be confused with specimens of the nominate subspecies.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="31" type="description">
<paragraph blockId="18.[140,1108,307,1355]" pageId="18" pageNumber="31">
<emphasis bold="true" box="[185,335,835,861]" pageId="18" pageNumber="31">
<emphasis box="[185,330,835,861]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[347,447,836,861]" italics="true" pageId="18" pageNumber="31">Forewing</emphasis>
: dorsal surface ground color charcoal to brownish gray, mottled with hoary scales; antemedial and postmedial lines black, modestly apparent; subreniform whitish to yellow, usually visible.
<emphasis box="[657,769,906,932]" italics="true" pageId="18" pageNumber="31">Hindwing</emphasis>
: pale areas dull yellow orange; bands black, apical spot dull yellow orange; anal spot separated from outer band; conspicuously narrow postmedial band (usually only
<quantity box="[694,790,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="18" pageNumber="31" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
near vein M2); anterior half of outer band wide with conspicuous rounded indentation in inner edge of outer band between M2 and Cu1; many dark scales at base.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="31" type="distribution">
<paragraph blockId="18.[140,1108,307,1355]" pageId="18" pageNumber="31">
<emphasis bold="true" box="[185,491,1081,1108]" pageId="18" pageNumber="31">
<emphasis box="[185,336,1081,1107]" inLineHeading="true" pageId="18" pageNumber="31" reason="1">Distribution</emphasis>
and biology.
</emphasis>
<emphasis box="[498,611,1082,1108]" italics="true" pageId="18" pageNumber="31">
<taxonomicName authorityName=", Barnes and McDunnough" authorityYear="1918" box="[498,593,1082,1108]" class="Insecta" family="Noctuidae" genus="Catocala" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="31" phylum="Arthropoda" rank="genus">Catocala</taxonomicName>
c
</emphasis>
.
<emphasis box="[625,748,1082,1108]" italics="true" pageId="18" pageNumber="31">occidentalis</emphasis>
occurs along the western desert edge in southern
<collectingRegion box="[336,450,1117,1143]" country="United States of America" name="California" pageId="18" pageNumber="31">California</collectingRegion>
and northward to at least Trinity County. County records for
<collectingCountry box="[180,235,1152,1178]" name="United States of America" pageId="18" pageNumber="31">USA</collectingCountry>
are as follows.
<collectingRegion box="[405,583,1152,1178]" country="United States of America" name="California" pageId="18" pageNumber="31">CALIFORNIA</collectingRegion>
: Kern, Lake, Los Angeles, Madera, Napa, Riverside, San Bernardino, San Diego, Santa Clara, Solano, Trinity.. The immature stages of
<emphasis box="[168,187,1223,1249]" italics="true" pageId="18" pageNumber="31">C</emphasis>
.
<emphasis box="[201,211,1223,1248]" italics="true" pageId="18" pageNumber="31">c</emphasis>
.
<emphasis box="[224,347,1223,1249]" italics="true" pageId="18" pageNumber="31">occidentalis</emphasis>
have been described and the larva figured by
<bibRefCitation author="Johnson JW" box="[849,1029,1222,1249]" pageId="18" pageNumber="31" pagination="303 - 327" refId="ref10573" refString="Johnson JW (1985) Th e immature stages of six California Catocala (Lepidoptera: Noctuidae). Journal of Research on the Lepidoptera 23: 303 - 327." type="journal article" year="1985">Johnson (1985)</bibRefCitation>
. At the San Diego, Riverside, and San Bernardino County localities the larvae feed on
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="18" pageNumber="31" phylum="Tracheophyta" rank="species" species="cornelius-mulleri">
<emphasis italics="true" pageId="18" pageNumber="31">Quercus cornelius-mulleri</emphasis>
</taxonomicName>
. Adults have been collected from early June to late August, with the peak flight period in late June at Pinyon Flat.
</paragraph>
</subSubSection>
</treatment>
</document>
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