<documentid="E8ACEC69C4F7BA235146E28A11BA1AAE"ID-DOI="10.5852/ejt.2021.737.1259"ID-GBIF-Dataset="4b7a838b-1449-4ae4-90ff-5a7f113868e0"ID-ISSN="2118-9773"ID-Zenodo-Dep="4594357"ID-ZooBank="FE4ED74A-3FE6-4CA6-A116-CB3AF46826F7"IM.materialsCitations_approvedBy="valdenar"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="valdenar"IM.treatments_approvedBy="valdenar"checkinTime="1615409071345"checkinUser="plazi"docAuthor="Goulding, Tricia C., Bourke, Adam J., Comendador, Joseph, Khalil, Munawar, Quang, Ngo Xuan, Tan, Shau Hwai, Tan, Siong Kiat & Dayrat, Benoît"docDate="2021"docId="03A6D248FFC38B34DD73FC0AFC14D373"docLanguage="en"docName="ejt-737_1-133.pdf"docOrigin="European Journal of Taxonomy 737 (1)"docSource="http://dx.doi.org/10.5852/ejt.2021.737.1259"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Platevindex martensi Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat 2021"docType="treatment"docVersion="15"lastPageNumber="103"masterDocId="FF9FAA30FF9C8B52DF38FFAFFFC3D05C"masterDocTitle="Systematic revision of Platevindex Baker, 1938 (Gastropoda: Euthyneura: Onchidiidae)"masterLastPageNumber="133"masterPageNumber="1"pageNumber="96"updateTime="1733327641084"updateUser="valdenar"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:affiliationid="A804438BFF303FD0A551ABDA12784534">Department of Biology, Pennsylvania State University, University Park, PA 16802, USA. & Smithsonian Institution, National Museum of Natural History, PO Box 37012, MRC 163, Washington, DC, 20013, USA.</mods:affiliation>
<mods:affiliationid="43201A69DAFDF5FDB3729B1CEC55617A">College of Engineering, Information Technology and the Environment, Charles Darwin University, Ellengowan Dr., Casuarina, NT 0810, Australia.</mods:affiliation>
<mods:affiliationid="C49EC2DBC4F3CC7AD4BC7FCF5DFED6BD">National Museum of the Philippines, Taft Ave., Ermita, Manila, 1000 Metro Manila, Philippines.</mods:affiliation>
<mods:affiliationid="4A68A67A6FDC52AD8042F3227C6806DE">Department of Marine Science, Faculty of Agriculture, Universitas Malikussaleh, Reuleut Main Campus, Kecamatan Muara Batu, North Aceh, Aceh, 24355, Indonesia.</mods:affiliation>
<mods:affiliationid="52D2B8F3FDBB73B7F0DA1ECF43418309">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
<mods:affiliationid="C31AE7122DAF7F64C2525DE7912EAFA7">Centre for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 Minden Penang, Malaysia. & Marine Science Laboratory, School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden Penang, Malaysia.</mods:affiliation>
<mods:affiliationid="C7621C5893B6E2D7B205DC33C235D09E">Lee Kong Chian Natural History Museum, 2 Conservatory Dr, National University of Singapore, 117377, Singapore.</mods:affiliation>
<mods:affiliationid="380AB42257B432AF472BA9CE24EEDB72">Department of Biology, Pennsylvania State University, University Park, PA 16802, USA.</mods:affiliation>
<taxonomicNameid="4C0F18DDFFC38B0DDD73FC0AFC2BD39C"ID-CoL="77NX3"authority="(Plate, 1893)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[587,1000,933,960]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="95"pageNumber="96"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFC38B0DDC61FC09FC23D39C"author="Plate L. H."box="[857,992,934,960]"pageId="95"pageNumber="96"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893</bibRefCitation>
<figureCitationid="13347FDBFFC38B0DDDE0FC66FC98D3B8"box="[728,859,969,996]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStart-3="Fig"captionStart-4="Fig"captionStartId-0="97.[189,232,1749,1775]"captionStartId-1="98.[189,232,1644,1670]"captionStartId-2="99.[191,234,1916,1942]"captionStartId-3="100.[190,233,1921,1947]"captionStartId-4="102.[189,232,1810,1836]"captionTargetBox-0="[208,1380,264,1720]"captionTargetBox-1="[204,1340,271,1622]"captionTargetBox-2="[189,1398,1041,1889]"captionTargetBox-3="[189,1398,1495,1898]"captionTargetBox-4="[189,1398,864,1775]"captionTargetId-0="figure-144@97.[208,1380,264,1748]"captionTargetId-1="figure-187@98.[189,1346,265,1625]"captionTargetId-2="figure-355@99.[189,1398,1041,1895]"captionTargetId-3="figure-480@100.[189,1398,1495,1903]"captionTargetId-4="figure-331@102.[189,1398,864,1775]"captionTargetPageId-0="97"captionTargetPageId-1="98"captionTargetPageId-2="99"captionTargetPageId-3="100"captionTargetPageId-4="102"captionText-0="Fig. 54. Platevindex martensi (Plate, 1893), live animals. A. Dorsal view, 34 mm long [1748], Indonesia, Sumatra (UMIZ 00109). B. Dorsal view, 24 mm long [3358], Philippines, Bohol (PNM 041251). C. Dorsal view, 34 mm long [3364], Philippines, Bohol (PNM 041251). D. Dorsal view, 13 mm long [1707], Australia, Darwin (NTM P.57611). E. Dorsal view, 38 mm long [#1], Peninsular Malaysia, Langkawi (USMMC 00032). F. Dorsal view, 25 mm long [1709], Australia, Darwin (NTM P.57611). G. Ventral view, 36 mm long [2334], Indonesia, Sulawesi (UMIZ 00111). H. Ventral view, same as C. I. Ventral view, same as F. J. Ventral view, 31 mm long [934], Peninsular Malaysia, Balok (USMMC 00030). K. Ventral view, 30 mm long, [964], Peninsular Malaysia, Penang (USMMC 00029)."captionText-1="Fig. 55 Platevindex martensi (Plate, 1893), external morphology, digestive and reproductive systems. A. Dorsal notum, entire animal, Bohol, Philippines [3365] (PNM 041251). B. Digestive system, dorsal view, type II with a transitional loop oriented at 9 o’clock, Sulawesi, Indonesia [2333] (UMIZ 00111). C. Digestive system, dorsal view, type II with a transitional loop oriented at 6 o’clock, Peninsular Malaysia [964] (USMMC 00029). D. Posterior reproductive system (remaining parts), Phetchaburi, Gulf of Thailand, holotype (ZMB/Moll 8454). E. Posterior reproductive system, Darwin, Northern Territory, Australia [1711] (NTM P.57611). F. Anterior, male copulatory apparatus, same as E. Abbreviations: dd = deferent duct; ddg = dorsal digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior digestive gland; ps = penial sheath; rg = rectal gland; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scale bars: A = 10 mm; B–C = 3 mm; D–F = 2 mm."captionText-2="Fig. 56.Platevindex martensi (Plate, 1893),radula, Philippines, Bohol[3336].A. Rachidian and innermost lateral teeth. B. Innermost lateral teeth. C. Lateral teeth. D. Outermost lateral teeth. Abrreviations: lp = lateral protuberance. Scale bars: A, D = 20 μm; B–C = 30 μm."captionText-3="Fig. 57.Platevindex martensi (Plate, 1893), penial hooks.A. Penis everted with hooks outside, Peninsular Malaysia, Penang [964] (USMMC 00029). B. Australia, Darwin [1711] (NTM P.57611). Scale bars: A = 50 μm; B = 30 μm."captionText-4="Fig. 58. Habitats of Platevindex martensi (Plate, 1893). A. Philippines, Bohol, narrow mangrove will tall trees on edge of fish ponds, cement ditches between the mangrove patches and the ponds (station 195). B. Philippines, Bohol, narrow mangrove on the edge of fish ponds, tall Rhizophora and Avicennia, many dead logs (station 194). C. Indonesia, Sulawesi, small mangrove used as outhouse by village, very impacted by trash (station 92). D. Australia, Darwin, Ceriops and old logs in Rhizophora forest (station 70)."figureDoi-0="http://doi.org/10.5281/zenodo.4594465"figureDoi-1="http://doi.org/10.5281/zenodo.4601464"figureDoi-2="http://doi.org/10.5281/z
<bibRefCitationid="EF9E1EAFFFC38B0DDE4FFBBDFDB0D470"author="Plate L. H."box="[375,627,1042,1068]"pageId="95"pageNumber="96"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893: 196–197</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFC38B0DDE9CFB9AFD61D413"author="Plate L. H."box="[420,674,1077,1103]"pageId="95"pageNumber="96"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893: 198–199</bibRefCitation>
<taxonomicNameid="4C0F18DDFFC38B0DDF85FAA4FE75D579"ID-CoL="77NX3"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[189,438,1291,1317]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="95"pageNumber="96"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFC38B0DDEEBFAA3FD46D57A"author="Dayrat B."box="[467,645,1291,1318]"pageId="95"pageNumber="96"pagination="1 - 26"refId="ref70452"refString="Dayrat B. 2009. Review of the current knowledge of the systematics of Onchidiidae (Mollusca: Gastropoda: Pulmonata) with a checklist of nominal species. Zootaxa 2068: 1 - 26. https: // doi. org / 10.11646 / zootaxa. 2068.1.1"type="journal article"year="2009">Dayrat 2009: 5</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFC38B0DDEF1FA81FD5FD514"author="Zhang K. & Wang D. & Shen H. & Qian J. & Guan H. & Wu H. & Gao Y."box="[457,668,1326,1353]"pageId="95"pageNumber="96"pagination="72 - 78"refId="ref75598"refString="Zhang K., Wang D., Shen H., Qian J., Guan H., Wu H. & Gao Y. 2017. Redescription of Platevindex mortoni (Gastropoda: Eupulmonata: Onchidiidae) from China. Molluscan Research 37: 72 - 78. https: // doi. org / 10.1080 / 13235818.2016.1223535"type="journal article"year="2017">
<bibRefCitationid="EF9E1EAFFFC38B0DDB13FA81FB0FD514"author="Britton K. M."box="[1067,1228,1326,1352]"pageId="95"pageNumber="96"pagination="179 - 191"refId="ref69860"refString="Britton K. M. 1984. The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera. Journal of Molluscan Studies 50: 179 - 191. https: // doi. org / 10.1093 / oxfordjournals. mollus. a 065863"type="journal article"year="1984">Britton, 1984</bibRefCitation>
<materialsCitationid="3B676903FFC38B0DDF85FA66FDCAD65B"ID-GBIF-Occurrence="3051754362"collectionCode="ZMB"country="Thailand"location="Gulf of Siam"municipality="Petshaburi"pageId="95"pageNumber="96"specimenCode="ZMB/Moll 8454"specimenCount="1"stateProvince="Phetchaburi"typeStatus="holotype">
<locationid="8ED03585FFC38B0DDCFEFA66FBABD5BF"LSID="urn:lsid:plazi:treatment:03A6D248FFC38B34DD73FC0AFC14D373:8ED03585FFC38B0DDCFEFA66FBABD5BF"box="[966,1128,1481,1507]"country="Thailand"municipality="Petshaburi"name="Gulf of Siam"pageId="95"pageNumber="96"stateProvince="Phetchaburi">Gulf of Siam</location>
was previously dissected. Internal organs are mostly missing; only a few pieces of the female (posterior) reproductive system and digestive system remain. The penial complex and buccal mass are missing.
was previously dissected. Internal organs are mostly missing. The buccal mass remains, as well as pieces of the deferent duct, oviduct and spermatheca.
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<collectedFromid="381320A3FFFC8B32DBF3FDF7FD03D2C8"pageId="96"pageNumber="97">mangrove just behind abandoned buildings, high intertidal</collectedFrom>
<materialsCitationid="3B676903FFFC8B32DCB8FDD5FDFED287"ID-GBIF-Occurrence="3051754457"collectedFrom="mounds of Thalassina, Rhizophora, dead logs in the high intertidal"collectingDate="2012-10-19"collectionCode="UMIZ"latitude="-5.6695666"location="Sungai Lubuk"longLatPrecision="1"longitude="105.56828"municipality="Sungai Lubuk"pageId="96"pageNumber="97"specimenCode="UMIZ 00109"specimenCount="1">
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<materialsCitationid="3B676903FFFC8B32DF85FC7DFE78D46E"ID-GBIF-Occurrence="3051754402"collectedFrom="deep Rhizophora forest with old, tall trees, hard mud, many small creeks and dead logs"collectingDate="2011-07-18"collectionCode="USMMC"latitude="5.6505833"location="Merbok"longLatPrecision="1"longitude="100.4297"pageId="96"pageNumber="97"specimenCode="USMMC 00031"specimenCount="2">
<materialsCitationid="3B676903FFFC8B32DEECFBB7FAA5D409"ID-GBIF-Occurrence="3051754337"collectedFrom="mangrove of mostly Rhizophora, some Avicennia with soft mud near creek"collectingDate="2011-07-19"collectionCode="USMMC"latitude="5.66905"location="Merbok"longLatPrecision="1"longitude="100.4363"pageId="96"pageNumber="97"specimenCode="USMMC 00077"specimenCount="1">
<materialsCitationid="3B676903FFFC8B32DF85FBF1FE0FD4E3"ID-GBIF-Occurrence="3051754356"collectedFrom="open forest of Rhizophora, Sonneratia, Bruguiera with high mounds of mud around some trees"collectingDate="2011-07-15"collectionCode="USMMC"latitude="6.42195"location="Langkawi"longLatPrecision="1"longitude="99.8351"municipality="Tanjung Rhu"pageId="96"pageNumber="97"specimenCode="USMMC 00032"specimenCount="1">
<materialsCitationid="3B676903FFFC8B32DED6FB0AFE41D559"ID-GBIF-Occurrence="3051754314"collectedFrom="old Rhizophora and Acrostichum ferns very high in tidal zone, near boardwalk"collectingDate="2011-07-19"collectionCode="USMMC"latitude="4.8405666"location="Kuala Sepatang"longLatPrecision="1"longitude="100.63627"municipality="Kuala Sepatang"pageId="96"pageNumber="97"specimenCode="USMMC 00033"specimenCount="1">
<materialsCitationid="3B676903FFFC8B32DF85FA86FC77D5D6"ID-GBIF-Occurrence="3051754406"collectedFrom="narrow Rhizophora and Avicennia mangrove by the sea with fish ponds built on landward side, many dead logs"collectingDate="2014-07-17"collectionCode="PNM"country="Philippines"latitude="9.858867"location="Mabini"longLatPrecision="1"longitude="124.52808"municipality="Mabini"pageId="96"pageNumber="97"specimenCode="PNM 041250"specimenCount="2"stateProvince="Bohol">
<materialsCitationid="3B676903FFFC8B32DCF4FAC0FC22D5AF"ID-GBIF-Occurrence="3051754425"collectedFrom="narrow Rhizophora and Avicennia mangrove by the sea with fish ponds built on landward side, cement ditches between the mangrove patches and the ponds"collectingDate="2014-07-18"collectionCode="PNM"latitude="9.8567"location="Mabini"longLatPrecision="1"longitude="124.516365"municipality="Mabini"pageId="96"pageNumber="97"specimenCode="PNM 041251"specimenCount="4">
<materialsCitationid="3B676903FFFC8B32DF85F9B9FC66D608"ID-GBIF-Occurrence="3051754443"collectedFrom="mud and dead logs in upper tidal"collectingDate="2010-03-30"collectionCode="ZRC"country="Singapore"latitude="1.3806667"location="Pasir Ris Park"longLatPrecision="1"longitude="103.953735"pageId="96"pageNumber="97"specimenCode="ZRC.MOL.10480"specimenCount="1">
<collectingDateid="EFF5BC76FFFC8B32DBE9F9B9FAB1D66C"box="[1233,1394,1558,1585]"pageId="96"pageNumber="97"value="2010-03-30">30 Mar. 2010</collectingDate>
</date>
; station 5;
<collectedFromid="381320A3FFFC8B32DE0DF996FD06D60F"box="[309,709,1593,1620]"pageId="96"pageNumber="97">mud and dead logs in upper tidal</collectedFrom>
<materialsCitationid="3B676903FFFC8B32DCFAF995FC18D6C6"ID-GBIF-Occurrence="3051754367"collectedFrom="piece of wood in Avicennia and Rhizophora mangrove"collectingDate="2010-04-05"collectionCode="ZRC"latitude="1.4464166"location="Lim Chu Kang"longLatPrecision="1"longitude="103.70885"municipality="Lim Chu Kang"pageId="96"pageNumber="97"specimenCode="ZRC.MOL.10481"specimenCount="3">
<emphasisid="B97BBF4CFFFC8B32DFD2F95FFD00D756"bold="true"box="[234,707,1776,1802]"pageId="96"pageNumber="97">Color and morphology of live animals</emphasis>
(
<figureCitationid="13347FDBFFFC8B32DDEBF95FFCADD757"box="[723,878,1776,1803]"captionStart-0="Fig"captionStart-1="Fig"captionStartId-0="97.[189,232,1749,1775]"captionStartId-1="98.[189,232,1644,1670]"captionTargetBox-0="[208,1380,264,1720]"captionTargetBox-1="[204,1340,271,1622]"captionTargetId-0="figure-144@97.[208,1380,264,1748]"captionTargetId-1="figure-187@98.[189,1346,265,1625]"captionTargetPageId-0="97"captionTargetPageId-1="98"captionText-0="Fig. 54. Platevindex martensi (Plate, 1893), live animals. A. Dorsal view, 34 mm long [1748], Indonesia, Sumatra (UMIZ 00109). B. Dorsal view, 24 mm long [3358], Philippines, Bohol (PNM 041251). C. Dorsal view, 34 mm long [3364], Philippines, Bohol (PNM 041251). D. Dorsal view, 13 mm long [1707], Australia, Darwin (NTM P.57611). E. Dorsal view, 38 mm long [#1], Peninsular Malaysia, Langkawi (USMMC 00032). F. Dorsal view, 25 mm long [1709], Australia, Darwin (NTM P.57611). G. Ventral view, 36 mm long [2334], Indonesia, Sulawesi (UMIZ 00111). H. Ventral view, same as C. I. Ventral view, same as F. J. Ventral view, 31 mm long [934], Peninsular Malaysia, Balok (USMMC 00030). K. Ventral view, 30 mm long, [964], Peninsular Malaysia, Penang (USMMC 00029)."captionText-1="Fig. 55 Platevindex martensi (Plate, 1893), external morphology, digestive and reproductive systems. A. Dorsal notum, entire animal, Bohol, Philippines [3365] (PNM 041251). B. Digestive system, dorsal view, type II with a transitional loop oriented at 9 o’clock, Sulawesi, Indonesia [2333] (UMIZ 00111). C. Digestive system, dorsal view, type II with a transitional loop oriented at 6 o’clock, Peninsular Malaysia [964] (USMMC 00029). D. Posterior reproductive system (remaining parts), Phetchaburi, Gulf of Thailand, holotype (ZMB/Moll 8454). E. Posterior reproductive system, Darwin, Northern Territory, Australia [1711] (NTM P.57611). F. Anterior, male copulatory apparatus, same as E. Abbreviations: dd = deferent duct; ddg = dorsal digestive gland; fgm = female gland mass; hd = hermaphroditic duct; hg = hermaphroditic gland; i = intestine; ov = oviduct; pdg = posterior digestive gland; ps = penial sheath; rg = rectal gland; rm = retractor muscle; rs = receptaculum seminis; sp = spermatheca; st = stomach; v = vestibule. Scale bars: A = 10 mm; B–C = 3 mm; D–F = 2 mm."figureDoi-0="http://doi.org/10.5281/zenodo.4594465"figureDoi-1="http://doi.org/10.5281/zenodo.4601464"httpUri-0="https://zenodo.org/record/4594465/files/figure.png"httpUri-1="https://zenodo.org/record/4601464/files/figure.png"pageId="96"pageNumber="97">Figs 54–55A</figureCitation>
Live animals are not usually covered with mud, and their natural color is visible without washing. The dorsal notum is brown to dark brown or black, often mottled with two colors. The notum bears small elongated (longitudinal) ridges. The ventral color varies but the hyponotum is yellow, beige or yellow-orange, with distinct darker spots of variable size (small to large) and color (faintly tan to black). When those spots are large and dark enough, the hyponotum appears nearly black (which was most common in the Strait of
, the edge of the hyponotum commonly is orange, which was also occasionally observed in other populations. The foot is distinctively orange. Papillae with dorsal eyes are present. Their exact number is variable (generally between 20 and 65). Each papilla bears one dorsal eye. Eyes are distributed across the notum and many eyes are found at the margin (i.e., eyes can be <
<taxonomicNameid="4C0F18DDFFFD8B33DE25F97AFD70D6B3"authority="(Plate, 1893)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[285,691,1749,1775]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="97"pageNumber="98"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFFD8B33DD1AF97AFD6AD6B3"author="Plate L. H."box="[546,681,1749,1775]"pageId="97"pageNumber="98"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893</bibRefCitation>
<taxonomicNameid="4C0F18DDFFFE8B30DE24F9C3FD7FD6DB"authority="(Plate, 1893)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[284,700,1644,1671]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="98"pageNumber="99"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFFE8B30DD10F9C3FD71D6DB"author="Plate L. H."box="[552,690,1644,1671]"pageId="98"pageNumber="99"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893</bibRefCitation>
)
</taxonomicName>
, external morphology, digestive and reproductive systems.
<figureCitationid="13347FDBFFFF8B31DEF0FE78FDD2D1AD"box="[456,529,471,497]"captionStart="Fig"captionStartId="15.[189,232,1538,1564]"captionTargetBox="[235,1348,269,1504]"captionTargetId="figure-243@15.[231,1357,265,1524]"captionTargetPageId="15"captionText="Fig. 2. Intestinal loops of type II. Small black arrows show the direction of the intestinal transport. The clockwise loop is in blue. The counterclockwise loop is in yellow. The transitional loop (between clockwise and counterclockwise loops) is in green. Red arrows indicate the orientation of the transitional loop. A. Type II (as defined by Plate 1893), with a transitional loop oriented at 9 o’clock, redrawn from Plate (1893: pl. 8, fig. 31). B. Type II, with a transitional loop oriented at 9 o’clock, Platevindex coriaceus coriaceus (Semper, 1880) unit #2 [3170] (PNM 041236), Luzon, Philippines. C. Type II, with a transitional loop oriented at 9 o’clock, P. martensi (Plate, 1893) [2333] (UMIZ 00111), Sulawesi, Indonesia. D. Type II, with a transitional loop oriented at 6 o’clock, P. martensi [964] (USMMC 00029), Peninsular Malaysia. E. Type II, with a transitional loop oriented at 8 o’clock, P. luteus (Semper, 1880) [3367] (PNM 041246), Bohol, Philippines. F. Type II, with a transitional loop oriented at 8 o’clock, P. aptei Goulding & Dayrat sp. nov., [5955] (USMMC 00078), Peninsular Malaysia. G. Type II, with a transitional loop oriented at 7 o’clock, P. burnupi (Collinge, 1902) [3150] (MNHN-IM-2019-1398), Madagascar. H. Type II, with a transitional loop oriented at 7 o’clock, P. coriaceus coriaceus unit #2 [3256] (PNM 041238), Bohol, Philippines. Scale bars: B–H = 2 mm."figureDoi="http://doi.org/10.5281/zenodo.4594361"httpUri="https://zenodo.org/record/4594361/files/figure.png"pageId="99"pageNumber="100">Figs 2</figureCitation>
in length. Examples of radular formulae are presented in
<tableCitationid="C68D56E5FFFF8B31DBE7FE55FAF9D249"box="[1247,1338,506,533]"captionStart="Table 5"captionStartId="42.[189,258,265,291]"captionTargetBox="[223,1387,323,1985]"captionTargetPageId="42"captionText="Table 5. Radular formulae in species of Platevindex Baker, 1938."httpUri="http://table.plazi.org/id/DF7033D6FFB68B78DF85FEA6FC7DD17F"pageId="99"pageNumber="100"tableUuid="DF7033D6FFB68B78DF85FEA6FC7DD17F">Table 5</tableCitation>
In the posterior part of the reproductive system, the oviduct is wider than the deferent duct (approximately up to twice as wide). Its distal section (distal to the spermatheca) is approximately up to twice as long as its proximal section. The deferent duct is longer than the oviduct, not attached to it, and highly coiled with tight, long, U-shaped loops. However, the deferent duct tends to be shorter in
apart. The distal, flexible region of the penis with hooks is approximately
<quantityid="4CF7CEBBFFFF8B31DEE0FCF9FDABD32C"box="[472,616,853,880]"metricMagnitude="-3"metricUnit="m"metricValue="2.25"metricValueMax="2.5"metricValueMin="2.0"pageId="99"pageNumber="100"unit="mm"value="2.25"valueMax="2.5"valueMin="2.0">2 to 2.5 mm</quantity>
wide. Penial hooks are relatively large, approximately
<quantityid="4CF7CEBBFFFF8B31DE80FCD7FD8BD3CF"box="[440,584,888,915]"metricMagnitude="-2"metricUnit="m"metricValue="4.5"metricValueMax="5.5"metricValueMin="3.5"pageId="99"pageNumber="100"unit="mm"value="45.0"valueMax="55.0"valueMin="35.0">35 to 55 µm</quantity>
long (
<figureCitationid="13347FDBFFFF8B31DDABFCD6FD3CD3CF"box="[659,767,888,915]"captionStart="Fig"captionStartId="100.[190,233,1921,1947]"captionTargetBox="[189,1398,1495,1898]"captionTargetId="figure-480@100.[189,1398,1495,1903]"captionTargetPageId="100"captionText="Fig. 57.Platevindex martensi (Plate, 1893), penial hooks.A. Penis everted with hooks outside, Peninsular Malaysia, Penang [964] (USMMC 00029). B. Australia, Darwin [1711] (NTM P.57611). Scale bars: A = 50 μm; B = 30 μm."figureDoi="http://doi.org/10.5281/zenodo.4594469"httpUri="https://zenodo.org/record/4594469/files/figure.png"pageId="99"pageNumber="100">Fig. 57B</figureCitation>
), and can be seen inside the semi-transparent penis or when the penis is evaginated like the finger of a glove in the vestibule (
<figureCitationid="13347FDBFFFF8B31DCCAFC33FB9CD3EB"box="[1010,1119,924,951]"captionStart="Fig"captionStartId="100.[190,233,1921,1947]"captionTargetBox="[189,1398,1495,1898]"captionTargetId="figure-480@100.[189,1398,1495,1903]"captionTargetPageId="100"captionText="Fig. 57.Platevindex martensi (Plate, 1893), penial hooks.A. Penis everted with hooks outside, Peninsular Malaysia, Penang [964] (USMMC 00029). B. Australia, Darwin [1711] (NTM P.57611). Scale bars: A = 50 μm; B = 30 μm."figureDoi="http://doi.org/10.5281/zenodo.4594469"httpUri="https://zenodo.org/record/4594469/files/figure.png"pageId="99"pageNumber="100">Fig. 57A</figureCitation>
). The posterior retractor muscle of the penis inserts on the right side, within the third (posterior) quarter of the visceral cavity. The length of the retractor muscle varies from half the length of the penial sheath to slightly longer than the penial sheath. The deferent duct is highly convoluted (but is less convoluted in juvenile specimens).
<taxonomicNameid="4C0F18DDFFFF8B31DE21F8D2FD6BD7CB"authority="(Plate, 1893)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[281,680,1917,1943]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="99"pageNumber="100"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFFF8B31DD22F8D2FD5CD7CB"author="Plate L. H."box="[538,671,1917,1943]"pageId="99"pageNumber="100"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893</bibRefCitation>
<tableCitationid="C68D56E5FFF88B36DD7CFEC1FD5DD1D4"box="[580,670,366,392]"captionStart="Table 4"captionStartId="22.[184,253,658,686]"captionTargetBox="[198,1916,827,1626]"captionTargetPageId="22"captionText="Table 4 (continued on next page).Summary of internaland external traits in speciesofPlatevindex Baker, 1938.Traits are described in the corresponding species descriptions and vary among individuals. External traits include the length of the animal, number of dorsal eyes, color of the foot and other distinguishing characters such as the color of the hyponotum. Internal traits include the insertion of the retractor muscle (RM), the length of the retractor muscle compared to that of the penial sheath (PS), the length of the region with penial hooks and the type of the intestinal loops."httpUri="http://table.plazi.org/id/DF7033D6FF8A8B44DF80FD3DF8C6D344"pageId="100"pageNumber="101"tableUuid="DF7033D6FF8A8B44DF80FD3DF8C6D344">Table 4</tableCitation>
<taxonomicNameid="4C0F18DDFFF88B36DD94FE33FC6BD1EA"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[684,936,412,438]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
. The small, elongated ridges on its dorsal notum are also present in
<taxonomicNameid="4C0F18DDFFF88B36DCCAFE6FFB8ED185"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[1010,1101,448,473]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
<taxonomicNameid="4C0F18DDFFF88B36DF85FE4CFEDBD1A0"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[189,280,483,508]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
<taxonomicNameid="4C0F18DDFFF88B36DC3BFDA8FC4AD27C"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[771,905,519,544]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF88B36DCFFFDA8FBE1D27C"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[967,1058,519,544]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
Internally, the long deferent duct with elongated loops in the anterior penial apparatus (which is not closely attached to the oviduct) differentiate
<taxonomicNameid="4C0F18DDFFF88B36DEFAFDE2FD8BD23A"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[450,584,589,614]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
. Also, the penial retractor muscle inserts between ½ to ¾ down the length of the visceral cavity in
<taxonomicNameid="4C0F18DDFFF88B36DCD9FDDFFBAAD2D5"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[993,1129,624,649]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
while it inserts at the posterior end of the cavity in
<taxonomicNameid="4C0F18DDFFF88B36DD2EFD3CFDADD2F0"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[534,622,659,684]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
<figureCitationid="13347FDBFFF88B36DE5EFD7BFE0CD2B2"box="[358,463,724,750]"captionStart="Fig"captionStartId="36.[189,232,1716,1742]"captionTargetBox="[189,1398,741,1688]"captionTargetId="figure-232@36.[189,1398,741,1688]"captionTargetPageId="36"captionText="Fig. 10. Geographic distributions of species of Platevindex. Colored areas correspond to hypothetical ranges proposed based on all known records, and the colors are the same as those used for each species in the phylogenetic trees (Figs 1–5). Distinct colors are used for each subspecies and mitochondrial unit in P. coriaceus (Semper, 1880) as well as for each subspecies of P. burnupi (Collinge, 1902). Black dots show sites sampled for this study and red dots (China and Japan) show localities of previously published genetic sequences included in the phylogenetic analyses. A. Platevindex luteus (Semper, 1880), P. applanatus (Simroth, 1920), P. burnupi. B. Platevindex tigrinus (Stoliczka, 1869), P. coriaceus coriaceus (units #1, #2, and #3) and P. coriaceus darwinensis subsp. nov. C. Platevindex martensi (Plate, 1893), P. aptei Goulding & Dayrat sp. nov., P. amboinae (Plate, 1893) and P. latus (Plate, 1893)."figureDoi="http://doi.org/10.5281/zenodo.4594373"httpUri="https://zenodo.org/record/4594373/files/figure.png"pageId="100"pageNumber="101">Fig. 10C</figureCitation>
<taxonomicNameid="4C0F18DDFFF88B36DED7FCE5FDB0D33F"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[495,627,842,867]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
<figureCitationid="13347FDBFFF88B36DE15FC25FE40D3F9"box="[301,387,906,933]"captionStart="Fig"captionStartId="102.[189,232,1810,1836]"captionTargetBox="[189,1398,864,1775]"captionTargetId="figure-331@102.[189,1398,864,1775]"captionTargetPageId="102"captionText="Fig. 58. Habitats of Platevindex martensi (Plate, 1893). A. Philippines, Bohol, narrow mangrove will tall trees on edge of fish ponds, cement ditches between the mangrove patches and the ponds (station 195). B. Philippines, Bohol, narrow mangrove on the edge of fish ponds, tall Rhizophora and Avicennia, many dead logs (station 194). C. Indonesia, Sulawesi, small mangrove used as outhouse by village, very impacted by trash (station 92). D. Australia, Darwin, Ceriops and old logs in Rhizophora forest (station 70)."figureDoi="http://doi.org/10.5281/zenodo.4594471"httpUri="https://zenodo.org/record/4594471/files/figure.png"pageId="100"pageNumber="101">Fig. 58</figureCitation>
<taxonomicNameid="4C0F18DDFFF88B36DF85FC16FE74D38F"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[189,439,953,979]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
is typically found in or near mangroves, on tree roots and trunks as well as dead logs. It usually lives in the high intertidal only reached by the highest tides (with
slugs normally are not found directly on mud or sand.
<taxonomicNameid="4C0F18DDFFF88B36DE39FBE9FE39D403"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[257,506,1094,1120]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
, including a flattened and hard body, a narrow foot, a male opening to the right of the midline between the two eye tentacles and no accessory penial gland. In the original description of
<bibRefCitationid="EF9E1EAFFFF88B36DB94FB53FAB4D54B"author="Plate L. H."box="[1196,1399,1276,1303]"pageId="100"pageNumber="101"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate (1893: 196)</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFF98B37DBC8FE83FF2FD135"author="Plate L. H."pageId="101"pageNumber="102"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate 1893: 196</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFF98B37DB32FEE0FB4DD135"author="Plate L. H."box="[1034,1166,335,361]"pageId="101"pageNumber="102"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate 1893</bibRefCitation>
<taxonomicNameid="4C0F18DDFFF88B36DE23F82DFD6DD7C0"authority="(Plate, 1893)"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[283,686,1922,1948]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="100"pageNumber="101"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFF88B36DD26F82DFD67D7C0"author="Plate L. H."box="[542,676,1922,1948]"pageId="100"pageNumber="101"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate, 1893</bibRefCitation>
<taxonomicNameid="4C0F18DDFFF98B37DC80FE1CFB83D190"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[952,1088,435,460]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
is largely destroyed. The hermaphroditic gland is in pieces and disconnected from the hermaphroditic duct. A large part of the female gland mass is separated from the oviduct and the deferent duct. However, the long and convoluted deferent duct is intact and is only consistent with the species described here.
<bibRefCitationid="EF9E1EAFFFF98B37DB51FDB3FAF8D26A"author="Plate L. H."box="[1129,1339,540,566]"pageId="101"pageNumber="102"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate (1893: 197)</bibRefCitation>
also described the color of the live animal (based on field notes from von Martens) as having an intense yellow hyponotum with black marbling as well as a light gray-brown foot. A yellow hyponotum is only found in
<taxonomicNameid="4C0F18DDFFF98B37DE1EFD29FE6AD2C3"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[294,425,646,671]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DED9FD29FDFBD2C3"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[481,568,646,671]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
The color of the foot is the external trait which can distinguish these two species, and Plate’s description of “light gray-brown” is a closer match to the bright orange foot of
<taxonomicNameid="4C0F18DDFFF98B37DE2CFD62FE5AD2BA"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[276,409,717,742]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DDD8FD62FCFBD2BA"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[736,824,717,742]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
The number of dorsal eyes he described (90) is higher than what we have observed in
<taxonomicNameid="4C0F18DDFFF98B37DDE9FD5FFC97D355"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[721,852,752,777]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
, but we could not check this as the dorsal eyes have lost their pigment.
<bibRefCitationid="EF9E1EAFFFF98B37DEEBFCBDFD6CD371"author="Plate L. H."box="[467,687,786,813]"pageId="101"pageNumber="102"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate’s (1893: 197)</bibRefCitation>
description of the dorsal eyes that “sit alone on the big warts” is consistent with this species, but his comment that “this rule does not apply without exception, because on about 12 warts there are 2 eye spots, so that one can speak in these cases of a group” is surprising. We have found no evidence that dorsal eyes occur in groups of two in this species; Plate’s statement that dorsal eyes occurred in pairs suggests that his count of dorsal eyes is likely an overestimate.
<bibRefCitationid="EF9E1EAFFFF98B37DF85FC70FE1DD3A5"author="Plate L. H."box="[189,478,991,1017]"pageId="101"pageNumber="102"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate’s (1893: 198–199)</bibRefCitation>
, but many specimens were observed with a nearly solid black hyponotum (due to a high density of black spots) along the north-western coast of peninsular
belongs to the species described here due to the remains of its female (posterior) reproductive system. The length of the deferent duct adjacent to the oviduct is long and convoluted with elongated loops, which is indistinguishable from
<taxonomicNameid="4C0F18DDFFF98B37DD7BFAEFFD09D505"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[579,714,1344,1369]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DC83FA29FBFDD5C3"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[955,1086,1414,1439]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DB02FA05FB7DD59F"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[1082,1214,1450,1475]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DDA8F99FFCD7D615"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[656,788,1584,1609]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DDBDF9FCFCC2D630"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Stoliczka"baseAuthorityYear="1869"box="[645,769,1619,1644]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="tigrinus">
<taxonomicNameid="4C0F18DDFFF98B37DF85F9D9FEFCD6D3"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[189,319,1654,1679]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
(the wide, long, highly-convoluted deferent duct adjacent to the oviduct). In addition, Semper rightly noted that dorsal eyes are present across the notum including on its margin, and dorsal eyes very close to the notum edge (<
<taxonomicNameid="4C0F18DDFFF98B37DC26F912FC61D68A"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[798,930,1725,1750]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFF98B37DCE3F912FBF1D68A"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[987,1074,1725,1750]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
<bibRefCitationid="EF9E1EAFFFF98B37DF85F953FE69D74A"author="Zhang K. & Wang D. & Shen H. & Qian J. & Guan H. & Wu H. & Gao Y."box="[189,426,1788,1814]"pageId="101"pageNumber="102"pagination="72 - 78"refId="ref75598"refString="Zhang K., Wang D., Shen H., Qian J., Guan H., Wu H. & Gao Y. 2017. Redescription of Platevindex mortoni (Gastropoda: Eupulmonata: Onchidiidae) from China. Molluscan Research 37: 72 - 78. https: // doi. org / 10.1080 / 13235818.2016.1223535"type="journal article"year="2017">
is considered here a nomen dubium and their description is based on specimens that belong to at least two different species. One individual illustrated with a bright orange foot (
<bibRefCitationid="EF9E1EAFFFF98B37DE6AF8CAFDEBD7DC"author="Zhang K. & Wang D. & Shen H. & Qian J. & Guan H. & Wu H. & Gao Y."box="[338,552,1893,1920]"pageId="101"pageNumber="102"pagination="72 - 78"refId="ref75598"refString="Zhang K., Wang D., Shen H., Qian J., Guan H., Wu H. & Gao Y. 2017. Redescription of Platevindex mortoni (Gastropoda: Eupulmonata: Onchidiidae) from China. Molluscan Research 37: 72 - 78. https: // doi. org / 10.1080 / 13235818.2016.1223535"type="journal article"year="2017">
<taxonomicNameid="4C0F18DDFFF98B37DC40F8C9FC3FD723"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[888,1020,1894,1919]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFF98B37DF85F80AFE07D79C"author="Plate L. H."box="[189,452,1957,1984]"pageId="101"pageNumber="102"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate (1893: 196–199)</bibRefCitation>
did not comment on the very close similarity between
<taxonomicNameid="4C0F18DDFFF98B37DFC9F866FEBBD7BE"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[241,376,1993,2018]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="101"pageNumber="102"phylum="Mollusca"rank="species"species="martensi">
<bibRefCitationid="EF9E1EAFFFFA8B34DE69FEA6FDD2D17F"author="Plate L. H."box="[337,529,265,291]"pageId="102"pageNumber="103"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate 1893: 199</bibRefCitation>
). This may be due to a similarity in their hyponotum color; the preserved hyponota of both
<bibRefCitationid="EF9E1EAFFFFA8B34DC04FEDDFBDFD1D1"author="Plate L. H."box="[828,1052,370,397]"pageId="102"pageNumber="103"pagination="93 - 234"refId="ref73697"refString="Plate L. H. 1893. Studien ʾ ber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbucher, Abtheilung fur Anatomie und Ontogenie der Thiere 7: 93 - 234. Available from https: // biodiversitylibrary. org / page / 11175213 [accessed 27 Sep. 2016]."type="journal article"year="1893">Plate’s (1893: 193)</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFFA8B34DC6FFE16FB96D18F"author="Hoffmann H."box="[855,1109,441,467]"pageId="102"pageNumber="103"pagination="29 - 118"refId="ref72283"refString="Hoffmann H. 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbucher (Jena) 55: 29 - 118."type="journal article"year="1928">Hoffmann (1928: 88)</bibRefCitation>
and
<bibRefCitationid="EF9E1EAFFFFA8B34DBA9FE16FAB1D188"author="Labbe A."box="[1169,1394,441,468]"pageId="102"pageNumber="103"pagination="173 - 246"refId="ref72690"refString="Labbe A. 1934. Les Silicodermes (Labbe) du Museum d'Histoire naturelle de Paris. Premiere partie: Classification, formes nouvelles ou peu connues. Annales de l'Institut oceanographique 14: 173 - 246."type="journal article"year="1934">Labbé (1934: 235)</bibRefCitation>
<bibRefCitationid="EF9E1EAFFFFA8B34DEEDFE73FD60D1AA"author="Westerlund C. A."box="[469,675,476,502]"pageId="102"pageNumber="103"pagination="164 - 166"refId="ref75434"refString="Westerlund C. A. 1883. Noch einige von der Vega-Expedition gesammelte Mollusken. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 15: 164 - 166."type="journal article"year="1883">Westerlund, 1883</bibRefCitation>
</taxonomicName>
actually belongs to the genus
<taxonomicNameid="4C0F18DDFFFA8B34DB38FE73FAB1D1AB"authority="(Dayrat et al. 2016)"baseAuthorityName="Dayrat"baseAuthorityYear="2016"box="[1024,1394,476,503]"class="Gastropoda"family="Onchidiidae"genus="Onchidium"kingdom="Animalia"order="Systellommatophora"pageId="102"pageNumber="103"phylum="Mollusca"rank="genus">
<bibRefCitationid="EF9E1EAFFFFA8B34DBAAFE73FAA8D1AB"author="Dayrat B. & Goulding T. C. & Apte D. & Bhave V. & Comendador J. & Ngo X. Q. & Tan S. K. & Tan S. H."box="[1170,1387,476,503]"pageId="102"pageNumber="103"pagination="1 - 40"refId="ref70682"refString="Dayrat B., Goulding T. C., Apte D., Bhave V., Comendador J., Ngo X. Q., Tan S. K. & Tan S. H. 2016. Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca: Gastropoda: Pulmonata: Onchidiidae). ZooKeys 636: 1 - 40. https: // doi. org / 10.3897 / zookeys. 636.8879"type="journal article"year="2016">
<taxonomicNameid="4C0F18DDFFFA8B34DE5AFDB1FD9ED26B"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[354,605,542,568]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="102"pageNumber="103"phylum="Mollusca"rank="species"species="martensi">
slug with elongated dorsal ridges similar to those found in
<taxonomicNameid="4C0F18DDFFFA8B34DBCBFD27FAB5D2FD"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[1267,1398,648,673]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="102"pageNumber="103"phylum="Mollusca"rank="species"species="martensi">
<taxonomicNameid="4C0F18DDFFFA8B34DFC9FD04FE89D298"authority="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat, 2021"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"box="[241,330,683,708]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="102"pageNumber="103"phylum="Mollusca"rank="species"species="aptei"status="sp. nov.">
Also, its description mentions that it is an “orange footed mangrove onch slug.” Because no photograph of the ventral surface is available, the identification of that slug cannot be confirmed. Until this geographic record is confirmed by the examination of actual specimens,
is not considered part of the geographic distribution for
<taxonomicNameid="4C0F18DDFFFA8B34DC75FCBAFC12D372"authorityName="Goulding & Bourke & Comendador & Khalil & Quang & Tan & Tan & Dayrat"authorityYear="2021"baseAuthorityName="Plate"baseAuthorityYear="1893"box="[845,977,789,814]"class="Gastropoda"family="Onchidiidae"genus="Platevindex"higherTaxonomySource="GBIF"kingdom="Animalia"order="Systellommatophora"pageId="102"pageNumber="103"phylum="Mollusca"rank="species"species="martensi">
