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225 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.867.28728" ID-GBIF-Dataset="bff0bb63-a934-4fa2-ae76-50c4db24d98d" ID-PMC="PMC6685932" ID-Pensoft-Pub="1313-2970-867-139" ID-Pensoft-UUID="729147F7504F521DB514FA3017CB4CD3" ID-PubMed="31406481" ID-ZooBank="9765595E1A204418B5C6AC4B6F587CE5" ModsDocID="1313-2970-867-139" checkinTime="1564648495948" checkinUser="pensoft" docAuthor="Goldstein, Paul Z. & Zilli, Alberto" docDate="2019" docId="D3B78251E7015C3E9BD09793B275D12C" docLanguage="en" docName="ZooKeys 867: 139-160" docOrigin="ZooKeys 867" docSource="http://dx.doi.org/10.3897/zookeys.867.28728" docTitle="Thraumata peruviensia Goldstein & Zilli, 2019, sp. nov." docType="treatment" docVersion="4" lastPageNumber="139" masterDocId="414CFF867946142B7D60E5509002FFFB" masterDocTitle="Thraumata, a new genus from South America with description of a new species from Peru (Lepidoptera, Noctuidae)" masterLastPageNumber="160" masterPageNumber="139" pageNumber="139" updateTime="1668167635196" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Thraumata, a new genus from South America with description of a new species from Peru (Lepidoptera, Noctuidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Goldstein, Paul Z.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zilli, Alberto</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>867</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>139</mods:start>
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<mods:end>160</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.867.28728</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.867.28728</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-867-139</mods:identifier>
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<mods:identifier type="ZooBank">9765595E1A204418B5C6AC4B6F587CE5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">729147F7504F521DB514FA3017CB4CD3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159293021" LSID="urn:lsid:plazi:treatment:D3B78251E7015C3E9BD09793B275D12C" httpUri="http://treatment.plazi.org/id/D3B78251E7015C3E9BD09793B275D12C" lastPageNumber="139" pageId="0" pageNumber="139">
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<subSubSection pageId="0" pageNumber="139" type="nomenclature">
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<paragraph pageId="0" pageNumber="139">
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<taxonomicName LSID="d3b78251-e701-5c3e-9bd0-9793b275d12c" class="Insecta" family="Noctuidae" genus="Thraumata" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraumata peruviensia" order="Lepidoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="peruviensia">Thraumata peruviensia</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="139">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Dorsal habitus of Thraumata species. 1 T. subvenata Holotype ♂, Guyana, USNMENT 00973242 2 T. subvenata, USNMENT 01440366 ♀, Suriname 3 T. petrovna ♂ USNMENT 01440358, Brazil 4 T. petrovna Holotype ♀ USNMENT 00973124, Costa Rica 5 T. peruviensia Holotype ♂ USNMENT 01440371, Peru 6 T. peruviensia ♀ USNMENT 01440374, Peru 7 - 9 Holotype specimen labels of Thraumata species 7 Phuphena subvenata Schaus 8 Leptina petrovna Schaus 9 Thraumata peruviensia sp. nov." figureDoi="10.3897/zookeys.867.28728.figures1-9" httpUri="https://binary.pensoft.net/fig/322322" pageId="0" pageNumber="139">Figs 5</figureCitation>
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,
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<figureCitation captionStart="Figures 1–9" captionStartId="F1" captionText="Figures 1 - 9. Dorsal habitus of Thraumata species. 1 T. subvenata Holotype ♂, Guyana, USNMENT 00973242 2 T. subvenata, USNMENT 01440366 ♀, Suriname 3 T. petrovna ♂ USNMENT 01440358, Brazil 4 T. petrovna Holotype ♀ USNMENT 00973124, Costa Rica 5 T. peruviensia Holotype ♂ USNMENT 01440371, Peru 6 T. peruviensia ♀ USNMENT 01440374, Peru 7 - 9 Holotype specimen labels of Thraumata species 7 Phuphena subvenata Schaus 8 Leptina petrovna Schaus 9 Thraumata peruviensia sp. nov." figureDoi="10.3897/zookeys.867.28728.figures1-9" httpUri="https://binary.pensoft.net/fig/322322" pageId="0" pageNumber="139">6</figureCitation>
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,
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<figureCitation captionStart="Figures 10–15" captionStartId="F2" captionText="Figures 10 - 15. Ventral habitus of Thraumata species. 10 T. subvenata Holotype ♂, Peru, NHMUK 010606200 11 T. subvenata, USNMENT 01440366 ♀, Suriname 12 T. petrovna ♂ USNMENT 01440358, Brazil 13 T. petrovna Holotype ♀ USNMENT 00973124, Costa Rica 14 T. peruviensia Holotype ♂ USNMENT 01440371, Peru 15 T. peruviensia ♀ USNMENT 01440374, Peru." figureDoi="10.3897/zookeys.867.28728.figures10-15" httpUri="https://binary.pensoft.net/fig/322323" pageId="0" pageNumber="139">14</figureCitation>
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,
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<figureCitation captionStart="Figures 10–15" captionStartId="F2" captionText="Figures 10 - 15. Ventral habitus of Thraumata species. 10 T. subvenata Holotype ♂, Peru, NHMUK 010606200 11 T. subvenata, USNMENT 01440366 ♀, Suriname 12 T. petrovna ♂ USNMENT 01440358, Brazil 13 T. petrovna Holotype ♀ USNMENT 00973124, Costa Rica 14 T. peruviensia Holotype ♂ USNMENT 01440371, Peru 15 T. peruviensia ♀ USNMENT 01440374, Peru." figureDoi="10.3897/zookeys.867.28728.figures10-15" httpUri="https://binary.pensoft.net/fig/322323" pageId="0" pageNumber="139">15</figureCitation>
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<figureCitation captionStart="Figures 18–22" captionStartId="F5" captionText="Figures 18 - 22. Thraumata male genitalia. 18 - 20 T. petrovna dissection 148363, USNMENT 01422625, Brazil 18 Apparatus 19, 20 Phallus 21, 22 T. peruviensia, dissection 148129, USNMENT 01440371 21 Apparatus 22 Phallus." figureDoi="10.3897/zookeys.867.28728.figures18-22" httpUri="https://binary.pensoft.net/fig/322326" pageId="0" pageNumber="139">(habitus); Figs 21</figureCitation>
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,
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<figureCitation captionStart="Figures 18–22" captionStartId="F5" captionText="Figures 18 - 22. Thraumata male genitalia. 18 - 20 T. petrovna dissection 148363, USNMENT 01422625, Brazil 18 Apparatus 19, 20 Phallus 21, 22 T. peruviensia, dissection 148129, USNMENT 01440371 21 Apparatus 22 Phallus." figureDoi="10.3897/zookeys.867.28728.figures18-22" httpUri="https://binary.pensoft.net/fig/322326" pageId="0" pageNumber="139">22</figureCitation>
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<figureCitation captionStart="Figures 25, 26" captionStartId="F7" captionText="Figures 25, 26. Thraumata male abdomens. 25 T. peruviensia dissection 148134, USNMENT 01440385 26 T. peruviensia dissection 148129, USNMENT 01440371, Brazil." figureDoi="10.3897/zookeys.867.28728.figures25-26" httpUri="https://binary.pensoft.net/fig/322328" pageId="0" pageNumber="139">(male genitalia); Fig. 26</figureCitation>
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<figureCitation captionStart="Figures 28, 29" captionStartId="F9" captionText="Figures 28, 29. Thraumata female genitalia. 28 T. peruviensia dissection 148130, USNMENT 01440374 USNM, Brazil 29 T. petrovna dissection 148127, USNMENT 01440363, Peru." figureDoi="10.3897/zookeys.867.28728.figures28-29" httpUri="https://binary.pensoft.net/fig/322330" pageId="0" pageNumber="139">(male abdomen); Fig. 28 (female genitalia)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="139" type="materials_examined">
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<paragraph pageId="0" pageNumber="139">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="139">
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<emphasis bold="true" pageId="0" pageNumber="139">Type material</emphasis>
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.
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<emphasis bold="true" pageId="0" pageNumber="139">PERU</emphasis>
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(11♂, 6♀):
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<emphasis bold="true" pageId="0" pageNumber="139">Holotype</emphasis>
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♂ (USNM), Oconeque Carahaya Perou S. E.; Dognin Collection; USNMENT01440384.
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<emphasis bold="true" pageId="0" pageNumber="139">Paratypes</emphasis>
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(10♂, 6♀; USNM, NHMUK, AMNH):
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<emphasis italics="true" pageId="0" pageNumber="139">Males</emphasis>
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: same data as holotype, Dissection 148129, USNMENT01440371, BOLD Process ID LNAUW3137-17; same data as holotype, USNMENT01440341; Quinton, Carabaya, Peru, 5000 ft, i 1905. G. Ockenden [Joicey Bequest Brit. Mus. 1934-120] [NHMUK]; 2♂ Oconeque, Carabaya, Peru. 5000 ft, ii 1905, G. Ockenden; 1♂, Joicey Bequest Brit. Mus. 1934-120; 1♂, Rothschild Bequest B.M. 1939-1 [NHMUK]; 2♂ Oconeque, Carabaya, 7000 ft., dry s., July 1904 (G. Ockenden), Rothschild Bequest B.M. 1939-1 [NHMUK]; 3♂ Oconeque S. E. Peru,
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<normalizedToken originalValue="Feb’">Feb'</normalizedToken>
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05 7000 ft, G. Ockenden [2♂, 67. 20. Ex Coll. Ed. Brabant 1920 / Rothschild Bequest B.M. 1939-1; 1♂ - 1909-89. [NHMUK].
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<emphasis italics="true" pageId="0" pageNumber="139">Females</emphasis>
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: same data as holotype, Dissection 148130, USNMENT01440374; PERU: Cusco Machu Picchu 2885 m 5.II.1959 J.F.G. Clarke, USNMENT01440443; Macchu Pichu [sic] Peru II-11 1966 B. Heineman [AMNH]; Oconeque, Carabaya, Peru. 5000 ft, ii 1905, G. Ockenden; Joicey Bequest Brit. Mus. 1934-120 [NHMUK]; Oconeque, Carabaya, 7000 ft., dry s., July 1904 (G. Ockenden), Rothschild Bequest B.M. 1939-1 [NHMUK]; Oconeque S. E. Peru
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<normalizedToken originalValue="Feb’">Feb'</normalizedToken>
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05, 7000 ft G. Ockenden, 67. 20. Ex Coll. Ed. Brabant 1920/Rothschild Bequest B.M. 1939-1 [NHMUK]
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</paragraph>
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<caption doi="10.3897/zookeys.867.28728.figure16" httpUri="https://binary.pensoft.net/fig/322324" pageId="0" pageNumber="139" start="Figure 16" startId="F3">
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<paragraph pageId="0" pageNumber="139">
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Figure 16. Known distribution of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Thraumata" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraumata" order="Lepidoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="139">Thraumata</emphasis>
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</taxonomicName>
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species.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="139" type="diagnosis">
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<paragraph pageId="0" pageNumber="139">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="139">
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<taxonomicName class="Insecta" family="Noctuidae" genus="Thraumata" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraumata peruviensia" order="Lepidoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="peruviensia">
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<emphasis italics="true" pageId="0" pageNumber="139">Thraumata peruviensia</emphasis>
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</taxonomicName>
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is most readily distinguished from the other two species by the combination of its larger size and distinctive vesica. In this and the previous species, the dominant ground color of forewing is gray, darker in the present species than in
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<taxonomicName lsidName="T. petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
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<emphasis italics="true" pageId="0" pageNumber="139">T. petrovna</emphasis>
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</taxonomicName>
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. This feature, the less pronounced white lining of pattern elements, particularly in basal and medial fields of forewing, and the comparatively reduced abdominal scent tufts in the male most readily distinguish this species pair from the more uniformly brown ground coloration of
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<taxonomicName lsidName="T. subvenata" pageId="0" pageNumber="139" rank="species" species="subvenata">
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<emphasis italics="true" pageId="0" pageNumber="139">T. subvenata</emphasis>
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</taxonomicName>
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. In the new species the triangular tooth projecting from the dark brown distal fascia of the forewing is longer than in
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<taxonomicName lsidName="T. petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
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<emphasis italics="true" pageId="0" pageNumber="139">T. petrovna</emphasis>
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</taxonomicName>
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, and there are at least two distinct white streaks on the hindwing underside instead of one.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="139" type="description">
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<paragraph pageId="0" pageNumber="139">Description.</paragraph>
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<paragraph pageId="0" pageNumber="139">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Head.</emphasis>
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Vestiture as in
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<taxonomicName lsidName="T. petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
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<emphasis italics="true" pageId="0" pageNumber="139">T. petrovna</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Thorax</emphasis>
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. Vestiture a mix of grayish-brown scales intermixed with white, especially within tegulae.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Wings.</emphasis>
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Forewing lengths: males (including holotype 15.3 mm) 14.2-15.3 mm (
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<emphasis italics="true" pageId="0" pageNumber="139">N</emphasis>
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= 3,
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<emphasis italics="true" pageId="0" pageNumber="139">x</emphasis>
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¯ = 15.0 mm, M = 12.5 mm); females, 15.5-16.8 mm (
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<emphasis italics="true" pageId="0" pageNumber="139">N</emphasis>
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= 3,
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<emphasis italics="true" pageId="0" pageNumber="139">x</emphasis>
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¯ = 16.1 mm, M = 16.1 mm). Forewing coloration dominated by gray scaling suffused with white, lines for the most part obscured, although in some individuals antemedial and postmedial lines still discernible as faint, thin pale gray lines. Four areas of more uniform, chocolate-brown scaling: an elongate, obtuse subtriangular smudge in lower basal field from wing base to antemedial line; an oblique band in place of median shading between from reniform stigma and inner margin, dark filling between orbicular and reniform stigmata, and in fascia bearing outwardly projected median tooth of distal field, this fascia lined externally with silvery white. Forewing underside nearly uniform chocolate brown excepting paler white inner marginal area; the shading on the hindwing underside interrupted by two-three conspicuous pale streaks, fading towards wing base.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Legs.</emphasis>
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White outward, otherwise as brown as in previous species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Abdomen</emphasis>
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. Abdominal segments predominantly gray, ringed with pale cream-colored scales distally; abdominal brushes lacking.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Male genitalia</emphasis>
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. Sacculus squared. Sclerotized lateral band on phallus less pronounced than in
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<taxonomicName lsidName="petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
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<emphasis italics="true" pageId="0" pageNumber="139">petrovna</emphasis>
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</taxonomicName>
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, vesica with three rudimentary subbasal diverticula and reduced para-basal plate from distal end of the phallus.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="139">Female genitalia</emphasis>
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. Ductus bursae quite short relative to distended corpus bursae. Anterior and posterior apophyses roughly co-equal. Papillae anales densely setose. As in other
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<taxonomicName class="Insecta" family="Noctuidae" genus="Thraumata" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraumata" order="Lepidoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="139">Thraumata</emphasis>
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</taxonomicName>
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, the ductus seminalis arises from a bulge at the posterior end of the corpus bursae.
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</paragraph>
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<caption doi="10.3897/zookeys.867.28728.figure17" httpUri="https://binary.pensoft.net/fig/322325" pageId="0" pageNumber="139" start="Figure 17" startId="F4">
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<paragraph pageId="0" pageNumber="139">
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Figure 17. Wing venation of
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<taxonomicName lsidName="T. petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
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<emphasis italics="true" pageId="0" pageNumber="139">T. petrovna</emphasis>
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</taxonomicName>
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, male, USNMENT01440439, wing preparation 148365.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="139" type="immature stages">
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<paragraph pageId="0" pageNumber="139">Immature stages.</paragraph>
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<paragraph pageId="0" pageNumber="139">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="139" type="etymology">
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<paragraph pageId="0" pageNumber="139">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="139">
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The name
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<taxonomicName lsidName="peruviensia" pageId="0" pageNumber="139" rank="species" species="peruviensia">
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<emphasis italics="true" pageId="0" pageNumber="139">peruviensia</emphasis>
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</taxonomicName>
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is a neuter pleural adjective, in reference to the country of origin of the new species, in coordination with the generic name
|
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<taxonomicName class="Insecta" family="Noctuidae" genus="Thraumata" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraumata" order="Lepidoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="139">Thraumata</emphasis>
|
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</taxonomicName>
|
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.
|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="139" type="distribution">
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<paragraph pageId="0" pageNumber="139">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="139">Southeastern Peru.</paragraph>
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="139" type="remarks">
|
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<paragraph pageId="0" pageNumber="139">Remarks.</paragraph>
|
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<paragraph pageId="0" pageNumber="139">
|
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This species is very similar in appearance to
|
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<taxonomicName lsidName="T. petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
|
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|
|
<emphasis italics="true" pageId="0" pageNumber="139">T. petrovna</emphasis>
|
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</taxonomicName>
|
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|
|
but consistently larger and occurring at higher altitudes. An incomplete barcode sequence (658bp - 88n = 570bp) was generated from a 1905 male specimen, USNMENT01440371 corresponding to USNM dissection 148129 (
|
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<figureCitation captionStart="Figures 18–22" captionStartId="F5" captionText="Figures 18 - 22. Thraumata male genitalia. 18 - 20 T. petrovna dissection 148363, USNMENT 01422625, Brazil 18 Apparatus 19, 20 Phallus 21, 22 T. peruviensia, dissection 148129, USNMENT 01440371 21 Apparatus 22 Phallus." figureDoi="10.3897/zookeys.867.28728.figures18-22" httpUri="https://binary.pensoft.net/fig/322326" pageId="0" pageNumber="139">Figs 21</figureCitation>
|
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,
|
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|
<figureCitation captionStart="Figures 18–22" captionStartId="F5" captionText="Figures 18 - 22. Thraumata male genitalia. 18 - 20 T. petrovna dissection 148363, USNMENT 01422625, Brazil 18 Apparatus 19, 20 Phallus 21, 22 T. peruviensia, dissection 148129, USNMENT 01440371 21 Apparatus 22 Phallus." figureDoi="10.3897/zookeys.867.28728.figures18-22" httpUri="https://binary.pensoft.net/fig/322326" pageId="0" pageNumber="139">22</figureCitation>
|
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|
|
), BOLD sample ID USNM_PG_H03.
|
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|
|
</paragraph>
|
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|
|
<caption doi="10.3897/zookeys.867.28728.figures18-22" httpUri="https://binary.pensoft.net/fig/322326" pageId="0" pageNumber="139" start="Figures 18–22" startId="F5">
|
|||
|
|
<paragraph pageId="0" pageNumber="139">
|
|||
|
|
Figures 18-22.
|
|||
|
|
<taxonomicName class="Insecta" family="Noctuidae" genus="Thraumata" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thraumata" order="Lepidoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="139">Thraumata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
male genitalia.
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="139">18-20</emphasis>
|
|||
|
|
<taxonomicName lsidName="T. petrovna" pageId="0" pageNumber="139" rank="species" species="petrovna">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="139">T. petrovna</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
dissection 148363, USNMENT01422625, Brazil
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="139">18</emphasis>
|
|||
|
|
Apparatus
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="139">19, 20</emphasis>
|
|||
|
|
Phallus
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="139">21, 22</emphasis>
|
|||
|
|
<taxonomicName lsidName="T. peruviensia" pageId="0" pageNumber="139" rank="species" species="peruviensia">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="139">T. peruviensia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, dissection 148129, USNMENT01440371
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="139">21</emphasis>
|
|||
|
|
Apparatus
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="139">22</emphasis>
|
|||
|
|
Phallus.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|