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<document id= "B032F14E4F7F9CFAF4DF08466D2373B1" ID-DOI= "10.15298/rusentj.32.1.03" ID-ISSN= "0132-8069" ID-Zenodo-Dep= "10949096" IM.bibliography_approvedBy= "tatiana" IM.illustrations_approvedBy= "tatiana" IM.materialsCitations_approvedBy= "tatiana" IM.taxonomicNames_approvedBy= "julia" IM.treatmentCitations_approvedBy= "tatiana" IM.treatments_approvedBy= "tatiana" checkinTime= "1712198792598" checkinUser= "felipe" docAuthor= "Fedorenko, D. N." docDate= "2023" docId= "2F4187C3FFCF1669784FFCB020AE77F7" docLanguage= "en" docName= "RussEntomolJ.32.1.16-39.pdf" docOrigin= "Russian Entomological Journal 32 (1)" docSource= "http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle= "DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.4:RussEntomolJ.2019-.journal_article" docStyleId= "0D5BAC5AA0C130B3809AF578A601D837" docStyleName= "RussEntomolJ.2019-.journal_article" docStyleVersion= "4" docTitle= "Pterostichus (Morphohaptoderus) unisetosus Fedorenko 2023, sp.n." docType= "treatment" docVersion= "4" lastPageNumber= "20" masterDocId= "D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle= "New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber= "39" masterPageNumber= "16" pageNumber= "20" updateTime= "1731592677940" updateUser= "julia" >
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<mods:title id= "7A41A306F9A2C941E4DFC62963065070" > New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:namePart id= "785FA76C704780E6F1FD345166CB8C38" > Fedorenko, D. N.</mods:namePart>
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<paragraph id= "A75736D5FFCF1669784FFCB0231B716F" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
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<taxonomicName id= "60E84D56FFCF1669784FFCB020D97150" ID-CoL= "8MBT6" authority= "Fedorenko, 2023" authorityName= "Fedorenko" authorityYear= "2023" class= "Insecta" family= "Carabidae" genus= "Pterostichus" kingdom= "Animalia" order= "Coleoptera" pageId= "4" pageNumber= "20" phylum= "Arthropoda" rank= "species" species= "unisetosus" subGenus= "Morphohaptoderus" >
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<emphasis id= "959CEAC7FFCF1669784FFCB020CC714D" box= "[297,431,810,834]" italics= "true" pageId= "4" pageNumber= "20" > Pterostichus</emphasis>
(
<emphasis id= "959CEAC7FFCF166978A5FCB023F2714D" box= "[451,657,810,834]" italics= "true" pageId= "4" pageNumber= "20" > Morphohaptoderus</emphasis>
)
<emphasis id= "959CEAC7FFCF1669782DFCD220D97150" box= "[331,442,840,863]" italics= "true" pageId= "4" pageNumber= "20" > unisetosus</emphasis>
</taxonomicName>
Fedorenko,
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<taxonomicNameLabel id= "8EAF57BCFFCF16697B25FCD2231B716F" box= "[579,632,840,864]" pageId= "4" pageNumber= "20" rank= "species" > sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id= "EFF2655EFFCF16697848FCFF23F67172" box= "[302,661,869,893]" pageId= "4" pageNumber= "20" type= "description" >
<paragraph id= "A75736D5FFCF16697848FCFF23F67172" blockId= "4.[189,776,810,1528]" box= "[302,661,869,893]" pageId= "4" pageNumber= "20" >
<figureCitation id= "3FD32A50FFCF16697848FCFF20107172" box= "[302,371,869,893]" captionStart= "Figs 1– 5" captionStartId= "2.[226,263,1969,1988]" captionTargetBox= "[198,1388,321,1922]" captionTargetId= "figure-102@2.[188,1398,317,1927]" captionTargetPageId= "2" captionText= "Figs 1– 5. Dorsal habitus: 1 — Pterostichus glabellus sp.n., paratype ♀; 2 — P. unisetosus sp.n., holotype; 3 — P. primitivus sp.n., holotype; 4 — P. subirideus sp.n., paratype ♂; 5 — P. batdaiensis sp.n., paratype ♂ from Sa Pa env. Рис. 1– 5. Габитус дорЗально: 1 — Pterostichus glabellus sp.n., паратип ♀; 2 — P. unisetosus sp.n., голотип; 3 — P. primitivus sp.n., голотип; P. subirideus sp.n., паратип ♂; 5 — P. batdaiensis sp.n., паратип ♂ иЗ окрестностей Sa Pa." figureDoi= "http://doi.org/10.5281/zenodo.10949098" httpUri= "https://zenodo.org/record/10949098/files/figure.png" pageId= "4" pageNumber= "20" > Figs 2</figureCitation>
,
<figureCitation id= "3FD32A50FFCF16697819FCFF20A77172" box= "[383,452,869,893]" captionStart= "Figs 12– 21" captionStartId= "5.[226,263,1945,1964]" captionTargetBox= "[189,1399,319,1919]" captionTargetId= "figure-122@5.[189,1399,314,1923]" captionTargetPageId= "5" captionText= "Figs 12– 21. Aedeagus median lobe: 12– 13 — Pterostichus glabellus sp.n.; 14– 15 — P. unisetosus sp.n.; 16– 17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20– 21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm. Рис. 12– 21. СреднЯЯ долЯ Эдеагуса: 12– 13 — Pterostichus glabellus sp.n.; 14– 15 — P. unisetosus sp.n.; 16– 17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20– 21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм." figureDoi= "http://doi.org/10.5281/zenodo.10949104" httpUri= "https://zenodo.org/record/10949104/files/figure.png" pageId= "4" pageNumber= "20" > 14– 15</figureCitation>
,
<figureCitation id= "3FD32A50FFCF166978B6FCFF23757172" box= "[464,534,869,893]" captionStart= "Figs 22– 34" captionStartId= "7.[226,263,1921,1940]" captionTargetBox= "[190,1399,697,1904]" captionTargetId= "figure-163@7.[189,1399,694,1904]" captionText= "Figs 22– 34. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28– 29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22– 26 — median lobe; 28– 34 — right paramere, scale as for respective median lobe; 22– 25, 27 — ventrobasal aspect; 29– 34 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 22– 34. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28– 29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22– 26 — среднЯЯ долЯ; 28– 34 — праваЯ парамера в масштабе соответствуюЩей средней доли; 22– 25, 27 — вентробаЗально; 29– 34 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi= "http://doi.org/10.5281/zenodo.10949106" httpUri= "https://zenodo.org/record/10949106/files/figure.png" pageId= "4" pageNumber= "20" > 23, 31</figureCitation>
, 56– 57, 68.
</paragraph>
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<subSubSection id= "EFF2655EFFCF16697984FC10232A71F2" pageId= "4" pageNumber= "20" type= "materials_examined" >
<paragraph id= "A75736D5FFCF16697984FC1023F871EA" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
<materialsCitation id= "17803C88FFCF16697984FC1023F871EA" collectingDate= "2018-10" collectionCode= "ZMMU, SIEE" collectorName= "Y Ty & Bat Xat" elevation= "1750" location= "Lao Cai" pageId= "4" pageNumber= "20" specimenCount= "5" specimenCount-female= "1" specimenCount-male= "2" stateProvince= "Lao Cai" typeStatus= "holotype" >
MATERIAL.
<typeStatus id= "78538877FFCF16697838FC1020CD7192" box= "[350,430,906,925]" pageId= "4" pageNumber= "20" type= "holotype" > Holotype</typeStatus>
<specimenCount id= "B1EEFD5CFFCF166978D0FC1D20A97192" box= "[438,458,903,925]" count= "1" pageId= "4" pageNumber= "20" type= "male" > ♂</specimenCount>
(
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) and
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,
<specimenCount id= "B1EEFD5CFFCF16697BBFFC1D238E7192" box= "[729,749,903,925]" count= "1" pageId= "4" pageNumber= "20" type= "male" > ♂</specimenCount>
,
<specimenCount id= "B1EEFD5CFFCF16697B9DFC1D22647192" box= "[763,775,903,925]" count= "1" pageId= "4" pageNumber= "20" type= "female" > ♀</specimenCount>
(
<collectionCode id= "C1F9AE10FFCF166979A5FC38219071BA" box= "[195,243,930,949]" pageId= "4" pageNumber= "20" > SIEE</collectionCode>
) with label: ‘ N-Vietnam,
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WNW of/
<collectingRegion id= "652CF837FFCF16697B39FC3823C671BA" box= "[607,677,930,949]" country= "Vietnam" name= "Lao Cai" pageId= "4" pageNumber= "20" > Lao Cai</collectingRegion>
, env.
<collectorName id= "0A1D5303FFCF16697BB3FC38226071BA" box= "[725,771,930,949]" pageId= "4" pageNumber= "20" > Y Ty</collectorName>
,
<collectorName id= "0A1D5303FFCF166979DBFC20206471C2" box= "[189,263,954,973]" pageId= "4" pageNumber= "20" > Bat Xat</collectorName>
/ Nat[io]n.[al] Park, h=
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/ 22°37´ -37.6´ N/ 103°37.3´ -38.3´ E, 16-21./ leg. D.Fedorenko
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<collectingDate id= "C312E9FDFFCF16697B2BFC4823F071EA" box= "[589,659,978,997]" pageId= "4" pageNumber= "20" value= "2018-10" > X.2018</collectingDate>
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’ .
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</paragraph>
<paragraph id= "A75736D5FFCF16697984FC70232A71F2" blockId= "4.[189,776,810,1528]" box= "[226,585,1002,1021]" pageId= "4" pageNumber= "20" > Genitalia examined in all the specimens.</paragraph>
</subSubSection>
<subSubSection id= "EFF2655EFFCF16697984FB9823DB7642" pageId= "4" pageNumber= "20" type= "diagnosis" >
<paragraph id= "A75736D5FFCF16697984FB9823DB7642" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
DIAGNOSIS. No significant differences from
<taxonomicName id= "60E84D56FFCF16697BC1FB9921BA763D" authority= "Fedorenko, 2023" authorityName= "Fedorenko" authorityYear= "2023" class= "Insecta" family= "Carabidae" genus= "Pterostichus" kingdom= "Animalia" order= "Coleoptera" pageId= "4" pageNumber= "20" phylum= "Arthropoda" rank= "species" species= "glabellus" status= "sp.n." >
<emphasis id= "959CEAC7FFCF16697BC1FB9921BA763D" italics= "true" pageId= "4" pageNumber= "20" > P. glabellus</emphasis>
</taxonomicName>
<emphasis id= "959CEAC7FFCF166979BBFB87206D763D" bold= "true" box= "[221,270,1053,1074]" pageId= "4" pageNumber= "20" >
<taxonomicNameLabel id= "8EAF57BCFFCF166979BBFB87206D763D" box= "[221,270,1053,1074]" pageId= "4" pageNumber= "20" rank= "species" > sp.n.</taxonomicNameLabel>
</emphasis>
other than the elytron with middle discal seta, d2,
<emphasis id= "959CEAC7FFCF16697B8FFB872399763D" box= "[745,762,1053,1074]" italics= "true" pageId= "4" pageNumber= "20" > fe</emphasis>
3 not sexually dimorphic, and aedeagus is distinctive.
</paragraph>
</subSubSection>
<subSubSection id= "EFF2655EFFCF16697984FBC820957757" pageId= "4" pageNumber= "20" type= "description" >
<paragraph id= "A75736D5FFCF16697984FBC820FE76DD" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
DESCRIPTION. Body as in
<figureCitation id= "3FD32A50FFCF16697B68FBC8232E7668" box= "[526,589,1106,1128]" captionStart= "Figs 1– 5" captionStartId= "2.[226,263,1969,1988]" captionTargetBox= "[198,1388,321,1922]" captionTargetId= "figure-102@2.[188,1398,317,1927]" captionTargetPageId= "2" captionText= "Figs 1– 5. Dorsal habitus: 1 — Pterostichus glabellus sp.n., paratype ♀; 2 — P. unisetosus sp.n., holotype; 3 — P. primitivus sp.n., holotype; 4 — P. subirideus sp.n., paratype ♂; 5 — P. batdaiensis sp.n., paratype ♂ from Sa Pa env. Рис. 1– 5. Габитус дорЗально: 1 — Pterostichus glabellus sp.n., паратип ♀; 2 — P. unisetosus sp.n., голотип; 3 — P. primitivus sp.n., голотип; P. subirideus sp.n., паратип ♂; 5 — P. batdaiensis sp.n., паратип ♂ иЗ окрестностей Sa Pa." figureDoi= "http://doi.org/10.5281/zenodo.10949098" httpUri= "https://zenodo.org/record/10949098/files/figure.png" pageId= "4" pageNumber= "20" > Fig. 2</figureCitation>
. BL 8.4– 8.8 mm. Pronotal base nearly straight, two fifths wider than apex. Elytral stria 7 inside humeral angle; parascutellar striole entire, stria 1 adjoining basal ridge. D2/EL 0.50– 0.57 (0.55, n = 3× 2). USS: 13– 15.
</paragraph>
<paragraph id= "A75736D5FFCF16697984FB4D20E8772C" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
Aedeagus (
<figureCitation id= "3FD32A50FFCF16697834FB4D20A676E2" box= "[338,453,1239,1261]" captionStart= "Figs 12– 21" captionStartId= "5.[226,263,1945,1964]" captionTargetBox= "[189,1399,319,1919]" captionTargetId= "figure-122@5.[189,1399,314,1923]" captionTargetPageId= "5" captionText= "Figs 12– 21. Aedeagus median lobe: 12– 13 — Pterostichus glabellus sp.n.; 14– 15 — P. unisetosus sp.n.; 16– 17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20– 21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm. Рис. 12– 21. СреднЯЯ долЯ Эдеагуса: 12– 13 — Pterostichus glabellus sp.n.; 14– 15 — P. unisetosus sp.n.; 16– 17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20– 21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм." figureDoi= "http://doi.org/10.5281/zenodo.10949104" httpUri= "https://zenodo.org/record/10949104/files/figure.png" pageId= "4" pageNumber= "20" > Figs 14– 15</figureCitation>
,
<figureCitation id= "3FD32A50FFCF166978B4FB4D237676E2" box= "[466,533,1239,1261]" captionStart= "Figs 22– 34" captionStartId= "7.[226,263,1921,1940]" captionTargetBox= "[190,1399,697,1904]" captionTargetId= "figure-163@7.[189,1399,694,1904]" captionText= "Figs 22– 34. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28– 29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22– 26 — median lobe; 28– 34 — right paramere, scale as for respective median lobe; 22– 25, 27 — ventrobasal aspect; 29– 34 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 22– 34. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28– 29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22– 26 — среднЯЯ долЯ; 28– 34 — праваЯ парамера в масштабе соответствуюЩей средней доли; 22– 25, 27 — вентробаЗально; 29– 34 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi= "http://doi.org/10.5281/zenodo.10949106" httpUri= "https://zenodo.org/record/10949106/files/figure.png" pageId= "4" pageNumber= "20" > 23, 31</figureCitation>
): median lobe in lateral view geniculate, apex in ventral view lamellate, triangular. Right paramere oval.
</paragraph>
<paragraph id= "A75736D5FFCF16697984FABD20957757" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
Abdominal urites VIII and IX, with reproductive tract, in female as in
<figureCitation id= "3FD32A50FFCF1669785CFAD820C57757" box= "[314,422,1346,1368]" captionStart= "Figs 52– 62" captionStartId= "13.[226,263,1873,1892]" captionTargetBox= "[193,1399,824,1838]" captionTargetId= "figure-208@13.[189,1399,820,1839]" captionTargetPageId= "13" captionText= "Figs 52– 62. Abdominal urites VIII– IX in female: 52– 53, 61 — Pterostichus austrinus sp.n.; 54– 55 — P. glabellus sp.n.; 56– 57 — P. unisetosus sp.n.; 58– 60 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 61– 62 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 52– 62. Уриты VIII– IX брюшка самки: 52– 53, 61 — Pterostichus austrinus sp.n.; 54– 55 — P. glabellus sp.n.; 56– 57 — P. unisetosus sp.n.; 58– 60 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 61– 62 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм." figureDoi= "http://doi.org/10.5281/zenodo.10949112" httpUri= "https://zenodo.org/record/10949112/files/figure.png" pageId= "4" pageNumber= "20" > Figs 56– 57</figureCitation>
and
<figureCitation id= "3FD32A50FFCF166978B1FAD820917757" box= "[471,498,1346,1368]" captionStart= "Figs 63– 68" captionStartId= "15.[226,263,1873,1892]" captionTargetBox= "[193,1392,817,1839]" captionTargetId= "figure-183@15.[188,1398,813,1843]" captionTargetPageId= "15" captionText= "Figs 63– 68. Reproductive tract in female: 63 — Pterostichus austrinus sp.n.; 64– 65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n.; 63, 65– 67 — ventral aspect; 64, 68 — dorsal aspect; bc — bursa copulatrix; bs — bulbous structure; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 63– 68. Репродуктивный тракт самки: 63 — Pterostichus austrinus sp.n.; 64– 65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n; 63, 65– 67 — вентрально; 64, 68 — дорЗально; bc — копулЯтивнаЯ сумка; bs — бульбарное обраЗование; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi= "http://doi.org/10.5281/zenodo.10949114" httpUri= "https://zenodo.org/record/10949114/files/figure.png" pageId= "4" pageNumber= "20" > 68</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id= "EFF2655EFFCF16697984FAC7238077CD" pageId= "4" pageNumber= "20" type= "distribution" >
<paragraph id= "A75736D5FFCF16697984FAC7234F77A8" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" >
DISTRIBUTION. Only known from the
<typeStatus id= "78538877FFCF16697BEFFAC723D0777D" box= "[649,691,1373,1394]" pageId= "4" pageNumber= "20" > type</typeStatus>
locality which is on the Hoang Lien [Son] mountain ridge
<quantity id= "60109B30FFCF16697BA0FAED22647782" box= "[710,775,1399,1421]" metricMagnitude= "4" metricUnit= "m" metricValue= "3.3" pageId= "4" pageNumber= "20" unit= "km" value= "33.0" > 33 km</quantity>
distant from that of
<taxonomicName id= "60E84D56FFCF166978E6FA09209777A8" authority= "Fedorenko, 2023" authorityName= "Fedorenko" authorityYear= "2023" box= "[384,500,1426,1448]" class= "Insecta" family= "Carabidae" genus= "Pterostichus" kingdom= "Animalia" order= "Coleoptera" pageId= "4" pageNumber= "20" phylum= "Arthropoda" rank= "species" species= "glabellus" status= "sp.n." >
<emphasis id= "959CEAC7FFCF166978E6FA09209777A8" box= "[384,500,1426,1448]" italics= "true" pageId= "4" pageNumber= "20" > P. glabellus</emphasis>
</taxonomicName>
<emphasis id= "959CEAC7FFCF1669789DFA08234F77A8" bold= "true" box= "[507,556,1426,1447]" pageId= "4" pageNumber= "20" >
<taxonomicNameLabel id= "8EAF57BCFFCF1669789DFA08234F77A8" box= "[507,556,1426,1447]" pageId= "4" pageNumber= "20" rank= "species" > sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id= "A75736D5FFCF16697984FA37238077CD" blockId= "4.[189,776,810,1528]" box= "[226,739,1453,1475]" pageId= "4" pageNumber= "20" > NAME. Refers to the elytron with single discal seta.</paragraph>
</subSubSection>
<subSubSection id= "EFF2655EFFCF16697984FA5D20AE77F7" pageId= "4" pageNumber= "20" type= "description" >
<paragraph id= "A75736D5FFCF16697984FA5D20AE77F7" blockId= "4.[189,776,810,1528]" pageId= "4" pageNumber= "20" > HABITATS AND HABITS. All the specimens were hand collected in a cloudy forest.</paragraph>
</subSubSection>
</treatment>
</document>