<taxonomicNameid="0FEA6EE207611016FFACB9C8FE9794C1"authorityName="Zhan & Jin"authorityYear="2005"box="[102,350,914,942]"class="Strophomenata"family="Leptestiidae"genus="Leptestiina"higherTaxonomySource="GBIF"kingdom="Animalia"order="Strophomenatida"pageId="20"pageNumber="385"phylum="Brachiopoda"rank="species"species="veturna"status="sp. nov.">
<figureCitationid="50D109E407611016FFACB9E3FF0F94A1"box="[102,198,953,974]"captionStart="Fig"captionStartId="17.[102,135,1843,1863]"captionText="Fig. 12. A. Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press); NIGP139131, ventral internal mould (A) and latex cast (A) preserved to− + 1 2 gether with a bryozoan colony (top left). B–D. Leptellinidae gen. nov., sp. B. NIGP139132, dorsal internal mould (B1) and latex cast (B2). C. NIGP139133, ventral external mould (C1) and latex cast (C2). D. NIGP139134, ventral external mould. E–M. Calyptolepta huanghuaensis (Chang, 1983). E. NIGP139135, ventral internal mould. F. NIGP139136, broken ventral internal mould. G. NIGP139137, dorsal internal mould. H. NIGP139138, holotype, dorsal internal mould (H1) and latex cast (H2) with local enlargement (H3) showing cardinalia and bema. I. NIGP139139, dorsal external mould. J. NIGP139140, dorsal internal mould of a juvenile shell. K. NIGP139141, dorsal internal mould. L. NIGP139142, two dorsal internal moulds. M. NIGP139143, dorsal internal mould. N. Leptestiina veturna sp. nov.; NIGP139144, paratype, ventral internal mould."pageId="20"pageNumber="385">Figs. 12N</figureCitation>
<figureCitationid="50D109E407611016FF1AB9E3FED994A1"box="[208,272,953,974]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">13A–I</figureCitation>
<specimenCodeid="984CBD1A07611016FF00BE47FE99935D"box="[202,336,1053,1074]"collectionCode="NIGP"country="China"httpUri="http://grbio.org/cool/j51y-avsb"name="Naking Institute of Geology and Palaeontology"pageId="20"pageNumber="385">NIGP 139147</specimenCode>
<figureCitationid="50D109E407611016FE97BE47FE7C935D"box="[349,437,1053,1074]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">Fig. 13C</figureCitation>
<materialsCitationid="78821F3C07611016FF3ABE65FF77931F"location="Weixin County town"municipality="Shizigou Valley"pageId="20"pageNumber="385"specimenCount="1"stateProvince="Yunnan">
<locationid="CD3543BA07611016FE2EBE65FD79933B"LSID="urn:lsid:plazi:treatment:4043A47707611016FFACB9C8FB8D9105:CD3543BA07611016FE2EBE65FD79933B"box="[484,688,1087,1108]"municipality="Shizigou Valley"name="Weixin County town"pageId="20"pageNumber="385"stateProvince="Yunnan">Weixin County town</location>
.—Shell small, concavoconvex, subsemicircular to subcircular; maximum width along hinge line. Cardinal extremities acute to rounded. Anterior commissure rectimarginate. Ventral valve about three−quarters as long as wide, strongly convex, deepest in central part; visceral area more convex than marginal area of shell; interarea high, about one−eighth length of shell, apsacline with planar or slightly curved surface; delthyrium relatively wide, covered by weakly arched pseudodeltidium in its posterior two−thirds (
<figureCitationid="50D109E407611016FFA4BC2BFF1B91E5"box="[110,210,1649,1674]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">Fig. 13C</figureCitation>
<figureCitationid="50D109E407611016FF18BC27FF1291E2"box="[210,219,1661,1677]"captionStart="Fig"captionStartId="2.[102,135,1189,1209]"captionTargetBox="[119,1466,226,1162]"captionTargetId="graphics-319@2.[152,1466,243,1162]"captionTargetPageId="2"captionText="Fig. 2. Middle Ordovician sedimentary facies on the Yangtze Platform, South China palaeoplate (modified from Zhang et al. 2002), with three fossil localities discussed in the text: 1, Shihtzupu Formation at Shizigou Valley near the Weixin County town, Yunnan Province; 2, Type area of Shihtzupu Formation at Donggongsi, Zunyi, Guizhou Province; 3, Dashaba Formation at Shuanghe, Changning, Sichuan Province."figureDoi="http://doi.org/10.5281/zenodo.13620321"httpUri="https://zenodo.org/record/13620321/files/figure.png"pageId="20"pageNumber="385">
). Dorsal valve about three−fifths as long as wide, deeply concave, with greatest concavity at junction between visceral area and geniculation; interarea low, shorter than one−tenth length of shell, anacline; notothyrium narrow, short, covered by arched chilidium (
<figureCitationid="50D109E407611016FDDDBCABFD489065"box="[535,641,1777,1802]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">Fig. 13H</figureCitation>
<figureCitationid="50D109E407611016FD4BBCA7FD439062"box="[641,650,1789,1805]"captionStart="Fig"captionStartId="2.[102,135,1189,1209]"captionTargetBox="[119,1466,226,1162]"captionTargetId="graphics-319@2.[152,1466,243,1162]"captionTargetPageId="2"captionText="Fig. 2. Middle Ordovician sedimentary facies on the Yangtze Platform, South China palaeoplate (modified from Zhang et al. 2002), with three fossil localities discussed in the text: 1, Shihtzupu Formation at Shizigou Valley near the Weixin County town, Yunnan Province; 2, Type area of Shihtzupu Formation at Donggongsi, Zunyi, Guizhou Province; 3, Dashaba Formation at Shuanghe, Changning, Sichuan Province."figureDoi="http://doi.org/10.5281/zenodo.13620321"httpUri="https://zenodo.org/record/13620321/files/figure.png"pageId="20"pageNumber="385">
). Unequal parvicostellae; 7–9 accentuated costae, with finer costellae inserting three times; about 9–11 fine costellae between two adjacent costae at shell anterior margin. Concentric fila evenly spaced, about 15–16 per mm.
Teeth small, wedge−shaped; dental plates short, high, subparallel, continuous with anterior bounding ridge of muscle field. Muscle field subtriangular to subpentagonal, about one−fifth length and width of shell, elevated anteriorly; adductor scars small, narrow, mainly in posteromedial part of muscle field, separated from diductor scars by pair of thin plates in some shells (
<figureCitationid="50D109E407611016FBEBB973FB41942D"box="[1057,1160,809,834]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">Fig. 13D</figureCitation>
); diductor scars much larger, subcircular, surrounding adductor scars laterally and anteriorly. Mantle canal system saccate; vascula media originating from anterior ends of diductor scars, with long, straight, weakly divergent main trunks. Bema relatively large, well−developed; dorsal platform marked by weak, discrete septules.
.—The convexity of ventral valves has a certain degree of variation: the visceral area is usually much more convex than the shell marginal area, giving the valve a galeate shape (e.g.,
<figureCitationid="50D109E407611016FBD8BE2BFBBF93E5"box="[1042,1142,1137,1162]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">Fig. 13E</figureCitation>
<figureCitationid="50D109E407611016FBBCBE27FBB693E2"box="[1142,1151,1149,1165]"captionStart="Fig"captionStartId="1.[102,135,1706,1726]"captionTargetBox="[102,773,226,1679]"captionTargetId="graphics-523@1.[102,773,226,1679]"captionTargetPageId="1"captionText="Fig. 1. Location map of the study area. A. Map of China, with the South China palaeoplate outlined by thick line; inset box enlarged as B. B. The border region between Yunnan, Guizhou and Sichuan provinces; inset enlarged as C. C. Detailed map of the Shizigou Valley section near the Weixin County town, with the fossil locality marked by solid triangle."figureDoi="http://doi.org/10.5281/zenodo.13620319"httpUri="https://zenodo.org/record/13620319/files/figure.png"pageId="20"pageNumber="385">
<figureCitationid="50D109E407611016FC14BECBFB9893C5"box="[990,1105,1169,1194]"captionStart="Fig"captionStartId="17.[102,135,1843,1863]"captionText="Fig. 12. A. Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press); NIGP139131, ventral internal mould (A) and latex cast (A) preserved to− + 1 2 gether with a bryozoan colony (top left). B–D. Leptellinidae gen. nov., sp. B. NIGP139132, dorsal internal mould (B1) and latex cast (B2). C. NIGP139133, ventral external mould (C1) and latex cast (C2). D. NIGP139134, ventral external mould. E–M. Calyptolepta huanghuaensis (Chang, 1983). E. NIGP139135, ventral internal mould. F. NIGP139136, broken ventral internal mould. G. NIGP139137, dorsal internal mould. H. NIGP139138, holotype, dorsal internal mould (H1) and latex cast (H2) with local enlargement (H3) showing cardinalia and bema. I. NIGP139139, dorsal external mould. J. NIGP139140, dorsal internal mould of a juvenile shell. K. NIGP139141, dorsal internal mould. L. NIGP139142, two dorsal internal moulds. M. NIGP139143, dorsal internal mould. N. Leptestiina veturna sp. nov.; NIGP139144, paratype, ventral internal mould."pageId="20"pageNumber="385">Figs. 12N</figureCitation>
<figureCitationid="50D109E407611016FB96BECBFB4293C5"box="[1116,1163,1169,1194]"captionStart="Fig"captionStartId="21.[102,135,1426,1446]"captionTargetBox="[102,1485,219,1398]"captionTargetId="figure-161@21.[102,1484,219,1392]"captionTargetPageId="21"captionText="Fig. 13. A–I. Leptestiina veturna sp. nov. A. NIGP139145, paratype, ventral internal mould. B. NIGP139146, paratype, ventral internal mould. C. NIGP139147, holotype, ventral internal mould (C1) and latex cast (C2). D. NIGP139148, paratype, ventral internal mould. E. NIGP139149, paratype, ventral internal mould (E1) and latex cast (E2). F. NIGP139150, paratype, dorsal internal mould. G. NIGP139151, paratype, three dorsal external moulds, with dorsal external mould of Leptellinidae gen. et sp. nov. (to be erected by Zhan and Jin in press) (second from right). H. NIGP139152, paratype, dorsal external mould (H1) and latex cast (H2); note also ventral interarea. I. NIGP139153, paratype, broken dorsal internal mould. J. Strophomenid gen. et sp. indeterminate; NIGP139154, dorsal internal mould (J1) and latex cast (J2), together with a dorsal internal mould of Leptellina spatiosa sp. nov. (top)."figureDoi="http://doi.org/10.5281/zenodo.13620331"httpUri="https://zenodo.org/record/13620331/files/figure.png"pageId="20"pageNumber="385">13C</figureCitation>
<figureCitationid="50D109E407611016FB41BEC7FB5D93C2"box="[1163,1172,1181,1197]"captionStart="Fig"captionStartId="1.[102,135,1706,1726]"captionTargetBox="[102,773,226,1679]"captionTargetId="graphics-523@1.[102,773,226,1679]"captionTargetPageId="1"captionText="Fig. 1. Location map of the study area. A. Map of China, with the South China palaeoplate outlined by thick line; inset box enlarged as B. B. The border region between Yunnan, Guizhou and Sichuan provinces; inset enlarged as C. C. Detailed map of the Shizigou Valley section near the Weixin County town, with the fossil locality marked by solid triangle."figureDoi="http://doi.org/10.5281/zenodo.13620319"httpUri="https://zenodo.org/record/13620319/files/figure.png"pageId="20"pageNumber="385">
, ranging from early Caradocian to mid−Ashgillian (
<bibRefCitationid="AC7B689007611016FAC4BE8BFCAE9265"author="Cocks, L. R. M. & Rong Jia-yu"pageId="20"pageNumber="385"pagination="77 - 163"refId="ref17978"refString="Cocks, L. R. M. and Rong Jia-yu 1989. Classification and review of the brachiopod superfamily Plectambonitacea. Bulletin of the British Museum (Natural History), Geology 45: 77 - 163."type="journal article"year="1989">Cocks and Rong 1989</bibRefCitation>
,
<bibRefCitationid="AC7B689007611016FCB8BEABFC669265"author="Cocks, L. R. M. & Rong Jia-yu"box="[882,943,1265,1290]"pageId="20"pageNumber="385"pagination="216 - 349"refId="ref18017"refString="Cocks, L. R. M. and Rong Jia-yu 2000. Strophomenida. In: R. L. Kaesler (ed.), Treatise on Invertebrate Paleontology, Part H. Brachiopoda (revised), Vol. 2, 216 - 349. Geological Society of America and University of Kansas Press, Lawrence."type="book chapter"year="2000">2000</bibRefCitation>
fauna in the upper Pingliang (upper Caradocian) and the Linhsiang (lower Ashgillian) formations (
<bibRefCitationid="AC7B689007611016FC3ABF0BFB9B9205"author="Fu Li-pu"box="[1008,1106,1361,1386]"pageId="20"pageNumber="385"pagination="95 - 178"refId="ref18403"refString="Fu Li-pu 1982. Brachiopoda [in Chinese]. In: Xian Institute of Geology and Mineral Resources (ed.), Paleontological Atlas of Northwest China, Shaanxi-Gansu-Ningxia Volume, pt. 1, Precambrian and Early Palaeozoic, 95 - 178. Geological Publishing House, Beijing."type="book chapter"year="1982">Fu 1982</bibRefCitation>
; Rong et al. 1994). The new species of late Darriwilian age is probably the oldest known of the genus. In comparison, the
<bibRefCitationid="AC7B689007611016FCE6BFEBFC1592A5"author="Havlicek, V."box="[812,988,1457,1482]"pageId="20"pageNumber="385"pagination="397 - 428"refId="ref18712"refString="Havlicek, V. 1952. On the Ordovician representatives of the family Plectambonitidae (Brachiopoda). Sbornik Ustredniho ustavu geologickeho 19: 397 - 428."type="book chapter"year="1952">Havlíček, 1952</bibRefCitation>
</taxonomicName>
from the Králův Dvůr Formation (middle Ashgillian) of
