treatments-xml/data/1C/11/87/1C1187FCEB7EED5DFF9EFA70602582C3.xml

294 lines
44 KiB
XML
Raw Normal View History

<document id="3DD5B7BABC2FFC375F5B19DA6B4A6441" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB7EED5DFF9EFA70602582C3" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus philippinus Heller" docType="treatment" docVersion="5" lastPageNumber="517" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="515" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="938B2F29FF9473CD0813512F1FF91F58" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C573A32F8282931C2A06F6F4B5E19346">
<mods:title id="940AAEB21E87DD3F89B920B3CDF2EF12">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
2024-06-21 12:30:48 +02:00
</mods:titleInfo>
<mods:name id="C3A2E16C72953B9316319AF35B929E4E" type="personal">
<mods:role id="B469A641D1529AE4DCF79F78298D3F5B">
<mods:roleTerm id="FCC05AA38E5E29A24C9166516B1564DD">Author</mods:roleTerm>
2024-06-21 12:30:48 +02:00
</mods:role>
<mods:namePart id="1B8A259D8AAE64EB5AB83765D1B20C4D">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="B1429D40FB6E3C9DD0B7E676302431F7">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
2024-06-21 12:30:48 +02:00
</mods:name>
<mods:name id="EFF891A4AA10B8BD15BC2FC8E979F0E1" type="personal">
<mods:role id="1B8E64AC5773B4F04C073FD54AEC2B64">
<mods:roleTerm id="04CE2E457F4978D17B3DCCD77653CF6A">Author</mods:roleTerm>
2024-06-21 12:30:48 +02:00
</mods:role>
<mods:namePart id="63318CCD742F2E4ECF4C8D02B22367A4">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="D27669CDB0147972A8A980ACCB5CD133">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
2024-06-21 12:30:48 +02:00
</mods:name>
<mods:typeOfResource id="02622B4F3FE6BF3C57D87BD6B4DC9B7F">text</mods:typeOfResource>
<mods:relatedItem id="583BC26664A1FB5DE37396C3B6E9CE49" type="host">
<mods:titleInfo id="DAE3C7AEDF3B5F0D2540B19051069038">
<mods:title id="C3A95EEBE007554BDFF1030467426E5C">Proceedings of the California Academy of Sciences</mods:title>
2024-06-21 12:30:48 +02:00
</mods:titleInfo>
<mods:part id="7728EB023D84F3352CA6BEFE73ACD36D">
<mods:date id="FACCB3F469B94D58C4543D483B9893CF">2023</mods:date>
<mods:detail id="34E180969892A50C3698ADA6F7DA518F" type="pubDate">
<mods:number id="6638CE2FACE0E35814AF526CF62AE1E8">2023-02-15</mods:number>
2024-06-21 12:30:48 +02:00
</mods:detail>
<mods:detail id="55CBC03BCCEA60E974846EFA7945CC0A" type="volume">
<mods:number id="97EB1AF3F2B68D07F51993DE049E6F24">67</mods:number>
2024-06-21 12:30:48 +02:00
</mods:detail>
<mods:detail id="33E09B6267789FAD9AC1B6864F955C04" type="issue">
<mods:number id="AC05C821DCCD251F6EDD8EB5D391F4AE">21</mods:number>
2024-06-21 12:30:48 +02:00
</mods:detail>
<mods:extent id="15D0E600C1AFDBA7CC29082E477972CF" unit="page">
<mods:start id="804A1443844B0E54A5BE334A625B0BF9">493</mods:start>
<mods:end id="60DCB42BAD2151E3E44C0363EE5808F0">537</mods:end>
2024-06-21 12:30:48 +02:00
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1F8827F1231E4B332A5256548C014B87">journal article</mods:classification>
<mods:identifier id="24F2A580738DF0DEED30346885CDECD7" type="CLB-Dataset">299696</mods:identifier>
<mods:identifier id="FDBE5178AC1F8BE7D671F48CBA6253C6" type="DOI">10.5281/zenodo.11512952</mods:identifier>
<mods:identifier id="40321755305A9595CDAAF492F3BF622F" type="GBIF-Dataset">2b56e3b0-2274-4963-b243-9c991b07072c</mods:identifier>
<mods:identifier id="E229BBC9F72F72DDE0B7E1D2F8931D1B" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="96B7769EC88FF8037D4C2290444482DE" type="Zenodo-Dep">11512952</mods:identifier>
2024-06-21 12:30:48 +02:00
</mods:mods>
<treatment id="1C1187FCEB7EED5DFF9EFA70602582C3" ID-DOI="http://doi.org/10.5281/zenodo.11513122" ID-GBIF-Taxon="235039984" ID-Zenodo-Dep="11513122" LSID="urn:lsid:plazi:treatment:1C1187FCEB7EED5DFF9EFA70602582C3" httpUri="http://treatment.plazi.org/id/1C1187FCEB7EED5DFF9EFA70602582C3" lastPageId="24" lastPageNumber="517" pageId="22" pageNumber="515">
2024-06-21 12:30:48 +02:00
<subSubSection id="DCA26561EB7EED53FF9EFA7062F6838D" box="[123,465,1503,1527]" pageId="22" pageNumber="515" type="nomenclature">
<paragraph id="940736EAEB7EED53FF9EFA7062F6838D" blockId="22.[123,1174,1503,1725]" box="[123,465,1503,1527]" pageId="22" pageNumber="515">
<heading id="CF4F8186EB7EED53FF9EFA7062F6838D" box="[123,465,1503,1527]" fontSize="10" level="2" pageId="22" pageNumber="515" reason="4">
<taxonomicName id="53B84D69EB7EED53FF9EFA7062F6838D" ID-CoL="6C5XG" authority="Heller" authorityName="Heller" box="[123,465,1503,1527]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="515" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB7EED53FF9EFA7062F6838D" bold="true" box="[123,465,1503,1527]" pageId="22" pageNumber="515">
<emphasis id="A6CCEAF8EB7EED53FF9EFA7062A4838C" bold="true" box="[123,387,1503,1527]" italics="true" pageId="22" pageNumber="515">Cyclosomus philippinus</emphasis>
Heller
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB7EED53FF9EFA5062BB806E" box="[123,412,1535,1557]" pageId="22" pageNumber="515" type="description">
<paragraph id="940736EAEB7EED53FF9EFA5062BB806E" blockId="22.[123,1174,1503,1725]" box="[123,412,1535,1557]" pageId="22" pageNumber="515">
<figureCitation id="0C832A6FEB7EED53FF9EFA5063C4806E" box="[123,227,1535,1557]" captionStart="FIGURE" captionStartId="23.[171,183,1573,1592]" captionTargetBox="[156,1139,216,1547]" captionTargetId="figure-14@23.[156,1139,216,1560]" captionTargetPageId="23" captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="22" pageNumber="515">Figures 10</figureCitation>
,
<figureCitation id="0C832A6FEB7EED53FF0BFA506234806E" box="[238,275,1535,1557]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="22" pageNumber="515">14F</figureCitation>
,
<figureCitation id="0C832A6FEB7EED53FEFBFA506260806E" box="[286,327,1535,1557]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="22" pageNumber="515">16E</figureCitation>
,
<figureCitation id="0C832A6FEB7EED53FEB4FA50625D806E" box="[337,378,1535,1557]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="22" pageNumber="515">17E</figureCitation>
,
<figureCitation id="0C832A6FEB7EED53FE61FA5062BB806E" box="[388,412,1535,1557]" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="22" pageNumber="515">21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB7EED53FF9EF98B6709802C" pageId="22" pageNumber="515" type="reference_group">
<paragraph id="940736EAEB7EED53FF9EF98B6709802C" blockId="22.[123,1174,1503,1725]" pageId="22" pageNumber="515">
<treatmentCitationGroup id="B4A811C4EB7EED53FF9EF98B6709802C" pageId="22" pageNumber="515">
<treatmentCitation id="151910FBEB7EED53FF9EF98B612B8041" author="HELLER, K. M." box="[123,524,1572,1594]" page="302" pageId="22" pageNumber="515" year="1923">
<taxonomicName id="53B84D69EB7EED53FF9EF98B612B8041" ID-CoL="6C5XG" authority="Heller, 1923: 302" authorityName="Heller" authorityPageNumber="302" authorityYear="1923" box="[123,524,1572,1594]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="515" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB7EED53FF9EF98B62468041" box="[123,353,1572,1594]" italics="true" pageId="22" pageNumber="515">Cyclosomus philippinus</emphasis>
<bibRefCitation id="F0294B1BEB7EED53FE82F98B612B8041" author="HELLER, K. M." box="[359,524,1572,1594]" pageId="22" pageNumber="515" pagination="295 - 307" refId="ref23510" refString="HELLER, K. M. 1923. Some new Malayan Carabidae, especially Philippine. The Philippine Journal of Science 23: 295 - 307 + 1 pl." type="journal article" year="1923">Heller, 1923:302</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="4B038848EB7EED53FDF3F98B61B08041" box="[534,663,1572,1594]" pageId="22" pageNumber="515" type="holotype">HOLOTYPE</typeStatus>
, a female, deposited in
<collectionCode id="F2A9AE2FEB7EED53FC9FF98B60E78041" box="[890,960,1572,1594]" pageId="22" pageNumber="515">SMTD</collectionCode>
. Type locality:
<collectingCountry id="ECAF767AEB7EED53FBB1F98B63C7802C" name="Philippines" pageId="22" pageNumber="515">Philippines</collectingCountry>
, Luzon Island,
<collectingRegion id="567CF808EB7EED53FE90F9EE613B802C" box="[373,540,1601,1623]" country="Philippines" name="Laguna" pageId="22" pageNumber="515">Laguna Province</collectingRegion>
, Los Baños.
<treatmentCitation id="151910FBEB7EED53FD72F9EE604A802C" author="ANDREWES, H. E." box="[663,877,1601,1623]" page="349" pageId="22" pageNumber="515" year="1926">
<bibRefCitation id="F0294B1BEB7EED53FD72F9EE604A802C" author="ANDREWES, H. E." box="[663,877,1601,1623]" pageId="22" pageNumber="515" pagination="345 - 361" refId="ref23053" refString="ANDREWES, H. E. 1926. A catalogue of Philippine Carabidae. Philippine Journal of Science 31: 345 - 361." type="journal article" year="1926">Andrewes (1926:349)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="151910FBEB7EED53FC92F9EE670D802C" author="CSIKI, E." box="[887,1066,1601,1623]" page="1295" pageId="22" pageNumber="515" year="1932">
<bibRefCitation id="F0294B1BEB7EED53FC92F9EE670D802C" author="CSIKI, E." box="[887,1066,1601,1623]" pageId="22" pageNumber="515" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1295)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB7EED5DFF4EF9C9602582C3" lastPageId="24" lastPageNumber="517" pageId="22" pageNumber="515" type="diagnosis">
<paragraph id="940736EAEB7EED5DFF4EF9C9605D846F" blockId="22.[123,1174,1503,1725]" lastBlockId="24.[123,1174,219,1208]" lastPageId="24" lastPageNumber="517" pageId="22" pageNumber="515">
<emphasis id="A6CCEAF8EB7EED53FF4EF9C962388006" bold="true" box="[171,287,1638,1661]" pageId="22" pageNumber="515">Diagnosis.</emphasis>
Adults of
<taxonomicName id="53B84D69EB7EED53FE79F9C961108006" box="[412,567,1637,1661]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="515" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB7EED53FE79F9C961108006" box="[412,567,1637,1661]" italics="true" pageId="22" pageNumber="515">C. philippinus</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="53B84D69EB7EED53FBF1F9CA63EF80E6" pageId="22" pageNumber="515">
<emphasis id="A6CCEAF8EB7EED53FBF1F9CA67B28006" box="[1044,1173,1637,1661]" italics="true" pageId="22" pageNumber="515">Cyclosomus</emphasis>
species
</taxonomicName>
in Asia by the following combination of character states: Body size medium for genus, BL males =
<quantity id="53409B0FEB7EED53FF3FF90A625E80C6" box="[218,377,1701,1725]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.6" metricValueMin="7.2" pageId="22" pageNumber="515" unit="mm" value="7.4" valueMax="7.6" valueMin="7.2">7.2 to 7.6 mm</quantity>
, females = 7.0 to
<quantity id="53409B0FEB7EED53FDA6F90A61BD80C6" box="[579,666,1701,1725]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="22" pageNumber="515" unit="mm" value="7.5">7.5 mm</quantity>
; pronotum (
<figureCitation id="0C832A6FEB7EED53FCCCF90A60AF80C6" box="[809,904,1701,1725]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="22" pageNumber="515">Fig. 14F</figureCitation>
) relatively narrow (ratio PWM/PL = 2.06 to 2.33) and less broadened basally (ratio PWM/PWA = 1.45 to 1.59), with disc piceous to black and lateral pale bands wide and well-defined, anterior angles broadly rounded, lateral margins not sinuate near anterior angles; free apex of prosternal intercoxal process long (as in
<figureCitation id="0C832A6FEB70ED5DFF9EFE9463FB8728" box="[123,220,315,339]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="24" pageNumber="517">Fig. 15C</figureCitation>
); elytral dark pattern varied (
<figureCitation id="0C832A6FEB70ED5DFDFDFE9461828728" box="[536,677,315,339]" captionStart="FIGURE" captionStartId="23.[171,183,1573,1592]" captionTargetBox="[156,1139,216,1547]" captionTargetId="figure-14@23.[156,1139,216,1560]" captionTargetPageId="23" captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="24" pageNumber="517">Figs. 10A, C</figureCitation>
) [slight blue cast to the dark elements seen in
<figureCitation id="0C832A6FEB70ED5DFF9EFEF463FD8708" box="[123,218,347,371]" captionStart="FIGURE" captionStartId="23.[171,183,1573,1592]" captionTargetBox="[156,1139,216,1547]" captionTargetId="figure-14@23.[156,1139,216,1560]" captionTargetPageId="23" captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="24" pageNumber="517">Fig. 10A</figureCitation>
is an imaging artifact], with basal and middle bands of average to greater than average thickness for the genus, preapical dark spot absent; elytral epipleura with long setae over entire length; median lobe of male genitalia with shaft arcuate basally and nearly straight subapically, tapered to a long and pointed apical lamella in lateral aspect (
<figureCitation id="0C832A6FEB70ED5DFCC6FE1360A187AF" box="[803,902,444,468]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="24" pageNumber="517">Fig. 16E</figureCitation>
), shaft moderately broad basally, gradually tapered to moderately broad, apically blunted, and sublanceolate apical lamella in dorsal aspect (
<figureCitation id="0C832A6FEB70ED5DFED4FE5362B5846F" box="[305,402,508,532]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="24" pageNumber="517">Fig. 17E</figureCitation>
); specimen from
<collectingCountry id="ECAF767AEB70ED5DFDA8FE536029846F" box="[589,782,508,532]" name="Philippines" pageId="24" pageNumber="517">Philippine Islands</collectingCountry>
(
<figureCitation id="0C832A6FEB70ED5DFCF8FE536049846F" box="[797,878,508,532]" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="24" pageNumber="517">Fig. 21</figureCitation>
).
</paragraph>
<caption id="C0C76662EB7FED52FF4EF98A67188029" ID-DOI="http://doi.org/10.5281/zenodo.11512978" ID-Zenodo-Dep="11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="23" pageNumber="516" startId="23.[171,183,1573,1592]" targetBox="[156,1139,216,1547]" targetPageId="23" targetType="figure">
<paragraph id="940736EAEB7FED52FF4EF98A67188029" blockId="23.[123,1173,1572,1618]" pageId="23" pageNumber="516">
<smallCapsWord id="92E1A036EB7FED52FF4EF98A63CF804C" baselines="1587,1587" box="[171,232,1573,1592]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="23" pageNumber="516">FIGURE</smallCapsWord>
10.
<taxonomicName id="53B84D69EB7FED52FEE8F98B6113804C" authority="Heller. A." authorityName="Heller. A." box="[269,564,1572,1591]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="516" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB7FED52FEE8F98B62FD804C" box="[269,474,1572,1591]" italics="true" pageId="23" pageNumber="516">Cyclosomus philippinus</emphasis>
Heller. A.
</taxonomicName>
Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="940736EAEB70ED5DFF4EFDB2677884CE" blockId="24.[123,1174,219,1208]" pageId="24" pageNumber="517">
Only
<taxonomicName id="53B84D69EB70ED5DFF0FFDB262A3844E" box="[234,388,541,565]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB70ED5DFF0FFDB262A3844E" box="[234,388,541,565]" italics="true" pageId="24" pageNumber="517">C. philippinus</emphasis>
</taxonomicName>
and
<taxonomicName id="53B84D69EB70ED5DFE5BFDB26164844E" authorityName=", Kavanaugh" authorityYear="2015" box="[446,579,541,565]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB70ED5DFE5BFDB26164844E" box="[446,579,541,565]" italics="true" pageId="24" pageNumber="517">C. flexuosus</emphasis>
</taxonomicName>
members share elytral epipleura with long setae over their entire length. Specimens of these two species can be distinguished by the presence (in
<taxonomicName id="53B84D69EB70ED5DFBA2FD92639F840E" authorityName=", Kavanaugh" authorityYear="2015" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB70ED5DFBA2FD92639F840E" italics="true" pageId="24" pageNumber="517">C. flexuosus</emphasis>
</taxonomicName>
) or absence (in
<taxonomicName id="53B84D69EB70ED5DFE80FDF162DA840E" box="[357,509,605,629]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB70ED5DFE80FDF162DA840E" box="[357,509,605,629]" italics="true" pageId="24" pageNumber="517">C. philippinus</emphasis>
</taxonomicName>
) of a preapical dark spot. Males of the two species also differ distinctly in the shape and curvature of the median lobe in lateral aspect (compare
<figureCitation id="0C832A6FEB70ED5DFC1AFDD2677984EE" box="[1023,1118,637,661]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="24" pageNumber="517">Fig. 16B</figureCitation>
with
<figureCitation id="0C832A6FEB70ED5DFF9EFD3263FF84CE" box="[123,216,669,693]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="24" pageNumber="517">Fig. 16E</figureCitation>
) and shape of the apical lamella in dorsal aspect (compare
<figureCitation id="0C832A6FEB70ED5DFCB3FD32609384CE" box="[854,948,669,693]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="24" pageNumber="517">Fig. 17B</figureCitation>
with
<figureCitation id="0C832A6FEB70ED5DFC17FD32677584CE" box="[1010,1106,669,693]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="24" pageNumber="517">Fig. 17E</figureCitation>
).
</paragraph>
<paragraph id="940736EAEB70ED5DFF4EFD1160F8848D" blockId="24.[123,1174,219,1208]" pageId="24" pageNumber="517">
<emphasis id="A6CCEAF8EB70ED5DFF4EFD1162B084AE" bold="true" box="[171,407,702,725]" pageId="24" pageNumber="517">Habitat distribution.</emphasis>
Unknown, but presumed to be restricted to the sandy shores of medium to large rivers at low elevation, like members of most other species of the genus.
</paragraph>
<paragraph id="940736EAEB70ED5DFF4EFD5060A685EC" blockId="24.[123,1174,219,1208]" pageId="24" pageNumber="517">
<emphasis id="A6CCEAF8EB70ED5DFF4EFD5062F8856D" bold="true" box="[171,479,767,790]" pageId="24" pageNumber="517">Geographical distribution.</emphasis>
<location id="91676031EB70ED5DFE0EFD506129856D" LSID="urn:lsid:plazi:treatment:1C1187FCEB7EED5DFF9EFA70602582C3:91676031EB70ED5DFE0EFD506129856D" box="[491,526,767,790]" name="We" pageId="24" pageNumber="517">We</location>
examined a total of
<specimenCount id="82BEFD63EB70ED5DFCE3FD516086856D" box="[774,929,766,790]" count="6" pageId="24" pageNumber="517" type="generic">six specimens</specimenCount>
(
<specimenCount id="82BEFD63EB70ED5DFC50FD51670A856D" box="[949,1069,766,790]" count="4" pageId="24" pageNumber="517" type="male">four males</specimenCount>
and
<specimenCount id="82BEFD63EB70ED5DFB88FD5063F7854D" count="2" pageId="24" pageNumber="517" type="female">two females</specimenCount>
) from the following localities:
<materialsCitation id="24D03CB7EB70ED5DFDC6FCB16290852D" ID-GBIF-Occurrence="4905152308" collectingDate="1998-11-05" collectingDateMax="1998-11-07" collectingDateMin="1998-11-05" collectionCode="SMTD, MFNB" collectorName="C. F. Baker" country="Philippines" county="Luzon Island" elevation="150" location="Mountains" municipality="Los Banos" pageId="24" pageNumber="517" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Zambales">
2024-06-21 12:30:48 +02:00
<collectingCountry id="ECAF767AEB70ED5DFDC6FCB161E3854D" box="[547,708,798,822]" name="Philippines" pageId="24" pageNumber="517">PHILIPPINES</collectingCountry>
:
<collectingCounty id="7D664E66EB70ED5DFD2AFCB06046854D" box="[719,865,798,822]" pageId="24" pageNumber="517">Luzon Island</collectingCounty>
:
<collectingMunicipality id="7463AC90EB70ED5DFC89FCB060F8854D" box="[876,991,799,822]" pageId="24" pageNumber="517">Los Baños</collectingMunicipality>
(
<collectorName id="394D533CEB70ED5DFC15FCB16743854D" box="[1008,1124,798,822]" pageId="24" pageNumber="517">C.F. Baker</collectorName>
collector [
<specimenCount id="82BEFD63EB70ED5DFF2DFC906260852D" box="[200,327,830,854]" count="1" pageId="24" pageNumber="517" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB70ED5DFEB0FC916282852D" box="[341,421,830,854]" pageId="24" pageNumber="517">SMTD</collectionCode>
]),
</materialsCitation>
<materialsCitation id="24D03CB7EB70ED5DFE27FC9161C7850D" ID-GBIF-Occurrence="4905152372" collectingDate="1998-11-05" collectingDateMax="1998-11-07" collectingDateMin="1998-11-05" collectionCode="SMTD, MFNB" collectorName="Mey &amp; Spiedel " country="Philippines" county="Luzon Island" elevation="150" location="Mountains" municipality="Los Banos" pageId="24" pageNumber="517" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Zambales">
2024-06-21 12:30:48 +02:00
<collectingRegion id="567CF808EB70ED5DFE27FC91610D852D" box="[450,554,830,854]" country="Philippines" name="Zambales" pageId="24" pageNumber="517">Zambales</collectingRegion>
<location id="91676031EB70ED5DFDD6FC916182852D" LSID="urn:lsid:plazi:treatment:1C1187FCEB7EED5DFF9EFA70602582C3:91676031EB70ED5DFDD6FC916182852D" box="[563,677,830,854]" country="Philippines" county="Luzon Island" municipality="Los Banos" name="Mountains" pageId="24" pageNumber="517" stateProvince="Zambales">Mountains</location>
(
<location id="91676031EB70ED5DFD52FC9161C7852D" LSID="urn:lsid:plazi:treatment:1C1187FCEB7EED5DFF9EFA70602582C3:91676031EB70ED5DFD52FC9161C7852D" box="[695,736,830,854]" country="Philippines" county="Luzon Island" municipality="Los Banos" name="Pili" pageId="24" pageNumber="517" stateProvince="Zambales">Pili</location>
,
<quantity id="53409B0FEB70ED5DFD0BFC916010852D" box="[750,823,830,854]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="517" unit="m" value="150.0">
<elevation id="1F95D1D9EB70ED5DFD0BFC916010852D" box="[750,823,830,854]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="517" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="E006102AEB70ED5DFCA3FC916701852D" box="[838,1062,830,854]" pageId="24" pageNumber="517" value="1998-11-05" valueMax="1998-11-07" valueMin="1998-11-05">
<collectingDate id="F042E9C2EB70ED5DFCA3FC916701852D" box="[838,1062,830,854]" pageId="24" pageNumber="517" value="1998-11-05" valueMax="1998-11-07" valueMin="1998-11-05">5-7 November 1998</collectingDate>
</date>
,
<collectorName id="394D533CEB70ED5DFBD1FC906742852C" box="[1076,1125,831,855]" pageId="24" pageNumber="517">Mey</collectorName>
and
<collectorName id="394D533CEB70ED5DFF9EFCF163EC850D" box="[123,203,862,886]" pageId="24" pageNumber="517">Spiedel</collectorName>
collectors [
<specimenCount id="82BEFD63EB70ED5DFEAFFCF0629C850D" box="[330,443,862,886]" count="2" pageId="24" pageNumber="517" type="male">two males</specimenCount>
and
<specimenCount id="82BEFD63EB70ED5DFE11FCF06148850D" box="[500,623,862,886]" count="1" pageId="24" pageNumber="517" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB70ED5DFD99FCF161E8850D" box="[636,719,862,886]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="24" pageNumber="517">MFNB</collectionCode>
]).
</materialsCitation>
<materialsCitation id="24D03CB7EB70ED5DFD0DFCF1605985EC" ID-GBIF-Occurrence="4905152320" collectingDate="1998-11-05" collectingDateMax="1998-11-07" collectingDateMin="1998-11-05" collectionCode="SMTD, MFNB" collectorName=" C. S. Banks" country="Philippines" county="Luzon Island" elevation="150" location="Mountains" municipality="Los Banos" pageId="24" pageNumber="517" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Zambales">
2024-06-21 12:30:48 +02:00
<collectorName id="394D533CEB70ED5DFD0DFCF1607B850D" box="[744,860,862,886]" pageId="24" pageNumber="517">Specimens</collectorName>
without locality: (“
<collectorName id="394D533CEB70ED5DFBD7FCF167B1850D" box="[1074,1174,862,886]" pageId="24" pageNumber="517">Acc. No.</collectorName>
4696” and “
<collectorName id="394D533CEB70ED5DFF1EFCD0627985ED" box="[251,350,895,919]" pageId="24" pageNumber="517">Acc. No.</collectorName>
5451”,
<collectorName id="394D533CEB70ED5DFE51FCD0610B85EC" box="[436,556,895,919]" pageId="24" pageNumber="517">C.S. Banks</collectorName>
collector [
<specimenCount id="82BEFD63EB70ED5DFD7BFCD0603585EC" box="[670,786,895,919]" count="2" pageId="24" pageNumber="517" type="male">two males</specimenCount>
;
<collectionCode id="F2A9AE2FEB70ED5DFCF8FCD0605785EC" box="[797,880,895,919]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="24" pageNumber="517">MFNB</collectionCode>
])
</materialsCitation>
.
</paragraph>
<paragraph id="940736EAEB70ED5DFF4EFC3060B785CC" blockId="24.[123,1174,219,1208]" box="[171,912,927,951]" pageId="24" pageNumber="517">
This species is known only from Luzon Island,
<collectingCountry id="ECAF767AEB70ED5DFD4EFC30600485CC" box="[683,803,927,951]" name="Philippines" pageId="24" pageNumber="517">Philippines</collectingCountry>
(
<figureCitation id="0C832A6FEB70ED5DFCD6FC3060A485CC" box="[819,899,927,951]" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="24" pageNumber="517">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="940736EAEB70ED5DFF4EFC6F611A8243" blockId="24.[123,1174,219,1208]" pageId="24" pageNumber="517">
<emphasis id="A6CCEAF8EB70ED5DFF4EFC6F629B85AC" bold="true" box="[171,444,960,983]" pageId="24" pageNumber="517">Geographical variation.</emphasis>
The range of variation in the thickness of the middle dark band of the elytra seen among specimens examined is shown in
<figureCitation id="0C832A6FEB70ED5DFD2FFC7060B4858C" box="[714,915,991,1015]" captionStart="FIGURE" captionStartId="23.[171,183,1573,1592]" captionTargetBox="[156,1139,216,1547]" captionTargetId="figure-14@23.[156,1139,216,1560]" captionTargetPageId="23" captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="24" pageNumber="517">Figs. 10A and 10C</figureCitation>
, with the
<typeStatus id="4B038848EB70ED5DFC1FFC706770858C" box="[1018,1111,991,1015]" pageId="24" pageNumber="517" type="holotype">holotype</typeStatus>
specimen presenting the thickest band seen. We were unable to identify any geographical component to this variation in our sample of specimens.
</paragraph>
<paragraph id="940736EAEB70ED5DFF4EFBEE602582C3" blockId="24.[123,1174,219,1208]" pageId="24" pageNumber="517">
<emphasis id="A6CCEAF8EB70ED5DFF4EFBEE60628223" bold="true" box="[171,837,1088,1112]" pageId="24" pageNumber="517">
Geographical relationships with other
<taxonomicName id="53B84D69EB70ED5DFD81FBEF60628223" box="[612,837,1088,1112]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A6CCEAF8EB70ED5DFD81FBEF61CF8223" bold="true" box="[612,744,1088,1112]" italics="true" pageId="24" pageNumber="517">Cyclosomus</emphasis>
species.
</taxonomicName>
</emphasis>
No other congeneric species is known to occur in the
<collectingCountry id="ECAF767AEB70ED5DFE8AFBCF62CC8203" box="[367,491,1120,1144]" name="Philippines" pageId="24" pageNumber="517">Philippines</collectingCountry>
; and the nearest
<taxonomicName id="53B84D69EB70ED5DFD44FBCF60058203" authorityName="Latreille" authorityYear="1829" box="[673,802,1120,1144]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="A6CCEAF8EB70ED5DFD44FBCF60058203" box="[673,802,1120,1144]" italics="true" pageId="24" pageNumber="517">Cyclosomus</emphasis>
</taxonomicName>
records are for
<taxonomicName id="53B84D69EB70ED5DFC2AFBCE671C8203" box="[975,1083,1121,1144]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="inustus">
<emphasis id="A6CCEAF8EB70ED5DFC2AFBCE671C8203" box="[975,1083,1121,1144]" italics="true" pageId="24" pageNumber="517">C. inustus</emphasis>
</taxonomicName>
, on
<collectingCountry id="ECAF767AEB70ED5DFB8EFBCF638082E3" name="Taiwan" pageId="24" pageNumber="517">Taiwan</collectingCountry>
(about
<quantity id="53409B0FEB70ED5DFF1AFB2F624782E3" box="[255,352,1152,1176]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="517" unit="km" value="1000.0">1000 km</quantity>
to the north) and
<collectingCountry id="ECAF767AEB70ED5DFDCBFB2E618982E2" box="[558,686,1153,1177]" name="Hong Kong" pageId="24" pageNumber="517">Hong Kong</collectingCountry>
and
<collectingRegion id="567CF808EB70ED5DFD0CFB2F60E982E3" box="[745,974,1152,1176]" country="China" name="Guangdong" pageId="24" pageNumber="517">Guangdong Province</collectingRegion>
(across the South
<collectingCountry id="ECAF767AEB70ED5DFF9EFB0F639C82C3" box="[123,187,1184,1208]" name="China" pageId="24" pageNumber="517">China</collectingCountry>
Sea and more than
<quantity id="53409B0FEB70ED5DFE74FB0F62C982C3" box="[401,494,1184,1208]" metricMagnitude="6" metricUnit="m" metricValue="1.1" pageId="24" pageNumber="517" unit="km" value="1100.0">1100 km</quantity>
distant to the northwest).
</paragraph>
</subSubSection>
</treatment>
</document>