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<document id="11A74AAC3BDBA91B32275A15569A30A1" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. & Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB7AED51FF9EFC0C6743826D" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus marginatus Motschulsky" docType="treatment" docVersion="5" lastPageNumber="513" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="511" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B72D4941EAD5EAFA5890D7008AA1F78B">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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<mods:namePart id="4F0F525D3C250FEBE508EA0B537D2996">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="C1380596171B43AD864587BB972ACDD7">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:namePart id="64AA3C331EBC4D6A780421FF78834D16">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="681EEED4ADB82865EE9E430C79D79F2C">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:title id="E1035EC9A00A3882B733611122AEE15B">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="1C238F46620B8FB81536A341BB6B00F7">
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<mods:date id="0BB3E385E53E0D1A71913F1AF83D8B4A">2023</mods:date>
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<mods:number id="CCE49EDC549EBCD96560DFC04678215C">2023-02-15</mods:number>
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<mods:number id="4EB4F254F64CA293A42647DD0D182B64">67</mods:number>
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<mods:number id="E967CB55A9A65ABA0253734B6CE425B6">21</mods:number>
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<mods:identifier id="C86AC6EF3E0DF57FAFAA6E8D4A168409" type="DOI">10.5281/zenodo.11512952</mods:identifier>
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<treatment id="1C1187FCEB7AED51FF9EFC0C6743826D" ID-DOI="http://doi.org/10.5281/zenodo.12726725" ID-GBIF-Taxon="235039985" ID-Zenodo-Dep="12726725" LSID="urn:lsid:plazi:treatment:1C1187FCEB7AED51FF9EFC0C6743826D" httpUri="http://treatment.plazi.org/id/1C1187FCEB7AED51FF9EFC0C6743826D" lastPageId="20" lastPageNumber="513" pageId="18" pageNumber="511">
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<subSubSection id="DCA26561EB7AED57FF9EFC0C613B85C1" box="[123,540,931,955]" pageId="18" pageNumber="511" type="nomenclature">
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<paragraph id="940736EAEB7AED57FF9EFC0C613B85C1" blockId="18.[123,1175,931,1725]" box="[123,540,931,955]" pageId="18" pageNumber="511">
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<heading id="CF4F8186EB7AED57FF9EFC0C613B85C1" box="[123,540,931,955]" fontSize="10" level="2" pageId="18" pageNumber="511" reason="4">
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<taxonomicName id="53B84D69EB7AED57FF9EFC0C613B85C1" ID-CoL="32X59" authority="Motschulsky" authorityName="Motschulsky" box="[123,540,931,955]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">
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<emphasis id="A6CCEAF8EB7AED57FF9EFC0C613B85C1" bold="true" box="[123,540,931,955]" pageId="18" pageNumber="511">
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<emphasis id="A6CCEAF8EB7AED57FF9EFC0C62A385C0" bold="true" box="[123,388,931,955]" italics="true" pageId="18" pageNumber="511">Cyclosomus marginatus</emphasis>
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Motschulsky
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB7AED57FF9EFC6C62BB85A2" box="[123,412,963,985]" pageId="18" pageNumber="511" type="description">
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<paragraph id="940736EAEB7AED57FF9EFC6C62BB85A2" blockId="18.[123,1175,931,1725]" box="[123,412,963,985]" pageId="18" pageNumber="511">
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<figureCitation id="0C832A6FEB7AED57FF9EFC6C63F185A2" box="[123,214,963,985]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="18" pageNumber="511">Figures 8</figureCitation>
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,
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<figureCitation id="0C832A6FEB7AED57FF07FC6C622A85A2" box="[226,269,963,985]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="18" pageNumber="511">14D</figureCitation>
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,
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<figureCitation id="0C832A6FEB7AED57FEFDFC6C626485A2" box="[280,323,963,985]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="18" pageNumber="511">16D</figureCitation>
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,
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<figureCitation id="0C832A6FEB7AED57FEABFC6C625E85A2" box="[334,377,963,985]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="18" pageNumber="511">17D</figureCitation>
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,
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<figureCitation id="0C832A6FEB7AED57FE61FC6C62BB85A2" box="[388,412,963,985]" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="18" pageNumber="511">21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB7AED57FF9EFC4762FF8363" pageId="18" pageNumber="511" type="reference_group">
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<paragraph id="940736EAEB7AED57FF9EFC4762408208" blockId="18.[123,1175,931,1725]" pageId="18" pageNumber="511">
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<treatmentCitationGroup id="B4A811C4EB7AED57FF9EFC4767B18585" box="[123,1174,1000,1022]" pageId="18" pageNumber="511">
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<treatmentCitation id="151910FBEB7AED57FF9EFC47617B8585" author="MOTSCHULSKY, V." box="[123,604,1000,1022]" page="200" pageId="18" pageNumber="511" year="1864">
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<taxonomicName id="53B84D69EB7AED57FF9EFC47617B8585" ID-CoL="32X59" authority="Motschulsky, 1864: 200" authorityName="Motschulsky" authorityPageNumber="200" authorityYear="1864" box="[123,604,1000,1022]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">
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<emphasis id="A6CCEAF8EB7AED57FF9EFC4762418585" box="[123,358,1000,1022]" italics="true" pageId="18" pageNumber="511">Cyclosomus marginatus</emphasis>
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<bibRefCitation id="F0294B1BEB7AED57FE97FC47617B8585" author="MOTSCHULSKY, V." box="[370,604,1000,1022]" pageId="18" pageNumber="511" refId="ref23886" refString="MOTSCHULSKY, V. 1864. Enumeration des nouvelles especes de coleopteres rapports de des voyages. 4 - eme. Carabicines. Bulletin de la Societe Imperiale des Naturalistes de Moscou 37: 171 - 40." type="book" year="1864">Motschulsky, 1864:200</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="4B038848EB7AED57FD8EFC4761D18585" box="[619,758,1000,1022]" pageId="18" pageNumber="511" type="lectotype">LECTOTYPE</typeStatus>
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, here designated, a female, deposited in
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</treatmentCitationGroup>
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<collectionCode id="F2A9AE2FEB7AED57FF4EFBA963DF8260" box="[171,248,1030,1051]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="18" pageNumber="511" type="Museum">ZMMU</collectionCode>
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, labeled: “Ind. Or.” [handwritten]/ “
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<taxonomicName id="53B84D69EB7AED57FDBDFBAA60CB8260" ID-CoL="32X59" authority="Motsch. Ind. Or." authorityName="Motsch. Ind. Or." box="[600,1004,1029,1051]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">Cyclosomus marginatus Motsch. Ind. Or.</taxonomicName>
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”/ “
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<taxonomicName id="53B84D69EB7AED57FBEBFBAA67B28260" ID-CoL="32X59" authority="Motschulsky, 1864: 200" authorityName="Motschulsky" authorityPageNumber="200" authorityYear="1864" box="[1038,1173,1029,1051]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">C. marginatus</taxonomicName>
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Motch =
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<taxonomicName id="53B84D69EB7AED57FEE0FB8C616A8242" authority="Wied. H. E. Andrewes" box="[261,589,1059,1081]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="suturalis">C. suturalis Wied. H.E. Andrewes</taxonomicName>
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det.”/ “
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<typeStatus id="4B038848EB7AED57FD7CFB8C60038242" box="[665,804,1059,1081]" pageId="18" pageNumber="511" type="lectotype">LECTOTYPE</typeStatus>
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<taxonomicName id="53B84D69EB7AED57FCCEFB8C67B28242" authority="Motschulsky 1864" authorityName="Motschulsky" authorityYear="1864" box="[811,1173,1059,1081]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">
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<emphasis id="A6CCEAF8EB7AED57FCCEFB8C67358242" box="[811,1042,1059,1081]" italics="true" pageId="18" pageNumber="511">Cyclosomus marginatus</emphasis>
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<bibRefCitation id="F0294B1BEB7AED57FBFCFB8C67B28242" author="MOTSCHULSKY, V." box="[1049,1173,1059,1081]" pageId="18" pageNumber="511" refId="ref23886" refString="MOTSCHULSKY, V. 1864. Enumeration des nouvelles especes de coleopteres rapports de des voyages. 4 - eme. Carabicines. Bulletin de la Societe Imperiale des Naturalistes de Moscou 37: 171 - 40." type="book" year="1864">Motschulsky</bibRefCitation>
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</taxonomicName>
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1864 design. by D.H. Kavanaugh & M. Cueva-Dabkoski 2023” [red label].
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<typeStatus id="4B038848EB7AED57FC6DFBEE609F822D" box="[904,952,1089,1110]" pageId="18" pageNumber="511">Type</typeStatus>
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locality: eastern
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<collectingCountry id="ECAF767AEB7AED57FBBBFBEF67B5822D" box="[1118,1170,1088,1110]" name="India" pageId="18" pageNumber="511">India</collectingCountry>
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.
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<bibRefCitation id="F0294B1BEB7AED57FF4EFBF262448208" author="CHAUDOIR, M." box="[171,355,1117,1139]" pageId="18" pageNumber="511" pagination="1 - 84" refId="ref23225" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir 1876: 32</bibRefCitation>
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.
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</paragraph>
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<paragraph id="940736EAEB7AED57FF9EFBD5670782EB" blockId="18.[123,1175,931,1725]" box="[123,1056,1146,1168]" pageId="18" pageNumber="511">
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<treatmentCitationGroup id="B4A811C4EB7AED57FF9EFBD5670782EB" box="[123,1056,1146,1168]" pageId="18" pageNumber="511">
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<treatmentCitation id="151910FBEB7AED57FF9EFBD5610682EB" author="ANDREWES, H. E." box="[123,545,1146,1168]" page="166" pageId="18" pageNumber="511" year="1921">
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<taxonomicName id="53B84D69EB7AED57FF9EFBD5610682EB" ID-CoL="32X5F" authority=", Andrewes (1921: 166" authorityName="Andrewes" authorityYear="1921" box="[123,545,1146,1168]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FF9EFBD5626E82EB" box="[123,329,1146,1168]" italics="true" pageId="18" pageNumber="511">Cyclosomus suturalis</emphasis>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="F0294B1BEB7AED57FEB0FBD5610682EB" author="ANDREWES, H. E." box="[341,545,1146,1168]" pageId="18" pageNumber="511" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:166</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="F0294B1BEB7AED57FDC9FBD5617882EB" author="ANDREWES, H. E." box="[556,607,1146,1168]" pageId="18" pageNumber="511" pagination="459 - 468" refId="ref23019" refString="ANDREWES, H. E. 1924. XXII. On the Oriental Carabidae of the " Reise Novara. " Transactions of the Entomological Society of London 1923: 459 - 468." type="journal article" year="1924">1924</bibRefCitation>
|
|||
|
|
a:464);
|
|||
|
|
<treatmentCitation id="151910FBEB7AED57FD43FBD5607E82EB" author="CSIKI, E." box="[678,857,1146,1168]" page="1295" pageId="18" pageNumber="511" year="1932">
|
|||
|
|
<bibRefCitation id="F0294B1BEB7AED57FD43FBD5607E82EB" author="CSIKI, E." box="[678,857,1146,1168]" pageId="18" pageNumber="511" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1295)</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
;
|
|||
|
|
<treatmentCitation id="151910FBEB7AED57FC81FBD4673B82EB" author="LORENZ, W." box="[868,1052,1146,1168]" page="452" pageId="18" pageNumber="511" year="2005">
|
|||
|
|
<bibRefCitation id="F0294B1BEB7AED57FC81FBD4673B82EB" author="LORENZ, W." box="[868,1052,1146,1168]" pageId="18" pageNumber="511" refId="ref23840" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005:452)</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
.
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB7AED57FF4EFB0F62FF8363" blockId="18.[123,1175,931,1725]" pageId="18" pageNumber="511">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FF4EFB0F610082CC" bold="true" box="[171,551,1184,1207]" pageId="18" pageNumber="511">
|
|||
|
|
Notes on
|
|||
|
|
<typeStatus id="4B038848EB7AED57FEF7FB0F626A82CC" box="[274,333,1184,1207]" pageId="18" pageNumber="511">types</typeStatus>
|
|||
|
|
and nomenclature.
|
|||
|
|
</emphasis>
|
|||
|
|
See below under
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FD07FB0F60D282CC" ID-CoL="32X5F" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[738,1013,1183,1207]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FD07FB0F607A82CC" box="[738,861,1183,1207]" italics="true" pageId="18" pageNumber="511">C. suturalis</emphasis>
|
|||
|
|
(Wiedemann)
|
|||
|
|
</taxonomicName>
|
|||
|
|
for further discussion about
|
|||
|
|
<typeStatus id="4B038848EB7AED57FEF3FB6F627582A3" box="[278,338,1216,1240]" pageId="18" pageNumber="511">types</typeStatus>
|
|||
|
|
. We must note here that if, as discussed above,
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FCBFFB6F60D482AC" ID-CoL="32X59" authority="Motschulsky, 1864: 200" authorityName="Motschulsky" authorityPageNumber="200" authorityYear="1864" box="[858,1011,1216,1239]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FCBFFB6F60D482AC" box="[858,1011,1216,1239]" italics="true" pageId="18" pageNumber="511">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FBCCFB6F67B182AC" ID-CoL="32X57" box="[1065,1174,1216,1239]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FBCCFB6F67B182AC" box="[1065,1174,1216,1239]" italics="true" pageId="18" pageNumber="511">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were to be considered conspecific, then the name
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FD6FFB4E60058283" ID-CoL="32X59" authority="Motschulsky, 1864: 200" authorityName="Motschulsky" authorityPageNumber="200" authorityYear="1864" box="[650,802,1249,1272]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FD6FFB4E60058283" box="[650,802,1249,1272]" italics="true" pageId="18" pageNumber="511">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has priority. We choose not to propose that synonymy at this time.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB7AED56FF4EFA8E60AC8019" lastPageId="19" lastPageNumber="512" pageId="18" pageNumber="511" type="diagnosis">
|
|||
|
|
<paragraph id="940736EAEB7AED56FF4EFA8E67518379" blockId="18.[123,1175,931,1725]" lastBlockId="19.[123,1174,1162,1699]" lastPageId="19" lastPageNumber="512" pageId="18" pageNumber="511">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FF4EFA8E62388343" bold="true" box="[171,287,1313,1336]" pageId="18" pageNumber="511">Diagnosis.</emphasis>
|
|||
|
|
Adults of
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FE79FA8E611F8342" box="[412,568,1313,1337]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FE79FA8E611F8342" box="[412,568,1313,1337]" italics="true" pageId="18" pageNumber="511">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be distinguished from those of other
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FBF1FA8E63EF8322" pageId="18" pageNumber="511">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FBF1FA8E67B28342" box="[1044,1173,1313,1337]" italics="true" pageId="18" pageNumber="511">Cyclosomus</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
in Asia by the following combination of character states: Body size medium for genus, BL male =
|
|||
|
|
<quantity id="53409B0FEB7AED57FF2FFACE623A8302" box="[202,285,1377,1401]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="18" pageNumber="511" unit="mm" value="7.3">7.3 mm</quantity>
|
|||
|
|
, female = 7.0 mm; body form (
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FD83FACE619E8302" box="[614,697,1377,1401]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="18" pageNumber="511">Fig. 8A</figureCitation>
|
|||
|
|
) roundly ovoid (ratio BL/EW = 1.46 to 1.49; ratio EL/EW = 0.94 to 0.96), with elytra widest at or near mid-length; dorsal surface (
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FBCBFA2D67AA83E1" box="[1070,1165,1410,1434]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="18" pageNumber="511">Figs. 8A</figureCitation>
|
|||
|
|
) with distinctly contrasting pale and dark areas; pronotum (
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FCC4FA0D60AD83C1" box="[801,906,1442,1466]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="18" pageNumber="511">Fig. 14D</figureCitation>
|
|||
|
|
) relatively broad (ratio PWM/PL = 2.67 to 2.92) and more broadened basally (ratio PWM/PWA = 1.65 to 1.67), with disc rufopiceous to black, lateral pale bands broad and well-defined, anterior angles broader, lateral margins not sinuate or very faintly sinuate near anterior angles; free apex of prosternal intercoxal process medium in length (as in
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FE32F98C61108040" box="[471,567,1571,1595]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="18" pageNumber="511">Fig. 15B</figureCitation>
|
|||
|
|
); elytra (
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FD72F98C61CD8040" box="[663,746,1571,1595]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="18" pageNumber="511">Fig. 8A</figureCitation>
|
|||
|
|
) pale yellowish-brown with basal, middle and longitudinal dark bands present and dark reddish brown to black, preapical dark spot absent, basal and middle bands of average thickness, middle transverse dark band narrow, extended laterally onto interval 6, portion on intervals 2 to 4 connected to portion on intervals 5 and 6 by a narrow, oblique dark line; elytral striae moderately impressed, elytral intervals flat or nearly so; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; median lobe of male genitalia with shaft of approximately equal thickness throughout, ventral curvature straightened subapically in lateral aspect (
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FC63FB6560CD8299" box="[902,1002,1226,1250]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="19" pageNumber="512">Fig. 16D</figureCitation>
|
|||
|
|
), apical lamella short and smoothly rounded in dorsal aspect (
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FD80FB4561EE8379" box="[613,713,1258,1282]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="19" pageNumber="512">Fig. 17D</figureCitation>
|
|||
|
|
); specimen not from
|
|||
|
|
<collectingCountry id="ECAF767AEB7BED56FC4BFB4567558379" box="[942,1138,1258,1282]" name="Philippines" pageId="19" pageNumber="512">Philippine Islands</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="C0C76662EB7BED56FF4EFB8261BA8221" ID-DOI="http://doi.org/10.5281/zenodo.11512974" ID-Zenodo-Dep="11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="19" pageNumber="512" startId="19.[171,183,1069,1088]" targetBox="[144,1131,234,1045]" targetPageId="19" targetType="figure">
|
|||
|
|
<paragraph id="940736EAEB7BED56FF4EFB8261BA8221" blockId="19.[123,1173,1069,1114]" pageId="19" pageNumber="512">
|
|||
|
|
<smallCapsWord id="92E1A036EB7BED56FF4EFB8263CF823B" baselines="1083,1084" box="[171,232,1069,1088]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="19" pageNumber="512">FIGURE</smallCapsWord>
|
|||
|
|
8.
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FEE5FB82617B823B" authority="Motschulsky. A." authorityName="Motschulsky. A." box="[256,604,1069,1088]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FEE5FB8262EB823B" box="[256,460,1069,1088]" italics="true" pageId="19" pageNumber="512">Cyclosomus marginatus</emphasis>
|
|||
|
|
Motschulsky.A.
|
|||
|
|
</taxonomicName>
|
|||
|
|
Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="940736EAEB7BED56FF4EFAA560AC8019" blockId="19.[123,1174,1162,1699]" pageId="19" pageNumber="512">
|
|||
|
|
Specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FEB4FAA462D68359" box="[337,497,1291,1314]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FEB4FAA462D68359" box="[337,497,1291,1314]" italics="true" pageId="19" pageNumber="512">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are most similar to those of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FCB3FAA460EE8359" box="[854,969,1291,1314]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FCB3FAA460EE8359" box="[854,969,1291,1314]" italics="true" pageId="19" pageNumber="512">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FC38FAA4677D8359" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[989,1114,1290,1314]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="pallidus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FC38FAA4677D8359" box="[989,1114,1290,1314]" italics="true" pageId="19" pageNumber="512">C. pallidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FF9EFA8462298339" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[123,270,1322,1346]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF9EFA8462298339" box="[123,270,1322,1346]" italics="true" pageId="19" pageNumber="512">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Features distinguishing specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FD5EFA8460718339" box="[699,854,1323,1346]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FD5EFA8460718339" box="[699,854,1323,1346]" italics="true" pageId="19" pageNumber="512">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from those of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FC1BFA84674B8339" box="[1022,1132,1323,1346]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FC1BFA84674B8339" box="[1022,1132,1323,1346]" italics="true" pageId="19" pageNumber="512">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are discussed above in the Diagnosis section for the latter species. The elytra of all known specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FF7CFAC4621483F9" box="[153,307,1387,1410]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF7CFAC4621483F9" box="[153,307,1387,1410]" italics="true" pageId="19" pageNumber="512">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have the middle dark band present and in the form of a compound-W, whereas
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FF9EFA2463D783D9" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[123,240,1418,1442]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="pallidus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF9EFA2463D783D9" box="[123,240,1418,1442]" italics="true" pageId="19" pageNumber="512">C. pallidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
members have the middle band absent. Members of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FCC0FA24609983D9" box="[805,958,1419,1442]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FCC0FA24609983D9" box="[805,958,1419,1442]" italics="true" pageId="19" pageNumber="512">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differ from those of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FF9EFA04622C83B9" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[123,267,1450,1474]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF9EFA04622C83B9" box="[123,267,1450,1474]" italics="true" pageId="19" pageNumber="512">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having the free apex of the prosternal intercoxal process moderate in length, as in
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FF9EFA6563FB8399" box="[123,220,1482,1506]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="19" pageNumber="512">Fig. 15B</figureCitation>
|
|||
|
|
, rather than short, as in
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FE00FA65616E8399" box="[485,585,1482,1506]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="19" pageNumber="512">Fig. 15D</figureCitation>
|
|||
|
|
. Most members of these two species also differ in the development of the middle dark band. In the all three known specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FC73FA4467098079" box="[918,1070,1515,1538]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FC73FA4467098079" box="[918,1070,1515,1538]" italics="true" pageId="19" pageNumber="512">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the band is relatively thin and in the compound-W form (
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FD70F9A561C88059" box="[661,751,1546,1570]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="19" pageNumber="512">Fig. 8A</figureCitation>
|
|||
|
|
). In contrast, the middle dark band in
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FF9EF984622C8039" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[123,267,1578,1602]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF9EF984622C8039" box="[123,267,1578,1602]" italics="true" pageId="19" pageNumber="512">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is similarly shaped but distinctly thicker (
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FD2FF98560488039" box="[714,879,1578,1602]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="19" pageNumber="512">Figs. 13A, 13B</figureCitation>
|
|||
|
|
) in most specimens, as thin as in
|
|||
|
|
<taxonomicName id="53B84D69EB7BED56FF51F9E4626A8019" box="[180,333,1611,1634]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="512" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF51F9E4626A8019" box="[180,333,1611,1634]" italics="true" pageId="19" pageNumber="512">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in only
|
|||
|
|
<specimenCount id="82BEFD63EB7BED56FE42F9E461628019" box="[423,581,1610,1634]" count="2" pageId="19" pageNumber="512" type="generic">two specimens</specimenCount>
|
|||
|
|
that we have seen (
|
|||
|
|
<figureCitation id="0C832A6FEB7BED56FCFEF9E560598019" box="[795,894,1610,1634]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="19" pageNumber="512">Fig. 13D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB7BED56FF4EF9C460F880D8" pageId="19" pageNumber="512" type="biology_ecology">
|
|||
|
|
<paragraph id="940736EAEB7BED56FF4EF9C460F880D8" blockId="19.[123,1174,1162,1699]" pageId="19" pageNumber="512">
|
|||
|
|
<emphasis id="A6CCEAF8EB7BED56FF4EF9C462B080F9" bold="true" box="[171,407,1643,1666]" pageId="19" pageNumber="512">Habitat distribution.</emphasis>
|
|||
|
|
Unknown, but presumed to be restricted to the sandy shores of medium to large rivers at low elevation, like members of most other species of the genus.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB7CED51FF4EFF7363E884AE" pageId="20" pageNumber="513" type="materials_examined">
|
|||
|
|
<paragraph id="940736EAEB7CED51FF4EFF7362FE8728" blockId="20.[123,1174,219,1046]" pageId="20" pageNumber="513">
|
|||
|
|
<emphasis id="A6CCEAF8EB7CED51FF4EFF7362FD8688" bold="true" box="[171,474,220,243]" pageId="20" pageNumber="513">Geographical distribution.</emphasis>
|
|||
|
|
<location id="91676031EB7CED51FE05FF7461248689" LSID="urn:lsid:plazi:treatment:1C1187FCEB7AED51FF9EFC0C6743826D:91676031EB7CED51FE05FF7461248689" box="[480,515,219,242]" name="We" pageId="20" pageNumber="513">We</location>
|
|||
|
|
have examined a total of
|
|||
|
|
<specimenCount id="82BEFD63EB7CED51FCFEFF7460E18688" box="[795,966,219,243]" count="3" pageId="20" pageNumber="513" type="generic">three specimens</specimenCount>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="82BEFD63EB7CED51FC30FF74671F8688" box="[981,1080,219,243]" count="1" pageId="20" pageNumber="513" type="male">one male</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB7CED51FB88FF7463F78768" count="2" pageId="20" pageNumber="513" type="female">two females</specimenCount>
|
|||
|
|
) from the following localities:
|
|||
|
|
<materialsCitation id="24D03CB7EB7CED51FDC3FF5362FE8728" ID-GBIF-Occurrence="4905152350" collectingDate="1809-05" collectionCode="ZMUC, ZMMU, NHMUK" collectorName="India Or. & N India" country="Bangladesh" location="Eastern India" municipality="Northern India" pageId="20" pageNumber="513" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
|
|||
|
|
<emphasis id="A6CCEAF8EB7CED51FDC3FF5360F88768" bold="true" box="[550,991,252,275]" pageId="20" pageNumber="513">
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FDC3FF5361F88768" box="[550,735,252,275]" name="Bangladesh" pageId="20" pageNumber="513">BANGLADESH</collectingCountry>
|
|||
|
|
/
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FD14FF5360198768" box="[753,830,252,275]" name="India" pageId="20" pageNumber="513">INDIA</collectingCountry>
|
|||
|
|
/
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FCAAFF5360F88768" box="[847,991,252,275]" name="Myanmar" pageId="20" pageNumber="513">MYANMAR</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
: “Bengal” (
|
|||
|
|
<date id="E006102AEB7CED51FB86FF5463978748" pageId="20" pageNumber="513" value="1809-05">
|
|||
|
|
<collectingDate id="F042E9C2EB7CED51FB86FF5463978748" pageId="20" pageNumber="513" value="1809-05">May 1809</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
[
|
|||
|
|
<specimenCount id="82BEFD63EB7CED51FF5AFEB362008748" box="[191,295,283,307]" count="1" pageId="20" pageNumber="513" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB7CED51FED1FEB462AF8748" box="[308,392,283,307]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="513">ZMUC</collectionCode>
|
|||
|
|
]).
|
|||
|
|
<location id="91676031EB7CED51FE44FEB361168748" LSID="urn:lsid:plazi:treatment:1C1187FCEB7AED51FF9EFC0C6743826D:91676031EB7CED51FE44FEB361168748" box="[417,561,283,307]" country="Bangladesh" municipality="Northern India" name="Eastern India" pageId="20" pageNumber="513">Eastern India</location>
|
|||
|
|
(“
|
|||
|
|
<collectorName id="394D533CEB7CED51FDAFFEB461968748" box="[586,689,283,307]" pageId="20" pageNumber="513">India Or.</collectorName>
|
|||
|
|
”) [
|
|||
|
|
<specimenCount id="82BEFD63EB7CED51FD34FEB360688748" box="[721,847,283,307]" count="1" pageId="20" pageNumber="513" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB7CED51FCBEFEB360938748" box="[859,948,284,307]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="20" pageNumber="513" type="Museum">ZMMU</collectionCode>
|
|||
|
|
].
|
|||
|
|
<collectingMunicipality id="7463AC90EB7CED51FC23FEB467428748" box="[966,1125,283,307]" pageId="20" pageNumber="513">Northern India</collectingMunicipality>
|
|||
|
|
(“
|
|||
|
|
<collectorName id="394D533CEB7CED51FB65FEB463938728" pageId="20" pageNumber="513">N India</collectorName>
|
|||
|
|
” ([
|
|||
|
|
<specimenCount id="82BEFD63EB7CED51FF36FE9462778728" box="[211,336,315,339]" count="1" pageId="20" pageNumber="513" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB7CED51FEB9FE9462EF8728" box="[348,456,315,339]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="20" pageNumber="513" type="Museum">NHMUK</collectionCode>
|
|||
|
|
]).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB7CED51FF4EFEF463E884AE" blockId="20.[123,1174,219,1046]" pageId="20" pageNumber="513">
|
|||
|
|
The
|
|||
|
|
<typeStatus id="4B038848EB7CED51FF39FEF462198708" box="[220,318,347,371]" pageId="20" pageNumber="513" type="lectotype">lectotype</typeStatus>
|
|||
|
|
specimen from ‘eastern India’ could have been collected anywhere in present-day
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FF4CFED463C587E8" box="[169,226,379,403]" name="India" pageId="20" pageNumber="513">India</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FF0BFED4624A87E8" box="[238,365,379,403]" name="Bangladesh" pageId="20" pageNumber="513">Bangladesh</collectingCountry>
|
|||
|
|
, or
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FE72FED3612787EF" box="[407,512,380,404]" name="Myanmar" pageId="20" pageNumber="513">Myanmar</collectingCountry>
|
|||
|
|
, the specimen from “Bengal” from anywhere in
|
|||
|
|
<collectingRegion id="567CF808EB7CED51FBEBFED367B187E8" box="[1038,1174,379,403]" country="India" name="West Bengal" pageId="20" pageNumber="513">West Bengal</collectingRegion>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FF67FE34639B87C8" box="[130,188,411,435]" name="India" pageId="20" pageNumber="513">India</collectingCountry>
|
|||
|
|
) or
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FF09FE34624C87C8" box="[236,363,411,435]" name="Bangladesh" pageId="20" pageNumber="513">Bangladesh</collectingCountry>
|
|||
|
|
, and the specimen from ‘northern India’ from anywhere from
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FBF8FE34675087C8" box="[1053,1143,411,435]" name="Pakistan" pageId="20" pageNumber="513">Pakistan</collectingCountry>
|
|||
|
|
to
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FF9EFE1363C487AF" box="[123,227,444,468]" name="Myanmar" pageId="20" pageNumber="513">Myanmar</collectingCountry>
|
|||
|
|
along the southern base of the Himalaya. Consequently, the range of this species remains undefined (
|
|||
|
|
<figureCitation id="0C832A6FEB7CED51FF1DFE73626E878F" box="[248,329,476,500]" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="20" pageNumber="513">Fig. 21</figureCitation>
|
|||
|
|
) and problematic. The record of its occurrence in “Indo-China” (
|
|||
|
|
<bibRefCitation id="F0294B1BEB7CED51FBCDFE736396846F" author="ANDREWES, H. E." pageId="20" pageNumber="513" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes 1921</bibRefCitation>
|
|||
|
|
) probably is based on a misidentification of specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB7CED51FCFFFE5260A3846F" box="[794,900,509,532]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7CED51FCFFFE5260A3846F" box="[794,900,509,532]" italics="true" pageId="20" pageNumber="513">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which species Andrewes subsequently described (
|
|||
|
|
<bibRefCitation id="F0294B1BEB7CED51FE60FDB36114844F" author="ANDREWES, H. E." box="[389,563,540,564]" pageId="20" pageNumber="513" pagination="459 - 468" refId="ref23019" refString="ANDREWES, H. E. 1924. XXII. On the Oriental Carabidae of the " Reise Novara. " Transactions of the Entomological Society of London 1923: 459 - 468." type="journal article" year="1924">Andrewes 1924</bibRefCitation>
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) as distinct (see above). Some other records from
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<collectingCountry id="ECAF767AEB7CED51FBBAFDB367B1844F" box="[1119,1174,540,564]" name="India" pageId="20" pageNumber="513">India</collectingCountry>
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may have been based on misidentified specimens of
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<taxonomicName id="53B84D69EB7CED51FD5AFD926076842F" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[703,849,572,596]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="vespertilio">
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<emphasis id="A6CCEAF8EB7CED51FD5AFD926076842F" box="[703,849,572,596]" italics="true" pageId="20" pageNumber="513">C. vespertilio</emphasis>
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</taxonomicName>
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, which is described below. If
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<taxonomicName id="53B84D69EB7CED51FF9EFDF263C1840F" box="[123,230,605,628]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="inustus">
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<emphasis id="A6CCEAF8EB7CED51FF9EFDF263C1840F" box="[123,230,605,628]" italics="true" pageId="20" pageNumber="513">C. inustus</emphasis>
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</taxonomicName>
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is actually restricted to sandy sea coast areas everywhere it occurs as it is in
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<collectingCountry id="ECAF767AEB7CED51FBFFFDF3674F840F" box="[1050,1128,604,628]" name="Taiwan" pageId="20" pageNumber="513">Taiwan</collectingCountry>
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and apparently in southeast coastal
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<collectingCountry id="ECAF767AEB7CED51FE2CFDD3612C84EF" box="[457,523,636,660]" name="China" pageId="20" pageNumber="513">China</collectingCountry>
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, then records from inland parts of Indochina (i.e., in
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<collectingCountry id="ECAF767AEB7CED51FBDAFDD363B984CE" name="Cambodia" pageId="20" pageNumber="513">Cambodia</collectingCountry>
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,
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<collectingCountry id="ECAF767AEB7CED51FF4BFD3263C384CF" box="[174,228,669,692]" name="Laos" pageId="20" pageNumber="513">Laos</collectingCountry>
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, and
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<collectingCountry id="ECAF767AEB7CED51FEC3FD3262A084CE" box="[294,391,669,693]" name="Thailand" pageId="20" pageNumber="513">Thailand</collectingCountry>
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) may, in fact, represent either
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<taxonomicName id="53B84D69EB7CED51FD0CFD3260A184CE" box="[745,902,669,693]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="marginatus">
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<emphasis id="A6CCEAF8EB7CED51FD0CFD3260A184CE" box="[745,902,669,693]" italics="true" pageId="20" pageNumber="513">C. marginatus</emphasis>
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</taxonomicName>
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or another, undescribed species.
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB7CED51FF4EFD716743826D" pageId="20" pageNumber="513" type="description">
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<paragraph id="940736EAEB7CED51FF4EFD71611C852E" blockId="20.[123,1174,219,1046]" pageId="20" pageNumber="513">
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<emphasis id="A6CCEAF8EB7CED51FF4EFD71629B848E" bold="true" box="[171,444,734,757]" pageId="20" pageNumber="513">Geographical variation.</emphasis>
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There is insufficient material available at present to recognize any possible geographical variation in this species, unless the specimens from inland part of Indochina are considered as conspecific with
|
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<taxonomicName id="53B84D69EB7CED51FE14FCB161AD854E" box="[497,650,798,821]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="marginatus">
|
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<emphasis id="A6CCEAF8EB7CED51FE14FCB161AD854E" box="[497,650,798,821]" italics="true" pageId="20" pageNumber="513">C. marginatus</emphasis>
|
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</taxonomicName>
|
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rather than
|
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<taxonomicName id="53B84D69EB7CED51FCEEFCB16051854E" box="[779,886,798,821]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="inustus">
|
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|
<emphasis id="A6CCEAF8EB7CED51FCEEFCB16051854E" box="[779,886,798,821]" italics="true" pageId="20" pageNumber="513">C. inustus</emphasis>
|
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</taxonomicName>
|
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|
|
(see comments about geographical variation in that species above).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB7CED51FF4EFCF16743826D" blockId="20.[123,1174,219,1046]" pageId="20" pageNumber="513">
|
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|
|
<emphasis id="A6CCEAF8EB7CED51FF4EFCF1606E850E" bold="true" box="[171,841,862,886]" pageId="20" pageNumber="513">
|
|||
|
|
Geographical relationships with other
|
|||
|
|
<taxonomicName id="53B84D69EB7CED51FD82FCF1606E850E" box="[615,841,862,886]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="undetermined">
|
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|
|
<emphasis id="A6CCEAF8EB7CED51FD82FCF161CC850D" bold="true" box="[615,747,862,886]" italics="true" pageId="20" pageNumber="513">Cyclosomus</emphasis>
|
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|
|
species.
|
|||
|
|
</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
In the absence of any specific locality records for this species, its syntopy or even sympatry with any other congeners cannot be confirmed.
|
|||
|
|
<taxonomicName id="53B84D69EB7CED51FF10FC3162FD85CD" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[245,474,926,950]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="suturalis">
|
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|
|
<emphasis id="A6CCEAF8EB7CED51FF10FC3162FD85CD" box="[245,474,926,950]" italics="true" pageId="20" pageNumber="513">Cyclosomus suturalis</emphasis>
|
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|
</taxonomicName>
|
|||
|
|
is also known from “Bengala”, again without specific locality, so it is possible that these species co-occur. Also, eastern
|
|||
|
|
<collectingCountry id="ECAF767AEB7CED51FD5AFC1161D185AD" box="[703,758,958,982]" name="India" pageId="20" pageNumber="513">India</collectingCountry>
|
|||
|
|
is within the broad geographical range of
|
|||
|
|
<taxonomicName id="53B84D69EB7CED51FF7DFC70623D858D" authorityName=", Kavanaugh" authorityYear="2015" box="[152,282,990,1014]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7CED51FF7DFC70623D858D" box="[152,282,990,1014]" italics="true" pageId="20" pageNumber="513">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(recorded from northernmost
|
|||
|
|
<collectingRegion id="567CF808EB7CED51FDBBFC7061C0858D" box="[606,743,990,1014]" country="India" name="West Bengal" pageId="20" pageNumber="513">West Bengal</collectingRegion>
|
|||
|
|
) and at the western limit of the range of
|
|||
|
|
<taxonomicName id="53B84D69EB7CED51FF9EFC5063C1826D" box="[123,230,1023,1046]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="513" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7CED51FF9EFC5063C1826D" box="[123,230,1023,1046]" italics="true" pageId="20" pageNumber="513">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, so there is at least potential for co-occurrence with either or both of these species.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|