163 lines
23 KiB
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163 lines
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<document id="590AC7F71078EEF039DEC2878861C9B7" ID-DOI="10.15298/rusentj.32.1.03" ID-ISSN="0132-8069" ID-Zenodo-Dep="10949096" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198792598" checkinUser="felipe" docAuthor="Fedorenko, D. N." docDate="2023" docId="2F4187C3FFCF1669784FFCB020AE77F7" docLanguage="en" docName="RussEntomolJ.32.1.16-39.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.03" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="Pterostichus (Morphohaptoderus) unisetosus Fedorenko, 2023, sp.n." docType="treatment" docVersion="3" lastPageNumber="20" masterDocId="D378FFBBFFCB166D7966FF9A2163720F" masterDocTitle="New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam" masterLastPageNumber="39" masterPageNumber="16" pageNumber="20" updateTime="1712665435938" updateUser="ExternalLinkService">
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<mods:titleInfo id="1BD86332D3D69CCB1DA9B21A2383ED29">
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<mods:title id="CDD86520A35E9AA5C2978F38EF220DAC">New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam</mods:title>
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<mods:name id="0A8002EA84A5FC00A5C47ED8AE493403" type="personal">
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<mods:namePart id="190F8B1E41A4978F7A0CBF904AE663F7">Fedorenko, D. N.</mods:namePart>
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<mods:title id="EAE5DB09FB9DB48A456B1531F803C73B">Russian Entomological Journal</mods:title>
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</mods:titleInfo>
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<mods:part id="19AE7FB6B0A61D02163F03F454438A52">
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<mods:date id="3040C3D1A12927B0A25D6D97AC40081D">2023</mods:date>
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<mods:detail id="1ACD00B3A2B10CEDBF79410297256CE5" type="pubDate">
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<mods:number id="56221CE804252B8A992D3A69AF36B955">2023-03-31</mods:number>
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<mods:number id="7848ECDA9BF2713B5B0616ADCCE07204">32</mods:number>
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<mods:number id="4639BDA2B510168C27419DE3599F609F">1</mods:number>
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<mods:url id="9266306F91C44073A770485A4D0F7D0F">http://dx.doi.org/10.15298/rusentj.32.1.03</mods:url>
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<mods:classification id="674D8E81982B82A408B6509ECDCA004D">journal article</mods:classification>
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<mods:identifier id="05975B50D86411A4B9B4BAA3D9212B9C" type="DOI">10.15298/rusentj.32.1.03</mods:identifier>
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<mods:identifier id="514D728BC8BD2387362D5DBF98AD923D" type="ISSN">0132-8069</mods:identifier>
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<mods:identifier id="92579A383E3D9EC3503E070E070A1879" type="Zenodo-Dep">10949096</mods:identifier>
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</mods:mods>
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<treatment id="2F4187C3FFCF1669784FFCB020AE77F7" LSID="urn:lsid:plazi:treatment:2F4187C3FFCF1669784FFCB020AE77F7" httpUri="http://treatment.plazi.org/id/2F4187C3FFCF1669784FFCB020AE77F7" lastPageNumber="20" pageId="4" pageNumber="20">
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<subSubSection id="EFF2655EFFCF1669784FFCB0231B716F" pageId="4" pageNumber="20" type="nomenclature">
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<paragraph id="A75736D5FFCF1669784FFCB0231B716F" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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<heading id="FC1F81B9FFCF1669784FFCB0231B716F" centered="true" fontSize="10" level="2" pageId="4" pageNumber="20" reason="8">
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<taxonomicName id="60E84D56FFCF1669784FFCB020D97150" ID-CoL="8MBT6" authorityName="Fedorenko" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="unisetosus" subGenus="Morphohaptoderus">
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<emphasis id="959CEAC7FFCF1669784FFCB020CC714D" box="[297,431,810,834]" italics="true" pageId="4" pageNumber="20">Pterostichus</emphasis>
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(
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<emphasis id="959CEAC7FFCF166978A5FCB023F2714D" box="[451,657,810,834]" italics="true" pageId="4" pageNumber="20">Morphohaptoderus</emphasis>
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)
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<emphasis id="959CEAC7FFCF1669782DFCD220D97150" box="[331,442,840,863]" italics="true" pageId="4" pageNumber="20">unisetosus</emphasis>
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</taxonomicName>
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Fedorenko,
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<emphasis id="959CEAC7FFCF16697B25FCD2231B716F" bold="true" box="[579,632,840,864]" pageId="4" pageNumber="20">
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<taxonomicNameLabel id="8EAF57BCFFCF16697B25FCD2231B716F" box="[579,632,840,864]" pageId="4" pageNumber="20" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697848FCFF23F67172" box="[302,661,869,893]" pageId="4" pageNumber="20" type="description">
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<paragraph id="A75736D5FFCF16697848FCFF23F67172" blockId="4.[189,776,810,1528]" box="[302,661,869,893]" pageId="4" pageNumber="20">
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<figureCitation id="3FD32A50FFCF16697848FCFF20107172" box="[302,371,869,893]" captionStart="Figs 1–5" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[198,1388,321,1922]" captionTargetId="figure-102@2.[188,1398,317,1927]" captionTargetPageId="2" captionText="Figs 1–5. Dorsal habitus: 1 — Pterostichus glabellus sp.n., paratype ♀; 2 — P. unisetosus sp.n., holotype; 3 — P. primitivus sp.n., holotype; 4 — P. subirideus sp.n., paratype ♂; 5 — P. batdaiensis sp.n., paratype ♂ from Sa Pa env. Рис. 1–5. Габитус дорЗально: 1 — Pterostichus glabellus sp.n., паратип ♀; 2 — P. unisetosus sp.n., голотип; 3 — P. primitivus sp.n., голотип; P. subirideus sp.n., паратип ♂; 5 — P. batdaiensis sp.n., паратип ♂ иЗ окрестностей Sa Pa." figureDoi="http://doi.org/10.5281/zenodo.10949098" httpUri="https://zenodo.org/record/10949098/files/figure.png" pageId="4" pageNumber="20">Figs 2</figureCitation>
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,
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<figureCitation id="3FD32A50FFCF16697819FCFF20A77172" box="[383,452,869,893]" captionStart="Figs 12–21" captionStartId="5.[226,263,1945,1964]" captionTargetBox="[189,1399,319,1919]" captionTargetId="figure-122@5.[189,1399,314,1923]" captionTargetPageId="5" captionText="Figs 12–21. Aedeagus median lobe: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm. Рис. 12–21. СреднЯЯ долЯ Эдеагуса: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949104" httpUri="https://zenodo.org/record/10949104/files/figure.png" pageId="4" pageNumber="20">14–15</figureCitation>
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,
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<figureCitation id="3FD32A50FFCF166978B6FCFF23757172" box="[464,534,869,893]" captionStart="Figs 22–34" captionStartId="7.[226,263,1921,1940]" captionTargetBox="[190,1399,697,1904]" captionTargetId="figure-163@7.[189,1399,694,1904]" captionText="Figs 22–34. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — median lobe; 28–34 — right paramere, scale as for respective median lobe; 22–25, 27 — ventrobasal aspect; 29–34 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 22–34. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — среднЯЯ долЯ; 28–34 — праваЯ парамера в масштабе соответствуюЩей средней доли; 22–25, 27 — вентробаЗально; 29–34 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949106" httpUri="https://zenodo.org/record/10949106/files/figure.png" pageId="4" pageNumber="20">23, 31</figureCitation>
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, 56–57, 68.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697984FC10232A71F2" pageId="4" pageNumber="20" type="materials_examined">
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<paragraph id="A75736D5FFCF16697984FC1023F871EA" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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<materialsCitation id="17803C88FFCF16697984FC1023F871EA" collectingDate="2018-10" collectionCode="ZMMU, SIEE" collectorName="Y Ty & Bat Xat" elevation="1750" location="Lao Cai" pageId="4" pageNumber="20" specimenCount="5" specimenCount-female="1" specimenCount-male="2" stateProvince="Lao Cai" typeStatus="holotype">
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MATERIAL.
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<typeStatus id="78538877FFCF16697838FC1020CD7192" box="[350,430,906,925]" pageId="4" pageNumber="20" type="holotype">Holotype</typeStatus>
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<specimenCount id="B1EEFD5CFFCF166978D0FC1D20A97192" box="[438,458,903,925]" count="1" pageId="4" pageNumber="20" type="male">♂</specimenCount>
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(
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<collectionCode id="C1F9AE10FFCF166978BEFC10237D7192" box="[472,542,906,925]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="20" type="Museum">ZMMU</collectionCode>
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) and
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<specimenCount id="B1EEFD5CFFCF16697B37FC1023AD7192" box="[593,718,906,925]" count="2" pageId="4" pageNumber="20" type="generic" typeStatus="paratype">two paratypes</specimenCount>
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,
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<specimenCount id="B1EEFD5CFFCF16697BBFFC1D238E7192" box="[729,749,903,925]" count="1" pageId="4" pageNumber="20" type="male">♂</specimenCount>
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,
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<specimenCount id="B1EEFD5CFFCF16697B9DFC1D22647192" box="[763,775,903,925]" count="1" pageId="4" pageNumber="20" type="female">♀</specimenCount>
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(
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<collectionCode id="C1F9AE10FFCF166979A5FC38219071BA" box="[195,243,930,949]" pageId="4" pageNumber="20">SIEE</collectionCode>
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) with label: ‘N-Vietnam,
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<quantity id="60109B30FFCF166978ABFC38236171BA" box="[461,514,930,949]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="20" unit="km" value="40.0">40 km</quantity>
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WNW of/
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<collectingRegion id="652CF837FFCF16697B39FC3823C671BA" box="[607,677,930,949]" country="Vietnam" name="Lao Cai" pageId="4" pageNumber="20">Lao Cai</collectingRegion>
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, env.
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<collectorName id="0A1D5303FFCF16697BB3FC38226071BA" box="[725,771,930,949]" pageId="4" pageNumber="20">Y Ty</collectorName>
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,
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<collectorName id="0A1D5303FFCF166979DBFC20206471C2" box="[189,263,954,973]" pageId="4" pageNumber="20">Bat Xat</collectorName>
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/ Nat[io]n.[al] Park, h=
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<quantity id="60109B30FFCF1669788DFC20230A71C2" box="[491,617,954,973]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="4" pageNumber="20" unit="m" value="1750.0" valueMax="1900.0" valueMin="1600.0">
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<elevation id="2CC5D1E6FFCF1669788DFC20230A71C2" box="[491,617,954,973]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="4" pageNumber="20" unit="m" value="1750.0" valueMax="1900.0" valueMin="1600.0">1600-1900 m</elevation>
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</quantity>
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/ 22°37´-37.6´N/ 103°37.3´-38.3´E, 16-21./ leg. D.
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<taxonomicName id="60E84D56FFCF166978B9FC48235C71EA" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[479,575,978,997]" pageId="4" pageNumber="26" rank="subGenus" status="subgen. nov." subGenus="Fedorenko">Fedorenko</taxonomicName>
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<date id="D3561015FFCF16697B2BFC4823F071EA" box="[589,659,978,997]" pageId="4" pageNumber="20" value="2018-10">
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<collectingDate id="C312E9FDFFCF16697B2BFC4823F071EA" box="[589,659,978,997]" pageId="4" pageNumber="20" value="2018-10">X.2018</collectingDate>
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</date>
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’.
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</materialsCitation>
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</paragraph>
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<paragraph id="A75736D5FFCF16697984FC70232A71F2" blockId="4.[189,776,810,1528]" box="[226,585,1002,1021]" pageId="4" pageNumber="20">Genitalia examined in all the specimens.</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697984FB9823DB7642" pageId="4" pageNumber="20" type="diagnosis">
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<paragraph id="A75736D5FFCF16697984FB9823DB7642" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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DIAGNOSIS. No significant differences from
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<taxonomicName id="60E84D56FFCF16697BC1FB9921BA763D" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="glabellus" status="sp.n.">
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<emphasis id="959CEAC7FFCF16697BC1FB9921BA763D" italics="true" pageId="4" pageNumber="20">P. glabellus</emphasis>
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</taxonomicName>
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<emphasis id="959CEAC7FFCF166979BBFB87206D763D" bold="true" box="[221,270,1053,1074]" pageId="4" pageNumber="20">
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<taxonomicNameLabel id="8EAF57BCFFCF166979BBFB87206D763D" box="[221,270,1053,1074]" pageId="4" pageNumber="20" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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other than the elytron with middle discal seta, d2,
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<emphasis id="959CEAC7FFCF16697B8FFB872399763D" box="[745,762,1053,1074]" italics="true" pageId="4" pageNumber="20">fe</emphasis>
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3 not sexually dimorphic, and aedeagus is distinctive.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697984FBC820957757" pageId="4" pageNumber="20" type="description">
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<paragraph id="A75736D5FFCF16697984FBC820FE76DD" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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DESCRIPTION. Body as in
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<figureCitation id="3FD32A50FFCF16697B68FBC8232E7668" box="[526,589,1106,1128]" captionStart="Figs 1–5" captionStartId="2.[226,263,1969,1988]" captionTargetBox="[198,1388,321,1922]" captionTargetId="figure-102@2.[188,1398,317,1927]" captionTargetPageId="2" captionText="Figs 1–5. Dorsal habitus: 1 — Pterostichus glabellus sp.n., paratype ♀; 2 — P. unisetosus sp.n., holotype; 3 — P. primitivus sp.n., holotype; 4 — P. subirideus sp.n., paratype ♂; 5 — P. batdaiensis sp.n., paratype ♂ from Sa Pa env. Рис. 1–5. Габитус дорЗально: 1 — Pterostichus glabellus sp.n., паратип ♀; 2 — P. unisetosus sp.n., голотип; 3 — P. primitivus sp.n., голотип; P. subirideus sp.n., паратип ♂; 5 — P. batdaiensis sp.n., паратип ♂ иЗ окрестностей Sa Pa." figureDoi="http://doi.org/10.5281/zenodo.10949098" httpUri="https://zenodo.org/record/10949098/files/figure.png" pageId="4" pageNumber="20">Fig. 2</figureCitation>
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. BL 8.4–8.8 mm. Pronotal base nearly straight, two fifths wider than apex. Elytral stria 7 inside humeral angle; parascutellar striole entire, stria 1 adjoining basal ridge. D2/EL 0.50–0.57 (0.55, n = 3×2). USS: 13–15.
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</paragraph>
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<paragraph id="A75736D5FFCF16697984FB4D20E8772C" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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Aedeagus (
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<figureCitation id="3FD32A50FFCF16697834FB4D20A676E2" box="[338,453,1239,1261]" captionStart="Figs 12–21" captionStartId="5.[226,263,1945,1964]" captionTargetBox="[189,1399,319,1919]" captionTargetId="figure-122@5.[189,1399,314,1923]" captionTargetPageId="5" captionText="Figs 12–21. Aedeagus median lobe: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — left lateral aspect; 13, 15, 17, 19, 21 — right lateral aspect. Scale bars 1 mm. Рис. 12–21. СреднЯЯ долЯ Эдеагуса: 12–13 — Pterostichus glabellus sp.n.; 14–15 — P. unisetosus sp.n.; 16–17 — P. tonkinensis; 18– 19 — P. austrinus sp.n.; 20–21 — P. subirideus sp.n.; 12, 14, 16, 18, 20 — слева; 13, 15, 17, 19, 21 — справа. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949104" httpUri="https://zenodo.org/record/10949104/files/figure.png" pageId="4" pageNumber="20">Figs 14–15</figureCitation>
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,
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<figureCitation id="3FD32A50FFCF166978B4FB4D237676E2" box="[466,533,1239,1261]" captionStart="Figs 22–34" captionStartId="7.[226,263,1921,1940]" captionTargetBox="[190,1399,697,1904]" captionTargetId="figure-163@7.[189,1399,694,1904]" captionText="Figs 22–34. Aedeagus: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — median lobe; 28–34 — right paramere, scale as for respective median lobe; 22–25, 27 — ventrobasal aspect; 29–34 — left lateral aspect; 26, 28 — dorsal aspect. Scale bars 1 mm. Рис. 22–34. Эдеагус: 22, 30 — Pterostichus glabellus sp.n.; 23, 31 — P. unisetosus sp.n.; 24, 32 — P. tonkinensis; 25, 28–29 — P. austrinus sp.n.; 26, 33 — P. subirideus sp.n.; 27, 34 — P. laevibasis sp.n.; 22–26 — среднЯЯ долЯ; 28–34 — праваЯ парамера в масштабе соответствуюЩей средней доли; 22–25, 27 — вентробаЗально; 29–34 — слева; 26, 28 — сверху. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949106" httpUri="https://zenodo.org/record/10949106/files/figure.png" pageId="4" pageNumber="20">23, 31</figureCitation>
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): median lobe in lateral view geniculate, apex in ventral view lamellate, triangular. Right paramere oval.
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</paragraph>
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<paragraph id="A75736D5FFCF16697984FABD20957757" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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Abdominal urites VIII and IX, with reproductive tract, in female as in
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<figureCitation id="3FD32A50FFCF1669785CFAD820C57757" box="[314,422,1346,1368]" captionStart="Figs 52–62" captionStartId="13.[226,263,1873,1892]" captionTargetBox="[193,1399,824,1838]" captionTargetId="figure-208@13.[189,1399,820,1839]" captionTargetPageId="13" captionText="Figs 52–62. Abdominal urites VIII–IX in female: 52–53, 61 — Pterostichus austrinus sp.n.; 54–55 — P. glabellus sp.n.; 56–57 — P. unisetosus sp.n.; 58–60 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 61–62 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 52–62. Уриты VIII–IX брюшка самки: 52–53, 61 — Pterostichus austrinus sp.n.; 54–55 — P. glabellus sp.n.; 56–57 — P. unisetosus sp.n.; 58–60 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 61–62 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949112" httpUri="https://zenodo.org/record/10949112/files/figure.png" pageId="4" pageNumber="20">Figs 56–57</figureCitation>
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and
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<figureCitation id="3FD32A50FFCF166978B1FAD820917757" box="[471,498,1346,1368]" captionStart="Figs 63–68" captionStartId="15.[226,263,1873,1892]" captionTargetBox="[193,1392,817,1839]" captionTargetId="figure-183@15.[188,1398,813,1843]" captionTargetPageId="15" captionText="Figs 63–68. Reproductive tract in female: 63 — Pterostichus austrinus sp.n.; 64–65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n.; 63, 65–67 — ventral aspect; 64, 68 — dorsal aspect; bc — bursa copulatrix; bs — bulbous structure; dv — diverticulum; ov — common oviduct; rp — receptacle; sc — seminal canal; sg — spermathecal gland; sgd — spermathecal gland duct; sp — spermatheca. Scale bars 1 mm. Рис. 63–68. Репродуктивный тракт самки: 63 — Pterostichus austrinus sp.n.; 64–65 — P. glabellus sp.n.; 66 — P. strenuus (Panzer, 1796); 67 — P. chameleon; 68 — P. unisetosus sp.n; 63, 65–67 — вентрально; 64, 68 — дорЗально; bc — копулЯтивнаЯ сумка; bs — бульбарное обраЗование; dv — дивертикул; ov — непарный Яйцевод; rp — семеприемник; sc — семепровод; sg — ЖелеЗа сперматеки; sgd — проток ЖелеЗы сперматеки; sp — сперматека. Масштаб 1 мм." figureDoi="http://doi.org/10.5281/zenodo.10949114" httpUri="https://zenodo.org/record/10949114/files/figure.png" pageId="4" pageNumber="20">68</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697984FAC7238077CD" pageId="4" pageNumber="20" type="distribution">
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<paragraph id="A75736D5FFCF16697984FAC7234F77A8" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">
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DISTRIBUTION. Only known from the
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<typeStatus id="78538877FFCF16697BEFFAC723D0777D" box="[649,691,1373,1394]" pageId="4" pageNumber="20">type</typeStatus>
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locality which is on the Hoang Lien [Son] mountain ridge
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<quantity id="60109B30FFCF16697BA0FAED22647782" box="[710,775,1399,1421]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="20" unit="km" value="33.0">33 km</quantity>
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distant from that of
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<taxonomicName id="60E84D56FFCF166978E6FA09209777A8" authority="Fedorenko, 2023" authorityName="Fedorenko" authorityYear="2023" box="[384,500,1426,1448]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="glabellus" status="sp.n.">
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<emphasis id="959CEAC7FFCF166978E6FA09209777A8" box="[384,500,1426,1448]" italics="true" pageId="4" pageNumber="20">P. glabellus</emphasis>
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</taxonomicName>
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<emphasis id="959CEAC7FFCF1669789DFA08234F77A8" bold="true" box="[507,556,1426,1447]" pageId="4" pageNumber="20">
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<taxonomicNameLabel id="8EAF57BCFFCF1669789DFA08234F77A8" box="[507,556,1426,1447]" pageId="4" pageNumber="20" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="A75736D5FFCF16697984FA37238077CD" blockId="4.[189,776,810,1528]" box="[226,739,1453,1475]" pageId="4" pageNumber="20">NAME. Refers to the elytron with single discal seta.</paragraph>
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</subSubSection>
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<subSubSection id="EFF2655EFFCF16697984FA5D20AE77F7" pageId="4" pageNumber="20" type="description">
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<paragraph id="A75736D5FFCF16697984FA5D20AE77F7" blockId="4.[189,776,810,1528]" pageId="4" pageNumber="20">HABITATS AND HABITS. All the specimens were hand collected in a cloudy forest.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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