A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.
Author
Kavanaugh, David H.
Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.
Author
Cueva-Dabkoski, Mollie
Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.
text
Proceedings of the California Academy of Sciences
2023
2023-02-15
67
21
493
537
journal article
299696
10.5281/zenodo.11512952
2b56e3b0-2274-4963-b243-9c991b07072c
0068-547X
11512952
Cyclosomus philippinus
Heller
Figures 10
,
14F
,
16E
,
17E
,
21
Cyclosomus philippinus
Heller, 1923:302
.
HOLOTYPE
, a female, deposited in
SMTD
. Type locality:
Philippines
, Luzon Island,
Laguna Province
, Los Baños.
Andrewes (1926:349)
;
Csiki (1932:1295)
.
Diagnosis.
Adults of
C. philippinus
can be distinguished from those of other
Cyclosomus
species
in Asia by the following combination of character states: Body size medium for genus, BL males =
7.2 to 7.6 mm
, females = 7.0 to
7.5 mm
; pronotum (
Fig. 14F
) relatively narrow (ratio PWM/PL = 2.06 to 2.33) and less broadened basally (ratio PWM/PWA = 1.45 to 1.59), with disc piceous to black and lateral pale bands wide and well-defined, anterior angles broadly rounded, lateral margins not sinuate near anterior angles; free apex of prosternal intercoxal process long (as in
Fig. 15C
); elytral dark pattern varied (
Figs. 10A, C
) [slight blue cast to the dark elements seen in
Fig. 10A
is an imaging artifact], with basal and middle bands of average to greater than average thickness for the genus, preapical dark spot absent; elytral epipleura with long setae over entire length; median lobe of male genitalia with shaft arcuate basally and nearly straight subapically, tapered to a long and pointed apical lamella in lateral aspect (
Fig. 16E
), shaft moderately broad basally, gradually tapered to moderately broad, apically blunted, and sublanceolate apical lamella in dorsal aspect (
Fig. 17E
); specimen from
Philippine Islands
(
Fig. 21
).
FIGURE
10.
Cyclosomus philippinus
Heller. A.
Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm.
Only
C. philippinus
and
C. flexuosus
members share elytral epipleura with long setae over their entire length. Specimens of these two species can be distinguished by the presence (in
C. flexuosus
) or absence (in
C. philippinus
) of a preapical dark spot. Males of the two species also differ distinctly in the shape and curvature of the median lobe in lateral aspect (compare
Fig. 16B
with
Fig. 16E
) and shape of the apical lamella in dorsal aspect (compare
Fig. 17B
with
Fig. 17E
).
Habitat distribution.
Unknown, but presumed to be restricted to the sandy shores of medium to large rivers at low elevation, like members of most other species of the genus.
Geographical distribution.
We
examined a total of
six specimens
(
four males
and
two females
) from the following localities:
PHILIPPINES
:
Luzon Island
:
Los Baños
(
C.F. Baker
collector [
one female
;
SMTD
]),
Zambales
Mountains
(
Pili
,
150 m
,
5-7 November 1998
,
Mey
and
Spiedel
collectors [
two males
and
one female
;
MFNB
]).
Specimens
without locality: (“
Acc. No.
4696” and “
Acc. No.
5451”,
C.S. Banks
collector [
two males
;
MFNB
])
.
This species is known only from Luzon Island,
Philippines
(
Fig. 21
).
Geographical variation.
The range of variation in the thickness of the middle dark band of the elytra seen among specimens examined is shown in
Figs. 10A and 10C
, with the
holotype
specimen presenting the thickest band seen. We were unable to identify any geographical component to this variation in our sample of specimens.
Geographical relationships with other
Cyclosomus
species.
No other congeneric species is known to occur in the
Philippines
; and the nearest
Cyclosomus
records are for
C. inustus
, on
Taiwan
(about
1000 km
to the north) and
Hong Kong
and
Guangdong Province
(across the South
China
Sea and more than
1100 km
distant to the northwest).