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<document id="F9DF85396142C148C98956514C611A48" ID-DOI="10.5281/zenodo.13183351" ID-ISSN="2533-4069" ID-Zenodo-Dep="13183351" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722606197785" checkinUser="felipe" docAuthor="Kvaček, Zlatko &amp; Teodoridis, Vasilis" docDate="2011" docId="C64487CCFFC0FF85FF2AF99D30571EC4" docLanguage="en" docName="ActaMusNatPraSerB.67.3-4.83-144.pdf" docOrigin="Acta Musei Nationalis Pragae Series B 67 (3 - 4)" docStyle="DocumentStyle:2E5EC822A0C9CE74494687E2DCEF8E0C.4:ActaMusNatPraSerB.2007-2013.journal_article.0cover.type2" docStyleId="2E5EC822A0C9CE74494687E2DCEF8E0C" docStyleName="ActaMusNatPraSerB.2007-2013.journal_article.0cover.type2" docStyleVersion="4" docTitle="Sterculia labrusca UNGER" docType="treatment" docVersion="2" lastPageNumber="102" masterDocId="3A7DFFB4FFD3FF96FFC7FF8E356D1C1F" masterDocTitle="The Late Eocene Flora Of Kučlín Near Bílina In North Bohemia Revisited" masterLastPageNumber="144" masterPageNumber="83" pageNumber="102" updateTime="1722656648556" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="563B60EAD0D4C2E978C0C97301D16713">The Late Eocene Flora Of Kučlín Near Bílina In North Bohemia Revisited</mods:title>
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<mods:namePart id="C1A72735DE0E20C4D41DF0545F0232AA">Kvaček, Zlatko</mods:namePart>
<mods:affiliation id="E34BD8B9FCDFCC2BFB907AD47BC56A7A">Institute of Geology and Palaeontology, Faculty of Science, Charles University in Prague, Albertov 6, 128 43 Prague 2, Czech Republic; e-mail: kvacek @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="9E75CABB6D00F8956C44256CE1711A97">Teodoridis, Vasilis</mods:namePart>
<mods:affiliation id="53C02D61D4F8B75707987DEC2D2E659D">Department of Biology and Environmental Education Studies, Faculty of Education, Charles University in Prague, M. D. Rettigové 4,116 39 Prague 1, Czech Republic; e-mail: vasilis. teodoridis @ pedf. cuni. cz</mods:affiliation>
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<taxonomicName id="89ED4D59FFC0FF85FF2AF99D371C1A35" authority="(UNGER) UNGER" authorityName="UNGER" baseAuthorityName="UNGER" box="[237,625,1555,1578]" class="Magnoliopsida" family="Malvaceae" genus="Sterculia" kingdom="Plantae" order="Malvales" pageId="19" pageNumber="102" phylum="Tracheophyta" rank="species" species="labrusca">
<emphasis id="7C99EAC8FFC0FF85FF2AF99D371C1A35" bold="true" box="[237,625,1555,1578]" pageId="19" pageNumber="102">
<emphasis id="7C99EAC8FFC0FF85FF2AF99D34DB1A35" bold="true" box="[237,438,1555,1578]" italics="true" pageId="19" pageNumber="102">Sterculia labrusca</emphasis>
(
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) 
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<emphasis id="7C99EAC8FFC0FF85FDE4F99D371C1A35" bold="true" box="[547,625,1555,1578]" pageId="19" pageNumber="102">UNGER</emphasis>
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</paragraph>
<paragraph id="4E5236DAFFC0FF85FED8F9B337501A4C" blockId="19.[287,573,1596,1619]" box="[287,573,1596,1619]" pageId="19" pageNumber="102">Pl. 9, figs 5-6. pl. 13, figs 6-7</paragraph>
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<subSubSection id="06F76551FFC0FF85FFB6F9E430571EC4" pageId="19" pageNumber="102" type="description">
<paragraph id="4E5236DAFFC0FF85FFB6F9E434801A60" blockId="19.[113,748,1642,1781]" box="[113,493,1642,1663]" pageId="19" pageNumber="102">
1850a 
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<emphasis id="7C99EAC8FFC0FF85FF72F9E434391A60" box="[181,340,1642,1663]" italics="true" pageId="19" pageNumber="102">Laurus labrusca</emphasis>
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, p. 433.
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1850b 
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<emphasis id="7C99EAC8FFC0FF85FF72F90934081A83" box="[181,357,1671,1692]" italics="true" pageId="19" pageNumber="102">Sterculia labrusca</emphasis>
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; Unger, p. 175, pl. 49, figs 1-11.
</paragraph>
<paragraph id="4E5236DAFFC0FF85FFB6F92B34671AC7" blockId="19.[113,748,1642,1781]" pageId="19" pageNumber="102">
1869 
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<emphasis id="7C99EAC8FFC0FF85FF72F92B34071AA5" box="[181,362,1701,1722]" italics="true" pageId="19" pageNumber="102">Sterculia labrusca</emphasis>
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; Ettingshausen, p. 13, pl. 43, figs 4, 5.
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1990 
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<emphasis id="7C99EAC8FFC0FF85FF72F96E34081AEA" box="[181,357,1760,1781]" italics="true" pageId="19" pageNumber="102">Sterculia labrusca</emphasis>
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; Bůžek et al., p. 172, fig. 3.25.
</paragraph>
<paragraph id="4E5236DAFFC0FF85FF50F89C37541BD5" blockId="19.[113,749,1810,1994]" pageId="19" pageNumber="102">Leaves simple, with thick petiole, lamina trilobate, lobes narrow and partly apically directed, partly patent, margin entire, venation tri- palmate, primaries thick, arising directly from the lamina base, secondaris veins numerous, arising perpendicularly, intersecondaries thin and parallel, venation of higher order poorly preserved.</paragraph>
<paragraph id="4E5236DAFFC0FF85FF50F855307A1D8B" blockId="19.[113,748,2011,2098]" lastBlockId="19.[824,1459,155,692]" pageId="19" pageNumber="102">
D i s c u s s i o n: These trilobate leaves with sleder, almost parallel-sided lobes were rarely recovered at Kučlín (
<bibRefCitation id="2A7C4B2BFFC0FF85FFBFF7953423142D" author="Ettingshausen, C." box="[120,334,2075,2098]" pageId="19" pageNumber="102" refId="ref37302" refString="Ettingshausen, C. (1869): Die fossile Flora des Tertiarbeckens von Bilin, III. (Enthaltend die Diapetalen und die allgemeinen Resultate der Bearbeitung). - K. K. Hof- und Staatsdruckerei, Wien, 110 pp. (Separatum from Denkschr. k. Akad. Wiss., math. - nat. Cl., 29)." type="book" year="1869">Ettingshausen 1869</bibRefCitation>
, pl. 43, figs 4-5). This species varies in leaf size and lobes. Most of the specimens recovered in the Staré Sedlo Fm. (
<bibRefCitation id="2A7C4B2BFFC0FF85FC35FF3531BA1CCD" author="Knobloch, E. &amp; Konzalova, M. &amp; Kvacek, Z." box="[1010,1239,187,210]" pageId="19" pageNumber="102" pagination="1 - 260" refId="ref39420" refString="Knobloch, E., Konzalova, M., Kvacek, Z. (1996): Die obereozane Flora der Stare Sedlo-Schichtenfolge in Bohmen (Mitteleuropa). - Rozpr. Ces. Geol. Ust., 49: 1 - 260." type="journal article" year="1996">Knobloch et al. 1996</bibRefCitation>
) are smaller, but the leaf described by 
<bibRefCitation id="2A7C4B2BFFC0FF85FC34FF5531DD1CED" author="Engelhardt, H." box="[1011,1200,219,242]" pageId="19" pageNumber="102" pagination="341 - 440" refId="ref37003" refString="Engelhardt, H. (1876): Tertiarpflanzen aus dem Leitmeritzer Mittelgebirge. Ein Beitrag zur Kenntniss der fossilen Pflanzen Bohmens. - Nova Acta Ksl. Leop. - Carol. Deutsch. Akad. Naturforsch., 38 (4): 341 - 440." type="journal article" year="1876">Engelhardt (1876</bibRefCitation>
, pl. 27, fig. 17) from the Žitenice quartzite matches well that from Kučlín. The other occurrences in the European Palaeogene (e.g., Geiseltal – 
<bibRefCitation id="2A7C4B2BFFC0FF85FCFFFEB236951D4C" author="Ruffle, L. &amp; Muller-Stoll, W. R. &amp; Litke, R." box="[824,1016,316,339]" pageId="19" pageNumber="102" pagination="199 - 282" refId="ref43437" refString="Ruffle, L., Muller-Stoll, W. R., Litke, R. (1976): Eozane Floren des Geiseltales. Weitere Ranales, Fagaceae, Loranthaceae, Apocynaceae. - Abh. Zentr. Geol. Inst., 26: 199 - 282." type="journal article" year="1976">Rüffle et al. 1976</bibRefCitation>
) may differ specifically and suggest that this foliage taxon may fall into smaller morphotypes, as already recognized by 
<bibRefCitation id="2A7C4B2BFFC0FF85FBEBFEF3307E1D8B" author="Ettingshausen, C." box="[1068,1299,381,404]" pageId="19" pageNumber="102" refId="ref37302" refString="Ettingshausen, C. (1869): Die fossile Flora des Tertiarbeckens von Bilin, III. (Enthaltend die Diapetalen und die allgemeinen Resultate der Bearbeitung). - K. K. Hof- und Staatsdruckerei, Wien, 110 pp. (Separatum from Denkschr. k. Akad. Wiss., math. - nat. Cl., 29)." type="book" year="1869">Ettingshausen (1869)</bibRefCitation>
.
</paragraph>
<paragraph id="4E5236DAFFC0FF85FC9AFE1330791EAB" blockId="19.[824,1459,155,692]" pageId="19" pageNumber="102">
The affinity of 
<taxonomicName id="89ED4D59FFC0FF85FBC0FE1331A01DAB" box="[1031,1229,413,436]" class="Magnoliopsida" family="Malvaceae" genus="Sterculia" kingdom="Plantae" order="Malvales" pageId="19" pageNumber="102" phylum="Tracheophyta" rank="species" species="labrusca">
<emphasis id="7C99EAC8FFC0FF85FBC0FE1331A01DAB" box="[1031,1229,413,436]" italics="true" pageId="19" pageNumber="102">Sterculia labrusca</emphasis>
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is not fully clarified (
<bibRefCitation id="2A7C4B2BFFC0FF85FCF8FE33314A1DCB" author="Knobloch, E. &amp; Konzalova, M. &amp; Kvacek, Z." box="[831,1063,445,468]" pageId="19" pageNumber="102" pagination="1 - 260" refId="ref39420" refString="Knobloch, E., Konzalova, M., Kvacek, Z. (1996): Die obereozane Flora der Stare Sedlo-Schichtenfolge in Bohmen (Mitteleuropa). - Rozpr. Ces. Geol. Ust., 49: 1 - 260." type="journal article" year="1996">Knobloch et al. 1996</bibRefCitation>
) and the often suggested as 
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<emphasis id="7C99EAC8FFC0FF85FA9AFE3336161DEB" italics="true" pageId="19" pageNumber="102">Brachychiton</emphasis>
</taxonomicName>
(
<taxonomicName id="89ED4D59FFC0FF85FC4AFE53314C1DEB" box="[909,1057,477,500]" class="Magnoliopsida" family="Sterculiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="19" pageNumber="102" phylum="Tracheophyta" rank="family">Sterculiaceae</taxonomicName>
), is unlikely because of its modern distribution in 
<collectingCountry id="36FA764AFFC0FF85FC15FE73315A1E0B" box="[978,1079,509,532]" name="Australia" pageId="19" pageNumber="102">Australia</collectingCountry>
. In its epidermal anatomy the material from Geiseltal (
<bibRefCitation id="2A7C4B2BFFC0FF85FBD2FD9331B21E2B" author="Ruffle, L. &amp; Muller-Stoll, W. R. &amp; Litke, R." box="[1045,1247,541,564]" pageId="19" pageNumber="102" pagination="199 - 282" refId="ref43437" refString="Ruffle, L., Muller-Stoll, W. R., Litke, R. (1976): Eozane Floren des Geiseltales. Weitere Ranales, Fagaceae, Loranthaceae, Apocynaceae. - Abh. Zentr. Geol. Inst., 26: 199 - 282." type="journal article" year="1976">Rüffle et al. 1976</bibRefCitation>
) deviates from the standard pattern of living sterculias (lack of stellate trichomes, paracytic stomata). No other leaf anatomical data are available from other occurrences in Europe, in particular from the 
<typeStatus id="91568878FFC0FF85FC79FD1336811EAB" box="[958,1004,669,692]" pageId="19" pageNumber="102">type</typeStatus>
locality Socka in 
<collectingCountry id="36FA764AFFC0FF85FB68FD13307D1EAB" box="[1199,1296,669,692]" name="Slovenia" pageId="19" pageNumber="102">Slovenia</collectingCountry>
.
</paragraph>
<paragraph id="4E5236DAFFC0FF85FC9AFD4A30571EC4" blockId="19.[861,1338,708,731]" box="[861,1338,708,731]" pageId="19" pageNumber="102">M a t e r i a l: BP 55. 2401.1, NM G 8654.</paragraph>
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</treatment>
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