treatments-xml/data/BE/25/42/BE2542944F5EC52B4409F71CB0D2B0D3.xml

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<mods:title id="AB287D1432FDFB23E2420FF1A52C0A13">Review of the Berosus Leach of Venezuela (Coleoptera, Hydrophilidae, Berosini) with description of fourteen new species</mods:title>
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<mods:namePart id="59949F1AE604D68E7A933A0D79497CD3">Oliva, Adriana</mods:namePart>
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<mods:namePart id="F2BA82755476061125D9A52442DA587F">Short, Andrew E. Z.</mods:namePart>
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<treatment id="BE2542944F5EC52B4409F71CB0D2B0D3" ID-GBIF-Taxon="152036269" LSID="urn:lsid:plazi:treatment:BE2542944F5EC52B4409F71CB0D2B0D3" httpUri="http://treatment.plazi.org/id/BE2542944F5EC52B4409F71CB0D2B0D3" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" scope_family="Hydrophilidae" scope_order="Coleoptera" scope_tribe="Berosini">
<subSubSection id="2C6359B5E5D79374B885891E142CF360" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="733338D5BA3C40C3B818A590026A93A8" pageId="49" pageNumber="50">
<taxonomicName id="1770008FB0BC00DE82A70B9D57D0C670" ID-CoL="LM7P" LSID="urn:lsid:zoobank.org:act:C2EBCCA6-8CC0-4195-865B-BE4124552305" authority="Castelnau, 1840" authorityName="Castelnau" authorityYear="1840" class="Insecta" family="Hydrophilidae" genus="Berosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Berosus truncatipennis" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="truncatipennis">Berosus truncatipennis Castelnau, 1840</taxonomicName>
Fig. 21A
</paragraph>
</subSubSection>
<subSubSection id="465591DDEF4AC30CEFD0DE7D28B56958" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="19A5EF34B9B47366C98D32532762C98C" pageId="49" pageNumber="50">
<taxonomicName id="9EB14D9D219619ABFA73FC9C721F2290" class="Insecta" family="Hydrophilidae" genus="Berosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Berosus truncatipennis" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="truncatipennis">Berosus truncatipennis</taxonomicName>
Castelnau, 1840: 56.
</paragraph>
<paragraph id="73EADD71ACEE75757A5AFFA52232FEA9" pageId="49" pageNumber="50">
<taxonomicName id="6E6E3D9EEAC2EFC0050B06D1E648D6E4" class="Insecta" family="Hydrophilidae" genus="Berosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Berosus truncatipennis" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="truncatipennis">Berosus truncatipennis</taxonomicName>
Castelnau: Oliva, 1989: 184.
</paragraph>
</subSubSection>
<subSubSection id="79EF45CE2A0AC824F3827AB1AE547ED5" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="material examined">
<paragraph id="7C8AF7AFFFCA4665239F6AB2592E4B35" pageId="49" pageNumber="50">Material examined</paragraph>
<paragraph id="23978448D169AD56734D995938DA504A" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
(269):VENEZUELA: 
<normalizedToken id="0134A6538C421BC0C652033DD95B8F91" originalValue="Anzoátegui">Anzoategui</normalizedToken>
State: 
<geoCoordinate id="38D605B1427773C7139D1F2764ADB167" direction="north" orientation="latitude" precision="1" value="9.111834">09°06'42.6&quot;N</geoCoordinate>
, 
<geoCoordinate id="DA7DF87B329F66EE4A84026F67DA178E" direction="west" orientation="longitude" precision="1" value="-64.15561">64°09'20.2&quot;W</geoCoordinate>
, 228 m, Permanent pond, 12.viii.2009, Cordero, VZ09-812-06A (1 ex.); 
<geoCoordinate id="07BB96295167C17E23A8FEE710BCB7AD" direction="north" orientation="latitude" precision="1" value="9.328167">09°19'41.4&quot;N</geoCoordinate>
, 
<geoCoordinate id="AF40FAC195F06B977FCA0101454CA36B" direction="west" orientation="longitude" precision="1" value="-64.118645">64°07'07.1&quot;W</geoCoordinate>
, 289 m, Temporal rain pond, 13.viii.2009, leg. Cordero, VZ09-813-07A (1 ex.); 
<geoCoordinate id="114331C3F36C41C75C42CFD9428755B1" direction="north" orientation="latitude" precision="1" value="9.276278">09°16'34.6&quot;N</geoCoordinate>
, 
<geoCoordinate id="887D8B2F72A8A2A2A056AF2665E231C2" direction="west" orientation="longitude" precision="1" value="-64.227585">64°13'39.3&quot;W</geoCoordinate>
, 259 m, Temporal pond aside the road, 14.viii.2009, leg. Cordero, VZ09-814-12A (4 exs.); 
<geoCoordinate id="CD207DA1F0BEC6A62F069CBF8E814330" direction="north" orientation="latitude" precision="1" value="9.287862">09°17'16.3&quot;N</geoCoordinate>
, 
<geoCoordinate id="16B7827C38001F72424B36B16B191B9C" direction="west" orientation="longitude" precision="1" value="-64.22753">64°13'39.1&quot;W</geoCoordinate>
, 274 m, Temporal pond aside the road, 15.viii.2009, leg. Cordero, VZ09-815-12A (2 exs.). Apure State: S. San Fernando, 
<geoCoordinate id="C202992D68183B5E54CB96E94989EB54" direction="north" orientation="latitude" precision="1" value="7.8693166">7°52.159'N</geoCoordinate>
, 
<geoCoordinate id="5FC25E81E258EA541F01B17A3D2531BA" direction="west" orientation="longitude" precision="1" value="-67.48766">67°29.260'W</geoCoordinate>
, 52 m, 3.i.2006, light on top of house, leg. Short, AS-06-006 (1 ex.); ca. 18 km S. San Fernando, 
<geoCoordinate id="517DE50E2F42A10C6CD3026BB0B8A28B" direction="north" orientation="latitude" precision="1" value="7.564483">7°33.869'N</geoCoordinate>
, 
<geoCoordinate id="592EBBB292425797D0AB9820B9351CF0" direction="west" orientation="longitude" precision="1" value="-67.64093">67°38.456'W</geoCoordinate>
, 50 m, marsh along road; 4.i.2006, leg. Short, AS-06-008 (6 exs.); Road between Rio Capanaparo and Rio Cinaruco; Medanos de la Soledad, 
<geoCoordinate id="C0F4CF24409C603D78F0C758BE24D167" direction="north" orientation="latitude" precision="1" value="7.33625">7°20.175'N</geoCoordinate>
, 
<geoCoordinate id="FFAD139A708EF3AEAD0515BB4CBCF381" direction="west" orientation="longitude" precision="1" value="-67.73113">67°43.868'W</geoCoordinate>
, 49 m, marshy area, 17.i.2009, leg. Short, Camacho, Miller, VZ09-0117-02X (2 exs.); Rt. 19, E. Apurito, 
<geoCoordinate id="FF872382B5A66F8BB6B027FD2AB39472" direction="north" orientation="latitude" precision="1" value="7.913717">7°54.823'N</geoCoordinate>
, 
<geoCoordinate id="6B8340535F4F84413CFBC09AFDC18377" direction="west" orientation="longitude" precision="1" value="-68.4297">68°25.782'W</geoCoordinate>
, shaded 
<normalizedToken id="06A8E67B1AB38E92D3FF2C38D567FCA3" originalValue="“pond”">&quot;pond&quot;</normalizedToken>
by road, 18.i.2009, leg. Short, 
<normalizedToken id="37112B3482D7252E5CA27E6934967158" originalValue="García">Garcia</normalizedToken>
, &amp; Camacho, VZ09-0118-01X (4 exs.); between &quot;La Ye&quot; &amp; Bruzual, 7 38.660'N, 69 18.004'W, 90 m, lagoon, 18.i.2009, Short, Camacho &amp; 
<normalizedToken id="61E06468DC55802F3C49F74FC843F331" originalValue="García">Garcia</normalizedToken>
, VZ09-0118-03X (1 ex.). Aragua State: &quot;Venez. Cagua/Edo. Aragua/28-VI-1961 Bordon.&quot; (32 exs., MSUC). Delta Amacuro State: 
<geoCoordinate id="C58F9B3E86ABD9BC89FED11E047BBF42" direction="north" orientation="latitude" precision="1" value="9.137222">09°08'14.0&quot;N</geoCoordinate>
, 
<geoCoordinate id="E381DB4B561C513BB46326CDF7E84FB9" direction="west" orientation="longitude" precision="1" value="-61.958694">61°57'31.3&quot;W</geoCoordinate>
, 6 m, Bank of a swamp formed by Orinoco flooding, 26.viii.2009, leg. Cordero, VZ09-826-27A (1 ex.); 
<geoCoordinate id="C71C4C8A3625E97A7DD1CF5C2391E8AB" direction="north" orientation="latitude" precision="1" value="9.101028">09°06'03.7&quot;N</geoCoordinate>
, 
<geoCoordinate id="37B245D9300A66FD789D2B5988814229" direction="west" orientation="longitude" precision="1" value="-61.97914">61°58'44.9&quot;W</geoCoordinate>
, 4 m, Bank of a swamp formed by Orinoco flooding, 27.viii.2009, leg. Cordero, VZ09-827-29A (3 exs.); 
<geoCoordinate id="42C3C8EEC062D33C9D25444B44139C40" direction="north" orientation="latitude" precision="1" value="9.144389">09°08'39.8&quot;N</geoCoordinate>
, 
<geoCoordinate id="F51DE7C6BB42B924E45A240DD11AE1F9" direction="west" orientation="longitude" precision="1" value="-61.929447">61°55'46.0&quot;W</geoCoordinate>
, 3 m, Permanent grass pond, 27.viii.2009, leg. Cordero, VZ09-827-31A (1 ex.); 09°012'00.1&quot;N, 
<geoCoordinate id="E7F9A123CF9CB5DFC0ABF1BC1AB5D737" direction="west" orientation="longitude" precision="1" value="-61.88925">61°53'21.3&quot;W</geoCoordinate>
, 3 m, Flooded pond, 
<pageBreakToken id="DCB3D0BA56A191509F6695DB171BFE21" pageId="50" pageNumber="51" start="start">28</pageBreakToken>
.viii.2009, leg. Cordero, VZ09-828-35A (2 exs.). 
<normalizedToken id="FACE3FF50D8CD0DEF7DB3E2C78388EBD" originalValue="Falcón">Falcon</normalizedToken>
State: E. Dabajuro, River at Bridge crossing on Rt. 3, 
<geoCoordinate id="87062123DECE1AFDFA73612F35E1AE5F" direction="north" orientation="latitude" precision="1" value="11.038417">11°02.305'N</geoCoordinate>
, 
<geoCoordinate id="346992EE9D6B9EE7B4CE4215D6B2A2DC" direction="west" orientation="longitude" precision="1" value="-70.64111">70°38.467'W</geoCoordinate>
, 98 m, Gravel and sand margins of river and associated side pools, 8.vii.2009, leg. Short, Gustafson, &amp; Inciarte, VZ09-0708-01A (1 ex.); 
<normalizedToken id="A46AC0971B4AEE19D472AC03CB8B5CAB" originalValue="Médanos">Medanos</normalizedToken>
de Coro, pond in dunes, 
<geoCoordinate id="AFB7F06B323658AB9FEE3B96D6507574" direction="north" orientation="latitude" precision="1" value="11.436916">11°26.215'N</geoCoordinate>
, 
<geoCoordinate id="64DC7D1DC84E841A9DD9018E5E96E76D" direction="west" orientation="longitude" precision="1" value="-69.66853">69°40.112'W</geoCoordinate>
, 8 m, pond in dunes, 9.vii.2009, leg. Shepard, VZ09-0709-03Z (13 exs.); SE of Tocopero, 
<geoCoordinate id="87F4D8FF4991CB3E5EA790649077918E" direction="north" orientation="latitude" precision="1" value="11.4487">11°26.922'N</geoCoordinate>
, 
<geoCoordinate id="2325D883B635121BB9253C66BA08B3F0" direction="west" orientation="longitude" precision="1" value="-69.21848">69°13.109'W</geoCoordinate>
, 12 m, Puddle in roadside ditch and culvert, 10.vii.2009, leg. Sites, VZ09-0710-03S (7 exs.); same locality but leg. Short et al., VZ09-0710-03B (23 exs.); same locality but leg. Short et al., VZ09-0710-03C (53 exs.). 
<normalizedToken id="6A1150AA9E726B40FDC22E308CFDAE9C" originalValue="Guárico">Guarico</normalizedToken>
State: Corozo Pando (8 km N.), 17-18.vi.1984, leg. F.W. Eiland &amp; V. Linares, blacklight (11 exs.); 15 km S. Calabozo, 9-13.ii.1969, Lago de los Patos, leg. P. &amp; P. Spangler (11 exs.); 20 km S. Calabozo, 8-13.ii.1969, Rio Orituco, leg. P. &amp; P. Spangler (2 exs., USNM); 44 km S. Calabozo, Hato Masaguaral, 6.iii.1986, P. Spangler &amp; S. Beaujon, Collection # 31 (9 exs., USNM); ca. 35 km N. San Fernando, 
<geoCoordinate id="7B5B8E9FCE8C01D0C4FDA1C57148DB9D" direction="north" orientation="latitude" precision="1" value="8.192966">8°11.578'N</geoCoordinate>
, 
<geoCoordinate id="710B35EE040AD002FB41C21F28967B11" direction="west" orientation="longitude" precision="1" value="-67.60478">67°36.287'W</geoCoordinate>
, 53 m, marshy area by road, 10.ix.2007, leg. A. Short, AS-07-002 (1 ex.); nr. Socorro, 
<geoCoordinate id="C1D6E5E7E672FFA0E279DCA211DFD281" direction="north" orientation="latitude" precision="1" value="8.983611">8°59'1.0&quot;N</geoCoordinate>
, 
<geoCoordinate id="50344124823E17F80B1BCDBA657CC4FC" direction="west" orientation="longitude" precision="1" value="-65.73855">65°44'18.8&quot;W</geoCoordinate>
, 100 m, muddy ditch, 29.vii.2008, leg. Short &amp; 
<normalizedToken id="91317362C9F0A350CA9376882A2889F7" originalValue="García">Garcia</normalizedToken>
, AS-08-050 (1 ex.); N. of Palenque, 
<geoCoordinate id="D60089C465398F26617B035F61CA7764" direction="north" orientation="latitude" precision="1" value="9.113234">9°6.794'N</geoCoordinate>
, 
<geoCoordinate id="A70BC8DA3E4F64CF1A2D52EE9870D633" direction="west" orientation="longitude" precision="1" value="-66.99325">66°59.595'W</geoCoordinate>
, 152 m, stream at road crossing, 8.i.2009, leg. Short, 
<normalizedToken id="35031F1C948FA3CFD75A4782EB5A20C7" originalValue="García">Garcia</normalizedToken>
, Miller, Camacho, Joly, VZ09-0108-03X (3 exs.); on side road ca. 10 km E. Mantecal, 
<geoCoordinate id="6891F5F44D8C2A34BDDF3D965E5A5F19" direction="north" orientation="latitude" precision="1" value="7.6216335">7°37.298'N</geoCoordinate>
, 
<geoCoordinate id="E2B04BB21955749D19A37063736CCB6C" direction="west" orientation="longitude" precision="1" value="-69.06132">69°3.679'W</geoCoordinate>
, 83 m, marshy area and pool by road, 18.i.2009, leg. Short, 
<normalizedToken id="69869CE663CEBA0F189B5106D0C475FD" originalValue="García">Garcia</normalizedToken>
, &amp; Camacho, VZ09-0118-02X (4 exs.); ca. 65 km S. Las Mercedes, 8 31.705'N, 66 22.602'W, 145 m, large lagoon with vegetation, 9.i.2009, leg. Short, Miller, 
<normalizedToken id="41038EE4066D113568D1276515D4979F" originalValue="García">Garcia</normalizedToken>
, &amp; Camacho, VZ09-0109-01X (2 ex.); W. of Las Mercedes, 
<geoCoordinate id="F9D4EC1C6A395A8028C026375E120793" direction="north" orientation="latitude" precision="1" value="9.08445">9°5.067'N</geoCoordinate>
, 
<geoCoordinate id="54CD9DBD34D5C86E36F475C98A6CE7D5" direction="west" orientation="longitude" precision="1" value="-66.475">66°28.500'W</geoCoordinate>
, 153 m, 8.i.2009, leg. Short &amp; Miller, roadside pond, VZ09-0108-04A (1 ex.); San Nicolasito Field Station, 
<geoCoordinate id="CD4015DC43C69EFDC86F3161082B5661" direction="north" orientation="latitude" precision="1" value="8.144617">8°8.677'N</geoCoordinate>
, 
<geoCoordinate id="0E3514935A56B65BEA277BCC5FD0E217" direction="west" orientation="longitude" precision="1" value="-66.40438">66°24.263'W</geoCoordinate>
, 50 m, pool without fish, 10.i.2009, leg. Miller &amp; 
<normalizedToken id="574FD005A5F8B121E23E9B2ADF760715" originalValue="García">Garcia</normalizedToken>
, VZ09-0110-03A (2 ex.). Portuguesa State: Guanare, 26.ii.1969, leg. P. &amp; P. Spangler (1 ex., USNM); Rio Guanare, N. of Guanare, 
<geoCoordinate id="1A5B1F73EC5530B7A0376BFC965999C5" direction="north" orientation="latitude" precision="1" value="8.42955">8°25.773'N</geoCoordinate>
, 
<geoCoordinate id="211C92C270ECD34CADC63DA9868BD134" direction="west" orientation="longitude" precision="1" value="-69.5867">69°35.202'W</geoCoordinate>
, 185 m, side rocky pools, 19.i.2009, leg. Short, VZ09-0119-02C (7 exs.). Sucre State: Finca Vuelta Larga, 
<geoCoordinate id="FAA1B4C657AAEEB96E6A328B6EFA8C12" direction="north" orientation="latitude" precision="1" value="10.50125">10°30.075'N</geoCoordinate>
, 
<geoCoordinate id="EFCE782894C153B105F9712549BFABA5" direction="west" orientation="longitude" precision="1" value="-63.103615">63°06.217'W</geoCoordinate>
, 10 m, muddy puddle in road with detritus, 29.i.2010, leg. Short, 
<normalizedToken id="EC6306C4371BF0F41C5B1B3EE2E447BA" originalValue="García">Garcia</normalizedToken>
, Joly, VZ10-0129-03A (2 exs.). Zulia State: 40 km SW Machiques, Rio Tucuco, 3.vii.1984, blacklight trap, leg. Eiland &amp; Linares (1 ex., USNM); Farm of the Universidad del Zulia (Ana Maria Campo), W. of Maracaibo, stock tank, 20.i.2006, leg. Short, AS-06-037 (1 ex.). When not indicated, material is distributed between MIZA, MALUZ, SEMC, and NMW.
</paragraph>
</subSubSection>
<subSubSection id="284090B97DE0703807E09D4AD0B9AFFA" pageId="50" pageNumber="51" type="distribution">
<paragraph id="9F19484F88A66A5D1DDA30CF03AB9A5B" pageId="50" pageNumber="51">Distribution.</paragraph>
<paragraph id="EBABE0A01905E62DABB59B10E2D50C48" pageId="50" pageNumber="51">
Widespread in tropical and subtropical South America, including NE Argentina, Bolivia, Brazil, Paraguay, and Venezuela (
<normalizedToken id="05D999F2EC2DD8EADAB2279EA12D7342" originalValue="Anzoátegui">Anzoategui</normalizedToken>
, Apure, Aragua, Delta Amacuro, 
<normalizedToken id="3C6DB4979EEFF088723721A8CC6CF0F2" originalValue="Falcón">Falcon</normalizedToken>
, 
<normalizedToken id="1A6845EA667C36206A0906FBABB3ACAA" originalValue="Guárico">Guarico</normalizedToken>
, Portuguesa, Sucre, Zulia). Records for Central America and the Caribbean need confirmation. Previously recorded from Venezuela (Apure) by 
<bibRefCitation id="3DE95090A33EE4EF71B2DAD115F69971" author="Oliva, A" journalOrPublisher="Belgian Journal of Entomology" pageId="58" pageNumber="59" pagination="97 - 103" title="A new species of Berosus (Coleoptera: Hydrophilidae) from Venezuela." volume="4" year="2002">Oliva (2002)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="81DEDA14D6A2FB6AFCAA29AA837917AA" pageId="50" pageNumber="51" type="remarks">
<paragraph id="6E12E6D7EAFD3998FDD67299E2DC9526" pageId="50" pageNumber="51">Remarks.</paragraph>
<paragraph id="35E3AC035D32B6BA50FADE32F0C9A999" pageId="50" pageNumber="51">
The apex of the median lobe is a fine membranous dome, which may be considerably deformed in dry-mounted specimens. See discussion of 
<taxonomicName id="C4D2821150117EBB3611015D19E47F1C" class="Insecta" family="Hydrophilidae" genus="Berosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Berosus megaphallus" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="megaphallus">Berosus megaphallus</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
</treatment>
</document>