<documentid="3DD5B7BABC2FFC375F5B19DA6B4A6441"ID-CLB-Dataset="299696"ID-DOI="10.5281/zenodo.11512952"ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c"ID-ISSN="0068-547X"ID-Zenodo-Dep="11512952"IM.bibliography_approvedBy="julia"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="julia"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="julia"IM.treatmentCitations_approvedBy="julia"IM.treatments_approvedBy="julia"checkinTime="1717721037103"checkinUser="felipe"docAuthor="Kavanaugh, David H. & Cueva-Dabkoski, Mollie"docDate="2023"docId="1C1187FCEB7EED5DFF9EFA70602582C3"docLanguage="en"docName="ProcCalifAcadSci.67.21.493-537.pdf"docOrigin="Proceedings of the California Academy of Sciences 67 (21)"docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article"docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B"docStyleName="ProcCalifAcadSci.2015-.journal_article"docStyleVersion="2"docTitle="Cyclosomus philippinus Heller"docType="treatment"docVersion="5"lastPageNumber="517"masterDocId="E028FF84EB68ED45FFE5FFAF6327867B"masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia."masterLastPageNumber="537"masterPageNumber="493"pageNumber="515"updateTime="1720721490288"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="940AAEB21E87DD3F89B920B3CDF2EF12">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
<mods:namePartid="1B8A259D8AAE64EB5AB83765D1B20C4D">Kavanaugh, David H.</mods:namePart>
<mods:affiliationid="B1429D40FB6E3C9DD0B7E676302431F7">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
<mods:affiliationid="D27669CDB0147972A8A980ACCB5CD133">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
<figureCitationid="0C832A6FEB7EED53FF9EFA5063C4806E"box="[123,227,1535,1557]"captionStart="FIGURE"captionStartId="23.[171,183,1573,1592]"captionTargetBox="[156,1139,216,1547]"captionTargetId="figure-14@23.[156,1139,216,1560]"captionTargetPageId="23"captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512978"httpUri="https://zenodo.org/record/11512978/files/figure.png"pageId="22"pageNumber="515">Figures 10</figureCitation>
,
<figureCitationid="0C832A6FEB7EED53FF0BFA506234806E"box="[238,275,1535,1557]"captionStart="FIGURE"captionStartId="33.[171,183,1573,1592]"captionTargetBox="[139,1158,216,1560]"captionTargetId="figure-14@33.[135,1162,216,1560]"captionTargetPageId="33"captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512988"httpUri="https://zenodo.org/record/11512988/files/figure.png"pageId="22"pageNumber="515">14F</figureCitation>
,
<figureCitationid="0C832A6FEB7EED53FEFBFA506260806E"box="[286,327,1535,1557]"captionStart="FIGURE"captionStartId="35.[171,183,1494,1513]"captionTargetBox="[120,1163,216,1464]"captionTargetId="figure-14@35.[120,1176,216,1479]"captionTargetPageId="35"captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512992"httpUri="https://zenodo.org/record/11512992/files/figure.png"pageId="22"pageNumber="515">16E</figureCitation>
,
<figureCitationid="0C832A6FEB7EED53FEB4FA50625D806E"box="[337,378,1535,1557]"captionStart="FIGURE"captionStartId="36.[171,183,1563,1582]"captionTargetBox="[120,1164,216,1526]"captionTargetId="figure-9@36.[120,1176,216,1548]"captionTargetPageId="36"captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512994"httpUri="https://zenodo.org/record/11512994/files/figure.png"pageId="22"pageNumber="515">17E</figureCitation>
,
<figureCitationid="0C832A6FEB7EED53FE61FA5062BB806E"box="[388,412,1535,1557]"captionStart="FIGURE"captionStartId="39.[171,183,1593,1612]"captionTargetBox="[168,1128,979,1582]"captionTargetId="figure-84@39.[168,1128,979,1582]"captionTargetPageId="39"captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km."figureDoi="http://doi.org/10.5281/zenodo.11513002"httpUri="https://zenodo.org/record/11513002/files/figure.png"pageId="22"pageNumber="515">21</figureCitation>
<treatmentCitationid="151910FBEB7EED53FF9EF98B612B8041"author="HELLER, K. M."box="[123,524,1572,1594]"page="302"pageId="22"pageNumber="515"year="1923">
<bibRefCitationid="F0294B1BEB7EED53FE82F98B612B8041"author="HELLER, K. M."box="[359,524,1572,1594]"pageId="22"pageNumber="515"pagination="295 - 307"refId="ref23510"refString="HELLER, K. M. 1923. Some new Malayan Carabidae, especially Philippine. The Philippine Journal of Science 23: 295 - 307 + 1 pl."type="journal article"year="1923">Heller, 1923:302</bibRefCitation>
<treatmentCitationid="151910FBEB7EED53FD72F9EE604A802C"author="ANDREWES, H. E."box="[663,877,1601,1623]"page="349"pageId="22"pageNumber="515"year="1926">
<bibRefCitationid="F0294B1BEB7EED53FD72F9EE604A802C"author="ANDREWES, H. E."box="[663,877,1601,1623]"pageId="22"pageNumber="515"pagination="345 - 361"refId="ref23053"refString="ANDREWES, H. E. 1926. A catalogue of Philippine Carabidae. Philippine Journal of Science 31: 345 - 361."type="journal article"year="1926">Andrewes (1926:349)</bibRefCitation>
<bibRefCitationid="F0294B1BEB7EED53FC92F9EE670D802C"author="CSIKI, E."box="[887,1066,1601,1623]"pageId="22"pageNumber="515"pagination="1279 - 1598"refId="ref23258"refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598."type="book chapter"year="1932">Csiki (1932:1295)</bibRefCitation>
in Asia by the following combination of character states: Body size medium for genus, BL males =
<quantityid="53409B0FEB7EED53FF3FF90A625E80C6"box="[218,377,1701,1725]"metricMagnitude="-3"metricUnit="m"metricValue="7.4"metricValueMax="7.6"metricValueMin="7.2"pageId="22"pageNumber="515"unit="mm"value="7.4"valueMax="7.6"valueMin="7.2">7.2 to 7.6 mm</quantity>
<figureCitationid="0C832A6FEB7EED53FCCCF90A60AF80C6"box="[809,904,1701,1725]"captionStart="FIGURE"captionStartId="33.[171,183,1573,1592]"captionTargetBox="[139,1158,216,1560]"captionTargetId="figure-14@33.[135,1162,216,1560]"captionTargetPageId="33"captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512988"httpUri="https://zenodo.org/record/11512988/files/figure.png"pageId="22"pageNumber="515">Fig. 14F</figureCitation>
) relatively narrow (ratio PWM/PL = 2.06 to 2.33) and less broadened basally (ratio PWM/PWA = 1.45 to 1.59), with disc piceous to black and lateral pale bands wide and well-defined, anterior angles broadly rounded, lateral margins not sinuate near anterior angles; free apex of prosternal intercoxal process long (as in
<figureCitationid="0C832A6FEB70ED5DFF9EFE9463FB8728"box="[123,220,315,339]"captionStart="FIGURE"captionStartId="34.[171,183,869,888]"captionTargetBox="[140,1176,216,846]"captionTargetId="figure-9@34.[120,1176,216,846]"captionTargetPageId="34"captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512990"httpUri="https://zenodo.org/record/11512990/files/figure.png"pageId="24"pageNumber="517">Fig. 15C</figureCitation>
); elytral dark pattern varied (
<figureCitationid="0C832A6FEB70ED5DFDFDFE9461828728"box="[536,677,315,339]"captionStart="FIGURE"captionStartId="23.[171,183,1573,1592]"captionTargetBox="[156,1139,216,1547]"captionTargetId="figure-14@23.[156,1139,216,1560]"captionTargetPageId="23"captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512978"httpUri="https://zenodo.org/record/11512978/files/figure.png"pageId="24"pageNumber="517">Figs. 10A, C</figureCitation>
) [slight blue cast to the dark elements seen in
<figureCitationid="0C832A6FEB70ED5DFF9EFEF463FD8708"box="[123,218,347,371]"captionStart="FIGURE"captionStartId="23.[171,183,1573,1592]"captionTargetBox="[156,1139,216,1547]"captionTargetId="figure-14@23.[156,1139,216,1560]"captionTargetPageId="23"captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512978"httpUri="https://zenodo.org/record/11512978/files/figure.png"pageId="24"pageNumber="517">Fig. 10A</figureCitation>
is an imaging artifact], with basal and middle bands of average to greater than average thickness for the genus, preapical dark spot absent; elytral epipleura with long setae over entire length; median lobe of male genitalia with shaft arcuate basally and nearly straight subapically, tapered to a long and pointed apical lamella in lateral aspect (
<figureCitationid="0C832A6FEB70ED5DFCC6FE1360A187AF"box="[803,902,444,468]"captionStart="FIGURE"captionStartId="35.[171,183,1494,1513]"captionTargetBox="[120,1163,216,1464]"captionTargetId="figure-14@35.[120,1176,216,1479]"captionTargetPageId="35"captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512992"httpUri="https://zenodo.org/record/11512992/files/figure.png"pageId="24"pageNumber="517">Fig. 16E</figureCitation>
), shaft moderately broad basally, gradually tapered to moderately broad, apically blunted, and sublanceolate apical lamella in dorsal aspect (
<figureCitationid="0C832A6FEB70ED5DFED4FE5362B5846F"box="[305,402,508,532]"captionStart="FIGURE"captionStartId="36.[171,183,1563,1582]"captionTargetBox="[120,1164,216,1526]"captionTargetId="figure-9@36.[120,1176,216,1548]"captionTargetPageId="36"captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512994"httpUri="https://zenodo.org/record/11512994/files/figure.png"pageId="24"pageNumber="517">Fig. 17E</figureCitation>
<figureCitationid="0C832A6FEB70ED5DFCF8FE536049846F"box="[797,878,508,532]"captionStart="FIGURE"captionStartId="39.[171,183,1593,1612]"captionTargetBox="[168,1128,979,1582]"captionTargetId="figure-84@39.[168,1128,979,1582]"captionTargetPageId="39"captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km."figureDoi="http://doi.org/10.5281/zenodo.11513002"httpUri="https://zenodo.org/record/11513002/files/figure.png"pageId="24"pageNumber="517">Fig. 21</figureCitation>
Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm.
) of a preapical dark spot. Males of the two species also differ distinctly in the shape and curvature of the median lobe in lateral aspect (compare
<figureCitationid="0C832A6FEB70ED5DFC1AFDD2677984EE"box="[1023,1118,637,661]"captionStart="FIGURE"captionStartId="35.[171,183,1494,1513]"captionTargetBox="[120,1163,216,1464]"captionTargetId="figure-14@35.[120,1176,216,1479]"captionTargetPageId="35"captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512992"httpUri="https://zenodo.org/record/11512992/files/figure.png"pageId="24"pageNumber="517">Fig. 16B</figureCitation>
with
<figureCitationid="0C832A6FEB70ED5DFF9EFD3263FF84CE"box="[123,216,669,693]"captionStart="FIGURE"captionStartId="35.[171,183,1494,1513]"captionTargetBox="[120,1163,216,1464]"captionTargetId="figure-14@35.[120,1176,216,1479]"captionTargetPageId="35"captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512992"httpUri="https://zenodo.org/record/11512992/files/figure.png"pageId="24"pageNumber="517">Fig. 16E</figureCitation>
) and shape of the apical lamella in dorsal aspect (compare
<figureCitationid="0C832A6FEB70ED5DFCB3FD32609384CE"box="[854,948,669,693]"captionStart="FIGURE"captionStartId="36.[171,183,1563,1582]"captionTargetBox="[120,1164,216,1526]"captionTargetId="figure-9@36.[120,1176,216,1548]"captionTargetPageId="36"captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512994"httpUri="https://zenodo.org/record/11512994/files/figure.png"pageId="24"pageNumber="517">Fig. 17B</figureCitation>
with
<figureCitationid="0C832A6FEB70ED5DFC17FD32677584CE"box="[1010,1106,669,693]"captionStart="FIGURE"captionStartId="36.[171,183,1563,1582]"captionTargetBox="[120,1164,216,1526]"captionTargetId="figure-9@36.[120,1176,216,1548]"captionTargetPageId="36"captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm."figureDoi="http://doi.org/10.5281/zenodo.11512994"httpUri="https://zenodo.org/record/11512994/files/figure.png"pageId="24"pageNumber="517">Fig. 17E</figureCitation>
Unknown, but presumed to be restricted to the sandy shores of medium to large rivers at low elevation, like members of most other species of the genus.
<collectingDateid="F042E9C2EB70ED5DFCA3FC916701852D"box="[838,1062,830,854]"pageId="24"pageNumber="517"value="1998-11-05"valueMax="1998-11-07"valueMin="1998-11-05">5-7 November 1998</collectingDate>
<collectionCodeid="F2A9AE2FEB70ED5DFD99FCF161E8850D"box="[636,719,862,886]"country="Italy"httpUri="http://grbio.org/cool/irpn-c0nq"name="Museo Friulano di Storia Naturale"pageId="24"pageNumber="517">MFNB</collectionCode>
<materialsCitationid="24D03CB7EB70ED5DFD0DFCF1605985EC"ID-GBIF-Occurrence="4905152320"collectingDate="1998-11-05"collectingDateMax="1998-11-07"collectingDateMin="1998-11-05"collectionCode="SMTD, MFNB"collectorName=" C. S. Banks"country="Philippines"county="Luzon Island"elevation="150"location="Mountains"municipality="Los Banos"pageId="24"pageNumber="517"specimenCount="6"specimenCount-female="2"specimenCount-male="4"stateProvince="Zambales">
<collectionCodeid="F2A9AE2FEB70ED5DFCF8FCD0605785EC"box="[797,880,895,919]"country="Italy"httpUri="http://grbio.org/cool/irpn-c0nq"name="Museo Friulano di Storia Naturale"pageId="24"pageNumber="517">MFNB</collectionCode>
<figureCitationid="0C832A6FEB70ED5DFCD6FC3060A485CC"box="[819,899,927,951]"captionStart="FIGURE"captionStartId="39.[171,183,1593,1612]"captionTargetBox="[168,1128,979,1582]"captionTargetId="figure-84@39.[168,1128,979,1582]"captionTargetPageId="39"captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km."figureDoi="http://doi.org/10.5281/zenodo.11513002"httpUri="https://zenodo.org/record/11513002/files/figure.png"pageId="24"pageNumber="517">Fig. 21</figureCitation>
The range of variation in the thickness of the middle dark band of the elytra seen among specimens examined is shown in
<figureCitationid="0C832A6FEB70ED5DFD2FFC7060B4858C"box="[714,915,991,1015]"captionStart="FIGURE"captionStartId="23.[171,183,1573,1592]"captionTargetBox="[156,1139,216,1547]"captionTargetId="figure-14@23.[156,1139,216,1560]"captionTargetPageId="23"captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.11512978"httpUri="https://zenodo.org/record/11512978/files/figure.png"pageId="24"pageNumber="517">Figs. 10A and 10C</figureCitation>