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<document ID-DOI="10.1093/zoolinnean/zlac034" ID-ISSN="0024-4082" ID-Zenodo-Dep="7187082" ID-ZooBank="26533E9F-C53F-4051-B6B4-990E63C08C74" checkinTime="1665471212239" checkinUser="plazi" docAuthor="Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich &amp; Harms, Danilo" docDate="2022" docId="03E7173D3524E323FCB0F402BC64DAEB" docLanguage="en" docName="zlac034.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Annazomus parvulus Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms 2022, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="801" masterDocId="FFDE6F453521E32AFFC0F00FB865DA12" masterDocTitle="Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane" masterLastPageNumber="844" masterPageNumber="792" pageNumber="797" updateTime="1665531758795" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Magnussen, Ilian De Francesco</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Müller, Sandro P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hammel, Jörg U.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kotthoff, Ulrich</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harms, Danilo</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>196</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>792</mods:start>
<mods:end>844</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1093/zoolinnean/zlac034</mods:identifier>
<mods:identifier type="ISSN">0024-4082</mods:identifier>
<mods:identifier type="Zenodo-Dep">7187082</mods:identifier>
<mods:identifier type="ZooBank">26533E9F-C53F-4051-B6B4-990E63C08C74</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7187094" ID-Zenodo-Dep="7187094" LSID="urn:lsid:plazi:treatment:03E7173D3524E323FCB0F402BC64DAEB" httpUri="http://treatment.plazi.org/id/03E7173D3524E323FCB0F402BC64DAEB" lastPageId="9" lastPageNumber="801" pageId="5" pageNumber="797">
<subSubSection pageId="5" pageNumber="797" type="nomenclature">
<taxonomicName authority="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms, 2022" authorityName="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms" authorityYear="2022" class="Arachnida" family="Hubbardiidae" genus="Annazomus" kingdom="Animalia" order="Schizomida" pageId="5" pageNumber="797" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
<paragraph blockId="5.[880,1353,1037,1093]" box="[880,1353,1037,1062]" pageId="5" pageNumber="797">
<heading box="[880,1353,1037,1062]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="797" reason="2">
†
<emphasis bold="true" box="[892,1170,1037,1061]" italics="true" pageId="5" pageNumber="797">ANNAZOMUS PARVULUS</emphasis>
DE FRANCESCO
</heading>
</paragraph>
<paragraph blockId="5.[880,1353,1037,1093]" box="[992,1242,1068,1093]" pageId="5" pageNumber="797">
<heading box="[992,1242,1068,1093]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="797" reason="3">
MAGNUSSEN
<emphasis bold="true" box="[1143,1242,1068,1092]" pageId="5" pageNumber="797">
<taxonomicNameLabel box="[1143,1242,1068,1092]" pageId="5" pageNumber="797" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</taxonomicName>
<paragraph blockId="5.[1057,1176,1106,1131]" box="[1057,1176,1106,1131]" pageId="5" pageNumber="797">
(
<figureCitation box="[1065,1168,1106,1131]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStartId-0="6.[163,241,1750,1772]" captionStartId-1="7.[144,222,1517,1539]" captionStartId-2="8.[163,241,1518,1540]" captionTargetBox-0="[371,1220,197,1709]" captionTargetBox-1="[209,1338,198,1474]" captionTargetBox-2="[165,1446,197,1477]" captionTargetId-0="figure-27@6.[369,1236,195,1710]" captionTargetId-1="figure-150@7.[206,1364,195,1477]" captionTargetId-2="figure-126@8.[163,1446,195,1478]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figure 3. Specimen GPIH04893, †Annazomus parvulus sp. nov., holotype. A, specimen in dorsal view, photograph. B, same, interpretative drawing. C, specimen in ventral view photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 1 mm." captionText-1="Figure 4. Specimen GPIH04893,†Annazomusparvulus sp.nov.,holotype,details.A, metapeltidium dorsalview,photograph. B, same, interpretative drawing. C, right pedipalp in mesal view, photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm (A, B); 0.5 mm (C, D)." captionText-2="Figure 5. Specimen GPIH04893, †Annazomus parvulus sp. nov., holotype, details. A, flagellum in dorsal view, photograph. B, same, interpretative drawing. C, flagellum in ventral view, photograph. D, same, interpretative drawing. E, flagellum in dorsal view, three-dimensional (3D) model based on synchrotron scan. F, flagellum in ventral view, 3D model based on X-rayscan. G, flagellum in lateral view, 3D model based on X-ray-scan. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7187092" figureDoi-1="http://doi.org/10.5281/zenodo.7187096" figureDoi-2="http://doi.org/10.5281/zenodo.7187098" httpUri-0="https://zenodo.org/record/7187092/files/figure.png" httpUri-1="https://zenodo.org/record/7187096/files/figure.png" httpUri-2="https://zenodo.org/record/7187098/files/figure.png" pageId="5" pageNumber="797">FIGS 3–5</figureCitation>
)
</paragraph>
<paragraph blockId="5.[809,1426,1168,1221]" pageId="5" pageNumber="797">
<emphasis box="[809,1090,1168,1190]" italics="true" pageId="5" pageNumber="797">Zoobank registration:</emphasis>
<uri pageId="5" pageNumber="797">
urn:lsid:zoobank.org:act:
<uuid box="[809,1322,1199,1221]" pageId="5" pageNumber="797">9221DD2F-0C17-48AC-8904-C18E18A05F96</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="797" type="materials_examined">
<paragraph blockId="5.[809,1424,1264,1348]" pageId="5" pageNumber="797">
<emphasis box="[809,1012,1264,1286]" italics="true" pageId="5" pageNumber="797">Type material:</emphasis>
<typeStatus box="[1029,1150,1264,1286]" pageId="5" pageNumber="797">Holotype</typeStatus>
(
<figureCitation box="[1178,1317,1264,1286]" captionStart="Figure 3" captionStartId="6.[163,241,1750,1772]" captionTargetBox="[371,1220,197,1709]" captionTargetId="figure-27@6.[369,1236,195,1710]" captionTargetPageId="6" captionText="Figure 3. Specimen GPIH04893, †Annazomus parvulus sp. nov., holotype. A, specimen in dorsal view, photograph. B, same, interpretative drawing. C, specimen in ventral view photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7187092" httpUri="https://zenodo.org/record/7187092/files/figure.png" pageId="5" pageNumber="797">Fig. 3A –D</figureCitation>
), adult homeomorphic
<specimenCount box="[1007,1023,1298,1316]" pageId="5" pageNumber="797" type="male">♂</specimenCount>
, collection no.
<emphasis box="[1229,1377,1295,1316]" italics="true" pageId="5" pageNumber="797">
<collectionCode box="[1229,1301,1295,1316]" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="5" pageNumber="797">GPIH</collectionCode>
04893
</emphasis>
; ex collection Patrick Müller (no. BUB556).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="797" type="diagnosis">
<paragraph blockId="5.[809,930,1413,1436]" box="[809,930,1413,1436]" pageId="5" pageNumber="797">
<emphasis box="[809,930,1413,1436]" italics="true" pageId="5" pageNumber="797">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="5.[809,1425,1452,1536]" pageId="5" pageNumber="797">
Pedipalps short; flagellum shape nearly rectangular, with a bulbous elevation on the left and right dorsal side; small body size of
<quantity box="[1076,1177,1513,1535]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="797" unit="mm" value="1.95">1.95 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="797" type="etymology">
<paragraph blockId="5.[809,938,1600,1624]" box="[809,938,1600,1624]" pageId="5" pageNumber="797">
<emphasis box="[809,938,1600,1624]" italics="true" pageId="5" pageNumber="797">Etymology</emphasis>
</paragraph>
<paragraph blockId="5.[809,1425,1640,1724]" pageId="5" pageNumber="797">
The specific epithet is Latin
<taxonomicName authorityName="De Francesco Magnussen" authorityYear="2022" box="[1133,1234,1640,1661]" class="Arachnida" family="Hubbardiidae" genus="Annazomus" kingdom="Animalia" order="Schizomida" pageId="5" pageNumber="797" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis box="[1133,1234,1640,1661]" italics="true" pageId="5" pageNumber="797">parvulus</emphasis>
</taxonomicName>
, small, referring to the small body size of the specimen. It is masculine in gender.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="801" pageId="5" pageNumber="797" type="description">
<paragraph blockId="5.[809,949,1789,1812]" box="[809,949,1789,1812]" pageId="5" pageNumber="797">
<emphasis box="[809,949,1789,1812]" italics="true" pageId="5" pageNumber="797">Description</emphasis>
</paragraph>
<paragraph blockId="5.[809,1425,1828,1881]" pageId="5" pageNumber="797">
Total length from the anterior process to the base of the flagellum
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.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7187092" ID-Zenodo-Dep="7187092" httpUri="https://zenodo.org/record/7187092/files/figure.png" pageId="6" pageNumber="798" startId="6.[163,241,1750,1772]" targetBox="[371,1220,197,1709]" targetPageId="6">
<paragraph blockId="6.[162,1443,1750,1831]" pageId="6" pageNumber="798">
<emphasis bold="true" box="[163,266,1750,1772]" pageId="6" pageNumber="798">Figure 3.</emphasis>
Specimen
<emphasis box="[384,508,1750,1772]" italics="true" pageId="6" pageNumber="798">GPIH04893</emphasis>
, †
<taxonomicName authority="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms, 2022" authorityName="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms" authorityYear="2022" box="[530,751,1750,1772]" class="Arachnida" family="Hubbardiidae" genus="Annazomus" kingdom="Animalia" order="Schizomida" pageId="6" pageNumber="798" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
<emphasis box="[530,751,1750,1772]" italics="true" pageId="6" pageNumber="798">Annazomus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[757,835,1750,1772]" pageId="6" pageNumber="798" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. A, specimen in dorsal view, photograph. B, same, interpretative drawing. C, specimen in ventral view photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 1 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7187096" ID-Zenodo-Dep="7187096" httpUri="https://zenodo.org/record/7187096/files/figure.png" pageId="7" pageNumber="799" startId="7.[144,222,1517,1539]" targetBox="[209,1338,198,1474]" targetPageId="7">
<paragraph blockId="7.[144,1426,1517,1597]" pageId="7" pageNumber="799">
<emphasis bold="true" box="[144,245,1517,1539]" pageId="7" pageNumber="799">Figure 4.</emphasis>
Specimen
<emphasis box="[363,487,1517,1539]" italics="true" pageId="7" pageNumber="799">GPIH04893</emphasis>
, †
<taxonomicName authority="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms, 2022" authorityName="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms" authorityYear="2022" box="[508,727,1517,1539]" class="Arachnida" family="Hubbardiidae" genus="Annazomus" kingdom="Animalia" order="Schizomida" pageId="7" pageNumber="799" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
<emphasis box="[508,727,1517,1539]" italics="true" pageId="7" pageNumber="799">Annazomus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[731,807,1517,1539]" pageId="7" pageNumber="799" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, details.A, metapeltidium dorsal view, photograph. B, same, interpretative drawing. C, right pedipalp in mesal view, photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm (A, B); 0.5 mm (C, D).
</paragraph>
</caption>
<paragraph blockId="7.[145,760,1660,1774]" pageId="7" pageNumber="799">
<emphasis box="[145,356,1660,1681]" italics="true" pageId="7" pageNumber="799">Colour (in amber):</emphasis>
Light brown and golden; prosoma, opisthosoma, flagellum, femur IV and pedipalps slightly darker brown golden; leg I golden and translucent, original colouring equivocal.
</paragraph>
<paragraph blockId="7.[145,763,1814,1898]" lastBlockId="8.[827,1443,1696,1779]" lastPageId="8" lastPageNumber="800" pageId="7" pageNumber="799">
<emphasis box="[145,261,1815,1836]" italics="true" pageId="7" pageNumber="799">Prosoma:</emphasis>
Length
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; propeltidium length
<quantity box="[145,254,1845,1867]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="7" pageNumber="799" unit="mm" value="0.68">0.68 mm</quantity>
, width
<quantity box="[343,450,1845,1866]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="7" pageNumber="799" unit="mm" value="0.32">0.32 mm</quantity>
. Anterior process sharply bent down, length not measurable; setation: one seta, sitting on the left side of the anterior process, no further setation visible; small round holes in the proximal part of the propeltidium prevent reconstruction of further setation, but there might have been a pair of setae with the right one now missing, definite setation remains unclear. Propeltidium setation prosoma: three pairs, first pair of setae still in place, second pair left one aborted, third pair both aborted with only left tubercle visible, right one not visible owing to a crack in the amber covering parts of the propeltidium. Corneate eyes and eyespots absent. Mesopeltidia width (right): ~
<quantity box="[177,281,1788,1809]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="800" unit="mm" value="0.11">0.11 mm</quantity>
, left one not visible owing to an air bubble in the amber. Metapeltidium not divided with a thin suture (
<figureCitation box="[258,377,1849,1871]" captionStart="Figure 4" captionStartId="7.[144,222,1517,1539]" captionTargetBox="[209,1338,198,1474]" captionTargetId="figure-150@7.[206,1364,195,1477]" captionTargetPageId="7" captionText="Figure 4. Specimen GPIH04893,†Annazomusparvulus sp.nov.,holotype,details.A, metapeltidium dorsalview,photograph. B, same, interpretative drawing. C, right pedipalp in mesal view, photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm (A, B); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7187096" httpUri="https://zenodo.org/record/7187096/files/figure.png" pageId="8" pageNumber="800">Fig. 4A, B</figureCitation>
), length
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, width
<quantity box="[669,777,1849,1871]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="800" unit="mm" value="0.42">0.42 mm</quantity>
. Anterior sternum with 10 + 2 sternapophysial setae, length
<quantity box="[912,1019,1727,1748]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="800" unit="mm" value="0.24">0.24 mm</quantity>
, width
<quantity box="[1108,1215,1727,1748]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="800" unit="mm" value="0.22">0.22 mm</quantity>
. Posterior sternum with six setae, length
<quantity box="[1080,1181,1757,1779]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="800" unit="mm" value="0.12">0.12 mm</quantity>
, width
<quantity box="[1265,1369,1757,1779]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="800" unit="mm" value="0.11">0.11 mm</quantity>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7187098" ID-Zenodo-Dep="7187098" httpUri="https://zenodo.org/record/7187098/files/figure.png" pageId="8" pageNumber="800" startId="8.[163,241,1518,1540]" targetBox="[165,1446,197,1477]" targetPageId="8">
<paragraph blockId="8.[161,1444,1518,1658]" pageId="8" pageNumber="800">
<emphasis bold="true" box="[163,267,1518,1540]" pageId="8" pageNumber="800">Figure 5.</emphasis>
Specimen
<emphasis box="[385,509,1518,1540]" italics="true" pageId="8" pageNumber="800">GPIH04893</emphasis>
, †
<taxonomicName authority="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms, 2022" authorityName="Magnussen &amp; Müller &amp; Hammel &amp; Kotthoff &amp; Harms" authorityYear="2022" box="[532,754,1518,1540]" class="Arachnida" family="Hubbardiidae" genus="Annazomus" kingdom="Animalia" order="Schizomida" pageId="8" pageNumber="800" phylum="Arthropoda" rank="species" species="parvulus" status="sp. nov.">
<emphasis box="[532,754,1518,1540]" italics="true" pageId="8" pageNumber="800">Annazomus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[760,838,1518,1540]" pageId="8" pageNumber="800" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, details. A, flagellum in dorsal view, photograph. B, same, interpretative drawing. C, flagellum in ventral view, photograph. D, same, interpretative drawing. E, flagellum in dorsal view, three-dimensional (3D) model based on synchrotron scan. F, flagellum in ventral view, 3D model based on X-rayscan. G, flagellum in lateral view, 3D model based on X-ray-scan. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph blockId="8.[826,1443,1820,1873]" pageId="8" pageNumber="800">
<emphasis box="[827,952,1820,1841]" italics="true" pageId="8" pageNumber="800">Chelicerae:</emphasis>
Setation of the mesal side of chelicerae not visible, except for G6-1.
</paragraph>
<paragraph blockId="9.[143,762,197,618]" pageId="9" pageNumber="801">
<emphasis box="[145,262,197,218]" italics="true" pageId="9" pageNumber="801">Pedipalps:</emphasis>
Short and slim, without armature (
<figureCitation captionStart="Figure 4" captionStartId="7.[144,222,1517,1539]" captionTargetBox="[209,1338,198,1474]" captionTargetId="figure-150@7.[206,1364,195,1477]" captionTargetPageId="7" captionText="Figure 4. Specimen GPIH04893,†Annazomusparvulus sp.nov.,holotype,details.A, metapeltidium dorsalview,photograph. B, same, interpretative drawing. C, right pedipalp in mesal view, photograph. D, same, interpretative drawing. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm (A, B); 0.5 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.7187096" httpUri="https://zenodo.org/record/7187096/files/figure.png" pageId="9" pageNumber="801">Fig. 4C, D</figureCitation>
), mesal spur on trochanter not visible; trochanter setation: five ventral strong setae, two ectal, four mesal setae; femur: longer than trochanter, setation: five dorsal, three ectal, two ventroectal, three mesal; patella: shorter than femur, setation: four mesal, two ectal, four dorsomesal, two dorsoectal, three dorsal; tibia: shorter than patella, setation: three dorsomesal, three ectal, two ventroectal, two dorsoectal, five mesal; tarsus: with tarsal spurs, spur length:
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, setation: three ectal, four dorsoectal, four ventroectal, two dorsal, three dorsomesal, three+ ventromesal, five mesal, two+ ventral; claw: straight, slightly longer than half the length of tarsus.
</paragraph>
<paragraph blockId="9.[144,761,658,834]" pageId="9" pageNumber="801">
<emphasis box="[145,202,659,680]" italics="true" pageId="9" pageNumber="801">Legs:</emphasis>
Right leg I aborted at the trochanter, left leg well preserved and complete, tarsus with six segments, leg I ~1.37 times longer than the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV 2.65 times longer than deep; leg IV&gt; 1.04 times longer than the total body length; leg formula: 1423.
</paragraph>
<paragraph blockId="9.[143,761,874,1326]" pageId="9" pageNumber="801">
<emphasis box="[145,315,874,895]" italics="true" pageId="9" pageNumber="801">Opisthosoma:</emphasis>
Length
<quantity box="[431,544,875,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" pageId="9" pageNumber="801" unit="mm" value="1.12">1.12 mm</quantity>
, width
<quantity box="[644,757,875,896]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="801" unit="mm" value="0.42">0.42 mm</quantity>
; tergites X–XII not elongated, tergite XII with a large, rectangular posterodorsal process. Abdominal tergites setation: I: Dm; II: Dm; III: Dm, left missing; IV: Dm, both missing; V: Dm, both missing; VI: Dm; VII: Dm, both missing; VIII: Dm, left missing, Dl1; IX: Dl1, Dl2; X: Dl1; XI: Dl1; XII: Dm, Dl1, setation pattern: 2:2:2:2:2:2:2:4:4:2:2:4. Flagellum: dorsoventrally flattened, rectangular with an apical tip at the posterior end, on the dorsal side with a bulbous elevation each left and right near the posterior end (
<figureCitation box="[557,669,1181,1203]" captionStart="Figure 5" captionStartId="8.[163,241,1518,1540]" captionTargetBox="[165,1446,197,1477]" captionTargetId="figure-126@8.[163,1446,195,1478]" captionTargetPageId="8" captionText="Figure 5. Specimen GPIH04893, †Annazomus parvulus sp. nov., holotype, details. A, flagellum in dorsal view, photograph. B, same, interpretative drawing. C, flagellum in ventral view, photograph. D, same, interpretative drawing. E, flagellum in dorsal view, three-dimensional (3D) model based on synchrotron scan. F, flagellum in ventral view, 3D model based on X-rayscan. G, flagellum in lateral view, 3D model based on X-ray-scan. See Material and methods for explanation of abbreviations. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7187098" httpUri="https://zenodo.org/record/7187098/files/figure.png" pageId="9" pageNumber="801">Fig. 5A–G</figureCitation>
), length
<quantity box="[143,254,1212,1233]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="801" unit="mm" value="0.17">0.17 mm</quantity>
, width
<quantity box="[348,458,1212,1233]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="801" unit="mm" value="0.22">0.22 mm</quantity>
; stalk length ~
<quantity box="[646,756,1212,1233]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="801" unit="mm" value="0.09">0.09 mm</quantity>
; setation: Dm1, Dl2 paired, Dm4, Dl3 paired and nearly at the posterior end of the flagellum, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired.
</paragraph>
<paragraph blockId="9.[143,763,1366,1910]" lastBlockId="9.[809,1423,197,250]" pageId="9" pageNumber="801">
<emphasis box="[145,461,1366,1388]" italics="true" pageId="9" pageNumber="801">Further measurements:</emphasis>
Pedipalp: trochanter
<quantity box="[145,255,1397,1419]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="801" unit="mm" value="0.16">0.16 mm</quantity>
; femur
<quantity box="[355,465,1397,1419]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="801" unit="mm" value="0.26">0.26 mm</quantity>
; patella
<quantity box="[578,688,1397,1419]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="801" unit="mm" value="0.23">0.23 mm</quantity>
; tibia
<quantity box="[145,259,1428,1449]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="801" unit="mm" value="0.2">0.20 mm</quantity>
; tarsus
<quantity box="[369,484,1428,1449]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="801" unit="mm" value="0.13">0.13 mm</quantity>
; claw
<quantity box="[570,684,1428,1449]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="801" unit="mm" value="0.07">0.07 mm</quantity>
; total length
<quantity box="[245,361,1458,1480]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="801" unit="mm" value="1.05">1.05 mm</quantity>
. Leg I: trochanter
<quantity box="[636,753,1459,1480]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="801" unit="mm" value="0.19">0.19 mm</quantity>
; femur
<quantity box="[229,340,1489,1511]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="801" unit="mm" value="0.61">0.61 mm</quantity>
; patella
<quantity box="[452,563,1489,1511]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="9" pageNumber="801" unit="mm" value="0.71">0.71 mm</quantity>
; tibia
<quantity box="[646,756,1489,1511]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="9" pageNumber="801" unit="mm" value="0.58">0.58 mm</quantity>
; metatarsus
<quantity box="[303,417,1520,1541]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="801" unit="mm" value="0.24">0.24 mm</quantity>
; tarsus (tarsus segments)
<quantity box="[145,252,1551,1572]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="801" unit="mm" value="0.34">0.34 mm</quantity>
(
<quantity box="[270,382,1551,1572]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="801" unit="mm" value="0.04">0.04 mm</quantity>
;
<quantity box="[397,507,1551,1572]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="801" unit="mm" value="0.04">0.04 mm</quantity>
;
<quantity box="[521,632,1551,1572]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="801" unit="mm" value="0.04">0.04 mm</quantity>
;
<quantity box="[646,755,1551,1572]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="801" unit="mm" value="0.04">0.04 mm</quantity>
;
<quantity box="[145,257,1581,1603]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="801" unit="mm" value="0.05">0.05 mm</quantity>
;
<quantity box="[268,380,1581,1602]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="801" unit="mm" value="0.13">0.13 mm</quantity>
); total length
<quantity box="[555,668,1581,1603]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" pageId="9" pageNumber="801" unit="mm" value="2.67">2.67 mm</quantity>
. Leg II: t r o c h a n t e r 0.1
<quantity box="[343,391,1612,1633]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="801" unit="m" value="2.0">2 m</quantity>
m; f e m u r 0.4
<quantity box="[571,621,1612,1633]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="801" unit="m" value="9.0">9 m</quantity>
m; p a t e l l a
<quantity box="[145,246,1643,1664]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="801" unit="mm" value="0.22">0.22 mm</quantity>
; tibia
<quantity box="[317,420,1642,1664]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="801" unit="mm" value="0.25">0.25 mm</quantity>
; metatarsus
<quantity box="[571,674,1643,1664]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="801" unit="mm" value="0.29">0.29 mm</quantity>
; tarsus (tarsus segments)
<quantity box="[383,494,1673,1695]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="801" unit="mm" value="0.26">0.26 mm</quantity>
(
<quantity box="[514,628,1673,1695]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="801" unit="mm" value="0.1">0.10 mm</quantity>
;
<quantity box="[644,755,1673,1695]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="801" unit="mm" value="0.06">0.06 mm</quantity>
;
<quantity box="[145,252,1704,1725]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="801" unit="mm" value="0.1">0.10 mm</quantity>
); total length
<quantity box="[420,526,1704,1726]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="9" pageNumber="801" unit="mm" value="1.63">1.63 mm</quantity>
. Leg III: trochanter
<quantity box="[146,255,1735,1756]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="801" unit="mm" value="0.13">0.13 mm</quantity>
; femur
<quantity box="[355,465,1735,1756]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="801" unit="mm" value="0.48">0.48 mm</quantity>
; patella
<quantity box="[578,688,1735,1756]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="801" unit="mm" value="0.17">0.17 mm</quantity>
; tibia
<quantity box="[145,261,1765,1787]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="801" unit="mm" value="0.19">0.19 mm</quantity>
; metatarsus
<quantity box="[439,555,1765,1787]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="801" unit="mm" value="0.25">0.25 mm</quantity>
; tarsus (tarsus segments)
<quantity box="[277,383,1796,1818]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="801" unit="mm" value="0.26">0.26 mm</quantity>
(
<quantity box="[399,507,1796,1817]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="801" unit="mm" value="0.12">0.12 mm</quantity>
;
<quantity box="[521,628,1796,1818]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="801" unit="mm" value="0.06">0.06 mm</quantity>
;
<quantity box="[641,749,1796,1817]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="801" unit="mm" value="0.08">0.08 mm</quantity>
); total length
<quantity box="[298,405,1827,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="9" pageNumber="801" unit="mm" value="1.48">1.48 mm</quantity>
. Leg IV: trochanter
<quantity box="[651,756,1827,1848]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="801" unit="mm" value="0.19">0.19 mm</quantity>
; femur
<quantity box="[225,328,1857,1879]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="801" unit="mm" value="0.7">0.70 mm</quantity>
; patella&gt;
<quantity box="[452,556,1857,1879]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="801" unit="mm" value="0.22">0.22 mm</quantity>
; tibia&gt;
<quantity box="[652,756,1857,1879]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="801" unit="mm" value="0.29">0.29 mm</quantity>
; metatarsus
<quantity box="[303,417,1888,1910]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="801" unit="mm" value="0.35">0.35 mm</quantity>
; tarsus (tarsus segments) 0.2
<quantity box="[849,899,197,219]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="801" unit="m" value="8.0">8 m</quantity>
m (0.1
<quantity box="[987,1036,197,219]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="801" unit="m" value="1.0">1 m</quantity>
m; 0.0
<quantity box="[1123,1172,197,219]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="801" unit="m" value="6.0">6 m</quantity>
m; 0.1
<quantity box="[1258,1307,197,219]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="801" unit="m" value="1.0">1 m</quantity>
m); t o t a l length&gt;
<quantity box="[915,1021,228,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="9" pageNumber="801" unit="mm" value="2.03">2.03 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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