treatments-xml/data/C2/73/87/C27387A7FFE0FFCF9A050201FB0A7162.xml

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<mods:title id="6503AE5144DA0EBA28E707FF6918F7BA">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:namePart id="D203A3659A833667FAFBFE68C77B28E2">Kamdńskd, Marcdn J.</mods:namePart>
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<mods:date id="B737F4E2972A53B8783BB54CA5D21A74">2024</mods:date>
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<paragraph id="4A6536B1FFE0FFCF9A050201FD767058" blockId="18.[113,762,1852,2009]" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9A050201FD767058" box="[119,757,1852,1879]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
<emphasis id="78AEEAA3FFE0FFCF9A050201FD767058" bold="true" box="[119,757,1852,1879]" pageId="18" pageNumber="376">
2. 
<taxonomicName id="8DDA4D32FFE0FFCF9AFC0201FDAB7059" authority="(Mulsant and Rey)" baseAuthorityName="Mulsant and Rey" box="[142,552,1852,1879]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9AFC0201FEE17059" bold="true" box="[142,354,1852,1879]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
(Mulsant and Rey)
</taxonomicName>
stat. res.,comb. nov.
</emphasis>
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<paragraph id="4A6536B1FFE0FFCF9B290246FD92709A" blockId="18.[113,762,1852,2009]" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
<heading id="112D81DDFFE0FFCF9B290246FD92709A" box="[347,529,1915,1941]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
<emphasis id="78AEEAA3FFE0FFCF9B290246FD92709A" bold="true" box="[347,529,1915,1941]" pageId="18" pageNumber="376">
(
<figureCitation id="D2E12A34FFE0FFCF9B140246FE30709A" box="[358,435,1915,1941]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." figureDoi="http://doi.org/10.5281/zenodo.13315277" httpUri="https://zenodo.org/record/13315277/files/figure.png" pageId="18" pageNumber="376">Figs 2B</figureCitation>
, 
<figureCitation id="D2E12A34FFE0FFCF9BCF0246FE5C709A" box="[445,479,1915,1941]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." figureDoi="http://doi.org/10.5281/zenodo.13315289" httpUri="https://zenodo.org/record/13315289/files/figure.png" pageId="18" pageNumber="376">6G</figureCitation>
, 
<figureCitation id="D2E12A34FFE0FFCF9B9B0246FD85709A" box="[489,518,1915,1941]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13315291" httpUri="https://zenodo.org/record/13315291/files/figure.png" pageId="18" pageNumber="376">8B</figureCitation>
)
</emphasis>
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</paragraph>
<paragraph id="4A6536B1FFE0FFCF9A03029FFD7970D6" blockId="18.[113,762,1852,2009]" pageId="18" pageNumber="376">
<pageTitle id="0A45EED6FFE0FFCF9A03029FFD7970D6" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9A03029FFED670B6" box="[113,341,1954,1977]" italics="true" pageId="18" pageNumber="376">Eurynotus asperipennis</emphasis>
Mulsant and Rey, 1854a: 23.– Mulsant and Rey 1854b: 169; Lacordadre 1859: 239; Gebden 1938: 414;
</pageTitle>
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<paragraph id="4A6536B1FFE0FFCF995805ADFC1B77C8" blockId="18.[810,1459,144,199]" pageId="18" pageNumber="376">
Koch 1954a: 280; 1954b: 2; 
<bibRefCitation id="2E4B4B40FFE0FFCF9E2205ADFAC477A7" author="Banaszkdewdcz M" box="[1104,1351,144,168]" pageId="18" pageNumber="376" pagination="59 - 77" refId="ref25294" refString="Banaszkdewdcz M. Comparatdve study of female gendtalda dn Peddndnd sens Iwan 2004 (Coleoptera: ºenebrdonddae: Peddndnd %, wdth notes on the classdficatdon. Annales Zoologici 2006; 56: 59 - 77." type="journal article" year="2006">Banaszkdewdcz 2006: 63</bibRefCitation>
; Kamdńskd 2016: 235.
</paragraph>
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<emphasis id="78AEEAA3FFE0FFCF995805D4FC13760F" box="[810,912,233,256]" italics="true" pageId="18" pageNumber="376">Diagnosis:</emphasis>
Lacks sclerdte dn bursa copulatrdx (present dn 
<taxonomicName id="8DDA4D32FFE0FFCF9FED05D4FC1E762F" authorityName="Koch" authorityYear="1954" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
<emphasis id="78AEEAA3FFE0FFCF9FED05D4FC1E762F" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="8DDA4D32FFE0FFCF99920434FBE9762F" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[992,1130,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF99920434FBE9762F" box="[992,1130,264,288]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%. 
<taxonomicName id="8DDA4D32FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9EF80435FACF762F" box="[1162,1356,264,288]" italics="true" pageId="18" pageNumber="376">Biolus asperipennis</emphasis>
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has two puncture 
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on ventrdte V (sdngle 
<typeStatus id="95618813FFE0FFCF9EC80415FB6B764F" box="[1210,1256,296,320]" pageId="18" pageNumber="376">type</typeStatus>
dn 
<taxonomicName id="8DDA4D32FFE0FFCF9F7D0415FAE77630" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1295,1380,296,319]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F7D0415FAE77630" box="[1295,1380,296,319]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
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and 
<taxonomicName id="8DDA4D32FFE0FFCF9FED0415FC127650" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="78AEEAA3FFE0FFCF9FED0415FC127650" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
</taxonomicName>
%, and golden setae on the tarsd (dark/black dn 
<taxonomicName id="8DDA4D32FFE0FFCF9F2F047AFA2E7651" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[1373,1453,327,350]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
<emphasis id="78AEEAA3FFE0FFCF9F2F047AFA2E7651" box="[1373,1453,327,350]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="8DDA4D32FFE0FFCF9958045AFC1C7671" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[810,927,358,382]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
<emphasis id="78AEEAA3FFE0FFCF9958045AFC1C7671" box="[810,927,358,382]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8DDA4D32FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF99A5045AFBB07671" box="[983,1075,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
%. Wdthout close dnspectdon, 
<taxonomicName id="8DDA4D32FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9F2A045AFA307671" box="[1368,1459,358,382]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
(length 
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; wddth 
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% specdmens may appear sdmdlar to smaller specdmens of 
<taxonomicName id="8DDA4D32FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0F0498FA9176B3" box="[1149,1298,421,444]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
(length 
<quantity id="8D229B54FFE0FFCF9F030498FCD576D4" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="18" pageNumber="376" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">19–22 mm</quantity>
; wddth 
<quantity id="8D229B54FFE0FFCF99ED04F9FC7D76D3" box="[927,1022,452,476]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="376" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
%; however, 
<taxonomicName id="8DDA4D32FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9E0504F8FA8476D3" box="[1143,1287,453,476]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
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has golden tarsal setae (black or dark dn 
<taxonomicName id="8DDA4D32FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E7D04D9FBEA76F4" box="[1039,1129,483,507]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
%, a weaker cdrcumocular depres* sdon, the pronotal process ds more pronounced, and the male tdbdae are also at most only sldghtly moddfied dn 
<taxonomicName id="8DDA4D32FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9E8D071EFADA7535" box="[1279,1369,546,570]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
, whdle dn 
<taxonomicName id="8DDA4D32FFE0FFCF9958077FFC387556" box="[810,955,578,601]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
<emphasis id="78AEEAA3FFE0FFCF9958077FFC387556" box="[810,955,578,601]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
</taxonomicName>
they are more strongly curved. See the ddagnosds of 
<taxonomicName id="8DDA4D32FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="brunoi">
<emphasis id="78AEEAA3FFE0FFCF9936075CFC1D7577" box="[836,926,608,632]" italics="true" pageId="18" pageNumber="376">B. brunoi</emphasis>
</taxonomicName>
below for more detadls.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF995807A7FC6D720A" blockId="18.[810,1460,666,1285]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF995807A7FC3E75BD" box="[810,957,666,690]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
Length 
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, wddth 
<quantity id="8D229B54FFE0FFCF9E8907A7FADC75BD" box="[1275,1375,666,690]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="18" pageNumber="376" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11mm</quantity>
. Head: Punctate; punctures closely spaced, separated by ≤1 punc* ture ddameter. Mentum wdthout apdcal notch. Eye wdth weakly dmpressed sulcus sdtuated around perdmeter of dorsal lobe. Prothorax: Pronotum punctate, punctures separated by ≥1 puncture ddameter. Hypomeron ldghtly sculptured/ wrdnkled wdth small punctatdon and mdcrotuberculatdon. Pterothorax: Elytra not costate, wdthout macrotubercles on ddsc. Elytral dntervals wdth or wdthout mdcrotuberculatdon. Epdpleuron wdth many small, fadnt mdcrotubercles. Abdomen: Fdnely punctate, punctures separated by&gt;1 puncture ddam* eter. Abdomdnal ventrdte V punctures separated by ≥ 1 punc* ture ddameter wdth larger, setdgerous punctures dntermdxed, females wdth meddan, apdcal notch. Legs: Male hdnd tdbdae sldghtly flattened/twdsted, appeardng bent. Female legs un* moddfded. ºarsd wdth golden setae. Males: pro/meso tarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres ‘bottle*shaped’, strongly tapered and truncate ap* dcally. Female termdnalda: Bursa copulatrdx wdthout accessory pouch or sclerdtes.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF9958001BFA267272" blockId="18.[810,1460,1318,1436]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF9958001BFC767231" box="[810,1013,1318,1342]" italics="true" pageId="18" pageNumber="376">Material examined:</emphasis>
<typeStatus id="95618813FFE0FFCF9E75001AFBF27230" box="[1031,1137,1319,1343]" pageId="18" pageNumber="376" type="lectotype">Lectotype</typeStatus>
, here desdgnated (MNHN%: ‘ 
<taxonomicName id="8DDA4D32FFE0FFCF995C007BFC1F7251" box="[814,924,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
? sp. cap.?; blue cdrcle label ‘ 
<taxonomicName id="8DDA4D32FFE0FFCF9ECF007BFAA87251" box="[1213,1323,1350,1374]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
asperdpennds Muls cap. b. sp.?’ also wdth a red label wdth handwrdtdng: ‘ºype’.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF993700B9FADD7293" blockId="18.[810,1460,1318,1436]" box="[837,1374,1412,1436]" pageId="18" pageNumber="376">(See Appenddx S1 for adddtdonal materdal examdned.%</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580083FA8D713B" blockId="18.[810,1459,1470,1588]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580083FCE272DA" box="[810,865,1470,1493]" italics="true" pageId="18" pageNumber="376">Note:</emphasis>
As the ordgdnal descrdptdon lacks dnformatdon defindng the number or specdficdty of specdmens used, one from the presumed 
<typeStatus id="95618813FFE0FFCF995800C0FCF8711A" box="[810,891,1533,1557]" pageId="18" pageNumber="376" type="syntype">syntype</typeStatus>
serdes ds here desdgnated as the 
<typeStatus id="95618813FFE0FFCF9EC900C1FA98711B" box="[1211,1307,1532,1556]" pageId="18" pageNumber="376" type="lectotype">lectotype</typeStatus>
to fix the taxo* nomdc status of the specdes and genus (Fdg. 2B%.
</paragraph>
<paragraph id="4A6536B1FFE0FFCF99580368FB0A7162" blockId="18.[810,1161,1621,1645]" box="[810,1161,1621,1645]" pageId="18" pageNumber="376">
<emphasis id="78AEEAA3FFE0FFCF99580368FC2B7162" box="[810,936,1621,1645]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
South Afrdca (Fdg. 9%.
</paragraph>
</subSubSection>
</treatment>
</document>