treatments-xml/data/F5/D4/3E/F5D43E8E278D5AE28249E292A2DB4ACF.xml

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<mods:title id="84405DF175DB9E51131197AAEED2A442">Revision of the southern Andean genus Sadocus Sorensen, 1886 (Opiliones, Gonyleptidae, Pachylinae)</mods:title>
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<mods:namePart id="3A6EF1521A9F3671E3951BC4E4B8BD67">Pessoa-Silva, Marilia</mods:namePart>
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<mods:namePart id="D0AAC043798CD34A7CB9B7A760B0048A">Hara, Marcos Ryotaro</mods:namePart>
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<mods:namePart id="E7A814B9C416436524DB3A53058C1F50">Pinto-da-Rocha, Ricardo</mods:namePart>
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<treatment id="F5D43E8E278D5AE28249E292A2DB4ACF" ID-GBIF-Taxon="180671092" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF" httpUri="http://treatment.plazi.org/id/F5D43E8E278D5AE28249E292A2DB4ACF" lastPageNumber="91" pageId="0" pageNumber="91" scope_family="Gonyleptidae" scope_order="Opiliones" scope_subFamily="Pachylinae">
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<paragraph id="210B2CE4C3D0F78DFC21DE50715E60B8" pageId="0" pageNumber="91">
<taxonomicName id="654CD3B698322E688A9233DA1406404B" ID-CoL="7YVYF" LSID="F5D43E8E-278D-5AE2-8249-E292A2DB4ACF" authority="(Gervais, 1847)" baseAuthorityName="Gervais" baseAuthorityYear="1847" class="Arachnida" family="Gonyleptidae" genus="Sadocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sadocus asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">Sadocus asperatus (Gervais, 1847)</taxonomicName>
<figureCitation id="B61218D3194DF7582FA617C57995D24F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geographical records of distribution of Sadocus species." figureDoi="10.3897/zookeys.1025.57806.figure2" httpUri="https://binary.pensoft.net/fig/522912" pageId="0" pageNumber="91">Figures 2</figureCitation>
<figureCitation id="CEF6EBFB24DF608155AE0DFDE873235A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sadocus asperatus (Gervais). Male (CAS 9055035) A Habitus, dorsal view B-E right leg IV, Trochanter-tibia IV B dorsal view C prolateral view D ventral view E retro-lateral view. Scale: 1 mm." figureDoi="10.3897/zookeys.1025.57806.figure3" httpUri="https://binary.pensoft.net/fig/522913" pageId="0" pageNumber="91">, 3A-E</figureCitation>
<figureCitation id="9B2E138B4450D8B5C1452D72CFB50CDC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Female specimens of Sadocus A, B S. asperatus (CAS- 9055035) A dorsal view B lateral view C, D S. dilatatus (MZSP- 8022) C dorsal view D lateral view E, F S. funestus (AMNH) E dorsal view F lateral view G, H S. ingens (MZSP- 36965) G dorsal view H lateral view I, J S. polyacanthus (AMNH) I dorsal view J lateral view K Drawing of species inquirenda Carampangue nigronotatum extracted from Mello-Leitao 1943." figureDoi="10.3897/zookeys.1025.57806.figure8" httpUri="https://binary.pensoft.net/fig/522918" pageId="0" pageNumber="91">, 8A, B</figureCitation>
<figureCitation id="625173F9847330064BF86E44A1FC88D6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photographs of living specimens of Sadocus A S. asperatus (ma) B S. asperatus (fe) C S. asperatus (ma) D S. dilatatus (ma) E S. asperatus (fe) F S. funestus (ma). A, B taken by R. Pinto-da-Rocha, C-F taken by Pedro H. Martins." figureDoi="10.3897/zookeys.1025.57806.figure9" httpUri="https://binary.pensoft.net/fig/522919" pageId="0" pageNumber="91">, 9A-C, E</figureCitation>
<figureCitation id="B08283B349C318E4E2311E3981DE5891" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distal part of penis Sadocus and Discocyrtus, dorsal and ventral views, respectively A, B S. asperatus, macrosetae colorized and numbered C, D S. ingens E, F S. polyacanthus G, H S. dilatatus I, J S. funestus K, L D. catharinensis. Scale bar of drawings: 0.1 mm." figureDoi="10.3897/zookeys.1025.57806.figure11" httpUri="https://binary.pensoft.net/fig/522921" pageId="0" pageNumber="91">, 11A, B</figureCitation>
<figureCitation id="90869553B28CC9734B11D9F29E010BAF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distal part of ovipositor of Sadocus A-C S. asperatus A upper view B villi C setae D-F S. funestus D upper view E lateral view F seta G-I S. ingens G upper view H, I setae J-L S. polyacanthus J upper view K lateral view L setae." figureDoi="10.3897/zookeys.1025.57806.figure12" httpUri="https://binary.pensoft.net/fig/522922" pageId="0" pageNumber="91">, 12A-C</figureCitation>
</paragraph>
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<paragraph id="B8F902FFB095D89F97CEC1558CB3607F" pageId="0" pageNumber="91">
<taxonomicName id="99CAC913FE7C8BF9F49B373600756CDE" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">Gonyleptes asperatus</taxonomicName>
Gervais, 1847: 577 [desc]; 1849: 26, pl. 1, fig 9 [rdesc]; 
<bibRefCitation id="FB699B51699FAF975C1F76E000E86172" DOI="https://doi.org/10.1080/00222937308696775" author="Butler, AG" journalOrPublisher="Cladistics" pageId="0" pageNumber="91" refId="B9" refString="Butler, AG, 1873. A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11(62): 112-117. https://doi.org/10.1080/00222937308696775" title="A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11 (62): 112 - 117." url="https://doi.org/10.1080/00222937308696775" year="1873">Butler 1873</bibRefCitation>
: 113 [cat]; 
<bibRefCitation id="8D608DF92205E3BFA384FE38B3A35573" author="Sorensen, WE" journalOrPublisher="Ergebnisse der Hamburger Magalhaensischen Sammelreise" pageId="0" pageNumber="91" pagination="1 - 36" refId="B99" refString="Sorensen, WE, 1902. Gonyleptiden (Opiliones, Laniatores). Ergebnisse der Hamburger Magalhaensischen Sammelreise 6 (5): 1 - 36" title="Gonyleptiden (Opiliones, Laniatores)." volume="6" year="1902">
<normalizedToken id="A36BF9D54BA3FCAF03A0F5C94205C23B" originalValue="Sørensen">Sorensen</normalizedToken>
1902
</bibRefCitation>
: 17 [syst]. Transferred to 
<taxonomicName id="5BEEA71AB0A47A599327E5EBBEAFC204" class="Arachnida" family="Gonyleptidae" genus="Lycomedes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedes" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="genus">Lycomedes</taxonomicName>
by 
<bibRefCitation id="3C104510BA849BCB32B734D4D2903D49" author="Sorensen, WE" journalOrPublisher="Ergebnisse der Hamburger Magalhaensischen Sammelreise" pageId="0" pageNumber="91" pagination="1 - 36" refId="B99" refString="Sorensen, WE, 1902. Gonyleptiden (Opiliones, Laniatores). Ergebnisse der Hamburger Magalhaensischen Sammelreise 6 (5): 1 - 36" title="Gonyleptiden (Opiliones, Laniatores)." volume="6" year="1902">
<normalizedToken id="D8298F9D3B0D4B3930340F643D4786CB" originalValue="Sørensen">Sorensen</normalizedToken>
1902
</bibRefCitation>
. (MNHN, type lost, not examined) (RPR visited the museum but the curator was unable to find the type material)
</paragraph>
<paragraph id="EA2E750B61488680F252A8BC44190744" pageId="0" pageNumber="91">
<taxonomicName id="5DD83823E47D16229B6AA553FF4B2CBB" class="Arachnida" family="Gonyleptidae" genus="Lycomedes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedes asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">Lycomedes asperatus</taxonomicName>
: 
<normalizedToken id="B630AFFCEF21791EC8482B6AC7D95AAF" originalValue="Sørensen">Sorensen</normalizedToken>
, 1902: 17 [syst]; 
<bibRefCitation id="6FA9AB3CCBEA036A4769738A4978DD59" author="Roewer, CF" journalOrPublisher="&quot; Ciencias Zoologicas" pageId="0" pageNumber="91" refId="B77" refString="Roewer, CF, 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79A(4-5): 1-473." title="Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79 A (4 - 5): 1 - 473." year="1913">Roewer 1913</bibRefCitation>
: 127-130, fig 57 [key, rdesc].
</paragraph>
<paragraph id="F526DA0E0E4C9A88AD3816A7CCFD75DD" pageId="0" pageNumber="91">
<taxonomicName id="201665D456FEBE6BB8F90736B76FB6FC" class="Arachnida" family="Gonyleptidae" genus="Lycomedicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedicus asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">Lycomedicus asperatus</taxonomicName>
: Roewer, 1923: 442, fig 556 [rdesc]; 1925: 17 [cit]; 1929: 213 [cat]; 
<bibRefCitation id="DEAC5A96D174D8AA018A6B79E586AF0D" author="Mello-Leitao, CF" journalOrPublisher="Annaes da Academia brasileira de Sciencias" pageId="0" pageNumber="91" pagination="83 - 97" refId="B50" refString="Mello-Leitao, CF, 1931b. Nota sobre arachnideos argentinos. III. Opilioes novos ou criticos. IV. Aranhas novas. Annaes da Academia brasileira de Sciencias 3 (2): 83 - 97" title="Nota sobre arachnideos argentinos. III. Opilioes novos ou criticos. IV. Aranhas novas." volume="3" year="1931 b">
<normalizedToken id="79E121DAF5762A9E27150A53E06396E0" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
1931b
</bibRefCitation>
: 84 [cit]; 
<bibRefCitation id="A782DE066001CB3034F35CC21D48D1F8" author="Canals, J" journalOrPublisher="Revista Chilena de Historia Natural" pageId="0" pageNumber="91" pagination="68 - 71" refId="B12" refString="Canals, J, 1936. Los Opiliones de Chile. Revista Chilena de Historia Natural 39: 68 - 71" title="Los Opiliones de Chile." volume="39" year="1936">Canals 1936</bibRefCitation>
: 69 [cat]; 
<bibRefCitation id="BA8D64BEB7157ED6F950F50BC29DFDA0" DOI="https://doi.org/10.5962/bhl.part.1498" author="Roewer, CF" journalOrPublisher="Abhandlungen der Naturwissenschaftlichen Verein zu Bremen" pageId="0" pageNumber="91" refId="B82" refString="Roewer, CF, 1938. https://doi.org/10.5962/bhl.part.1498" url="https://doi.org/10.5962/bhl.part.1498" year="1938">Roewer 1938</bibRefCitation>
: 6 [cat]; 
<bibRefCitation id="9A81833180DFDA26BD108C5536061FDA" author="Mello-Leitao, CF" journalOrPublisher="Physis" pageId="0" pageNumber="91" pagination="601 - 630" refId="B54" refString="Mello-Leitao, CF, 1939. Les arachnides et la zoogeographie de l'Argentine. Physis 17: 601 - 630" title="Les arachnides et la zoogeographie de l'Argentine." volume="17" year="1939">
<normalizedToken id="5A9BC9B18E38D4E90B9C0E56EBA9293A" originalValue="Mello-Leitão">Mello-Leitao</normalizedToken>
1939
</bibRefCitation>
: 624 [cat]; 
<bibRefCitation id="15573B4AB95913B58A6F88153A9FF14E" author="Soares, BAM" journalOrPublisher="Arquivos de zoologia do Estado de Sao Paulo" pageId="0" pageNumber="91" pagination="225 - 302" refId="B92" refString="Soares, BAM, Soares, HEM, 1954. Monografia dos generos de opilioes neotropicos III. Arquivos de zoologia do Estado de Sao Paulo 8 (9): 225 - 302" title="Monografia dos generos de opilioes neotropicos III." volume="8" year="1954">Soares and Soares 1954</bibRefCitation>
: 270 [cat]; 
<bibRefCitation id="EE7839053E959F5558CA07B55FE45851" author="Ringuelet, RA" journalOrPublisher="&quot; Ciencias Zoologicas" pageId="0" pageNumber="91" pagination="127 - 439" refId="B76" refString="Ringuelet, RA, 1959. Los aracnidos Argentinos del orden Opiliones. Revista del Museo Argentino de Ciencias Naturales &quot;Bernardino Rivadavia. &quot; Ciencias Zoologicas 5 (2): 127 - 439" title="Los aracnidos Argentinos del orden Opiliones. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia." volume="5" year="1959">Ringuelet 1959</bibRefCitation>
: 329, fig 44 [rdesc]; H. 
<bibRefCitation id="03094FD807D9902FCFE5829F8785C6BC" author="Soares, HEM" journalOrPublisher="Papeis avulsos de Zoologia" pageId="0" pageNumber="91" pagination="259 - 272" refId="B96" refString="Soares, HEM, 1968. Contribuicao ao estudo dos opilioes do Chile (Opiliones: Gonyleptidae, Triaenonychidae). Papeis avulsos de Zoologia 21 (27): 259 - 272" title="Contribuicao ao estudo dos opilioes do Chile (Opiliones: Gonyleptidae, Triaenonychidae)." volume="21" year="1968">Soares 1968</bibRefCitation>
: 264 [cit]; 
<bibRefCitation id="D50CBA4C59A1A83D251545EFDD9E98D4" author="Cekalovic-K, TN" journalOrPublisher="Noticiario Mensual" pageId="0" pageNumber="91" pagination="5 - 11" refId="B15" refString="Cekalovic-K, TN, 1968. Conocimiento actual de los opiliones chilenos. Noticiario Mensual 12 (138): 5 - 11" title="Conocimiento actual de los opiliones chilenos." volume="12" year="1968">Cekalovic 1968</bibRefCitation>
: 8 [cat]; 1985: 18 [cat].
</paragraph>
<paragraph id="727BA0752F0022D7067CB70A5FBB64A4" pageId="0" pageNumber="91">
<taxonomicName id="7DA19535FAEB855F1CC9DFB25D88FA5F" baseAuthorityName="Gervais" baseAuthorityYear="1847" class="Arachnida" family="Gonyleptidae" genus="Sadocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sadocus asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">Sadocus asperatus</taxonomicName>
: Kury, 2003: 191 [cat]; 
<bibRefCitation id="DF2BFD06E7B81840B53CE9B300C91CA9" author="Kury, AB" journalOrPublisher="Self published, Rio de Janeiro" pageId="0" pageNumber="91" publicationUrl="https://wcolite.com/" refId="B41" refString="Kury, AB, Mendes, AC, Cardoso, L, Kury, MS, Granado Ade, A, Giribet, G, 2020b. World Catalogue of Opiliones. WCO-Lite version 1.2.1. https://wcolite.com/" title="World Catalogue of Opiliones. WCO-Lite version 1.2.1." url="https://wcolite.com/" year="2020 b">Kury et al. 2020b</bibRefCitation>
[cat].
</paragraph>
<paragraph id="3F61232167074C5C1F7CCC55030F1F82" pageId="0" pageNumber="91">
<taxonomicName id="EA23B2422053BFC8334AE335C2C85E21" authorityName="Sorensen" authorityYear="1886" class="Arachnida" family="Gonyleptidae" genus="Sadocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sadocus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="genus">Sadocus</taxonomicName>
(?) 
<taxonomicName id="9687FB7C4CB5ED182CA805C4E1F44A2C" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes subsimilis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="subsimilis">Gonyleptes subsimilis</taxonomicName>
: 
<normalizedToken id="441B78B22B851092EE1DB156C497FC26" originalValue="Sørensen">Sorensen</normalizedToken>
, 1902: 17 [cit]; 
<bibRefCitation id="D640210E5E4CEF6F76A407A8B10423FB" author="Roewer, CF" journalOrPublisher="&quot; Ciencias Zoologicas" pageId="0" pageNumber="91" refId="B77" refString="Roewer, CF, 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79A(4-5): 1-473." title="Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79 A (4 - 5): 1 - 473." year="1913">Roewer 1913</bibRefCitation>
: 127-130, fig 57 [syst]. Synonymy with 
<taxonomicName id="F90E23D25BB4B279CF92BFD1E86941C5" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">L. asperatus</taxonomicName>
established by 
<bibRefCitation id="DC4085A143180E8094281B7A13DC782F" author="Roewer, CF" journalOrPublisher="&quot; Ciencias Zoologicas" pageId="0" pageNumber="91" refId="B77" refString="Roewer, CF, 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79A(4-5): 1-473." title="Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79 A (4 - 5): 1 - 473." year="1913">Roewer 1913</bibRefCitation>
.
</paragraph>
<paragraph id="70F2BE6480A0C76B651D82D7AC18D773" pageId="0" pageNumber="91">
<taxonomicName id="FADF879400518E01096299344B53FB5F" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes polyacanthoides" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="polyacanthoides">Gonyleptes polyacanthoides</taxonomicName>
Gervais, 1847: 577 [desc]; 
<bibRefCitation id="E76973DDE82F93DB512AE4CB12718F36" DOI="https://doi.org/10.1080/00222937308696775" author="Butler, AG" journalOrPublisher="Cladistics" pageId="0" pageNumber="91" refId="B9" refString="Butler, AG, 1873. A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11(62): 112-117. https://doi.org/10.1080/00222937308696775" title="A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11 (62): 112 - 117." url="https://doi.org/10.1080/00222937308696775" year="1873">Butler 1873</bibRefCitation>
: 114 [cat]. Synonymy with 
<taxonomicName id="1842E7CA5B604D91B5650C66F3C517DC" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="G. subsimilis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="subsimilis">G. subsimilis</taxonomicName>
established by 
<bibRefCitation id="4AFF2A1FCB8A489AEDCD281F722D22A8" DOI="https://doi.org/10.1080/00222937308696775" author="Butler, AG" journalOrPublisher="Cladistics" pageId="0" pageNumber="91" refId="B9" refString="Butler, AG, 1873. A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11(62): 112-117. https://doi.org/10.1080/00222937308696775" title="A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11 (62): 112 - 117." url="https://doi.org/10.1080/00222937308696775" year="1873">Butler 1873</bibRefCitation>
. (MNHN, type lost, material not examined) (RPR visited the Museum but the curator was unable to find the type material)
</paragraph>
<paragraph id="2AF44D91E8C3C70B84B76860FC547079" pageId="0" pageNumber="91">
<taxonomicName id="E7ABECD0AF3E39F39A54DFDB0427B508" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes subsimilis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="subsimilis">Gonyleptes subsimilis</taxonomicName>
Gervais, 1849 [desc]: 25, pl. 1, fig 8; 
<bibRefCitation id="28A71CD8200FBD5D91EA701B727D19A0" DOI="https://doi.org/10.1080/00222937308696775" author="Butler, AG" journalOrPublisher="Cladistics" pageId="0" pageNumber="91" refId="B9" refString="Butler, AG, 1873. A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11(62): 112-117. https://doi.org/10.1080/00222937308696775" title="A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11 (62): 112 - 117." url="https://doi.org/10.1080/00222937308696775" year="1873">Butler 1873</bibRefCitation>
: 114 [cat]; 
<bibRefCitation id="4BD540DB1B341AF1CE1F87066896CA02" author="Sorensen, WE" journalOrPublisher="Ergebnisse der Hamburger Magalhaensischen Sammelreise" pageId="0" pageNumber="91" pagination="1 - 36" refId="B99" refString="Sorensen, WE, 1902. Gonyleptiden (Opiliones, Laniatores). Ergebnisse der Hamburger Magalhaensischen Sammelreise 6 (5): 1 - 36" title="Gonyleptiden (Opiliones, Laniatores)." volume="6" year="1902">
<normalizedToken id="35925013245B680DC3BCD3DF46C16187" originalValue="Sørensen">Sorensen</normalizedToken>
1902
</bibRefCitation>
: 6 [syst], 16 [syst]. Synonymy with 
<taxonomicName id="C6E4914D85916D8F818A91EDA9F1EA1B" authorityName="Sorensen" authorityYear="1886" class="Arachnida" family="Gonyleptidae" genus="Sadocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sadocus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="genus">Sadocus</taxonomicName>
(?) 
<taxonomicName id="C7C171DD24DC198F2EA8190D896043B6" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes subsimilis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="subsimilis">Gonyleptes subsimilis</taxonomicName>
established by 
<bibRefCitation id="86B6ADE9C2B4D16DCB51FA965AE7A8BC" author="Sorensen, WE" journalOrPublisher="Ergebnisse der Hamburger Magalhaensischen Sammelreise" pageId="0" pageNumber="91" pagination="1 - 36" refId="B99" refString="Sorensen, WE, 1902. Gonyleptiden (Opiliones, Laniatores). Ergebnisse der Hamburger Magalhaensischen Sammelreise 6 (5): 1 - 36" title="Gonyleptiden (Opiliones, Laniatores)." volume="6" year="1902">
<normalizedToken id="3E9184360E6FCF095795818D5AD77F56" originalValue="Sørensen">Sorensen</normalizedToken>
1902
</bibRefCitation>
(MNHN, type lost, material not examined) (RPR visited the museum but the curator was unable to find the type material)
</paragraph>
<paragraph id="50F188546FA9DF00FC6BFAF6AEC4F8F6" pageId="0" pageNumber="91">
<taxonomicName id="CD36557A309E9EB41AC0C78EA18A67CB" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes bicornis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="bicornis">Gonyleptes bicornis</taxonomicName>
Gervais, 1849: 21, pl. 1, fig 4a-b [desc]; 
<bibRefCitation id="A1C4FF958FBDC49E462C6CA44913D04D" DOI="https://doi.org/10.1080/00222937308696775" author="Butler, AG" journalOrPublisher="Cladistics" pageId="0" pageNumber="91" refId="B9" refString="Butler, AG, 1873. A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11(62): 112-117. https://doi.org/10.1080/00222937308696775" title="A monographic list of the species of the genus Gonyleptes, with descriptions of three remarkable new species. Annals and Magazine of Natural History (series 4) 11 (62): 112 - 117." url="https://doi.org/10.1080/00222937308696775" year="1873">Butler 1873</bibRefCitation>
: 114 [cat]; 
<bibRefCitation id="3D722F53E8AE6DE50BD51732D3057A4C" author="Sorensen, WE" journalOrPublisher="Ergebnisse der Hamburger Magalhaensischen Sammelreise" pageId="0" pageNumber="91" pagination="1 - 36" refId="B99" refString="Sorensen, WE, 1902. Gonyleptiden (Opiliones, Laniatores). Ergebnisse der Hamburger Magalhaensischen Sammelreise 6 (5): 1 - 36" title="Gonyleptiden (Opiliones, Laniatores)." volume="6" year="1902">
<normalizedToken id="4205568E7C8A0C9054B2465E3AC96D7C" originalValue="Sørensen">Sorensen</normalizedToken>
1902
</bibRefCitation>
: 19 [syst]. (type material depository unknown). syn. nov.
</paragraph>
<paragraph id="D0F87EBC076C1EBF5CDD9731A045337C" pageId="0" pageNumber="91">
<taxonomicName id="42BEB58D2FC79DB7106559AFBD6A6703" class="Arachnida" family="Gonyleptidae" genus="Lycomedes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedes bicornis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="bicornis">Lycomedes bicornis</taxonomicName>
: 
<normalizedToken id="9FE950733156E735588D57BBD9A94DCA" originalValue="Sørensen">Sorensen</normalizedToken>
, 1902: 20, fig 4-4b [rdesc]; 
<bibRefCitation id="05955CC5381464046CFF764D6061EEF0" author="Roewer, CF" journalOrPublisher="&quot; Ciencias Zoologicas" pageId="0" pageNumber="91" refId="B77" refString="Roewer, CF, 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79A(4-5): 1-473." title="Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79 A (4 - 5): 1 - 473." year="1913">Roewer 1913</bibRefCitation>
: 136-137, fig 62 [rdesc].
</paragraph>
<paragraph id="C9B47774B83D1F7F9150A25A4F5981A3" pageId="0" pageNumber="91">
<taxonomicName id="5F2523FC3908683EAA267DA2646EE480" class="Arachnida" family="Gonyleptidae" genus="Lycomedicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedicus bicornis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="bicornis">Lycomedicus bicornis</taxonomicName>
: Roewer, 1923: 445, fig 561 [cit]; 
<bibRefCitation id="5DECBC34BBEDEFDD9245A6B2BAFFF886" author="Canals, J" journalOrPublisher="Revista Chilena de Historia Natural" pageId="0" pageNumber="91" pagination="68 - 71" refId="B12" refString="Canals, J, 1936. Los Opiliones de Chile. Revista Chilena de Historia Natural 39: 68 - 71" title="Los Opiliones de Chile." volume="39" year="1936">Canals 1936</bibRefCitation>
: 69 [cat]; 
<bibRefCitation id="342F7D7781A87EF25F0BFF0379A3BE52" author="Soares, BAM" journalOrPublisher="Arquivos de zoologia do Estado de Sao Paulo" pageId="0" pageNumber="91" pagination="225 - 302" refId="B92" refString="Soares, BAM, Soares, HEM, 1954. Monografia dos generos de opilioes neotropicos III. Arquivos de zoologia do Estado de Sao Paulo 8 (9): 225 - 302" title="Monografia dos generos de opilioes neotropicos III." volume="8" year="1954">Soares and Soares 1954</bibRefCitation>
a: 270 [cat]; 
<bibRefCitation id="04DAB8A9461162D34B0904592F74F4E9" author="Cekalovic-K, TN" journalOrPublisher="Noticiario Mensual" pageId="0" pageNumber="91" pagination="5 - 11" refId="B15" refString="Cekalovic-K, TN, 1968. Conocimiento actual de los opiliones chilenos. Noticiario Mensual 12 (138): 5 - 11" title="Conocimiento actual de los opiliones chilenos." volume="12" year="1968">Cekalovic 1968</bibRefCitation>
: 7 [cat], 1985: 18 [cat].
</paragraph>
<paragraph id="0FB6E38676E7A8BFBA08481673240C71" pageId="0" pageNumber="91">
<taxonomicName id="F42BFC563243DDD02AF35C89208ED3BF" class="Arachnida" family="Gonyleptidae" genus="Sadocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sadocus bicornis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="bicornis">Sadocus bicornis</taxonomicName>
: Kury, 2003: 191 [cat]; 
<bibRefCitation id="4E6EBE9355426DA2B8697E2AD8BEC7D9" author="Kury, AB" journalOrPublisher="Self published, Rio de Janeiro" pageId="0" pageNumber="91" publicationUrl="https://wcolite.com/" refId="B41" refString="Kury, AB, Mendes, AC, Cardoso, L, Kury, MS, Granado Ade, A, Giribet, G, 2020b. World Catalogue of Opiliones. WCO-Lite version 1.2.1. https://wcolite.com/" title="World Catalogue of Opiliones. WCO-Lite version 1.2.1." url="https://wcolite.com/" year="2020 b">Kury et al. 2020b</bibRefCitation>
[cat].
</paragraph>
<paragraph id="BC6DEAD40CCDA1FFDB3DBD0AE84C7462" pageId="0" pageNumber="91">
<taxonomicName id="5BD4DDBA6FE803880AD944334B99203D" authorityName="Loman" authorityYear="1899" class="Arachnida" family="Gonyleptidae" genus="Discocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocyrtus calcitrosus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="calcitrosus">Discocyrtus calcitrosus</taxonomicName>
Loman, 1899: 7, fig 5 [desc]; 
<bibRefCitation id="D2373402BCDC50B9AEAB4842C05AEC25" author="Sorensen, WE" journalOrPublisher="Ergebnisse der Hamburger Magalhaensischen Sammelreise" pageId="0" pageNumber="91" pagination="1 - 36" refId="B99" refString="Sorensen, WE, 1902. Gonyleptiden (Opiliones, Laniatores). Ergebnisse der Hamburger Magalhaensischen Sammelreise 6 (5): 1 - 36" title="Gonyleptiden (Opiliones, Laniatores)." volume="6" year="1902">
<normalizedToken id="A679243E30C3CFB7EA15A33D197D2823" originalValue="Sørensen">Sorensen</normalizedToken>
1902
</bibRefCitation>
: 19 [syst]. (type ZMB 7837, ma holotype - examined by detailed photographs).
</paragraph>
<paragraph id="966D90135576C995982B76C94FA01F6E" pageId="0" pageNumber="91">
<taxonomicName id="463BE1B4CBACC3AD525462A8FAEDEB92" class="Arachnida" family="Gonyleptidae" genus="Lycomedes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedes calcitrosus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="calcitrosus">Lycomedes calcitrosus</taxonomicName>
: 
<normalizedToken id="33F81D717C4B1BEBFD684C87BF91BE35" originalValue="Sørensen">Sorensen</normalizedToken>
, 1902: 19 [syst]. Synonymy with 
<taxonomicName id="F93F8C219947F504F884DB03C25D6E04" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">L. asperatus</taxonomicName>
established by 
<bibRefCitation id="871B8B0DD26C58126302F1C068B77B8E" author="Roewer, CF" journalOrPublisher="&quot; Ciencias Zoologicas" pageId="0" pageNumber="91" refId="B77" refString="Roewer, CF, 1913. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79A(4-5): 1-473." title="Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fuer Naturgeschichte. 79 A (4 - 5): 1 - 473." year="1913">Roewer 1913</bibRefCitation>
.
</paragraph>
<paragraph id="5D2EBC367CA9BD64D7DAE4A4BFB80719" pageId="0" pageNumber="91">
<taxonomicName id="558C944361EBD070B988AC2998394021" class="Arachnida" family="Gonyleptidae" genus="Lycomedicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lycomedicus calcitrosus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="calcitrosus">Lycomedicus calcitrosus</taxonomicName>
: Moritz, 1971: 193 [cat].
</paragraph>
</subSubSection>
<subSubSection id="641E2CC10DA8F1DF9FE4B29B3F4E4D21" pageId="0" pageNumber="91" type="materials_examined">
<paragraph id="FF51EBAB057552DDD2417526A01B4D84" pageId="0" pageNumber="91">Material examined.</paragraph>
<paragraph id="4129FAF4E509952821FBA3265348CCA1" pageId="0" pageNumber="91">
<materialsCitation id="1389BD03B2FEF7D769BF80320D3637F2" ID-GBIF-Occurrence="3059965308" accessionNumber="MCZ 31267, MCZ 31275, MCZ 38272, MCZ 38265, MCZ 31263, MCZ 31264, MCZ 31232, MCZ 31270, MCZ 31274" collectingDate="1951-01-08" collectingDateMax="2016-12-27" collectingDateMin="1951-01-08" collectorName="El Canelo, de Concepcion, N. I. Platnick, O. F. Francke, T. Cekalovic, I. Avila, S. Ocares, D. Silva, F. Marques, F. Cadiz, F. Carbayo, L. E. Pena, L. Pena, G. Moreno, de Cautin, S. Gonzalez, E. Ross, A. Michelbacher, A. Anker, P. H. Martins, R. Pinto-da-Rocha, D. Cadiz, Cadiz L., G. Giribet, G. Hormiga, A. Perez-Gonzalez, de Llanquihue, de Valdivia, E. Krahmer" country="Chile" county="Same" latitude="-39.81389" location="Same" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" specimenCount="1" stateProvince="Region Metropolitana de Santiago">
<collectingCountry id="A4C78C04FFA2832AD04737E0D03A6C0C" name="Chile">Chile</collectingCountry>
, date and collector unknown, 1 ma (
<accessionNumber id="656CB9B02BA192841875DED7DE368DFB" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31267">MCZ 31267</accessionNumber>
). 
<collectingRegion id="A80471F83D01981EE833851E621C4C86" country="Chile" name="Region Metropolitana">
<normalizedToken id="4257B3BDB0E0A0D1C0B142AAE607DB61" originalValue="Región">Region</normalizedToken>
Metropolitana de Santiago
</collectingRegion>
, 
<emphasis id="B0FC390915D5E545EF299CA8617FC7D0" italics="true" pageId="0" pageNumber="91">
<collectingRegion id="5E213CBF5A4311A4EC661504BAB87A33" country="Chile" name="Region Metropolitana">Santiago</collectingRegion>
</emphasis>
, date or collector unknown, 6 ma, 3 fe (SMF 1369); 
<collectingCounty id="518B722EBEF21E17422E25BE23011958">Same</collectingCounty>
, 
<emphasis id="E580727362E98472DBCA6DCCEC7BCEC2" italics="true" pageId="0" pageNumber="91">
<collectorName id="C49A0D8C00F81F3C0DF3C2F52CA5A0A0">
<collectingMunicipality id="16F7B7118323624F98348D9C0E617687">El Canelo</collectingMunicipality>
</collectorName>
</emphasis>
, 
<collectingDate id="4620015912DEE74C73A7C54F8E04AFF2" value="1980-01-16">16.I.1980</collectingDate>
, collector unknown, 2 ma (AMNH); 
<location id="09EDD88952E2211838F6CAA316D47AF4" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:09EDD88952E2211838F6CAA316D47AF4" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Same" stateProvince="Region Metropolitana de Santiago">Same</location>
, 
<collectingDate id="D7CF4943FD3198CC654EF596BBB6197D" value="1980-01-16">16.I.1980</collectingDate>
, collector unknown, 1 ma, 1 fe (AMNH). 
<collectingRegion id="46DC6537833ADAF232D5F0E14585A2C0" country="Chile" name="Valparaiso">
<normalizedToken id="2E5BA81E2F3BA01ACAE25CEEFF9DDE9E" originalValue="Región">Region</normalizedToken>
de 
<normalizedToken id="6C010E70F1D28A996931DE0560FD5D81" originalValue="Valparaíso">Valparaiso</normalizedToken>
</collectingRegion>
, collection date or collector unknown, 8 ma, 19 fe (SMF 5382). 
<collectingRegion id="D7BC5760231BFE6C76D9B6DFDD911D18" country="Chile" name="Biobio">
<normalizedToken id="0249C92EB5BE0E51B49C53E9A90FC8B8" originalValue="Región">Region</normalizedToken>
de 
<normalizedToken id="98A3483EA64144264CCF8A015004EB98" originalValue="Biobío">Biobio</normalizedToken>
</collectingRegion>
, 
<emphasis id="8B026C247226322593157446B922D0E6" italics="true" pageId="0" pageNumber="91">
<location id="710F8CBA42B9C0B84CDB8BCD8DE7BC3F" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:710F8CBA42B9C0B84CDB8BCD8DE7BC3F" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Provincia de Concepcion" stateProvince="Region Metropolitana de Santiago">
Provincia 
<collectorName id="B9A50160DCBB027274F73FD69C0867C9">
de 
<normalizedToken id="18A8CEF44CCA965F1232AA451B38F993" originalValue="Concepción">Concepcion</normalizedToken>
</collectorName>
</location>
</emphasis>
, 
<collectingDate id="8747BF49773087E00F188540D9FAF8DA" value="1985-01-22">22.I.1985</collectingDate>
, 
<collectorName id="48A265D1CC495FA188F6D3B8BE78C94D">N.I. Platnick</collectorName>
&amp; 
<collectorName id="E11844AF55A416FC95FC194169108EBA">O.F. Francke</collectorName>
coll., 1 ma (AMNH); 
<location id="20DDF5D6701CEDE9C7FCDB4F7D5BAB36" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:20DDF5D6701CEDE9C7FCDB4F7D5BAB36" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Same" stateProvince="Region Metropolitana de Santiago">Same</location>
, 
<location id="023020A6A4631448E45D5E22E65538D9" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:023020A6A4631448E45D5E22E65538D9" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Estero Nonguen" stateProvince="Region Metropolitana de Santiago">
Estero 
<normalizedToken id="E6D320C221942679FD09B97832129DB8" originalValue="Nonguén">Nonguen</normalizedToken>
</location>
(
<geoCoordinate id="7E18D6FB409703E92311776EFFF6A287" degrees="36.831501" direction="south" orientation="latitude" precision="1" value="-36.8315">-36.831501</geoCoordinate>
, 
<geoCoordinate id="75792ED8ED7A770A6DB3E43BEA72895F" degrees="73.008374" direction="west" orientation="longitude" precision="1" value="-73.00838">-73.008374</geoCoordinate>
), 
<collectingDate id="4569A57955757E82E6EC1557FB2C043E" value="1977-03-13">13.III.1977</collectingDate>
, 
<collectorName id="96C346324FB2E2BD4D251688FB0CD4A5">T. Cekalovic</collectorName>
coll., 1 ma (
<accessionNumber id="DD6786933C49D3C38B24FABB45171B1A" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31275">MCZ 31275</accessionNumber>
); Same, Cerro Caracol, Mirador Aleman (
<geoCoordinate id="D389561195D9E84202D2D44CE8A008C3" degrees="36.834167" direction="south" orientation="latitude" precision="1" value="-36.834167">-36.834167</geoCoordinate>
, 
<geoCoordinate id="B1EE4F572B799375BC4584573B67A1B3" degrees="73.047778" direction="west" orientation="longitude" precision="1" value="-73.047775">-73.047778</geoCoordinate>
), 
<collectingDate id="393115DC5A7D6E921FCEE0A706762733" value="1968-09-15">15.IX.1968</collectingDate>
, 
<collectorName id="30C239090F56DCAD50CE65AA2795B013">T. Cekalovic</collectorName>
coll., 1 ma, 3 fe (MZSP 9965); Same, 
<emphasis id="0BE66D7270913A275CDBD4DB275280B5" italics="true" pageId="0" pageNumber="91">
Provincia de 
<collectingRegion id="9C52A754F00363FB8A9D6DFF93C46296" country="Chile" name="Araucania">Arauco</collectingRegion>
</emphasis>
, Parque Nacional Nahuelbuta (
<geoCoordinate id="B6AD215ADE7C858E6B2BD0B7FA27D84A" degrees="37.800000" direction="south" orientation="latitude" precision="1" value="-37.8">-37.800000</geoCoordinate>
, 
<geoCoordinate id="3CFE65B0219F3BB421BA0B98646B9310" degrees="73.033333" direction="west" orientation="longitude" precision="1" value="-73.03333">-73.033333</geoCoordinate>
), 
<collectingDate id="65C1D8514F27A384C726458651499538" value="2003-11-30">30.XI.2003</collectingDate>
, 
<collectorName id="1D90F875694F1AA931115ABD6E446A8A">I. Avila</collectorName>
, 
<collectorName id="481BF814868191CBDB0735411381B662">S. Ocares</collectorName>
&amp; 
<collectorName id="FCE60B22BCDE32D29B019AB3408B8700">D. Silva</collectorName>
coll., 1 ma (CAS 9055050); Same, Parque Nacional Nahuelbuta (
<geoCoordinate id="107EB8A1DC53660D096F0C70F5E52480" degrees="37.8043374259" direction="south" orientation="latitude" precision="1" value="-37.804337">-37.8043374259</geoCoordinate>
, 
<geoCoordinate id="ADBAB480311F71D197B0E4146737E43E" degrees="73.0344813614" direction="west" orientation="longitude" precision="1" value="-73.034485">-73.0344813614</geoCoordinate>
, date or collector unknown, 1 ma, 1 fe (CAS 9052227); Same, Parque Nacional Nahuelbuta (
<geoCoordinate id="8B780E7FE8DD7B9F4778FFF1D446CE7E" degrees="37.827500" direction="south" orientation="latitude" precision="1" value="-37.8275">-37.827500</geoCoordinate>
, 
<geoCoordinate id="483F7E1F45C4B04D9A998E33D71C197F" degrees="73.009722" direction="west" orientation="longitude" precision="1" value="-73.00972">-73.009722</geoCoordinate>
), 
<collectingDate id="ABC6151A60A17FEDB86E71B3FBB82409" value="2010-12-09">9.XII.2010</collectingDate>
, 
<collectorName id="E750BE81940E2314591F8112B52B7D6E">F. Marques</collectorName>
, 
<collectorName id="82D2EE82D08A1F2F33F38407FE7B8A38">F. Cadiz</collectorName>
&amp; 
<collectorName id="119B5B1070686C1201365BFC65B77D1F">F. Carbayo</collectorName>
coll., 5 ma (MZSP 36839); Same, Parque Nacional Nahuelbuta, (
<geoCoordinate id="B7FE53B4BB53EFB809A5A3E984991234" degrees="37.827500" direction="south" orientation="latitude" precision="1" value="-37.8275">-37.827500</geoCoordinate>
, 
<geoCoordinate id="B5DA0595CA8BBA3CB59AFD64DE7DDA76" degrees="73.009722" direction="west" orientation="longitude" precision="1" value="-73.00972">-73.009722</geoCoordinate>
), 
<collectingDate id="2B5838CA8C2224A96B71E5AABC1F2CC4" value="2010-12-09">9.XII.2010</collectingDate>
, 
<collectorName id="0357D83AA969EB296D0267026BC74888">F. Marques</collectorName>
, 
<collectorName id="E476D9B5308007CC9B3877A13860A83C">F. Cadiz</collectorName>
&amp; 
<collectorName id="BF3215409E06CBAA4A4744E866369E1D">F. Carbayo</collectorName>
coll., 6 ma, 1 fe (MZSP 36840); Same, Cayucupil, Pichinahuel, 
<collectingDate id="DB988979CB249C96359223E314FA04A9" value="1976-12-23">23.XII.1976</collectingDate>
, collector unknown, 1 ma (AMNH); Same (
<geoCoordinate id="4E7E86F30C140EE422D5306C7C7DD455" degrees="37.766667" direction="south" orientation="latitude" precision="1" value="-37.766666">-37.766667</geoCoordinate>
, 
<geoCoordinate id="4F73EE794429E57300ECCF04E4799F64" degrees="73.333333" direction="west" orientation="longitude" precision="1" value="-73.333336">-73.333333</geoCoordinate>
), 
<collectingDate id="C5AA406BA4E1CD2072CB27D8C5C4D72A" value="1965-02-09">9.II.1965</collectingDate>
, 
<collectorName id="76ADCCF8EE4ADB0649811F9A8E6E95B6">T. Cekalovic</collectorName>
coll., 1 ma (MZSP 27695). 
<location id="E2EA213FD39D4334097C9A67DC0C98C2" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:E2EA213FD39D4334097C9A67DC0C98C2" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Region de Nuble" stateProvince="Region Metropolitana de Santiago">
<normalizedToken id="7D6F3728BB4B0E073BCF51B8166F2991" originalValue="Región">Region</normalizedToken>
de Nuble
</location>
, 
<emphasis id="E227C852175AA47E19E1FBCEBF55381A" italics="true" pageId="0" pageNumber="91">
Provincia 
<normalizedToken id="C4EC8EEC4655D3931756DDF5D15E91F9" originalValue="Diguillín">Diguillin</normalizedToken>
</emphasis>
, 
<normalizedToken id="DD43ADA49E9CD1C0B742A01A3B2861C3" originalValue="Chillán">Chillan</normalizedToken>
, Las trancas Nuble, 
<collectingDate id="FBB29FB14F243F0B2610119A5F1FA704" value="1976-12">XII.1976</collectingDate>
, collector unknown, 1 ma, 3 fe (AMNH); Same,?. 
<collectingDate id="053CCC1BED8AD55DCC6D1B5BD946F7C9" value="1976-12">XII.1976</collectingDate>
, collector unknown, 1 ma, 2 fe (AMNH); Same, 
<collectingDate id="B0D8768DC45B790C2F532FCFA64D4C8B" value="1983-01-11" valueMax="1983-01-17" valueMin="1983-01-11">11-17.I.1983</collectingDate>
, 
<collectorName id="E0746D5ABCFCCB2690A56A6F18804073">L.E. Pena</collectorName>
coll., 1 ma (AMNH); Same, 
<collectingDate id="9D381B5A9A33317BCBD2562427899AD6" value="1974-12">XII.1974</collectingDate>
, collector unknown, 1 ma, 3 fe (AMNH); Same, Las Trancas, (
<geoCoordinate id="8860CE1E367F5CDC077048D647FE8E1E" degrees="36.600" direction="south" orientation="latitude" precision="55" value="-36.6">-36.600</geoCoordinate>
, 
<geoCoordinate id="1337A0D3D2728BC73BC1508223B062B5" degrees="72.117" direction="west" orientation="longitude" precision="55" value="-72.117">-72.117</geoCoordinate>
), 
<collectingDate id="E69A729A6A3349B93B0E8E8F918120C6" value="1967-01">I.1967</collectingDate>
, 
<collectorName id="DDEA79BC12428C190EAF2D51F38E212D">
L. 
<normalizedToken id="2C31BC1A21041C25C5E57AEB63EA2B56" originalValue="Peña">Pena</normalizedToken>
</collectorName>
coll, 1 ma (
<accessionNumber id="BD3B62336BEA4F89E8FF8D641C932943" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ38272">MCZ 38272</accessionNumber>
); Same, Los Lleuques (
<geoCoordinate id="C8ADDDFD3A2C1E34886929678B47F3EF" degrees="36.85800" direction="south" orientation="latitude" precision="1" value="-36.858">-36.85800</geoCoordinate>
, 
<geoCoordinate id="9E8795E119A290666AAF89ABC32F13E4" degrees="71.61700" direction="west" orientation="longitude" precision="1" value="-71.617">-71.61700</geoCoordinate>
), 
<collectingDate id="B9E6E8626CB1C1F6856007CC4F66A8D0" value="1975-05-11" valueMax="1975-05-07" valueMin="1975-05-11">11-7.V.1975</collectingDate>
, 
<collectorName id="F8E3D81B8C7D0E43FDA6DA181C4370AE">G. Moreno</collectorName>
coll., 1 ma, 4 fe (MCZ 3125). 
<collectingRegion id="5496325A0208397E3DEB1E884135FA6C" country="Chile" name="Araucania">
<normalizedToken id="6C5D75190BE9A2951EA4CFE4C83B4B88" originalValue="Región">Region</normalizedToken>
de 
<normalizedToken id="29B733A41045F0D9265BC0AB2BB5015E" originalValue="Araucanía">Araucania</normalizedToken>
</collectingRegion>
, 
<emphasis id="2B71A787BDF4B18B7610849AE3C539C6" italics="true" pageId="0" pageNumber="91">
<location id="0F6B11F7388C482E1CDAF4143332B45F" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:0F6B11F7388C482E1CDAF4143332B45F" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Provincia de Cautin" stateProvince="Region Metropolitana de Santiago">
Provincia 
<collectorName id="AA30CC82161212A27EF2E63395375519">
de 
<normalizedToken id="8FB19EB1876DEE0E2971CAD4A942F78B" originalValue="Cautín">Cautin</normalizedToken>
</collectorName>
</location>
</emphasis>
, 
<collectingDate id="D4AE14A1EBAD92B1268F001B7120FEC8" value="1985-01-27">27.I.1985</collectingDate>
, 
<collectorName id="DCAD048C296899C6A7135CA62642369C">N.I. Platnick</collectorName>
&amp; 
<collectorName id="54490B7F5B921E66D5E752A0CF6E0E15">O.F. Francke</collectorName>
coll., 1 ma (AMNH); Same, 
<collectingDate id="8987D68C2A224AD186AE173FC20FE128" value="1992-12-06">6.XII.1992</collectingDate>
, 
<collectorName id="CD13FF93FBFFC9098534DA9AC2F5EBC6">T. Cekalovic</collectorName>
&amp; 
<collectorName id="898AD7822070B2A5998D363C75A40FBE">S. Gonzalez</collectorName>
coll., 1 ma (AMNH); Same, Fundo de las Selvas, 
<collectingDate id="F3F3007D25A45BB85C9FF77B91735FBF" value="1981-02-16" valueMax="1981-02-20" valueMin="1981-02-16">16-20.II.1981</collectingDate>
, 
<collectorName id="FEA681653E064C07AEF947B4453A15E2">L.E. Pena</collectorName>
coll., 3 ma, 1 fe (AMNH); Same, Fundo de las Selvas, 
<collectingDate id="5C267530531A006C7010AFB6DC7C95C9" value="1981-02-16" valueMax="1981-02-20" valueMin="1981-02-16">16-20.II.1981</collectingDate>
; 
<collectorName id="28E14657EAFF4527ADAE13F53E0EC93D">L.E. Pena</collectorName>
coll., 3 ma, 1 fe (AMNH); Same, Villarrica (-39.03, -72.121), 
<collectingDate id="A68ED97F75848957439087B5CA71B298" value="1965-01-01" valueMax="1965-01-30" valueMin="1965-01-01">1-30.I.1965</collectingDate>
, 
<collectorName id="E07129DF1DE68414F4D7011A50E22152">
L. 
<normalizedToken id="AE0C4BDA9B736C251F930406C7945DB9" originalValue="Peña">Pena</normalizedToken>
</collectorName>
coll., 1 ma (
<accessionNumber id="B90809337F814E8FCEC036606287C985" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ38265">MCZ 38265</accessionNumber>
); Same, (-39.03, -72.121), 
<collectingDate id="1748AF6233E34E3EBFA61279C34A6DFF" value="1965-01-01" valueMax="1965-01-30" valueMin="1965-01-01">1-30.I.1965</collectingDate>
, 
<collectorName id="61BFF28E3FA00A919F5694863A57272E">
L. 
<normalizedToken id="615FB5AA929F2D37813533CF26269912" originalValue="Peña">Pena</normalizedToken>
</collectorName>
coll., 5 ma (
<accessionNumber id="A9354690997613F29F7971A737202786" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31263">MCZ 31263</accessionNumber>
); Same, 1 ma (
<accessionNumber id="545B6967BE93278DC4B0ED86D346B488" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31264">MCZ 31264</accessionNumber>
); Same, 6 fe, 4 ma (
<accessionNumber id="916B82DA18437690AFBC440357B12A65" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31232">MCZ 31232</accessionNumber>
); Same, 1 ma, 3 fe (
<accessionNumber id="8326CCB1D88C94111C94990EFD4551D4" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31270">MCZ 31270</accessionNumber>
); Same, 
<location id="83D5E3E9397884F24B492EA14D9333A7" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:83D5E3E9397884F24B492EA14D9333A7" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Flor del Lago Ranch" stateProvince="Region Metropolitana de Santiago">Flor del Lago Ranch</location>
(
<geoCoordinate id="0D8AF564BC84E49A9F50AB892708B3B7" degrees="39.2044664427" direction="south" orientation="latitude" precision="1" value="-39.204468">-39.2044664427</geoCoordinate>
, 
<geoCoordinate id="C532B2DE548E263F239BC0354284C8AF" degrees="72.1279383487" direction="west" orientation="longitude" precision="1" value="-72.12794">-72.1279383487</geoCoordinate>
), 
<collectingDate id="36457BC8DB972E23BB49A60260E68970" value="2003-12-11">11.XII.2003</collectingDate>
, 
<collectorName id="2E77A7F4E5C18C1F5B17F9B9FC3CA047">I. Avila</collectorName>
&amp; 
<collectorName id="D90D06686A4B4D3F6D1C0E21D7B2EABB">D. Silva</collectorName>
coll., 1 ma, 2 fe (CAS 9055035); Same, 2 ma (CAS 9055009); Same, 
<normalizedToken id="DCA50D76589D41B86F6BA7A6A5A9647F" originalValue="Pucón">Pucon</normalizedToken>
, 
<collectingDate id="E967E61AB8C8FC5B691B89326895BCD6" value="1951-01-12">12.I.1951</collectingDate>
, 
<collectorName id="1E49F8A952D6BED3BFCD1D05F8C429A5">E. Ross</collectorName>
&amp; 
<collectorName id="5D82DBA3DCF90F57862C6DFD820A4A0A">A. Michelbacher</collectorName>
coll., 3 ma, 7 fe (CAS 9055052); Same, Angol, Sierra Nahuelbuta, 
<collectingDate id="2171F1E6D30A845CC8F8009592633938" value="1951-01-13">13.I.1951</collectingDate>
, 
<collectorName id="8FC8E3A8F6046976174814CBCE94D43A">E. Ross</collectorName>
&amp; 
<collectorName id="1108DDB936D5BE008F9398F44A8AFB1D">A. Michelbacher</collectorName>
coll., 1 ma (CAS, 9055400); Same, Angol (
<geoCoordinate id="E36930CEEBC9EAAB945935D87072E834" degrees="37.829000" direction="south" orientation="latitude" precision="1" value="-37.829">-37.829000</geoCoordinate>
, 
<geoCoordinate id="9EE916B490B74482F02F297849EBE9A5" degrees="73.007278" direction="west" orientation="longitude" precision="1" value="-73.00728">-73.007278</geoCoordinate>
), 
<collectingDate id="60CC9D539C416D9B7332EC8B4FBB0754" value="2016-12-25" valueMax="2016-12-27" valueMin="2016-12-25">25-27.XII.2016</collectingDate>
, 
<collectorName id="2E4A971334B6D50B3B4270549F37A916">A. Anker</collectorName>
&amp; 
<collectorName id="B27EF9E3FEA9EB6E671F9801C8D20220">P.H. Martins</collectorName>
coll., 1 fe (UFMG 22643); Same, 2 fe (UFMG 22644); same, 1 ma (UFMG 22645); Same, 1 ma, 1 fe (UFMG 22646); same, 2 fe (UFMG 22647); Same, 1 ma (UFMG 22649); Same, Temuco, 
<collectingDate id="ACFA813BAEC1DBFEA3CDA2A2FD66E2F0" value="1951-01-08">8.I.1951</collectingDate>
, 
<collectorName id="B272A0DCE40277E3C0B70D9461F63F2F">E. Ross</collectorName>
&amp; 
<collectorName id="500A8173F5CD974FFEC1DD668ABE5F12">A. Michelbacher</collectorName>
coll., 1 ma (CAS 9052224); Same, Cerro 
<normalizedToken id="695CC3E1523B47AF4AAFE4F2C4C7F29F" originalValue="Ñielol">Nielol</normalizedToken>
(
<geoCoordinate id="D3053B6A64744C570779ECCFADE21E0A" degrees="38.726389" direction="south" orientation="latitude" precision="1" value="-38.72639">-38.726389</geoCoordinate>
, 
<geoCoordinate id="23FDFFA95540762D6FC9BF5C2FACD706" degrees="72.590833" direction="west" orientation="longitude" precision="1" value="-72.590836">-72.590833</geoCoordinate>
), 
<collectingDate id="D01808A066C6CBF360D2899EC8707590" value="2010-01-26">26.I.2010</collectingDate>
, 
<collectorName id="56A5FBC3386B589FB955766B75EA5B73">R. Pinto-da-Rocha</collectorName>
, 
<collectorName id="8F49C3E89C248AE68CA4423ACBDEBC91">
F. 
<normalizedToken id="54141E8582E2207BF98F4C1848844468" originalValue="Cádiz">Cadiz</normalizedToken>
</collectorName>
L. &amp; 
<collectorName id="6BAC2C49FA05D62E88C901D7B6547099">
D. 
<collectorName id="EFB6B3B3B7FD67363CCC11DD1B084750">
<normalizedToken id="236FD4580D2D27D98C8356D82DCE70EB" originalValue="Cádiz">Cadiz</normalizedToken>
L.
</collectorName>
</collectorName>
coll., 5 ma, 5 fe (MZSP 36836); Same, 
<emphasis id="BBD2CC7DBEF81FE08DB2534FCFCBDB9D" italics="true" pageId="0" pageNumber="91">
<location id="0C3473A6A3AA5CE02703B9ADCFAC6921" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:0C3473A6A3AA5CE02703B9ADCFAC6921" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Provincia de Malleco" stateProvince="Region Metropolitana de Santiago">Provincia de Malleco</location>
</emphasis>
, 
<normalizedToken id="27FB75C52231D424A0445B8F4FE20BC9" originalValue="Caracatín">Caracatin</normalizedToken>
, 
<collectingDate id="6DC80BF9737EA7416F41AB726B5EDF55" value="1975-06">VI.1975</collectingDate>
, collector unknown, 1 ma, 1 fe (AMNH); Same, 
<normalizedToken id="2B6158A4856CED13DC6BA61625A3312C" originalValue="Purén">Puren</normalizedToken>
, Monumento Natural Contulmo, (
<geoCoordinate id="8322A2A57A0668E2A577B937492607AA" degrees="38.01256" direction="south" orientation="latitude" precision="1" value="-38.01256">-38.01256</geoCoordinate>
, 
<geoCoordinate id="B261ADF635CBF1B721FC6D7AC394C2F1" degrees="73.18517" direction="west" orientation="longitude" precision="1" value="-73.18517">-73.18517</geoCoordinate>
), 
<collectingDate id="82580CA9AA8D809A6E83A3C49F5913CF" value="2014-11-12">12.XI.2014</collectingDate>
, 
<collectorName id="0C9451E0ED966C6817233331341AFBCF">G. Giribet</collectorName>
, 
<collectorName id="6DE1B44A32C9D37A152A5925D843693E">G. Hormiga</collectorName>
&amp; 
<collectorName id="F48B0C3633A2CF017A782B710297B08C">
A. 
<normalizedToken id="85C9DB6DE56F335D304046597614FF6D" originalValue="Pérez-González">Perez-Gonzalez</normalizedToken>
</collectorName>
coll., 1 ma, 1 fe (MCZ 138066); Same, Monumento Nacional Contulmo (
<geoCoordinate id="070D511B842CFB71920AF93D686E3D10" degrees="38.012778" direction="south" orientation="latitude" precision="1" value="-38.01278">-38.012778</geoCoordinate>
, 
<geoCoordinate id="13EB6860123625C802880B753199BDEE" degrees="73.187500" direction="west" orientation="longitude" precision="1" value="-73.1875">-73.187500</geoCoordinate>
), 
<collectingDate id="3960C6AD960CD97181F953CDB209237B" value="2010-12-12">12.XII.2010</collectingDate>
, 
<collectorName id="58BA6F1FA24FC9D0AE479B4F487A16FC">F. Marques</collectorName>
, 
<collectorName id="B42ACA650199FB68379840EEA8F9B11D">
F. 
<normalizedToken id="2882165F6BAA690EFBEA2BE6EA806FB5" originalValue="Cádiz">Cadiz</normalizedToken>
</collectorName>
&amp; 
<collectorName id="BED365C0987D6E7A367776B51E6C4C12">F. Carbayo</collectorName>
coll., 9 ma, 10 fe, 3 juvs. (MZSP 36967). 
<collectingRegion id="AAF38D496E34D161BDE7720089A27DFA" country="Chile" name="Los Lagos">
<normalizedToken id="3D3A7C790FFE2780287F0503F30A903A" originalValue="Región">Region</normalizedToken>
de Los Lagos
</collectingRegion>
, 
<emphasis id="8B91883C37A8D89CABC8C28F634BB37F" italics="true" pageId="0" pageNumber="91">
<location id="6B07AFE525B802627AE4DDCFCC39F071" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:6B07AFE525B802627AE4DDCFCC39F071" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Provincia de Llanquihue" stateProvince="Region Metropolitana de Santiago">
Provincia 
<collectorName id="92385BCAF003D22EFACDF10E33A057C9">de Llanquihue</collectorName>
</location>
</emphasis>
, 
<collectingDate id="BC73DCCFE9F02B0AB8E7EFD3BD14E21A" value="1994-02-13">13.II.1994</collectingDate>
, 
<collectorName id="864D1D3474CF96A6D568CA6E227C526F">T. Cekalovic</collectorName>
coll., 6 ma, 9 fe (AMNH). 
<collectingRegion id="C55E2C58C297590EF8231325F0A1F5A5" country="Chile" name="Los Rios">
<normalizedToken id="D164817E6EF1D5304D8ADF0E88B21BEC" originalValue="Región">Region</normalizedToken>
de Los 
<normalizedToken id="A593155A1BAD5AEA6721B53AF6024CB1" originalValue="Ríos">Rios</normalizedToken>
</collectingRegion>
, 
<emphasis id="145FB34AE84E057ECF3884375A26354E" italics="true" pageId="0" pageNumber="91">
<location id="95D9D34842F4F2796F52060A27EF67CC" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:95D9D34842F4F2796F52060A27EF67CC" country="Chile" county="Same" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" municipality="El Canelo" name="Provincia de Valdivia" stateProvince="Region Metropolitana de Santiago">
Provincia 
<collectorName id="47C6BD8F44AC898E9197E199A5EF5DF9">de Valdivia</collectorName>
</location>
</emphasis>
, date or collector unknown, 2 ma, 4 fe (SMF 783); Same, 2 ma, 3 fe (SMF 778); Same, 
<collectingDate id="21D0484E53B46122F97DD292E4D745A7" value="1993-02-28">28.II.1993</collectingDate>
, 
<collectorName id="B55FA32CDB66FE7ED67E54F927F7A8DA">T. Cekalovic</collectorName>
coll., 2 ma (AMNH); Same, (
<geoCoordinate id="12E4F6021158B72DCFF7F146DEDFA454" degrees="39.81389" direction="south" orientation="latitude" precision="1" value="-39.81389">-39.81389</geoCoordinate>
, 
<geoCoordinate id="DEDF07B8043D28075FDDD9830C9732C7" degrees="73.24583" direction="west" orientation="longitude" precision="1" value="-73.24583">-73.24583</geoCoordinate>
), 
<collectingDate id="30131E4DEE5B587F97929DEFB6FC1C10" value="1978-11-15" valueMax="1978-11-20" valueMin="1978-11-15">15-20.XI.1978</collectingDate>
, 
<collectorName id="1C9E563197272C0DB6B551F7B3B6FAF5">E. Krahmer</collectorName>
coll., 3 ma, 4 fe (
<accessionNumber id="FE43449C0E5420912A7FBE09FD61C89D" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31274">MCZ 31274</accessionNumber>
); Same, Panguipulli, Puerto Fuy, 
<collectingDate id="087AA41A44D2881E8D19617E8DAE0C98" value="1978-02-24">24.II.1978</collectingDate>
, collector unknown, 1 ma (AMNH); Same, Valdivia
</materialsCitation>
, 
<materialsCitation id="9FCFE0E6217827D2EE4A98FE87568700" ID-GBIF-Occurrence="3059965301" accessionNumber="MCZ 31231" collectingDate="1965-03-06" collectorName="H. Levi" country="Isla" latitude="-39.81389" location="Teja" longLatPrecision="1" longitude="-73.24583" specimenCount="1">
<collectingCountry id="B818923BD984E8A9717CCA4C313B5A7B" name="Iceland">Isla</collectingCountry>
<location id="5E0F6E8BE46B84DD2E231E57F4A9C736" LSID="urn:lsid:plazi:treatment:F5D43E8E278D5AE28249E292A2DB4ACF:5E0F6E8BE46B84DD2E231E57F4A9C736" country="Isla" latitude="-39.81389" longLatPrecision="1" longitude="-73.24583" name="Teja">Teja</location>
(
<geoCoordinate id="0EA25E111FB273FF30385175A5C7600B" degrees="39.81389" direction="south" orientation="latitude" precision="1" value="-39.81389">-39.81389</geoCoordinate>
, 
<geoCoordinate id="49BDAE553C05B2FA9523742BD23F4EBB" degrees="73.24583" direction="west" orientation="longitude" precision="1" value="-73.24583">-73.24583</geoCoordinate>
), 
<collectingDate id="9A7E4E26756E3217BFB45717A1EDD40C" value="1965-03-06">6.III.1965</collectingDate>
, 
<collectorName id="1157A0FD577E004E30974D8AB1E4E9EC">H. Levi</collectorName>
coll., 1 ma (
<accessionNumber id="43C25B522AE184B53B684271DF363D8F" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ31231">MCZ 31231</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BFE8D77D4967B4C4D4C5BF58E01FE73" pageId="0" pageNumber="91" type="diagnosis">
<paragraph id="9768F4842216B7C4D5FC702222266CAB" pageId="0" pageNumber="91">Diagnosis.</paragraph>
<paragraph id="3F474B20FB6E9817CC1F91491CDF7ECE" pageId="0" pageNumber="91">
<taxonomicName id="0236B3EAE107554A3482D337EC702B9C" baseAuthorityName="Gervais" baseAuthorityYear="1847" class="Arachnida" family="Gonyleptidae" genus="Sadocus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sadocus asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="3D5349214DE4BAAB3DB3F418381A79D2" italics="true" pageId="0" pageNumber="91">Sadocus asperatus</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="43BC708CEE66C16CAE64970D9FC161D5" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. ingens" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="ingens">
<emphasis id="D1E8EED66BF26FA6A35BB628209694BB" italics="true" pageId="0" pageNumber="91">S. ingens</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6EDF18B267CA563BE5B110165CE75A1" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. polyacanthus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="polyacanthus">
<emphasis id="A7CAD4A5CD7421B89F86595B981B8068" italics="true" pageId="0" pageNumber="91">S. polyacanthus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="721104F53DD3B1755C720CBC0F9D2A89" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. dilatatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="dilatatus">
<emphasis id="ADE2595D15DD9771DCE2E11CCB3AA209" italics="true" pageId="0" pageNumber="91">S. dilatatus</emphasis>
</taxonomicName>
by the bifid prodorsal apical apophysis on coxa IV. 
<taxonomicName id="5B6F9C5E3F1A0536B4828C4F60E72979" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="A17594B5D892F10A6B0B24F6107D0AA2" italics="true" pageId="0" pageNumber="91">S. asperatus</emphasis>
</taxonomicName>
can be distinguished from the latter species by the combination of the following characters: lateral margin of dorsal scutum covered by granules; trochanter IV with a blunt retro-dorsal apical apophysis being half of the podomere length, and a rhombus retro-ventral apical tubercle; femur IV curved (in dorsal view), with a retro-lateral row of spiniform apophysis (the middle one longest).
</paragraph>
</subSubSection>
<subSubSection id="86FE74B15B0EDCDEB27B1A9D6AAA3923" pageId="0" pageNumber="91" type="redescription">
<paragraph id="8A5C6D8FAF83B943E66A8980D8982C12" pageId="0" pageNumber="91">Redescription.</paragraph>
<paragraph id="37AEAEE98C50F4C36DDCDB0BD383BA43" pageId="0" pageNumber="91">
<emphasis id="46EF53AC511C894C8EB42C54AD280004" bold="true" pageId="0" pageNumber="91">Male</emphasis>
(CAS 9055035). Measurements. Dorsal scutum maximum length 6.3; dorsal scutum maximum width 7.2; prosoma maximum length 2.5; prosoma maximum width 3.2; leg femora I 3.2; II 6.5; III 5.5; IV 7.2. 
<emphasis id="F3FCD814100190E5497FECF45402F17D" bold="true" italics="true" pageId="0" pageNumber="91">Dorsum</emphasis>
(Fig. 
<figureCitation id="7146A49D82B5469218D19E90120CE2F3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sadocus asperatus (Gervais). Male (CAS 9055035) A Habitus, dorsal view B-E right leg IV, Trochanter-tibia IV B dorsal view C prolateral view D ventral view E retro-lateral view. Scale: 1 mm." figureDoi="10.3897/zookeys.1025.57806.figure3" httpUri="https://binary.pensoft.net/fig/522913" pageId="0" pageNumber="91">3A</figureCitation>
). Dorsal scutum type gamma triangular. Carapace with granules sparsely distributed. Scutal areas I-IV with eight, four, two and four granules, respectively; scutal area III with one pair of paramedian spiniform tubercles; scutal area IV incompletely divided. Lateral margin of dorsal scutum mostly covered in granules (from posterior half of carapace to posterior margin of dorsal scutum). Posterior margin of dorsal scutum and free tergite I each with a pair of paramedian tubercles. 
<emphasis id="552DBF57F2D06769B27AC533B27BECCD" bold="true" pageId="0" pageNumber="91">
<emphasis id="A9C425C24B7609292A27601D5285A8C3" italics="true" pageId="0" pageNumber="91">Chelicerae</emphasis>
.
</emphasis>
Segment I with basal tubercle, bulla with small setae, each finger with five teeth. 
<emphasis id="A8F4F86CA06AFF35237CBD02F0EBED8B" bold="true" pageId="0" pageNumber="91">
<emphasis id="20CC341D49828CD100BD843DA9F2B58B" italics="true" pageId="0" pageNumber="91">Pedipalps</emphasis>
.
</emphasis>
Coxa dorsal face smooth, ventral face with two apical tubercles. Trochanter dorsal and ventral faces smooth. Femur ventral face granulate. Patella with sparsely distributed setae. Tibial setation: prolateral IiiiIi/IiiiIi; retro-lateral iIiIi/IiIi. Tarsal setation: prolateral and retro-lateral IiIi/IiIi. 
<emphasis id="B0209B2A9A043A90E423084B084E90C0" bold="true" italics="true" pageId="0" pageNumber="91">Legs</emphasis>
(Fig. 
<figureCitation id="543A2EA3B55AFD0602377EF101F8DF7B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sadocus asperatus (Gervais). Male (CAS 9055035) A Habitus, dorsal view B-E right leg IV, Trochanter-tibia IV B dorsal view C prolateral view D ventral view E retro-lateral view. Scale: 1 mm." figureDoi="10.3897/zookeys.1025.57806.figure3" httpUri="https://binary.pensoft.net/fig/522913" pageId="0" pageNumber="91">3B-E</figureCitation>
). Coxa IV with one long, oblique, bifid prodorsal apical apophysis and one retro-ventral apical spine. Trochanters I and II each with one pair of prodorsal spiniform tubercles. Trochanter III with one medio-ventral tubercle and three retro-lateral ones. Trochanter IV prodorsal and proventral faces with few granules, the prodorsal apical apophysis long (ca. half the podomere length); retro-lateral face with one basal, one central, and one apical tubercles; one retro-dorsal apical spiniform apophysis (ca. 
<normalizedToken id="A0E0A8A6A924ED0E4B94F1F0F5F78E8B" originalValue="¼">1/4</normalizedToken>
podomere length); ventral face with setiferous tubercles sparsely distributed. Femur III with one retro-basal tubercle. Femur IV sigmoid, with dorsal row of apophysis on the basal half abruptly decreasing in size apically, becoming granules; prolateral row with central-subapical tubercles; retro-lateral row of granules with spiniform apophysis (basal most and central one longer than the others, apical most oblique, curved ventrad); ventral face with two short retro-lateral sub-apical spiniform apophyses and one proapical spiniform apophysis. Patella IV ventral face mostly smooth, with one probasal, one proapical and one retro-apical large, spiniform tubercles. Tibia IV roughly with two ventral rows of granules increasing in size from central to apical becoming spines. Tarsal counts: 6, 9, 7, 8. 
<emphasis id="2FD294DC503517C331A931118B82C06C" bold="true" italics="true" pageId="0" pageNumber="91">Penis</emphasis>
(Fig. 
<figureCitation id="A82EBBCCD4906EE68E1BAACBDA5AF997" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distal part of penis Sadocus and Discocyrtus, dorsal and ventral views, respectively A, B S. asperatus, macrosetae colorized and numbered C, D S. ingens E, F S. polyacanthus G, H S. dilatatus I, J S. funestus K, L D. catharinensis. Scale bar of drawings: 0.1 mm." figureDoi="10.3897/zookeys.1025.57806.figure11" httpUri="https://binary.pensoft.net/fig/522921" pageId="0" pageNumber="91">11A, B</figureCitation>
). Ventral plate of penis with moderate cleft on anterior margin, three pairs of MS A, one pair of MS B, four or five pairs of MS C and one pair of MS D, without MS E.
</paragraph>
<paragraph id="D568A40F1A12896F6953D34E0A4504E9" pageId="0" pageNumber="91">
<emphasis id="C1302BDBAB59EFCF729FD63DDCCAA538" bold="true" pageId="0" pageNumber="91">
<emphasis id="B80CC6F7EE6D56AEADF915AD929380D8" italics="true" pageId="0" pageNumber="91">Coloration</emphasis>
.
</emphasis>
Immersed in ethanol: carapace, trochanter-patella IV brown, tibia light brown, legs I-III, pedipalps, and chelicerae yellowish-brown. Live specimens (Fig. 
<figureCitation id="2E10A365A4352B302BD9C317DEB8E5CF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Photographs of living specimens of Sadocus A S. asperatus (ma) B S. asperatus (fe) C S. asperatus (ma) D S. dilatatus (ma) E S. asperatus (fe) F S. funestus (ma). A, B taken by R. Pinto-da-Rocha, C-F taken by Pedro H. Martins." figureDoi="10.3897/zookeys.1025.57806.figure9" httpUri="https://binary.pensoft.net/fig/522919" pageId="0" pageNumber="91">9A-C, E</figureCitation>
): carapace, coxa, and trochanter black, light gray spot (inverted T shape) from ocularium to scutal area II; femora I-III orange; patellae-tibiae I-IV brown.
</paragraph>
<paragraph id="0C31446F051CFB1B22AE44B0CB5E345E" pageId="0" pageNumber="91">
<emphasis id="D7500893802B3581BF1C52B23D5E3E6B" bold="true" italics="true" pageId="0" pageNumber="91">Variations</emphasis>
(n = 56). Scutal area I-IV tubercles (5 minimum, 16 maximum per site), Measurements. Dorsal scutum maximum length 6.0-8.5; dorsal scutum maximum width 6.7-11.6; prosoma maximum length 2.5-3.5; prosoma maximum width 3.2-4.8; leg femora: I 3.2-5.0; II 6.5-10.0; III 5.5-8.5; IV 7.2-11.0.
</paragraph>
<paragraph id="E3E34D97F5A0A94B5B01E0D595E2617C" pageId="0" pageNumber="91">
<emphasis id="94C0E4B04ED5A51C02584E1B920EB238" italics="true" pageId="0" pageNumber="91">Female</emphasis>
(CAS 9055035). Measurements. Dorsal scutum maximum length 6.3; dorsal scutum maximum width 8.1; prosoma maximum length 2.6; prosoma maximum width 3.7; leg femora: I 3.7; II 7; III 6.1; IV 7.2.
</paragraph>
</subSubSection>
<subSubSection id="8B891EDEE937067C6182E1AA131B99F6" pageId="0" pageNumber="91" type="redescription">
<paragraph id="2975190A49D00233E5C80F0D2BD2F8A4" pageId="0" pageNumber="91">Redescription.</paragraph>
<paragraph id="8B7FC036DC93DBD874A029E9EB720BD1" pageId="0" pageNumber="91">
<emphasis id="DFD0AE9B04CAA39E2DC1974C5DDFCF07" bold="true" italics="true" pageId="0" pageNumber="91">Dorsum</emphasis>
(Fig. 
<figureCitation id="1A3C866C722CEA980720EBB21A69CD28" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Female specimens of Sadocus A, B S. asperatus (CAS- 9055035) A dorsal view B lateral view C, D S. dilatatus (MZSP- 8022) C dorsal view D lateral view E, F S. funestus (AMNH) E dorsal view F lateral view G, H S. ingens (MZSP- 36965) G dorsal view H lateral view I, J S. polyacanthus (AMNH) I dorsal view J lateral view K Drawing of species inquirenda Carampangue nigronotatum extracted from Mello-Leitao 1943." figureDoi="10.3897/zookeys.1025.57806.figure8" httpUri="https://binary.pensoft.net/fig/522918" pageId="0" pageNumber="91">8A, B</figureCitation>
). Scutal areas I, II, and IV with three, five, and two granules, respectively. 
<emphasis id="587D56B633F69B686E3406ABCD4AF6BF" bold="true" pageId="0" pageNumber="91">
<emphasis id="BBBBA97B561F8DAAE3EE1E04F1A0BAE2" italics="true" pageId="0" pageNumber="91">Legs</emphasis>
.
</emphasis>
Coxa IV with one prodorsal apical apophysis and one retro-ventral apical spine shorter than on the male, trochanter IV with retro-lateral row of tubercles, the apical one longest, femur IV with pro- and retro-ventral rows of tubercles, patella-metatarsi IV unarmed. Tarsal counts: 6, 9, 7, 8. 
<emphasis id="E906C4005798B34161AAE3B6A07783B1" bold="true" italics="true" pageId="0" pageNumber="91">Ovipositor</emphasis>
(Fig. 
<figureCitation id="C20DC494E0F7C7F8B69BA0F83EDC2608" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Distal part of ovipositor of Sadocus A-C S. asperatus A upper view B villi C setae D-F S. funestus D upper view E lateral view F seta G-I S. ingens G upper view H, I setae J-L S. polyacanthus J upper view K lateral view L setae." figureDoi="10.3897/zookeys.1025.57806.figure12" httpUri="https://binary.pensoft.net/fig/522922" pageId="0" pageNumber="91">12A-C</figureCitation>
). Two main groups of lobes delimited by a constriction, ovipositors peripheral setae inserted into sockets that are a mixture of dorsal and ventral sockets, the dorsal lobe with five setae and the ventral one with six; each main group of lobes divided by a fissure.
</paragraph>
<paragraph id="CE8BDFC558F08BD846AACC4E4CE80DC3" pageId="0" pageNumber="91">
<emphasis id="175E0FAD04510EA49AA673843407A471" bold="true" italics="true" pageId="0" pageNumber="91">Variations</emphasis>
(n = 66). Tubercle variation in the scutal areas: I 2-7; II 1-8; III 2-8; IV 3-5. Free tergites I-III each with one pair of blunt or pointed spiniform tubercles. Measurements. Dorsal scutum maximum length 5.9-8.2; dorsal scutum maximum width 8.0-9.6; prosoma maximum length 2.4-3.5; prosoma maximum width 3.7-4.5; leg femora: I 3.5-4.5; II 6.7-8.7; III 5.7-7.0; IV 7.0-8.9.
</paragraph>
</subSubSection>
<subSubSection id="A5C694C64F2BAF6B7407B0393567A360" pageId="0" pageNumber="91" type="type locality">
<paragraph id="8111A1CB28B1540FA4B7FBCE0E9084E5" pageId="0" pageNumber="91">Type locality.</paragraph>
<paragraph id="5397AD71AC5161767BE0875ECD2A74D9" pageId="0" pageNumber="91">
Of 
<taxonomicName id="3B75A61CA5B63904A2EF126BBD3EA784" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="FD60E468165403537890AD5D991852BC" italics="true" pageId="0" pageNumber="91">Gonyleptes asperatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="59F624B1356455E09600B22CE8025852" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes subsimilis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="CAC78CD1B5BBD8FFD7CCBA5F3724AED4" italics="true" pageId="0" pageNumber="91">Gonyleptes subsimilis</emphasis>
</taxonomicName>
: CHILE. Of 
<taxonomicName id="8D0080539FD1366F571339C403EE9BE6" authorityName="Loman" authorityYear="1899" class="Arachnida" family="Gonyleptidae" genus="Discocyrtus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discocyrtus calcitrosus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="calcitrosus">
<emphasis id="25CB0DF17613F1720C6980832BFA3984" italics="true" pageId="0" pageNumber="91">Discocyrtus calcitrosus</emphasis>
</taxonomicName>
: CHILE, 
<normalizedToken id="98AF7926F0075AD90447665826D76B3B" originalValue="Región">Region</normalizedToken>
de Los Rios, 
<emphasis id="BE858334D4BDA8A504467E91BA9FA1B1" italics="true" pageId="0" pageNumber="91">Provincia de Valdivia</emphasis>
, Corral.
</paragraph>
</subSubSection>
<subSubSection id="7BF9DFEC378CC4F0691913309C0F80B5" pageId="0" pageNumber="91" type="distribution">
<paragraph id="D9070AF922C31546CA0F8455455D7B79" pageId="0" pageNumber="91">Geographical distribution</paragraph>
<paragraph id="E8F4AE5A206937B11DADCC105F04B729" pageId="0" pageNumber="91">
<emphasis id="4803A8B99289CE824C7AAE932C0B4774" bold="true" pageId="0" pageNumber="91">
(Fig. 
<figureCitation id="21C791ED1935E4E86B133528C18FBAA2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geographical records of distribution of Sadocus species." figureDoi="10.3897/zookeys.1025.57806.figure2" httpUri="https://binary.pensoft.net/fig/522912" pageId="0" pageNumber="91">2</figureCitation>
).
</emphasis>
Chile, 
<normalizedToken id="156218726962DAFC695562DB05029F41" originalValue="Región">Region</normalizedToken>
de Los 
<normalizedToken id="7CC97A35D00B39EAE3F86AAA5FF9CA53" originalValue="Ríos">Rios</normalizedToken>
, 
<emphasis id="53D8492F66FC54639B9D4834BA1DB238" italics="true" pageId="0" pageNumber="91">Valdivia</emphasis>
, Corral; 
<normalizedToken id="93715AA99D48A3AC07E573878222969E" originalValue="Región">Region</normalizedToken>
Metropolitana de Santiago, 
<emphasis id="D1EF81B36C3028D5BAD2BDB33169678E" italics="true" pageId="0" pageNumber="91">Santiago</emphasis>
; 
<normalizedToken id="960879365F400020C80AC26A12078379" originalValue="Región">Region</normalizedToken>
de 
<normalizedToken id="4484F66D5A25FDE7676C3CC30A12B953" originalValue="Valparaíso">Valparaiso</normalizedToken>
; 
<normalizedToken id="18D01FB45F377585EB097565020D6E04" originalValue="Región">Region</normalizedToken>
de 
<normalizedToken id="8B12165E46FFC354BDF5D3626BE469CF" originalValue="Biobío">Biobio</normalizedToken>
, 
<emphasis id="40D76D7DAB9E3E209108D0A42325CCF0" italics="true" pageId="0" pageNumber="91">
Provincia de 
<normalizedToken id="B5DF0B13066814606D2BF06553662EC0" originalValue="Concepción">Concepcion</normalizedToken>
</emphasis>
, 
<emphasis id="55A7F9B67D9EF7A01BFF3D2F1DF1B51D" italics="true" pageId="0" pageNumber="91">Provincia de Arauco</emphasis>
, 
<emphasis id="E7308AD2009DFEDDAA4D2C9F99B00C49" italics="true" pageId="0" pageNumber="91">Provincia de Nuble</emphasis>
; 
<normalizedToken id="E4F039ADE88F6756DCB9B352A7181D25" originalValue="Región">Region</normalizedToken>
de 
<normalizedToken id="99226CFF117C2FFC1C42BB3A0E42B162" originalValue="Araucanía">Araucania</normalizedToken>
, 
<emphasis id="C79FCA3F78776C48EDAAB36326BEB6F4" italics="true" pageId="0" pageNumber="91">Provincia de Cautin</emphasis>
, 
<emphasis id="6CAFE286E85899FBDD90E2B319C91C31" italics="true" pageId="0" pageNumber="91">Provincia de Malleco</emphasis>
; 
<normalizedToken id="54E69525101220FF0CFC65E8D6D3A04C" originalValue="Región">Region</normalizedToken>
de Los Lagos, 
<emphasis id="24F67A05C4F5259C3FCF56994E54BA3B" italics="true" pageId="0" pageNumber="91">Provincia de Llanquihue</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="192FD7ABB7425406F11EB86F181F656D" pageId="0" pageNumber="91" type="taxonomic notes">
<paragraph id="A074E0608A111E4C93CBE50F81FBFAFB" pageId="0" pageNumber="91">Taxonomic notes.</paragraph>
<paragraph id="7EE1070C872CAD2C1F00455286CC6202" pageId="0" pageNumber="91">
After examining the original description and the drawing of 
<taxonomicName id="6F47A6AACEF1324F0D9A5061E0C81D71" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonyleptes bicornis" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="bicornis">
<emphasis id="7E0CDC5BBADCED15E5566E9507B8AECF" italics="true" pageId="0" pageNumber="91">Gonyleptes bicornis</emphasis>
</taxonomicName>
, we concluded that it is of a male of 
<taxonomicName id="D251A592C77A9FFF93028F6529E4328A" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="AC07B566939175748D3D353A843DE387" italics="true" pageId="0" pageNumber="91">S. asperatus</emphasis>
</taxonomicName>
. In the original description, the spines on the free tergite, the two apical apophyses on the trochanter IV and uneven spines in the inner part of the 
<normalizedToken id="1455E650EFF09D005FB31C8E93CE28D4" originalValue="“leg”">&quot;leg&quot;</normalizedToken>
(referring to the femur IV) are mentioned. Those characters lead us to conclude that it is 
<taxonomicName id="639C9D9F74DA38EDDE848BE3B10690B0" class="Arachnida" family="Gonyleptidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="S. asperatus" order="Opiliones" pageId="0" pageNumber="91" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="23A24670221127CEB1F46B8BB9657B02" italics="true" pageId="0" pageNumber="91">S. asperatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>