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<document id="746C71A46481320F06FB9B88AE8F0FFC" ID-DOI="10.5281/zenodo.3677235" ID-GBIF-Dataset="1ee19d5d-27cf-4c93-989c-3fe3eb467203" ID-ISSN="1942-1354" ID-Zenodo-Dep="3677235" ID-ZooBank="BA35690A-FC73-4E5A-A805-FE9550275FEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582218729488" checkinUser="felipe" docAuthor="Cong, Qian, Zhang, Jing, Shen, Jinhui &amp; Grishin, Nick V." docDate="2019" docId="038E29227E7BFFF4FF3F7178DBEBFC25" docLanguage="en" docName="InsectaMundi.0731.1-56.pdf.imf" docOrigin="Insecta Mundi 731 (731)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Aurivittia Grishin 2019, new genus" docType="treatment" docUuid="C2C0299C-67C8-4F0E-ACB8-29A7292E10BD" docUuidSource="ZooBank" docVersion="11" lastPageNumber="9" masterDocId="FFB7515A7E72FFFEFFFF732ADF1EFFFB" masterDocTitle="Fifty new genera of Hesperiidae (Lepidoptera)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="8" updateTime="1698720609646" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="D1DBB3918F79DBDCFF6A6245E380C0D3">Fifty new genera of Hesperiidae (Lepidoptera)</mods:title>
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<mods:namePart id="F1E40AD2CD8CF94A9A4591E3AB533D1A">Cong, Qian</mods:namePart>
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<mods:namePart id="4C65607512125A4CA8651F0649238CE5">Zhang, Jing</mods:namePart>
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<mods:namePart id="17262CE639110C94CA7F31B3B24FD9D4">Shen, Jinhui</mods:namePart>
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<mods:namePart id="7E7EBC55715A08A802C10DA510585ADB">Grishin, Nick V.</mods:namePart>
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<mods:title id="F4400454BE822553C85C6015287FCCD4">Insecta Mundi</mods:title>
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<mods:number id="3BA7F27BFFDEAC1AA80B9673985E9C14">2019-10-11</mods:number>
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<treatment id="038E29227E7BFFF4FF3F7178DBEBFC25" ID-DOI="http://doi.org/10.5281/zenodo.3681691" ID-GBIF-Taxon="162238038" ID-Zenodo-Dep="3681691" LSID="urn:lsid:zoobank.org:act:C2C0299C-67C8-4F0E-ACB8-29A7292E10BD" httpUri="http://treatment.plazi.org/id/038E29227E7BFFF4FF3F7178DBEBFC25" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<subSubSection id="C33DCBBF7E7BFFF7FF3F7178DD44FD91" box="[192,602,594,618]" pageId="9" pageNumber="8" type="nomenclature">
<paragraph id="8B9898347E7BFFF7FF3F7178DD44FD91" blockId="9.[192,602,594,618]" box="[192,602,594,618]" pageId="9" pageNumber="8">
<heading id="D0D02F587E7BFFF7FF3F7178DD44FD91" bold="true" box="[192,602,594,618]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
<emphasis id="B95344267E7BFFF7FF3F7178DD44FD91" bold="true" box="[192,602,594,618]" pageId="9" pageNumber="8">
<taxonomicName id="4C27E3B77E7BFFF7FF3F7178DEA4FD91" authority="Grishin" authorityName="Grishin" authorityYear="2019" box="[192,442,594,618]" class="Insecta" family="Hesperiidae" genus="Aurivittia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95344267E7BFFF7FF3F7178DE57FD91" bold="true" box="[192,329,594,618]" italics="true" pageId="9" pageNumber="8">Aurivittia</emphasis>
Grishin
</taxonomicName>
,
<taxonomicNameLabel id="A260F95D7E7BFFF7FE377178DD44FD91" box="[456,602,594,618]" pageId="9" pageNumber="8" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C33DCBBF7E7BFFF7FF3F7155DCC9FD6C" box="[192,983,639,663]" pageId="9" pageNumber="8" type="description">
<paragraph id="8B9898347E7BFFF7FF3F7155DCC9FD6C" blockId="9.[192,983,639,663]" box="[192,983,639,663]" pageId="9" pageNumber="8">
http://zoobank.org/
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<subSubSection id="C33DCBBF7E7BFFF4FF3F719EDB82FD90" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="type_taxon">
<paragraph id="8B9898347E7BFFF7FF3F719EDC21FD36" blockId="9.[192,831,692,717]" box="[192,831,692,717]" pageId="9" pageNumber="8">
<emphasis id="B95344267E7BFFF7FF3F719EDE69FD37" bold="true" box="[192,375,692,716]" pageId="9" pageNumber="8">
<typeStatus id="549C26967E7BFFF7FF3F719EDE1AFD37" box="[192,260,692,716]" pageId="9" pageNumber="8">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C27E3B77E7BFFF7FE81719EDC25FD36" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[382,827,692,717]" class="Insecta" family="Hesperiidae" genus="Plesioneura" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="aurivittata">
<emphasis id="B95344267E7BFFF7FE81719EDD84FD37" box="[382,666,692,716]" italics="true" pageId="9" pageNumber="8">Plesioneura aurivittata</emphasis>
Moore, 1878
</taxonomicName>
.
</paragraph>
<paragraph id="8B9898347E7BFFF4FF3F71F5DB82FD90" blockId="9.[192,1440,735,887]" lastBlockId="10.[192,1440,221,619]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<emphasis id="B95344267E7BFFF7FF3F71F5DE4DFD0C" bold="true" box="[192,339,735,759]" pageId="9" pageNumber="8">Definition.</emphasis>
Surprisingly, a possible sister genus to
<taxonomicName id="4C27E3B77E7BFFF7FCCB71F5DB01FD0C" ID-CoL="8ZZBP" authority="Evans, 1935" authorityName="Evans" authorityYear="1935" box="[820,1055,735,759]" class="Insecta" family="Hesperiidae" genus="Alenia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E7BFFF7FCCB71F5DC9DFD0C" box="[820,899,735,759]" italics="true" pageId="9" pageNumber="8">Alenia</emphasis>
Evans, 1935
</taxonomicName>
(
<typeStatus id="549C26967E7BFFF7FBD171F5DB7AFD0C" box="[1070,1124,735,759]" pageId="9" pageNumber="8">type</typeStatus>
species:
<taxonomicName id="4C27E3B77E7BFFF7FB3271CADE7DFCEC" ID-CoL="986RN" authority="Trimen, 1868" authorityName="Trimen" authorityYear="1868" class="Insecta" family="Hesperiidae" genus="Pyrgus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="sandaster">
<emphasis id="B95344267E7BFFF7FB3271CADABEFD0C" box="[1229,1440,735,759]" italics="true" pageId="9" pageNumber="8">Pyrgus sandaster</emphasis>
Trimen, 1868
</taxonomicName>
), and in the same clade with
<taxonomicName id="4C27E3B77E7BFFF7FD4971D5DCDDFCEC" authority="Strand, 1911" authorityName="Strand" authorityYear="1911" box="[694,963,767,791]" class="Insecta" family="Hesperiidae" genus="Apallaga" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E7BFFF7FD4971D5DC3AFCEC" box="[694,804,767,791]" italics="true" pageId="9" pageNumber="8">Apallaga</emphasis>
Strand, 1911
</taxonomicName>
(
<typeStatus id="549C26967E7BFFF7FC2F71D5DB1BFCEC" box="[976,1029,767,791]" pageId="9" pageNumber="8">type</typeStatus>
species
<taxonomicName id="4C27E3B77E7BFFF7FB9A71D5DFE3FCCC" authority="Strand, 1911" authorityName="Strand" authorityYear="1911" class="Insecta" family="Hesperiidae" genus="Apallaga" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="separata">
<emphasis id="B95344267E7BFFF7FB9A71D5DA5EFCEC" box="[1125,1344,767,791]" italics="true" pageId="9" pageNumber="8">Apallaga separata</emphasis>
Strand, 1911
</taxonomicName>
), but genetically far removed from these and all other taxa (
<figureCitation id="131C84B17E7BFFF7FC0B7035DB23FCCC" box="[1012,1085,799,823]" captionStart="Figure 3" captionStartId="10.[192,275,1906,1928]" captionTargetBox="[193,1437,1173,1872]" captionTargetId="figure@10.[192,1440,1172,1872]" captionTargetPageId="10" captionText="Figure 3. Genomic trees of Tagiadinae. See Fig. 2 legend for notations." figureDoi="http://doi.org/10.5281/zenodo.3677241" httpUri="https://zenodo.org/record/3677241/files/figure.png" pageId="9" pageNumber="8">Fig. 3</figureCitation>
). Species in this genus were formerly placed in
<taxonomicName id="4C27E3B77E7BFFF7FE5C7015DC00FCAC" ID-CoL="8ZXLT" authority="Hubner, [1819]" authorityName="Hubner" authorityYear="1819" box="[419,798,831,855]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E7BFFF7FE5C7015DD7EFCAC" box="[419,608,831,855]" italics="true" pageId="9" pageNumber="8">Celaenorrhinus</emphasis>
Hübner, [1819]
</taxonomicName>
(
<typeStatus id="549C26967E7BFFF7FCD37015DC7CFCAC" box="[812,866,831,855]" pageId="9" pageNumber="8">type</typeStatus>
species
<taxonomicName id="4C27E3B77E7BFFF7FC3A7015DA11FCAC" authority="Stoll, [1781]" authorityName="Stoll" authorityYear="1781" box="[965,1295,831,855]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="eligius">
<emphasis id="B95344267E7BFFF7FC3A7015DB6DFCAC" box="[965,1139,831,855]" italics="true" pageId="9" pageNumber="8">Papilio eligius</emphasis>
Stoll, [1781]
</taxonomicName>
;) and key to B.
<quantity id="4CDF35D17E7BFFF7FF287075DE33FC8C" box="[215,301,863,887]" metricMagnitude="-1" metricUnit="m" metricValue="1.6001999999999998" pageId="9" pageNumber="8" unit="in" value="6.3">6.30 in</quantity>
<bibRefCitation id="EFB6E5C57E7BFFF7FECD704ADECAFC8C" author="Evans" box="[306,468,863,887]" firstAuthor="Evans" pageId="9" pageNumber="8" refId="ref27372" refString="Evans, W. H. 1949. A catalogue of the Hesperiidae from Europe, Asia, and Australia in the British Museum (Natural History). British Museum (Natural History); London. xix + 502 p., 53 pl." type="book" year="1949">Evans (1949)</bibRefCitation>
, notably lacking hair pencil on hind tibiae, which other
<taxonomicName id="4C27E3B77E7BFFF7FB8E7075DA30FC8C" ID-CoL="8ZXLT" authorityName="Hubner" authorityYear="1819" box="[1137,1326,863,887]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E7BFFF7FB8E7075DA30FC8C" box="[1137,1326,863,887]" italics="true" pageId="9" pageNumber="8">Celaenorrhinus</emphasis>
</taxonomicName>
-like taxa possess. Morphologically, distinguished from
<taxonomicName id="4C27E3B77E78FFF4FD0C73F7DCAEFF0E" ID-CoL="8ZXLT" authorityName="Hubner" authorityYear="1819" box="[755,944,221,245]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E78FFF4FD0C73F7DCAEFF0E" box="[755,944,221,245]" italics="true" pageId="10" pageNumber="9">Celaenorrhinus</emphasis>
</taxonomicName>
(sensu stricto) by essentially undivided uncus, sometimes with small knobs (not processes) as arms and the lack of hair pencil on hind tibiae; and from
<taxonomicName id="4C27E3B77E78FFF4FEC8720BDE98FEC2" ID-CoL="8ZZBP" authorityName="Evans" authorityYear="1935" box="[311,390,289,313]" class="Insecta" family="Hesperiidae" genus="Alenia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E78FFF4FEC8720BDE98FEC2" box="[311,390,289,313]" italics="true" pageId="10" pageNumber="9">Alenia</emphasis>
</taxonomicName>
(a genus with undivided uncus) by the shape of valva that is not terminally split like a crab claw, and the presence of yellow band across the forewing. In male genitalia most similar to
<taxonomicName id="4C27E3B77E78FFF4FF3F724FDE31FE86" authorityName="Strand" authorityYear="1911" box="[192,303,357,381]" class="Insecta" family="Hesperiidae" genus="Apallaga" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E78FFF4FF3F724FDE31FE86" box="[192,303,357,381]" italics="true" pageId="10" pageNumber="9">Apallaga</emphasis>
</taxonomicName>
, a genus with many species characterized by undivided uncus, but differs in longer gnathos (not less than half of the uncus length), thicker penis, valva with more robust and broad harpe that is gradually curved dorsad (not ventrad), does not carry any processes and is not forked, but narrows to a single point, and ampulla with a long process (style) along the harpe. In wing patterns, hindwing is not prominently variegated dorsally and is mostly unmarked, forewing with a compact yellow discal band with regular edges that does not extend into the discal cell and frequently with a triplet of apical yellow spots. In DNA, a combination of the following base pairs is diagnostic: aly
<date id="FF99BEF47E78FFF4FB28711BDA5CFDB2" box="[1239,1346,561,585]" pageId="10" pageNumber="9" value="2532-02-01">2532.2.1</date>
:T488A, aly997.12.1:C310A, aly527.19.4:T178A, aly822.15.1:C589A, aly235.8.18:G1178A.
</paragraph>
</subSubSection>
<caption id="DF58C8BC7E7BFFF7FF3F751EDD43F87C" ID-DOI="http://doi.org/10.5281/zenodo.3677239" ID-Zenodo-Dep="3677239" httpUri="https://zenodo.org/record/3677239/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,275,1588,1610]" targetBox="[192,1440,939,1553]" targetPageId="9">
<paragraph id="8B9898347E7BFFF7FF3F751EDD43F87C" blockId="9.[192,1440,1588,1927]" pageId="9" pageNumber="8">
<emphasis id="B95344267E7BFFF7FF3F751EDE30F9B2" bold="true" box="[192,302,1588,1610]" pageId="9" pageNumber="8">Figure 2.</emphasis>
Genomic trees of
<taxonomicName id="4C27E3B77E7BFFF7FE0A751FDD6AF9B0" authorityName="Mabille" authorityYear="1877" box="[501,628,1589,1611]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Eudaminae">Eudaminae</taxonomicName>
and representatives of other
<taxonomicName id="4C27E3B77E7BFFF7FC45751FDB21F9B0" box="[954,1087,1589,1611]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="family">Hesperiidae</taxonomicName>
subfamilies. The trees are built from protein-coding regions in different genomic partitions:
<emphasis id="B95344267E7BFFF7FCB6757BDC7FF99C" bold="true" box="[841,865,1617,1639]" pageId="9" pageNumber="8">a)</emphasis>
Nuclear genome.
<emphasis id="B95344267E7BFFF7FBDC757BDB25F99C" bold="true" box="[1059,1083,1617,1639]" pageId="9" pageNumber="8">b)</emphasis>
Z chromosome.
<emphasis id="B95344267E7BFFF7FB16757BDBE1F99C" bold="true" box="[1257,1279,1617,1639]" pageId="9" pageNumber="8">c)</emphasis>
Mitochondrial genome;
<emphasis id="B95344267E7BFFF7FEDC7544DE22F97F" bold="true" box="[291,316,1646,1668]" pageId="9" pageNumber="8">d)</emphasis>
COI barcode. The trees are rooted with
<taxonomicName id="4C27E3B77E7BFFF7FCFA7545DCD0F978" box="[773,974,1646,1668]" class="Insecta" family="Papilionidae" genus="Pterourus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="glaucus">
<emphasis id="B95344267E7BFFF7FCFA7545DCD0F978" box="[773,974,1646,1668]" italics="true" pageId="9" pageNumber="8">Pterourus glaucus</emphasis>
</taxonomicName>
(NVG-1670), not shown. See Table 1 and Table S1 in the Supplemental file deposited at https://osf.io/5cfht/?view_only=21eb53b6f8f344afaee3de2be90bf 5d2 for additional data about these specimens. Where possible, taxa are ordered similarly to that in the nuclear genome tree. Names of species placed in new genera described in this work are highlighted in yellow and clades of these genera are colored in red. Clades for genera where these species were placed previously are colored in blue and green arrow points from the former genus to the new genus (only on nuclear genome tree). Names of genera resurrected from synonymy are highlighted in cyan. Names of genera that form new genus-species combinations proposed in this work are highlighted in magenta. Statistical support values are shown by nodes in all but the COI barcode trees. COI barcode NJ dendrogram is given for comparison and is not expected to reflect phylogeny. The same notations are used in Fig. 2–13.
</paragraph>
</caption>
<subSubSection id="C33DCBBF7E78FFF4FF3F71AFDC4FFCBD" pageId="10" pageNumber="9" type="etymology">
<paragraph id="8B9898347E78FFF4FF3F71AFDC72FD3B" blockId="10.[192,1440,645,704]" pageId="10" pageNumber="9">
<emphasis id="B95344267E78FFF4FF3F71AFDE45FD66" bold="true" box="[192,347,645,669]" pageId="10" pageNumber="9">Etymology.</emphasis>
The name is a feminine noun in the nominative singular, formed to reflect the golden stripe on the forewing, similar to the name of the
<typeStatus id="549C26967E78FFF4FD2E7182DC18FD3B" box="[721,774,680,704]" pageId="10" pageNumber="9">type</typeStatus>
species.
</paragraph>
<paragraph id="8B9898347E78FFF4FF3F71F3DD1EFCEF" blockId="10.[192,1440,729,788]" pageId="10" pageNumber="9">
<emphasis id="B95344267E78FFF4FF3F71F3DEADFD0A" bold="true" box="[192,435,729,753]" pageId="10" pageNumber="9">Species included.</emphasis>
The
<typeStatus id="549C26967E78FFF4FE0971F0DD35FD09" box="[502,555,730,754]" pageId="10" pageNumber="9">type</typeStatus>
species,
<taxonomicName id="4C27E3B77E78FFF4FD6271F3DB43FD09" authority="Distant, 1882" authorityName="Distant" authorityYear="1882" box="[669,1117,729,754]" class="Insecta" family="Hesperiidae" genus="Plesioneura" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="cameroni">
<emphasis id="B95344267E78FFF4FD6271F3DCB9FD0A" box="[669,935,729,753]" italics="true" pageId="10" pageNumber="9">Plesioneura cameroni</emphasis>
Distant, 1882
</taxonomicName>
, and
<taxonomicName id="4C27E3B77E78FFF4FB5B71F3DEE2FCEF" authority="Devyatkin, 1998" authorityName="Devyatkin" authorityYear="1998" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B95344267E78FFF4FB5B71F3DE36FCE8" italics="true" pageId="10" pageNumber="9">Celaenorrhinus vietnamicus</emphasis>
Devyatkin, 1998
</taxonomicName>
.
</paragraph>
<paragraph id="8B9898347E78FFF4FF3F7004DC4FFCBD" blockId="10.[192,849,814,838]" box="[192,849,814,838]" pageId="10" pageNumber="9">
<emphasis id="B95344267E78FFF4FF3F7004DE62FCBD" bold="true" box="[192,380,814,838]" pageId="10" pageNumber="9">Parent taxon.</emphasis>
Tribe
<taxonomicName id="4C27E3B77E78FFF4FE337004DC53FCBD" authority="Swinhoe, 1912" authorityName="Swinhoe" authorityYear="1912" box="[460,845,814,838]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Celaenorrhinini">Celaenorrhinini Swinhoe, 1912</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33DCBBF7E78FFF4FF3F7075DBEBFC25" pageId="10" pageNumber="9" type="discussion">
<paragraph id="8B9898347E78FFF4FF3F7075DBEBFC25" blockId="10.[192,1440,863,990]" pageId="10" pageNumber="9">
<emphasis id="B95344267E78FFF4FF3F7075DE47FC8C" bold="true" box="[192,345,863,887]" pageId="10" pageNumber="9">Comments.</emphasis>
These south Asian butterflies are not so prominently distinct from the striped species of
<taxonomicName id="4C27E3B77E78FFF4FF3F70ABDE63FC62" authorityName="Hubner" authorityYear="1819" box="[192,381,897,921]" class="Insecta" family="Hesperiidae" genus="Celaenorrhinus" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E78FFF4FF3F70ABDE63FC62" box="[192,381,897,921]" italics="true" pageId="10" pageNumber="9">Celaenorrhinus</emphasis>
</taxonomicName>
. Therefore, their uniqueness was not apparent prior to our genomic study, although genitalic features revealed
<emphasis id="B95344267E78FFF4FDF9708EDD90FC40" box="[518,654,932,955]" italics="true" pageId="10" pageNumber="9">a posteriori</emphasis>
can diagnose the genus morphologically. Phylogenetic placement of this Asian genus in the African clade of
<taxonomicName id="4C27E3B77E78FFF4FD3970ECDB5DFC25" authority="Swinhoe, 1912" authorityName="Swinhoe" authorityYear="1912" box="[710,1091,966,990]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Celaenorrhinini">Celaenorrhinini Swinhoe, 1912</taxonomicName>
is interesting.
</paragraph>
</subSubSection>
</treatment>
</document>
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