<documentid="88C3B79CCB944CE91A7D42AA21F55EBB"ID-DOI="10.11646/zootaxa.5583.2.3"ID-ISSN="1175-5326"ID-Zenodo-Dep="14798833"ID-ZooBank="FC4C2C8C-B379-4E7C-9FBB-116783DDD4A0"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1738625443653"checkinUser="plazi"docAuthor="Heppner, J. B., Arita, Yutaka & Bae, Yang-Seop"docDate="2025"docId="03946562FFC8FF94FF1562EE616AF8A8"docLanguage="en"docName="zootaxa.5583.2.3.pdf"docOrigin="Zootaxa 5583 (2)"docSource="https://doi.org/10.11646/zootaxa.5583.2.3"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Thaumatographa fennchihwu Heppner & Arita 2025, new species"docType="treatment"docVersion="2"lastPageNumber="283"masterDocId="FFAD1D1AFFC2FF98FF8264576327FFD9"masterDocTitle="Review of Thaumatographa tortricids in Taiwan (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini)"masterLastPageNumber="292"masterPageNumber="271"pageNumber="281"updateTime="1738628064399"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="F137393D632A4B31FA7E4641F1F8B949">Review of Thaumatographa tortricids in Taiwan (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini)</mods:title>
<mods:namePartid="37DA922DA72C997529F03360B681A8EA">Heppner, J. B.</mods:namePart>
<mods:affiliationid="50038A3F2F90FF2F613AFE1265F914D2">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA. & Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:affiliationid="1A6F854565F87C25031A579CE176DF8B">Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. & Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
<taxonomicNameid="4C3DAFF7FFC8FF92FF1562EE61C3F90D"authority="Heppner & Arita"authorityName="Heppner & Arita"authorityYear="2025"box="[151,740,1721,1748]"class="Insecta"family="Tortricidae"genus="Thaumatographa"kingdom="Animalia"order="Lepidoptera"pageId="10"pageNumber="281"phylum="Arthropoda"rank="species"species="fennchihwu"status="sp. nov.">
<figureCitationid="1306C8F1FFC8FF92FF1D6355622CF8C5"box="[159,267,1794,1820]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="10"pageNumber="281">Figs. 3–4</figureCitation>
<figureCitationid="1306C8F1FFC8FF92FE9563556213F8C5"box="[279,308,1794,1820]"captionStart="FIGURE 11"captionStartId="9.[151,250,1310,1334]"captionTargetBox="[151,1437,181,1287]"captionTargetId="figure-20@9.[151,1437,181,1287]"captionTargetPageId="9"captionText="FIGURE 11. Thaumatographa fennchihwu Heppner & Arita, sp. nov., holotype male (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus enlarged (a) and with ductus ejaculatorius (with typical proximal elongated hood found in many Thaumatographa species), with enlargement of uncus to juxta (b), and structures of the pregenital abdomen and coremata (c) (gen. slide JBH- 4646) (MGCL)."figureDoi="http://doi.org/10.5281/zenodo.14798845"httpUri="https://zenodo.org/record/14798845/files/figure.png"pageId="10"pageNumber="281">11</figureCitation>
<figureCitationid="1306C8F1FFC8FF92FEC26355627AF8C5"box="[320,349,1794,1820]"captionStart="FIGURE 14"captionStartId="13.[151,250,1341,1365]"captionTargetBox="[152,1436,181,1317]"captionTargetId="figure-20@13.[151,1436,181,1317]"captionTargetPageId="13"captionText="FIGURE 14. Thaumatographa fennchihwu Heppner & Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)."figureDoi="http://doi.org/10.5281/zenodo.14798851"httpUri="https://zenodo.org/record/14798851/files/figure.png"pageId="10"pageNumber="281">14</figureCitation>
<materialsCitationid="3B55DE29FFC9FF93FE4164EC605DFF0C"box="[451,890,187,213]"collectionCode="MGCL"collectorName="J. B. Heppner"country="Taiwan"elevation="400"location="Wulai"pageId="11"pageNumber="282"specimenCount="1"stateProvince="Taipei">
<collectorNameid="26C8B1A2FFC9FF93FE4164EC617BFF0C"box="[451,604,187,213]"pageId="11"pageNumber="282">J. B. Heppner</collectorName>
<collectingDateid="EFC70B5CFFC9FF93FC0464EC6739FF0C"box="[902,1054,186,213]"pageId="11"pageNumber="282"value="1985-08-14">14 Aug. 1985</collectingDate>
<collectionCodeid="ED2C4CB1FFC9FF93FB7664EC666FFF0C"box="[1268,1352,187,213]"country="Japan"httpUri="http://grbio.org/cool/f678-ci92"name="National Science Museum (Natural History)"pageId="11"pageNumber="282">NSMT</collectionCode>
<taxonomicNameid="4C3DAFF7FFC9FF93FEAF61F861D0FA1E"authority="Diakonoff"authorityName="Diakonoff & Arita"authorityYear="1977"box="[301,759,1455,1479]"class="Insecta"family="Tortricidae"genus="Thaumatographa"kingdom="Animalia"order="Lepidoptera"pageId="11"pageNumber="282"phylum="Arthropoda"rank="species"species="mesostigmatias">
, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL).
<figureCitationid="1306C8F1FFC9FF93FD2362696028F981"box="[673,783,1598,1624]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="11"pageNumber="282">Figs. 3–4</figureCitation>
), with forewings black-brown with orange-marked striae, and a large orange tornal patch with two black spots; the hindwings are black-brown with a prominent yellow patch on middle of wing to base and a narrow cubital narrow pale orange patch, and fringe is prominent and white; and both sexes with equal maculation. The male genitalia have the valvae oblong-quadratic, with a truncated termen, the uncus is short with a wide base, the hami are short and somewhat flattened, and the socii are short, wide and very setose. The female genitalia are typical for the genus but the ductus bursae has a large diverticular sac on its caudal end.
<figureCitationid="1306C8F1FFC9FF93FE9563096281F8A1"box="[279,422,1886,1912]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="11"pageNumber="282">Figs. 3a, 4a</figureCitation>
): dark brown, with orange-yellow scales laterally by neck and medially on vertex and frons; eyes large, black-brown; ocellus large, dark brown; antennal scape amber to brown; antenna brown (orange ventrally); labial palpus mostly white, with brown to yellow dorsally on basal segment and somewhat on middle segment, with bluish iridescence on apical segment; head venter white, with neck fringe yellow to orange-yellow.
<figureCitationid="1306C8F1FFCEFF94FEA564C062B3FF68"box="[295,404,151,177]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="12"pageNumber="283">Figs. 3–4</figureCitation>
): dark brown, with light orange as lateral margin on prothorax, becoming more tawny orange on mesothorax and metathorax, with orange caudal margin; patagia dark brown with orange from prothoax; tegula dark brown with orange laterally; venter white; legs white, with black-brown markings, as in the related species.
<figureCitationid="1306C8F1FFCEFF94FEC065546297FEC4"box="[322,432,259,285]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="12"pageNumber="283">Figs. 3–4</figureCitation>
): black-brown, with orange striae and lines from costa and dorsal margin (in generic form like
<taxonomicNameid="4C3DAFF7FFCEFF94FF48657062B3FE99"authorityName="Diakonoff & Arita"authorityYear="1977"box="[202,404,295,320]"class="Insecta"family="Tortricidae"genus="Thaumatographa"kingdom="Animalia"order="Lepidoptera"pageId="12"pageNumber="283"phylum="Arthropoda"rank="species"species="mesostigmatias">
but darker orange); five costal bars angled to termen, with pale orange-yellow on margin, then silvery toward middle of wing, and dull orange suffusion on dorsal 1/3 of forewing from middle to apex and interrupted by black curved preapical bar; forewing apex with an orange marginal line, sharply bent inwards before a pale yellow to white bar near mid-termen; tornus with a large elongated marginal orange patch with two black spots in middle and a pair of tiny black spots dorsad near white mark of termen; fringe brown with yellow at tornus and near mid-termen; venter dark brown with pale yellow in cell area and marks of dorsum very faintly repeated, except distinct on costa, with dorsal margin white.
<figureCitationid="1306C8F1FFCEFF94FEC56674629FFDE4"box="[327,440,547,573]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="12"pageNumber="283">Figs. 3–4</figureCitation>
): black-brown with a large oblong orange-yellow patch, narrowing to wingbase, and a similar more orange and narrower patch on the cubitus near wingbase; costa white; fringe widely white to pale yellow at wing margin, and pale orange along anal margin; venter like dorsum but more suffused, and costa brown.
<figureCitationid="1306C8F1FFCEFF94FEC266D8628AFD70"box="[320,429,655,681]"captionStart="FIGURES 1–6"captionStartId="4.[151,264,1560,1584]"captionTargetBox="[151,1436,555,1536]"captionTargetId="figure-133@4.[151,1436,555,1536]"captionTargetPageId="4"captionText="FIGURES 1–6. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff & Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner & Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner & Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a."figureDoi="http://doi.org/10.5281/zenodo.14798835"httpUri="https://zenodo.org/record/14798835/files/figure.png"pageId="12"pageNumber="283">Figs. 3–4</figureCitation>
): brown (dark brown on T-1), with orange posterior margins on tergites; venter tan; genital tufts light brown with tan-white venter; pregenital segment with sternite strongly invaginated caudally, with strong coremata from lateral base pouches; caudal tergite with similar median invagination.
<figureCitationid="1306C8F1FFCEFF94FE0266AC62F1FCCC"box="[384,470,763,789]"captionStart="FIGURE 11"captionStartId="9.[151,250,1310,1334]"captionTargetBox="[151,1437,181,1287]"captionTargetId="figure-20@9.[151,1437,181,1287]"captionTargetPageId="9"captionText="FIGURE 11. Thaumatographa fennchihwu Heppner & Arita, sp. nov., holotype male (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus enlarged (a) and with ductus ejaculatorius (with typical proximal elongated hood found in many Thaumatographa species), with enlargement of uncus to juxta (b), and structures of the pregenital abdomen and coremata (c) (gen. slide JBH- 4646) (MGCL)."figureDoi="http://doi.org/10.5281/zenodo.14798845"httpUri="https://zenodo.org/record/14798845/files/figure.png"pageId="12"pageNumber="283">Fig. 11</figureCitation>
): tegumen quadratic, slightly narrowed dorsally with rounded dorsal corners; uncus short, narrowing to semi-acute decumbent apex from widened base; hamus short and widened, somewhat flattened; socius short but very wide and setose; gnathos membranous and rounded; transtilla relatively straight and narrow, with moderate ventral appendage each side of anellus with acute end; juxta small and spoon-like; anellus membranous, undeveloped; aedeagus small and short tubular, with small decumbent phallobase; vesica scobinate; cornuti undeveloped; ductus ejaculatorius short, with long proximal hood appendage; valva oblong quadratic, with round-blunt termen and dorsal apex slightly angular and more setose; saccular margin slightly convex near base; vinculum triangular; saccus very short, with blunt apex.
<figureCitationid="1306C8F1FFCEFF94FE11604C62C1FBEC"box="[403,486,1051,1077]"captionStart="FIGURE 14"captionStartId="13.[151,250,1341,1365]"captionTargetBox="[152,1436,181,1317]"captionTargetId="figure-20@13.[151,1436,181,1317]"captionTargetPageId="13"captionText="FIGURE 14. Thaumatographa fennchihwu Heppner & Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)."figureDoi="http://doi.org/10.5281/zenodo.14798851"httpUri="https://zenodo.org/record/14798851/files/figure.png"pageId="12"pageNumber="283">Fig. 14</figureCitation>
): ovipositor very short (2/3 length of segment 7), with elongate-oval horizontal dorsal median flap; papilla anales large, setose and widened caudally; apophyses subequal in length; ostium (
<figureCitationid="1306C8F1FFCEFF94FA986068665CFB80"box="[1306,1403,1087,1113]"captionStart="FIGURE 14"captionStartId="13.[151,250,1341,1365]"captionTargetBox="[152,1436,181,1317]"captionTargetId="figure-20@13.[151,1436,181,1317]"captionTargetPageId="13"captionText="FIGURE 14. Thaumatographa fennchihwu Heppner & Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)."figureDoi="http://doi.org/10.5281/zenodo.14798851"httpUri="https://zenodo.org/record/14798851/files/figure.png"pageId="12"pageNumber="283">Fig. 14a</figureCitation>
), a deep funnel, gradually narrowing to junction with ductus bursae, with a band of sclerotized scobination on narrower section; sterigma a concave membranous margin; ductus bursae long and evenly narrow, with a prominent ventral diverticular sac (or small pouch) with internally spiculate walls and bifid distally from junction with antrum; ductus seminalis from ductus bursae proximal end; corpus bursae oblong-ovate, with one end more acute; oblong-ovate accessory diverticulum from thin tube to near ductus bursae base; signum a large typical form for the genus.
<bibRefCitationid="EFACA985FFCEFF94FAB6616C63C7FAA0"author="Heppner, J. B."pageId="12"pageNumber="283"pagination="85 - 106"refId="ref11384"refString="Heppner, J. B. (2012 b) Historical review of Lepidoptera studies in Taiwan. Lepidoptera Novae, 5, 85-106, 1 pl."type="journal article"year="2012">Heppner, 2012b</bibRefCitation>
, c, d). This locality, Fennchihwu, was formerly a Japanese logging camp and a railway stop on the Alishan rail line going higher to the town of Alishan, on the slopes of the high mountain of the same name (Mt. Ali portion of Yushan): this mountain has been heavily logged for over 100 years but small patches of original forest still remain and are now expanding with new conservation efforts.
will be distributed also to other museum collections (RMNH, TFRI, USNM). The species is one of the larger species of the genus. Its habitus superficially resembles species of the tropical Asian tortricid genus
<bibRefCitationid="EFACA985FFCEFF94FF1563586268F8F0"author="Diakonoff, A."box="[151,335,1806,1833]"pageId="12"pageNumber="283"pagination="297 - 320"refId="ref10870"refString="Diakonoff, A. (1975) New Tortricoidea (Lepidoptera) from Southeast Asia in the British Museum (Natural History). Zoologische Mededelingen, Leiden, 48, 297-320, 8 pls."type="book chapter"year="1975">Diakonoff, 1975</bibRefCitation>
), which also have the yellow-patched hindwing, but differ in forewing maculation. The bifid ventral diverticulum of the caudal end on the ductus bursae has no known function but is possibly some kind of gland apparatus.
