treatments-xml/data/03/F8/87/03F8878BFF94FFC2F5EAF88E4642FAE9.xml

<document id="11D6CAA22DD1AC0EB096BF3851D1D195" ID-DOI="10.11646/zootaxa.3860.1.1" ID-GBIF-Dataset="ceb30f74-c770-4952-859a-9d3a84a5e087" ID-ISSN="1175-5326" ID-Zenodo-Dep="287059" ID-ZooBank="FDFD1014-8DDA-4EED-A385-95FA4F964CFC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460903466920" checkinUser="plazi" docAuthor="Eguchi, Katsuyuki, Viet, Bui Tuan &amp; Yamane, Seiki" docDate="2014" docId="03F8878BFF94FFC2F5EAF88E4642FAE9" docLanguage="en" docName="zt03860p046.pdf" docOrigin="Zootaxa 3860 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protanilla Taylor 1990" docType="treatment" docVersion="6" lastPageNumber="23" masterDocId="FFC1FFF3FF81FFD4F57DFFFF445AFFC9" masterDocTitle="Generic Synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), Part II — Cerapachyinae, Aenictinae, Dorylinae, Leptanillinae, Amblyoponinae, Ponerinae, Ectatomminae and Proceratiinae" masterLastPageNumber="46" masterPageNumber="1" pageNumber="22" updateTime="1698360472837" updateUser="plazi">
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<mods:title id="7A1C254AF0978332B4FA33CAF1AED081">Generic Synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), Part II — Cerapachyinae, Aenictinae, Dorylinae, Leptanillinae, Amblyoponinae, Ponerinae, Ectatomminae and Proceratiinae</mods:title>
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<mods:namePart id="A65A6BC0B9D3303B9DB6324EEF18FA93">Eguchi, Katsuyuki</mods:namePart>
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<mods:namePart id="1C6E0D48FCE988455142EFB37D1FD9DD">Viet, Bui Tuan</mods:namePart>
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<mods:namePart id="49E36F75153B693CE752EC5DA2EA0897">Yamane, Seiki</mods:namePart>
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<mods:date id="21E39D170814868ABA3E8202C15F5C49">2014</mods:date>
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<mods:number id="C58A9FCF8A605DD4E095CF35F760FE6B">3860</mods:number>
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<treatment id="03F8878BFF94FFC2F5EAF88E4642FAE9" ID-DOI="http://doi.org/10.5281/zenodo.6124671" ID-GBIF-Taxon="127669770" ID-Zenodo-Dep="6124671" LSID="urn:lsid:plazi:treatment:03F8878BFF94FFC2F5EAF88E4642FAE9" httpUri="http://treatment.plazi.org/id/03F8878BFF94FFC2F5EAF88E4642FAE9" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="C34B6516FF94FFC2F5EAF88E4642FAE9" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BEE369DFF94FFC1F5EAF88E45EAF842" blockId="21.[151,432,1905,1931]" box="[151,432,1905,1931]" pageId="21" pageNumber="22">
<heading id="D0A681F1FF94FFC1F5EAF88E45EAF842" bold="true" box="[151,432,1905,1931]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="4C514D1EFF94FFC1F5EAF88E45EAF842" ID-CoL="6XDG" authority="Taylor, 1990" authorityName="Taylor" authorityYear="1990" box="[151,432,1905,1931]" class="Insecta" family="Formicidae" genus="Protanilla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B925EA8FFF94FFC1F5EAF88E45EAF842" bold="true" box="[151,432,1905,1931]" pageId="21" pageNumber="22">
<emphasis id="B925EA8FFF94FFC1F5EAF88E454BF842" bold="true" box="[151,273,1905,1931]" italics="true" pageId="21" pageNumber="22">Protanilla</emphasis>
Taylor, 1990
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BEE369DFF94FFC1F5EAF847409FF83D" blockId="21.[151,1436,1976,2037]" pageId="21" pageNumber="22">
<emphasis id="B925EA8FFF94FFC1F5EAF8474540F818" bold="true" box="[151,282,1976,2001]" pageId="21" pageNumber="22">Taxonomy.</emphasis>
The genus 
<taxonomicName id="4C514D1EFF94FFC1F4D7F8474646F819" box="[426,540,1976,2000]" class="Insecta" family="Formicidae" genus="Protanilla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B925EA8FFF94FFC1F4D7F8474646F819" box="[426,540,1976,2000]" italics="true" pageId="21" pageNumber="22">Protanilla</emphasis>
</taxonomicName>
is assigned to the tribe 
<taxonomicName id="4C514D1EFF94FFC1F63EF8474056F818" box="[835,1036,1976,2001]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Anomalomyrmini">Anomalomyrmini</taxonomicName>
(
<bibRefCitation id="EFC04B6CFF94FFC1F162F8474096F819" author="Bolton" box="[1055,1228,1976,2001]" pageId="21" pageNumber="22" refString="Bolton, B. (1990 c) The higher classification of the ant subfamily Leptanillinae (Hymenoptera: Formicidae). Systematic Entomology, 15, 267 - 282. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1990. tb 00063. x" type="journal article" year="1990" yearSuffix="c">Bolton 1990c</bibRefCitation>
, 
<bibRefCitation id="EFC04B6CFF94FFC1F1A0F8474143F819" author="Bolton" box="[1245,1305,1976,2000]" pageId="21" pageNumber="22" refString="Bolton, B. (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute, 71, 1 - 370." type="journal article" year="2003">2003</bibRefCitation>
). The tribe 
<taxonomicName id="4C514D1EFF94FFC1F5EAF823453AF83C" box="[151,352,2012,2037]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Anomalomyrmini">Anomalomyrmini</taxonomicName>
was recently revised, and 
<taxonomicName id="4C514D1EFF94FFC1F7F5F82346A0F83D" box="[648,762,2012,2036]" class="Insecta" family="Formicidae" genus="Protanilla" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B925EA8FFF94FFC1F7F5F82346A0F83D" box="[648,762,2012,2036]" italics="true" pageId="21" pageNumber="22">Protanilla</emphasis>
</taxonomicName>
was redefined by 
<bibRefCitation id="EFC04B6CFF94FFC1F6B5F823409AF83D" box="[968,1216,2012,2037]" pageId="21" pageNumber="22" refString="Borowiec, M. L., Schulz, A., Alpert, G. D. &amp; Banar, P. (2011) Discovery of the worker caste and descriptions of two new species of Anomalomyrma (Hymenoptera: Formicidae: Leptanillinae) with unique abdominal morphology. Zootaxa, 2810, 1 - 14." type="journal article">
Borowiec 
<emphasis id="B925EA8FFF94FFC1F141F8224037F83D" box="[1084,1133,2012,2036]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
(2011)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFF6845C8FF1D" blockId="22.[151,1437,151,1005]" pageId="22" pageNumber="23">
<emphasis id="B925EA8FFF97FFC2F5BAFF68453AFF79" bold="true" box="[199,352,151,176]" pageId="22" pageNumber="23">Morphology.</emphasis>
Workers of Vietnamese species have the following features (see also 
<bibRefCitation id="EFC04B6CFF97FFC2F1B8FF68413FFF79" author="Bolton" box="[1221,1381,151,176]" pageId="22" pageNumber="23" refString="Bolton, B. (1990 c) The higher classification of the ant subfamily Leptanillinae (Hymenoptera: Formicidae). Systematic Entomology, 15, 267 - 282. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1990. tb 00063. x" type="journal article" year="1990" yearSuffix="c">Bolton 1990c</bibRefCitation>
and 
<bibRefCitation id="EFC04B6CFF97FFC2F5EAFF4345DCFF1D" author="Borowiec" box="[151,390,188,213]" pageId="22" pageNumber="23" refString="Borowiec, M. L., Schulz, A., Alpert, G. D. &amp; Banar, P. (2011) Discovery of the worker caste and descriptions of two new species of Anomalomyrma (Hymenoptera: Formicidae: Leptanillinae) with unique abdominal morphology. Zootaxa, 2810, 1 - 14." type="journal article" year="2011">
Borowiec 
<emphasis id="B925EA8FFF97FFC2F476FF424519FF1D" box="[267,323,188,212]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2011
</bibRefCitation>
):
</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFF2045ADFCF1" blockId="22.[151,1437,151,1005]" pageId="22" pageNumber="23">Worker monomorphic; head in full-face view somewhat elongate trapezoidal; preoccipital carina present but weak dorsally and laterally on head; frontal lobe, frontal carina and antennal scrobe absent; antennal sockets in fullface view fully exposed, very close to anterior margin of head, widely separated from each other by clypeal disc; clypeal disc trapezoidal, raised, and flattened or weakly concave medially, margined laterally with a sharp edge running from mandibular insertion to junction with frons; mandibles subtriangular; masticatory margin of mandible with small peg-like teeth; inner surface of mandible with peg-like teeth (modified setae) and thick setae; antenna 12-segmented; eye absent (but intercaste often with reduced eye consisting of a few ommatidia); mesosoma in lateral view with almost straight dorsal outline, or promesonotum feebly raised; promesonotal suture present and flexible; metanotal groove conspicuous dorsally; propodeum unarmed; orifice of propodeal spiracle round, situated relatively low on lateral face of propodeum; propodeal lobe absent; waist consisting of two small segments (petiole and postpetiole) which have clearly differentiated posterior faces; petiole without anterior peduncle; its tergosternal fusion complete, without trace of sutrue; its sternite swollen strongly and forming anteroventral flange; petiolar and postpetiolar spiracles located on the side of anteriormost part of the petiolar and postpetiolar nodes, respectively; postpetiole almost as long as or longer than broad, not distinctly broader than petiole, lacking tergosternal fusion; its sternite swollen strongly; abdominal spiracles V–VII concealed by the posterior margins of the preceding segments; pygidium large, convex across its width, laterally and posteriorly unarmed; sting well developed.</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFCBB460FFC24" blockId="22.[151,1437,151,1005]" pageId="22" pageNumber="23">
<emphasis id="B925EA8FFF97FFC2F5BAFCBB4525FC94" bold="true" box="[199,383,836,861]" pageId="22" pageNumber="23">Differentiation.</emphasis>
The worker of 
<taxonomicName id="4C514D1EFF97FFC2F74AFCBB46F3FC95" box="[567,681,836,860]" class="Insecta" family="Formicidae" genus="Protanilla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B925EA8FFF97FFC2F74AFCBB46F3FC95" box="[567,681,836,860]" italics="true" pageId="22" pageNumber="23">Protanilla</emphasis>
</taxonomicName>
is easily distinguished from that of the other genera known from 
<collectingCountry id="F346760DFF97FFC2F5EAFC9844ACFC49" box="[151,246,871,896]" name="Vietnam" pageId="22" pageNumber="23">Vietnam</collectingCountry>
by a combination of features: antennal sockets in full-face view fully exposed; clypeal disc trapezoidal, raised and flattened, margined laterally with a sharp edge; inner surface of mandible with peg-like teeth; promesonotal suture present and flexible; waist consisting of two small segments (petiole and postpetiole) which have clearly diffintiated posterior faces.
</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFBE3464AFBFC" blockId="22.[199,528,1052,1077]" box="[199,528,1052,1077]" pageId="22" pageNumber="23">
<emphasis id="B925EA8FFF97FFC2F5BAFBE3464AFBFC" bold="true" box="[199,528,1052,1077]" pageId="22" pageNumber="23">Vietnamese species (3 spp.).</emphasis>
</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFB9A45FFFB51" blockId="22.[199,489,1124,1207]" pageId="22" pageNumber="23">
<emphasis id="B925EA8FFF97FFC2F5BAFB9A4482FBB3" box="[199,216,1125,1146]" italics="true" pageId="22" pageNumber="23">P.</emphasis>
sp. eg-1 (Tay Yen Tu). 
<emphasis id="B925EA8FFF97FFC2F5BAFB7C4482FB51" box="[199,216,1155,1176]" italics="true" pageId="22" pageNumber="23">P.</emphasis>
sp. eg-3 (Cat Tien).
</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFB5D45B3FB7E" blockId="22.[199,489,1124,1207]" box="[199,489,1185,1207]" pageId="22" pageNumber="23">
<emphasis id="B925EA8FFF97FFC2F5BAFB5D4482FB7E" box="[199,216,1186,1207]" italics="true" pageId="22" pageNumber="23">P.</emphasis>
sp. eg-4 (Bidoup-Nui Ba).
</paragraph>
<paragraph id="8BEE369DFF97FFC2F5BAFB1B4642FAE9" blockId="22.[151,1436,1252,1312]" pageId="22" pageNumber="23">
<emphasis id="B925EA8FFF97FFC2F5BAFB1B4510FB34" bold="true" box="[199,330,1252,1277]" pageId="22" pageNumber="23">Bionomics.</emphasis>
<taxonomicName id="4C514D1EFF97FFC2F42CFB1B4599FB35" box="[337,451,1252,1276]" class="Insecta" family="Formicidae" genus="Protanilla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B925EA8FFF97FFC2F42CFB1B4599FB35" box="[337,451,1252,1276]" italics="true" pageId="22" pageNumber="23">Protanilla</emphasis>
</taxonomicName>
spp. usually occur in well-developed forests but sometimes in forest edges. They nest in rotten wood fragments and in soil.
</paragraph>
</subSubSection>
</treatment>
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