333 lines
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333 lines
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<mods:title id="BA64A2E584F60A47CC98F09F8758A8CE">Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary</mods:title>
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<mods:namePart id="EB0860E6010D0DA6354DCB7F7CF14FDC">Augier, Dominique</mods:namePart>
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<mods:namePart id="F21CAE1D048A85C10D5C89FEEEB80E02">Bailly, Gilles</mods:namePart>
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<mods:title id="EE9CC4AE33121A21522FD6B6C23709E1">Geodiversitas</mods:title>
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<mods:date id="EBAF97E96CBB5885DBC7735FB6A04A9D">2022</mods:date>
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<mods:number id="5ABBDD6200F9358C15D7D1968A05780D">2022-07-21</mods:number>
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<mods:number id="057D293674F367DCE4743BE6F61D0834">44</mods:number>
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<mods:number id="98C690718DB152EE40A8577CD9AFDA23">25</mods:number>
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<mods:identifier id="0EF8A47B4237C1325DC069B2E13D58CA" type="DOI">10.5252/geodiversitas2022v44a25</mods:identifier>
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<treatment id="03C887B9FFB8FFE57405A06EFABAFABC" ID-DOI="http://doi.org/10.5281/zenodo.6929025" ID-GBIF-Taxon="197866737" ID-Zenodo-Dep="6929025" LSID="urn:lsid:plazi:treatment:03C887B9FFB8FFE57405A06EFABAFABC" httpUri="http://treatment.plazi.org/id/03C887B9FFB8FFE57405A06EFABAFABC" lastPageNumber="695" pageId="14" pageNumber="695">
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<subSubSection id="C37B6524FFB8FFE57405A06EFDA6FDC1" box="[319,589,598,624]" pageId="14" pageNumber="695" type="nomenclature">
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<paragraph id="8BDE36AFFFB8FFE57405A06EFDA6FDC1" blockId="14.[319,589,598,656]" box="[319,589,598,624]" pageId="14" pageNumber="695">
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<taxonomicName id="4C614D2CFFB8FFE57405A06EFDEDFDC1" authorityName="Fox & Naylor" authorityYear="1982" box="[319,518,598,624]" class="Amphibia" family="Albanerpetontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="family">Albanerpetontidae</taxonomicName>
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indet.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6524FFB8FFE574A5A04EFABAFABC" pageId="14" pageNumber="695" type="description">
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<paragraph id="8BDE36AFFFB8FFE574A5A04EFE06FD21" blockId="14.[319,589,598,656]" box="[415,493,630,656]" pageId="14" pageNumber="695">
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(
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<figureCitation id="135A2A2AFFB8FFE5749DA04EFE0FFD21" box="[423,484,630,656]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9</figureCitation>
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)
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</paragraph>
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<paragraph id="8BDE36AFFFB8FFE575BEA08EFEFEFD61" blockId="14.[130,777,694,2028]" box="[132,277,694,720]" pageId="14" pageNumber="695">DESCRIPTION</paragraph>
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<paragraph id="8BDE36AFFFB8FFE575BEA0EDFD81FA9F" blockId="14.[130,777,694,2028]" pageId="14" pageNumber="695">
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Albanerpetontids are represented in Angeac-Charente by numerous and diverse bones, including dentaries, premaxillae, maxillae, vertebrae and forelimb bones (
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<figureCitation id="135A2A2AFFB8FFE5775EA12DFD48FC9E" box="[612,675,789,815]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9</figureCitation>
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A-K). All the material was recovered from screen washing residues. It is always disarticulated, and almost always fragmentary. Several diagnostic characters allow their assignment to the
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<taxonomicName id="4C614D2CFFB8FFE577A9A14DFF37FC1E" authorityName="Fox & Naylor" authorityYear="1982" class="Amphibia" family="Albanerpetontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="family">Albanerpetontidae</taxonomicName>
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, including: 1) intertonguing symphyseal joint between dentaries, in a mortise-and-tenon style (
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<bibRefCitation id="EFF04B5EFFB8FFE57716A18DFCEBFC7E" author="Fox & Naylor" box="[556,768,949,975]" firstAuthor="Fox" pageId="14" pageNumber="695" pagination="118 - 128" refId="ref39749" refString="FOX R. C. & NAYLOR M. 1982. - A reconsideration of the relationships of the fossil amphibian Albanerpeton. Canadian Journal of Earth Sciences 19: 118 - 128. https: // doi. org / 10.1139 / e 82 - 009" type="journal article" year="1982">Fox & Naylor 1982</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575BEA1ECFEFBFC5E" author="Milner" box="[132,272,980,1007]" firstAuthor="Milner" pageId="14" pageNumber="695" refId="ref44242" refString="MILNER A. R. 1988. - The relationships and origin of living amphibians, in BENTON M. J. (ed.), The Phylogeny and Classification of the Tetrapods. Volume 1. Amphibians, Reptiles, Birds. Clarendon Press, Oxford, 380 p." type="book" year="1988">Milner 1988</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE5741AA1EDFDD0FC5E" author="McGowan & Evans" box="[288,571,980,1007]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="143 - 145" refId="ref44053" refString="MCGOWAN G. J. & EVANS S. E. 1995. - Albanerpetontid ampibians from the Cretaceous of Spain. Nature 373: 143 - 145. https: // doi. org / 10.1038 / 373143 a 0" type="journal article" year="1995">McGowan & Evans 1995</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE57771A1EDFD14FC5F" author="McGowan" box="[587,767,980,1007]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="227 - 234" refId="ref43887" refString="MCGOWAN G. J. 1996. - Albanerpetontid amphibians from the Jurassic (Bathonian) of southern England. Museum of Northern Arizona Bulletin 60: 227 - 234." type="journal article" year="1996">McGowan 1996</bibRefCitation>
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,
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<bibRefCitation id="EFF04B5EFFB8FFE575BEA1CDFF56FBBE" author="McGowan" box="[132,189,1013,1039]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="1 - 32" refId="ref43957" refString="MCGOWAN G. J. 2002. - Albanerpetontid amphibians from the Lower Cretaceous of Spain and Italy: a description and reconsideration of their systematics. Zoological Journal of the Linnean Society 135: 1 - 32. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.00013. x" type="journal article" year="2002">2002</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575F7A1CCFE92FBBF" author="Gardner" box="[205,377,1012,1038]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="529 - 544" refId="ref40256" refString="GARDNER J. D. 1999 b. - The amphibian Albanerpeton arthridion and the Aptian - Albian biogeography of albanerpetontids. Palaeontology 42: 529 - 544. https: // doi. org / 10.1111 / 1475 - 4983.00083" type="journal article" year="1999" yearSuffix="b">Gardner 1999b</bibRefCitation>
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,
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<bibRefCitation id="EFF04B5EFFB8FFE574B3A1CDFE29FBBE" author="Gardner" box="[393,450,1013,1039]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">2000</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE574E8A1CCFD44FBBE" author="Gardner" box="[466,687,1012,1039]" etAl="et al." firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="301 - 319" refId="ref40323" refString="GARDNER J. D., EVANS S. E. & SIGOGNEAU- RUSSELL D. 2003. - New albanerpetontid amphibians from the Early Cretaceous of Morocco and Middle Jurassic of England. Acta Palaeontologica Polonica 48 (2): 301 - 319." type="journal article" year="2003">
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Gardner
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<emphasis id="B915EABDFFB8FFE5770DA1CDFD86FBBF" box="[567,621,1012,1038]" italics="true" pageId="14" pageNumber="695">et al.</emphasis>
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2003
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</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE57785A1CDFE9CFB9F" author="SWEETMAN S. C. & GARDNER J. D." journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE574BDA62DFD47FB9F" author="Matsumoto & Evans" box="[391,684,1044,1071]" firstAuthor="Matsumoto" pageId="14" pageNumber="695" pagination="0189767" refId="ref43724" refString="MATSUMOTO R. & EVANS S. E. 2018. - The first record of albanerpetontid amphibians (Amphibia: Albanerpetontidae) from East Asia. PLoS ONE 13: e 0189767. https: // doi. org / 10.1371 / journal. pone. 0189767" type="journal article" year="2018">Matsumoto & Evans 2018</bibRefCitation>
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). In the Angeac-Charente specimens, there are two symphyseal prongs (
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<figureCitation id="135A2A2AFFB8FFE575B7A66CFF11FBDE" box="[141,250,1108,1135]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9A, B</figureCitation>
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); 2) pleurodont, chisel-like and regularly arranged non-pedicellate teeth, labiolingually compressed and bearing three faint, mesiodistally aligned cuspules (
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<figureCitation id="135A2A2AFFB8FFE57773A6ACFD6DFB1F" box="[585,646,1172,1198]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9</figureCitation>
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A-G;
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<bibRefCitation id="EFF04B5EFFB8FFE577F8A6ACFEE6FB7F" author="Fox & Naylor" firstAuthor="Fox" pageId="14" pageNumber="695" pagination="118 - 128" refId="ref39749" refString="FOX R. C. & NAYLOR M. 1982. - A reconsideration of the relationships of the fossil amphibian Albanerpeton. Canadian Journal of Earth Sciences 19: 118 - 128. https: // doi. org / 10.1139 / e 82 - 009" type="journal article" year="1982">Fox & Naylor 1982</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE57426A68CFE54FB7F" author="Gardner" box="[284,447,1204,1230]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="57 - 84" refId="ref40194" refString="GARDNER J. D. 1999 a. - Redescription of the geologically youngest albanerpetontid (? Lissamphibia): Albanerpeton inexpectatum Estes and Hoffstetter, 1976, from the Miocene of France. Annales de Paleontologie 85: 57 - 84. https: // doi. org / 10.1016 / S 0753 - 3969 (99) 80008 - 1" type="journal article" year="1999" yearSuffix="a">Gardner 1999a</bibRefCitation>
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, b,
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<bibRefCitation id="EFF04B5EFFB8FFE574D1A68CFDC9FB7F" author="GARDNER J. D." box="[491,546,1204,1230]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="55 - 70" part="45" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." title="Revised taxonomy of albanerpetontid amphibians" type="journal article" year="2000">2000</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE5770BA68CFF51FB5F" author="McGowan & Evans" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="143 - 145" refId="ref44053" refString="MCGOWAN G. J. & EVANS S. E. 1995. - Albanerpetontid ampibians from the Cretaceous of Spain. Nature 373: 143 - 145. https: // doi. org / 10.1038 / 373143 a 0" type="journal article" year="1995">McGowan & Evans 1995</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575FFA6ECFE34FB5F" author="SWEETMAN S. C. & GARDNER J. D." box="[197,479,1236,1262]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE574D3A6ECFD13FB5F" author="Matsumoto & Evans" box="[489,760,1236,1262]" firstAuthor="Matsumoto" pageId="14" pageNumber="695" pagination="0189767" refId="ref43724" refString="MATSUMOTO R. & EVANS S. E. 2018. - The first record of albanerpetontid amphibians (Amphibia: Albanerpetontidae) from East Asia. PLoS ONE 13: e 0189767. https: // doi. org / 10.1371 / journal. pone. 0189767" type="journal article" year="2018">Matsumoto & Evans 2018</bibRefCitation>
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); and 3) foramina arranged in line on external face of dentary (
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<figureCitation id="135A2A2AFFB8FFE575B7A72BFF31FA9F" box="[141,218,1299,1326]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9F</figureCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575DFA72CFE5CFA9F" author="Fox & Naylor" box="[229,439,1299,1326]" firstAuthor="Fox" pageId="14" pageNumber="695" pagination="118 - 128" refId="ref39749" refString="FOX R. C. & NAYLOR M. 1982. - A reconsideration of the relationships of the fossil amphibian Albanerpeton. Canadian Journal of Earth Sciences 19: 118 - 128. https: // doi. org / 10.1139 / e 82 - 009" type="journal article" year="1982">Fox & Naylor 1982</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE574FEA72BFDB1FA9F" author="Gardner" box="[452,602,1299,1326]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">Gardner 2000</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BDE36AFFFB8FFE575A1A70CFEE6F8FA" blockId="14.[130,777,694,2028]" pageId="14" pageNumber="695">
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In Angeac-Charente albanerpetontids, the maxilla is unornamented labially, except for scattered external nutritive foramina that are characteristic of the group (
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<figureCitation id="135A2A2AFFB8FFE574D9A74BFDD9FA3F" box="[483,562,1395,1422]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig.9D</figureCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE57701A74CFCEBFA3C" author="Fox & Naylor" box="[571,768,1395,1422]" firstAuthor="Fox" pageId="14" pageNumber="695" pagination="118 - 128" refId="ref39749" refString="FOX R. C. & NAYLOR M. 1982. - A reconsideration of the relationships of the fossil amphibian Albanerpeton. Canadian Journal of Earth Sciences 19: 118 - 128. https: // doi. org / 10.1139 / e 82 - 009" type="journal article" year="1982">Fox & Naylor 1982</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575BEA7ABFEF0FA1C" author="Gardner" box="[132,283,1427,1453]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">Gardner 2000</bibRefCitation>
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). In this respect, they differ from
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<taxonomicName id="4C614D2CFFB8FFE577BBA7ABFEE0FA7C" baseAuthorityName="Estes & Hoffstetter" baseAuthorityYear="1976" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="species" species="inexpectatum">
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<emphasis id="B915EABDFFB8FFE577BBA7ABFEE0FA7C" italics="true" pageId="14" pageNumber="695">Albanerpeton inexpectatum</emphasis>
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</taxonomicName>
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from the Miocene of
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<collectingCountry id="F376763FFFB8FFE574C6A78BFDA9FA7C" box="[508,578,1459,1485]" name="France" pageId="14" pageNumber="695">France</collectingCountry>
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(
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<bibRefCitation id="EFF04B5EFFB8FFE5776EA78BFD1CFA7C" author="Gardner" box="[596,759,1459,1485]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="57 - 84" refId="ref40194" refString="GARDNER J. D. 1999 a. - Redescription of the geologically youngest albanerpetontid (? Lissamphibia): Albanerpeton inexpectatum Estes and Hoffstetter, 1976, from the Miocene of France. Annales de Paleontologie 85: 57 - 84. https: // doi. org / 10.1016 / S 0753 - 3969 (99) 80008 - 1" type="journal article" year="1999" yearSuffix="a">Gardner 1999a</bibRefCitation>
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). As in other albanerpetontid for which trunk vertebrae have been described (
|
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<bibRefCitation id="EFF04B5EFFB8FFE57408A7CBFDAAF9BC" author="Estes & Hoffstetter" box="[306,577,1523,1549]" firstAuthor="Estes" pageId="14" pageNumber="695" pagination="297 - 343" refId="ref39143" refString="ESTES R. & HOFFSTETTER R. 1976. - Les urodeles du Miocene de La Grive-Saint-Alban (Isere, France). Bulletins du Museum national d'Histoire naturelle, 3 eme serie, Sciences de la Terre 398: 297 - 343. https: // www. biodiversitylibrary. org / page / 55505209" type="journal article" year="1976">Estes & Hoffstetter 1976</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE5776AA7CBFCEBF9BC" author="McGowan" box="[592,768,1523,1549]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="227 - 234" refId="ref43887" refString="MCGOWAN G. J. 1996. - Albanerpetontid amphibians from the Jurassic (Bathonian) of southern England. Museum of Northern Arizona Bulletin 60: 227 - 234." type="journal article" year="1996">McGowan 1996</bibRefCitation>
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,
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<bibRefCitation id="EFF04B5EFFB8FFE575BEA42BFF51F99C" author="McGowan" box="[132,186,1555,1581]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="1 - 32" refId="ref43957" refString="MCGOWAN G. J. 2002. - Albanerpetontid amphibians from the Lower Cretaceous of Spain and Italy: a description and reconsideration of their systematics. Zoological Journal of the Linnean Society 135: 1 - 32. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.00013. x" type="journal article" year="2002">2002</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575FCA42BFE32F99C" author="McGowan & Ensom" box="[198,473,1555,1581]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="113 - 117" refId="ref44012" refString="MCGOWAN G. & ENSOM P. C. 1997. - Albanerpetontid amphibians from the Lower Cretaceous of the Isle of Purbeck, Dorset. Proceedings of the Dorset Natural History and Archaeological Society 118: 113 - 117." type="journal article" year="1997">McGowan & Ensom 1997</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE574DEA42BFCEBF99C" author="SWEETMAN S. C. & GARDNER J. D." box="[484,768,1554,1581]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFB8FFE575BEA40BFE76F9FC" author="Matsumoto & Evans" box="[132,413,1587,1613]" firstAuthor="Matsumoto" pageId="14" pageNumber="695" pagination="0189767" refId="ref43724" refString="MATSUMOTO R. & EVANS S. E. 2018. - The first record of albanerpetontid amphibians (Amphibia: Albanerpetontidae) from East Asia. PLoS ONE 13: e 0189767. https: // doi. org / 10.1371 / journal. pone. 0189767" type="journal article" year="2018">Matsumoto & Evans 2018</bibRefCitation>
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), those from the Angeac-Charente material (
|
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<figureCitation id="135A2A2AFFB8FFE575D0A46AFEC3F9DD" box="[234,296,1618,1644]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9</figureCitation>
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I-K) are amphicoelous, hourglass-shaped and bear a short unicipital transverse process. The centrum is narrowly constricted at its center (
|
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<figureCitation id="135A2A2AFFB8FFE574F6A4AAFDDFF91C" box="[460,564,1682,1709]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9I, K</figureCitation>
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) and the cotyles are circular in outline and have thickened rims (
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|
<figureCitation id="135A2A2AFFB8FFE57757A48AFD52F97C" box="[621,697,1714,1741]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9J</figureCitation>
|
||
|
|
). As in other albanerpetontid trunk vertebrae described elsewhere, the notochordal canal is anteroposteriorly continuous (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE57786A4CAFCE9F8BD" box="[700,770,1778,1804]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9J</figureCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE575BEA52AFE46F89D" author="SWEETMAN S. C. & GARDNER J. D." box="[132,429,1810,1836]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
|
||
|
|
) and thus the vertebrae are fully notochordal.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BDE36AFFFB8FFE575A1A569FA45FD21" blockId="14.[130,777,694,2028]" lastBlockId="14.[810,1456,215,1294]" pageId="14" pageNumber="695">
|
||
|
|
Angeac-Charente specimens also show numerous features that have been described in other albanerpetontids and allow the distinction from other lissamphibians and lizards.The premaxilla (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE575B7A589FF12F87D" box="[141,249,1969,1996]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig.9D, E</figureCitation>
|
||
|
|
) and maxilla (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE574B1A589FE32F87A" box="[395,473,1969,1995]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig.9C</figureCitation>
|
||
|
|
) have a deep
|
||
|
|
<emphasis id="B915EABDFFB8FFE57764A58AFD32F87A" box="[606,729,1969,1995]" italics="true" pageId="14" pageNumber="695">pars dentalis</emphasis>
|
||
|
|
and the dentary (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE57432A5E9FE4DF85A" box="[264,422,2001,2028]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9A, B, F, G</figureCitation>
|
||
|
|
) has a tall dental parapet, allowing the attachment of highly pleurodont teeth (
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE5703CA2EFFA74FF40" author="Gardner" box="[1286,1439,215,241]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">Gardner 2000</bibRefCitation>
|
||
|
|
). Upper jaws are also characterized by a prominent, shelf-like
|
||
|
|
<emphasis id="B915EABDFFB8FFE57614A320FC2CFE80" box="[814,967,279,305]" italics="true" pageId="14" pageNumber="695">pars palatinum</emphasis>
|
||
|
|
lingually (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE57109A32FFB69FE83" box="[1075,1154,279,306]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9E</figureCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE571B1A32FFACBFE80" author="Gardner" box="[1163,1312,279,305]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">Gardner 2000</bibRefCitation>
|
||
|
|
). The maxilla has a low, posteriorly tapered
|
||
|
|
<emphasis id="B915EABDFFB8FFE57155A300FB0DFEE0" box="[1135,1254,311,337]" italics="true" pageId="14" pageNumber="695">pars facialis</emphasis>
|
||
|
|
(
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE571CCA30FFAA2FEE0" box="[1270,1353,311,337]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9C</figureCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE5706CA30FFC84FEC0" author="Gardner" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="57 - 84" refId="ref40194" refString="GARDNER J. D. 1999 a. - Redescription of the geologically youngest albanerpetontid (? Lissamphibia): Albanerpeton inexpectatum Estes and Hoffstetter, 1976, from the Miocene of France. Annales de Paleontologie 85: 57 - 84. https: // doi. org / 10.1016 / S 0753 - 3969 (99) 80008 - 1" type="journal article" year="1999" yearSuffix="a">Gardner 1999a</bibRefCitation>
|
||
|
|
,
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE57647A36FFC5FFEC0" author="Gardner" box="[893,948,343,369]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">2000</bibRefCitation>
|
||
|
|
). On the dentary, the Meckelian canal is closed anteriorly, and the subdental shelf is low, narrow and gutterlike anteriorly (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE576F7A3AEFBF6FE00" box="[973,1053,406,433]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9A</figureCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE5711DA3AEFB23FE00" author="Gardner" box="[1063,1224,406,433]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="57 - 84" refId="ref40194" refString="GARDNER J. D. 1999 a. - Redescription of the geologically youngest albanerpetontid (? Lissamphibia): Albanerpeton inexpectatum Estes and Hoffstetter, 1976, from the Miocene of France. Annales de Paleontologie 85: 57 - 84. https: // doi. org / 10.1016 / S 0753 - 3969 (99) 80008 - 1" type="journal article" year="1999" yearSuffix="a">Gardner 1999a</bibRefCitation>
|
||
|
|
, b,
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE571D5A3AFFACDFE00" author="GARDNER J. D." box="[1263,1318,407,433]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="55 - 70" part="45" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." title="Revised taxonomy of albanerpetontid amphibians" type="journal article" year="2000">2000</bibRefCitation>
|
||
|
|
). As in other known albanerpetontid species, the humeral condyle is spherical, fully ossified and larger than the adjacent radial epicondyle (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE5760FA3CEFC6DFDA0" box="[821,902,502,529]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9H</figureCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE576AAA3CEFB40FDA1" author="SWEETMAN S. C. & GARDNER J. D." box="[912,1195,502,528]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
|
||
|
|
). Above this humeral ball is a triangular and well-defined cubital fossa, at the proximal extremity of which a small foramen can be seen (
|
||
|
|
<figureCitation id="135A2A2AFFB8FFE57008A00EFA6DFDE0" box="[1330,1414,566,593]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-15@15.[132,1455,215,1878]" captionTargetPageId="15" captionText="FIG. 9. — Albanerpetontid remains from Angeac-Charente: A, B, left dentary of Albanerpetontidae indet. (ANG M-101) in lingual (A) and dorsal (B) views; C, premaxillary of Albanerpetontidae indet. (ANG M-102) in ventrolingual view; D, E, left maxillary of Albanerpetontidae indet. (ANG M-51) in labial (D) and lingual (E) views; F, G, dentary fragment of Albanerpetontidae indet. (ANG M-35) in labial view showing foramina discussed in the text (F) and lingual (G) view; H, distal end of a left humerus of Albanerpetontidae indet. (ANG M-36) in ventral view; I-K, dorsal vertebra of Albanerpetontidae indet. (ANG M-103) in dorsal (I), anterior (J) and lateral (K) views. Scale bars: A-E, H-K, 400 µm; F, G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6902053" httpUri="https://zenodo.org/record/6902053/files/figure.png" pageId="14" pageNumber="695">Fig. 9H</figureCitation>
|
||
|
|
), as also reported for
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE576E9A06EFC87FD21" authority="(Estes & Hoffstetter 1976)" baseAuthorityName="Estes & Hoffstetter" baseAuthorityYear="1976" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="species" species="inexpectatum">
|
||
|
|
<emphasis id="B915EABDFFB8FFE576E9A06EFB36FDC1" box="[979,1245,598,624]" italics="true" pageId="14" pageNumber="695">Albanerpeton inexpectatum</emphasis>
|
||
|
|
(
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE571D6A06EFC8AFD21" author="Estes & Hoffstetter" firstAuthor="Estes" pageId="14" pageNumber="695" pagination="297 - 343" refId="ref39143" refString="ESTES R. & HOFFSTETTER R. 1976. - Les urodeles du Miocene de La Grive-Saint-Alban (Isere, France). Bulletins du Museum national d'Histoire naturelle, 3 eme serie, Sciences de la Terre 398: 297 - 343. https: // www. biodiversitylibrary. org / page / 55505209" type="journal article" year="1976">Estes & Hoffstetter 1976</bibRefCitation>
|
||
|
|
)
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE5769BA04FFA41FD21" authority="(Sweetman & Gardner 2013)" baseAuthorityName="Sweetman & Gardner" baseAuthorityYear="2013" box="[929,1450,630,656]" class="Amphibia" family="Albanerpetontidae" genus="Wesserpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="species" species="evansae">
|
||
|
|
<emphasis id="B915EABDFFB8FFE5769BA04FFB81FD21" box="[929,1130,631,656]" italics="true" pageId="14" pageNumber="695">Wesserpeton evansae</emphasis>
|
||
|
|
(
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE57143A04EFA74FD21" author="SWEETMAN S. C. & GARDNER J. D." box="[1145,1439,630,656]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
|
||
|
|
)
|
||
|
|
</taxonomicName>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BDE36AFFFB8FFE57679A0AEFA71FB3F" blockId="14.[810,1456,215,1294]" pageId="14" pageNumber="695">
|
||
|
|
The albanerpetontids from Angeac-Charente differ from
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE57614A08DFC2BFD7E" authorityName="Gardner, Evans & Sigogneau-Russell" authorityYear="2003" box="[814,960,693,719]" class="Amphibia" family="Albanerpetontidae" genus="Anoualerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="B915EABDFFB8FFE57614A08DFC2BFD7E" box="[814,960,693,719]" italics="true" pageId="14" pageNumber="695">Anoualerpeton</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE576D5A08DFB34FD7E" authorityName="Estes" authorityYear="1981" box="[1007,1247,693,719]" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="species" species="nexuosum">
|
||
|
|
<emphasis id="B915EABDFFB8FFE576D5A08DFB34FD7E" box="[1007,1247,693,719]" italics="true" pageId="14" pageNumber="695">Albanerpeton nexuosum</emphasis>
|
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</taxonomicName>
|
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|
, but resemble other
|
||
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<taxonomicName id="4C614D2CFFB8FFE57611A0EDFBE9FD5E" box="[811,1026,725,751]" pageId="14" pageNumber="695">
|
||
|
|
<emphasis id="B915EABDFFB8FFE57611A0EDFC5AFD5E" box="[811,945,725,751]" italics="true" pageId="14" pageNumber="695">Albanerpeton</emphasis>
|
||
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|
species
|
||
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|
</taxonomicName>
|
||
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,
|
||
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<taxonomicName id="4C614D2CFFB8FFE57130A0EDFB83FD5E" baseAuthorityName="Gardner" baseAuthorityYear="2000" box="[1034,1128,725,751]" class="Amphibia" family="Albanerpetontidae" genus="Celtedens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="B915EABDFFB8FFE57130A0EDFB83FD5E" box="[1034,1128,725,751]" italics="true" pageId="14" pageNumber="695">Celtedens</emphasis>
|
||
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</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE571A1A0EEFAF9FD5E" authorityName="Sweetman & Gardner" authorityYear="2013" box="[1179,1298,726,751]" class="Amphibia" family="Albanerpetontidae" genus="Wesserpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="B915EABDFFB8FFE571A1A0EEFAF9FD5E" box="[1179,1298,726,751]" italics="true" pageId="14" pageNumber="695">Wesserpeton</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
in having dentaries and maxillae with relatively straight occlusal margins, and teeth weakly heterodont in size (
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE57147A12DFA4BFC9E" author="SWEETMAN S. C. & GARDNER J. D." box="[1149,1440,789,815]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="295 - 324" part="58" refId="ref49766" refString="SWEETMAN S. C. & GARDNER J. D. 2013. - A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England. Acta Palaeontologica Polonica 58: 295 - 324. https: // doi. org / 10.4202 / app. 2011.0109" title="A new albanerpetontid amphibian from the Barremian (Early Cretaceous) Wessex Formation of the Isle of Wight, southern England" type="journal article" year="2013">Sweetman & Gardner 2013</bibRefCitation>
|
||
|
|
). Unfortunately, the diagnostic characters necessary for identification at the generic level are lacking. Neither the maxilla nor the dentary is diagnostic for
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE57115A14DFB5EFC3E" authorityName="Estes & Hoffstetter" authorityYear="1976" box="[1071,1205,885,911]" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="B915EABDFFB8FFE57115A14DFB5EFC3E" box="[1071,1205,885,911]" italics="true" pageId="14" pageNumber="695">Albanerpeton</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE571D3A14DFC80FC1E" authority="(Gardner 2000)" baseAuthorityName="Gardner" baseAuthorityYear="2000" class="Amphibia" family="Albanerpetontidae" genus="Celtedens" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="B915EABDFFB8FFE571D3A14DFAACFC3E" box="[1257,1351,885,911]" italics="true" pageId="14" pageNumber="695">Celtedens</emphasis>
|
||
|
|
(
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE5706CA14DFC89FC1E" author="Gardner" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">Gardner 2000</bibRefCitation>
|
||
|
|
)
|
||
|
|
</taxonomicName>
|
||
|
|
, as their diagnoses are based on frontal characters (
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE57060A1ADFC84FC7E" author="Gardner" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="57 - 84" refId="ref40194" refString="GARDNER J. D. 1999 a. - Redescription of the geologically youngest albanerpetontid (? Lissamphibia): Albanerpeton inexpectatum Estes and Hoffstetter, 1976, from the Miocene of France. Annales de Paleontologie 85: 57 - 84. https: // doi. org / 10.1016 / S 0753 - 3969 (99) 80008 - 1" type="journal article" year="1999" yearSuffix="a">Gardner 1999a</bibRefCitation>
|
||
|
|
, b,
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE576AFA18DFC27FC7E" author="GARDNER J. D." box="[917,972,949,975]" journalOrPublisher="Acta Palaeontologica Polonica" pageId="14" pageNumber="695" pagination="55 - 70" part="45" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." title="Revised taxonomy of albanerpetontid amphibians" type="journal article" year="2000">2000</bibRefCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE576E3A18DFB0DFC7E" author="McGowan & Evans" box="[985,1254,948,975]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="143 - 145" refId="ref44053" refString="MCGOWAN G. J. & EVANS S. E. 1995. - Albanerpetontid ampibians from the Cretaceous of Spain. Nature 373: 143 - 145. https: // doi. org / 10.1038 / 373143 a 0" type="journal article" year="1995">McGowan & Evans 1995</bibRefCitation>
|
||
|
|
;
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE571C8A18DFA4BFC7E" author="McGowan" box="[1266,1440,949,975]" firstAuthor="McGowan" pageId="14" pageNumber="695" pagination="1 - 32" refId="ref43957" refString="MCGOWAN G. J. 2002. - Albanerpetontid amphibians from the Lower Cretaceous of Spain and Italy: a description and reconsideration of their systematics. Zoological Journal of the Linnean Society 135: 1 - 32. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.00013. x" type="journal article" year="2002">McGowan 2002</bibRefCitation>
|
||
|
|
). We did not find any frontal bones in the Angeac-Charente material, and so we cannot attribute the albanerpetontid material to these genera with certainty. Within
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE57019A62CFA42FB9F" authorityName="Estes & Hoffstetter" authorityYear="1976" box="[1315,1449,1044,1070]" class="Amphibia" family="Albanerpetontidae" genus="Albanerpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="B915EABDFFB8FFE57019A62CFA42FB9F" box="[1315,1449,1044,1070]" italics="true" pageId="14" pageNumber="695">Albanerpeton</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
, the premaxilla is the most taxonomically informative bone for species (
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE576BFA66CFBC1FBDF" author="Gardner" box="[901,1066,1108,1134]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="529 - 544" refId="ref40256" refString="GARDNER J. D. 1999 b. - The amphibian Albanerpeton arthridion and the Aptian - Albian biogeography of albanerpetontids. Palaeontology 42: 529 - 544. https: // doi. org / 10.1111 / 1475 - 4983.00083" type="journal article" year="1999" yearSuffix="b">Gardner 1999b</bibRefCitation>
|
||
|
|
,
|
||
|
|
<bibRefCitation id="EFF04B5EFFB8FFE57103A66CFB9BFBDF" author="Gardner" box="[1081,1136,1108,1134]" firstAuthor="Gardner" pageId="14" pageNumber="695" pagination="55 - 70" refId="ref40297" refString="GARDNER J. D. 2000. - Revised taxonomy of albanerpetontid amphibians. Acta Palaeontologica Polonica 45 (1): 55 - 70." type="journal article" year="2000">2000</bibRefCitation>
|
||
|
|
), but the specimens found to date at Angeac-Charente are too incomplete to be relevant.
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="8BDE36AFFFB8FFE57679A6ACFABAFABC" blockId="14.[810,1456,215,1294]" pageId="14" pageNumber="695">
|
||
|
|
Thus, considering the characters mentioned above, we can attribute the Angeac-Charente specimens to the family
|
||
|
|
<taxonomicName id="4C614D2CFFB8FFE5704CA68CFC2BFB5F" authorityName="Fox & Naylor" authorityYear="1982" class="Amphibia" family="Albanerpetontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gymnophiona" pageId="14" pageNumber="695" phylum="Chordata" rank="family">Albanerpetontidae</taxonomicName>
|
||
|
|
, but so far the material is too incomplete to allow a further attribution at the generic and specific level.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|