<documentid="DBA028A50BF0C1F7C3D9CF131F9B080D"ID-CLB-Dataset="305014"ID-DOI="10.11646/zootaxa.5527.1.1"ID-GBIF-Dataset="0d9d4ebd-5dd8-473c-80b9-19960053d8b4"ID-ISSN="1175-5334"ID-Zenodo-Dep="14021823"ID-ZooBank="771D2E7B-4025-45BF-B328-6EC8A8851ECD"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="guilherme"IM.materialsCitations_approvedBy="guilherme"IM.metadata_approvedBy="guilherme"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="guilherme"IM.treatmentCitations_approvedBy="guilherme"IM.treatments_approvedBy="guilherme"checkinTime="1730451569409"checkinUser="plazi"docAuthor="Ellenrieder, Natalia Von & Gill, Raymond J."docDate="2024"docId="039787AAFFA1FFFFFF45039DFAB2B715"docLanguage="en"docName="zootaxa.5527.1.1.pdf"docOrigin="Zootaxa 5527 (1)"docSource="http://dx.doi.org/10.11646/zootaxa.5527.1.1"docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph"docStyleId="5EBBA59367AD13919D70D935FA04F6A3"docStyleName="Zootaxa.2013-.monograph"docStyleVersion="19"docTitle="Tetralicia hoelmeri von Ellenrieder & Gill 2024, sp. nov."docType="treatment"docVersion="8"lastPageNumber="35"masterDocId="FFAEFFD2FFBFFFDDFFD2062DFFF1B368"masterDocTitle="The genus Tetralicia Harrison (Hemiptera: Sternorrhyncha: Aleyrodidae) in California, U. S. A., with the description of five new species and a redescription of Tetralicia granulata Sampson & Drews, 1941"masterLastPageNumber="129"masterPageNumber="1"pageNumber="31"updateTime="1730914976340"updateUser="ExternalLinkService">
<mods:titleid="6DDD547F61642FFF7E65DB8B28EB6753">The genus Tetralicia Harrison (Hemiptera: Sternorrhyncha: Aleyrodidae) in California, U. S. A., with the description of five new species and a redescription of Tetralicia granulata Sampson & Drews, 1941</mods:title>
<mods:affiliationid="E2704311EB10F6A970802D4CDF5C73C1">Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832 - 1448, U. S. A. & natalia. von. ellenrieder @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 1159 - 2019</mods:affiliation>
<mods:affiliationid="6B9CD2EE2E8F1D64935372C6D4C1D55C">Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832 - 1448, U. S. A. & Plant Pest Diagnostics Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832 - 1448, U. S. A. & rayromagill @ gmail. com; https: // orcid. org / 0009 - 0005 - 9047 - 2742 * Corresponding author</mods:affiliation>
<taxonomicNameid="4C3E4D3FFFA1FFC3FF45039DFD6FB6A2"authority="von Ellenrieder & Gill"authorityName="von Ellenrieder & Gill"authorityYear="2024"box="[151,670,1456,1483]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="30"pageNumber="31"phylum="Arthropoda"rank="species"species="hoelmeri"status="sp. nov.">
<figureCitationid="13052A39FFA1FFC3FF4503F9FEE7B687"box="[151,278,1492,1519]"captionStart="FIGURES 13–16"captionStartId="32.[151,275,1659,1685]"captionTargetBox="[185,1403,192,1622]"captionTargetId="figure-30@32.[172,1416,179,1635]"captionText="FIGURES 13–16. Puparia of Tetralicia in situ: 13, 14: T. hoelmeri, Riverside Co.,Agua Caliente Indian Reservation, Palm Springs, on native shrub, 13: dorsal view, 14: posterior view; 15: T. hyptisemoryi, Imperial Co., Shell Canyon, on desert lavender, dorsolateral view; 16: T. laingi, Orange Co., Trabuco, on Salvia. Figs 13–16 by N. von Ellenrieder & R.W. Garrison."figureDoi="http://doi.org/10.5281/zenodo.14046326"httpUri="https://zenodo.org/record/14046326/files/figure.png"pageId="30"pageNumber="31">Figs 13, 14</figureCitation>
<treatmentCitationid="0A9F10ADFFA1FFC3FDE20034FCC9B559"ID-DOI="http://doi.org/10.5281/zenodo.6194113"ID-Zenodo-Dep="6194113"author="Polaszek, A. & Gill, R."box="[560,824,1561,1585]"httpUri="http://treatment.plazi.org/id/591087F34367F925198F013D06B9ACE2"page="52"pageId="30"pageNumber="31"year="2011">
<bibRefCitationid="EFAF4B4DFFA1FFC3FDE20034FCC9B559"author="Polaszek, A. & Gill, R."box="[560,824,1561,1585]"pageId="30"pageNumber="31"pagination="51 - 59"refId="ref60115"refString="Polaszek, A. & Gill, R. (2011) A new species of whitefly (Hemiptera: Aleyrodidae) and its parasitoid (Hymenoptera: Aphelinidae) from desert lavender in California. Zootaxa, 2750 (1), 51 - 59. https: // doi. org / 10.11646 / zootaxa. 2750.1.5"type="journal article"year="2011">Polaszek & Gill 2011: 52</bibRefCitation>
<figureCitationid="13052A39FFA1FFC3FC4E0034FC0BB559"box="[924,1018,1561,1585]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="30"pageNumber="31">Figs 2, 3</figureCitation>
. Named after Kim A. Hoelmer, one of the collectors of this species, who provided several whitefly species on native host plants resulting from his Southern
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desert survey of native whiteflies and their parasitoids to CSCA.
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75 slide mounted puparia:
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92 dry mounted puparia:
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12,
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<figureCitationid="13052A39FFA0FFC2FB1507D2FAE9B171"box="[1223,1304,511,537]"captionStart="FIGURES 13–16"captionStartId="32.[151,275,1659,1685]"captionTargetBox="[185,1403,192,1622]"captionTargetId="figure-30@32.[172,1416,179,1635]"captionText="FIGURES 13–16. Puparia of Tetralicia in situ: 13, 14: T. hoelmeri, Riverside Co.,Agua Caliente Indian Reservation, Palm Springs, on native shrub, 13: dorsal view, 14: posterior view; 15: T. hyptisemoryi, Imperial Co., Shell Canyon, on desert lavender, dorsolateral view; 16: T. laingi, Orange Co., Trabuco, on Salvia. Figs 13–16 by N. von Ellenrieder & R.W. Garrison."figureDoi="http://doi.org/10.5281/zenodo.14046326"httpUri="https://zenodo.org/record/14046326/files/figure.png"pageId="31"pageNumber="32">Fig. 14</figureCitation>
) not visible dorsally, VO located on a promontory, five to seven lobate to strongly conical projections on each side of T3 and abdomen along apparent margin pointing laterally, and three to five (usually four) dorsolateral conical papillae along submargin of abdomen (
<figureCitationid="13052A39FFA0FFC2FE7A0446FE0DB1ED"box="[424,508,619,645]"captionStart="FIGURES 13–16"captionStartId="32.[151,275,1659,1685]"captionTargetBox="[185,1403,192,1622]"captionTargetId="figure-30@32.[172,1416,179,1635]"captionText="FIGURES 13–16. Puparia of Tetralicia in situ: 13, 14: T. hoelmeri, Riverside Co.,Agua Caliente Indian Reservation, Palm Springs, on native shrub, 13: dorsal view, 14: posterior view; 15: T. hyptisemoryi, Imperial Co., Shell Canyon, on desert lavender, dorsolateral view; 16: T. laingi, Orange Co., Trabuco, on Salvia. Figs 13–16 by N. von Ellenrieder & R.W. Garrison."figureDoi="http://doi.org/10.5281/zenodo.14046326"httpUri="https://zenodo.org/record/14046326/files/figure.png"pageId="31"pageNumber="32">Fig. 13</figureCitation>
<figureCitationid="13052A39FFA0FFC2FACD0446FA83B1ED"box="[1311,1394,619,645]"captionStart="FIGURES 13–16"captionStartId="32.[151,275,1659,1685]"captionTargetBox="[185,1403,192,1622]"captionTargetId="figure-30@32.[172,1416,179,1635]"captionText="FIGURES 13–16. Puparia of Tetralicia in situ: 13, 14: T. hoelmeri, Riverside Co.,Agua Caliente Indian Reservation, Palm Springs, on native shrub, 13: dorsal view, 14: posterior view; 15: T. hyptisemoryi, Imperial Co., Shell Canyon, on desert lavender, dorsolateral view; 16: T. laingi, Orange Co., Trabuco, on Salvia. Figs 13–16 by N. von Ellenrieder & R.W. Garrison."figureDoi="http://doi.org/10.5281/zenodo.14046326"httpUri="https://zenodo.org/record/14046326/files/figure.png"pageId="31"pageNumber="32">Fig. 13</figureCitation>
); longitudinal molting suture not lined with tubercles; eyespots absent; Ce setae absent; T2 and T3 setae long and flattened, arising from bulbous bases (107, 108, 112); T3 setae arising just behind anterior margin of metanotum (107, 108, 112); medial area with large tubercles on head lateral to mouth, along T1/T2 and T2/T3 sutures (
<figureCitationid="13052A39FFA0FFC2FE69054AFE16B0E9"box="[443,487,871,897]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">112</figureCitation>
), with large tubercles in a row along anterior area of A1–A7, and smaller in a row along posterior area of each segment; abdominal depressions indistinct (
<figureCitationid="13052A39FFA0FFC2FC6B05A6FB7AB0CD"box="[953,1163,906,933]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">Figs 112, 114, 115</figureCitation>
); with pairs of pores and porettes on each side of submedial area of head (1–2), T2 (0–1), T3 (0), A1 (0), A2 (0), A3 (0–1), A4 (1), A5 (0–1), A6 (0–1), A7 (0), and A8 (0); dorsal disc with large tubercles and pairs of pores and porettes along sides (
<figureCitationid="13052A39FFA0FFC2FF4505DAFF0BB779"box="[151,250,1015,1041]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">114, 116</figureCitation>
); dorsal submargin with tubercles along inner portion, with irregular darkened depressions, with pores, and with 3–5 [4] (usually 4) conical papillae on each side on abdomen (
<figureCitationid="13052A39FFA0FFC2FB910236FB57B75D"box="[1091,1190,1051,1077]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">114–116</figureCitation>
), some of which with a marginal disc pore; deflexed submargin with a row of pores, with dark granulations extending over entire width lacking a particular pattern (
<figureCitationid="13052A39FFA0FFC2FE05024EFDC8B715"box="[471,569,1123,1149]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">Fig. 113</figureCitation>
); marginal glandular teeth subquadrangular with tips smooth; VO oval, inset from posterior margin by more than its own length or about its own length; operculum cordate, its dorsal surface with a few longitudinal ridges and with microspinulae across distal third to fourth; lingula concealed by operculum; VO ring subtriangular, anteriorly wide and open (
<figureCitationid="13052A39FFA0FFC2FD0A02E2FC8AB781"box="[728,891,1231,1257]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">Figs 116, 117</figureCitation>
), with dorsal setae of A8 arising from bulbous bases anterior to level of anterior margin of operculum (
<figureCitationid="13052A39FFA0FFC2FCC102DEFC40B665"box="[787,945,1267,1293]"captionStart="FIGURES 112–117"captionStartId="82.[151,264,1480,1504]"captionTargetBox="[173,1416,194,1443]"captionTargetId="figure-30@82.[162,1425,181,1456]"captionTargetPageId="82"captionText="FIGURES 112–117. Tetralicia hoelmeri sp. nov.: 112–114, 116, 117: Riverside Co., Agua Caliente Indian Reservation, Palm Springs, on native shrub, 112, 113: Paratype; 114, 116, 117: Holotype; 115: Riverside Co., Living Desert, Palm Desert, on desert lavender. 112: Thorax andA1–A6 dorsal disc; 113: Detail deflexed submargin; 114, 115: Dorsal disc and submargin of abdomen; 116: Caudal end, dorsal view; 117: Detail vasiform orifice."figureDoi="http://doi.org/10.5281/zenodo.14021886"httpUri="https://zenodo.org/record/14021886/files/figure.png"pageId="31"pageNumber="32">Figs 116, 117</figureCitation>
); bases of caudal setae widely separated, at level of lateral margins of VO ring (
) with 4 pairs of wax plates on abdomen. Upper and lower compound eyes connected by one ommatidium (
<figureCitationid="13052A39FFA0FFC2FD2E038AFC6BB6A9"box="[764,922,1446,1473]"captionStart="FIGURES 235–242"captionStartId="121.[151,264,1674,1698]"captionTargetBox="[176,1412,194,1637]"captionTargetId="figure-25@121.[163,1425,181,1650]"captionTargetPageId="121"captionText="FIGURES 235–242. Tetralicia hoelmeri sp. nov., adults, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 235: female compound eye and antenna; 236: male compound eye and antenna; 237: female metatibia; 238: female mesotibia; 239: female protibia; 240: male metatibia; 241: male mesotibia; 242: male protibia."figureDoi="http://doi.org/10.5281/zenodo.14021993"httpUri="https://zenodo.org/record/14021993/files/figure.png"pageId="31"pageNumber="32">Figs 235, 236</figureCitation>
). Antennae 7 segmented (
<figureCitationid="13052A39FFA0FFC2FB6B038AFAA6B6A9"box="[1209,1367,1446,1473]"captionStart="FIGURES 235–242"captionStartId="121.[151,264,1674,1698]"captionTargetBox="[176,1412,194,1637]"captionTargetId="figure-25@121.[163,1425,181,1650]"captionTargetPageId="121"captionText="FIGURES 235–242. Tetralicia hoelmeri sp. nov., adults, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 235: female compound eye and antenna; 236: male compound eye and antenna; 237: female metatibia; 238: female mesotibia; 239: female protibia; 240: male metatibia; 241: male mesotibia; 242: male protibia."figureDoi="http://doi.org/10.5281/zenodo.14021993"httpUri="https://zenodo.org/record/14021993/files/figure.png"pageId="31"pageNumber="32">Figs 235, 236</figureCitation>
), with an elongate third segment with two primary round sensoria near the apex, fifth, sixth, and seventh segments each with 1 round primary sensorium located at apex on fifth and sixth, and at about two thirds of its length on seventh, and sixth and seventh segments with an elongated rod-like sensorium each. Hind tibiae (
<figureCitationid="13052A39FFA0FFC2FBBC003EFAFBB545"box="[1134,1290,1555,1581]"captionStart="FIGURES 235–242"captionStartId="121.[151,264,1674,1698]"captionTargetBox="[176,1412,194,1637]"captionTargetId="figure-25@121.[163,1425,181,1650]"captionTargetPageId="121"captionText="FIGURES 235–242. Tetralicia hoelmeri sp. nov., adults, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 235: female compound eye and antenna; 236: male compound eye and antenna; 237: female metatibia; 238: female mesotibia; 239: female protibia; 240: male metatibia; 241: male mesotibia; 242: male protibia."figureDoi="http://doi.org/10.5281/zenodo.14021993"httpUri="https://zenodo.org/record/14021993/files/figure.png"pageId="31"pageNumber="32">Figs 237, 240</figureCitation>
) with a tibial comb of 12–15 setae in male, 15–18 setae in female; metatibial and mesotibial brushes not defined. Female cement gland not discernible (
<figureCitationid="13052A39FFA0FFC2FD93008EFD52B5D5"box="[577,675,1699,1725]"captionStart="FIGURES 261–264"captionStartId="126.[151,264,1649,1673]"captionTargetBox="[176,1412,194,1612]"captionTargetId="figure-30@126.[163,1424,181,1625]"captionTargetPageId="126"captionText="FIGURES 261–264. Tetralicia hoelmeri sp. nov., nymphal instars 1–3, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 261: first instar nymph; 262: second instar nymph, stacked between first and third instars; 263: third instar nymph, with cephalothorax folded over; 264: third instar nymph, detail of unobstructed abdominal view."figureDoi="http://doi.org/10.5281/zenodo.14022006"httpUri="https://zenodo.org/record/14022006/files/figure.png"pageId="31"pageNumber="32">Fig. 261</figureCitation>
): with 6 pairs of long submarginal setae along sides of cephalothorax and 1 pair posterior to level of VO. VO oval, wider than long, with anterior margin straight; operculum oval occupying most of VO; lingula head bulbous and covered with microsetae; other characters not discernible. Second and third instars: lacking long submarginal setae, with button-like structures with a central pore along margins of cephalothorax and abdomen (25–32 on third instar; exact number not discernible on second instar) which are set close together, separated from each other by a space subequal to width of each structure (
<figureCitationid="13052A39FFA0FFC2FB54017AFAD7B419"box="[1158,1318,1879,1905]"captionStart="FIGURES 261–264"captionStartId="126.[151,264,1649,1673]"captionTargetBox="[176,1412,194,1612]"captionTargetId="figure-30@126.[163,1424,181,1625]"captionTargetPageId="126"captionText="FIGURES 261–264. Tetralicia hoelmeri sp. nov., nymphal instars 1–3, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 261: first instar nymph; 262: second instar nymph, stacked between first and third instars; 263: third instar nymph, with cephalothorax folded over; 264: third instar nymph, detail of unobstructed abdominal view."figureDoi="http://doi.org/10.5281/zenodo.14022006"httpUri="https://zenodo.org/record/14022006/files/figure.png"pageId="31"pageNumber="32">Figs 263, 264</figureCitation>
), and with conical submarginal abdominal papillae (4–6 on third instar; exact number not discernible on second instar), VO located on a promontory, and operculum subcordate (
<figureCitationid="13052A39FFA0FFC2FD3E01B2FC7DB4D1"box="[748,908,1951,1977]"captionStart="FIGURES 261–264"captionStartId="126.[151,264,1649,1673]"captionTargetBox="[176,1412,194,1612]"captionTargetId="figure-30@126.[163,1424,181,1625]"captionTargetPageId="126"captionText="FIGURES 261–264. Tetralicia hoelmeri sp. nov., nymphal instars 1–3, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 261: first instar nymph; 262: second instar nymph, stacked between first and third instars; 263: third instar nymph, with cephalothorax folded over; 264: third instar nymph, detail of unobstructed abdominal view."figureDoi="http://doi.org/10.5281/zenodo.14022006"httpUri="https://zenodo.org/record/14022006/files/figure.png"pageId="31"pageNumber="32">Figs 262–264</figureCitation>
<taxonomicNameid="4C3E4D3FFF9FFFFDFD210051FEDEB5D1"authority=", Riverside Co., Agua Caliente Indian Reservation, Palm Springs"authorityName="Riverside Co., Agua Caliente Indian Reservation, Palm Springs"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="32"pageNumber="33"phylum="Arthropoda"rank="species"species="hoelmeri">
, and the presence of lateral projections along apparent margin, VO ring open anteriorly, and all T2, T3, and dorsal A8 setae originating from bulbous bases with
share the presence of dorsal A8 setae longer than caudal setae (in puparia and earlier nymphs) with puparia of
<taxonomicNameid="4C3E4D3FFF9EFFFCFDEC056EFD3BB035"authorityName="von Ellenrieder & Gill"authorityYear="2024"box="[574,714,835,861]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="33"pageNumber="34"phylum="Arthropoda"rank="species"species="agrifoliae">
. Both species are the only known members of this genus with submarginal papillae (in puparia and second and third instar nymphs) and marginal button-like structures (in second and third instar nymphs). The latter resemble the gland-tubercles described by
<bibRefCitationid="EFAF4B4DFF9EFFFCFBC305A6FB3EB0CD"author="Nakahara, S."box="[1041,1231,906,933]"pageId="33"pageNumber="34"pagination="105 - 150"refId="ref60069"refString="Nakahara, S. (1995) Taxonomic studies of the genus Tetraleurodes (Homoptera: Aleyrodidae). Insecta Mundi, 9, 105 - 150. [https: // journals. flvc. org / mundi / article / view / 24778]"type="journal article"year="1995">Nakahara (1995)</bibRefCitation>
<figureCitationid="13052A39FF9EFFFCFB8F05DBFB7BB779"box="[1117,1162,1014,1041]"captionStart="FIGURE 105"captionStartId="80.[151,250,1691,1715]"captionTargetBox="[204,1384,194,1652]"captionTargetId="figure-30@80.[191,1396,181,1667]"captionTargetPageId="80"captionText="FIGURE 105. Tetralicia hoelmeri sp. nov., Paratype puparium, Riverside Co., Living Desert, Palm Desert, on desert lavender: Habitus."figureDoi="http://doi.org/10.5281/zenodo.14021882"httpUri="https://zenodo.org/record/14021882/files/figure.png"pageId="33"pageNumber="34">105</figureCitation>
), three to five—usually four—on each side of abdomen only [six to eight on each side of thorax and abdomen—usually seven—and usually also a medial one on head;
in parenthesis): length 650 [570–760] (740 [665–850]); maximum width 490 [410–550] (540 [485–630]), dorsal A8 setae and caudal setae are relatively longer in
: dorsal A8 setae length 90 [70–130] (65 [55–90]); caudal setae length: 60 [45–80] (50 [40–55]), and caudal setae are about as long as 1.5 times to 2.5 times the length of operculum in
have button-like structures along margins of cephalothorax and abdomen set close together, separated from each other by a space subequal to the width of each structure (
<figureCitationid="13052A39FF9EFFFCFAB80076FF0FB5F1"captionStart="FIGURES 261–264"captionStartId="126.[151,264,1649,1673]"captionTargetBox="[176,1412,194,1612]"captionTargetId="figure-30@126.[163,1424,181,1625]"captionTargetPageId="126"captionText="FIGURES 261–264. Tetralicia hoelmeri sp. nov., nymphal instars 1–3, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 261: first instar nymph; 262: second instar nymph, stacked between first and third instars; 263: third instar nymph, with cephalothorax folded over; 264: third instar nymph, detail of unobstructed abdominal view."figureDoi="http://doi.org/10.5281/zenodo.14022006"httpUri="https://zenodo.org/record/14022006/files/figure.png"pageId="33"pageNumber="34">Figs 262–264</figureCitation>
), whereas these structures are separated by a space subequal to two to four times the width of each structure in those nymphs of
<figureCitationid="13052A39FF9EFFFCFDE4008EFD26B5D5"box="[566,727,1699,1725]"captionStart="FIGURES 265–267"captionStartId="127.[151,264,1604,1628]"captionTargetBox="[180,1408,194,1567]"captionTargetId="figure-25@127.[167,1421,181,1580]"captionTargetPageId="127"captionText="FIGURES 265–267. Tetralicia hyptisemoryi, set of stacked nymphal instars 1–3 exuviae overlaying holotype puparium: 265: first instar nymph; 266: submargin of third instar nymph; 267: Caudal end of second and third instar nymphs (third instar below)."figureDoi="http://doi.org/10.5281/zenodo.14022008"httpUri="https://zenodo.org/record/14022008/files/figure.png"pageId="33"pageNumber="34">Figs 266, 267</figureCitation>
). The submarginal papillae are accompanied each by a pore in those nymphs of
<figureCitationid="13052A39FF9EFFFCFDC900EAFD8FB589"box="[539,638,1735,1761]"captionStart="FIGURES 261–264"captionStartId="126.[151,264,1649,1673]"captionTargetBox="[176,1412,194,1612]"captionTargetId="figure-30@126.[163,1424,181,1625]"captionTargetPageId="126"captionText="FIGURES 261–264. Tetralicia hoelmeri sp. nov., nymphal instars 1–3, Paratypes, Riverside Co., Living Desert, Palm Desert, on desert lavender: 261: first instar nymph; 262: second instar nymph, stacked between first and third instars; 263: third instar nymph, with cephalothorax folded over; 264: third instar nymph, detail of unobstructed abdominal view."figureDoi="http://doi.org/10.5281/zenodo.14022006"httpUri="https://zenodo.org/record/14022006/files/figure.png"pageId="33"pageNumber="34">Fig. 264</figureCitation>
<figureCitationid="13052A39FF9EFFFCFB6E00EAFAD1B589"box="[1212,1312,1735,1761]"captionStart="FIGURES 265–267"captionStartId="127.[151,264,1604,1628]"captionTargetBox="[180,1408,194,1567]"captionTargetId="figure-25@127.[167,1421,181,1580]"captionTargetPageId="127"captionText="FIGURES 265–267. Tetralicia hyptisemoryi, set of stacked nymphal instars 1–3 exuviae overlaying holotype puparium: 265: first instar nymph; 266: submargin of third instar nymph; 267: Caudal end of second and third instar nymphs (third instar below)."figureDoi="http://doi.org/10.5281/zenodo.14022008"httpUri="https://zenodo.org/record/14022008/files/figure.png"pageId="33"pageNumber="34">Fig. 266</figureCitation>
). As in the puparia, they also differ by their relative size, with
earlier nymphal instars being smaller (first instar 177–211 long, second instar 224–243 long, third instar 335–398 long, vs. 243, 296, and 446 long respectively in
Adults are diagnosed from other known species under
<taxonomicNameid="4C3E4D3FFF9EFFFCFCF80156FC5CB4FC"authorityName="von Ellenrieder & Gill"authorityYear="2024"box="[810,941,1915,1940]"class="Insecta"family="Aleyrodidae"genus="Tetralicia"kingdom="Animalia"order="Hemiptera"pageId="33"pageNumber="34"phylum="Arthropoda"rank="species"species="guajavae">
, sharing only with it the presence of submarginal papillae. However, numerous differences were observed between them (see diagnosis above), and the combination of characters by which they differ is consistent in all specimens examined. Specimens of this species were included as
<bibRefCitationid="EFAF4B4DFF9DFFFFFCCD0696FBD8B3BD"author="Polaszek, A. & Gill, R."box="[799,1065,187,213]"pageId="34"pageNumber="35"pagination="51 - 59"refId="ref60115"refString="Polaszek, A. & Gill, R. (2011) A new species of whitefly (Hemiptera: Aleyrodidae) and its parasitoid (Hymenoptera: Aphelinidae) from desert lavender in California. Zootaxa, 2750 (1), 51 - 59. https: // doi. org / 10.11646 / zootaxa. 2750.1.5"type="journal article"year="2011">Polaszek & Gill (2011)</bibRefCitation>
, where the presence of tubercles on the dorsal disc and submargin was interpreted as ontogenetic variability, hypothesizing that the tubercles would develop as the pupa aged (
<bibRefCitationid="EFAF4B4DFF9DFFFFFE18072EFD04B275"author="Polaszek, A. & Gill, R."box="[458,757,258,285]"pageId="34"pageNumber="35"pagination="51 - 59"refId="ref60115"refString="Polaszek, A. & Gill, R. (2011) A new species of whitefly (Hemiptera: Aleyrodidae) and its parasitoid (Hymenoptera: Aphelinidae) from desert lavender in California. Zootaxa, 2750 (1), 51 - 59. https: // doi. org / 10.11646 / zootaxa. 2750.1.5"type="journal article"year="2011">Polaszek & Gill 2011: 52</bibRefCitation>
). However, no intermediates were found, with pupae being either entirely smooth (
), and slide mounting and examination of numerous additional specimens of both species on samples of the CSCA dry collection and CDA herbarium showed that pupae otherwise matching
always have these tubercles irrespective of being young or already eclosed, disproving that hypothesis. Furthermore, both species were found to coexist on the same host plant in two localities (Imperial National Wildlife Refuge and Vidal Junction in Imperial County), indicating that the differences considered here to be interspecific are not due to environmental or geographical variability. The descriptions of nymphal instars 1–3 and adults of
<bibRefCitationid="EFAF4B4DFF9DFFFFFF66046AFE46B109"author="Polaszek, A. & Gill, R."box="[180,439,583,609]"pageId="34"pageNumber="35"pagination="51 - 59"refId="ref60115"refString="Polaszek, A. & Gill, R. (2011) A new species of whitefly (Hemiptera: Aleyrodidae) and its parasitoid (Hymenoptera: Aphelinidae) from desert lavender in California. Zootaxa, 2750 (1), 51 - 59. https: // doi. org / 10.11646 / zootaxa. 2750.1.5"type="journal article"year="2011">Polaszek & Gill (2011)</bibRefCitation>
<figureCitationid="13052A39FF9DFFFFFBA9046AFB28B109"box="[1147,1241,583,609]"captionStart="FIGURES 1–4"captionStartId="12.[151,275,1849,1875]"captionTargetBox="[183,1405,391,1811]"captionTargetId="figure-135@12.[170,1418,378,1824]"captionText="FIGURES 1–4. Puparia of Tetralicia in situ: 1: T. abnormis, Santa Barbara Co., Los Olivos, on Quercus agrifolia; 2, 3: T. acaudata, 2: Riverside Co., Tin Canyon, on manzanita, 3: San Bernardino Co., Mountain Home Village, Mill Creek Canyon, on manzanita; 4: T. agrifoliae, Riverside Co., Blue Jay Road & Ortega highway, on Quercus agrifolia. Fig. 2, by N. von Ellenrieder & R.W. Garrison, 1, 3, 4 by R.J. Gill."figureDoi="http://doi.org/10.5281/zenodo.14046317"httpUri="https://zenodo.org/record/14046317/files/figure.png"pageId="34"pageNumber="35">Figure 2</figureCitation>
<bibRefCitationid="EFAF4B4DFF9DFFFFFB3A046AFF0BB1ED"author="Polaszek, A. & Gill, R."pageId="34"pageNumber="35"pagination="51 - 59"refId="ref60115"refString="Polaszek, A. & Gill, R. (2011) A new species of whitefly (Hemiptera: Aleyrodidae) and its parasitoid (Hymenoptera: Aphelinidae) from desert lavender in California. Zootaxa, 2750 (1), 51 - 59. https: // doi. org / 10.11646 / zootaxa. 2750.1.5"type="journal article"year="2011">Polaszek & Gill 2011: 54</bibRefCitation>
from the Living Desert Museum with two submarginal papillae on thorax is inaccurate, since reexamination of that particular specimen (photographed here in
<figureCitationid="13052A39FF9DFFFFFB4E04A2FAF1B1C1"box="[1180,1280,654,681]"captionStart="FIGURE 105"captionStartId="80.[151,250,1691,1715]"captionTargetBox="[204,1384,194,1652]"captionTargetId="figure-30@80.[191,1396,181,1667]"captionTargetPageId="80"captionText="FIGURE 105. Tetralicia hoelmeri sp. nov., Paratype puparium, Riverside Co., Living Desert, Palm Desert, on desert lavender: Habitus."figureDoi="http://doi.org/10.5281/zenodo.14021882"httpUri="https://zenodo.org/record/14021882/files/figure.png"pageId="34"pageNumber="35">Figs 105</figureCitation>
,
<figureCitationid="13052A39FF9DFFFFFADE04A2FAC8B1C1"box="[1292,1337,655,681]"captionStart="FIGURES 106–111"captionStartId="81.[151,264,1633,1657]"captionTargetBox="[167,1420,194,1596]"captionTargetId="figure-25@81.[154,1433,181,1609]"captionTargetPageId="81"captionText="FIGURES 106–111. Tetralicia hoelmeri sp. nov.: 106, 108, 110: Holotype; 107, 109, 111: Paratype, Riverside Co., Living Desert, Palm Desert, on desert lavender. 106: Cephalothorax; 107: T2–A3 dorsal disc and submargin; 108: Detail submargin T3–A2; 109: Detail submargin A2–A5; 110: Venter cephalothorax; 111: Detail venter of T2 and T3."figureDoi="http://doi.org/10.5281/zenodo.14021884"httpUri="https://zenodo.org/record/14021884/files/figure.png"pageId="34"pageNumber="35">107</figureCitation>
in CSCA, shows papillae to be restricted to submargin of abdomen and absent from submargin of thorax in this species.
</paragraph>
<paragraphid="8B8136BCFF9DFFFFFF1504D6FBFBB0A1"blockId="34.[151,1437,151,1149]"pageId="34"pageNumber="35">The projections along apparent margin can be lobate to markedly conical and their development responds most likely to environmental conditions; specimens from Living Desert and Vidal Junction presented projections going from lobate to markedly conical, whereas specimens from Agua Caliente and Imperial Wildlife Refuge presented only lobate projections. Although their position corresponds to the position of the submarginal papillae for the most part, there are usually two additional submarginal projections on each side that are not accompanied by papillae, one on T3 and one on caudal abdomen between the last papilla and the caudal seta.</paragraph>
<bibRefCitationid="EFAF4B4DFF9DFFFFFF1505FEFE04B085"author="Carapia-Ruiz, V. E. & Sanchez-Flores, O. A. & Garcia-Martinez, O. & Castillo-Gutierrez, A."box="[199,501,979,1005]"pageId="34"pageNumber="35"pagination="1 - 5"refId="ref59151"refString="Carapia-Ruiz, V. E., Sanchez-Flores, O. A., Garcia-Martinez, O. & Castillo-Gutierrez, A. (2018 a) Estudio morfologico de Aleuropleurocelus abnormis (Quaintance) (Hemiptera: Aleyrodidae) y nuevos registros de especies del genero para Mexico. Acta Zoologica Mexicana, Nueva Serie, 34 (1), 1 - 5. https: // doi. org / 10.21829 / azm. 2018.3411177"type="journal article"year="2018">
<collectingRegionid="49FAF85EFF9DFFFFFB2005FEFF4FB779"country="Mexico"name="Baja California Sur"pageId="34"pageNumber="35">Baja California Sur</collectingRegion>
<collectingRegionid="49FAF85EFF9DFFFFFD53024EFD02B715"box="[641,755,1123,1149]"country="United States of America"name="California"pageId="34"pageNumber="35">California</collectingRegion>
(Imperial, Riverside, and San Bernardino Counties).
