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<document id="AC2C7A1087FF5CF1DB96A35404B3BEB7" ID-DOI="10.11646/zootaxa.5583.2.2" ID-ISSN="1175-5326" ID-ZooBank="E5168208-01D7-4218-BE79-67B2E0E61E5E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738625326105" checkinUser="plazi" docAuthor="Vasanth, M., Kubendran, T., Subramanian, K. A. &amp; Babu, R." docDate="2025" docId="034C87EDFFEDFF84D8B5B5FC8E27FC14" docLanguage="en" docName="zootaxa.5583.2.2.pdf" docOrigin="Zootaxa 5583 (2)" docSource="https://doi.org/10.11646/zootaxa.5583.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thalerosphyrus flowersi Venkataraman &amp; Sivaramakrishnan 1987" docType="treatment" docVersion="2" lastPageNumber="266" masterDocId="FF75FF95FFFFFF97D822B4578A12FFE7" masterDocTitle="Two new species of the genus Thalerosphyrus Eaton, 1881 (Ephemeroptera: Heptageniidae: Ecdyonurinae) from India" masterLastPageNumber="270" masterPageNumber="247" pageNumber="265" updateTime="1738625987165" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="48549B68E649F207254472DC2F0D9023">Two new species of the genus Thalerosphyrus Eaton, 1881 (Ephemeroptera: Heptageniidae: Ecdyonurinae) from India</mods:title>
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<mods:namePart id="782A3DABF10ED3743196546E8E85F921">Vasanth, M.</mods:namePart>
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<mods:namePart id="F5E00DF6E11D07BC1F832957C25B6499">Kubendran, T.</mods:namePart>
<mods:affiliation id="299BD4D9F43F279DFA42185C19FFD394">Zoological Survey of India, Prani Vigyan Bhavan, M-Block, New Alipore, Kolkata- 700 053, India.</mods:affiliation>
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<mods:namePart id="9D1FF951CCDDDF5BB02DA7DF1359236A">Subramanian, K. A.</mods:namePart>
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<mods:affiliation id="73769BDD27A5ABB2AFB3B101E026792F">Southern Regional Centre, Zoological Survey of India, Chennai- 600 028, India.</mods:affiliation>
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<mods:namePart id="BE08F35F79FB6A8E13DC6514F1B04A65">Babu, R.</mods:namePart>
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<mods:affiliation id="5459B8C192C56777333D7B1A1800442C">Southern Regional Centre, Zoological Survey of India, Chennai- 600 028, India.</mods:affiliation>
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<mods:date id="53AF4EA999ED6A7E0FE1C80348E8C881">2025</mods:date>
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<paragraph id="8B5A36FBFFEDFF85D8B5B5FC89CBFE21" blockId="18.[151,985,427,490]" box="[151,985,427,454]" pageId="18" pageNumber="265">
<heading id="D0128197FFEDFF85D8B5B5FC89CBFE21" bold="true" box="[151,985,427,454]" fontSize="11" level="1" pageId="18" pageNumber="265" reason="1">
<taxonomicName id="4CE54D78FFEDFF85D8B5B5FC89CBFE21" ID-CoL="55ZWP" authority="Venkataraman &amp; Sivaramakrishnan, 1987" authorityName="Venkataraman &amp; Sivaramakrishnan" authorityYear="1987" box="[151,985,427,454]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="265" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B991EAE9FFEDFF85D8B5B5FC89CBFE21" bold="true" box="[151,985,427,454]" pageId="18" pageNumber="265">
<emphasis id="B991EAE9FFEDFF85D8B5B5FC8BAFFE21" bold="true" box="[151,445,427,454]" italics="true" pageId="18" pageNumber="265">Thalerosphyrus flowersi</emphasis>
<bibRefCitation id="EF744B0AFFEDFF85D9E7B5FB89CBFE21" author="Venkataraman, K. &amp; Sivaramakrishnan, K. G." box="[453,985,428,454]" pageId="18" pageNumber="265" pagination="1126 - 1129" refId="ref8076" refString="Venkataraman, K. &amp; Sivaramakrishnan, K. G. (1987) A new species of Thalerosphyrus for South India (Ephemeroptera: Heptageniidae). Current Science, 56 (21), 1126-1129." type="journal article" year="1987">Venkataraman &amp; Sivaramakrishnan, 1987</bibRefCitation>
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<subSubSection id="C3FF6570FFEDFF85D8B5B5878B26FE0D" box="[151,308,464,490]" pageId="18" pageNumber="265" type="description">
<paragraph id="8B5A36FBFFEDFF85D8B5B5878B26FE0D" blockId="18.[151,985,427,490]" box="[151,308,464,490]" pageId="18" pageNumber="265">
(
<figureCitation id="13DE2A7EFFEDFF85D8BDB5878B3EFE0D" box="[159,300,464,490]" captionStart-0="FIGURES 96103" captionStart-1="FIGURES 104110" captionStartId-0="17.[151,264,1883,1907]" captionStartId-1="18.[151,264,1982,2006]" captionTargetBox-0="[151,1435,181,1859]" captionTargetBox-1="[311,1276,910,1958]" captionTargetId-0="figure-19@17.[151,1435,181,1859]" captionTargetId-1="figure-267@18.[311,1276,910,1958]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 96103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." captionText-1="FIGURES 104110. Larva of Thalerosphyrus flowersi: 104. Maxillae; 105. Galea-lacinea closer view; 106. Claw. 107. Gill I; 108. Protopenis; 109. Tergum V posterolateral spine; 110. Type species label." pageId="18" pageNumber="265">Figs 96110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFEDFF85D8B5B64F88EEFDB1" pageId="18" pageNumber="265" type="materials_examined">
<paragraph id="8B5A36FBFFEDFF85D8B5B64F88EEFDB1" blockId="18.[151,1437,536,887]" pageId="18" pageNumber="265">
<emphasis id="B991EAE9FFEDFF85D8B5B64F8B96FDD5" bold="true" box="[151,388,536,562]" pageId="18" pageNumber="265">Material examined.</emphasis>
<materialsCitation id="3B8D3CA6FFEDFF85D9ACB64F88EAFDB1" country="India" county="Theni district" elevation="397" latitude="10.18025" location="Kumbakkarai falls" longLatPrecision="1" longitude="77.53047" municipality="Palni" pageId="18" pageNumber="265" specimenCount="397" stateProvince="Tamil Nadu" typeStatus="paratype">
<emphasis id="B991EAE9FFEDFF85D9ACB64F8816FDD5" bold="true" box="[398,516,536,562]" pageId="18" pageNumber="265">
<typeStatus id="545E8859FFEDFF85D9ACB64F8BEDFDD5" box="[398,511,536,562]" pageId="18" pageNumber="265" type="paratype">Paratype</typeStatus>
:
</emphasis>
<collectingCountry id="F3F2766BFFEDFF85DA2CB64F8858FDD5" box="[526,586,536,562]" name="India" pageId="18" pageNumber="265">India</collectingCountry>
,
<collectingRegion id="4921F819FFEDFF85DA7AB64F88F0FDD5" box="[600,738,536,562]" country="India" name="Tamil Nadu" pageId="18" pageNumber="265">Tamil Nadu</collectingRegion>
,
<collectingCounty id="623B4E77FFEDFF85DAD2B64F899EFDD5" box="[752,908,536,562]" pageId="18" pageNumber="265">Theni district</collectingCounty>
,
<collectingMunicipality id="6B3EAC81FFEDFF85DBB8B64F89C6FDD5" box="[922,980,536,562]" pageId="18" pageNumber="265">Palni</collectingMunicipality>
hills,
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,
<geoCoordinate id="EED1503CFFEDFF85DD21B64F8F8AFDD4" box="[1283,1432,536,563]" degrees="10.18025" direction="north" orientation="latitude" pageId="18" pageNumber="265" precision="1" value="10.18025">10.18025° N</geoCoordinate>
,
<geoCoordinate id="EED1503CFFEDFF85D8B5B66B8B37FDB0" box="[151,293,572,599]" degrees="77.53047" direction="east" orientation="longitude" pageId="18" pageNumber="265" precision="1" value="77.53047">77.53047° E</geoCoordinate>
,
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<quantity id="4C1D9B1EFFEDFF85D913B66B8B69FDB0" box="[305,379,572,599]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="18" pageNumber="265" unit="m" value="397.0">
<elevation id="00C8D1C8FFEDFF85D913B66B8B69FDB0" box="[305,379,572,599]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="18" pageNumber="265" unit="m" value="397.0">397 m</elevation>
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.
</specimenCount>
(Type specimen label (
<figureCitation id="13DE2A7EFFEDFF85DAA7B66B88F7FDB1" box="[645,741,572,598]" captionStart="FIGURES 104110" captionStartId="18.[151,264,1982,2006]" captionTargetBox="[311,1276,910,1958]" captionTargetId="figure-267@18.[311,1276,910,1958]" captionTargetPageId="18" captionText="FIGURES 104110. Larva of Thalerosphyrus flowersi: 104. Maxillae; 105. Galea-lacinea closer view; 106. Claw. 107. Gill I; 108. Protopenis; 109. Tergum V posterolateral spine; 110. Type species label." pageId="18" pageNumber="265">Fig. 110</figureCitation>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFEDFF85D8E5B6378807FD9D" box="[199,533,608,634]" pageId="18" pageNumber="265" type="distribution">
<paragraph id="8B5A36FBFFEDFF85D8E5B6378807FD9D" blockId="18.[151,1437,536,887]" box="[199,533,608,634]" pageId="18" pageNumber="265">
<emphasis id="B991EAE9FFEDFF85D8E5B6378B73FD9D" bold="true" box="[199,353,608,634]" pageId="18" pageNumber="265">Distribution.</emphasis>
Southern
<collectingCountry id="F3F2766BFFEDFF85D9F6B6378803FD9D" box="[468,529,608,634]" name="India" pageId="18" pageNumber="265">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFEDFF84D8E5B6D38E27FC14" lastPageId="19" lastPageNumber="266" pageId="18" pageNumber="265" type="diagnosis">
<paragraph id="8B5A36FBFFEDFF85D8E5B6D38E04FC91" blockId="18.[151,1437,536,887]" pageId="18" pageNumber="265">
<emphasis id="B991EAE9FFEDFF85D8E5B6D38B50FD79" bold="true" box="[199,322,644,670]" pageId="18" pageNumber="265">Diagnosis.</emphasis>
<taxonomicName id="4CE54D78FFEDFF85D968B6D38849FD79" authorityName="Venkataraman &amp; Sivaramakrishnan" authorityYear="1987" box="[330,603,644,670]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="265" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B991EAE9FFEDFF85D968B6D38849FD79" box="[330,603,644,670]" italics="true" pageId="18" pageNumber="265">Thalerosphyrus flowersi</emphasis>
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by the following combination of characters: (i) labrum greatly expanded laterally and shallow medially (
<figureCitation id="13DE2A7EFFEDFF85DA22B6FF8845FD25" box="[512,599,680,706]" captionStart="FIGURES 96103" captionStartId="17.[151,264,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-19@17.[151,1435,181,1859]" captionTargetPageId="17" captionText="FIGURES 96103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." pageId="18" pageNumber="265">Fig. 99</figureCitation>
); (ii) gill I asymmetrical and ovoid, 1.5× as long as wide (
<figureCitation id="13DE2A7EFFEDFF85DCDAB6FF8F4EFD25" box="[1272,1372,680,706]" captionStart="FIGURES 104110" captionStartId="18.[151,264,1982,2006]" captionTargetBox="[311,1276,910,1958]" captionTargetId="figure-267@18.[311,1276,910,1958]" captionTargetPageId="18" captionText="FIGURES 104110. Larva of Thalerosphyrus flowersi: 104. Maxillae; 105. Galea-lacinea closer view; 106. Claw. 107. Gill I; 108. Protopenis; 109. Tergum V posterolateral spine; 110. Type species label." pageId="18" pageNumber="265">Fig. 107</figureCitation>
); (iii) femora of legs yellowish brown with zig-zag pale yellow band; (iv) glossae clearly concave with surface three long setae (
<figureCitation id="13DE2A7EFFEDFF85D8FCB6A78B52FCED" box="[222,320,752,778]" captionStart="FIGURES 96103" captionStartId="17.[151,264,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-19@17.[151,1435,181,1859]" captionTargetPageId="17" captionText="FIGURES 96103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." pageId="18" pageNumber="265">Fig. 101</figureCitation>
); (v) maxillae of galea-lacinia with 1718 comb-shaped setae (
<figureCitation id="13DE2A7EFFEDFF85DC22B6A78E8EFCEC" box="[1024,1180,752,779]" captionStart="FIGURES 104110" captionStartId="18.[151,264,1982,2006]" captionTargetBox="[311,1276,910,1958]" captionTargetId="figure-267@18.[311,1276,910,1958]" captionTargetPageId="18" captionText="FIGURES 104110. Larva of Thalerosphyrus flowersi: 104. Maxillae; 105. Galea-lacinea closer view; 106. Claw. 107. Gill I; 108. Protopenis; 109. Tergum V posterolateral spine; 110. Type species label." pageId="18" pageNumber="265">Figs 104105</figureCitation>
) (vi) pronotum greatly enlarged laterally and posteriorly (
<figureCitation id="13DE2A7EFFEDFF85DA37B743887BFCC9" box="[533,617,788,814]" captionStart="FIGURES 96103" captionStartId="17.[151,264,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-19@17.[151,1435,181,1859]" captionTargetPageId="17" captionText="FIGURES 96103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." pageId="18" pageNumber="265">Fig. 96</figureCitation>
); (vii) ornamentation of hind tibia with 1314 pointed bristles in marginal or submarginal position; (viii) posterolateral expansion not developed on tergum I, weakly developed on tergum II, strongly developed on tergum III and increasing in size up to VIII (
<figureCitation id="13DE2A7EFFEDFF85DBA5B70B8E18FC91" box="[903,1034,860,886]" captionStart="FIGURES 96103" captionStartId="17.[151,264,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-19@17.[151,1435,181,1859]" captionTargetPageId="17" captionText="FIGURES 96103. Larva of Thalerosphyrus flowersi: 96. Dorsal view; 97. Abdomen ventral view; 98. Abdomen dorsal view; 99. Labrum; 100. Labium; 101. Glossae closer view (arrow indicates long setae); 102. Left mandible; 103. Right mandible." pageId="18" pageNumber="265">Figs 9798</figureCitation>
).
</paragraph>
<caption id="DF9A6673FFEDFF85D8B5B3E989B5F81D" pageId="18" pageNumber="265" startId="18.[151,264,1982,2006]" targetBox="[311,1276,910,1958]" targetPageId="18" targetType="figure">
<paragraph id="8B5A36FBFFEDFF85D8B5B3E989B5F81D" blockId="18.[151,1436,1982,2042]" pageId="18" pageNumber="265">
<emphasis id="B991EAE9FFEDFF85D8B5B3E98B7FF831" bold="true" box="[151,365,1982,2006]" pageId="18" pageNumber="265">FIGURES 104110.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFEDFF85D9F3B3E988DAF831" authorityName="Venkataraman &amp; Sivaramakrishnan" authorityYear="1987" box="[465,712,1982,2006]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="265" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B991EAE9FFEDFF85D9F3B3E988DAF831" box="[465,712,1982,2006]" italics="true" pageId="18" pageNumber="265">Thalerosphyrus flowersi</emphasis>
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: 104. Maxillae; 105. Galea-lacinea closer view; 106. Claw. 107. Gill I; 108. Protopenis; 109. Tergum V posterolateral spine; 110. Type species label.
</paragraph>
</caption>
<caption id="DF9A6673FFECFF84D8B5B7718BDFFC84" pageId="19" pageNumber="266" startId="19.[151,264,806,830]" targetBox="[313,1274,183,780]" targetPageId="19" targetType="figure">
<paragraph id="8B5A36FBFFECFF84D8B5B7718BDFFC84" blockId="19.[151,1436,806,867]" pageId="19" pageNumber="266">
<emphasis id="B991EAE9FFECFF84D8B5B7718B7BFCD9" bold="true" box="[151,361,806,830]" pageId="19" pageNumber="266">FIGURES 111114.</emphasis>
Gill I of dorsal plate: 111.
<taxonomicName id="4CE54D78FFECFF84DAA2B77188E8FCD9" authorityName="Venkataraman &amp; Sivaramakrishnan" authorityYear="1987" box="[640,762,806,830]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="266" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B991EAE9FFECFF84DAA2B77188E8FCD9" box="[640,762,806,830]" italics="true" pageId="19" pageNumber="266">Th. flowersi</emphasis>
</taxonomicName>
(Paratype); 112.
<taxonomicName id="4CE54D78FFECFF84DB89B7718E37FCD9" authorityName="Venkataraman &amp; Sivaramakrishnan" authorityYear="1987" box="[939,1061,806,830]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="266" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B991EAE9FFECFF84DB89B7718E37FCD9" box="[939,1061,806,830]" italics="true" pageId="19" pageNumber="266">Th. flowersi</emphasis>
</taxonomicName>
(Thailand); 113.
<taxonomicName id="4CE54D78FFECFF84DCF5B7718F65FCD9" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[1239,1399,806,830]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="266" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFECFF84DCF5B7718F65FCD9" box="[1239,1399,806,830]" italics="true" pageId="19" pageNumber="266">Th. bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFECFF84DD5CB7708AD1FC84" bold="true" pageId="19" pageNumber="266">
<taxonomicNameLabel id="A2A25792FFECFF84DD5CB7708AD1FC84" pageId="19" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 114.
<taxonomicName id="4CE54D78FFECFF84D921B71D8B64FC85" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[259,374,842,866]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="266" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFECFF84D921B71D8B64FC85" box="[259,374,842,866]" italics="true" pageId="19" pageNumber="266">Th. sartorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2A25792FFECFF84D95EB71C8BDFFC84" box="[380,461,843,867]" pageId="19" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B5A36FBFFECFF84D8E5B7E38E27FC14" blockId="19.[151,1436,948,1011]" pageId="19" pageNumber="266">
<emphasis id="B991EAE9FFECFF84D8E5B7E38B20FC29" bold="true" box="[199,306,948,974]" pageId="19" pageNumber="266">Genitals.</emphasis>
Protopenis (
<figureCitation id="13DE2A7EFFECFF84D9E2B7E28809FC28" box="[448,539,949,975]" captionStart="FIGURES 104110" captionStartId="18.[151,264,1982,2006]" captionTargetBox="[311,1276,910,1958]" captionTargetId="figure-267@18.[311,1276,910,1958]" captionTargetPageId="18" captionText="FIGURES 104110. Larva of Thalerosphyrus flowersi: 104. Maxillae; 105. Galea-lacinea closer view; 106. Claw. 107. Gill I; 108. Protopenis; 109. Tergum V posterolateral spine; 110. Type species label." pageId="19" pageNumber="266">Fig.108</figureCitation>
): male genitalia yellowish, penis fused, apically blunt, and laterally with tapered square, tergum X rounded and apically with small stout setae. Forceps rectangular.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,409 @@
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<mods:title id="82B1CE06F46CABDB17C8B5746DC1ECFB">Two new species of the genus Thalerosphyrus Eaton, 1881 (Ephemeroptera: Heptageniidae: Ecdyonurinae) from India</mods:title>
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<mods:namePart id="75D216199EABC2DD1DD0D0496AEC7E94">Vasanth, M.</mods:namePart>
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<mods:namePart id="3C638874631F77AD558293B39B564549">Subramanian, K. A.</mods:namePart>
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<mods:date id="A2AF4039DA97C59996AA495CC6994E18">2025</mods:date>
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<subSubSection id="C3FF6570FFF6FF9ED8B5B05689EBFBFB" box="[151,1017,1025,1052]" pageId="9" pageNumber="256" type="nomenclature">
<paragraph id="8B5A36FBFFF6FF9ED8B5B05689EBFBFB" blockId="9.[151,1017,1025,1089]" box="[151,1017,1025,1052]" pageId="9" pageNumber="256">
<heading id="D0128197FFF6FF9ED8B5B05689EBFBFB" bold="true" box="[151,1017,1025,1052]" fontSize="11" level="1" pageId="9" pageNumber="256" reason="1">
<emphasis id="B991EAE9FFF6FF9ED8B5B05689EBFBFB" bold="true" box="[151,1017,1025,1052]" pageId="9" pageNumber="256">
<taxonomicName id="4CE54D78FFF6FF9ED8B5B0568986FBFC" authority="Vasanth, Kubendran &amp; Subramanian" authorityName="Vasanth, Kubendran &amp; Subramanian" authorityYear="2025" box="[151,916,1025,1052]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="256" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFF6FF9ED8B5B0568BA7FBFB" bold="true" box="[151,437,1025,1052]" italics="true" pageId="9" pageNumber="256">Thalerosphyrus sartorii</emphasis>
Vasanth, Kubendran &amp; Subramanian
</taxonomicName>
<taxonomicNameLabel id="A2A25792FFF6FF9EDBB9B05589EBFBFB" box="[923,1017,1026,1052]" pageId="9" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8B5A36FBFFF6FF9ED8B5B0718B35FBA6" blockId="9.[151,1017,1025,1089]" box="[151,295,1062,1089]" pageId="9" pageNumber="256">
(
<figureCitation id="13DE2A7EFFF6FF9ED8BDB0718B32FBA6" box="[159,288,1062,1089]" captionStart-0="FIGURES 5152" captionStart-1="FIGURES 5354" captionStart-2="FIGURES 5562" captionStart-3="FIGURES 6367" captionStart-4="FIGURES 6877" captionStart-5="FIGURES 7887" captionStart-6="FIGURES 8895" captionStartId-0="10.[151,264,1632,1656]" captionStartId-1="11.[151,264,1385,1409]" captionStartId-2="12.[151,264,1920,1944]" captionStartId-3="13.[151,264,1415,1439]" captionStartId-4="14.[151,264,1943,1967]" captionStartId-5="15.[151,264,1856,1880]" captionStartId-6="16.[151,264,1914,1938]" captionTargetBox-0="[164,1423,273,1609]" captionTargetBox-1="[164,1423,273,1362]" captionTargetBox-2="[151,1435,181,1897]" captionTargetBox-3="[164,1423,181,1391]" captionTargetBox-4="[184,1407,181,1919]" captionTargetBox-5="[151,1435,185,1820]" captionTargetBox-6="[151,1435,181,1890]" captionTargetId-0="figure-68@10.[164,1423,273,1609]" captionTargetId-1="figure-62@11.[164,1423,273,1362]" captionTargetId-2="figure-30@12.[151,1435,181,1897]" captionTargetId-3="figure-19@13.[164,1423,181,1391]" captionTargetId-4="figure-30@14.[179,1407,181,1919]" captionTargetId-5="figure-19@15.[151,1435,181,1833]" captionTargetId-6="figure-30@16.[151,1435,181,1890]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="14" captionTargetPageId-5="15" captionTargetPageId-6="16" captionText-0="FIGURES 5152. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." captionText-1="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." captionText-2="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" captionText-3="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." captionText-4="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." captionText-5="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." captionText-6="FIGURES 8895. Larva of Thalerosphyrus sartorii sp. nov. 88. Gill I; 89. Gill II; 90. Gill III; 91. Gill IV; 92. Gill V; 93. Gill VI; 94. Gill VII; 95. Antenna." pageId="9" pageNumber="256">Figs 5195</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF6FF9ED8B5B0398E79FB13" pageId="9" pageNumber="256" type="materials_examined">
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<emphasis id="B991EAE9FFF6FF9ED8B5B0398B96FB6F" bold="true" box="[151,388,1134,1160]" pageId="9" pageNumber="256">Material examined.</emphasis>
<materialsCitation id="3B8D3CA6FFF6FF9ED9ACB0398EBBFB4B" collectingDate="2023-02-02" collectionCode="R" collectorName="Babu &amp; Party &amp; Reg. No. I" country="India" county="Mature" elevation="314" latitude="11.5728" location="Gunderipallam" longLatPrecision="1" longitude="77.37006" municipality="Erode district" pageId="9" pageNumber="256" specimenCount="314" stateProvince="Tamil Nadu" typeStatus="holotype">
<emphasis id="B991EAE9FFF6FF9ED9ACB0398810FB6F" bold="true" box="[398,514,1134,1160]" pageId="9" pageNumber="256">
<typeStatus id="545E8859FFF6FF9ED9ACB0398BEFFB6F" box="[398,509,1134,1160]" pageId="9" pageNumber="256" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="623B4E77FFF6FF9EDA2FB039884CFB6F" box="[525,606,1134,1160]" pageId="9" pageNumber="256">Mature</collectingCounty>
larva,
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,
<collectingRegion id="4921F819FFF6FF9EDB30B039898EFB6F" box="[786,924,1134,1160]" country="India" name="Tamil Nadu" pageId="9" pageNumber="256">Tamil Nadu</collectingRegion>
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<collectingMunicipality id="6B3EAC81FFF6FF9EDB89B0398E5BFB6F" box="[939,1097,1134,1160]" pageId="9" pageNumber="256">Erode district</collectingMunicipality>
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, coll.
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<collectorName id="2610532DFFF6FF9EDB10B0C5897FFB4B" box="[818,877,1170,1196]" pageId="9" pageNumber="256">Babu</collectorName>
&amp;
<collectorName id="2610532DFFF6FF9EDBB3B0C589DEFB4B" box="[913,972,1170,1196]" pageId="9" pageNumber="256">Party</collectorName>
(
<collectorName id="2610532DFFF6FF9EDBFEB0C58E43FB4B" box="[988,1105,1170,1196]" pageId="9" pageNumber="256">Reg. No. I</collectorName>
/E/866)
</materialsCitation>
.
<materialsCitation id="3B8D3CA6FFF6FF9EDC96B0C588ECFB37" collectingDate="2023-02-02" collectionCode="R" collectorName="Reg. No. I" country="India" county="Mature" elevation="314" latitude="11.5728" location="Gunderipallam" longLatPrecision="1" longitude="77.37006" municipality="Erode district" pageId="9" pageNumber="256" specimenCount="4" specimenCount-larva="4" stateProvince="Tamil Nadu" typeStatus="paratype">
<emphasis id="B991EAE9FFF6FF9EDC96B0C58F27FB4B" bold="true" box="[1204,1333,1170,1196]" pageId="9" pageNumber="256">
<typeStatus id="545E8859FFF6FF9EDC96B0C58F22FB4B" box="[1204,1328,1170,1196]" pageId="9" pageNumber="256" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9DE3FD72FFF6FF9EDD1EB0C58F8AFB4B" box="[1340,1432,1170,1196]" count="4" pageId="9" pageNumber="256" type="larva">4 larvae</specimenCount>
, same place and data as holotype (
<collectorName id="2610532DFFF6FF9EDA0AB0E188B4FB37" box="[552,678,1206,1232]" pageId="9" pageNumber="256">Reg. No. I</collectorName>
/E/867)
</materialsCitation>
;
<materialsCitation id="3B8D3CA6FFF6FF9EDB2FB0E18818FB13" country="India" county="Kottadai" elevation="948" latitude="11.65037" location="Erode district" longLatPrecision="1" longitude="77.13364" municipality="Matheshwaram" pageId="9" pageNumber="256" specimenCount="3" specimenCount-larva="3" stateProvince="Tamil Nadu" typeStatus="paratype">
<specimenCount id="9DE3FD72FFF6FF9EDB2FB0E1897FFB37" box="[781,877,1206,1232]" count="3" pageId="9" pageNumber="256" type="larva">3 larvae</specimenCount>
,
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,
<collectingMunicipality id="6B3EAC81FFF6FF9EDC09B0E18ECAFB37" box="[1067,1240,1206,1232]" pageId="9" pageNumber="256">Matheshwaram</collectingMunicipality>
kuttai,
<collectingCounty id="623B4E77FFF6FF9EDD17B0E18F8AFB37" box="[1333,1432,1206,1232]" pageId="9" pageNumber="256">Kottadai</collectingCounty>
,
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,
<geoCoordinate id="EED1503CFFF6FF9ED90FB08D8BA6FB13" box="[301,436,1242,1268]" degrees="77.13364" direction="east" orientation="longitude" pageId="9" pageNumber="256" precision="1" value="77.13364">77.13364°E</geoCoordinate>
,
<quantity id="4C1D9B1EFFF6FF9ED9E2B08D8818FB13" box="[448,522,1242,1268]" metricMagnitude="2" metricUnit="m" metricValue="9.48" pageId="9" pageNumber="256" unit="m" value="948.0">
<specimenCount id="9DE3FD72FFF6FF9ED9E2B08D8818FB13" box="[448,522,1242,1268]" count="948" pageId="9" pageNumber="256" type="generic">
<elevation id="00C8D1C8FFF6FF9ED9E2B08D8818FB13" box="[448,522,1242,1268]" metricMagnitude="2" metricUnit="m" metricValue="9.48" pageId="9" pageNumber="256" unit="m" value="948.0">948 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="3B8D3CA6FFF6FF9EDA28B08D8E75FB13" box="[522,1127,1242,1268]" collectingDate="2023-02-04" collectionCode="R" collectorName="Babu &amp; Party &amp; Reg. No. I" country="India" pageId="9" pageNumber="256" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="paratype">
.
<date id="FF5B103BFFF6FF9EDA37B08D8899FB13" box="[533,651,1242,1268]" pageId="9" pageNumber="256" value="2023-02-04">
<collectingDate id="EF1FE9D3FFF6FF9EDA37B08D8899FB13" box="[533,651,1242,1268]" pageId="9" pageNumber="256" value="2023-02-04">04.II.2023</collectingDate>
</date>
, coll.
<collectionCode id="EDF4AE3EFFF6FF9EDAEDB08D88F7FB13" box="[719,741,1242,1268]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="256">R</collectionCode>
.
<collectorName id="2610532DFFF6FF9EDAD2B08D8939FB13" box="[752,811,1242,1268]" pageId="9" pageNumber="256">Babu</collectorName>
&amp;
<collectorName id="2610532DFFF6FF9EDB6DB08D8998FB13" box="[847,906,1242,1268]" pageId="9" pageNumber="256">Party</collectorName>
(
<collectorName id="2610532DFFF6FF9EDBBBB08D8E1DFB13" box="[921,1039,1242,1268]" pageId="9" pageNumber="256">Reg. No. I</collectorName>
/E/868)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF6FF9AD8E5B0A98B82F9C8" lastPageId="13" lastPageNumber="260" pageId="9" pageNumber="256" type="description">
<paragraph id="8B5A36FBFFF6FF9ED8E5B0A98887FADB" blockId="9.[151,1437,1134,2024]" pageId="9" pageNumber="256">
<emphasis id="B991EAE9FFF6FF9ED8E5B0A98816FAFF" bold="true" box="[199,516,1278,1304]" pageId="9" pageNumber="256">Description. Mature larva</emphasis>
(
<figureCitation id="13DE2A7EFFF6FF9EDA34B0A9887EFAFE" box="[534,620,1278,1305]" captionStart="FIGURES 5152" captionStartId="10.[151,264,1632,1656]" captionTargetBox="[164,1423,273,1609]" captionTargetId="figure-68@10.[164,1423,273,1609]" captionTargetPageId="10" captionText="FIGURES 5152. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." pageId="9" pageNumber="256">Fig. 51</figureCitation>
). Length (mm): body 1111.5; cerci 1313.5.
<specimenCount id="9DE3FD72FFF6FF9EDCA6B0A98E8BFAFF" box="[1156,1177,1278,1304]" count="1" pageId="9" pageNumber="256" type="female"></specimenCount>
body 12.513.5; cerci 14.515.5; prothorax width 4.24.5 (
<figureCitation id="13DE2A7EFFF6FF9EDA16B175889AFADB" box="[564,648,1314,1340]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="9" pageNumber="256">Fig. 78</figureCitation>
).
</paragraph>
<paragraph id="8B5A36FBFFF6FF9ED8E5B1118AFFFA2B" blockId="9.[151,1437,1134,2024]" pageId="9" pageNumber="256">
<emphasis id="B991EAE9FFF6FF9ED8E5B1118B32FA87" bold="true" box="[199,288,1350,1376]" pageId="9" pageNumber="256">Colour.</emphasis>
Generally pale brownish. Antennae brown and much shorter than the width of head (
<figureCitation id="13DE2A7EFFF6FF9EDCD5B1118F5EFA86" box="[1271,1356,1350,1377]" captionStart="FIGURES 8895" captionStartId="16.[151,264,1914,1938]" captionTargetBox="[151,1435,181,1890]" captionTargetId="figure-30@16.[151,1435,181,1890]" captionTargetPageId="16" captionText="FIGURES 8895. Larva of Thalerosphyrus sartorii sp. nov. 88. Gill I; 89. Gill II; 90. Gill III; 91. Gill IV; 92. Gill V; 93. Gill VI; 94. Gill VII; 95. Antenna." pageId="9" pageNumber="256">Fig. 95</figureCitation>
). Head circular, brown, and flattened; anterior margin of the head with numerous small setae; a pair of pale maculae on the front of head, and dorsal surface has four pale dots; ocelli with three black dots mixed with pale color in the middle.
</paragraph>
<paragraph id="8B5A36FBFFF6FF9DD8E5B1818AFCFF1E" blockId="9.[151,1437,1134,2024]" lastBlockId="10.[151,1436,150,249]" lastPageId="10" lastPageNumber="257" pageId="9" pageNumber="256">
<emphasis id="B991EAE9FFF6FF9ED8E5B1818BBFFA17" bold="true" box="[199,429,1494,1520]" pageId="9" pageNumber="256">Head. Mouthparts:</emphasis>
Labrum moderately broad, 3.5× wider than long, anterior margin straight, medio-ventrally with a curved row composed of 78 long stout setae (
<figureCitation id="13DE2A7EFFF6FF9EDACCB1AD8963F9F2" box="[750,881,1530,1557]" captionStart="FIGURES 5562" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" pageId="9" pageNumber="256">Figs 5758</figureCitation>
); dorsal surface and anterior margin covered with long, hair-like and stout setae; ventral surface of anterior margin with numerous long hair-like setae lateral margin slightly concave. Mandibles strongly bent in distal part; anterolateral margins with several rows of dense long hair-like setae; anterolateral margins medially bulged (
<figureCitation id="13DE2A7EFFF6FF9EDAEAB2318959F967" box="[712,843,1638,1664]" captionStart-0="FIGURES 5562" captionStart-1="FIGURES 6367" captionStartId-0="12.[151,264,1920,1944]" captionStartId-1="13.[151,264,1415,1439]" captionTargetBox-0="[151,1435,181,1897]" captionTargetBox-1="[164,1423,181,1391]" captionTargetId-0="figure-30@12.[151,1435,181,1897]" captionTargetId-1="figure-19@13.[164,1423,181,1391]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" captionText-1="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="9" pageNumber="256">Figs 6063</figureCitation>
). Right mandible with outer incisor serrated on inner margin, composed of 8 blunt teeth; inner incisor with two teeth, apically sharp; prostheca with 1011 thick fimbriate submarginal setae; mola joined, posterior margin with 56 hair-like setae (
<figureCitation id="13DE2A7EFFF6FF9EDBC5B2F98E7EF92F" box="[999,1132,1710,1736]" captionStart="FIGURES 5562" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" pageId="9" pageNumber="256">Figs 6061</figureCitation>
). Left mandible with outer incisor serrated with 13 blunt teeth; inner incisor apically bulged with trifid teeth, anterior tooth with small stout setae at the tip; prostheca with 11 thick fimbriate setae; mola with 67 setae and posterior median margin with 5 hair-like setae (
<figureCitation id="13DE2A7EFFF6FF9ED965B34D8BDFF8D3" box="[327,461,1818,1844]" captionStart-0="FIGURES 5562" captionStart-1="FIGURES 6367" captionStartId-0="12.[151,264,1920,1944]" captionStartId-1="13.[151,264,1415,1439]" captionTargetBox-0="[151,1435,181,1897]" captionTargetBox-1="[164,1423,181,1391]" captionTargetId-0="figure-30@12.[151,1435,181,1897]" captionTargetId-1="figure-19@13.[164,1423,181,1391]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" captionText-1="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="9" pageNumber="256">Figs 6263</figureCitation>
). Maxilla with long, simple setae on inner margin of galea-lacinia and with fimbriate setae scattered on ventral surface. Segment I of maxillary palp with rows of thin and simple setae on outer margins, dorsal surface with 4 small simple setae; segment II large with rows of numerous hair-like setae on outer margin; apical segment small, pointed, with rows of simple setae on outer margin. Crown of the galea-lacinia of maxillae with 1415 comb-shaped setae (
<figureCitation id="13DE2A7EFFF6FF9ED9D9B3FD885DF823" box="[507,591,1962,1988]" captionStart="FIGURES 6367" captionStartId="13.[151,264,1415,1439]" captionTargetBox="[164,1423,181,1391]" captionTargetId="figure-19@13.[164,1423,181,1391]" captionTargetPageId="13" captionText="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="9" pageNumber="256">Fig. 64</figureCitation>
). Hypopharynx narrow, lingua with tuft of small robust setae (lingua slightly damaged), superlinguae slightly expanded laterally, densely covered with long and thin setae replaced apex with small setae up to lower part of the superlinguae (
<figureCitation id="13DE2A7EFFF5FF9DDA92B4C08911FF56" box="[688,771,150,177]" captionStart="FIGURES 5562" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" pageId="10" pageNumber="257">Fig. 59</figureCitation>
). Labium well developed, glossae dorsally with two distinct long setae and median patch of feathered setae; inner margin slightly concave, outer margin straight or serrated (
<figureCitation id="13DE2A7EFFF5FF9DDD48B4EC8AF3FF1E" captionStart="FIGURES 6367" captionStartId="13.[151,264,1415,1439]" captionTargetBox="[164,1423,181,1391]" captionTargetId="figure-19@13.[164,1423,181,1391]" captionTargetPageId="13" captionText="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="10" pageNumber="257">Figs 6567</figureCitation>
).
</paragraph>
<caption id="DF9A6673FFF5FF9DD8B5B2378E4DF99E" box="[151,1119,1632,1657]" pageId="10" pageNumber="257" startId="10.[151,264,1632,1656]" targetBox="[164,1423,273,1609]" targetPageId="10" targetType="figure">
<paragraph id="8B5A36FBFFF5FF9DD8B5B2378E4DF99E" blockId="10.[151,1119,1632,1657]" box="[151,1119,1632,1657]" pageId="10" pageNumber="257">
<emphasis id="B991EAE9FFF5FF9DD8B5B2378B46F99E" bold="true" box="[151,340,1632,1657]" pageId="10" pageNumber="257">FIGURES 5152.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFF5FF9DD995B23688B5F99F" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[439,679,1633,1657]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="257" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFF5FF9DD995B23688B5F99F" box="[439,679,1633,1657]" italics="true" pageId="10" pageNumber="257">Thalerosphyrus sartorii</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFF5FF9DDA8CB23688ECF99E" bold="true" box="[686,766,1633,1657]" pageId="10" pageNumber="257">
<taxonomicNameLabel id="A2A25792FFF5FF9DDA8CB23688ECF99E" box="[686,766,1633,1657]" pageId="10" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
51. Dorsal view; 52. Ventral view.
</paragraph>
</caption>
<paragraph id="8B5A36FBFFF5FF9CD8E5B29C88E7FF1E" blockId="10.[151,1437,1739,2017]" lastBlockId="11.[151,1436,151,249]" lastPageId="11" lastPageNumber="258" pageId="10" pageNumber="257">
<emphasis id="B991EAE9FFF5FF9DD8E5B29C8B35F902" bold="true" box="[199,295,1739,1765]" pageId="10" pageNumber="257">Thorax.</emphasis>
Pronotum moderately enlarged laterally, medially concave, wider than the head, and basally not fused to the mesonotum (
<figureCitation id="13DE2A7EFFF5FF9DD951B2B88BE5F8ED" box="[371,503,1775,1802]" captionStart="FIGURES 5152" captionStartId="10.[151,264,1632,1656]" captionTargetBox="[164,1423,273,1609]" captionTargetId="figure-68@10.[164,1423,273,1609]" captionTargetPageId="10" captionText="FIGURES 5152. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." pageId="10" pageNumber="257">Figs 5152</figureCitation>
,
<figureCitation id="13DE2A7EFFF5FF9DDA21B2B88830F8EE" box="[515,546,1775,1801]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="10" pageNumber="257">78</figureCitation>
), anterior margin of pronotum with a small projection at apex, anteromedially dark black indentation marking, and posterior part of medially with a pair of black spots (
<figureCitation id="13DE2A7EFFF5FF9DDC8BB3448F12F8CA" box="[1193,1280,1811,1837]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="10" pageNumber="257">Fig. 78</figureCitation>
). Mesonotum pale and brown color mixed, anterolaterally with deep concavity (
<figureCitation id="13DE2A7EFFF5FF9DDBA5B36089CFF8B6" box="[903,989,1847,1873]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="10" pageNumber="257">Fig. 78</figureCitation>
). Legs brown with pale markings (
<figureCitation id="13DE2A7EFFF5FF9DDD48B3608B1EF891" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="10" pageNumber="257">Figs 68, 71, 75</figureCitation>
). Foreleg: femur with a row of long strong setae on dorsal margin, anterior surface with numerous blunt stout setae, medially with small indentation of middle, and distally with a black spot (
<figureCitation id="13DE2A7EFFF5FF9DDC43B3288EF7F87E" box="[1121,1253,1919,1945]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="10" pageNumber="257">Figs 6870</figureCitation>
). Midleg: femur anterior surface with numerous stout setae, median and distally with black spot; tibia dorsal surface with small stout feathered setae, inner side with small thick setae, and outer margin with a row of hair-like setae (
<figureCitation id="13DE2A7EFFF5FF9DDD2FB3908F83F806" box="[1293,1425,1991,2017]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="10" pageNumber="257">Figs 7174</figureCitation>
). Hindleg: femur with dorsal surface triangle-shaped setae, clearly pointed median, and distally with black spot; tibia with dorsally two rows of stout setae, inner side without setae and marginal and submarginal 1718 pointed setae (
<figureCitation id="13DE2A7EFFF4FF9CD8BDB4888B31FF1E" box="[159,291,222,249]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="11" pageNumber="258">Figs 7577</figureCitation>
). Claw with 34 blunt denticles (
<figureCitation id="13DE2A7EFFF4FF9CDAB6B48888FBFF1E" box="[660,745,223,249]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="11" pageNumber="258">Fig. 77</figureCitation>
).
</paragraph>
<caption id="DF9A6673FFF4FF9CD8B5B13E8919FA42" pageId="11" pageNumber="258" startId="11.[151,264,1385,1409]" targetBox="[164,1423,273,1362]" targetPageId="11" targetType="figure">
<paragraph id="8B5A36FBFFF4FF9CD8B5B13E8919FA42" blockId="11.[151,1436,1385,1445]" pageId="11" pageNumber="258">
<emphasis id="B991EAE9FFF4FF9CD8B5B13E8B46FA66" bold="true" box="[151,340,1385,1409]" pageId="11" pageNumber="258">FIGURES 5354.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFF4FF9CD996B13E88B6FA66" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[436,676,1385,1409]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="258" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFF4FF9CD996B13E88B6FA66" box="[436,676,1385,1409]" italics="true" pageId="11" pageNumber="258">Thalerosphyrus sartorii</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFF4FF9CDA88B13E891FFA66" bold="true" box="[682,781,1385,1409]" pageId="11" pageNumber="258">
<taxonomicNameLabel id="A2A25792FFF4FF9CDA88B13E88EBFA66" box="[682,761,1385,1409]" pageId="11" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
5
</emphasis>
3. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male).
</paragraph>
</caption>
<paragraph id="8B5A36FBFFF4FF9CD8E5B1A08997F89D" blockId="11.[151,1437,1527,2022]" pageId="11" pageNumber="258">
<emphasis id="B991EAE9FFF4FF9CD8E5B1A08B51F9F6" bold="true" box="[199,323,1527,1553]" pageId="11" pageNumber="258">Abdomen.</emphasis>
Posterolateral expansions on terga IIIVIII parallel to abdomen, poorly developed on tergum III, weakly developed on terga IVV, moderately developed on terga VIVIII (
<figureCitation id="13DE2A7EFFF4FF9CDBD8B24C8E92F9D1" box="[1018,1152,1563,1590]" captionStart="FIGURES 5354" captionStartId="11.[151,264,1385,1409]" captionTargetBox="[164,1423,273,1362]" captionTargetId="figure-62@11.[164,1423,273,1362]" captionTargetPageId="11" captionText="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." pageId="11" pageNumber="258">Figs 5354</figureCitation>
). Dorsal surface of terga IIII pale brown, medially a brown spot; tergum III with laterally black spot; terga IVV medially with a distinct pale-yellow spot (
<figureCitation id="13DE2A7EFFF4FF9CD947B2348BA9F999" box="[357,443,1635,1662]" captionStart="FIGURES 5152" captionStartId="10.[151,264,1632,1656]" captionTargetBox="[164,1423,273,1609]" captionTargetId="figure-68@10.[164,1423,273,1609]" captionTargetPageId="10" captionText="FIGURES 5152. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." pageId="11" pageNumber="258">Fig. 51</figureCitation>
); terga IVX brown mixed with pale yellow (
<figureCitation id="13DE2A7EFFF4FF9CDBECB2348E31F999" box="[974,1059,1635,1662]" captionStart="FIGURES 5354" captionStartId="11.[151,264,1385,1409]" captionTargetBox="[164,1423,273,1362]" captionTargetId="figure-62@11.[164,1423,273,1362]" captionTargetPageId="11" captionText="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." pageId="11" pageNumber="258">Fig. 53</figureCitation>
); sternum pale yellowish or pale brownish (mature larva), sterna IIIVII laterally with brown spot (
<figureCitation id="13DE2A7EFFF4FF9CDBBDB2D08E09F945" box="[927,1051,1671,1698]" captionStart="FIGURES 5354" captionStartId="11.[151,264,1385,1409]" captionTargetBox="[164,1423,273,1362]" captionTargetId="figure-62@11.[164,1423,273,1362]" captionTargetPageId="11" captionText="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." pageId="11" pageNumber="258">Fig 5354</figureCitation>
). Posterior margins of terga with irregular pointed teeth and a row of 23 microdenticles (
<figureCitation id="13DE2A7EFFF4FF9CDB3CB2FC89B2F922" box="[798,928,1707,1733]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="11" pageNumber="258">Figs 8486</figureCitation>
) and each tergite laterally with small simple setae (
<figureCitation id="13DE2A7EFFF4FF9CD8FFB2988B3DF90D" box="[221,303,1743,1770]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="11" pageNumber="258">Fig. 85</figureCitation>
). Gill I convex, asymmetrical, elongated, apically moderately sharp, 2.5× longer than wide and ventral lamella long, fimbriate (
<figureCitation id="13DE2A7EFFF4FF9CD98BB2A48812F8EA" box="[425,512,1779,1805]" captionStart="FIGURES 8895" captionStartId="16.[151,264,1914,1938]" captionTargetBox="[151,1435,181,1890]" captionTargetId="figure-30@16.[151,1435,181,1890]" captionTargetPageId="16" captionText="FIGURES 8895. Larva of Thalerosphyrus sartorii sp. nov. 88. Gill I; 89. Gill II; 90. Gill III; 91. Gill IV; 92. Gill V; 93. Gill VI; 94. Gill VII; 95. Antenna." pageId="11" pageNumber="258">Figs 88</figureCitation>
,
<figureCitation id="13DE2A7EFFF4FF9CDA2FB2A48828F8EA" box="[525,570,1779,1805]" captionStart="FIGURES 111114" captionStartId="19.[151,264,806,830]" captionTargetBox="[313,1274,183,780]" captionTargetId="figure-19@19.[311,1276,181,782]" captionTargetPageId="19" captionText="FIGURES 111114. Gill I of dorsal plate: 111. Th. flowersi (Paratype); 112. Th. flowersi (Thailand); 113. Th. bengalensis sp. nov.; 114. Th. sartorii sp. nov." pageId="11" pageNumber="258">114</figureCitation>
); Gill IIVI strongly asymmetrical, length and width subequal (
<figureCitation id="13DE2A7EFFF4FF9CDD2EB2A48F82F8EA" box="[1292,1424,1779,1805]" captionStart="FIGURES 8895" captionStartId="16.[151,264,1914,1938]" captionTargetBox="[151,1435,181,1890]" captionTargetId="figure-30@16.[151,1435,181,1890]" captionTargetPageId="16" captionText="FIGURES 8895. Larva of Thalerosphyrus sartorii sp. nov. 88. Gill I; 89. Gill II; 90. Gill III; 91. Gill IV; 92. Gill V; 93. Gill VI; 94. Gill VII; 95. Antenna." pageId="11" pageNumber="258">Figs 8993</figureCitation>
). Gill VII long, 2.5× longer than wide (
<figureCitation id="13DE2A7EFFF4FF9CDA65B340888FF8D6" box="[583,669,1815,1841]" captionStart="FIGURES 8895" captionStartId="16.[151,264,1914,1938]" captionTargetBox="[151,1435,181,1890]" captionTargetId="figure-30@16.[151,1435,181,1890]" captionTargetPageId="16" captionText="FIGURES 8895. Larva of Thalerosphyrus sartorii sp. nov. 88. Gill I; 89. Gill II; 90. Gill III; 91. Gill IV; 92. Gill V; 93. Gill VI; 94. Gill VII; 95. Antenna." pageId="11" pageNumber="258">Fig. 94</figureCitation>
). Cerci pale brownish with medium brown bands increasing in size towards apex. Paracercus pale whitish with yellow, with a dark brown band on 34 segments and each segment surrounded with pointed small yellowish spine-like setae (
<figureCitation id="13DE2A7EFFF4FF9CDB06B308896BF89D" box="[804,889,1887,1914]" captionStart="FIGURES 5562" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" pageId="11" pageNumber="258">Fig. 55</figureCitation>
).
</paragraph>
<paragraph id="8B5A36FBFFF4FF9CD8E5B3D488B6F802" blockId="11.[151,1437,1527,2022]" pageId="11" pageNumber="258">
<emphasis id="B991EAE9FFF4FF9CD8E5B3D48B20F87A" bold="true" box="[199,306,1923,1949]" pageId="11" pageNumber="258">Genitals.</emphasis>
Protopenis (
<figureCitation id="13DE2A7EFFF4FF9CD9E6B3D48859F87A" box="[452,587,1923,1949]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="11" pageNumber="258">Figs 8082</figureCitation>
): male genitalia yellowish, penis fused, V-shaped medially, apically blunt, and laterally with tapered square, tergum X triangular and apically with numerous stout setae (
<figureCitation id="13DE2A7EFFF4FF9CDCFAB3F08F3FF826" box="[1240,1325,1959,1985]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="11" pageNumber="258">Fig. 81</figureCitation>
). Forceps proximally broad and towards narrow apically.
</paragraph>
<caption id="DF9A6673FFF3FF9BD8B5B3D7898BF807" pageId="12" pageNumber="259" startId="12.[151,264,1920,1944]" targetBox="[151,1435,181,1897]" targetPageId="12" targetType="figure">
<paragraph id="8B5A36FBFFF3FF9BD8B5B3D7898BF807" blockId="12.[151,1437,1920,2017]" pageId="12" pageNumber="259">
<emphasis id="B991EAE9FFF3FF9BD8B5B3D78B46F87F" bold="true" box="[151,340,1920,1944]" pageId="12" pageNumber="259">FIGURES 5562.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFF3FF9BD995B3D788B5F87F" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[439,679,1920,1944]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="259" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFF3FF9BD995B3D788B5F87F" box="[439,679,1920,1944]" italics="true" pageId="12" pageNumber="259">Thalerosphyrus sartorii</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFF3FF9BDA8FB3D688ECF87E" bold="true" box="[685,766,1921,1945]" pageId="12" pageNumber="259">
<taxonomicNameLabel id="A2A25792FFF3FF9BDA8FB3D688ECF87E" box="[685,766,1921,1945]" pageId="12" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;
</paragraph>
</caption>
<caption id="DF9A6673FFF2FF9AD8B5B1D08EC1FA24" pageId="13" pageNumber="260" startId="13.[151,264,1415,1439]" targetBox="[164,1423,181,1391]" targetPageId="13" targetType="figure">
<paragraph id="8B5A36FBFFF2FF9AD8B5B1D08EC1FA24" blockId="13.[151,1437,1415,1475]" pageId="13" pageNumber="260">
<emphasis id="B991EAE9FFF2FF9AD8B5B1D08B47FA78" bold="true" box="[151,341,1415,1439]" pageId="13" pageNumber="260">FIGURES 6367.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFF2FF9AD99AB1D088BBFA78" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[440,681,1415,1439]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="260" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFF2FF9AD99AB1D088BBFA78" box="[440,681,1415,1439]" italics="true" pageId="13" pageNumber="260">Thalerosphyrus sartorii</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFF2FF9ADA92B1D08913FA78" bold="true" box="[688,769,1415,1439]" pageId="13" pageNumber="260">
<taxonomicNameLabel id="A2A25792FFF2FF9ADA92B1D08913FA78" box="[688,769,1415,1439]" pageId="13" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view.
</paragraph>
</caption>
<paragraph id="8B5A36FBFFF2FF9AD8E5B2428B82F9C8" blockId="13.[151,1437,1557,2016]" box="[199,400,1557,1583]" pageId="13" pageNumber="260">
<emphasis id="B991EAE9FFF2FF9AD8E5B2428B01F9C8" bold="true" box="[199,275,1557,1583]" pageId="13" pageNumber="260">Adult.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF2FF9AD8E5B26D8BCEF990" pageId="13" pageNumber="260" type="etymology">
<paragraph id="8B5A36FBFFF2FF9AD8E5B26D8BCEF990" blockId="13.[151,1437,1557,2016]" pageId="13" pageNumber="260">
<emphasis id="B991EAE9FFF2FF9AD8E5B26D8B5CF9B3" bold="true" box="[199,334,1594,1620]" pageId="13" pageNumber="260">Etymology.</emphasis>
This species is named in honour of Dr. Michel Sartori in recognition of his significant contributions to the genus
<emphasis id="B991EAE9FFF2FF9AD907B20A8BCEF990" box="[293,476,1629,1655]" italics="true" pageId="13" pageNumber="260">
<taxonomicName id="4CE54D78FFF2FF9AD907B20A8BC5F990" authorityName="Eaton" authorityYear="1881" box="[293,471,1629,1655]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="260" phylum="Arthropoda" rank="genus">Thalerosphyrus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF2FF9AD8E5B2D68970F97B" box="[199,866,1665,1692]" pageId="13" pageNumber="260" type="distribution">
<paragraph id="8B5A36FBFFF2FF9AD8E5B2D68970F97B" blockId="13.[151,1437,1557,2016]" box="[199,866,1665,1692]" pageId="13" pageNumber="260">
<emphasis id="B991EAE9FFF2FF9AD8E5B2D68B73F97C" bold="true" box="[199,353,1665,1691]" pageId="13" pageNumber="260">Distribution.</emphasis>
Erode district,
<collectingRegion id="4921F819FFF2FF9ADA2FB2D68883F97C" box="[525,657,1665,1691]" country="India" name="Tamil Nadu" pageId="13" pageNumber="260">Tamil Nadu</collectingRegion>
(
<collectingCountry id="F3F2766BFFF2FF9ADA82B2D688CFF97C" box="[672,733,1665,1691]" name="India" pageId="13" pageNumber="260">India</collectingCountry>
) (
<figureCitation id="13DE2A7EFFF2FF9ADAD6B2D68947F97B" box="[756,853,1665,1692]" captionStart="FIGURES 111114" captionStartId="19.[151,264,806,830]" captionTargetBox="[313,1274,183,780]" captionTargetId="figure-19@19.[311,1276,181,782]" captionTargetPageId="19" captionText="FIGURES 111114. Gill I of dorsal plate: 111. Th. flowersi (Paratype); 112. Th. flowersi (Thailand); 113. Th. bengalensis sp. nov.; 114. Th. sartorii sp. nov." pageId="13" pageNumber="260">Fig. 112</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF2FF85D8E5B2F2889CFE82" lastPageId="18" lastPageNumber="265" pageId="13" pageNumber="260" type="diagnosis">
<paragraph id="8B5A36FBFFF2FF85D8E5B2F2889CFE82" blockId="13.[151,1437,1557,2016]" lastBlockId="18.[151,1437,151,357]" lastPageId="18" lastPageNumber="265" pageId="13" pageNumber="260">
<emphasis id="B991EAE9FFF2FF9AD8E5B2F28B50F958" bold="true" box="[199,322,1701,1727]" pageId="13" pageNumber="260">Diagnosis.</emphasis>
<taxonomicName id="4CE54D78FFF2FF9AD96AB2F289FDF958" authority="Vasanth, Kubendran &amp; Subramanian" authorityName="Vasanth, Kubendran &amp; Subramanian" authorityYear="2025" box="[328,1007,1701,1728]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="260" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B991EAE9FFF2FF9AD96AB2F28843F958" box="[328,593,1701,1727]" italics="true" pageId="13" pageNumber="260">Thalerosphyrus sartorii</emphasis>
Vasanth, Kubendran &amp; Subramanian
</taxonomicName>
<emphasis id="B991EAE9FFF2FF9ADBD7B2F18E5CF927" bold="true" box="[1013,1102,1702,1728]" pageId="13" pageNumber="260">
<taxonomicNameLabel id="A2A25792FFF2FF9ADBD7B2F18E5CF927" box="[1013,1102,1702,1728]" pageId="13" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4CE54D78FFF2FF9ADCFCB2F18F40F958" authorityName="Venkataraman &amp; Sivamarakrishnan" authorityYear="1987" box="[1246,1362,1701,1727]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="260" phylum="Arthropoda" rank="species" species="flowersi">
<emphasis id="B991EAE9FFF2FF9ADCFCB2F18F40F958" box="[1246,1362,1701,1727]" italics="true" pageId="13" pageNumber="260">T. flowersi</emphasis>
</taxonomicName>
and
<emphasis id="B991EAE9FFF2FF9ADDAAB2F18B51F904" italics="true" pageId="13" pageNumber="260">
<taxonomicName id="4CE54D78FFF2FF9ADDAAB2F18B2DF904" authorityName="Selvakumar &amp; Chandra" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="260" phylum="Arthropoda" rank="species" species="meghalayensis">T. meghalayensis</taxonomicName>
,
</emphasis>
but it can be distinguished from all known species of
<taxonomicName id="4CE54D78FFF2FF9ADBBEB29E8E59F904" authorityName="Eaton" authorityYear="1881" box="[924,1099,1737,1763]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B991EAE9FFF2FF9ADBBEB29E8E59F904" box="[924,1099,1737,1763]" italics="true" pageId="13" pageNumber="260">Thalerosphyrus</emphasis>
</taxonomicName>
by the following combination of characters: (i) posterolateral expansions on terga IIIVIII, terga VIVIII moderately developed, tergum III poorly developed, weakly developed on terga IVV (
<figureCitation id="13DE2A7EFFF2FF9ADA80B3468934F8CB" box="[674,806,1809,1836]" captionStart-0="FIGURES 5152" captionStart-1="FIGURES 5354" captionStartId-0="10.[151,264,1632,1656]" captionStartId-1="11.[151,264,1385,1409]" captionTargetBox-0="[164,1423,273,1609]" captionTargetBox-1="[164,1423,273,1362]" captionTargetId-0="figure-68@10.[164,1423,273,1609]" captionTargetId-1="figure-62@11.[164,1423,273,1362]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 5152. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." captionText-1="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." pageId="13" pageNumber="260">Figs 5154</figureCitation>
); (ii) gill I convex, asymmetrical, elongate, 2.5× longer than wide (
<figureCitation id="13DE2A7EFFF2FF9AD934B3628B78F8B7" box="[278,362,1845,1872]" captionStart="FIGURES 8895" captionStartId="16.[151,264,1914,1938]" captionTargetBox="[151,1435,181,1890]" captionTargetId="figure-30@16.[151,1435,181,1890]" captionTargetPageId="16" captionText="FIGURES 8895. Larva of Thalerosphyrus sartorii sp. nov. 88. Gill I; 89. Gill II; 90. Gill III; 91. Gill IV; 92. Gill V; 93. Gill VI; 94. Gill VII; 95. Antenna." pageId="13" pageNumber="260">Fig. 88</figureCitation>
); (iii) abdominal tergum III dorsolaterally with black markings (
<figureCitation id="13DE2A7EFFF2FF9ADC63B3628E87F8B7" box="[1089,1173,1845,1872]" captionStart="FIGURES 5354" captionStartId="11.[151,264,1385,1409]" captionTargetBox="[164,1423,273,1362]" captionTargetId="figure-62@11.[164,1423,273,1362]" captionTargetPageId="11" captionText="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." pageId="13" pageNumber="260">Fig. 53</figureCitation>
); (iv) abdominal sterna IIIVII with laterally black streaks (
<figureCitation id="13DE2A7EFFF2FF9ADA0BB30E8865F893" box="[553,631,1881,1908]" captionStart="FIGURES 5354" captionStartId="11.[151,264,1385,1409]" captionTargetBox="[164,1423,273,1362]" captionTargetId="figure-62@11.[164,1423,273,1362]" captionTargetPageId="11" captionText="FIGURES 5354. Larva of Thalerosphyrus sartorii sp. nov. 53. Abdomen dorsal view (arrows indicate tergum III black spots); 54. Posterolateral spine (left-side female and right-side male)." pageId="13" pageNumber="260">Fig 54</figureCitation>
); (v) posterior margins of terga with irregular pointed teeth and a row of 23 micro denticles (
<figureCitation id="13DE2A7EFFF2FF9AD9A5B32A881EF870" box="[391,524,1917,1943]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="13" pageNumber="260">Figs 8486</figureCitation>
); (vi) pronotum moderately enlarged laterally, with anteromedian black marking and submedian pair of black spots (
<figureCitation id="13DE2A7EFFF2FF9ADA1BB3F68886F85B" box="[569,660,1953,1980]" captionStart="FIGURES 5152" captionStartId="10.[151,264,1632,1656]" captionTargetBox="[164,1423,273,1609]" captionTargetId="figure-68@10.[164,1423,273,1609]" captionTargetPageId="10" captionText="FIGURES 5152. Larva of Thalerosphyrus sartorii sp. nov. 51. Dorsal view; 52. Ventral view." pageId="13" pageNumber="260">Figs 51</figureCitation>
,
<figureCitation id="13DE2A7EFFF2FF9ADA81B3F588D1F85B" box="[675,707,1954,1980]" captionStart="FIGURES 7887" captionStartId="15.[151,264,1856,1880]" captionTargetBox="[151,1435,185,1820]" captionTargetId="figure-19@15.[151,1435,181,1833]" captionTargetPageId="15" captionText="FIGURES 7887. Larva of Thalerosphyrus sartorii sp. nov. 78. Notum (arrows indicate pro-and metanotum); 79. Pronotum with a small projection; 80. Protopenis (arrow indicates penis); 81. Protopenis closer view; 82. Penis closer view; 8384. Posterior margin of tergum VI; 85. Dorsal surface of tergum VI with small simple setae; 86. Posterior margin of tergum VIII with posteromarginal spines; 87. Tergum VIII posterolateral spine closer view." pageId="13" pageNumber="260">78</figureCitation>
); (vii) all femora medially with black maculae and blunt stout setae (
<figureCitation id="13DE2A7EFFF2FF9AD8C0B3928B8BF807" box="[226,409,1989,2016]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="13" pageNumber="260">Figs 68, 71, 75</figureCitation>
); (viii) hind femur triangular-shaped setae, clearly pointed (
<figureCitation id="13DE2A7EFFF2FF9ADC71B3928EB9F807" box="[1107,1195,1989,2016]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="13" pageNumber="260">Fig. 75</figureCitation>
); (xi) claw with 34 blunt denticles (
<figureCitation id="13DE2A7EFFEDFF85D96BB4C08B8FFF56" box="[329,413,151,177]" captionStart="FIGURES 6877" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[184,1407,181,1919]" captionTargetId="figure-30@14.[179,1407,181,1919]" captionTargetPageId="14" captionText="FIGURES 6877. Larva of Thalerosphyrus sartorii sp. nov. 68. Forefemur; 69. Forefemur black macula; 70. Closer view of stout setae; 71. Midleg; 72. Stout setae closer view; 73. Midtibia setae; 74. Tibia setae closer view; 75. Hindleg; 76. Hindtibia (arrow indicates long stout setae); 77. Claw (arrow indicates denticles)." pageId="18" pageNumber="265">Fig. 77</figureCitation>
); (x) labrum moderately expanded laterally, anterior margin straight, medio-ventrally with a curved row composed of 78 long stout setae each; lateral margin slightly concave (
<figureCitation id="13DE2A7EFFEDFF85DC61B4EC8ED4FF32" box="[1091,1222,186,213]" captionStart="FIGURES 5562" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" pageId="18" pageNumber="265">Figs 5758</figureCitation>
); (xi) hypopharynx narrow and superlinguae slightly expanded laterally with apically with small setae (
<figureCitation id="13DE2A7EFFEDFF85DC6EB4888EB0FF1E" box="[1100,1186,222,249]" captionStart="FIGURES 5562" captionStartId="12.[151,264,1920,1944]" captionTargetBox="[151,1435,181,1897]" captionTargetId="figure-30@12.[151,1435,181,1897]" captionTargetPageId="12" captionText="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" pageId="18" pageNumber="265">Fig. 59</figureCitation>
); (xii) mandibles with lateral margin medially bulged, of mandibles and left mandible inner incisor with three teeth with small stout setae joined with first teeth (
<figureCitation id="13DE2A7EFFEDFF85D9BFB5708831FEA6" box="[413,547,295,321]" captionStart-0="FIGURES 5562" captionStart-1="FIGURES 6367" captionStartId-0="12.[151,264,1920,1944]" captionStartId-1="13.[151,264,1415,1439]" captionTargetBox-0="[151,1435,181,1897]" captionTargetBox-1="[164,1423,181,1391]" captionTargetId-0="figure-30@12.[151,1435,181,1897]" captionTargetId-1="figure-19@13.[164,1423,181,1391]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 5562. Larva of Thalerosphyrus sartorii sp. nov. 55. Caudal filaments; 56. Dorsal view of protopenis; 57. Labrum; 58. Labrum closer view (arrow indicates dorsal stout setae); 59. Hypopharynx; 60. Right mandible; 61. Right mandible closer view (arrow indicates incisor teeth and associated setae). 62. Left mandible;" captionText-1="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="18" pageNumber="265">Figs 6063</figureCitation>
); (xiii) maxillae broad, galea-lacinia with 1415 comb-shaped setae (
<figureCitation id="13DE2A7EFFEDFF85DD1BB5708F9DFEA6" box="[1337,1423,295,321]" captionStart="FIGURES 6367" captionStartId="13.[151,264,1415,1439]" captionTargetBox="[164,1423,181,1391]" captionTargetId="figure-19@13.[164,1423,181,1391]" captionTargetPageId="13" captionText="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="18" pageNumber="265">Fig. 64</figureCitation>
); (xiv) glossae with slightly concave (
<figureCitation id="13DE2A7EFFEDFF85DA0FB51C8893FE82" box="[557,641,331,357]" captionStart="FIGURES 6367" captionStartId="13.[151,264,1415,1439]" captionTargetBox="[164,1423,181,1391]" captionTargetId="figure-19@13.[164,1423,181,1391]" captionTargetPageId="13" captionText="FIGURES 6367. Larva of Thalerosphyrus sartorii sp. nov. 63. Left mandible closer view (arrow indicates incisor teeth); 64. Maxilla; 65. Labium; 66. Glossae (arrows indicate dorsal setae closer view); 67. Glossae setae closer view." pageId="18" pageNumber="265">Fig. 66</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,429 @@
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<mods:title id="BDC684C694F2608360B4B4B6226C0087">Two new species of the genus Thalerosphyrus Eaton, 1881 (Ephemeroptera: Heptageniidae: Ecdyonurinae) from India</mods:title>
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<taxonomicName id="4CE54D78FFFEFF96D8B5B5FC89DBFE21" authority="Vasanth, Kubendran &amp; Subramanian" authorityName="Vasanth, Kubendran &amp; Subramanian" authorityYear="2025" box="[151,969,427,454]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFFEFF96D8B5B5FC8BF9FE21" bold="true" box="[151,491,427,454]" italics="true" pageId="1" pageNumber="248">Thalerosphyrus bengalensis</emphasis>
Vasanth, Kubendran &amp; Subramanian
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<paragraph id="8B5A36FBFFFEFF96D8B5B5878B0BFE0C" blockId="1.[151,1070,427,491]" box="[151,281,464,491]" pageId="1" pageNumber="248">
(
<figureCitation id="13DE2A7EFFFEFF96D8BDB5878B03FE0C" box="[159,273,464,491]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 45" captionStart-2="FIGURES 611" captionStart-3="FIGURES 1219" captionStart-4="FIGURES 2024" captionStart-5="FIGURES 2530" captionStart-6="FIGURES 3137" captionStart-7="FIGURES 3843" captionStart-8="FIGURES 4450" captionStartId-0="1.[151,264,1999,2023]" captionStartId-1="2.[151,264,1438,1462]" captionStartId-2="3.[151,264,1930,1954]" captionStartId-3="4.[151,264,1909,1933]" captionStartId-4="5.[151,264,1421,1445]" captionStartId-5="6.[151,264,1357,1381]" captionStartId-6="7.[151,264,1466,1490]" captionStartId-7="8.[151,264,1937,1961]" captionStartId-8="9.[151,264,883,907]" captionTargetBox-0="[214,1374,656,1973]" captionTargetBox-1="[225,1363,181,1414]" captionTargetBox-2="[213,1374,489,1907]" captionTargetBox-3="[164,1423,181,1884]" captionTargetBox-4="[189,1398,181,1397]" captionTargetBox-5="[201,1386,181,1333]" captionTargetBox-6="[236,1386,181,1443]" captionTargetBox-7="[218,1386,561,1914]" captionTargetBox-8="[177,1411,181,860]" captionTargetId-0="figure-130@1.[214,1374,656,1973]" captionTargetId-1="figure-30@2.[225,1363,181,1414]" captionTargetId-2="figure-183@3.[213,1374,489,1907]" captionTargetId-3="figure-30@4.[164,1423,181,1884]" captionTargetId-4="figure-19@5.[189,1398,181,1397]" captionTargetId-5="figure-30@6.[201,1386,181,1333]" captionTargetId-6="figure-19@7.[201,1386,181,1443]" captionTargetId-7="figure-206@8.[201,1386,561,1914]" captionTargetId-8="figure-19@9.[177,1411,181,860]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="7" captionTargetPageId-7="8" captionTargetPageId-8="9" captionText-0="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." captionText-1="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." captionText-2="FIGURES 611. Larva of Thalerosphyrus bengalensis sp. nov. 6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament." captionText-3="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." captionText-4="FIGURES 2024. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." captionText-5="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." captionText-6="FIGURES 3137. Larva of Thalerosphyrus bengalensis sp. nov. 31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae." captionText-7="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." captionText-8="FIGURES 4450. Larva of Thalerosphyrus bengalensis sp. nov. 44. Gill I; 45. Gill II; 46. Gill III; 47. Gill IV; 48. Gill V; 49. Gill VI; 50. Gill VII." pageId="1" pageNumber="248">Figs 150</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFFEFF96D8B5B64F8BABFD9C" pageId="1" pageNumber="248" type="materials_examined">
<paragraph id="8B5A36FBFFFEFF96D8B5B64F8BABFD9C" blockId="1.[151,1437,536,635]" pageId="1" pageNumber="248">
<emphasis id="B991EAE9FFFEFF96D8B5B64F8B65FDD5" bold="true" box="[151,375,536,562]" pageId="1" pageNumber="248">Material examined.</emphasis>
<materialsCitation id="3B8D3CA6FFFEFF96D95DB64F898BFDB1" collectingDate="2022-10-14" collectionCode="T" country="India" county="Mature" elevation="311" latitude="23.41131" location="Jalika village" longLatPrecision="1" longitude="86.19734" municipality="Purulia district" pageId="1" pageNumber="248" specimenCount="311" stateProvince="West Bengal" typeStatus="holotype">
<emphasis id="B991EAE9FFFEFF96D95DB64F8BFFFDD5" bold="true" box="[383,493,536,562]" pageId="1" pageNumber="248">
<typeStatus id="545E8859FFFEFF96D95DB64F8BFBFDD5" box="[383,489,536,562]" pageId="1" pageNumber="248" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="623B4E77FFFEFF96D9D7B64E8850FDD4" box="[501,578,537,563]" pageId="1" pageNumber="248">Mature</collectingCounty>
larva,
<collectingCountry id="F3F2766BFFFEFF96DAAFB64F88C9FDD5" box="[653,731,536,562]" name="India" pageId="1" pageNumber="248">INDIA</collectingCountry>
,
<collectingRegion id="4921F819FFFEFF96DAC5B64E8963FDD5" box="[743,881,536,563]" country="India" name="West Bengal" pageId="1" pageNumber="248">West Bengal</collectingRegion>
,
<collectingMunicipality id="6B3EAC81FFFEFF96DB5CB64F8E0CFDD5" box="[894,1054,536,562]" pageId="1" pageNumber="248">Purulia district</collectingMunicipality>
,
<location id="8E3A6020FFFEFF96DC08B64F8EA5FDD5" LSID="urn:lsid:plazi:treatment:034C87EDFFFEFF9FD8B5B5FC88CCFDFE:8E3A6020FFFEFF96DC08B64F8EA5FDD5" box="[1066,1207,536,562]" country="India" county="Mature" latitude="23.41131" longLatPrecision="1" longitude="86.19734" municipality="Purulia district" name="Jalika village" pageId="1" pageNumber="248" stateProvince="West Bengal">Jalika village</location>
stream,
<geoCoordinate id="EED1503CFFFEFF96DD37B64F8F8AFDD5" box="[1301,1432,536,562]" degrees="23.41131" direction="north" orientation="latitude" pageId="1" pageNumber="248" precision="1" value="23.41131">23.41131°N</geoCoordinate>
,
<geoCoordinate id="EED1503CFFFEFF96D8B5B66B8B0AFDB1" box="[151,280,572,598]" degrees="86.19734" direction="east" orientation="longitude" pageId="1" pageNumber="248" precision="1" value="86.19734">86.19734°E</geoCoordinate>
,
<specimenCount id="9DE3FD72FFFEFF96D904B66B8B60FDB0" box="[294,370,572,599]" count="311" pageId="1" pageNumber="248" type="generic">
<quantity id="4C1D9B1EFFFEFF96D904B66B8B7CFDB0" box="[294,366,572,599]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="248" unit="m" value="311.0">
<elevation id="00C8D1C8FFFEFF96D904B66B8B7CFDB0" box="[294,366,572,599]" metricMagnitude="2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="248" unit="m" value="311.0">311 m</elevation>
</quantity>
.
</specimenCount>
<date id="FF5B103BFFFEFF96D959B66B8BFCFDB1" box="[379,494,572,598]" pageId="1" pageNumber="248" value="2022-10-14">
<collectingDate id="EF1FE9D3FFFEFF96D959B66B8BFCFDB1" box="[379,494,572,598]" pageId="1" pageNumber="248" value="2022-10-14">14.X.2022</collectingDate>
</date>
, coll.
<collectionCode id="EDF4AE3EFFFEFF96DA10B66A8851FDB0" box="[562,579,573,599]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="248">T</collectionCode>
. Kubendran (Reg. no. I/E/864)
</materialsCitation>
.
<materialsCitation id="3B8D3CA6FFFEFF96DB84B66B8BABFD9C" collectingDate="2022-10-14" collectionCode="T" country="India" county="Mature" elevation="311" latitude="23.41131" location="Jalika village" longLatPrecision="1" longitude="86.19734" municipality="Purulia district" pageId="1" pageNumber="248" specimenCount="15" specimenCount-larva="15" stateProvince="West Bengal" typeStatus="paratype">
<emphasis id="B991EAE9FFFEFF96DB84B66B8E33FDB1" bold="true" box="[934,1057,572,598]" pageId="1" pageNumber="248">
<typeStatus id="545E8859FFFEFF96DB84B66B8E0EFDB1" box="[934,1052,572,598]" pageId="1" pageNumber="248" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9DE3FD72FFFEFF96DC08B66B8E83FDB1" box="[1066,1169,572,599]" count="15" pageId="1" pageNumber="248" type="larva">15 larvae</specimenCount>
, same place and data as holotype (Reg. no. I/E/865)
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF9A6673FFFEFF96D8B5B3988EECF800" box="[151,1278,1999,2023]" pageId="1" pageNumber="248" startId="1.[151,264,1999,2023]" targetBox="[214,1374,656,1973]" targetPageId="1" targetType="figure">
<paragraph id="8B5A36FBFFFEFF96D8B5B3988EECF800" blockId="1.[151,1278,1999,2023]" box="[151,1278,1999,2023]" pageId="1" pageNumber="248">
<emphasis id="B991EAE9FFFEFF96D8B5B3988B28F800" bold="true" box="[151,314,1999,2023]" pageId="1" pageNumber="248">FIGURES 13.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFFEFF96D9BCB39888A8F800" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[414,698,1999,2023]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="248" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFFEFF96D9BCB39888A8F800" box="[414,698,1999,2023]" italics="true" pageId="1" pageNumber="248">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFFEFF96DAE2B3988903F800" bold="true" box="[704,785,1999,2023]" pageId="1" pageNumber="248">
<taxonomicNameLabel id="A2A25792FFFEFF96DAE2B3988903F800" box="[704,785,1999,2023]" pageId="1" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1. Dorsal view; 2. Ventral view; 3. Lateral view.
</paragraph>
</caption>
<caption id="DF9A6673FFFDFF95D8B5B1C98F67FA51" box="[151,1397,1438,1463]" pageId="2" pageNumber="249" startId="2.[151,264,1438,1462]" targetBox="[225,1363,181,1414]" targetPageId="2" targetType="figure">
<paragraph id="8B5A36FBFFFDFF95D8B5B1C98F67FA51" blockId="2.[151,1397,1438,1463]" box="[151,1397,1438,1463]" pageId="2" pageNumber="249">
<emphasis id="B991EAE9FFFDFF95D8B5B1C98B29FA51" bold="true" box="[151,315,1438,1462]" pageId="2" pageNumber="249">FIGURES 45.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFFDFF95D9BCB1C988A8FA51" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[414,698,1438,1462]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="249" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFFDFF95D9BCB1C988A8FA51" box="[414,698,1438,1462]" italics="true" pageId="2" pageNumber="249">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFFDFF95DAE2B1C88903FA50" bold="true" box="[704,785,1439,1463]" pageId="2" pageNumber="249">
<taxonomicNameLabel id="A2A25792FFFDFF95DAE2B1C88903FA50" box="[704,785,1439,1463]" pageId="2" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
4. Dorsal view (middle instar); 5. Dorsal view (final instar).
</paragraph>
</caption>
<subSubSection id="C3FF6570FFFDFF90D8E5B25F8B82F928" lastPageId="7" lastPageNumber="254" pageId="2" pageNumber="249" type="description">
<paragraph id="8B5A36FBFFFDFF95D8E5B25F8876F9A0" blockId="2.[151,1436,1544,2039]" pageId="2" pageNumber="249">
<emphasis id="B991EAE9FFFDFF95D8E5B25F8BEDF9C4" bold="true" box="[199,511,1544,1571]" pageId="2" pageNumber="249">
Description.
<emphasis id="B991EAE9FFFDFF95D942B25E8BEAF9C4" bold="true" box="[352,504,1544,1571]" italics="true" pageId="2" pageNumber="249">Mature larva</emphasis>
.
</emphasis>
(
<figureCitation id="13DE2A7EFFFDFF95DA32B25E885FF9C4" box="[528,589,1545,1571]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="2" pageNumber="249">Fig.1</figureCitation>
). Length (mm): body 1314; cerci 9.510.5; head length 1.52; head width 3.33.5; prothorax width 3.54 (
<figureCitation id="13DE2A7EFFFDFF95DA21B27A8845F9A0" box="[515,599,1580,1607]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="2" pageNumber="249">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8B5A36FBFFFDFF95D8E5B2078F34F954" blockId="2.[151,1436,1544,2039]" pageId="2" pageNumber="249">
<emphasis id="B991EAE9FFFDFF95D8E5B2078B32F98D" bold="true" box="[199,288,1616,1642]" pageId="2" pageNumber="249">Colour.</emphasis>
Generally brownish with pale white (
<figureCitation id="13DE2A7EFFFDFF95DAF2B2068901F98C" box="[720,787,1617,1643]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="2" pageNumber="249">Fig. 1</figureCitation>
). Antennae are pale brown and very short (
<figureCitation id="13DE2A7EFFFDFF95DD26B2068F58F98C" box="[1284,1354,1616,1643]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="2" pageNumber="249">Fig. 5</figureCitation>
). Head sub quadrangular, brown with pale dots, and flattened; anterior margin of head with numerous small setae; dorsal surface with four pale dots; ocelli with three black dots mixed with pale colour in the middle (
<figureCitation id="13DE2A7EFFFDFF95DC91B2CE8F0BF954" box="[1203,1305,1688,1715]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 45" captionStartId-0="1.[151,264,1999,2023]" captionStartId-1="2.[151,264,1438,1462]" captionTargetBox-0="[214,1374,656,1973]" captionTargetBox-1="[225,1363,181,1414]" captionTargetId-0="figure-130@1.[214,1374,656,1973]" captionTargetId-1="figure-30@2.[225,1363,181,1414]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." captionText-1="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="2" pageNumber="249">Figs 15</figureCitation>
).
</paragraph>
<paragraph id="8B5A36FBFFFDFF94D8E5B2EA8F09FE36" blockId="2.[151,1436,1544,2039]" lastBlockId="3.[151,1437,150,465]" lastPageId="3" lastPageNumber="250" pageId="2" pageNumber="249">
<emphasis id="B991EAE9FFFDFF95D8E5B2EA8BA1F930" bold="true" box="[199,435,1725,1751]" pageId="2" pageNumber="249">Head. Mouthparts:</emphasis>
Labrum expanded laterally, 4.5× wider than long, anterior margin straight, medially bulged with a concave shape and small blunt crenations on either side (
<figureCitation id="13DE2A7EFFFDFF95DBE1B2B68E54F91C" box="[963,1094,1761,1787]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="2" pageNumber="249">Figs 1214</figureCitation>
), dorsal surface with a curved row composed of 1011 long stout setae (
<figureCitation id="13DE2A7EFFFDFF95DA4DB35288D1F8F8" box="[623,707,1797,1823]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="2" pageNumber="249">Fig. 14</figureCitation>
); dorsal surface and anterior margin covered with long, hair-like and stout setae; ventral surface of anterior margin with numerous long setae. Mandibles slightly bent in distal part; anterolateral margins with two rows of dense long hair-like setae; anterolateral margins with median bulge. Right mandible with two outer incisors broad with serrations on the inner margin, composed of 7 blunt teeth; inner incisor with two blunt teeth; prostheca with 67 thick setae surrounded with small setae; mola joined, posterior margin with 56 hair-like setae (
<figureCitation id="13DE2A7EFFFDFF95D95AB3EE8BEEF835" box="[376,508,1976,2003]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="2" pageNumber="249">Figs 1617</figureCitation>
). Left mandible with single outer incisor serrated with 6 blunt teeth; inner incisor with two blunt teeth, anterior teeth smaller than posterior; prostheca with 8 thick fimbriate submarginal setae; mola with 78 setae and posterior median margin with 5 hair-like setae (
<figureCitation id="13DE2A7EFFFCFF94DBA1B4C08E14FF56" box="[899,1030,151,177]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="3" pageNumber="250">Figs 1819</figureCitation>
). Maxilla with long, simple setae on inner margin on the lateral margin and dorsal surface with stout setae. Segment I of maxillary palp on dorsal surface proximally with numerous long thread-like setae and rows of thin and simple setae on outer margin; segment II long and apically broad with rows of numerous long hair-like setae on outer margin; apical segment small, pointed, with rows of simple setae on outer margin. Crown of the galea-lacinia of maxillae with 1516 comb-shaped bristles (
<figureCitation id="13DE2A7EFFFCFF94DD48B5708AF0FE82" captionStart="FIGURES 2024" captionStartId="5.[151,264,1421,1445]" captionTargetBox="[189,1398,181,1397]" captionTargetId="figure-19@5.[189,1398,181,1397]" captionTargetPageId="5" captionText="FIGURES 2024. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." pageId="3" pageNumber="250">Figs 2022</figureCitation>
). Hypopharynx of lingua with tuft of small robust setae on anterior margin, superlinguae expanded laterally, densely covered with long and thin setae replaced before the apex by very small setae down to the lower part of the superlinguae (
<figureCitation id="13DE2A7EFFFCFF94D917B5C48B95FE4A" box="[309,391,402,429]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="3" pageNumber="250">Fig. 15</figureCitation>
). Labium well developed, glossae dorsally with two distinct long setae (
<figureCitation id="13DE2A7EFFFCFF94DC86B5C48EE4FE4A" box="[1188,1270,403,429]" captionStart="FIGURES 2024" captionStartId="5.[151,264,1421,1445]" captionTargetBox="[189,1398,181,1397]" captionTargetId="figure-19@5.[189,1398,181,1397]" captionTargetPageId="5" captionText="FIGURES 2024. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." pageId="3" pageNumber="250">Fig. 23</figureCitation>
); glossae inner margin concave, outer margin serrated, and dorsal surface with numerous long hair-like setae (
<figureCitation id="13DE2A7EFFFCFF94DC98B5E08F1CFE36" box="[1210,1294,439,465]" captionStart="FIGURES 2024" captionStartId="5.[151,264,1421,1445]" captionTargetBox="[189,1398,181,1397]" captionTargetId="figure-19@5.[189,1398,181,1397]" captionTargetPageId="5" captionText="FIGURES 2024. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." pageId="3" pageNumber="250">Fig. 24</figureCitation>
).
</paragraph>
<caption id="DF9A6673FFFCFF94D8B5B3DD884EF80C" pageId="3" pageNumber="250" startId="3.[151,264,1930,1954]" targetBox="[213,1374,489,1907]" targetPageId="3" targetType="figure">
<paragraph id="8B5A36FBFFFCFF94D8B5B3DD884EF80C" blockId="3.[151,1437,1930,2027]" pageId="3" pageNumber="250">
<emphasis id="B991EAE9FFFCFF94D8B5B3DD8B5BF845" bold="true" box="[151,329,1930,1954]" pageId="3" pageNumber="250">FIGURES 611.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFFCFF94D994B3DC88C4F844" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[438,726,1931,1955]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="250" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFFCFF94D994B3DC88C4F844" box="[438,726,1931,1955]" italics="true" pageId="3" pageNumber="250">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFFCFF94DAFDB3DC8921F844" bold="true" box="[735,819,1931,1955]" pageId="3" pageNumber="250">
<taxonomicNameLabel id="A2A25792FFFCFF94DAFDB3DC8921F844" box="[735,819,1931,1955]" pageId="3" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament.
</paragraph>
</caption>
<caption id="DF9A6673FFFBFF93D8B5B32288DEF81E" pageId="4" pageNumber="251" startId="4.[151,264,1909,1933]" targetBox="[164,1423,181,1884]" targetPageId="4" targetType="figure">
<paragraph id="8B5A36FBFFFBFF93D8B5B32288DEF81E" blockId="4.[151,1437,1909,2041]" pageId="4" pageNumber="251">
<emphasis id="B991EAE9FFFBFF93D8B5B3228B4AF86A" bold="true" box="[151,344,1909,1933]" pageId="4" pageNumber="251">FIGURES 1219.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFFBFF93D9E7B32288F7F86A" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[453,741,1909,1933]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="251" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFFBFF93D9E7B32288F7F86A" box="[453,741,1909,1933]" italics="true" pageId="4" pageNumber="251">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFFBFF93DACCB3228950F86A" bold="true" box="[750,834,1909,1933]" pageId="4" pageNumber="251">
<taxonomicNameLabel id="A2A25792FFFBFF93DACCB3228950F86A" box="[750,834,1909,1933]" pageId="4" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view).
</paragraph>
</caption>
<caption id="DF9A6673FFFAFF92D8B5B1DA89E1FA2D" pageId="5" pageNumber="252" startId="5.[151,264,1421,1445]" targetBox="[189,1398,181,1397]" targetPageId="5" targetType="figure">
<paragraph id="8B5A36FBFFFAFF92D8B5B1DA89E1FA2D" blockId="5.[151,1436,1421,1482]" pageId="5" pageNumber="252">
<emphasis id="B991EAE9FFFAFF92D8B5B1DA8B46FA41" bold="true" box="[151,340,1421,1446]" pageId="5" pageNumber="252">FIGURES 2024.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFFAFF92D994B1D988C0FA41" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[438,722,1422,1446]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="252" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFFAFF92D994B1D988C0FA41" box="[438,722,1422,1446]" italics="true" pageId="5" pageNumber="252">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFFAFF92DAFAB1D9893AFA41" bold="true" box="[728,808,1422,1446]" pageId="5" pageNumber="252">
<taxonomicNameLabel id="A2A25792FFFAFF92DAFAB1D9893AFA41" box="[728,808,1422,1446]" pageId="5" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin).
</paragraph>
</caption>
<paragraph id="8B5A36FBFFFAFF92D8E5B24B899AF801" blockId="5.[151,1437,1564,2022]" pageId="5" pageNumber="252">
<emphasis id="B991EAE9FFFAFF92D8E5B24B8B35F9D1" bold="true" box="[199,295,1564,1590]" pageId="5" pageNumber="252">Thorax.</emphasis>
Pronotum moderately enlarged laterally, deeply concave medially, wider than the head, and basally not fused to the mesonotum (
<figureCitation id="13DE2A7EFFFAFF92D98CB2178BE1F9BD" box="[430,499,1600,1626]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="5" pageNumber="252">Figs 1</figureCitation>
,
<figureCitation id="13DE2A7EFFFAFF92DA22B2178802F9BD" box="[512,528,1600,1626]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="5" pageNumber="252">4</figureCitation>
,
<figureCitation id="13DE2A7EFFFAFF92DA39B2178828F9BC" box="[539,570,1600,1627]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="5" pageNumber="252">25</figureCitation>
), anterior margin of pronotum with a small projection at apex.Anteromedially with black or brown marking and posterior margin medially with black line (
<figureCitation id="13DE2A7EFFFAFF92DBD0B2338E28F999" box="[1010,1082,1636,1662]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="5" pageNumber="252">Figs 4</figureCitation>
,
<figureCitation id="13DE2A7EFFFAFF92DC64B2338E74F998" box="[1094,1126,1636,1663]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="5" pageNumber="252">25</figureCitation>
). Mesonotum with a mix of pale and brown color, anterolaterally with deep concavity (
<figureCitation id="13DE2A7EFFFAFF92DB07B2DF896AF944" box="[805,888,1672,1699]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="5" pageNumber="252">Fig. 25</figureCitation>
), and black posteriorly at lateroparapsidal suture. Legs yellowish brown with black markings (
<figureCitation id="13DE2A7EFFFAFF92DAABB2FB891EF921" box="[649,780,1708,1734]" captionStart="FIGURES 3137" captionStartId="7.[151,264,1466,1490]" captionTargetBox="[236,1386,181,1443]" captionTargetId="figure-19@7.[201,1386,181,1443]" captionTargetPageId="7" captionText="FIGURES 3137. Larva of Thalerosphyrus bengalensis sp. nov. 31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae." pageId="5" pageNumber="252">Figs 3137</figureCitation>
). Foreleg: femur with marginal row of long strong bristles, anterior and dorsal surface with numerous blunt stout setae, medially with small black maculae and distally with a small black spot (
<figureCitation id="13DE2A7EFFFAFF92D97DB2A38BF0F8E9" box="[351,482,1780,1806]" captionStart="FIGURES 3137" captionStartId="7.[151,264,1466,1490]" captionTargetBox="[236,1386,181,1443]" captionTargetId="figure-19@7.[201,1386,181,1443]" captionTargetPageId="7" captionText="FIGURES 3137. Larva of Thalerosphyrus bengalensis sp. nov. 31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae." pageId="5" pageNumber="252">Figs 3133</figureCitation>
). Tibia inner margin with few sharp stout setae (
<figureCitation id="13DE2A7EFFFAFF92DC2AB2A38E4FF8E9" box="[1032,1117,1780,1806]" captionStart="FIGURES 3843" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." pageId="5" pageNumber="252">Fig. 38</figureCitation>
). Tarsi without any setation. Midleg: femur dorsal surface with numerous long stout setae along with dorsal margin, medially with small black maculae, and distally with small black maculae (
<figureCitation id="13DE2A7EFFFAFF92DAE3B36B8954F8B0" box="[705,838,1852,1879]" captionStart="FIGURES 3137" captionStartId="7.[151,264,1466,1490]" captionTargetBox="[236,1386,181,1443]" captionTargetId="figure-19@7.[201,1386,181,1443]" captionTargetPageId="7" captionText="FIGURES 3137. Larva of Thalerosphyrus bengalensis sp. nov. 31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae." pageId="5" pageNumber="252">Figs 3435</figureCitation>
); tibia outer margin with rows of numerous hair-like setae and dorsal surface with U-shaped patella-tibial suture (
<figureCitation id="13DE2A7EFFFAFF92DB6AB337898CF89D" box="[840,926,1888,1914]" captionStart="FIGURES 3843" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." pageId="5" pageNumber="252">Fig. 39</figureCitation>
). Hindleg: similar to fore and midleg but the dorsal surface with arrow-shaped, blunt and strong setae (
<figureCitation id="13DE2A7EFFFAFF92DB34B3D3898BF879" box="[790,921,1924,1950]" captionStart="FIGURES 3137" captionStartId="7.[151,264,1466,1490]" captionTargetBox="[236,1386,181,1443]" captionTargetId="figure-19@7.[201,1386,181,1443]" captionTargetPageId="7" captionText="FIGURES 3137. Larva of Thalerosphyrus bengalensis sp. nov. 31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae." pageId="5" pageNumber="252">Figs 3637</figureCitation>
). Tibia with dorsally 1516 pointed bristles on marginal or submarginal position and U-shaped patella-tibial suture and tarsi outer margin with few hair-like setae (
<figureCitation id="13DE2A7EFFFAFF92D8BDB39B8B30F801" box="[159,290,1996,2022]" captionStart="FIGURES 3843" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." pageId="5" pageNumber="252">Figs 4041</figureCitation>
). Claws of all legs with 3 blunt denticles (
<figureCitation id="13DE2A7EFFFAFF92DADAB39B896EF801" box="[760,892,1996,2022]" captionStart="FIGURES 3843" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." pageId="5" pageNumber="252">Figs 4243</figureCitation>
).
</paragraph>
<caption id="DF9A6673FFF9FF91D8B5B11A8B9DFA4A" pageId="6" pageNumber="253" startId="6.[151,264,1357,1381]" targetBox="[201,1386,181,1333]" targetPageId="6" targetType="figure">
<paragraph id="8B5A36FBFFF9FF91D8B5B11A8B9DFA4A" blockId="6.[151,1436,1357,1453]" pageId="6" pageNumber="253">
<emphasis id="B991EAE9FFF9FF91D8B5B11A8B46FA82" bold="true" box="[151,340,1357,1381]" pageId="6" pageNumber="253">FIGURES 2530.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFF9FF91D995B11A88C1FA82" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[439,723,1357,1381]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="253" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFF9FF91D995B11A88C1FA82" box="[439,723,1357,1381]" italics="true" pageId="6" pageNumber="253">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFF9FF91DAFBB11A893BFA82" bold="true" box="[729,809,1357,1381]" pageId="6" pageNumber="253">
<taxonomicNameLabel id="A2A25792FFF9FF91DAFBB11A893BFA82" box="[729,809,1357,1381]" pageId="6" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament.
</paragraph>
</caption>
<paragraph id="8B5A36FBFFF9FF91D8E5B1A88850F809" blockId="6.[151,1437,1535,2030]" pageId="6" pageNumber="253">
<emphasis id="B991EAE9FFF9FF91D8E5B1A88B51F9FE" bold="true" box="[199,323,1535,1561]" pageId="6" pageNumber="253">Abdomen.</emphasis>
Posterolateral expansions on terga IIVIII and directed parallel to abdomen, slightly developed on terga IIIII and well developed on terga IVVIII (
<figureCitation id="13DE2A7EFFF9FF91DA9DB2748936F9D9" box="[703,804,1571,1598]" captionStart-0="FIGURES 45" captionStart-1="FIGURES 611" captionStartId-0="2.[151,264,1438,1462]" captionStartId-1="3.[151,264,1930,1954]" captionTargetBox-0="[225,1363,181,1414]" captionTargetBox-1="[213,1374,489,1907]" captionTargetId-0="figure-30@2.[225,1363,181,1414]" captionTargetId-1="figure-183@3.[213,1374,489,1907]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." captionText-1="FIGURES 611. Larva of Thalerosphyrus bengalensis sp. nov. 6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament." pageId="6" pageNumber="253">Figs 47</figureCitation>
). Dorsal surface of terga III pale with brown; tergum III with black with white spot; terga IVVI with distinct four pale spots (
<figureCitation id="13DE2A7EFFF9FF91DBBBB21089CCF985" box="[921,990,1607,1634]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="6" pageNumber="253">Fig. 4</figureCitation>
); terga VIIVIII medially pale band and laterally brown; tergum IX with anteriorly pale and posterior brown band; tergum X 1/3 brown and pale bands (
<figureCitation id="13DE2A7EFFF9FF91DD48B23C8AD6F94D" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="6" pageNumber="253">Figs 12</figureCitation>
); middle instar and mature larvae in slightly variable coloration (
<figureCitation id="13DE2A7EFFF9FF91DB82B2D889F7F94D" box="[928,997,1679,1706]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="6" pageNumber="253">Fig. 4</figureCitation>
). Dorsal surface of sternum commonly pale, medially brown, and laterally light brown spots (
<figureCitation id="13DE2A7EFFF9FF91DB22B2E48954F929" box="[768,838,1715,1742]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="6" pageNumber="253">Fig. 2</figureCitation>
); abdominal sterna IIIVII with laterally pale brown streaks; sterna VIIIX medially very dark brown maculae (
<figureCitation id="13DE2A7EFFF9FF91DB63B28089B8F916" box="[833,938,1751,1777]" captionStart="FIGURES 611" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[213,1374,489,1907]" captionTargetId="figure-183@3.[213,1374,489,1907]" captionTargetPageId="3" captionText="FIGURES 611. Larva of Thalerosphyrus bengalensis sp. nov. 6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament." pageId="6" pageNumber="253">Figs 67</figureCitation>
). Sternum I, joining to the metanotum, has brown streaked maculae (
<figureCitation id="13DE2A7EFFF9FF91D999B2AC8820F8F1" box="[443,562,1787,1814]" captionStart="FIGURES 611" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[213,1374,489,1907]" captionTargetId="figure-183@3.[213,1374,489,1907]" captionTargetPageId="3" captionText="FIGURES 611. Larva of Thalerosphyrus bengalensis sp. nov. 6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament." pageId="6" pageNumber="253">Figs 910</figureCitation>
). Posterior margins of each tergum with irregular pointed teeth and a row of two microdenticles (
<figureCitation id="13DE2A7EFFF9FF91D95DB3488813F8DD" box="[383,513,1823,1850]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="6" pageNumber="253">Figs 2829</figureCitation>
) and each tergum laterally with small simple setae. Gill I convex, asymmetrical, elongated, apically blunt, 2.5× longer than wide; ventral lamella, long fimbriate (
<figureCitation id="13DE2A7EFFF9FF91DC06B3148E6EF8B9" box="[1060,1148,1859,1886]" captionStart="FIGURES 4450" captionStartId="9.[151,264,883,907]" captionTargetBox="[177,1411,181,860]" captionTargetId="figure-19@9.[177,1411,181,860]" captionTargetPageId="9" captionText="FIGURES 4450. Larva of Thalerosphyrus bengalensis sp. nov. 44. Gill I; 45. Gill II; 46. Gill III; 47. Gill IV; 48. Gill V; 49. Gill VI; 50. Gill VII." pageId="6" pageNumber="253">Figs 44</figureCitation>
,
<figureCitation id="13DE2A7EFFF9FF91DCA5B3138EA6F8B9" box="[1159,1204,1860,1886]" captionStart="FIGURES 111114" captionStartId="19.[151,264,806,830]" captionTargetBox="[313,1274,183,780]" captionTargetId="figure-19@19.[311,1276,181,782]" captionTargetPageId="19" captionText="FIGURES 111114. Gill I of dorsal plate: 111. Th. flowersi (Paratype); 112. Th. flowersi (Thailand); 113. Th. bengalensis sp. nov.; 114. Th. sartorii sp. nov." pageId="6" pageNumber="253">113</figureCitation>
); Gill IIVI strongly asymmetrical, length longer than width or subequal, tracheae well developed (
<figureCitation id="13DE2A7EFFF9FF91DC04B3308EBFF865" box="[1062,1197,1895,1922]" captionStart="FIGURES 4450" captionStartId="9.[151,264,883,907]" captionTargetBox="[177,1411,181,860]" captionTargetId="figure-19@9.[177,1411,181,860]" captionTargetPageId="9" captionText="FIGURES 4450. Larva of Thalerosphyrus bengalensis sp. nov. 44. Gill I; 45. Gill II; 46. Gill III; 47. Gill IV; 48. Gill V; 49. Gill VI; 50. Gill VII." pageId="6" pageNumber="253">Figs 4549</figureCitation>
). Gill VII broad and apically blunt (
<figureCitation id="13DE2A7EFFF9FF91D960B3DC8B8AF841" box="[322,408,1931,1958]" captionStart="FIGURES 4450" captionStartId="9.[151,264,883,907]" captionTargetBox="[177,1411,181,860]" captionTargetId="figure-19@9.[177,1411,181,860]" captionTargetPageId="9" captionText="FIGURES 4450. Larva of Thalerosphyrus bengalensis sp. nov. 44. Gill I; 45. Gill II; 46. Gill III; 47. Gill IV; 48. Gill V; 49. Gill VI; 50. Gill VII." pageId="6" pageNumber="253">Fig. 50</figureCitation>
). Cerci dark brownish with band pale increasing in size towards the apex. Paracercus dark brownish with yellow, with a dark brown band every 56 segments and each segment surrounded with pointed small yellowish spine-like bristles (
<figureCitation id="13DE2A7EFFF9FF91D9C3B3848827F809" box="[481,565,2003,2030]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="6" pageNumber="253">Fig. 30</figureCitation>
).
</paragraph>
<caption id="DF9A6673FFF8FF90D8B5B1ED883CF9FD" pageId="7" pageNumber="254" startId="7.[151,264,1466,1490]" targetBox="[236,1386,181,1443]" targetPageId="7" targetType="figure">
<paragraph id="8B5A36FBFFF8FF90D8B5B1ED883CF9FD" blockId="7.[151,1436,1466,1563]" pageId="7" pageNumber="254">
<emphasis id="B991EAE9FFF8FF90D8B5B1ED8B41FA35" bold="true" box="[151,339,1466,1490]" pageId="7" pageNumber="254">FIGURES 3137.</emphasis>
Larva of
<taxonomicName id="4CE54D78FFF8FF90D996B1ED88C2FA35" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[436,720,1466,1490]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="254" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFF8FF90D996B1ED88C2FA35" box="[436,720,1466,1490]" italics="true" pageId="7" pageNumber="254">Thalerosphyrus bengalensis</emphasis>
</taxonomicName>
<emphasis id="B991EAE9FFF8FF90DAF7B1EC8936FA34" bold="true" box="[725,804,1467,1491]" pageId="7" pageNumber="254">
<taxonomicNameLabel id="A2A25792FFF8FF90DAF7B1EC8936FA34" box="[725,804,1467,1491]" pageId="7" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae.
</paragraph>
</caption>
<paragraph id="8B5A36FBFFF8FF90D8E5B23B8EE5F94C" blockId="7.[151,1437,1644,2032]" pageId="7" pageNumber="254">
<emphasis id="B991EAE9FFF8FF90D8E5B23B8B20F961" bold="true" box="[199,306,1644,1670]" pageId="7" pageNumber="254">Genitals.</emphasis>
Protopenis: male genitalia yellowish, apically blunt, and laterally expanded, segment X triangular and apically with numerous small setae (
<figureCitation id="13DE2A7EFFF8FF90DA12B2C68896F94C" box="[560,644,1681,1707]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="7" pageNumber="254">Fig. 27</figureCitation>
). Forceps proximally broad and distally narrow, simple.
</paragraph>
<paragraph id="8B5A36FBFFF8FF90D8E5B2E38B82F928" blockId="7.[151,1437,1644,2032]" box="[199,400,1716,1743]" pageId="7" pageNumber="254">
<emphasis id="B991EAE9FFF8FF90D8E5B2E38B01F929" bold="true" box="[199,275,1716,1742]" pageId="7" pageNumber="254">Adult.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF8FF90D8E5B28E8846F8F0" pageId="7" pageNumber="254" type="etymology">
<paragraph id="8B5A36FBFFF8FF90D8E5B28E8846F8F0" blockId="7.[151,1437,1644,2032]" pageId="7" pageNumber="254">
<emphasis id="B991EAE9FFF8FF90D8E5B28E8B5CF914" bold="true" box="[199,334,1753,1779]" pageId="7" pageNumber="254">Etymology.</emphasis>
The species name
<taxonomicName id="4CE54D78FFF8FF90DA03B28E88A3F914" authorityName="Vasanth &amp; Kubendran &amp; Subramanian &amp; Babu" authorityYear="2025" box="[545,689,1753,1779]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="254" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B991EAE9FFF8FF90DA0FB28E88A3F914" box="[557,689,1753,1779]" italics="true" pageId="7" pageNumber="254">bengalensis</emphasis>
</taxonomicName>
” refers to the state of
<collectingRegion id="4921F819FFF8FF90DB85B28E8E25F914" box="[935,1079,1753,1779]" country="India" name="West Bengal" pageId="7" pageNumber="254">West Bengal</collectingRegion>
,
<collectingCountry id="F3F2766BFFF8FF90DC60B28E8E6CF914" box="[1090,1150,1753,1779]" name="India" pageId="7" pageNumber="254">India</collectingCountry>
, from where the
<typeStatus id="545E8859FFF8FF90DD18B28E8F8EF914" box="[1338,1436,1753,1779]" pageId="7" pageNumber="254" type="holotype">holotype</typeStatus>
and
<typeStatus id="545E8859FFF8FF90D8E5B2AA8B35F8F0" box="[199,295,1789,1815]" pageId="7" pageNumber="254" type="paratype">paratype</typeStatus>
specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF8FF90D8E5B377896AF8DC" box="[199,888,1824,1851]" pageId="7" pageNumber="254" type="distribution">
<paragraph id="8B5A36FBFFF8FF90D8E5B377896AF8DC" blockId="7.[151,1437,1644,2032]" box="[199,888,1824,1851]" pageId="7" pageNumber="254">
<emphasis id="B991EAE9FFF8FF90D8E5B3778B73F8DD" bold="true" box="[199,353,1824,1850]" pageId="7" pageNumber="254">Distribution.</emphasis>
Purulia district,
<collectingRegion id="4921F819FFF8FF90DA39B37688BAF8DC" box="[539,680,1825,1851]" country="India" name="West Bengal" pageId="7" pageNumber="254">West Bengal</collectingRegion>
(
<collectingCountry id="F3F2766BFFF8FF90DA95B37688E6F8DC" box="[695,756,1825,1851]" name="India" pageId="7" pageNumber="254">India</collectingCountry>
) (
<figureCitation id="13DE2A7EFFF8FF90DB29B3768979F8DC" box="[779,875,1825,1851]" captionStart="FIGURES 111114" captionStartId="19.[151,264,806,830]" captionTargetBox="[313,1274,183,780]" captionTargetId="figure-19@19.[311,1276,181,782]" captionTargetPageId="19" captionText="FIGURES 111114. Gill I of dorsal plate: 111. Th. flowersi (Paratype); 112. Th. flowersi (Thailand); 113. Th. bengalensis sp. nov.; 114. Th. sartorii sp. nov." pageId="7" pageNumber="254">Fig. 111</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3FF6570FFF8FF9FD8E5B31388CCFDFE" lastPageId="8" lastPageNumber="255" pageId="7" pageNumber="254" type="diagnosis">
<paragraph id="8B5A36FBFFF8FF9FD8E5B31388CCFDFE" blockId="7.[151,1437,1644,2032]" lastBlockId="8.[151,1437,151,537]" lastPageId="8" lastPageNumber="255" pageId="7" pageNumber="254">
<emphasis id="B991EAE9FFF8FF90D8E5B3138B50F8B9" bold="true" box="[199,322,1860,1886]" pageId="7" pageNumber="254">Diagnosis.</emphasis>
<taxonomicName id="4CE54D78FFF8FF90D970B3128E4FF8B8" authority="Vasanth, Kubendran &amp; Subramanian" authorityName="Vasanth, Kubendran &amp; Subramanian" authorityYear="2025" box="[338,1117,1861,1887]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="254" phylum="Arthropoda" rank="species" species="bengalensis" status="sp. nov.">
<emphasis id="B991EAE9FFF8FF90D970B3128887F8B8" box="[338,661,1861,1887]" italics="true" pageId="7" pageNumber="254">Thalerosphyrus bengalensis</emphasis>
Vasanth, Kubendran &amp; Subramanian
</taxonomicName>
<emphasis id="B991EAE9FFF8FF90DC4FB3128EC2F8B8" bold="true" box="[1133,1232,1861,1887]" pageId="7" pageNumber="254">
<taxonomicNameLabel id="A2A25792FFF8FF90DC4FB3128EC2F8B8" box="[1133,1232,1861,1887]" pageId="7" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4CE54D78FFF8FF90DDAAB3118B2EF864" authorityName="Selvakumar &amp; Chandra" authorityYear="2017" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="254" phylum="Arthropoda" rank="species" species="meghalayensis">
<emphasis id="B991EAE9FFF8FF90DDAAB3118B2EF864" italics="true" pageId="7" pageNumber="254">T. meghalayensis</emphasis>
</taxonomicName>
but can be distinguished from all known species of
<taxonomicName id="4CE54D78FFF8FF90DBB1B33E8E50F864" authorityName="Eaton" authorityYear="1881" box="[915,1090,1897,1923]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="B991EAE9FFF8FF90DBB1B33E8E50F864" box="[915,1090,1897,1923]" italics="true" pageId="7" pageNumber="254">Thalerosphyrus</emphasis>
</taxonomicName>
by the following combination of characters: (i) posterolateral expansions on terga IIIII slightly developed and terga VIVIII well-developed (
<figureCitation id="13DE2A7EFFF8FF90D8BDB3E68AFBF82C" box="[159,233,1969,1995]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="7" pageNumber="254">Figs 2</figureCitation>
,
<figureCitation id="13DE2A7EFFF8FF90D8D5B3E68B36F82C" box="[247,292,1969,1995]" captionStart="FIGURES 611" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[213,1374,489,1907]" captionTargetId="figure-183@3.[213,1374,489,1907]" captionTargetPageId="3" captionText="FIGURES 611. Larva of Thalerosphyrus bengalensis sp. nov. 6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament." pageId="7" pageNumber="254">67</figureCitation>
); (ii) tergum III bears a broad black with white spot (
<figureCitation id="13DE2A7EFFF8FF90DBA4B3E689DCF82C" box="[902,974,1969,1995]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="7" pageNumber="254">Figs 1</figureCitation>
,
<figureCitation id="13DE2A7EFFF8FF90DBFFB3E68E1EF82B" box="[989,1036,1969,1996]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="7" pageNumber="254">45</figureCitation>
); (iii) terga VIIVIII medially with a pale white band and laterally brown; tergum IX with anteriorly pale and posteriorly brown band (
<figureCitation id="13DE2A7EFFF8FF90DCD5B3828F4DF817" box="[1271,1375,2005,2032]" captionStart="FIGURES 45" captionStartId="2.[151,264,1438,1462]" captionTargetBox="[225,1363,181,1414]" captionTargetId="figure-30@2.[225,1363,181,1414]" captionTargetPageId="2" captionText="FIGURES 45. Larva of Thalerosphyrus bengalensis sp. nov. 4. Dorsal view (middle instar); 5. Dorsal view (final instar)." pageId="7" pageNumber="254">Figs 45</figureCitation>
); (iv) abdominal sterna IIIVII with laterally brown streaks; sterna VIIIX medially very dark brown maculae (
<figureCitation id="13DE2A7EFFF7FF9FDD6FB4C08F85FF56" box="[1357,1431,151,177]" captionStart="FIGURES 13" captionStartId="1.[151,264,1999,2023]" captionTargetBox="[214,1374,656,1973]" captionTargetId="figure-130@1.[214,1374,656,1973]" captionTargetPageId="1" captionText="FIGURES 13. Larva of Thalerosphyrus bengalensis sp. nov. 1. Dorsal view; 2. Ventral view; 3. Lateral view." pageId="8" pageNumber="255">Figs 2</figureCitation>
,
<figureCitation id="13DE2A7EFFF7FF9FD8B5B4EC8AD7FF32" box="[151,197,187,213]" captionStart="FIGURES 611" captionStartId="3.[151,264,1930,1954]" captionTargetBox="[213,1374,489,1907]" captionTargetId="figure-183@3.[213,1374,489,1907]" captionTargetPageId="3" captionText="FIGURES 611. Larva of Thalerosphyrus bengalensis sp. nov. 6. Sternum VIX (middle instar); 7. Sternum VIX (final instar); 8. Head and thorax ventral view (LRM- Labrum, MX- Maxillae, LIM- Labium); 9. Sternum I (middle instar); 10. Sternum I (final instar); 11. Caudal filament." pageId="8" pageNumber="255">67</figureCitation>
); (v) posterior margins of abdominal terga with irregular pointed teeth and a row of few micro denticles (
<figureCitation id="13DE2A7EFFF7FF9FDD48B4EC8AF0FF1E" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="8" pageNumber="255">Figs 2829</figureCitation>
); (vi) pronotum moderately enlarged laterally, with ornamentation of anteromedian black marking (
<figureCitation id="13DE2A7EFFF7FF9FDD19B4888F9CFF1E" box="[1339,1422,222,249]" captionStart="FIGURES 2530" captionStartId="6.[151,264,1357,1381]" captionTargetBox="[201,1386,181,1333]" captionTargetId="figure-30@6.[201,1386,181,1333]" captionTargetPageId="6" captionText="FIGURES 2530. Larva of Thalerosphyrus bengalensis sp. nov. 25. Dorsal surface of notum; 26. Hindwing pad with segment I posterior margin; 27. Segment IX abdominal sterna of male; 28. Posterior margin of tergum V; 29. Posterior margin of tergum VI; 30. Caudal filament." pageId="8" pageNumber="255">Fig. 25</figureCitation>
); (vii) all femora with various stout setae, medially black maculae (
<figureCitation id="13DE2A7EFFF7FF9FDB50B55489E6FEFA" box="[882,1012,259,285]" captionStart="FIGURES 3137" captionStartId="7.[151,264,1466,1490]" captionTargetBox="[236,1386,181,1443]" captionTargetId="figure-19@7.[201,1386,181,1443]" captionTargetPageId="7" captionText="FIGURES 3137. Larva of Thalerosphyrus bengalensis sp. nov. 31. Forefemur; 32. Anterior marginal setae; 33. Dorsal surface with setae; 34. Midfemur (arrow indicates black maculae); 35. Dorsal surface with setae; 36. Hindfemur (arrow indicates black maculae); 37. Dorsal surface with setae." pageId="8" pageNumber="255">Figs 3137</figureCitation>
); (viii) hind femur dorsal surface with triangle-shaped blunt setae tibia 1516 pointed bristles on marginal or submarginal position (
<figureCitation id="13DE2A7EFFF7FF9FDCB9B5708EFCFEA6" box="[1179,1262,295,321]" captionStart="FIGURES 3843" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." pageId="8" pageNumber="255">Fig. 40</figureCitation>
); (xi) claw with 3 blunt denticles (
<figureCitation id="13DE2A7EFFF7FF9FD979B51C8BCFFE82" box="[347,477,331,357]" captionStart="FIGURES 3843" captionStartId="8.[151,264,1937,1961]" captionTargetBox="[218,1386,561,1914]" captionTargetId="figure-206@8.[201,1386,561,1914]" captionTargetPageId="8" captionText="FIGURES 3843. Larva of Thalerosphyrus bengalensis sp. nov. 38. Foretibia (arrow indicates inner marginal setae); 39. Midtibia (arrow indicates dorsal patella-tibial suture); 40. Hindtibia (arrow indicates dorsal patella-tibial suture); 41. Hindtibia (arrow indicates inner and outer rows of setae); 42. Claw; 43. Claw denticles closer view." pageId="8" pageNumber="255">Figs 4243</figureCitation>
); (x) gill I convex, asymmetrical, elongated, apically blunt, 2.5× longer than wide (
<figureCitation id="13DE2A7EFFF7FF9FDD4DB51C8AA5FE6E" captionStart="FIGURES 4450" captionStartId="9.[151,264,883,907]" captionTargetBox="[177,1411,181,860]" captionTargetId="figure-19@9.[177,1411,181,860]" captionTargetPageId="9" captionText="FIGURES 4450. Larva of Thalerosphyrus bengalensis sp. nov. 44. Gill I; 45. Gill II; 46. Gill III; 47. Gill IV; 48. Gill V; 49. Gill VI; 50. Gill VII." pageId="8" pageNumber="255">Fig. 44</figureCitation>
); (xi) labrum expanded laterally, 4.5× wider than long, anterior margin straight, medially bulged with a concave shape and small blunt crenations on either side, medio-ventrally with a curved row composed of 1011 long stout setae (
<figureCitation id="13DE2A7EFFF7FF9FD8FDB5E08B71FE36" box="[223,355,439,465]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="8" pageNumber="255">Figs 1214</figureCitation>
); (xii) hypopharynx broad, lingua apically with small setae and superlinguae expanded laterally (
<figureCitation id="13DE2A7EFFF7FF9FD8BDB58C8AE1FE12" box="[159,243,474,501]" captionStart="FIGURES 1219" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[164,1423,181,1884]" captionTargetId="figure-30@4.[164,1423,181,1884]" captionTargetPageId="4" captionText="FIGURES 1219. Larva of Thalerosphyrus bengalensis sp. nov. 12. Labrum; 13. Labrum closer view of lateral part; 14. Labrum closer view of medial part (arrows indicate medial concavity and marginal crenation); 15. Hypopharynx (arrow indicates superlinua setae); 16. Right mandible (arrow indicates incisor closer view); 17. Right mandible closer view; 18. Left mandible; 19. Left mandible (arrow indicates incisor closer view)." pageId="8" pageNumber="255">Fig. 15</figureCitation>
); (xiii) maxillae broad, galea-lacinia with 1516 comb-shaped bristles (
<figureCitation id="13DE2A7EFFF7FF9FDC37B58C8E78FE12" box="[1045,1130,475,501]" captionStart="FIGURES 2024" captionStartId="5.[151,264,1421,1445]" captionTargetBox="[189,1398,181,1397]" captionTargetId="figure-19@5.[189,1398,181,1397]" captionTargetPageId="5" captionText="FIGURES 2024. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." pageId="8" pageNumber="255">Fig. 20</figureCitation>
); (xvi) glossae broad, inner margin concave and outer margin serrated (
<figureCitation id="13DE2A7EFFF7FF9FDA5FB5A888C0FDFE" box="[637,722,511,537]" captionStart="FIGURES 2024" captionStartId="5.[151,264,1421,1445]" captionTargetBox="[189,1398,181,1397]" captionTargetId="figure-19@5.[189,1398,181,1397]" captionTargetPageId="5" captionText="FIGURES 2024. Larva of Thalerosphyrus bengalensis sp. nov. 20. Maxilla; 21. Segment III of maxillary palp; 22. Segment I of maxillary palp; 23. Labium; 24. Glossae (arrow indicates inner and outer margin)." pageId="8" pageNumber="255">Fig. 24</figureCitation>
).
</paragraph>
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(s. str.)
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TL = 12.0
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(mean =
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, n = 164) in males,
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<paragraph id="8B9136FA0329FFCCCED4E6816D72FC90" blockId="3.[151,1437,655,1978]" pageId="3" pageNumber="204">Coloration highly variable from unicoloured black, brown, blue, green, olive, cupreous-bronze or purple to bicolored with blue head and pronotum and blue-green or green elytra, or with blue or blue-green head and pronotum and cupreous or cupreous-green elytra.</paragraph>
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Head wider than pronotum, with protruding eyes; genae glabrous, deeply striated; clypeus asetose; frons long, virtually flat, finely striated; vertex with sparse soft hairs, shallowly rugose, with two wide shallow longitudinal depressions in the base of ocular plates; latter shallowly striated, with two long anterior and posterior setae; occiput indistinctly rugose. Labrum pale with dark brown or black narrowly infuscate wave-shaped anterior margin, unidentate, transverse, slightly trapeziform, especially in females, LW/LL =
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(mean =
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, n = 131) in females, with 414 (mean 8, n = 299) submarginal setae. Mandibles medium length, in males relatively stout, ML/MW = 5.438.0 (mean = 6.58, n = 63), in females thinner, ML/MW = 6.08.5 (mean = 7.38, n = 47); black with greenish-blue tinge except long pale lateral margin up to base of apical tooth. Maxillary palpi metallic blue, bluish-green, green or cuprous-green with purple or golden reflection. Labial palpomeres 2 and 3 pale, palpomere 1 brown, palpomere 4 metallic green or bluish green.Antennae relatively long, reach basal third of elytra, antennomeres 14 metallic blue, bluish-green or green with purple or golden reflection, antennomeres 511 black, densely covered by numerous very short white setae, each with two longer apical setae; scape besides 35 apical setae often with 13 rarely with 45 additional setae or glabrous; antennomere 3 with 410 and antennomere 4 with 16 short white setae.
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Aedeagus relatively long, distinctly depressed on apical third in right side (see in ventral view), EL/AL = 1.64 2.0 (mean = 1.80, n = 81), apex lanceolate, clearly curved with sharp hook, short lateral carina and small groove on dorsal side. Internal sac large, distinctly rounded towards to ventral side,
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long,
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long and smoothly curved,
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long, rarely short, as a rule with smoothly curved apex, VA and DA large, B distinctly larger than DLL, VLR prolonged, VB and VLL as flat membranous fields, BLR virtually invisible.
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</mods:part>
</mods:relatedItem>
<mods:location id="FFF3692B12C6AD4DAA7E12517A43B6BC">
<mods:url id="B3E747865190CB0FECFAEB687905AB98">https://doi.org/10.11646/zootaxa.5583.2.3</mods:url>
</mods:location>
<mods:classification id="7AFCB1C6A932470C12CCBFD2B0B8DACC">journal article</mods:classification>
<mods:identifier id="B9E5487BFDD5FEC9A9EA2D4E27E8F6FD" type="DOI">10.11646/zootaxa.5583.2.3</mods:identifier>
<mods:identifier id="6F6BFF4624E3C1AFB71FFD8593DC8E03" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6923F4044E058EE796F9184E9B74FB76" type="ZooBank">FC4C2C8C-B379-4E7C-9FBB-116783DDD4A0</mods:identifier>
</mods:mods>
<treatment id="03946562FFC8FF94FF1562EE616AF8A8" LSID="urn:lsid:plazi:treatment:03946562FFC8FF94FF1562EE616AF8A8" httpUri="http://treatment.plazi.org/id/03946562FFC8FF94FF1562EE616AF8A8" lastPageId="12" lastPageNumber="283" pageId="10" pageNumber="281">
<subSubSection id="C32787FFFFC8FF92FF1562EE6709F920" pageId="10" pageNumber="281" type="nomenclature">
<paragraph id="8B82D474FFC8FF92FF1562EE60A7F90D" blockId="10.[151,1070,1721,1820]" box="[151,896,1721,1748]" pageId="10" pageNumber="281">
<heading id="D0CA6318FFC8FF92FF1562EE60A7F90D" bold="true" box="[151,896,1721,1748]" fontSize="11" level="1" pageId="10" pageNumber="281" reason="1">
<emphasis id="B9490866FFC8FF92FF1562EE60A7F90D" bold="true" box="[151,896,1721,1748]" pageId="10" pageNumber="281">
<taxonomicName id="4C3DAFF7FFC8FF92FF1562EE61C3F90D" authority="Heppner &amp; Arita" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[151,740,1721,1748]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="281" phylum="Arthropoda" rank="species" species="fennchihwu" status="sp. nov.">
<emphasis id="B9490866FFC8FF92FF1562EE6124F90D" bold="true" box="[151,515,1721,1748]" italics="true" pageId="10" pageNumber="281">Thaumatographa fennchihwu</emphasis>
Heppner &amp; Arita
</taxonomicName>
,
<taxonomicNameLabel id="A27AB51DFFC8FF92FD7262ED60A7F90D" box="[752,896,1722,1748]" pageId="10" pageNumber="281" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B82D474FFC8FF92FF1562896709F920" blockId="10.[151,1070,1721,1820]" box="[151,1070,1758,1785]" pageId="10" pageNumber="281">
LSID:
<uri id="FFACD876FFC8FF92FF6662896709F920" box="[228,1070,1758,1785]" pageId="10" pageNumber="281">
urn:lsid:zoobank.org:act:
<uuid id="FF9BEEA1FFC8FF92FE7762896709F920" box="[501,1070,1758,1785]" pageId="10" pageNumber="281">FB01099C-EFFD-4AD5-A3F5-B8DABCB82F82</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC8FF92FF156355615CF887" pageId="10" pageNumber="281" type="description">
<paragraph id="8B82D474FFC8FF92FF1563556242F8C5" blockId="10.[151,1070,1721,1820]" box="[151,357,1794,1820]" pageId="10" pageNumber="281">
(
<figureCitation id="1306C8F1FFC8FF92FF1D6355622CF8C5" box="[159,267,1794,1820]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="10" pageNumber="281">Figs. 34</figureCitation>
,
<figureCitation id="1306C8F1FFC8FF92FE9563556213F8C5" box="[279,308,1794,1820]" captionStart="FIGURE 11" captionStartId="9.[151,250,1310,1334]" captionTargetBox="[151,1437,181,1287]" captionTargetId="figure-20@9.[151,1437,181,1287]" captionTargetPageId="9" captionText="FIGURE 11. Thaumatographa fennchihwu Heppner &amp; Arita, sp. nov., holotype male (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus enlarged (a) and with ductus ejaculatorius (with typical proximal elongated hood found in many Thaumatographa species), with enlargement of uncus to juxta (b), and structures of the pregenital abdomen and coremata (c) (gen. slide JBH- 4646) (MGCL)." pageId="10" pageNumber="281">11</figureCitation>
,
<figureCitation id="1306C8F1FFC8FF92FEC26355627AF8C5" box="[320,349,1794,1820]" captionStart="FIGURE 14" captionStartId="13.[151,250,1341,1365]" captionTargetBox="[152,1436,181,1317]" captionTargetId="figure-20@13.[151,1436,181,1317]" captionTargetPageId="13" captionText="FIGURE 14. Thaumatographa fennchihwu Heppner &amp; Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)." pageId="10" pageNumber="281">14</figureCitation>
)
</paragraph>
<paragraph id="8B82D474FFC8FF92FF156311615CF887" blockId="10.[151,635,1862,1886]" box="[151,635,1862,1886]" pageId="10" pageNumber="281">
Type-locality:
<collectingCountry id="F32A94E4FFC8FF92FEA96311625EF887" box="[299,377,1862,1886]" name="Taiwan" pageId="10" pageNumber="281">Taiwan</collectingCountry>
: Fennchihwu,
<collectingRegion id="49F91A96FFC8FF92FD8E63116177F887" box="[524,592,1862,1886]" country="Taiwan" name="Chiayi" pageId="10" pageNumber="281">Chiayi</collectingRegion>
Co.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC8FF93FF1563DA6673FF0C" lastPageId="11" lastPageNumber="282" pageId="10" pageNumber="281" type="materials_examined">
<paragraph id="8B82D474FFC8FF92FF1563DA66BAF836" blockId="10.[151,1437,1932,2031]" pageId="10" pageNumber="281">
<emphasis id="B9490866FFC8FF92FF1563DA6261F87F" bold="true" box="[151,326,1932,1959]" pageId="10" pageNumber="281">Type material.</emphasis>
<materialsCitation id="3B55DE29FFC8FF92FECD63DA61BDF812" collectingDate="1984-07-14" collectionCode="MGCL" collectorName="Y. Arita" country="Taiwan" elevation="1400" location="Fennchihwu" pageId="10" pageNumber="281" specimenCount="1" specimenCount-male="1" stateProvince="Chiayi" typeStatus="holotype">
<emphasis id="B9490866FFC8FF92FECD63DA6111F87F" bold="true" box="[335,566,1932,1959]" pageId="10" pageNumber="281">
<typeStatus id="54866AD6FFC8FF92FECD63DA6299F87E" box="[335,446,1933,1959]" pageId="10" pageNumber="281" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F32A94E4FFC8FF92FE5063DB6108F87F" box="[466,559,1932,1958]" name="Taiwan" pageId="10" pageNumber="281">Taiwan</collectingCountry>
]
</emphasis>
<specimenCount id="9D3B1FFDFFC8FF92FDBC63DA6174F87E" box="[574,595,1933,1959]" count="1" pageId="10" pageNumber="281" type="male"></specimenCount>
“Fennchihhu” [
<location id="8EE282AFFFC8FF92FC8563DB60BFF87F" LSID="urn:lsid:plazi:treatment:03946562FFC8FF94FF1562EE616AF8A8:8EE282AFFFC8FF92FC8563DB60BFF87F" box="[775,920,1932,1958]" country="Taiwan" name="Fennchihwu" pageId="10" pageNumber="281" stateProvince="Chiayi">Fennchihwu</location>
] (
<quantity id="4CC57991FFC8FF92FC2C63DA672EF87E" box="[942,1033,1933,1959]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="281" unit="m" value="1400.0">
<elevation id="00103347FFC8FF92FC2C63DA672EF87E" box="[942,1033,1933,1959]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="281" unit="m" value="1400.0">1400 m</elevation>
</quantity>
),
<collectingRegion id="49F91A96FFC8FF92FB9D63DB674DF87F" box="[1055,1130,1932,1958]" country="Taiwan" name="Chiayi" pageId="10" pageNumber="281">Chiayi</collectingRegion>
Co.,
<date id="FF83F2B4FFC8FF92FB2B63DA661EF87E" box="[1193,1337,1932,1959]" pageId="10" pageNumber="281" value="1984-07-14">
<collectingDate id="EFC70B5CFFC8FF92FB2B63DA661EF87E" box="[1193,1337,1932,1959]" pageId="10" pageNumber="281" value="1984-07-14">14 Jul. 1984</collectingDate>
</date>
,
<collectorName id="26C8B1A2FFC8FF92FAC663DA66BAF87F" box="[1348,1437,1932,1959]" pageId="10" pageNumber="281">Y. Arita</collectorName>
(adult photo 5570; gen. JBH-4646) (
<collectionCode id="ED2C4CB1FFC8FF92FDB963E661B5F812" box="[571,658,1969,1995]" pageId="10" pageNumber="281">MGCL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B55DE29FFC8FF92FD2563E661E3F836" collectingDate="1984-07-14" country="Taiwan" elevation="1158" location="Taiwan" pageId="10" pageNumber="281" specimenCount="34" specimenCount-female="17" specimenCount-male="17" stateProvince="Chiayi" typeStatus="paratype">
<emphasis id="B9490866FFC8FF92FD2563E66038F812" bold="true" box="[679,799,1969,1995]" pageId="10" pageNumber="281">
<typeStatus id="54866AD6FFC8FF92FD2563E66038F812" box="[679,799,1969,1995]" pageId="10" pageNumber="281" type="paratype">Paratypes</typeStatus>
</emphasis>
. (
<specimenCount id="9D3B1FFDFFC8FF92FCBA63E7604BF813" box="[824,876,1968,1994]" count="16" pageId="10" pageNumber="281" type="male">16♂</specimenCount>
,
<specimenCount id="9D3B1FFDFFC8FF92FCF863E76089F812" box="[890,942,1968,1995]" count="15" pageId="10" pageNumber="281" type="female">15♀</specimenCount>
; n = 31).
<location id="8EE282AFFFC8FF92FBA063E76750F813" LSID="urn:lsid:plazi:treatment:03946562FFC8FF94FF1562EE616AF8A8:8EE282AFFFC8FF92FBA063E76750F813" box="[1058,1143,1968,1994]" country="Taiwan" name="Taiwan" pageId="10" pageNumber="281" stateProvince="Chiayi">Taiwan</location>
,
<collectingRegion id="49F91A96FFC8FF92FB0763E767F7F813" box="[1157,1232,1968,1994]" country="Taiwan" name="Chiayi" pageId="10" pageNumber="281">Chiayi</collectingRegion>
Co.: same site as
<typeStatus id="54866AD6FFC8FF92FF15638363DBF837" box="[151,252,2004,2030]" pageId="10" pageNumber="281" type="holotype">holotype</typeStatus>
,
<date id="FF83F2B4FFC8FF92FE84638262AAF836" box="[262,397,2004,2031]" pageId="10" pageNumber="281" value="1984-07-14">
<collectingDate id="EFC70B5CFFC8FF92FE84638262AAF836" box="[262,397,2004,2031]" pageId="10" pageNumber="281" value="1984-07-14">14 Jul. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFC8FF92FE1F638262E6F836" box="[413,449,2005,2031]" count="1" pageId="10" pageNumber="281" type="male">1♂</specimenCount>
,
<specimenCount id="9D3B1FFDFFC8FF92FE4E638262C9F836" box="[460,494,2005,2031]" count="2" pageId="10" pageNumber="281" type="female">2♀</specimenCount>
) (gen. YA-
<quantity id="4CC57991FFC8FF92FDEC6383619BF836" box="[622,700,2004,2031]" metricMagnitude="3" metricUnit="m" metricValue="1.158" pageId="10" pageNumber="281" unit="m" value="1158.0">
<elevation id="00103347FFC8FF92FDEC6383619BF836" box="[622,700,2004,2031]" metricMagnitude="3" metricUnit="m" metricValue="1.158" pageId="10" pageNumber="281" unit="m" value="1158.0">1158m</elevation>
</quantity>
)
</materialsCitation>
;
<date id="FF83F2B4FFC8FF92FD4F63836073F836" box="[717,852,2004,2031]" pageId="10" pageNumber="281" value="1984-07-16">16 Jul. 1984</date>
(
<specimenCount id="9D3B1FFDFFC8FF92FCE1638260AFF836" box="[867,904,2005,2031]" count="3" pageId="10" pageNumber="281" type="male">3♂</specimenCount>
,
<specimenCount id="9D3B1FFDFFC8FF92FC1163826092F836" box="[915,949,2005,2031]" count="7" pageId="10" pageNumber="281" type="female">7♀</specimenCount>
) (gen. JBH-4647
<specimenCount id="9D3B1FFDFFC8FF92FB00638367B7F837" box="[1154,1168,2004,2030]" count="1" pageId="10" pageNumber="281" type="female"></specimenCount>
);
<date id="FF83F2B4FFC8FF92FB206382660EF836" box="[1186,1321,2004,2031]" pageId="10" pageNumber="281" value="1984-07-18">18 Jul. 1984</date>
(
<specimenCount id="9D3B1FFDFFC8FF92FABA6383667AF837" box="[1336,1373,2004,2030]" count="6" pageId="10" pageNumber="281" type="male">6♂</specimenCount>
,
<specimenCount id="9D3B1FFDFFC8FF92FAEA638266AEF836" box="[1384,1417,2005,2031]" count="1" pageId="10" pageNumber="281" type="female">1♀</specimenCount>
);
</paragraph>
<paragraph id="8B82D474FFC9FF93FF1564C06673FF0C" blockId="11.[151,1436,150,213]" pageId="11" pageNumber="282">
<materialsCitation id="3B55DE29FFC9FF93FF1564C06767FF68" box="[151,1088,150,177]" collectingDate="1984-07-19" collectionCode="MGCL" country="Taiwan" elevation="200" location="Wulai" municipality="Co." pageId="11" pageNumber="282" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Taipei">
<date id="FF83F2B4FFC9FF93FF1564C06238FF68" box="[151,287,151,177]" pageId="11" pageNumber="282" value="1984-07-19">
<collectingDate id="EFC70B5CFFC9FF93FF1564C06238FF68" box="[151,287,151,177]" pageId="11" pageNumber="282" value="1984-07-19">19 Jul. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFC9FF93FEAC64C16274FF68" box="[302,339,150,177]" count="5" pageId="11" pageNumber="282" type="male">5♂</specimenCount>
,
<specimenCount id="9D3B1FFDFFC9FF93FEDC64C062A6FF68" box="[350,385,151,177]" count="4" pageId="11" pageNumber="282" type="female">4♀</specimenCount>
) (gen. YA-1192f) (
<collectionCode id="ED2C4CB1FFC9FF93FDD764C0618BFF68" box="[597,684,151,177]" pageId="11" pageNumber="282">MGCL</collectionCode>
).
<collectingCountry id="F32A94E4FFC9FF93FD3F64C06035FF68" box="[701,786,151,177]" name="Taiwan" pageId="11" pageNumber="282">Taiwan</collectingCountry>
,
<collectingRegion id="49F91A96FFC9FF93FC9E64C06045FF68" box="[796,866,151,177]" country="Taiwan" name="Taipei" pageId="11" pageNumber="282">Taipei</collectingRegion>
<collectingMunicipality id="6BE64E0EFFC9FF93FCE564C060B4FF68" box="[871,915,151,177]" pageId="11" pageNumber="282">Co.</collectingMunicipality>
:
<location id="8EE282AFFFC9FF93FC1E64C060F8FF68" LSID="urn:lsid:plazi:treatment:03946562FFC8FF94FF1562EE616AF8A8:8EE282AFFFC9FF93FC1E64C060F8FF68" box="[924,991,151,177]" country="Taiwan" municipality="Co." name="Wulai" pageId="11" pageNumber="282" stateProvince="Taipei">Wulai</location>
(
<quantity id="4CC57991FFC9FF93FC6F64C0671FFF68" box="[1005,1080,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="282" unit="m" value="200.0">
<elevation id="00103347FFC9FF93FC6F64C0671FFF68" box="[1005,1080,151,177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="282" unit="m" value="200.0">200 m</elevation>
</quantity>
)
</materialsCitation>
,
<date id="FF83F2B4FFC9FF93FBC964C067F4FF68" box="[1099,1235,151,177]" pageId="11" pageNumber="282" value="1988-08-02">2 Aug. 1988</date>
(
<specimenCount id="9D3B1FFDFFC9FF93FB6164C06621FF68" box="[1251,1286,151,177]" count="1" pageId="11" pageNumber="282" type="male">1♂</specimenCount>
) (adult photo 14171; gen. JBH-
<quantity id="4CC57991FFC9FF93FEDD64EC6297FF0C" box="[351,432,187,213]" metricMagnitude="3" metricUnit="m" metricValue="3.763" pageId="11" pageNumber="282" unit="m" value="3763.0">3763m</quantity>
),
<materialsCitation id="3B55DE29FFC9FF93FE4164EC605DFF0C" box="[451,890,187,213]" collectionCode="MGCL" collectorName="J. B. Heppner" country="Taiwan" elevation="400" location="Wulai" pageId="11" pageNumber="282" specimenCount="1" stateProvince="Taipei">
<collectorName id="26C8B1A2FFC9FF93FE4164EC617BFF0C" box="[451,604,187,213]" pageId="11" pageNumber="282">J. B. Heppner</collectorName>
(
<collectionCode id="ED2C4CB1FFC9FF93FDE964EC61E5FF0C" box="[619,706,187,213]" pageId="11" pageNumber="282">MGCL</collectionCode>
).
<location id="8EE282AFFFC9FF93FD5764EC603FFF0C" LSID="urn:lsid:plazi:treatment:03946562FFC8FF94FF1562EE616AF8A8:8EE282AFFFC9FF93FD5764EC603FFF0C" box="[725,792,187,213]" country="Taiwan" name="Wulai" pageId="11" pageNumber="282" stateProvince="Taipei">Wulai</location>
(
<quantity id="4CC57991FFC9FF93FCA564EC6055FF0C" box="[807,882,187,213]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="282" unit="m" value="400.0">
<elevation id="00103347FFC9FF93FCA564EC6055FF0C" box="[807,882,187,213]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="282" unit="m" value="400.0">400 m</elevation>
</quantity>
)
</materialsCitation>
,
<materialsCitation id="3B55DE29FFC9FF93FC0464EC6677FF0C" box="[902,1360,186,213]" collectingDate="1985-08-14" collectionCode="NSMT" collectorName="Kawabe" country="Taiwan" pageId="11" pageNumber="282" specimenCount="1" specimenCount-female="1" stateProvince="Taipei">
<date id="FF83F2B4FFC9FF93FC0464EC6739FF0C" box="[902,1054,186,213]" pageId="11" pageNumber="282" value="1985-08-14">
<collectingDate id="EFC70B5CFFC9FF93FC0464EC6739FF0C" box="[902,1054,186,213]" pageId="11" pageNumber="282" value="1985-08-14">14 Aug. 1985</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFC9FF93FBB264EC6777FF0C" box="[1072,1104,187,213]" count="1" pageId="11" pageNumber="282" type="female">1♀</specimenCount>
), A.
<collectorName id="26C8B1A2FFC9FF93FB0B64EC67C2FF0C" box="[1161,1253,187,213]" pageId="11" pageNumber="282">Kawabe</collectorName>
(
<collectionCode id="ED2C4CB1FFC9FF93FB7664EC666FFF0C" box="[1268,1352,187,213]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="11" pageNumber="282">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF4284FCFFC9FF93FF1561F8670FF9C9" pageId="11" pageNumber="282" startId="11.[151,250,1455,1479]" targetBox="[160,1436,237,1432]" targetPageId="11" targetType="figure">
<paragraph id="8B82D474FFC9FF93FF1561F8670FF9C9" blockId="11.[151,1436,1455,1552]" pageId="11" pageNumber="282">
<emphasis id="B9490866FFC9FF93FF1561F86204FA1E" bold="true" box="[151,291,1455,1479]" pageId="11" pageNumber="282">FIGURE 13.</emphasis>
<taxonomicName id="4C3DAFF7FFC9FF93FEAF61F861D0FA1E" authority="Diakonoff" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[301,759,1455,1479]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="282" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFC9FF93FEAF61F861A4FA1E" box="[301,643,1455,1479]" italics="true" pageId="11" pageNumber="282">Thaumatographa mesostigmatias</emphasis>
Diakonoff
</taxonomicName>
, female genitalia (Taiwan: Chilan Nsy., Ilan Co.) (with remnant spermatophore intact inside bursa), with details of ostium, sterigma and colliculum bulge (a), signum (b), and added view of colliculum bulge interior (c) (gen. slide JBH-4645, except YA-1193 for detail c) (MGCL).
</paragraph>
</caption>
<subSubSection id="C32787FFFFC9FF93FF45626A6230F8E9" pageId="11" pageNumber="282" type="diagnosis">
<paragraph id="8B82D474FFC9FF93FF45626A6230F8E9" blockId="11.[151,1437,1597,2020]" pageId="11" pageNumber="282">
<emphasis id="B9490866FFC9FF93FF45626A621DF98E" bold="true" box="[199,314,1597,1623]" pageId="11" pageNumber="282">Diagnosis</emphasis>
. A large species for the genus (
<figureCitation id="1306C8F1FFC9FF93FD2362696028F981" box="[673,783,1598,1624]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="11" pageNumber="282">Figs. 34</figureCitation>
), with forewings black-brown with orange-marked striae, and a large orange tornal patch with two black spots; the hindwings are black-brown with a prominent yellow patch on middle of wing to base and a narrow cubital narrow pale orange patch, and fringe is prominent and white; and both sexes with equal maculation. The male genitalia have the valvae oblong-quadratic, with a truncated termen, the uncus is short with a wide base, the hami are short and somewhat flattened, and the socii are short, wide and very setose. The female genitalia are typical for the genus but the ductus bursae has a large diverticular sac on its caudal end.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFC9FF94FF45636E6674FAD4" lastPageId="12" lastPageNumber="283" pageId="11" pageNumber="282" type="description">
<paragraph id="8B82D474FFC9FF93FF45636E67EFF88C" blockId="11.[151,1437,1597,2020]" box="[199,1224,1849,1877]" pageId="11" pageNumber="282">
<emphasis id="B9490866FFC9FF93FF45636E6276F88A" bold="true" box="[199,337,1849,1875]" pageId="11" pageNumber="282">Description</emphasis>
. Wing expanse: males12.819.0 mm (n = 17), females 14.819.8 mm (n = 15).
</paragraph>
<paragraph id="8B82D474FFC9FF93FF45630966B7F83D" blockId="11.[151,1437,1597,2020]" pageId="11" pageNumber="282">
<emphasis id="B9490866FFC9FF93FF4563096223F8A1" box="[199,260,1886,1912]" italics="true" pageId="11" pageNumber="282">Head</emphasis>
(
<figureCitation id="1306C8F1FFC9FF93FE9563096281F8A1" box="[279,422,1886,1912]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="11" pageNumber="282">Figs. 3a, 4a</figureCitation>
): dark brown, with orange-yellow scales laterally by neck and medially on vertex and frons; eyes large, black-brown; ocellus large, dark brown; antennal scape amber to brown; antenna brown (orange ventrally); labial palpus mostly white, with brown to yellow dorsally on basal segment and somewhat on middle segment, with bluish iridescence on apical segment; head venter white, with neck fringe yellow to orange-yellow.
</paragraph>
<paragraph id="8B82D474FFCEFF94FF4564C066A4FF20" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4564C0623FFF68" box="[199,280,151,177]" italics="true" pageId="12" pageNumber="283">Thorax</emphasis>
(
<figureCitation id="1306C8F1FFCEFF94FEA564C062B3FF68" box="[295,404,151,177]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="12" pageNumber="283">Figs. 34</figureCitation>
): dark brown, with light orange as lateral margin on prothorax, becoming more tawny orange on mesothorax and metathorax, with orange caudal margin; patagia dark brown with orange from prothoax; tegula dark brown with orange laterally; venter white; legs white, with black-brown markings, as in the related species.
</paragraph>
<paragraph id="8B82D474FFCEFF94FF45655461ECFDC0" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4565546214FEC5" box="[199,307,259,284]" italics="true" pageId="12" pageNumber="283">Forewing</emphasis>
(
<figureCitation id="1306C8F1FFCEFF94FEC065546297FEC4" box="[322,432,259,285]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="12" pageNumber="283">Figs. 34</figureCitation>
): black-brown, with orange striae and lines from costa and dorsal margin (in generic form like
<taxonomicName id="4C3DAFF7FFCEFF94FF48657062B3FE99" authorityName="Diakonoff &amp; Arita" authorityYear="1977" box="[202,404,295,320]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="283" phylum="Arthropoda" rank="species" species="mesostigmatias">
<emphasis id="B9490866FFCEFF94FF48657062B3FE99" box="[202,404,295,320]" italics="true" pageId="12" pageNumber="283">T. mesostigmatias</emphasis>
</taxonomicName>
but darker orange); five costal bars angled to termen, with pale orange-yellow on margin, then silvery toward middle of wing, and dull orange suffusion on dorsal 1/3 of forewing from middle to apex and interrupted by black curved preapical bar; forewing apex with an orange marginal line, sharply bent inwards before a pale yellow to white bar near mid-termen; tornus with a large elongated marginal orange patch with two black spots in middle and a pair of tiny black spots dorsad near white mark of termen; fringe brown with yellow at tornus and near mid-termen; venter dark brown with pale yellow in cell area and marks of dorsum very faintly repeated, except distinct on costa, with dorsal margin white.
</paragraph>
<paragraph id="8B82D474FFCEFF94FF4566746668FD5C" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4566746212FDE4" box="[199,309,547,573]" italics="true" pageId="12" pageNumber="283">Hindwing</emphasis>
(
<figureCitation id="1306C8F1FFCEFF94FEC56674629FFDE4" box="[327,440,547,573]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="12" pageNumber="283">Figs. 34</figureCitation>
): black-brown with a large oblong orange-yellow patch, narrowing to wingbase, and a similar more orange and narrower patch on the cubitus near wingbase; costa white; fringe widely white to pale yellow at wing margin, and pale orange along anal margin; venter like dorsum but more suffused, and costa brown.
</paragraph>
<paragraph id="8B82D474FFCEFF94FF4566D8676CFD28" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4566D86216FD70" box="[199,305,655,681]" italics="true" pageId="12" pageNumber="283">Abdomen</emphasis>
(
<figureCitation id="1306C8F1FFCEFF94FEC266D8628AFD70" box="[320,429,655,681]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="12" pageNumber="283">Figs. 34</figureCitation>
): brown (dark brown on T-1), with orange posterior margins on tergites; venter tan; genital tufts light brown with tan-white venter; pregenital segment with sternite strongly invaginated caudally, with strong coremata from lateral base pouches; caudal tergite with similar median invagination.
</paragraph>
<paragraph id="8B82D474FFCEFF94FF4566AC602FFBC8" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4566AC6248FCCC" box="[199,367,763,789]" italics="true" pageId="12" pageNumber="283">Male genitalia</emphasis>
(
<figureCitation id="1306C8F1FFCEFF94FE0266AC62F1FCCC" box="[384,470,763,789]" captionStart="FIGURE 11" captionStartId="9.[151,250,1310,1334]" captionTargetBox="[151,1437,181,1287]" captionTargetId="figure-20@9.[151,1437,181,1287]" captionTargetPageId="9" captionText="FIGURE 11. Thaumatographa fennchihwu Heppner &amp; Arita, sp. nov., holotype male (Taiwan: Fennchihwu, Chiayi Co.), with aedeagus enlarged (a) and with ductus ejaculatorius (with typical proximal elongated hood found in many Thaumatographa species), with enlargement of uncus to juxta (b), and structures of the pregenital abdomen and coremata (c) (gen. slide JBH- 4646) (MGCL)." pageId="12" pageNumber="283">Fig. 11</figureCitation>
): tegumen quadratic, slightly narrowed dorsally with rounded dorsal corners; uncus short, narrowing to semi-acute decumbent apex from widened base; hamus short and widened, somewhat flattened; socius short but very wide and setose; gnathos membranous and rounded; transtilla relatively straight and narrow, with moderate ventral appendage each side of anellus with acute end; juxta small and spoon-like; anellus membranous, undeveloped; aedeagus small and short tubular, with small decumbent phallobase; vesica scobinate; cornuti undeveloped; ductus ejaculatorius short, with long proximal hood appendage; valva oblong quadratic, with round-blunt termen and dorsal apex slightly angular and more setose; saccular margin slightly convex near base; vinculum triangular; saccus very short, with blunt apex.
</paragraph>
<paragraph id="8B82D474FFCEFF94FF45604C6674FAD4" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF45604C62A2FBEC" box="[199,389,1051,1077]" italics="true" pageId="12" pageNumber="283">Female genitalia</emphasis>
(
<figureCitation id="1306C8F1FFCEFF94FE11604C62C1FBEC" box="[403,486,1051,1077]" captionStart="FIGURE 14" captionStartId="13.[151,250,1341,1365]" captionTargetBox="[152,1436,181,1317]" captionTargetId="figure-20@13.[151,1436,181,1317]" captionTargetPageId="13" captionText="FIGURE 14. Thaumatographa fennchihwu Heppner &amp; Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)." pageId="12" pageNumber="283">Fig. 14</figureCitation>
): ovipositor very short (2/3 length of segment 7), with elongate-oval horizontal dorsal median flap; papilla anales large, setose and widened caudally; apophyses subequal in length; ostium (
<figureCitation id="1306C8F1FFCEFF94FA986068665CFB80" box="[1306,1403,1087,1113]" captionStart="FIGURE 14" captionStartId="13.[151,250,1341,1365]" captionTargetBox="[152,1436,181,1317]" captionTargetId="figure-20@13.[151,1436,181,1317]" captionTargetPageId="13" captionText="FIGURE 14. Thaumatographa fennchihwu Heppner &amp; Arita, sp. nov., paratype female (Taiwan: Fennchichwu, Chiayi Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum (b) (gen. slide JBH-4647, except YA-1192 for detail b) (MGCL)." pageId="12" pageNumber="283">Fig. 14a</figureCitation>
), a deep funnel, gradually narrowing to junction with ductus bursae, with a band of sclerotized scobination on narrower section; sterigma a concave membranous margin; ductus bursae long and evenly narrow, with a prominent ventral diverticular sac (or small pouch) with internally spiculate walls and bifid distally from junction with antrum; ductus seminalis from ductus bursae proximal end; corpus bursae oblong-ovate, with one end more acute; oblong-ovate accessory diverticulum from thin tube to near ductus bursae base; signum a large typical form for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCEFF94FF45614061D0FA3C" pageId="12" pageNumber="283" type="etymology">
<paragraph id="8B82D474FFCEFF94FF45614061D0FA3C" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF456140626FFAE8" bold="true" box="[199,328,1303,1329]" pageId="12" pageNumber="283">Etymology</emphasis>
. The species is named for the
<typeStatus id="54866AD6FFCEFF94FD28614061FDFAE8" box="[682,730,1303,1329]" pageId="12" pageNumber="283">type</typeStatus>
locality, “Fennchihwu” in
<collectingRegion id="49F91A96FFCEFF94FB8D6140677DFAE8" box="[1039,1114,1303,1329]" country="Taiwan" name="Chiayi" pageId="12" pageNumber="283">Chiayi</collectingRegion>
Co.,
<collectingCountry id="F32A94E4FFCEFF94FB18614067D7FAE8" box="[1178,1264,1303,1329]" name="Taiwan" pageId="12" pageNumber="283">Taiwan</collectingCountry>
. There is some discrepancy with the English spelling of the village, but we use the spelling used by the
<collectingCountry id="F32A94E4FFCEFF94FBE9616C679AFA8C" box="[1131,1213,1339,1365]" name="Taiwan" pageId="12" pageNumber="283">Taiwan</collectingCountry>
Museum (
<bibRefCitation id="EFACA985FFCEFF94FAB6616C63C7FAA0" author="Heppner, J. B." pageId="12" pageNumber="283" pagination="85 - 106" refId="ref11384" refString="Heppner, J. B. (2012 b) Historical review of Lepidoptera studies in Taiwan. Lepidoptera Novae, 5, 85-106, 1 pl." type="journal article" year="2012">Heppner, 2012b</bibRefCitation>
, c, d). This locality, Fennchihwu, was formerly a Japanese logging camp and a railway stop on the Alishan rail line going higher to the town of Alishan, on the slopes of the high mountain of the same name (Mt. Ali portion of Yushan): this mountain has been heavily logged for over 100 years but small patches of original forest still remain and are now expanding with new conservation efforts.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCEFF94FF4561B96020F9F4" pageId="12" pageNumber="283" type="biology_ecology">
<paragraph id="8B82D474FFCEFF94FF4561B96020F9F4" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4561B96206F9D1" bold="true" box="[199,289,1518,1544]" pageId="12" pageNumber="283">Biology</emphasis>
. Unknown. Recorded elevations in
<collectingCountry id="F32A94E4FFCEFF94FD4261B86035F9D0" box="[704,786,1519,1545]" name="Taiwan" pageId="12" pageNumber="283">Taiwan</collectingCountry>
range from
<quantity id="4CC57991FFCEFF94FC2161B860CAF9D0" box="[931,1005,1519,1545]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="283" unit="m" value="200.0">200 m</quantity>
to
<quantity id="4CC57991FFCEFF94FB9761B8674AF9D0" box="[1045,1133,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="283" unit="m" value="1400.0">1400 m</quantity>
in montane forest habitat. Flight period (n = 47 records): mid-July to mid-August.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCEFF94FF456261628DF940" pageId="12" pageNumber="283" type="distribution">
<paragraph id="8B82D474FFCEFF94FF456261628DF940" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF456261627EF989" bold="true" box="[199,345,1590,1616]" pageId="12" pageNumber="283">Distribution</emphasis>
. Known only from
<collectingCountry id="F32A94E4FFCEFF94FDBE626061B6F988" box="[572,657,1591,1617]" name="Taiwan" pageId="12" pageNumber="283">Taiwan</collectingCountry>
, in
<collectingRegion id="49F91A96FFCEFF94FD3E62606020F988" box="[700,775,1591,1617]" country="Taiwan" name="Chiayi" pageId="12" pageNumber="283">Chiayi</collectingRegion>
Co., and also the low mountain site near
<collectingRegion id="49F91A96FFCEFF94FB666260660AF988" box="[1252,1325,1591,1617]" country="Taiwan" name="Taipei" pageId="12" pageNumber="283">Taipei</collectingRegion>
, in
<collectingRegion id="49F91A96FFCEFF94FAD5626066BAF988" box="[1367,1437,1591,1617]" country="Taiwan" name="Taipei" pageId="12" pageNumber="283">Taipei</collectingRegion>
Co., called Wulai. The Wulai area has various elevations, with the main village elevation at
<quantity id="4CC57991FFCEFF94FB0D620C67F2F9AC" box="[1167,1237,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="283" unit="m" value="200.0">200 m</quantity>
and forest habitat going higher from there.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCEFF94FF4562F5616AF8A8" pageId="12" pageNumber="283" type="discussion">
<paragraph id="8B82D474FFCEFF94FF4562F5616AF8A8" blockId="12.[151,1437,151,1905]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FF4562F56262F965" bold="true" box="[199,325,1698,1724]" pageId="12" pageNumber="283">Discussion</emphasis>
. This is one of the few
<taxonomicName id="4C3DAFF7FFCEFF94FDDE62F4605FF964" box="[604,888,1699,1725]" pageId="12" pageNumber="283">
<emphasis id="B9490866FFCEFF94FDDE62F46039F964" box="[604,798,1699,1725]" italics="true" pageId="12" pageNumber="283">Thaumatographa</emphasis>
species
</taxonomicName>
with an ample specimen sample, and
<typeStatus id="54866AD6FFCEFF94FAB362F466BBF964" box="[1329,1436,1699,1725]" pageId="12" pageNumber="283" type="paratype">paratypes</typeStatus>
will be distributed also to other museum collections (RMNH, TFRI, USNM). The species is one of the larger species of the genus. Its habitus superficially resembles species of the tropical Asian tortricid genus
<taxonomicName id="4C3DAFF7FFCEFF94FB6C62BC6268F8F0" authority="Diakonoff, 1975" authorityName="Diakonoff" authorityYear="1975" class="Insecta" family="Tortricidae" genus="Eucosmogastra" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFCEFF94FB6C62BC66BBF8DD" box="[1262,1436,1771,1796]" italics="true" pageId="12" pageNumber="283">Eucosmogastra</emphasis>
<bibRefCitation id="EFACA985FFCEFF94FF1563586268F8F0" author="Diakonoff, A." box="[151,335,1806,1833]" pageId="12" pageNumber="283" pagination="297 - 320" refId="ref10870" refString="Diakonoff, A. (1975) New Tortricoidea (Lepidoptera) from Southeast Asia in the British Museum (Natural History). Zoologische Mededelingen, Leiden, 48, 297-320, 8 pls." type="book chapter" year="1975">Diakonoff, 1975</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C3DAFF7FFCEFF94FEDE635862DCF8F0" authorityName="Walsingham" authorityYear="1895" box="[348,507,1807,1833]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="274" phylum="Arthropoda" rank="subFamily" subFamily="Olethreutinae">Olethreutinae</taxonomicName>
:
<taxonomicName id="4C3DAFF7FFCEFF94FD86635861B2F8F0" box="[516,661,1807,1833]" class="Insecta" family="Tortricidae" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="283" phylum="Arthropoda" rank="tribe" tribe="Enarmoniini">Enarmoniini</taxonomicName>
), which also have the yellow-patched hindwing, but differ in forewing maculation. The bifid ventral diverticulum of the caudal end on the ductus bursae has no known function but is possibly some kind of gland apparatus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,294 @@
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<mods:title id="BB443EAAEB8F456326878BE67BA7B962">Review of Thaumatographa tortricids in Taiwan (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini)</mods:title>
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<mods:namePart id="B04AAB06B3C383A0C4D36806A17D5335">Heppner, J. B.</mods:namePart>
<mods:affiliation id="BAA137673BD7E36752707F0D1DBAED6D">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida, USA. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="5019A15A774F79FEBBF45313C8AD6F7C">Arita, Yutaka</mods:namePart>
<mods:affiliation id="2AF7C85669FBC61B007833CED0BF4F14">Zoological Laboratory, Faculty of Agriculture, Meijo University, Tempaku-ku, Nagoya 468 - 85 - 2, Japan.</mods:affiliation>
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</mods:role>
<mods:namePart id="64C0DF21B315694EF0F3D01062B2016F">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="BEAC31FC087DE13AADEDF5F5DEC5E795" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="6C621B4518290A1EF1B2F16D16001054">Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
<mods:nameIdentifier id="6AF39F59085357ADAC164507CDE3CFCA" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<treatment id="03946562FFCCFF97FF1564C06760FE50" LSID="urn:lsid:plazi:treatment:03946562FFCCFF97FF1564C06760FE50" httpUri="http://treatment.plazi.org/id/03946562FFCCFF97FF1564C06760FE50" lastPageId="15" lastPageNumber="286" pageId="14" pageNumber="285">
<subSubSection id="C32787FFFFCCFF96FF1564C0673EFF0F" pageId="14" pageNumber="285" type="nomenclature">
<paragraph id="8B82D474FFCCFF96FF1564C060F2FF68" blockId="14.[151,1049,151,250]" box="[151,981,151,178]" pageId="14" pageNumber="285">
<heading id="D0CA6318FFCCFF96FF1564C060F2FF68" bold="true" box="[151,981,151,178]" fontSize="11" level="1" pageId="14" pageNumber="285" reason="1">
<emphasis id="B9490866FFCCFF96FF1564C060F2FF68" bold="true" box="[151,981,151,178]" pageId="14" pageNumber="285">
<taxonomicName id="4C3DAFF7FFCCFF96FF1564C0602AFF68" authority="Heppner &amp; Arita" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[151,781,151,178]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="subSpecies" species="aurosa" status="subsp. nov." subSpecies="taiwana">
<emphasis id="B9490866FFCCFF96FF1564C0610AFF68" bold="true" box="[151,557,151,178]" italics="true" pageId="14" pageNumber="285">Thaumatographa aurosa taiwana</emphasis>
Heppner &amp; Arita
</taxonomicName>
,
<taxonomicNameLabel id="A27AB51DFFCCFF96FC9864C060F2FF68" box="[794,981,151,177]" pageId="14" pageNumber="285" rank="subSpecies">new subspecies</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B82D474FFCCFF96FF1564EC673EFF0F" blockId="14.[151,1049,151,250]" box="[151,1049,187,214]" pageId="14" pageNumber="285">
LSID:
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urn:lsid:zoobank.org:act:
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<subSubSection id="C32787FFFFCCFF96FF1564886017FEE5" pageId="14" pageNumber="285" type="description">
<paragraph id="8B82D474FFCCFF96FF156488623EFF23" blockId="14.[151,1049,151,250]" box="[151,281,223,250]" pageId="14" pageNumber="285">
(
<figureCitation id="1306C8F1FFCCFF96FF1D648863CFFF23" box="[159,232,223,250]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Figs 5</figureCitation>
,
<figureCitation id="1306C8F1FFCCFF96FF7664886236FF23" box="[244,273,223,250]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." pageId="14" pageNumber="285">15</figureCitation>
)
</paragraph>
<paragraph id="8B82D474FFCCFF96FF1565736017FEE5" blockId="14.[151,816,292,316]" box="[151,816,292,316]" pageId="14" pageNumber="285">
Type-locality:
<collectingCountry id="F32A94E4FFCCFF96FEA96573625EFEE5" box="[299,377,292,316]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
: Lien-Hua-Chih For. Sta., Puli,
<collectingRegion id="49F91A96FFCCFF96FD3965736021FEE2" box="[699,774,292,315]" country="Taiwan" name="Nantou" pageId="14" pageNumber="285">Nantou</collectingRegion>
Co.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF15653D6786FE29" pageId="14" pageNumber="285" type="materials_examined">
<paragraph id="8B82D474FFCCFF96FF15653D6786FE29" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF15653D6261FE5A" bold="true" box="[151,326,361,388]" pageId="14" pageNumber="285">Type material.</emphasis>
<materialsCitation id="3B55DE29FFCCFF96FECC653D6015FE71" collectingDate="1983-09-07" collectingDateMax="1983-09-12" collectingDateMin="1983-09-07" collectionCode="MGCL" collectorName="J. B. Heppner" country="Taiwan" elevation="700" location="Nantou" pageId="14" pageNumber="285" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="holotype">
<emphasis id="B9490866FFCCFF96FECC653D6112FE5A" bold="true" box="[334,565,361,388]" pageId="14" pageNumber="285">
<typeStatus id="54866AD6FFCCFF96FECC653D629AFE5D" box="[334,445,362,388]" pageId="14" pageNumber="285" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F32A94E4FFCCFF96FE53653E6109FE5A" box="[465,558,361,387]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
]
</emphasis>
<specimenCount id="9D3B1FFDFFCCFF96FDBF653D6175FE5D" box="[573,594,362,388]" count="1" pageId="14" pageNumber="285" type="female"></specimenCount>
Lien-Hua-Chih For. Sta. (
<quantity id="4CC57991FFCCFF96FCFE653D60EEFE5D" box="[892,969,362,388]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="285" unit="m" value="700.0">
<elevation id="00103347FFCCFF96FCFE653D60EEFE5D" box="[892,969,362,388]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="285" unit="m" value="700.0">700 m</elevation>
</quantity>
), [nr. Puli],
<collectingRegion id="49F91A96FFCCFF96FBD0653D6783FE5D" box="[1106,1188,362,388]" country="Taiwan" name="Nantou" pageId="14" pageNumber="285">Nantou</collectingRegion>
Co.,
<date id="FF83F2B4FFCCFF96FB66653D66BFFE5D" box="[1252,1432,362,388]" pageId="14" pageNumber="285" value="1983-09-07" valueMax="1983-09-12" valueMin="1983-09-07">
<collectingDate id="EFC70B5CFFCCFF96FB66653D66BFFE5D" box="[1252,1432,362,388]" pageId="14" pageNumber="285" value="1983-09-07" valueMax="1983-09-12" valueMin="1983-09-07">712 Sep. 1983</collectingDate>
</date>
,
<collectorName id="26C8B1A2FFCCFF96FF1565D96216FE71" box="[151,305,398,424]" pageId="14" pageNumber="285">J. B. Heppner</collectorName>
(adult photo 5572; gen. JBH-4643) (
<collectionCode id="ED2C4CB1FFCCFF96FD5065D9600EFE71" box="[722,809,398,424]" pageId="14" pageNumber="285">MGCL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B55DE29FFCCFF96FCBE65D967BAFE29" collectingDate="1984-10-11" collectingDateMax="1984-10-13" collectingDateMin="1984-10-11" collectionCode="MGCL" collectorName="J. B. Heppner &amp; H. - Y. Wang" country="Taiwan" county="Ilan Co." elevation="100" location="Taiwan" municipality="Taiwan" pageId="14" pageNumber="285" specimenCount="2" specimenCount-female="2" stateProvince="Nantou" typeStatus="paratype">
<emphasis id="B9490866FFCCFF96FCBE65D96093FE71" bold="true" box="[828,948,398,424]" pageId="14" pageNumber="285">
<typeStatus id="54866AD6FFCCFF96FCBE65D96093FE71" box="[828,948,398,424]" pageId="14" pageNumber="285" type="paratype">Paratypes</typeStatus>
</emphasis>
. (n = 2).
<collectingMunicipality id="6BE64E0EFFCCFF96FB9F65D96755FE71" box="[1053,1138,398,424]" pageId="14" pageNumber="285">Taiwan</collectingMunicipality>
,
<collectingRegion id="49F91A96FFCCFF96FBFC65D967F7FE71" box="[1150,1232,398,424]" country="Taiwan" name="Nantou" pageId="14" pageNumber="285">Nantou</collectingRegion>
Co.: same data as holotype (
<specimenCount id="9D3B1FFDFFCCFF96FE8F65E5620BFE15" box="[269,300,434,460]" count="1" pageId="14" pageNumber="285" type="female">1♀</specimenCount>
) (adult photo 16013; gen. JBH-4644) (
<collectionCode id="ED2C4CB1FFCCFF96FD6C65E56062FE15" box="[750,837,434,460]" pageId="14" pageNumber="285">MGCL</collectionCode>
).
<location id="8EE282AFFFCCFF96FCDB65E56089FE15" LSID="urn:lsid:plazi:treatment:03946562FFCCFF97FF1564C06760FE50:8EE282AFFFCCFF96FCDB65E56089FE15" box="[857,942,434,460]" county="Ilan Co." municipality="Taiwan" name="Taiwan" pageId="14" pageNumber="285" stateProvince="Nantou">Taiwan</location>
,
<collectingCounty id="62E3ACF8FFCCFF96FC3965E5673CFE15" box="[955,1051,434,460]" pageId="14" pageNumber="285">Ilan Co.</collectingCounty>
:
<location id="8EE282AFFFCCFF96FBA565E56757FE15" LSID="urn:lsid:plazi:treatment:03946562FFCCFF97FF1564C06760FE50:8EE282AFFFCCFF96FBA565E56757FE15" box="[1063,1136,434,460]" county="Ilan Co." municipality="Taiwan" name="Nanao" pageId="14" pageNumber="285" stateProvince="Nantou">Nanao</location>
(
<quantity id="4CC57991FFCCFF96FB0065E56799FE15" box="[1154,1214,434,461]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="14" pageNumber="285" unit="mi" value="5.0">5 mi.</quantity>
N) (
<quantity id="4CC57991FFCCFF96FB7665E56666FE15" box="[1268,1345,434,460]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="285" unit="m" value="100.0">
<elevation id="00103347FFCCFF96FB7665E56666FE15" box="[1268,1345,434,460]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="285" unit="m" value="100.0">100 m</elevation>
</quantity>
),
<date id="FF83F2B4FFCCFF96FAD565E56221FE29" pageId="14" pageNumber="285" value="1984-10-11" valueMax="1984-10-13" valueMin="1984-10-11">
<collectingDate id="EFC70B5CFFCCFF96FAD565E56221FE29" pageId="14" pageNumber="285" value="1984-10-11" valueMax="1984-10-13" valueMin="1984-10-11">1113 Oct. 1984</collectingDate>
</date>
(
<specimenCount id="9D3B1FFDFFCCFF96FE9A6581621FFE29" box="[280,312,470,496]" count="1" pageId="14" pageNumber="285" type="female">1♀</specimenCount>
),
<collectorName id="26C8B1A2FFCCFF96FED2658162CDFE29" box="[336,490,470,496]" pageId="14" pageNumber="285">J. B. Heppner</collectorName>
&amp;
<collectorName id="26C8B1A2FFCCFF96FD8C658161B5FE29" box="[526,658,470,496]" pageId="14" pageNumber="285">H.-Y. Wang</collectorName>
(adult photo 16014; gen. JBH-4642) (
<collectionCode id="ED2C4CB1FFCCFF96FBBC658167B2FE29" box="[1086,1173,470,496]" pageId="14" pageNumber="285">MGCL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF4565AE62C0FD35" pageId="14" pageNumber="285" type="diagnosis">
<paragraph id="8B82D474FFCCFF96FF4565AE62C0FD35" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4565AE621DFDCA" bold="true" box="[199,314,505,531]" pageId="14" pageNumber="285">Diagnosis</emphasis>
. This new subspecies is very similar to
<taxonomicName id="4C3DAFF7FFCCFF96FC9265AD667BFDCD" authority="Diakonoff &amp; Arita (1976)" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[784,1372,506,532]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFCCFF96FC9265AD670DFDCD" box="[784,1066,506,532]" italics="true" pageId="14" pageNumber="285">Thaumatographa aurosa</emphasis>
<bibRefCitation id="EFACA985FFCCFF96FBB165AD667BFDCD" author="Diakonoff, A. &amp; Arita, Y." box="[1075,1372,506,532]" pageId="14" pageNumber="285" pagination="179 - 219" refId="ref11038" refString="Diakonoff, A. &amp; Arita, Y. (1976) New and old species of Glyphipterigidae and a new species of Acrolepiidae from Japan and the Far East (Lepidoptera). Zoologische Mededelingen, 50, 179-219." type="journal article" year="1976">Diakonoff &amp; Arita (1976)</bibRefCitation>
</taxonomicName>
from southern
<collectingCountry id="F32A94E4FFCCFF96FE8066496265FDE1" box="[258,322,542,568]" name="Japan" pageId="14" pageNumber="285">Japan</collectingCountry>
(Kyushu and Yakushima), and has the following distinctions: on the forewing the basal pair of vertical orange lines are more curved distally from the dorsal margin (straight in nominate
<taxonomicName id="4C3DAFF7FFCCFF96FB09661467D4FD85" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[1163,1267,579,604]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFCCFF96FB09661467D4FD85" box="[1163,1267,579,604]" italics="true" pageId="14" pageNumber="285">T. aurosa</emphasis>
</taxonomicName>
) and the costal angled streaks are white on the costal margin (silvery blue in nominate
<taxonomicName id="4C3DAFF7FFCCFF96FC2F66306733FD59" authorityName="Diakonoff &amp; Arita" authorityYear="1976" box="[941,1044,615,640]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="species" species="aurosa">
<emphasis id="B9490866FFCCFF96FC2F66306733FD59" box="[941,1044,615,640]" italics="true" pageId="14" pageNumber="285">T. aurosa</emphasis>
</taxonomicName>
). The female genitalia are virtually identical and
<taxonomicName id="4C3DAFF7FFCCFF96FEAC66DC62DFFD7D" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[302,504,651,676]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="285" phylum="Arthropoda" rank="subSpecies" species="aurosa" subSpecies="taiwana">
<emphasis id="B9490866FFCCFF96FEAC66DC62DFFD7D" box="[302,504,651,676]" italics="true" pageId="14" pageNumber="285">T. aurosa taiwana</emphasis>
</taxonomicName>
has the ovipositor somewhat more slender caudally, especially very narrow papilla anales (slightly wider in the nominate subspecies from
<collectingCountry id="F32A94E4FFCCFF96FC8566F9606BFD11" box="[775,844,686,712]" name="Japan" pageId="14" pageNumber="285">Japan</collectingCountry>
). Male genitalia could not be compared, lacking any male specimens from
<collectingCountry id="F32A94E4FFCCFF96FE0F668562C4FD35" box="[397,483,722,748]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCCFF96FF4566A26209F86D" pageId="14" pageNumber="285" type="description">
<paragraph id="8B82D474FFCCFF96FF4566A26785FCC9" blockId="14.[151,1437,361,1972]" box="[199,1186,757,784]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4566A26276FCD6" bold="true" box="[199,337,757,783]" pageId="14" pageNumber="285">Description</emphasis>
. Wing expanse: male 10.014.0 mm (
<collectingCountry id="F32A94E4FFCCFF96FD7966A16067FCC9" box="[763,832,758,784]" name="Japan" pageId="14" pageNumber="285">Japan</collectingCountry>
), female 11.013.6 mm (n = 3).
</paragraph>
<paragraph id="8B82D474FFCCFF96FF45674D6109FCEC" blockId="14.[151,1437,361,1972]" box="[199,558,794,821]" pageId="14" pageNumber="285">
Female (
<figureCitation id="1306C8F1FFCCFF96FEAB674D624AFCEC" box="[297,365,794,821]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Fig. 5</figureCitation>
).
<emphasis id="B9490866FFCCFF96FE01674D62E7FCED" box="[387,448,794,820]" italics="true" pageId="14" pageNumber="285">Head</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FE4D674D6106FCEC" box="[463,545,794,821]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Fig. 5a</figureCitation>
).
</paragraph>
<paragraph id="8B82D474FFCCFF96FF45676860E1FB45" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4567686214FC81" box="[199,307,831,856]" italics="true" pageId="14" pageNumber="285">Forewing</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEC6676962ADFC80" box="[324,394,830,857]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Fig. 5</figureCitation>
): overall black-brown, with usual geneedric striae pattern, with five thin costal striae white and slightly silvery from margin (more silvery in nominate subspecies), with basal two strongly bent to termen on radius; basal streaks straight and long, with dorsal on orange-yellow and middle streak yellow-white, and at end of forewing cell is a prominent tan-white spot; three curved light yellow-white lines from dorsal margin at middle of forewing, with a 4
<superScript id="7C48793CFFCCFF96FE07679A62B5FC02" attach="left" box="[389,402,973,987]" fontSize="6" pageId="14" pageNumber="285">th</superScript>
slightly further distad; tornal quarter with four series of interrupted striae curved slightly to termen; forewing apex with a white subapical line convergent and recurved as a loop to similar preapical line, both margined with orange; termen orange, with mid-termen white margin on termen and toward tornus 3 black spots; fringe white; venter dark brown with light yellow in cell and dorsal markings faintly repeated except for distinct white marks on costa, and dorsal margin white; venter brown with dorsal lines repeated as dull light tan lines, with white liens of apex repeated, and likewise the tornal markings but duller.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4560F163E5FAD1" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4560F16212FB19" box="[199,309,1190,1216]" italics="true" pageId="14" pageNumber="285">Hindwing</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEC660F162A0FB18" box="[324,391,1190,1217]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Fig. 5</figureCitation>
): dark brown with thin orange-yellow streak diffusely margined in cell area, and diffuse tawny orange area on anal sector to base; fringe white; venter bronze-brown, with pale white markings on costa on basal 1/3.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4561456149FAAD" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF456145623FFAF5" box="[199,280,1298,1324]" italics="true" pageId="14" pageNumber="285">Thorax</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEAB61456257FAF4" box="[297,368,1298,1325]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Fig. 5</figureCitation>
): like nominate subspecies, dark brown, lacking any longitudinal lines; tegula slightly lighter brown than thorax; venter white; legs white, with dark brown markings dorsally on tibiae like related species, as dorsal spots on tibiae and tarsal segments.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF45612966AAFA41" blockId="14.[151,1437,361,1972]" box="[199,1421,1406,1433]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4561296216FA41" box="[199,305,1406,1432]" italics="true" pageId="14" pageNumber="285">Abdomen</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FEC2612962A2FA40" box="[320,389,1406,1433]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="14" pageNumber="285">Fig. 5</figureCitation>
): dark brown, with posterior margins of tergites tan; venter cream-white; genital tuft brown.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4561F560AFF961" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
<emphasis id="B9490866FFCCFF96FF4561F562AEFA65" box="[199,393,1442,1468]" italics="true" pageId="14" pageNumber="285">Female genitalia</emphasis>
(
<figureCitation id="1306C8F1FFCCFF96FE1961F562D6FA64" box="[411,497,1442,1469]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." pageId="14" pageNumber="285">Fig. 15</figureCitation>
): ovipositor short (subequal to length of segment 7) (dorsal flap lacking); papilla anales narrow, with caudal ends semi-acute; apophyses with anterior pair slightly longer; ostium (
<figureCitation id="1306C8F1FFCCFF96FB5F6191661BFA38" box="[1245,1340,1478,1505]" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." pageId="14" pageNumber="285">Fig. 15a</figureCitation>
), a short funnel, narrowed proximally with a sclerotized and scobinate proximal ventral wall (divided into two halves); sterigma membranous, widened from ostial funnel and slightly concave; ductus bursae evenly narrow and long (ca. 4x as long as width of bursa), with slight sclerotization on caudal 1/10, and with small ventral diverticular sac (internally partially spiculate) by antrum; corpus bursae oblong-ovate; ductus seminalis from junction of ductus bursae with bursa; accessory sac elongate-oblong from short and thin tube at ductus bursae junction; signum (
<figureCitation id="1306C8F1FFCCFF96FAED622D63E3F960" captionStart="FIGURE 15" captionStartId="15.[151,250,1858,1882]" captionTargetBox="[151,1436,453,1835]" captionTargetId="figure-128@15.[151,1436,453,1835]" captionTargetPageId="15" captionText="FIGURE 15. Thaumatographa aurosa taiwana Heppner &amp; Arita, subsp. nov., paratype female (Taiwan: Lien-Hua-Chih For. Sta., Nantou Co.), with details of ostium, sterigma, and antrum diverticulum (a), and signum, (b) (gen. slide JBH-4644 (MGCL)." pageId="14" pageNumber="285">Fig. 15b</figureCitation>
), a typical form for genus but in inverted V-shaped spine array.
</paragraph>
<paragraph id="8B82D474FFCCFF96FF4562956209F86D" blockId="14.[151,1437,361,1972]" pageId="14" pageNumber="285">
Male: unknown in
<collectingCountry id="F32A94E4FFCCFF96FE1F629562D4F905" box="[413,499,1730,1756]" name="Taiwan" pageId="14" pageNumber="285">Taiwan</collectingCountry>
. The male genitalia for the Japanese nominate subspecies are figured by
<bibRefCitation id="EFACA985FFCCFF96FAA8629563C1F8D9" author="Diakonoff, A." pageId="14" pageNumber="285" pagination="1 - 436" refId="ref10974" refString="Diakonoff, A. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Reisser, H. &amp; Gregor, F. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1-436, 175 pls." type="book chapter" year="1986">Diakonoff (1986)</bibRefCitation>
: uncus short and narrow, very evenly recurved from base to blunt apex; hamus stout and somewhat upwardly curved to acute apex, with small basal process; socius nearly subequal to hamus and relatively straight and setose; gnathos membranous; transtilla quadratic, thin; juxta a small concave plate, dorsally blunt; aedeagus moderate, thin, with short decumbent phallobase; vesica scobinate, with narrow cornutus tube; valva oblong, with costal margin somewhat convex with a slight invagination at middle, and termen very truncated; vinculum with saccus undeveloped.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF4564C060A2FF68" box="[199,901,151,177]" pageId="15" pageNumber="286" type="etymology">
<paragraph id="8B82D474FFCDFF97FF4564C060A2FF68" blockId="15.[151,1436,151,393]" box="[199,901,151,177]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF4564C0626FFF68" bold="true" box="[199,328,151,177]" pageId="15" pageNumber="286">Etymology</emphasis>
. The subspecies is named for the island of
<collectingCountry id="F32A94E4FFCDFF97FCA964C060A6FF68" box="[811,897,151,177]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF4564ED67CFFF0C" box="[199,1256,186,213]" pageId="15" pageNumber="286" type="biology_ecology">
<paragraph id="8B82D474FFCDFF97FF4564ED67CFFF0C" blockId="15.[151,1436,151,393]" box="[199,1256,186,213]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF4564ED6206FF0D" bold="true" box="[199,289,186,212]" pageId="15" pageNumber="286">Biology</emphasis>
. Unknown. Flight period (n = 3 records): mid-September to mid-October (in
<collectingRegion id="49F91A96FFCDFF97FB0764EC67C3FF0C" box="[1157,1252,187,213]" country="China" name="Taiwan)" pageId="15" pageNumber="286">Taiwan)</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF456489608AFF20" box="[199,941,222,249]" pageId="15" pageNumber="286" type="distribution">
<paragraph id="8B82D474FFCDFF97FF456489608AFF20" blockId="15.[151,1436,151,393]" box="[199,941,222,249]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF456489627EFF21" bold="true" box="[199,345,222,248]" pageId="15" pageNumber="286">Distribution</emphasis>
. The
<collectingCountry id="F32A94E4FFCDFF97FE1B648862CCFF20" box="[409,491,223,249]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
subspecies is known only from
<collectingCountry id="F32A94E4FFCDFF97FCD16488608EFF20" box="[851,937,223,249]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFCDFF97FF4565556760FE50" pageId="15" pageNumber="286" type="discussion">
<paragraph id="8B82D474FFCDFF97FF4565556760FE50" blockId="15.[151,1436,151,393]" pageId="15" pageNumber="286">
<emphasis id="B9490866FFCDFF97FF4565556262FEC5" bold="true" box="[199,325,258,284]" pageId="15" pageNumber="286">Discussion</emphasis>
. While some maculation distinctions are evident between the two subspecies, further study is needed once males are found of
<taxonomicName id="4C3DAFF7FFCDFF97FE2E65706152FE99" authorityName="Heppner &amp; Arita" authorityYear="2025" box="[428,629,295,320]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="286" phylum="Arthropoda" rank="subSpecies" species="aurosa" subSpecies="taiwana">
<emphasis id="B9490866FFCDFF97FE2E65706152FE99" box="[428,629,295,320]" italics="true" pageId="15" pageNumber="286">T. aurosa taiwana</emphasis>
</taxonomicName>
to compare the male genitalia of the two subspecies, or to verify if they should be considered distinct species. All available
<collectingCountry id="F32A94E4FFCDFF97FD56651C6001FEBC" box="[724,806,331,357]" name="Taiwan" pageId="15" pageNumber="286">Taiwan</collectingCountry>
specimens are females. The purpose of the small ventral flap (or pouch) external to the base of the ostial funnel near the antrum, is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5F8A24CC0DB04FB3FEC67DA49B550E90">Review of Thaumatographa tortricids in Taiwan (Lepidoptera: Tortricidae: Chlidanotinae: Hilarographini)</mods:title>
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<emphasis id="B9490866FFCFFF95FF15626062E2F98B" bold="true" box="[151,453,1591,1618]" italics="true" pageId="13" pageNumber="284">Thaumatographa aurosa</emphasis>
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: Miyanoura, Yakushima Id.
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@ -0,0 +1,221 @@
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Heppner &amp; Bae
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(
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,
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Type-locality:
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<emphasis id="B9490866FFD2FF88FF15653D6263FE5A" bold="true" box="[151,324,361,388]" pageId="16" pageNumber="287">Type material.</emphasis>
<materialsCitation id="3B55DE29FFD2FF88FECB653D62F2FE71" collectingDate="1980-07-22" collectingDateMax="1980-07-25" collectingDateMin="1980-07-22" collectionCode="USNM" collectorName="D. R. Davis" country="Taiwan" location="Kenting" pageId="16" pageNumber="287" specimenCode="USNM-78387" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung" typeStatus="holotype">
<emphasis id="B9490866FFD2FF88FECB653D6109FE5A" bold="true" box="[329,558,361,388]" pageId="16" pageNumber="287">
<typeStatus id="54866AD6FFD2FF88FECB653D629FFE5D" box="[329,440,362,388]" pageId="16" pageNumber="287" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F32A94E4FFD2FF88FE48653E6100FE5A" box="[458,551,361,387]" name="Taiwan" pageId="16" pageNumber="287">Taiwan</collectingCountry>
]
</emphasis>
<specimenCount id="9D3B1FFDFFD2FF88FDB1653D616FFE5D" box="[563,584,362,388]" count="1" pageId="16" pageNumber="287" type="male"></specimenCount>
<location id="8EE282AFFFD2FF88FDCC653D618CFE5D" LSID="urn:lsid:plazi:treatment:03946562FFD2FF88FF1564C0623BF86D:8EE282AFFFD2FF88FDCC653D618CFE5D" box="[590,683,362,388]" country="Taiwan" name="Kenting" pageId="16" pageNumber="287" stateProvince="Pingtung">Kenting</location>
,
<collectingRegion id="49F91A96FFD2FF88FD37653D603DFE5D" box="[693,794,362,388]" country="Taiwan" name="Pingtung" pageId="16" pageNumber="287">Pingtung</collectingRegion>
Co.,
<date id="FF83F2B4FFD2FF88FCD7653D672EFE5D" box="[853,1033,362,389]" pageId="16" pageNumber="287" value="1980-07-22" valueMax="1980-07-25" valueMin="1980-07-22">
<collectingDate id="EFC70B5CFFD2FF88FCD7653D672EFE5D" box="[853,1033,362,389]" pageId="16" pageNumber="287" value="1980-07-22" valueMax="1980-07-25" valueMin="1980-07-22">2225 Jul. 1980</collectingDate>
</date>
,
<collectorName id="26C8B1A2FFD2FF88FB91653D67B3FE5D" box="[1043,1172,362,388]" pageId="16" pageNumber="287">D. R. Davis</collectorName>
(adult photo 5562; gen. JBH-2194) (
<specimenCode id="DB9B7C0FFFD2FF88FEA065D962EEFE71" box="[290,457,398,424]" collectionCode="USNM-" pageId="16" pageNumber="287">USNM-78387</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFD2FF88FF4565E66101FD85" pageId="16" pageNumber="287" type="diagnosis">
<paragraph id="8B82D474FFD2FF88FF4565E66101FD85" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4565E6621DFE12" bold="true" box="[199,314,433,459]" pageId="16" pageNumber="287">Diagnosis</emphasis>
. This minute species is among the smallest
<taxonomicName id="4C3DAFF7FFD2FF88FC9C65E560C7FE15" authorityName="Walsignham" authorityYear="1897" box="[798,992,434,460]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="B9490866FFD2FF88FC9C65E560C7FE15" box="[798,992,434,460]" italics="true" pageId="16" pageNumber="287">Thaumatographa</emphasis>
</taxonomicName>
known. Maculation is very dark brown, with white marks on the costa and white striae from the dorsal margin strongly curved to middle of wing directed to apex, and the termen is orange with two widely-spaced black spots; the hindwing is dark brown with only faint tawny orange streaks in cell and on anal sector to wingbase. The male genitalia are distinctive for the very stout and thick uncus and only stubby socius.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFD2FF88FF45663262B6F94D" pageId="16" pageNumber="287" type="description">
<paragraph id="8B82D474FFD2FF88FF45663261DEFD59" blockId="16.[151,1437,361,1972]" box="[199,761,613,641]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4566326276FDA6" bold="true" box="[199,337,613,639]" pageId="16" pageNumber="287">Description</emphasis>
. Wing expanse: male 7.5 mm (n = 1).
</paragraph>
<paragraph id="8B82D474FFD2FF88FF4566DD613BFCC9" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
Male (
<figureCitation id="1306C8F1FFD2FF88FE9366DD6272FD7D" box="[273,341,650,676]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="16" pageNumber="287">Fig. 6</figureCitation>
).
<emphasis id="B9490866FFD2FF88FEE966DD628FFD7D" box="[363,424,650,676]" italics="true" pageId="16" pageNumber="287">Head</emphasis>
(
<figureCitation id="1306C8F1FFD2FF88FE3566DD6123FD7D" box="[439,516,650,676]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="16" pageNumber="287">Fig 6a</figureCitation>
): brown to dark brown, with orange-yellow scales on neck and on frons; antennal scape dull tan-yellow, with antenna brown mixed with tan-yellow scales; eye margins pale yellow; labial palpus short, yellow-white, with more yellow on middle segment and somewhat on apical segment, and distally brown on apical segment; head venter white.
</paragraph>
<paragraph id="8B82D474FFD2FF88FF45674D6160FCA5" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF45674D623FFCED" box="[199,280,794,820]" italics="true" pageId="16" pageNumber="287">Thorax</emphasis>
(
<figureCitation id="1306C8F1FFD2FF88FEAA674D6248FCED" box="[296,367,794,820]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="16" pageNumber="287">Fig. 6</figureCitation>
): dark brown, with dull yellow lateral line and on tegula; venter white; legs white, with black-brown on dorsum of foretibia and ends of mid- and hindtibia, and tarsi yellow-white with black-brown on first and distal tarsal segments; leg spurs white.
</paragraph>
<paragraph id="8B82D474FFD2FF88FF4567D062DCFB19" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4567D06214FC79" box="[199,307,903,928]" italics="true" pageId="16" pageNumber="287">Forewing</emphasis>
(
<figureCitation id="1306C8F1FFD2FF88FEC767D162AAFC79" box="[325,397,902,928]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="16" pageNumber="287">Fig. 6</figureCitation>
): dark brown to black-brown, with hilarographine striae and marks, orange near wingbase and between each of the costal angled white wedges, all directed toward termen, and with orange striae, partially interrupted, near wingbase and past middle of wing from dorsal margin; the prominent white curved striae from the dorsal margin at middle of wing to cell are very approximate to each other, almost merging at their distal ends; the end of the cell has a black-brown spot and also a tiny pale yellow-white dot; the termen and apex are distally orange, with a small white bar on termen below apex, and two black spots in the orange termen patch, with three subterminal curved pale yellow-white striae; fringe dark brown and white distally; venter with dorsal pattern repeated as diffuse alternating dull tan-white and light brown markings, with dull white in cell and yellow near apex, and termen with black spots of dorsum repeated.
</paragraph>
<paragraph id="8B82D474FFD2FF88FF45609D675AFAD1" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF45609D6212FB3D" box="[199,309,1226,1252]" italics="true" pageId="16" pageNumber="287">Hindwing</emphasis>
(
<figureCitation id="1306C8F1FFD2FF88FEC7609D62ADFB3D" box="[325,394,1226,1252]" captionStart="FIGURES 16" captionStartId="4.[151,264,1560,1584]" captionTargetBox="[151,1436,555,1536]" captionTargetId="figure-133@4.[151,1436,555,1536]" captionTargetPageId="4" captionText="FIGURES 16. Adults of Taiwan Thaumatographa species, with lateral views of heads (a) for each moth: 1) T. mesostigmatias Diakonoff &amp; Arita, male (Chiayi Co.: Fennchihwu, photo 5569) (WS = 13.0mm); 2) same data, female (photo14173) (WS = 14.8 mm) (MGCL). 3) T. fennchihwu Heppner &amp; Arita, sp. nov., male holotype (Chiayi Co.: Fennchihwu, photo 5570) (WS = 15.5 mm); 4) same data, female paratype (photo 5571) (WS = 19.0 mm) (MGCL). 5) T. aurosa taiwana Heppner &amp;Arita, subsp. nov., female paratype (Ilan Co.: Nanao, photo 16014) (WS = 13.6 mm) (MGCL). 6) T. kentingana Heppner &amp; Bae, sp. nov., male holotype (Pingtung Co.: Kenting, photo 5562) (WS = 7.5 mm) (USNM). Note: the white mark in middle of wing Fig. 2 is partially descaled, thus otherwise as small as in the male Fig. 1. Also, note the bluish iridescence of the labial palpi in Fig. 1a and 2a." pageId="16" pageNumber="287">Fig. 6</figureCitation>
): dark brown, with faint diffuse tawny orange scales in cell area and along anal sector; venter gray-brown, with costa dull gray-white between brown marginal marks and along radius.
</paragraph>
<paragraph id="8B82D474FFD2FF88FF45614567BCFA89" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4561456216FAF5" box="[199,305,1298,1324]" italics="true" pageId="16" pageNumber="287">Abdomen</emphasis>
: gray-brown; venter white with gray-brown basally on each sternite and caudally as genital tufts; pregenital sternite unmodified with simple curved caudal margin and lateral corematal sacs.
</paragraph>
<paragraph id="8B82D474FFD2FF88FF45610D61D9F9A9" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF45610D624CFAAD" box="[199,363,1370,1396]" italics="true" pageId="16" pageNumber="287">Male genitalia</emphasis>
(
<figureCitation id="1306C8F1FFD2FF88FEF8610D62E9FAAD" box="[378,462,1370,1396]" captionStart="FIGURE 12" captionStartId="10.[151,250,1507,1531]" captionTargetBox="[151,1436,417,1484]" captionTargetId="figure-135@10.[151,1436,417,1484]" captionTargetPageId="10" captionText="FIGURE 12. Thaumatographa kentingana Heppner &amp; Bae, sp. nov., holotype male (Taiwan: Kenting Park, Pingtung Co.), with aedeagus greatly enlarged (a) and with ductus ejaculatorius remnant (right) and part of juxta attached below (left); with enlargement of uncus to juxta (b), and structures of the pregenital abdomen and corematal sockets (c) (corematal setae removed) (gen. slide JBH-2194) (MGCL)." pageId="16" pageNumber="287">Fig. 12</figureCitation>
): tegumen globular and very sclerotized; uncus very stout, tubular and straight, slightly tapering distally to acute apex; hamus subequal in length to uncus and stout, slightly flattened and slightly curved to acute apex, but abruptly bent dorsally from strongly sclerotized basal arm and basal arm ventrally with an acute thin spine (longer than socius); socius very small and stubby setose process; gnathos undeveloped, membranous; transtilla a thin band, slightly curved; juxta and small oval concave plate, dorsally truncated; aedeagus short and thick, slightly curved and tapering to acute apex, with caudal phallobase small and bulbous, decumbent but ventrally even with distal curving; valva oblong and setose, with truncated termen angled to saccular apex; vinculum triangular; saccus undeveloped, merged as acute end of vinculum.
</paragraph>
<paragraph id="8B82D474FFD2FF88FF45622D62B6F94D" blockId="16.[151,1437,361,1972]" box="[199,401,1658,1684]" pageId="16" pageNumber="287">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFD2FF88FF4562C96263F905" pageId="16" pageNumber="287" type="etymology">
<paragraph id="8B82D474FFD2FF88FF4562C96263F905" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4562C9626FF961" bold="true" box="[199,328,1694,1720]" pageId="16" pageNumber="287">Etymology</emphasis>
. The species is named for Kenting Park (now Kenting National Park),
<collectingRegion id="49F91A96FFD2FF88FBC062C96780F961" box="[1090,1191,1694,1720]" country="Taiwan" name="Pingtung" pageId="16" pageNumber="287">Pingtung</collectingRegion>
Co., the southernmost area of
<collectingCountry id="F32A94E4FFD2FF88FF6962956266F905" box="[235,321,1730,1756]" name="Taiwan" pageId="16" pageNumber="287">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFD2FF88FF4562B26075F8D9" box="[199,850,1765,1792]" pageId="16" pageNumber="287" type="biology_ecology">
<paragraph id="8B82D474FFD2FF88FF4562B26075F8D9" blockId="16.[151,1437,361,1972]" box="[199,850,1765,1792]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4562B26206F926" bold="true" box="[199,289,1765,1791]" pageId="16" pageNumber="287">Biology</emphasis>
. Unknown. Flight period (n = 1 record): late July.
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFD2FF88FF45635E6232F891" pageId="16" pageNumber="287" type="distribution">
<paragraph id="8B82D474FFD2FF88FF45635E6232F891" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF45635E627EF8FA" bold="true" box="[199,345,1801,1827]" pageId="16" pageNumber="287">Distribution</emphasis>
. Known only from the southernmost area of
<collectingCountry id="F32A94E4FFD2FF88FCFD635D60F3F8FD" box="[895,980,1802,1828]" name="Taiwan" pageId="16" pageNumber="287">Taiwan</collectingCountry>
, Kenting Park,
<collectingRegion id="49F91A96FFD2FF88FB10635D67D0F8FD" box="[1170,1271,1802,1828]" country="Taiwan" name="Pingtung" pageId="16" pageNumber="287">Pingtung</collectingRegion>
Co. (Oluanpi Peninsula).
</paragraph>
</subSubSection>
<subSubSection id="C32787FFFFD2FF88FF456306623BF86D" pageId="16" pageNumber="287" type="discussion">
<paragraph id="8B82D474FFD2FF88FF456306623BF86D" blockId="16.[151,1437,361,1972]" pageId="16" pageNumber="287">
<emphasis id="B9490866FFD2FF88FF4563066262F8B2" bold="true" box="[199,325,1873,1899]" pageId="16" pageNumber="287">Discussion</emphasis>
. This species is among the smallest of known
<taxonomicName id="4C3DAFF7FFD2FF88FCC163056744F8B5" box="[835,1123,1874,1900]" class="Insecta" family="Tortricidae" genus="Thaumatographa" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="287" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9490866FFD2FF88FCC163056722F8B5" box="[835,1029,1874,1900]" italics="true" pageId="16" pageNumber="287">Thaumatographa</emphasis>
species.
</taxonomicName>
The Kenting area of
<collectingCountry id="F32A94E4FFD2FF88FAC8630566BBF8B5" box="[1354,1436,1874,1900]" name="Taiwan" pageId="16" pageNumber="287">Taiwan</collectingCountry>
is floristically and faunally very much related to the
<collectingCountry id="F32A94E4FFD2FF88FD6463216043F849" box="[742,868,1910,1936]" name="Philippines" pageId="16" pageNumber="287">Philippines</collectingCountry>
(
<bibRefCitation id="EFACA985FFD2FF88FCF1632160F7F849" author="Li, H. - L." box="[883,976,1910,1936]" pageId="16" pageNumber="287" refId="ref11980" refString="Li, H. - L. (1963) Woody Flora of Taiwan. Livingston Publishing Company, Narberth, Pennsylvania, 974 pp." type="book" year="1963">Li 1963</bibRefCitation>
), thus the new species may possibly also occur there.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,176 @@
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<mods:title id="4AF1217881E4AC2EA0BADD36258D1C1F">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:namePart id="6E6527AF8A803B651472B5804EBC3A87">Sankaran, Pradeep M.</mods:namePart>
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<mods:affiliation id="8783C6231EC3C12A03314DABFA425394">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
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<mods:namePart id="F048D2307AD785B05DDAC2F51E9CC71F">Haddad, Charles R.</mods:namePart>
<mods:nameIdentifier id="E191A0945837149A310AD1A498897FA3" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="A4245BC57599E5762A7CDE7D303B7636">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="D9514036F5CEDC021AD6A1635254DC51" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name id="6D6D6755E3D44579A2A08CB3FBC8852F" type="personal">
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<mods:namePart id="1A469FAF9E82DC5BA179FB59EDB7EEF5">Tripathi, Rishikesh</mods:namePart>
<mods:nameIdentifier id="D4B54B7196E4DBCCDD7C900C4AFAE2CD" type="ORCID">0000-0002-9192-4609</mods:nameIdentifier>
<mods:affiliation id="57A449E8C8866A2E953056919FC120C2">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Thrissur, Kerala 680 125, India.</mods:affiliation>
<mods:nameIdentifier id="6D10C1A03F16C0962CDE8F1CC62CB3E8" type="email">rishikesh.tripathi14@gmail.com</mods:nameIdentifier>
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<mods:title id="3738C392A336B4C8858F2B1A6900A7DC">Zootaxa</mods:title>
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<mods:date id="CD80AE8AD7D19C19D12251AB539B3488">2025</mods:date>
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<heading id="D0966E185162F75B95E1FB02FD07FB6A" bold="true" box="[151,753,1219,1246]" fontSize="11" level="1" pageId="27" pageNumber="66" reason="1">
<emphasis id="B91505665162F75B95E1FB02FD07FB6A" bold="true" box="[151,753,1219,1246]" pageId="27" pageNumber="66">
<taxonomicName id="4C61A2F75162F75B95E1FB02FD92FB6A" authority="(Kritscher, 1996)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Kritscher" baseAuthorityYear="1996" box="[151,612,1219,1246]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="66" phylum="Arthropoda" rank="species" species="loricata" status="comb. nov.">
<emphasis id="B91505665162F75B95E1FB02FE7CFB6A" bold="true" box="[151,394,1219,1246]" italics="true" pageId="27" pageNumber="66">Trichothyse loricata</emphasis>
(
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)
</taxonomicName>
<taxonomicNameLabel id="A226B81D5162F75B971DFB05FD07FB6A" box="[619,753,1220,1246]" pageId="27" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5162F75B95E1FAC9FF08FAEA" pageId="27" pageNumber="66" type="reference_group">
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<emphasis id="B91505665162F75B95E1FAC9FE83FA94" box="[151,373,1288,1312]" italics="true" pageId="27" pageNumber="66">Poecilochroa loricata</emphasis>
<bibRefCitation id="EFF0A4855162F75B940DFAC9FDBBFA95" author="Kritscher, E." box="[379,589,1288,1313]" pageId="27" pageNumber="66" pagination="117 - 156" refId="ref19172" refString="Kritscher, E. (1996) Ein Beitrag zur Kenntnis der Spinnen-Fauna der Maltesischen Inseln (Chelicerata: Araneae). Annalen des Naturhistorischen Museums in Wien, Serie B, 98, 117-156." type="journal article" year="1996">Kritscher, 1996: 143</bibRefCitation>
</taxonomicName>
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, fig. 7 (
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and
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:
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<collectingCountry id="F37699E45162F75B96DBFAC9FBECFA94" box="[941,1050,1288,1312]" name="France" pageId="27" pageNumber="66">FRANCE</collectingCountry>
:
</emphasis>
Malta: east of Marsaxlokk [
<geoCoordinate id="EE55BFB35162F75B904AFAC9FA61FA94" box="[1340,1431,1288,1312]" degrees="35" direction="north" minutes="50" orientation="latitude" pageId="27" pageNumber="66" precision="925" value="35.833332">35°50N</geoCoordinate>
,
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,
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a.s.l.],
<date id="FFDFFFB45162F75B94EBFAE6FDE5FA8B" box="[413,531,1319,1343]" pageId="27" pageNumber="66" value="1990-06-08">08.VI.1990</date>
, leg. Erich Kritscher &amp; Karl Bilek, NHMW—not examined);
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</treatmentCitation>
, figs 1ae, 2ac.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5162F75B95E1FA4DFE2DF972" pageId="27" pageNumber="66" type="description">
<paragraph id="8BDED9745162F75B95E1FA4DFC8DFA12" blockId="27.[151,1437,1420,1770]" box="[151,891,1420,1446]" pageId="27" pageNumber="66">
<emphasis id="B91505665162F75B95E1FA4DFEDFFA12" bold="true" box="[151,297,1420,1446]" pageId="27" pageNumber="66">Description.</emphasis>
<emphasis id="B91505665162F75B9446FA4DFE9CFA12" box="[304,362,1420,1446]" italics="true" pageId="27" pageNumber="66">Male</emphasis>
. For description of the male, see
<bibRefCitation id="EFF0A4855162F75B97A9FA4DFC81FA12" author="Bauer, T." box="[735,887,1420,1446]" pageId="27" pageNumber="66" pagination="97 - 100" refId="ref18320" refString="Bauer, T. (2022) Remarks on the type material of Poecilochroa loricata Kritscher, 1996 (Araneae: Gnaphosidae). Zootaxa, 5169 (1), 97-100. https://doi.org/10.11646/zootaxa.5169.1.10" type="journal article" year="2022">Bauer (2022)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED9745162F75B95B1FA71FE69FA7E" blockId="27.[151,1437,1420,1770]" box="[199,415,1456,1482]" pageId="27" pageNumber="66">
<emphasis id="B91505665162F75B95B1FA71FEEDFA7E" box="[199,283,1456,1482]" italics="true" pageId="27" pageNumber="66">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BDED9745162F75B95B1FA15FE2DF972" blockId="27.[151,1437,1420,1770]" pageId="27" pageNumber="66">
<emphasis id="B91505665162F75B95B1FA15FE13FA5A" bold="true" box="[199,485,1492,1518]" pageId="27" pageNumber="66">Justification of transfer.</emphasis>
<typeStatus id="54DA67D65162F75B949DFA15FDA5FA5A" box="[491,595,1492,1518]" pageId="27" pageNumber="66" type="holotype">Holotype</typeStatus>
images are provided in
<bibRefCitation id="EFF0A4855162F75B9616FA15FC06FA5A" author="Bauer, T." box="[864,1008,1492,1518]" pageId="27" pageNumber="66" pagination="97 - 100" refId="ref18320" refString="Bauer, T. (2022) Remarks on the type material of Poecilochroa loricata Kritscher, 1996 (Araneae: Gnaphosidae). Zootaxa, 5169 (1), 97-100. https://doi.org/10.11646/zootaxa.5169.1.10" type="journal article" year="2022">Bauer (2022</bibRefCitation>
: fig. 1) and were used for comparison. These figures are consistent with the palps of male
<taxonomicName id="4C61A2F75162F75B97AFFA39FC4EF9A6" box="[729,952,1528,1554]" pageId="27" pageNumber="66">
<emphasis id="B91505665162F75B97AFFA39FCAAF9A6" box="[729,860,1528,1554]" italics="true" pageId="27" pageNumber="66">Trichothyse</emphasis>
species
</taxonomicName>
: distomedially originating broad conductor with prolateral fold, apically originating claw-like embolus (but lacks a broad base as per the illustrations referred to here), medially contiguous sperm duct with single loop, and robust RTA with apical bifurcation (cf.
<bibRefCitation id="EFF0A4855162F75B9065F981FA6EF9EE" author="Bauer, T." box="[1299,1432,1600,1626]" pageId="27" pageNumber="66" pagination="97 - 100" refId="ref18320" refString="Bauer, T. (2022) Remarks on the type material of Poecilochroa loricata Kritscher, 1996 (Araneae: Gnaphosidae). Zootaxa, 5169 (1), 97-100. https://doi.org/10.11646/zootaxa.5169.1.10" type="journal article" year="2022">Bauer 2022</bibRefCitation>
: figs 1ce, 2ac and
<bibRefCitation id="EFF0A4855162F75B9404F9A5FE0BF9CA" author="Simon, E." box="[370,509,1636,1662]" pageId="27" pageNumber="66" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon 1914</bibRefCitation>
: fig. 398,
<bibRefCitation id="EFF0A4855162F75B971CF9A5FD15F9CA" author="Fage, L." box="[618,739,1636,1662]" pageId="27" pageNumber="66" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage 1929</bibRefCitation>
: fig. 1,
<bibRefCitation id="EFF0A4855162F75B9642F9A5FC13F9CA" author="Di Franco, F." box="[820,997,1636,1662]" pageId="27" pageNumber="66" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco 2001</bibRefCitation>
: figs 34,
<bibRefCitation id="EFF0A4855162F75B9120F9A5FB24F9CA" author="Levy, G." box="[1110,1234,1636,1662]" pageId="27" pageNumber="66" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: fig. 10, 2009: fig. 45,
<bibRefCitation id="EFF0A4855162F75B95B4F949FE01F916" author="Fomichev, A. A. &amp; Marusik, Y. M." box="[194,503,1672,1698]" pageId="27" pageNumber="66" pagination="443 - 457" refId="ref18957" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2021) A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa, 4966 (4), 443-457. https://doi.org/10.11646/zootaxa.4966.4.3" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 23 and
<figureCitation id="135AC5F15162F75B97F4F949FCA3F916" box="[642,853,1672,1698]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="27" pageNumber="66">Figs 109110, 112</figureCitation>
,
<figureCitation id="135AC5F15162F75B9616F949FC36F916" box="[864,960,1672,1698]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="27" pageNumber="66">116117</figureCitation>
herein). We therefore propose the transfer of
<taxonomicName id="4C61A2F75162F75B95C0F96CFED8F972" authorityName="Kritscher" authorityYear="1996" box="[182,302,1708,1734]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="66" phylum="Arthropoda" rank="species" species="loricata">
<emphasis id="B91505665162F75B95C0F96CFF31F972" box="[182,199,1709,1734]" italics="true" pageId="27" pageNumber="66">P</emphasis>
.
<emphasis id="B91505665162F75B95A3F96DFED8F972" box="[213,302,1708,1734]" italics="true" pageId="27" pageNumber="66">loricata</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75162F75B9427F96DFE22F972" authorityName="Tucker" authorityYear="1923" box="[337,468,1708,1734]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665162F75B9427F96DFE22F972" box="[337,468,1708,1734]" italics="true" pageId="27" pageNumber="66">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5162F75B95B1F911FBE3F95E" box="[199,1045,1744,1770]" pageId="27" pageNumber="66" type="distribution">
<paragraph id="8BDED9745162F75B95B1F911FBE3F95E" blockId="27.[151,1437,1420,1770]" box="[199,1045,1744,1770]" pageId="27" pageNumber="66">
<emphasis id="B91505665162F75B95B1F911FE97F95E" bold="true" box="[199,353,1744,1770]" pageId="27" pageNumber="66">Distribution.</emphasis>
Known only from
<collectingCountry id="F37699E45162F75B9741F911FD8EF95E" box="[567,632,1744,1770]" name="Malta" pageId="27" pageNumber="66">Malta</collectingCountry>
in Mediterranean Europe (
<figureCitation id="135AC5F15162F75B96D1F911FBFEF95E" box="[935,1032,1744,1770]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="27" pageNumber="66">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,180 @@
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<mods:title id="E48405886A2FB645C2219870929C4C77">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:namePart id="18C39A0DE46578E031EE5E0F2C6F2970">Sankaran, Pradeep M.</mods:namePart>
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<mods:affiliation id="8F0016DB30A77436CFC4027DB559CBD4">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
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<mods:namePart id="A40DB4A7E46A58A790C4C6CF1EADC0A5">Haddad, Charles R.</mods:namePart>
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<mods:affiliation id="454A1828B94CCF56872F8B00BF6C00B0">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="D4F3FC79D6B85EBADFD709B4965501FA" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:namePart id="0304DD826D1A9BB06AFB5F5468E92CAA">Tripathi, Rishikesh</mods:namePart>
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<mods:affiliation id="BC37E40131095BDB51E4339A0BFFEC15">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Thrissur, Kerala 680 125, India.</mods:affiliation>
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<heading id="D0966E185162F75B95E1F8F2FD23F8FA" bold="true" box="[151,725,1843,1870]" fontSize="11" level="1" pageId="27" pageNumber="66" reason="1">
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<taxonomicName id="4C61A2F75162F75B95E1F8F2FDB1F8FA" authority="(Simon, 1914)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Simon" baseAuthorityYear="1914" box="[151,583,1843,1870]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="66" phylum="Arthropoda" rank="species" species="perversa" status="comb. nov.">
<emphasis id="B91505665162F75B95E1F8F2FE62F8FA" bold="true" box="[151,404,1843,1870]" italics="true" pageId="27" pageNumber="66">Trichothyse perversa</emphasis>
(
<bibRefCitation id="EFF0A4855162F75B94D5F8F5FDC9F8FA" author="Simon, E." box="[419,575,1844,1870]" pageId="27" pageNumber="66" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon, 1914</bibRefCitation>
)
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<taxonomicNameLabel id="A226B81D5162F75B9739F8F5FD23F8FA" box="[591,725,1844,1870]" pageId="27" pageNumber="66" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<treatmentCitation id="0AC0FF655162F75B95E1F8B9FDB7F825" author="Simon, E." box="[151,577,1912,1937]" page="184" pageId="27" pageNumber="66" year="1914">
<taxonomicName id="4C61A2F75162F75B95E1F8B9FDB7F825" ID-CoL="4KTC9" authority="Simon, 1914: 184" authorityName="Simon" authorityPageNumber="184" authorityYear="1914" box="[151,577,1912,1937]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="66" phylum="Arthropoda" rank="species" species="perversa">
<emphasis id="B91505665162F75B95E1F8B9FE76F824" box="[151,384,1912,1936]" italics="true" pageId="27" pageNumber="66">Poecilochroa perversa</emphasis>
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<bibRefCitation id="EFF0A4855162F75B94F1F8B9FDB7F825" author="Simon, E." box="[391,577,1912,1937]" pageId="27" pageNumber="66" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon, 1914: 184</bibRefCitation>
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, 221, figs 395396 (
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:
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:
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date and collector unknown, repository unknown, probably MNHN—not examined).
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<emphasis id="B91505665162F75B95E1F81CFEDFF843" bold="true" box="[151,297,2013,2039]" pageId="27" pageNumber="66">Description.</emphasis>
<emphasis id="B91505665162F75B9446F81CFE9CF843" box="[304,362,2013,2039]" italics="true" pageId="27" pageNumber="66">Male</emphasis>
. For description of the male, see
<bibRefCitation id="EFF0A4855162F75B97A9F81CFC8BF843" author="Simon, E." box="[735,893,2013,2039]" pageId="27" pageNumber="66" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon (1914)</bibRefCitation>
.
</paragraph>
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<emphasis id="B91505665165F75C95B1FF56FEEDFF05" box="[199,283,151,177]" italics="true" pageId="28" pageNumber="67">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BDED9745165F75C95B1FF7BFEC8FE19" blockId="28.[151,1437,151,465]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C95B1FF7BFE10FF60" bold="true" box="[199,486,186,212]" pageId="28" pageNumber="67">Justification of transfer.</emphasis>
The illustration of the palp provided in
<bibRefCitation id="EFF0A4855165F75C96D3FF7AFBCDFF61" author="Simon, E." box="[933,1083,187,213]" pageId="28" pageNumber="67" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon (1914</bibRefCitation>
: fig. 395) is diagnostic and was used for comparison. Even though we did not examine the
<typeStatus id="54DA67D65165F75C9647FF1EFC97FF4D" box="[817,865,223,249]" pageId="28" pageNumber="67">type</typeStatus>
of
<taxonomicName id="4C61A2F75165F75C96FEFF1EFBFFFF4C" authorityName="Simon" authorityYear="1914" box="[904,1033,223,249]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="perversa">
<emphasis id="B91505665165F75C96FEFF1EFC6FFF4C" box="[904,921,223,248]" italics="true" pageId="28" pageNumber="67">P</emphasis>
.
<emphasis id="B91505665165F75C96D1FF1EFBFFFF4C" box="[935,1033,223,248]" italics="true" pageId="28" pageNumber="67">perversa</emphasis>
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, the original illustration of the RTA of the male palp of this species shows that it has a robust RTA with an apical bifurcation, as seen in all known male
<taxonomicName id="4C61A2F75165F75C95E1FEE6FE8EFEF5" box="[151,376,295,321]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C95E1FEE6FEECFEF5" box="[151,282,295,321]" italics="true" pageId="28" pageNumber="67">Trichothyse</emphasis>
species
</taxonomicName>
, in contrast to
<emphasis id="B91505665165F75C9751FEE6FCA1FEF4" box="[551,855,295,321]" italics="true" pageId="28" pageNumber="67">
<taxonomicName id="4C61A2F75165F75C9751FEE6FD4BFEF5" authorityName="Westring" authorityYear="1874" box="[551,701,295,321]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus" sensu="stricto">Poecilochroa</taxonomicName>
<taxonomicNameLabel id="A226B81D5165F75C97B1FEE6FCA1FEF4" box="[711,855,295,320]" pageId="28" pageNumber="67" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C61A2F75165F75C96E5FEE6FBCCFEF5" authorityName="Wunderlich" authorityYear="2011" box="[915,1082,295,321]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665165F75C96E5FEE6FBCCFEF5" box="[915,1082,295,321]" italics="true" pageId="28" pageNumber="67">Macarophaeus</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFF0A4855165F75C9105FEE6FAF4FEF5" author="Simon, E." box="[1139,1282,295,321]" pageId="28" pageNumber="67" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon 1914</bibRefCitation>
: fig. 395 and
<bibRefCitation id="EFF0A4855165F75C95E1FE8AFED2FED1" author="Simon, E." box="[151,292,331,357]" pageId="28" pageNumber="67" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon 1914</bibRefCitation>
: fig. 398,
<bibRefCitation id="EFF0A4855165F75C94E3FE8AFDF9FED1" author="Fage, L." box="[405,527,331,357]" pageId="28" pageNumber="67" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage 1929</bibRefCitation>
: fig. 1,
<bibRefCitation id="EFF0A4855165F75C9713FE8AFCECFED1" author="Di Franco, F." box="[613,794,331,357]" pageId="28" pageNumber="67" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco 2001</bibRefCitation>
: figs 34,
<bibRefCitation id="EFF0A4855165F75C96F9FE8AFBFAFED1" author="Levy, G." box="[911,1036,331,357]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: fig. 10, 2009: fig. 45,
<bibRefCitation id="EFF0A4855165F75C9079FE8AFECFFE3D" author="Fomichev, A. A. &amp; Marusik, Y. M." pageId="28" pageNumber="67" pagination="443 - 457" refId="ref18957" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2021) A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa, 4966 (4), 443-457. https://doi.org/10.11646/zootaxa.4966.4.3" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 23, and
<figureCitation id="135AC5F15165F75C94B9FEAEFD52FE3D" box="[463,676,367,393]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="28" pageNumber="67">Figs 109110, 112</figureCitation>
,
<figureCitation id="135AC5F15165F75C97C6FEAEFCE6FE3D" box="[688,784,367,393]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="28" pageNumber="67">116117</figureCitation>
herein). We therefore propose the transfer of
<taxonomicName id="4C61A2F75165F75C906CFEAEFA6AFE3C" authorityName="Simon" authorityYear="1914" box="[1306,1436,367,393]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="perversa">
<emphasis id="B91505665165F75C906CFEAEFADDFE3C" box="[1306,1323,367,392]" italics="true" pageId="28" pageNumber="67">P</emphasis>
.
<emphasis id="B91505665165F75C904CFEAEFA6AFE3C" box="[1338,1436,367,392]" italics="true" pageId="28" pageNumber="67">perversa</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75165F75C95C2FE52FEC1FE19" authorityName="Tucker" authorityYear="1923" box="[180,311,403,429]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665165F75C95C2FE52FEC1FE19" box="[180,311,403,429]" italics="true" pageId="28" pageNumber="67">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<emphasis id="B91505665165F75C95B1FE77FE97FE64" bold="true" box="[199,353,438,464]" pageId="28" pageNumber="67">Distribution.</emphasis>
Known only from
<collectingCountry id="F37699E45165F75C9741FE76FD75FE65" box="[567,643,439,465]" name="France" pageId="28" pageNumber="67">France</collectingCountry>
in Mediterranean Europe (
<figureCitation id="135AC5F15165F75C96C7FE76FBE5FE65" box="[945,1043,439,465]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="28" pageNumber="67">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,190 @@
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<mods:title id="2F2CD55A02405FF68F4251C39156DEBB">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:namePart id="997F7CA5F8E8B37907EF03E99CA182B7">Haddad, Charles R.</mods:namePart>
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<heading id="D0966E185164F75D95E1FD87FCE1FDD5" bold="true" box="[151,791,582,609]" fontSize="11" level="1" pageId="29" pageNumber="68" reason="1">
<emphasis id="B91505665164F75D95E1FD87FCE1FDD5" bold="true" box="[151,791,582,609]" pageId="29" pageNumber="68">
<taxonomicName id="4C61A2F75164F75D95E1FD87FD7FFDD4" authority="(Simon, 1878)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Simon" baseAuthorityYear="1878" box="[151,649,582,609]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="subSpecies" species="senilis" status="comb. nov." subSpecies="auspex">
<emphasis id="B91505665164F75D95E1FD87FE20FDD5" bold="true" box="[151,470,582,609]" italics="true" pageId="29" pageNumber="68">Trichothyse senilis auspex</emphasis>
(
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)
</taxonomicName>
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</emphasis>
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<subSubSection id="C37B8AFF5164F75D95E1FD4DFCD5FD55" pageId="29" pageNumber="68" type="reference_group">
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<treatmentCitation id="0AC0FF655164F75D95E1FD4DFE1BFD17" author="Simon, E." box="[151,493,651,675]" page="154" pageId="29" pageNumber="68" year="1878">
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<emphasis id="B91505665164F75D95E1FD4DFEC3FD17" box="[151,309,652,675]" italics="true" pageId="29" pageNumber="68">Drassus auspex</emphasis>
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<bibRefCitation id="EFF0A4855164F75D944CFD4AFE1BFD17" author="Simon, E." box="[314,493,651,675]" pageId="29" pageNumber="68" refId="ref20068" refString="Simon, E. (1878) Les arachnides de France. Tome quatrieme, contenant la famille des Drassidae. Roret, Paris, 334 pp., pls. XIV - XVI." type="book" year="1878">Simon, 1878: 154</bibRefCitation>
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(
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:
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:
</emphasis>
<location id="8EBE8FAF5164F75D9781FD4AFC31FD17" LSID="urn:lsid:plazi:treatment:03C868625164F75D95E1FD87FBD6FB91:8EBE8FAF5164F75D9781FD4AFC31FD17" box="[759,967,651,675]" country="France" latitude="42.533333" longLatPrecision="1149" longitude="2.4" municipality="Vernet" name="Pyrenees-Orientales" pageId="29" pageNumber="68">Pyrénées-Orientales</location>
:
<collectingMunicipality id="6BBA430E5164F75D96B8FD4AFBE4FD17" box="[974,1042,651,675]" pageId="29" pageNumber="68">Vernet</collectingMunicipality>
/Vernet-Les-Bains [
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,
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], date unknown, leg.
<collectorName id="2694BCA25164F75D94F8FD6BFE29FD75" box="[398,479,682,705]" pageId="29" pageNumber="68">M. Nou</collectorName>
, repository unknown, probably MNHN—not examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745164F75D95E1FD09FCD5FD55" blockId="29.[151,1436,651,737]" box="[151,803,712,737]" pageId="29" pageNumber="68">
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<emphasis id="B91505665164F75D95E1FD09FE24FD54" box="[151,466,712,736]" italics="true" pageId="29" pageNumber="68">Poecilochroa dimidiata auspex</emphasis>
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, 221, fig. 398.
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<emphasis id="B91505665164F75D95E1FCCFFEDFFC9C" bold="true" box="[151,297,782,808]" pageId="29" pageNumber="68">Description.</emphasis>
<emphasis id="B91505665164F75D9446FCCEFE9CFC9D" box="[304,362,783,809]" italics="true" pageId="29" pageNumber="68">Male</emphasis>
and
<emphasis id="B91505665164F75D94D6FCCEFE1DFC9D" box="[416,491,783,809]" italics="true" pageId="29" pageNumber="68">female</emphasis>
. For redescription of the male and female, see
<bibRefCitation id="EFF0A4855164F75D968FFCCEFB60FC9D" author="Simon, E." box="[1017,1174,783,809]" pageId="29" pageNumber="68" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon (1914)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED9745164F75D95B1FCF3FCF2FBB5" blockId="29.[151,1437,782,1061]" pageId="29" pageNumber="68">
<emphasis id="B91505665164F75D95B1FCF3FE10FCF8" bold="true" box="[199,486,818,844]" pageId="29" pageNumber="68">Justification of transfer.</emphasis>
The illustration of the palp provided in
<bibRefCitation id="EFF0A4855164F75D96D3FCF2FBCDFCF9" author="Simon, E." box="[933,1083,819,845]" pageId="29" pageNumber="68" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon (1914</bibRefCitation>
: fig. 398) is diagnostic and was used for comparison. Even though we did not examine the
<typeStatus id="54DA67D65164F75D965EFC96FC95FCC5" box="[808,867,855,881]" pageId="29" pageNumber="68">types</typeStatus>
of
<taxonomicName id="4C61A2F75164F75D96F1FC96FBB4FCC4" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Simon" baseAuthorityYear="1878" box="[903,1090,855,881]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="subSpecies" species="senilis" subSpecies="auspex">
<emphasis id="B91505665164F75D96F1FC96FC6EFCC4" box="[903,920,855,880]" italics="true" pageId="29" pageNumber="68">P</emphasis>
.
<emphasis id="B91505665164F75D96D0FC96FBB4FCC4" box="[934,1090,855,881]" italics="true" pageId="29" pageNumber="68">senilis auspex</emphasis>
</taxonomicName>
, the original illustration shows that this species has a robust RTA with an apical bifurcation, which is consistent with the RTA of male
<taxonomicName id="4C61A2F75164F75D906CFCBAFF1EFC0D" pageId="29" pageNumber="68">
<emphasis id="B91505665164F75D906CFCBAFA6BFC21" box="[1306,1437,891,917]" italics="true" pageId="29" pageNumber="68">Trichothyse</emphasis>
species
</taxonomicName>
(cf.
<bibRefCitation id="EFF0A4855164F75D946CFC5EFE53FC0D" author="Simon, E." box="[282,421,927,953]" pageId="29" pageNumber="68" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon 1914</bibRefCitation>
: fig. 398 and
<bibRefCitation id="EFF0A4855164F75D974FFC5EFD32FC0D" author="Simon, E." box="[569,708,927,953]" pageId="29" pageNumber="68" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon 1914</bibRefCitation>
: fig. 395,
<bibRefCitation id="EFF0A4855164F75D9647FC5EFC5CFC0D" author="Fage, L." box="[817,938,927,953]" pageId="29" pageNumber="68" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage 1929</bibRefCitation>
: fig. 1,
<bibRefCitation id="EFF0A4855164F75D968CFC5EFB5AFC0D" author="Di Franco, F." box="[1018,1196,927,953]" pageId="29" pageNumber="68" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco 2001</bibRefCitation>
: figs 34,
<bibRefCitation id="EFF0A4855164F75D906BFC5EFA6EFC0D" author="Levy, G." box="[1309,1432,927,953]" pageId="29" pageNumber="68" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: fig. 10, 2009: fig. 45,
<bibRefCitation id="EFF0A4855164F75D94E5FC02FD3DFC69" author="Fomichev, A. A. &amp; Marusik, Y. M." box="[403,715,963,989]" pageId="29" pageNumber="68" pagination="443 - 457" refId="ref18957" refString="Fomichev, A. A. &amp; Marusik, Y. M. (2021) A survey of East Palaearctic Gnaphosidae (Araneae). 11. New data on Gnaphosidae (Araneae) from Tajikistan. Zootaxa, 4966 (4), 443-457. https://doi.org/10.11646/zootaxa.4966.4.3" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 23, and
<figureCitation id="135AC5F15164F75D9614FC02FBC1FC69" box="[866,1079,963,989]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="29" pageNumber="68">Figs 109110, 112</figureCitation>
,
<figureCitation id="135AC5F15164F75D9132FC02FB52FC69" box="[1092,1188,963,989]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="29" pageNumber="68">116117</figureCitation>
herein). We therefore propose the transfer of
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<emphasis id="B91505665164F75D94EDFC26FE5AFBB4" box="[411,428,999,1024]" italics="true" pageId="29" pageNumber="68">P</emphasis>
.
<emphasis id="B91505665164F75D94CCFC26FDA1FBB4" box="[442,599,999,1025]" italics="true" pageId="29" pageNumber="68">senilis auspex</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75164F75D970CFC26FD0BFBB5" authorityName="Tucker" authorityYear="1923" box="[634,765,999,1025]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665164F75D970CFC26FD0BFBB5" box="[634,765,999,1025]" italics="true" pageId="29" pageNumber="68">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5164F75D95B1FBCBFBD6FB91" box="[199,1056,1034,1061]" pageId="29" pageNumber="68" type="distribution">
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<emphasis id="B91505665164F75D95B1FBCBFE97FB90" bold="true" box="[199,353,1034,1060]" pageId="29" pageNumber="68">Distribution.</emphasis>
Known only from
<collectingCountry id="F37699E45164F75D9741FBCAFD75FB91" box="[567,643,1035,1061]" name="France" pageId="29" pageNumber="68">France</collectingCountry>
in Mediterranean Europe (
<figureCitation id="135AC5F15164F75D96C7FBCAFBE5FB91" box="[945,1043,1035,1061]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="29" pageNumber="68">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,300 @@
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<emphasis id="B91505665165F75C95E1FDE3FE42FD89" bold="true" box="[151,436,546,573]" italics="true" pageId="28" pageNumber="67">Trichothyse poonaensis</emphasis>
(
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)
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<emphasis id="B91505665165F75C95E1FDA6FE7AFDCB" box="[151,396,615,639]" italics="true" pageId="28" pageNumber="67">Phaeocedus poonaensis</emphasis>
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, figs 158163 (
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:
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:
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:
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near
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University [
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,
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,
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a.s.l.
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, leg.
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,
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(4493/18), examined
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(
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images are provided in
<bibRefCitation id="EFF0A4855165F75C976AFD65FD0AFD09" author="Sankaran, P. M. &amp; Caleb, J. T. D. &amp; Sebastian, P. A." box="[540,764,676,701]" pageId="28" pageNumber="67" pagination="1325 - 1336" refId="ref19852" refString="Sankaran, P. M., Caleb, J. T. D. &amp; Sebastian, P. A. (2020) On the taxonomic validity of Indian ground spiders: III. Genus Phaeocedus Simon, 1893 (Araneae: Gnaphosidae). Journal of Natural History, 54, 1325-1336. https://doi.org/10.1080/00222933.2020.1797201" type="journal article" year="2020">
Sankaran
<emphasis id="B91505665165F75C97F7FD64FD42FD08" box="[641,692,676,700]" italics="true" pageId="28" pageNumber="67">et al.</emphasis>
(2020)
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—not re-examined).
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<emphasis id="B91505665165F75C95E1FD02FE7BFD6F" box="[151,397,707,731]" italics="true" pageId="28" pageNumber="67">Poecilochroa taborensis</emphasis>
<bibRefCitation id="EFF0A4855165F75C94E5FD05FDCFFD6F" author="Levy, G." box="[403,569,707,732]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy, 1999: 436</bibRefCitation>
</taxonomicName>
, figs 1718, 2009: 20, figs 4446;
<bibRefCitation id="EFF0A4855165F75C96E0FD02FB36FD6F" author="Lissner, J. &amp; Chatzaki, M." box="[918,1216,707,731]" pageId="28" pageNumber="67" pagination="16 - 21" refId="ref19447" refString="Lissner, J. &amp; Chatzaki, M. (2016) First records of Poecilochroa taborensis Levy, 1999 (Araneae: Gnaphosidae) from Greece and Cyprus with notes on some closely related species. Nieuwsbrief SPINED, 36, 16-21." type="journal article" year="2016">Lissner &amp; Chatzaki, 2016: 17</bibRefCitation>
, figs 1, 2AC, 3AD,
</paragraph>
<paragraph id="8BDED9745165F75C95B1FD23FA6AFCE2" blockId="28.[151,1437,615,885]" pageId="28" pageNumber="67">
4AE, 5AC, 6AD;
<bibRefCitation id="EFF0A4855165F75C94D5FD23FD4AFD4E" author="Sankaran, P. M. &amp; Caleb, J. T. D. &amp; Sebastian, P. A." box="[419,700,738,762]" pageId="28" pageNumber="67" pagination="1325 - 1336" refId="ref19852" refString="Sankaran, P. M., Caleb, J. T. D. &amp; Sebastian, P. A. (2020) On the taxonomic validity of Indian ground spiders: III. Genus Phaeocedus Simon, 1893 (Araneae: Gnaphosidae). Journal of Natural History, 54, 1325-1336. https://doi.org/10.1080/00222933.2020.1797201" type="journal article" year="2020">
Sankaran
<emphasis id="B91505665165F75C977EFD22FDB7FD4E" box="[520,577,738,762]" italics="true" pageId="28" pageNumber="67">et al.,</emphasis>
2020: 1329
</bibRefCitation>
, fig. 3ac.
<taxonomicName id="4C61A2F75165F75C95E1FCC1FD99FCAD" authority="Wunderlich, 2011: 49" authorityName="Wunderlich" authorityPageNumber="49" authorityYear="2011" box="[151,623,768,793]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="sabulum">
<emphasis id="B91505665165F75C95E1FCC1FE7DFCAC" box="[151,395,768,792]" italics="true" pageId="28" pageNumber="67">Macarophaeus sabulum</emphasis>
<bibRefCitation id="EFF0A4855165F75C94E6FCC1FD99FCAD" author="Wunderlich, J." box="[400,623,768,793]" pageId="28" pageNumber="67" pagination="1 - 640" refId="ref20444" refString="Wunderlich, J. (2011) Extant and fossil spiders (Araneae). Beitrage zur Araneologie, 6, 1-640." type="journal article" year="2011">Wunderlich, 2011: 49</bibRefCitation>
</taxonomicName>
, fig. 68hi, 2012: 187, figs 811.
<taxonomicName id="4C61A2F75165F75C95E1FCDEFD79FC83" authority="Wunderlich, 2017: 315" authorityName="Wunderlich" authorityPageNumber="315" authorityYear="2017" box="[151,655,799,823]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="taborensis">
<emphasis id="B91505665165F75C95E1FCDEFE68FC83" box="[151,414,799,823]" italics="true" pageId="28" pageNumber="67">Macarophaeus taborensis</emphasis>
<bibRefCitation id="EFF0A4855165F75C94D5FCDEFD79FC83" author="Wunderlich, J." box="[419,655,799,823]" pageId="28" pageNumber="67" pagination="298 - 326" refId="ref20514" refString="Wunderlich, J. (2017) Descriptions, notes and synonyms of some mainly Mediterranean and Macaronesian spiders (Araneae) of various families. Beitrage zur Araneologie, 10, 298-326 + 354." type="journal article" year="2017">Wunderlich, 2017: 315</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C61A2F75165F75C95E1FCFFFDF6FCE2" authority="(Tikader)" baseAuthorityName="Tikader" baseAuthorityYear="1982" box="[151,512,830,854]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="poonaensis">
<emphasis id="B91505665165F75C95E1FCFFFE61FCE1" box="[151,407,830,854]" italics="true" pageId="28" pageNumber="67">Poecilochroa poonaensis</emphasis>
(Tikader)
</taxonomicName>
: Sankaran
<emphasis id="B91505665165F75C9718FCFEFD50FCE2" box="[622,678,830,854]" italics="true" pageId="28" pageNumber="67">et al.,</emphasis>
2021: 3003;
<bibRefCitation id="EFF0A4855165F75C965BFCFFFBBEFCE2" author="Domenech, M. &amp; Mendez, O. &amp; Espigule-Pons, J. &amp; Carbonell, R. &amp; Pinilla-Rosa, M." box="[813,1096,830,854]" pageId="28" pageNumber="67" pagination="159 - 166" refId="ref18871" refString="Domenech, M., Mendez, O., Espigule-Pons, J., Carbonell, R. &amp; Pinilla-Rosa, M. (2023) Nuevas citas aracnologicas (Araneae) en Cataluna (NE de la Peninsula Iberica). Revista Iberica de Aracnologia, 42, 159-166." type="journal article" year="2023">
Domènech
<emphasis id="B91505665165F75C96D7FCFEFC2CFCE2" box="[929,986,830,854]" italics="true" pageId="28" pageNumber="67">et al.,</emphasis>
2023: 160
</bibRefCitation>
, fig. 1AD;
<bibRefCitation id="EFF0A4855165F75C91B1FCFFFA6AFCE2" author="Bosmans, R. &amp; Gavalas, I." box="[1223,1436,830,854]" pageId="28" pageNumber="67" pagination="1 - 91" refId="ref18487" refString="Bosmans, R. &amp; Gavalas, I. (2023) The spiders (Araneae) of the tiny Greek island Iraklia (Kiklades), with the descriptions of 5 new Harpactea species (Araneae: Dysderidae), 3 species new to Europe and 8 new to Greece. Parnassiana Archives, 11 (Supplementum 1), 1-91." type="journal article" year="2023">Bosmans &amp; Gavalas,</bibRefCitation>
</paragraph>
<paragraph id="8BDED9745165F75C95B1FC9CFE50FCC0" blockId="28.[151,1437,615,885]" box="[199,422,860,885]" pageId="28" pageNumber="67">2023: 50, fig. 25AC.</paragraph>
<paragraph id="8BDED9745165F75C95E1FC63FF1FFC55" blockId="28.[151,1437,930,1353]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C95E1FC63FEDFFC08" bold="true" box="[151,297,930,956]" pageId="28" pageNumber="67">Description.</emphasis>
<emphasis id="B91505665165F75C9446FC62FE9CFC09" box="[304,362,931,957]" italics="true" pageId="28" pageNumber="67">Male</emphasis>
and
<emphasis id="B91505665165F75C94D7FC62FE1AFC09" box="[417,492,931,957]" italics="true" pageId="28" pageNumber="67">female</emphasis>
. For description of the male and female, see
<bibRefCitation id="EFF0A4855165F75C969CFC62FB99FC09" author="Levy, G." box="[1002,1135,931,957]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy (1999</bibRefCitation>
,
<bibRefCitation id="EFF0A4855165F75C910DFC62FB43FC09" author="Levy, G." box="[1147,1205,931,957]" pageId="28" pageNumber="67" pagination="1 - 49" refId="ref19403" refString="Levy, G. (2009) New ground-spider genera and species with annexed checklist of the Gnaphosidae (Araneae) of Israel. Zootaxa, 2066, 1-49. https://doi.org/10.11646/zootaxa.2066.1.1" type="journal article" year="2009">2009</bibRefCitation>
) and
<bibRefCitation id="EFF0A4855165F75C9182FC62FF13FC55" author="Sankaran, P. M. &amp; Caleb, J. T. D. &amp; Sebastian, P. A." pageId="28" pageNumber="67" pagination="1325 - 1336" refId="ref19852" refString="Sankaran, P. M., Caleb, J. T. D. &amp; Sebastian, P. A. (2020) On the taxonomic validity of Indian ground spiders: III. Genus Phaeocedus Simon, 1893 (Araneae: Gnaphosidae). Journal of Natural History, 54, 1325-1336. https://doi.org/10.1080/00222933.2020.1797201" type="journal article" year="2020">
Sankaran
<emphasis id="B91505665165F75C9012FC62FA6BFC09" box="[1380,1437,931,957]" italics="true" pageId="28" pageNumber="67">et al.</emphasis>
(2020)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED9745165F75C95B1FC2BFDF6FA91" blockId="28.[151,1437,930,1353]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C95B1FC2BFE11FBB0" bold="true" box="[199,487,1002,1028]" pageId="28" pageNumber="67">Justification of transfer.</emphasis>
An examination of the
<typeStatus id="54DA67D65165F75C9783FC2AFCA1FBB1" box="[757,855,1003,1029]" pageId="28" pageNumber="67" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C61A2F75165F75C9609FC2AFBEBFBB0" baseAuthorityName="Tikader" baseAuthorityYear="1982" box="[895,1053,1003,1029]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="poonaensis">
<emphasis id="B91505665165F75C9609FC2AFC66FBB0" box="[895,912,1003,1028]" italics="true" pageId="28" pageNumber="67">P</emphasis>
.
<emphasis id="B91505665165F75C96E9FC2AFBEBFBB0" box="[927,1053,1003,1028]" italics="true" pageId="28" pageNumber="67">poonaensis</emphasis>
</taxonomicName>
in a recent paper (
<bibRefCitation id="EFF0A4855165F75C9187FC2AFF25FB9D" author="Sankaran, P. M. &amp; Caleb, J. T. D. &amp; Sebastian, P. A." pageId="28" pageNumber="67" pagination="1325 - 1336" refId="ref19852" refString="Sankaran, P. M., Caleb, J. T. D. &amp; Sebastian, P. A. (2020) On the taxonomic validity of Indian ground spiders: III. Genus Phaeocedus Simon, 1893 (Araneae: Gnaphosidae). Journal of Natural History, 54, 1325-1336. https://doi.org/10.1080/00222933.2020.1797201" type="journal article" year="2020">
Sankaran
<emphasis id="B91505665165F75C9015FC2AFA6BFBB1" box="[1379,1437,1003,1029]" italics="true" pageId="28" pageNumber="67">et al.</emphasis>
2020
</bibRefCitation>
), as well as good illustrations of male and female genitalia available in
<bibRefCitation id="EFF0A4855165F75C917AFBCEFB62FB9D" author="Levy, G." box="[1036,1172,1039,1065]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy (1999</bibRefCitation>
: figs 1718, 2009: figs 4445),
<bibRefCitation id="EFF0A4855165F75C9581FBF2FDF5FBF9" author="Domenech, M. &amp; Mendez, O. &amp; Espigule-Pons, J. &amp; Carbonell, R. &amp; Pinilla-Rosa, M." box="[247,515,1075,1101]" pageId="28" pageNumber="67" pagination="159 - 166" refId="ref18871" refString="Domenech, M., Mendez, O., Espigule-Pons, J., Carbonell, R. &amp; Pinilla-Rosa, M. (2023) Nuevas citas aracnologicas (Araneae) en Cataluna (NE de la Peninsula Iberica). Revista Iberica de Aracnologia, 42, 159-166." type="journal article" year="2023">
Domènech
<emphasis id="B91505665165F75C940CFBF2FE43FBF9" box="[378,437,1075,1101]" italics="true" pageId="28" pageNumber="67">et al.</emphasis>
(2023
</bibRefCitation>
: fig. 1AB) and
<bibRefCitation id="EFF0A4855165F75C97B1FBF2FBF6FBF9" author="Bosmans, R. &amp; Gavalas, I." box="[711,1024,1075,1101]" pageId="28" pageNumber="67" pagination="1 - 91" refId="ref18487" refString="Bosmans, R. &amp; Gavalas, I. (2023) The spiders (Araneae) of the tiny Greek island Iraklia (Kiklades), with the descriptions of 5 new Harpactea species (Araneae: Dysderidae), 3 species new to Europe and 8 new to Greece. Parnassiana Archives, 11 (Supplementum 1), 1-91." type="journal article" year="2023">Bosmans &amp; Gavalas (2023</bibRefCitation>
: fig. 25BC), showed that it has all of the diagnostic features of
<taxonomicName id="4C61A2F75165F75C94ABFB96FD4AFBC5" box="[477,700,1111,1137]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C94ABFB96FD96FBC5" box="[477,608,1111,1137]" italics="true" pageId="28" pageNumber="67">Trichothyse</emphasis>
species
</taxonomicName>
, although at the time the association with this genus was unclear due to its distribution range. The male palp is typical of male
<taxonomicName id="4C61A2F75165F75C961AFBBAFBBAFB21" box="[876,1100,1147,1173]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C961AFBBAFC19FB21" box="[876,1007,1147,1173]" italics="true" pageId="28" pageNumber="67">Trichothyse</emphasis>
species
</taxonomicName>
, as described for
<taxonomicName id="4C61A2F75165F75C906DFBBAFA6AFB20" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[1307,1436,1147,1173]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="antineae" status="comb. nov.">
<emphasis id="B91505665165F75C906DFBBAFADDFB20" box="[1307,1323,1147,1172]" italics="true" pageId="28" pageNumber="67">T</emphasis>
.
<emphasis id="B91505665165F75C904AFBBAFA6AFB20" box="[1340,1436,1147,1172]" italics="true" pageId="28" pageNumber="67">antineae</emphasis>
</taxonomicName>
<emphasis id="B91505665165F75C95E1FB5EFEEEFB0D" bold="true" box="[151,280,1183,1209]" pageId="28" pageNumber="67">
<taxonomicNameLabel id="A226B81D5165F75C95E1FB5EFEEEFB0D" box="[151,280,1183,1209]" pageId="28" pageNumber="67" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
above, and the epigyne has a small median atrium, as well as a vulva with features as described for
<taxonomicName id="4C61A2F75165F75C90F0FB5EFED1FB69" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B91505665165F75C90F0FB5EFA60FB0C" box="[1414,1430,1183,1208]" italics="true" pageId="28" pageNumber="67">T</emphasis>
.
<emphasis id="B91505665165F75C95E1FB02FED1FB69" box="[151,295,1219,1245]" italics="true" pageId="28" pageNumber="67">jodhpurensis</emphasis>
</taxonomicName>
<emphasis id="B91505665165F75C945BFB02FE5AFB69" bold="true" box="[301,428,1219,1245]" pageId="28" pageNumber="67">
<taxonomicNameLabel id="A226B81D5165F75C945BFB02FE5AFB69" box="[301,428,1219,1245]" pageId="28" pageNumber="67" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
above (cf.
<figureCitation id="135AC5F15165F75C9751FB02FD6EFB69" box="[551,664,1219,1245]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 920" captionStartId-0="3.[151,264,1372,1396]" captionStartId-1="4.[151,264,1607,1631]" captionTargetBox-0="[152,1436,179,1346]" captionTargetBox-1="[151,1436,179,1581]" captionTargetId-0="figure-17@3.[151,1436,179,1346]" captionTargetId-1="figure-17@4.[151,1436,179,1581]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." captionText-1="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="28" pageNumber="67">Figs 120</figureCitation>
,
<figureCitation id="135AC5F15165F75C97D5FB03FD1DFB69" box="[675,747,1218,1245]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="28" pageNumber="67">5457</figureCitation>
,
<figureCitation id="135AC5F15165F75C9780FB02FCA1FB69" box="[758,855,1219,1245]" captionStart-0="FIGURES 109115" captionStart-1="FIGURES 116119" captionStartId-0="24.[151,257,1943,1967]" captionStartId-1="25.[151,264,1479,1503]" captionTargetBox-0="[190,1398,195,1913]" captionTargetBox-1="[161,1426,189,1443]" captionTargetId-0="figure-17@24.[179,1408,179,1917]" captionTargetId-1="figure-17@25.[151,1436,179,1453]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." captionText-1="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="28" pageNumber="67">109119</figureCitation>
and
<bibRefCitation id="EFF0A4855165F75C96FDFB02FBF1FB69" author="Levy, G." box="[907,1031,1219,1245]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: figs 1718, 2009: 4445). Based on the similarity in genitalic structure and somatic morphology of this species to
<taxonomicName id="4C61A2F75165F75C9172FB26FB71FAB5" authorityName="Tucker" authorityYear="1923" box="[1028,1159,1255,1281]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665165F75C9172FB26FB71FAB5" box="[1028,1159,1255,1281]" italics="true" pageId="28" pageNumber="67">Trichothyse</emphasis>
</taxonomicName>
, we propose the transfer of
<taxonomicName id="4C61A2F75165F75C95C0FACAFEA5FA90" baseAuthorityName="Tikader" baseAuthorityYear="1982" box="[182,339,1291,1317]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="poonaensis">
<emphasis id="B91505665165F75C95C0FACAFF31FA90" box="[182,199,1291,1316]" italics="true" pageId="28" pageNumber="67">P</emphasis>
.
<emphasis id="B91505665165F75C95A3FACAFEA5FA90" box="[213,339,1291,1316]" italics="true" pageId="28" pageNumber="67">poonaensis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75165F75C9400FACAFE0FFA91" authorityName="Tucker" authorityYear="1923" box="[374,505,1291,1317]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665165F75C9400FACAFE0FFA91" box="[374,505,1291,1317]" italics="true" pageId="28" pageNumber="67">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5165F75C95B1FAEFFB8FFAFD" box="[199,1145,1326,1353]" pageId="28" pageNumber="67" type="distribution">
<paragraph id="8BDED9745165F75C95B1FAEFFB8FFAFD" blockId="28.[151,1437,930,1353]" box="[199,1145,1326,1353]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C95B1FAEFFE97FAFC" bold="true" box="[199,353,1326,1352]" pageId="28" pageNumber="67">Distribution.</emphasis>
Known from
<collectingCountry id="F37699E45165F75C9488FAEEFDA4FAFD" box="[510,594,1327,1353]" name="Cyprus" pageId="28" pageNumber="67">Cyprus</collectingCountry>
,
<collectingCountry id="F37699E45165F75C972BFAEEFD59FAFD" box="[605,687,1327,1353]" name="Greece" pageId="28" pageNumber="67">Greece</collectingCountry>
,
<collectingCountry id="F37699E45165F75C97CCFAEEFD00FAFD" box="[698,758,1327,1353]" name="India" pageId="28" pageNumber="67">India</collectingCountry>
,
<collectingCountry id="F37699E45165F75C9674FAEEFCB5FAFD" box="[770,835,1327,1353]" name="Israel" pageId="28" pageNumber="67">Israel</collectingCountry>
,
<collectingCountry id="F37699E45165F75C9638FAEEFC46FAFD" box="[846,944,1327,1353]" name="Portugal" pageId="28" pageNumber="67">Portugal</collectingCountry>
,
<collectingCountry id="F37699E45165F75C96CDFAEEFC0DFAFD" box="[955,1019,1327,1353]" name="Spain" pageId="28" pageNumber="67">Spain</collectingCountry>
(
<figureCitation id="135AC5F15165F75C917CFAEEFB9AFAFD" box="[1034,1132,1327,1353]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="28" pageNumber="67">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,302 @@
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<mods:title id="27FA89A8D25A9A9D2D9119AA1CF61B0E">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<heading id="D0966E185165F75C95E1FA5BFC7AFA01" bold="true" box="[151,908,1434,1461]" fontSize="11" level="1" pageId="28" pageNumber="67" reason="1">
<emphasis id="B91505665165F75C95E1FA5BFC7AFA01" bold="true" box="[151,908,1434,1461]" pageId="28" pageNumber="67">
<taxonomicName id="4C61A2F75165F75C95E1FA5BFD08FA00" authority="(O. Pickard-Cambridge, 1872)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[151,766,1434,1461]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="senilis" status="comb. nov.">
<emphasis id="B91505665165F75C95E1FA5BFE8CFA01" bold="true" box="[151,378,1434,1461]" italics="true" pageId="28" pageNumber="67">Trichothyse senilis</emphasis>
(O.
<bibRefCitation id="EFF0A4855165F75C94C6FA5BFD00FA00" author="Pickard-Cambridge, O." box="[432,758,1434,1460]" pageId="28" pageNumber="67" pagination="212 - 354" refId="ref19729" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40, 212-354, pls. XIII - XVI. https://doi.org/10.1111/j.1469-7998.1872.tb00489.x" type="journal article" year="1872">Pickard-Cambridge, 1872</bibRefCitation>
)
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<emphasis id="B91505665165F75C95E1FA21FED9FA43" box="[151,303,1503,1527]" italics="true" pageId="28" pageNumber="67">Drassus senilis</emphasis>
<materialsCitation id="3B09D3295165F75C9445FA1EFD70FA43" box="[307,646,1503,1527]" collectionCode="OUMNH" country="Israel" latitude="30.583334" location="Alexandria" longLatPrecision="1220" longitude="36.233334" pageId="28" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Al Iskandariyah" typeStatus="syntype">
O.
<bibRefCitation id="EFF0A4855165F75C9426FA1EFD70FA43" author="Pickard-Cambridge, O." box="[336,646,1503,1527]" pageId="28" pageNumber="67" pagination="212 - 354" refId="ref19729" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40, 212-354, pls. XIII - XVI. https://doi.org/10.1111/j.1469-7998.1872.tb00489.x" type="journal article" year="1872">Pickard-Cambridge, 1872: 236</bibRefCitation>
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, plate XV, fig. 13 (
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:
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:
</emphasis>
Jordan [
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,
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], collector unknown, date unknown, repository OUMNH (B. 243, t. 51) and from
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:
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near
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, collector unknown, date unknown, repository OUMNH (B. 241, t. 102)—neither examined).
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<treatmentCitation id="0AC0FF655165F75C95E1F9FDFD2EF9E7" author="Pickard-Cambridge, O." box="[151,728,1595,1619]" page="392" pageId="28" pageNumber="67" year="1874">
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<emphasis id="B91505665165F75C95E1F9FDFE8EF9E7" box="[151,376,1596,1619]" italics="true" pageId="28" pageNumber="67">Drassus campestratus</emphasis>
O.
<bibRefCitation id="EFF0A4855165F75C94EBF9FAFD2EF9E7" author="Pickard-Cambridge, O." box="[413,728,1595,1619]" pageId="28" pageNumber="67" pagination="370 - 419" refId="ref19799" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42, 370-419, pls. LI - LII. https://doi.org/10.1111/j.1096-3642.1874.tb02495.x" type="journal article" year="1874">Pickard-Cambridge, 1874: 392</bibRefCitation>
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, plate LI, fig. 17.
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</paragraph>
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<treatmentCitation id="0AC0FF655165F75C95E1F99BFDD0F9C6" author="Thorell, T." box="[151,550,1626,1650]" page="82" pageId="28" pageNumber="67" year="1875">
<taxonomicName id="4C61A2F75165F75C95E1F99BFDD0F9C6" authority="Thorell, 1875 a: 82" authorityName="Thorell" authorityPageNumber="82" authorityYear="1875" box="[151,550,1626,1650]" class="Arachnida" family="Gnaphosidae" genus="Prosthesima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B91505665165F75C95E1F99BFE97F9C6" box="[151,353,1626,1650]" italics="true" pageId="28" pageNumber="67">Prosthesima rufipes</emphasis>
<bibRefCitation id="EFF0A4855165F75C9411F99BFDD0F9C6" author="Thorell, T." box="[359,550,1626,1650]" pageId="28" pageNumber="67" pagination="39 - 122" refId="ref20279" refString="Thorell, T. (1875 a) Verzeichniss sudrussischer Spinnen. Horae Societatis Entomologicae Rossicae, 11, 39-122." type="journal article" year="1875">Thorell, 1875a: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
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<bibRefCitation id="EFF0A4855165F75C9746F99BFD53F9C6" author="Thorell, T." box="[560,677,1626,1650]" pageId="28" pageNumber="67" pagination="1 - 204" refId="ref20302" refString="Thorell, T. (1875 b) Descriptions of several European and North African spiders. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 13, 1-204." type="journal article" year="1875">1875b: 110</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BDED9745165F75C95E1F9B8FDE8F924" blockId="28.[151,1437,1503,1926]" box="[151,542,1656,1681]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F9B8FDE8F924" box="[151,542,1656,1681]" pageId="28" pageNumber="67">
<treatmentCitation id="0AC0FF655165F75C95E1F9B8FDECF924" author="Simon, E." box="[151,538,1656,1681]" page="153" pageId="28" pageNumber="67" year="1878">
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<emphasis id="B91505665165F75C95E1F9B8FEAAF924" box="[151,348,1656,1680]" italics="true" pageId="28" pageNumber="67">Drassus dimidiatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
<paragraph id="8BDED9745165F75C95E1F959FD1FF91B" blockId="28.[151,1437,1503,1926]" box="[151,745,1687,1711]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F959FD1FF91B" box="[151,745,1687,1711]" pageId="28" pageNumber="67">
<treatmentCitation id="0AC0FF655165F75C95E1F959FDA0F91B" author="Koch, L." box="[151,598,1687,1711]" page="40" pageId="28" pageNumber="67" year="1878">
<taxonomicName id="4C61A2F75165F75C95E1F959FDA0F91B" authority="L. Koch, 1878: 40" authorityName="L. Koch" authorityPageNumber="40" authorityYear="1878" box="[151,598,1687,1711]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="flavo-maculatus">
<emphasis id="B91505665165F75C95E1F959FE62F91B" box="[151,404,1687,1711]" italics="true" pageId="28" pageNumber="67">Drassus flavo-maculatus</emphasis>
L.
<bibRefCitation id="EFF0A4855165F75C94C0F956FDA0F91B" author="Koch, L." box="[438,598,1687,1711]" pageId="28" pageNumber="67" pagination="36 - 71" refId="ref19116" refString="Koch, L. (1878) Kaukasische Arachnoiden. In: Schneider, O. (Ed.), Naturwissenschaftliche Beitrage zur Kenntniss der Kaukasuslander auf Grund seiner Sammelbeute. Sitzungs-Berichte der naturwissenschaftlichen Gesellschaft Isis in Dresden, 1878, 36-71 + 159 - 160, pls. I - II." type="journal article" year="1878">Koch, 1878: 40</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, plate I, fig. 2.
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</paragraph>
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<treatmentCitationGroup id="AB71FE5A5165F75C95E1F977FDCAF97A" box="[151,572,1718,1742]" pageId="28" pageNumber="67">
<treatmentCitation id="0AC0FF655165F75C95E1F977FDCEF97A" author="Pavesi, P." box="[151,568,1718,1742]" page="350" pageId="28" pageNumber="67" year="1880">
<taxonomicName id="4C61A2F75165F75C95E1F977FDCEF97A" authority="Pavesi, 1880: 350" authorityName="Pavesi" authorityPageNumber="350" authorityYear="1880" box="[151,568,1718,1742]" class="Arachnida" family="Gnaphosidae" genus="Prosthesima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="incompta">
<emphasis id="B91505665165F75C95E1F977FE8FF979" box="[151,377,1718,1742]" italics="true" pageId="28" pageNumber="67">Prosthesima incompta</emphasis>
<bibRefCitation id="EFF0A4855165F75C94F6F977FDCEF97A" author="Pavesi, P." box="[384,568,1718,1742]" pageId="28" pageNumber="67" pagination="283 - 388" refId="ref19646" refString="Pavesi, P. (1880) Studi sugli Aracnidi africani. I. Aracnidi di Tunisia. Annali del Museo Civico di Storia Naturale di Genova, 15, 283-388." type="journal article" year="1880">Pavesi, 1880: 350</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745165F75C95E1F915FCD2F959" blockId="28.[151,1437,1503,1926]" box="[151,804,1748,1773]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F915FCD2F959" box="[151,804,1748,1773]" pageId="28" pageNumber="67">
<taxonomicName id="4C61A2F75165F75C95E1F915FDA9F958" authority="(O. Pickard-Cambridge)" baseAuthorityName="O. Pickard-Cambridge" box="[151,607,1748,1773]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="senilis">
<emphasis id="B91505665165F75C95E1F915FEA9F958" box="[151,351,1748,1772]" italics="true" pageId="28" pageNumber="67">Scotophaeus senilis</emphasis>
(O. Pickard-Cambridge)
</taxonomicName>
:
<treatmentCitation id="0AC0FF655165F75C971EF915FCD6F959" author="Simon, E." box="[616,800,1748,1773]" page="371" pageId="28" pageNumber="67" year="1893">
<bibRefCitation id="EFF0A4855165F75C971EF915FCD6F959" author="Simon, E." box="[616,800,1748,1773]" pageId="28" pageNumber="67" pagination="257 - 488" refId="ref20103" refString="Simon, E. (1893) Histoire naturelle des araignees. Deuxieme edition, tome premier. Roret, Paris, 232 pp. [pp. 257-488] https://doi.org/10.5962/bhl.title.51973" type="book chapter" year="1893">Simon, 1893: 371</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745165F75C95E1F932FD93F8BF" blockId="28.[151,1437,1503,1926]" box="[151,613,1779,1803]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F932FD93F8BF" box="[151,613,1779,1803]" pageId="28" pageNumber="67">
<treatmentCitation id="0AC0FF655165F75C95E1F932FD97F8BF" author="Simon, E." box="[151,609,1779,1803]" page="372" pageId="28" pageNumber="67" year="1893">
<taxonomicName id="4C61A2F75165F75C95E1F932FD97F8BF" ID-CoL="4KTBK" authority="Simon, 1893: 372" authorityName="Simon" authorityPageNumber="372" authorityYear="1893" box="[151,609,1779,1803]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="campestrata">
<emphasis id="B91505665165F75C95E1F932FE54F8BF" box="[151,418,1779,1803]" italics="true" pageId="28" pageNumber="67">Poecilochroa campestrata</emphasis>
<bibRefCitation id="EFF0A4855165F75C94DFF932FD97F8BF" author="Simon, E." box="[425,609,1779,1803]" pageId="28" pageNumber="67" pagination="257 - 488" refId="ref20103" refString="Simon, E. (1893) Histoire naturelle des araignees. Deuxieme edition, tome premier. Roret, Paris, 232 pp. [pp. 257-488] https://doi.org/10.5962/bhl.title.51973" type="book chapter" year="1893">Simon, 1893: 372</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745165F75C95E1F8D3FDAAF89E" blockId="28.[151,1437,1503,1926]" box="[151,604,1810,1834]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F8D3FDAAF89E" box="[151,604,1810,1834]" pageId="28" pageNumber="67">
<treatmentCitation id="0AC0FF655165F75C95E1F8D3FDAEF89E" author="Strand, E." box="[151,600,1810,1834]" page="66" pageId="28" pageNumber="67" year="1908">
<taxonomicName id="4C61A2F75165F75C95E1F8D3FDAEF89E" authority="Strand, 1908: 66" authorityName="Strand" authorityPageNumber="66" authorityYear="1908" box="[151,600,1810,1834]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="subincompta">
<emphasis id="B91505665165F75C95E1F8D3FE50F89E" box="[151,422,1810,1834]" italics="true" pageId="28" pageNumber="67">Poecilochroa subincompta</emphasis>
<bibRefCitation id="EFF0A4855165F75C94DBF8D3FDAEF89E" author="Strand, E." box="[429,600,1810,1834]" pageId="28" pageNumber="67" pagination="11 - 101" refId="ref20228" refString="Strand, E. (1908) Nordafrikanische, hauptsachlich von Carlo Freiherr von Erlanger gesammelte Aviculariidae, Drassidae und Theridiidae. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg, 64, 11-101. https://doi.org/10.5962/bhl.part.27907" type="journal article" year="1908">Strand, 1908: 66</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745165F75C95E1F8F1FCFBF8FD" blockId="28.[151,1437,1503,1926]" box="[151,781,1840,1865]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F8F1FCFBF8FD" box="[151,781,1840,1865]" pageId="28" pageNumber="67">
<treatmentCitation id="0AC0FF655165F75C95E1F8F1FDB5F8FC" author="Simon, E." box="[151,579,1840,1865]" page="185" pageId="28" pageNumber="67" year="1914">
<taxonomicName id="4C61A2F75165F75C95E1F8F1FDB5F8FC" ID-CoL="4KTBQ" authority="Simon, 1914: 185" authorityName="Simon" authorityPageNumber="185" authorityYear="1914" box="[151,579,1840,1865]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B91505665165F75C95E1F8F1FE70F8FC" box="[151,390,1840,1864]" italics="true" pageId="28" pageNumber="67">Poecilochroa dimidiata</emphasis>
<bibRefCitation id="EFF0A4855165F75C94FAF8F1FDB5F8FC" author="Simon, E." box="[396,579,1840,1865]" pageId="28" pageNumber="67" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon, 1914: 185</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 221, figs 347, 400.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745165F75C95E1F88EFEFFF832" blockId="28.[151,1437,1503,1926]" pageId="28" pageNumber="67">
<treatmentCitationGroup id="AB71FE5A5165F75C95E1F88EFEFFF832" pageId="28" pageNumber="67">
<taxonomicName id="4C61A2F75165F75C95E1F88EFD91F8D3" ID-CoL="4KTCG" authority="(O. Pickard-Cambridge)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[151,615,1871,1895]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="67" phylum="Arthropoda" rank="species" species="senilis">
<emphasis id="B91505665165F75C95E1F88EFE90F8D3" box="[151,358,1871,1895]" italics="true" pageId="28" pageNumber="67">Poecilochroa senilis</emphasis>
(O. Pickard-Cambridge)
</taxonomicName>
:
<treatmentCitation id="0AC0FF655165F75C9707F891FCEEF8D3" author="Levy, G." box="[625,792,1871,1896]" page="431" pageId="28" pageNumber="67" year="1999">
<bibRefCitation id="EFF0A4855165F75C9707F891FCEEF8D3" author="Levy, G." box="[625,792,1871,1896]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy, 1999: 431</bibRefCitation>
</treatmentCitation>
, figs 69 (for complete bibliography, see
<bibRefCitation id="EFF0A4855165F75C91C9F88EFF08F832" author="World Spider Catalog" pageId="28" pageNumber="67" refId="ref20403" refString="World Spider Catalog (2024) Version 25.5. Natural History Museum, Bern. online at http://wsc.nmbe.ch (accessed 22 November 2024) https://doi.org/10.24436/2" type="url" year="2024">World Spider Catalog 2024</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5165F75D95E1F875FBF1FE19" lastPageId="29" lastPageNumber="68" pageId="28" pageNumber="67" type="description">
<paragraph id="8BDED9745165F75C95E1F875FB83F87A" blockId="28.[151,1141,1972,1999]" box="[151,1141,1972,1999]" pageId="28" pageNumber="67">
<emphasis id="B91505665165F75C95E1F875FEDFF87A" bold="true" box="[151,297,1972,1998]" pageId="28" pageNumber="67">Description.</emphasis>
<emphasis id="B91505665165F75C9446F875FE9CF87A" box="[304,362,1972,1998]" italics="true" pageId="28" pageNumber="67">Male</emphasis>
and
<emphasis id="B91505665165F75C94D6F875FE1DF87A" box="[416,491,1972,1998]" italics="true" pageId="28" pageNumber="67">female</emphasis>
. For description of the male and female, see
<bibRefCitation id="EFF0A4855165F75C9695F874FB87F87A" author="Levy, G." box="[995,1137,1972,1999]" pageId="28" pageNumber="67" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy (1999)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED9745164F75D95B1FF57FBF1FE19" blockId="29.[151,1437,150,501]" pageId="29" pageNumber="68">
<emphasis id="B91505665164F75D95B1FF57FE16FF04" bold="true" box="[199,480,150,176]" pageId="29" pageNumber="68">Justification of transfer.</emphasis>
The illustrations of the male and female genitalia in
<bibRefCitation id="EFF0A4855164F75D9165FF56FB60FF05" author="Levy, G." box="[1043,1174,151,177]" pageId="29" pageNumber="68" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy(1999</bibRefCitation>
:figs 69) are diagnostic and were used for comparison. Even though we did not examine the
<typeStatus id="54DA67D65164F75D96E8FF7AFC2FFF61" box="[926,985,187,213]" pageId="29" pageNumber="68">types</typeStatus>
of
<taxonomicName id="4C61A2F75164F75D9176FF7AFB91FF61" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[1024,1127,187,213]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="species" species="senilis">
<emphasis id="B91505665164F75D9176FF7AFBE7FF60" box="[1024,1041,187,212]" italics="true" pageId="29" pageNumber="68">P</emphasis>
.
<emphasis id="B91505665164F75D9169FF7AFB91FF61" box="[1055,1127,187,213]" italics="true" pageId="29" pageNumber="68">senilis</emphasis>
</taxonomicName>
, subsequent illustrations of its genitalia are diagnostic and share all of the diagnostic features of male and female
<taxonomicName id="4C61A2F75164F75D9115FF1EFAB7FF4D" box="[1123,1345,223,249]" pageId="29" pageNumber="68">
<emphasis id="B91505665164F75D9115FF1EFB10FF4D" box="[1123,1254,223,249]" italics="true" pageId="29" pageNumber="68">Trichothyse</emphasis>
species
</taxonomicName>
, instead of
<emphasis id="B91505665164F75D95C1FEC2FE12FEA8" box="[183,484,259,285]" italics="true" pageId="29" pageNumber="68">
<taxonomicName id="4C61A2F75164F75D95C1FEC2FEBBFEA9" authorityName="Westring" authorityYear="1874" box="[183,333,259,285]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="genus" sensu="stricto">Poecilochroa</taxonomicName>
<taxonomicNameLabel id="A226B81D5164F75D9423FEC2FE12FEA8" box="[341,484,259,284]" pageId="29" pageNumber="68" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C61A2F75164F75D976BFEC2FD32FEA9" authorityName="Wunderlich" authorityYear="2011" box="[541,708,259,285]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665164F75D976BFEC2FD32FEA9" box="[541,708,259,285]" italics="true" pageId="29" pageNumber="68">Macarophaeus</emphasis>
</taxonomicName>
: distomedially originating broad conductor with prolateral fold, apically originating claw-like embolus with broad embolic base, medially contiguous sperm duct with single loop, robust RTA with apical bifurcation, epigyne with median extension and vulva with short copulatory ducts, large, bean-shaped primary, and small secondary spermathecae (cf.
<figureCitation id="135AC5F15164F75D9632FEAEFC43FE3D" box="[836,949,367,393]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 920" captionStartId-0="3.[151,264,1372,1396]" captionStartId-1="4.[151,264,1607,1631]" captionTargetBox-0="[152,1436,179,1346]" captionTargetBox-1="[151,1436,179,1581]" captionTargetId-0="figure-17@3.[151,1436,179,1346]" captionTargetId-1="figure-17@4.[151,1436,179,1581]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." captionText-1="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="29" pageNumber="68">Figs 120</figureCitation>
,
<figureCitation id="135AC5F15164F75D96B7FEAFFBFFFE3D" box="[961,1033,366,393]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="29" pageNumber="68">5457</figureCitation>
,
<figureCitation id="135AC5F15164F75D9162FEAEFB83FE3D" box="[1044,1141,367,393]" captionStart-0="FIGURES 109115" captionStart-1="FIGURES 116119" captionStartId-0="24.[151,257,1943,1967]" captionStartId-1="25.[151,264,1479,1503]" captionTargetBox-0="[190,1398,195,1913]" captionTargetBox-1="[161,1426,189,1443]" captionTargetId-0="figure-17@24.[179,1408,179,1917]" captionTargetId-1="figure-17@25.[151,1436,179,1453]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." captionText-1="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="29" pageNumber="68">109119</figureCitation>
and
<bibRefCitation id="EFF0A4855164F75D91DDFEAEFAD1FE3D" author="Levy, G." box="[1195,1319,367,393]" pageId="29" pageNumber="68" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: figs 69). Based on these observations, we propose transferring
<taxonomicName id="4C61A2F75164F75D9785FE52FCACFE19" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[755,858,403,429]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="species" species="senilis">
<emphasis id="B91505665164F75D9785FE52FCF2FE18" box="[755,772,403,428]" italics="true" pageId="29" pageNumber="68">P</emphasis>
.
<emphasis id="B91505665164F75D9664FE52FCACFE19" box="[786,858,403,429]" italics="true" pageId="29" pageNumber="68">senilis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75164F75D960BFE52FBF6FE19" authorityName="Tucker" authorityYear="1923" box="[893,1024,403,429]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665164F75D960BFE52FBF6FE19" box="[893,1024,403,429]" italics="true" pageId="29" pageNumber="68">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5164F75D95B1FE77FDE3FE41" pageId="29" pageNumber="68" type="distribution">
<paragraph id="8BDED9745164F75D95B1FE77FDE3FE41" blockId="29.[151,1437,150,501]" pageId="29" pageNumber="68">
<emphasis id="B91505665164F75D95B1FE77FE97FE64" bold="true" box="[199,353,438,464]" pageId="29" pageNumber="68">Distribution.</emphasis>
Known from Asia, Africa and Europe (
<figureCitation id="135AC5F15164F75D9655FE76FC70FE65" box="[803,902,439,465]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="29" pageNumber="68">Fig. 121</figureCitation>
). This is the only Palaearctic species that has a distribution extending into Africa.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,195 @@
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<mods:title id="431516C336B7672B1AE360327897A064">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:roleTerm id="E94F87A6FA5AA648893450513FCCE00F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BCF2599C4AA06D14285B396839481E7">Sankaran, Pradeep M.</mods:namePart>
<mods:nameIdentifier id="4E150CD6A36A124D22C8D860B4E668C4" type="ORCID">0000-0003-0884-5625</mods:nameIdentifier>
<mods:affiliation id="4AAB4D8F92F4575BA7D8D67752D7168A">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
<mods:nameIdentifier id="B78A3CFD36583EA7E7B06F1E3C6D002F" type="email">pradeepmspala@rediffmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="1E7396617951E519BDCC43471F441300" type="personal">
<mods:role id="87333A0F39EBFDF894BD5899C9869E30">
<mods:roleTerm id="20238F9DB56E2BD974F2E284184ACCDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="54B0529CA9316BA55906EAEB84C08CD4">Haddad, Charles R.</mods:namePart>
<mods:nameIdentifier id="CCB45B82B561A11AC71BA41C83BE3692" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="82E2C89594EC1079560CE42404888B80">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="FBA0C5172B079557147F87BC5CE9A750" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name id="A9040EF13C17B2C7BF1F414CDA0051E1" type="personal">
<mods:role id="B3D145A3E4EA2D455294A6CFFE57B743">
<mods:roleTerm id="B4B4FF3B887ABB18CCEB27922F19EB9B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="640A9A8CF9C37C26E19AF1193389C904">Tripathi, Rishikesh</mods:namePart>
<mods:nameIdentifier id="E9FDC7FE1C515657FA42B09E730FA5C2" type="ORCID">0000-0002-9192-4609</mods:nameIdentifier>
<mods:affiliation id="43C3CC95138F08151DDE8CFC4F77194E">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Thrissur, Kerala 680 125, India.</mods:affiliation>
<mods:nameIdentifier id="2643951A5E0EBE813F13AB0052368716" type="email">rishikesh.tripathi14@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="F69D2DB63B0868B09DC6AE59CAD11F4D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3FCA36DF74430908E7B2EC8DECE56676">
<mods:date id="EC472D000AF2FD86F6C70F16470A88C9">2025</mods:date>
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<mods:number id="D6B28E1C4C26B3E81EDF4E9A72E04A71">2025-01-31</mods:number>
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<mods:number id="C3D07C99F4F6C3A7B36091C62B6BA593">5583</mods:number>
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<mods:number id="02B70B053601C6A422D974135F7FC099">1</mods:number>
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<mods:start id="473F0C7464798B1FA7835100C2BE26A0">39</mods:start>
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<mods:identifier id="2B569F74036D605089B1FD150A26FD46" type="ISSN">1175-5326</mods:identifier>
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<heading id="D0966E18516CF75595E1FEABFBF9FE31" bold="true" box="[151,1039,362,389]" fontSize="11" level="1" pageId="21" pageNumber="60" reason="1">
<emphasis id="B9150566516CF75595E1FEABFBF9FE31" bold="true" box="[151,1039,362,389]" pageId="21" pageNumber="60">
<taxonomicName id="4C61A2F7516CF75595E1FEABFC77FE30" authority="(Cosar, Danisman &amp; Marusik, 2024)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Cosar, Danisman &amp; Marusik" baseAuthorityYear="2024" box="[151,897,362,389]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="ilkerakkusi" status="comb. nov.">
<emphasis id="B9150566516CF75595E1FEABFE42FE31" bold="true" box="[151,436,362,389]" italics="true" pageId="21" pageNumber="60">Trichothyse ilkerakkusi</emphasis>
(
<bibRefCitation id="EFF0A485516CF75594B5FEABFC8FFE30" author="Cosar, I. &amp; Danisman, T. &amp; Marusik, Y. M." box="[451,889,362,389]" pageId="21" pageNumber="60" pagination="112 - 124" refId="ref18552" refString="Cosar, I., Danisman, T. &amp; Marusik, Y. M. (2024) Two new species belonging to Coreodrassus and Poecilochroa (Aranei: Gnaphosidae) from Anatolia with comments on taxonomy of both genera. Arthropoda Selecta, 33 (1), 112-124. https://doi.org/10.15298/arthsel.33.1.11" type="journal article" year="2024">Coşar, Danişman &amp; Marusik, 2024</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A226B81D516CF75596FEFEAAFBF9FE31" box="[904,1039,363,389]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C37B8AFF516CF75595E1FE6EFB07FDB0" pageId="21" pageNumber="60" type="reference_group">
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<taxonomicName id="4C61A2F7516CF75595E1FE6EFC91FE73" authority="Cosar, Danisman &amp; Marusik, 2024: 120" authorityName="Cosar, Danisman &amp; Marusik" authorityPageNumber="120" authorityYear="2024" box="[151,871,431,455]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="ilkerakkusi">
<emphasis id="B9150566516CF75595E1FE6EFE6AFE73" box="[151,412,431,455]" italics="true" pageId="21" pageNumber="60">Poecilochroa ilkerakkusi</emphasis>
<materialsCitation id="3B09D329516CF75594DCFE6EFC91FE73" box="[426,871,431,455]" collectingDate="2020-06-29" collectionCode="KUAM" collectorName="Danisman &amp; I. Cosar" country="Turkey" elevation="951" latitude="37.61758" location="Kanliboget" longLatPrecision="1" longitude="36.44625" municipality="Andirin Dist." pageId="21" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Kahramanmaras" typeStatus="holotype">
<bibRefCitation id="EFF0A485516CF75594DCFE6EFC91FE73" author="Cosar, I. &amp; Danisman, T. &amp; Marusik, Y. M." box="[426,871,431,455]" pageId="21" pageNumber="60" pagination="112 - 124" refId="ref18552" refString="Cosar, I., Danisman, T. &amp; Marusik, Y. M. (2024) Two new species belonging to Coreodrassus and Poecilochroa (Aranei: Gnaphosidae) from Anatolia with comments on taxonomy of both genera. Arthropoda Selecta, 33 (1), 112-124. https://doi.org/10.15298/arthsel.33.1.11" type="journal article" year="2024">Coşar, Danişman &amp; Marusik, 2024: 120</bibRefCitation>
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, figs 5CF, 7AE, 8AD (
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<specimenCount id="9D6712FD516CF7559074FE71FAEEFE73" box="[1282,1304,432,455]" count="1" pageId="21" pageNumber="60" type="male"></specimenCount>
:
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<collectingCountry id="F37699E4516CF755905CFE6EFA6EFE73" box="[1322,1432,431,455]" name="Turkey" pageId="21" pageNumber="60">TURKEY</collectingCountry>
:
</emphasis>
<collectingRegion id="49A51796516CF75595B1FE0FFE20FE52" box="[199,470,462,486]" country="Turkey" name="Kahramanmaras" pageId="21" pageNumber="60">Kahramanmaraş Province</collectingRegion>
:
<collectingMunicipality id="6BBA430E516CF7559495FE0FFD84FE52" box="[483,626,462,486]" pageId="21" pageNumber="60">Andirin Dist.</collectingMunicipality>
: Emirler Vill.:
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promenade [
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,
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;
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<quantity id="4C997491516CF755902EFE0FFA6BFE51" box="[1368,1437,462,486]" metricMagnitude="2" metricUnit="m" metricValue="9.51" pageId="21" pageNumber="60" unit="m" value="951.0">951 m</quantity>
a.s.l.
</elevation>
],
<date id="FFDFFFB4516CF755947DFE2CFE77FDB1" box="[267,385,493,517]" pageId="21" pageNumber="60" value="2020-06-29">
<collectingDate id="EF9B065C516CF755947DFE2CFE77FDB1" box="[267,385,493,517]" pageId="21" pageNumber="60" value="2020-06-29">29.VI.2020</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2516CF75594C1FE2DFDE8FDB0" box="[439,542,492,516]" pageId="21" pageNumber="60">Danişman</collectorName>
&amp;
<collectorName id="2694BCA2516CF7559748FE2CFD66FDB1" box="[574,656,493,517]" pageId="21" pageNumber="60">I. Coşar</collectorName>
, KUAM (no registration number specified)—not examined.
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</paragraph>
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<subSubSection id="C37B8AFF516CF75595E1FDF3FBB0FCFD" pageId="21" pageNumber="60" type="description">
<paragraph id="8BDED974516CF75595E1FDF3FC4FFDF9" blockId="21.[151,1436,562,877]" box="[151,953,562,589]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595E1FDF3FEDFFDF8" bold="true" box="[151,297,562,588]" pageId="21" pageNumber="60">Description.</emphasis>
<emphasis id="B9150566516CF7559446FDF2FE9CFDF9" box="[304,362,563,589]" italics="true" pageId="21" pageNumber="60">Male</emphasis>
. For description of the male, see
<bibRefCitation id="EFF0A485516CF75597A9FDF2FC42FDF9" author="Cosar, I. &amp; Danisman, T. &amp; Marusik, Y. M." box="[735,948,563,589]" pageId="21" pageNumber="60" pagination="112 - 124" refId="ref18552" refString="Cosar, I., Danisman, T. &amp; Marusik, Y. M. (2024) Two new species belonging to Coreodrassus and Poecilochroa (Aranei: Gnaphosidae) from Anatolia with comments on taxonomy of both genera. Arthropoda Selecta, 33 (1), 112-124. https://doi.org/10.15298/arthsel.33.1.11" type="journal article" year="2024">
Coşar
<emphasis id="B9150566516CF7559651FDF2FC96FDF9" box="[807,864,563,589]" italics="true" pageId="21" pageNumber="60">et al.</emphasis>
(2024)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED974516CF75595B1FD96FE69FDC5" blockId="21.[151,1436,562,877]" box="[199,415,599,625]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595B1FD96FEEDFDC5" box="[199,283,599,625]" italics="true" pageId="21" pageNumber="60">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BDED974516CF75595B1FDBBFBB0FCFD" blockId="21.[151,1436,562,877]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595B1FDBBFE12FD20" bold="true" box="[199,484,634,660]" pageId="21" pageNumber="60">Justification of transfer.</emphasis>
Although we could not examine the
<typeStatus id="54DA67D6516CF755960BFDBAFC29FD21" box="[893,991,635,661]" pageId="21" pageNumber="60" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C61A2F7516CF7559175FDBAFB6AFD21" authorityName="Cosar, Danisman &amp; Marusik" authorityYear="2024" box="[1027,1180,635,661]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="ilkerakkusi">
<emphasis id="B9150566516CF7559175FDBAFBE2FD20" box="[1027,1044,635,660]" italics="true" pageId="21" pageNumber="60">P</emphasis>
.
<emphasis id="B9150566516CF7559157FDBAFB6AFD21" box="[1057,1180,635,661]" italics="true" pageId="21" pageNumber="60">ilkerakkusi</emphasis>
</taxonomicName>
, the illustrations of the palp of this species are diagnostic, and present all typical features of the male
<taxonomicName id="4C61A2F7516CF7559161FD5EFB0EFD0D" box="[1047,1272,671,697]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF7559161FD5EFB6CFD0D" box="[1047,1178,671,697]" italics="true" pageId="21" pageNumber="60">Trichothyse</emphasis>
species
</taxonomicName>
: distomedially originating broad conductor with prolateral fold, apically originating claw-like embolus with broad embolic base, medially contiguous sperm duct with single loop, and robust RTA with apical bifurcation (cf.
<bibRefCitation id="EFF0A485516CF75591D0FD26FA9EFCB5" author="Cosar, I. &amp; Danisman, T. &amp; Marusik, Y. M." box="[1190,1384,743,769]" pageId="21" pageNumber="60" pagination="112 - 124" refId="ref18552" refString="Cosar, I., Danisman, T. &amp; Marusik, Y. M. (2024) Two new species belonging to Coreodrassus and Poecilochroa (Aranei: Gnaphosidae) from Anatolia with comments on taxonomy of both genera. Arthropoda Selecta, 33 (1), 112-124. https://doi.org/10.15298/arthsel.33.1.11" type="journal article" year="2024">
Coşar
<emphasis id="B9150566516CF7559198FD26FAD0FCB5" box="[1262,1318,743,769]" italics="true" pageId="21" pageNumber="60">et al.</emphasis>
2024
</bibRefCitation>
: figs 5CF, 8AD and
<figureCitation id="135AC5F1516CF7559417FCCAFE30FC91" box="[353,454,779,805]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="21" pageNumber="60">Figs 18</figureCitation>
,
<figureCitation id="135AC5F1516CF75594A5FCCBFDEAFC91" box="[467,540,778,805]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="21" pageNumber="60">5455</figureCitation>
,
<figureCitation id="135AC5F1516CF755975FFCCAFD7AFC91" box="[553,652,779,805]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="21" pageNumber="60">109112</figureCitation>
,
<figureCitation id="135AC5F1516CF75597EFFCCAFD08FC91" box="[665,766,779,805]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="21" pageNumber="60">116117</figureCitation>
). Based on the similarity in genitalic structure and somatic morphology of this species to
<taxonomicName id="4C61A2F7516CF755949FFCEEFD9AFCFD" authorityName="Tucker" authorityYear="1923" box="[489,620,815,841]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B9150566516CF755949FFCEEFD9AFCFD" box="[489,620,815,841]" italics="true" pageId="21" pageNumber="60">Trichothyse</emphasis>
</taxonomicName>
, we propose the transfer of
<taxonomicName id="4C61A2F7516CF75596D3FCEEFBB6FCFD" authorityName="Cosar, Danisman &amp; Marusik" authorityYear="2024" box="[933,1088,815,841]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="ilkerakkusi">
<emphasis id="B9150566516CF75596D3FCEEFC40FCFC" box="[933,950,815,840]" italics="true" pageId="21" pageNumber="60">P</emphasis>
.
<emphasis id="B9150566516CF75596B3FCEEFBB6FCFD" box="[965,1088,815,841]" italics="true" pageId="21" pageNumber="60">ilkerakkusi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516CF75595B1FC93FB66FCD9" box="[199,1168,850,877]" pageId="21" pageNumber="60" type="distribution">
<paragraph id="8BDED974516CF75595B1FC93FB66FCD9" blockId="21.[151,1436,562,877]" box="[199,1168,850,877]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595B1FC93FE97FCD8" bold="true" box="[199,353,850,876]" pageId="21" pageNumber="60">Distribution.</emphasis>
Known only from the
<typeStatus id="54DA67D6516CF7559716FC92FD66FCD9" box="[608,656,851,877]" pageId="21" pageNumber="60">type</typeStatus>
locality in
<collectingCountry id="F37699E4516CF7559679FC92FCA9FCD9" box="[783,863,851,877]" name="Turkey" pageId="21" pageNumber="60">Turkey</collectingCountry>
in western Asia (
<figureCitation id="135AC5F1516CF7559154FC92FB75FCD9" box="[1058,1155,851,877]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="21" pageNumber="60">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,503 @@
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<mods:title id="755907D09DAFCE8BF73BB93D2D0EB7FC">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<subSubSection id="C37B8AFF516CF75595E1FC7BFCFEFC61" box="[151,776,954,981]" pageId="21" pageNumber="60" type="nomenclature">
<paragraph id="8BDED974516CF75595E1FC7BFCFEFC61" blockId="21.[151,776,954,1017]" box="[151,776,954,981]" pageId="21" pageNumber="60">
<heading id="D0966E18516CF75595E1FC7BFCFEFC61" bold="true" box="[151,776,954,981]" fontSize="11" level="1" pageId="21" pageNumber="60" reason="1">
<emphasis id="B9150566516CF75595E1FC7BFCFEFC61" bold="true" box="[151,776,954,981]" pageId="21" pageNumber="60">
<taxonomicName id="4C61A2F7516CF75595E1FC7BFD8CFC60" authority="(Gajbe, 1993)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[151,634,954,981]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B9150566516CF75595E1FC7BFE3FFC61" bold="true" box="[151,457,954,981]" italics="true" pageId="21" pageNumber="60">Trichothyse jodhpurensis</emphasis>
(
<bibRefCitation id="EFF0A485516CF75594AFFC7BFD84FC60" author="Gajbe, U. A." box="[473,626,954,980]" pageId="21" pageNumber="60" pagination="231 - 233" refId="ref19018" refString="Gajbe, U. A. (1993) A new Megamyrmecion spider from India (Araneae: Gnaphosidae). Records of the Zoological Survey of India, 91, 231-233. https://doi.org/10.26515/rzsi/v91/i2/1992/160945" type="journal article" year="1993">Gajbe, 1993</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A226B81D516CF75597F7FC7AFCFEFC61" box="[641,776,955,981]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516CF75595E1FC1EFED7FC4D" box="[151,289,991,1017]" pageId="21" pageNumber="60" type="description">
<paragraph id="8BDED974516CF75595E1FC1EFED7FC4D" blockId="21.[151,776,954,1017]" box="[151,289,991,1017]" pageId="21" pageNumber="60">
<figureCitation id="135AC5F1516CF75595E1FC1EFED7FC4D" box="[151,289,991,1017]" captionStart-0="FIGURES 9099" captionStart-1="FIGURES 100108" captionStart-2="FIGURES 109115" captionStart-3="FIGURES 116119" captionStartId-0="22.[151,264,1875,1899]" captionStartId-1="23.[151,264,1683,1707]" captionStartId-2="24.[151,257,1943,1967]" captionStartId-3="25.[151,264,1479,1503]" captionTargetBox-0="[155,1432,451,1843]" captionTargetBox-1="[155,1431,185,1654]" captionTargetBox-2="[190,1398,195,1913]" captionTargetBox-3="[161,1426,189,1443]" captionTargetId-0="figure-197@22.[151,1436,446,1852]" captionTargetId-1="figure-17@23.[151,1436,179,1657]" captionTargetId-2="figure-17@24.[179,1408,179,1917]" captionTargetId-3="figure-17@25.[151,1436,179,1453]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionText-0="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." captionText-1="FIGURES 100108. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., female from Jaisalmer. 100. Habitus, dorsal view; 101. Same, retrolateral view; 102. Same, frontal view; 103. Eye region, dorsal view; 104. Left chelicera, prolateral view; 105. Same, retrolateral view; 106. Spinnerets, ventral view; 107. Leg I, prolateral view; 108. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 100101 = 1.0 mm; 102105 = 0.1 mm, 106108 = 0.5 mm." captionText-2="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." captionText-3="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="21" pageNumber="60">Figs 90119</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516CF75595E1FBE5FC0CFB23" pageId="21" pageNumber="60" type="reference_group">
<paragraph id="8BDED974516CF75595E1FBE5FB81FBCC" blockId="21.[151,1437,1059,1175]" pageId="21" pageNumber="60">
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<treatmentCitation id="0AC0FF65516CF75595E1FBE5FD76FB8F" author="Gajbe, U. A." box="[151,640,1059,1083]" page="231" pageId="21" pageNumber="60" year="1993">
<taxonomicName id="4C61A2F7516CF75595E1FBE5FD76FB8F" authority="Gajbe, 1993: 231" authorityName="Gajbe" authorityPageNumber="231" authorityYear="1993" box="[151,640,1059,1083]" class="Arachnida" family="Gnaphosidae" genus="Megamyrmecion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="jodhpurensis">
<emphasis id="B9150566516CF75595E1FBE5FE3FFB8F" box="[151,457,1059,1083]" italics="true" pageId="21" pageNumber="60">Megamyrmecion jodhpurensis</emphasis>
<materialsCitation id="3B09D329516CF75594A6FBE2FD76FB8F" box="[464,640,1059,1083]" collectingDate="1962-08-03" collectionCode="NZC-ZSI" collectorName="Motilal &amp; Sankaran &amp; Caleb" country="India" county="Jodhpur" latitude="26.183332" location="Sangriya" longLatPrecision="1243" longitude="73.01667" municipality="Sangariya" pageId="21" pageNumber="60" specimenCount="3" specimenCount-female="3" stateProvince="Rajasthan" typeStatus="holotype">
<bibRefCitation id="EFF0A485516CF75594A6FBE2FD76FB8F" author="Gajbe, U. A." box="[464,640,1059,1083]" pageId="21" pageNumber="60" pagination="231 - 233" refId="ref19018" refString="Gajbe, U. A. (1993) A new Megamyrmecion spider from India (Araneae: Gnaphosidae). Records of the Zoological Survey of India, 91, 231-233. https://doi.org/10.26515/rzsi/v91/i2/1992/160945" type="journal article" year="1993">Gajbe, 1993: 231</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, figs 15 (
<emphasis id="B9150566516CF755979DFBE2FCDAFB8F" box="[747,812,1059,1083]" italics="true" pageId="21" pageNumber="60">lapsus</emphasis>
) (
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<specimenCount id="9D6712FD516CF75596D4FBE5FC43FB8F" box="[930,949,1060,1083]" count="1" pageId="21" pageNumber="60" type="female"></specimenCount>
and
<specimenCount id="9D6712FD516CF7559693FBE2FBF0FB8F" box="[997,1030,1059,1083]" count="2" pageId="21" pageNumber="60" type="female">2♀</specimenCount>
<typeStatus id="54DA67D6516CF755917DFBE5FB86FB88" box="[1035,1136,1060,1084]" pageId="21" pageNumber="60" type="paratype">paratypes</typeStatus>
:
<emphasis id="B9150566516CF755910FFBE2FB3BFB8F" bold="true" box="[1145,1229,1059,1083]" pageId="21" pageNumber="60">
<collectingCountry id="F37699E4516CF755910FFBE2FB31FB8F" box="[1145,1223,1059,1083]" name="India" pageId="21" pageNumber="60">INDIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49A51796516CF75591A5FBE2FACDFB8F" box="[1235,1339,1059,1083]" country="India" name="Rajasthan" pageId="21" pageNumber="60">Rajasthan</collectingRegion>
:
<collectingCounty id="62BFA1F8516CF7559032FBE2FA6CFB8F" box="[1348,1434,1059,1083]" pageId="21" pageNumber="60">Jodhpur</collectingCounty>
:
<collectingMunicipality id="6BBA430E516CF75595B1FB83FEDAFBEE" box="[199,300,1090,1114]" pageId="21" pageNumber="60">Sangariya</collectingMunicipality>
/
<location id="8EBE8FAF516CF7559442FB83FE78FBEE" LSID="urn:lsid:plazi:treatment:03C86862516CF75A95E1FC7BFB65FC7D:8EBE8FAF516CF7559442FB83FE78FBEE" box="[308,398,1090,1114]" country="India" county="Jodhpur" latitude="26.183332" longLatPrecision="1243" longitude="73.01667" municipality="Sangariya" name="Sangriya" pageId="21" pageNumber="60" stateProvince="Rajasthan">Sangriya</location>
[
<geoCoordinate id="EE55BFB3516CF75594E8FB83FE0EFBEE" box="[414,504,1090,1114]" degrees="26" direction="north" minutes="11" orientation="latitude" pageId="21" pageNumber="60" precision="925" value="26.183332">26°11N</geoCoordinate>
,
<geoCoordinate id="EE55BFB3516CF7559770FB83FDA9FBEE" box="[518,607,1090,1114]" degrees="73" direction="east" minutes="01" orientation="longitude" pageId="21" pageNumber="60" precision="925" value="73.01667">73°01E</geoCoordinate>
],
<date id="FFDFFFB4516CF7559705FB83FD1BFBEE" box="[627,749,1090,1114]" pageId="21" pageNumber="60" value="1962-08-03">
<collectingDate id="EF9B065C516CF7559705FB83FD1BFBEE" box="[627,749,1090,1114]" pageId="21" pageNumber="60" value="1962-08-03">3.VIII.1962</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2516CF755965FFB83FC80FBEE" box="[809,886,1090,1114]" pageId="21" pageNumber="60">Motilal</collectorName>
, NZC-ZSI (no register number specified), examined (
<typeStatus id="54DA67D6516CF75595B8FBA1FEDEFBCC" box="[206,296,1120,1144]" pageId="21" pageNumber="60" type="holotype">holotype</typeStatus>
and
<typeStatus id="54DA67D6516CF755942CFBA0FE47FBCD" box="[346,433,1121,1145]" pageId="21" pageNumber="60" type="paratype">paratype</typeStatus>
images are provided in
<collectorName id="2694BCA2516CF75597D3FBA1FCF5FBCC" box="[677,771,1120,1144]" pageId="21" pageNumber="60">Sankaran</collectorName>
&amp;
<collectorName id="2694BCA2516CF7559652FBA1FCA9FBCC" box="[804,863,1120,1144]" pageId="21" pageNumber="60">Caleb</collectorName>
(2021)—not re-examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED974516CF75595E1FBBEFC0CFB23" blockId="21.[151,1437,1059,1175]" box="[151,1018,1151,1175]" pageId="21" pageNumber="60">
<taxonomicName id="4C61A2F7516CF75595E1FBBEFE0FFB23" ID-CoL="737VV" authority="(Gajbe)" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[151,505,1151,1175]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="jodhpurense">
<emphasis id="B9150566516CF75595E1FBBEFE57FB23" box="[151,417,1151,1175]" italics="true" pageId="21" pageNumber="60">Poecilochroa jodhpurense</emphasis>
(Gajbe)
</taxonomicName>
: Sankaran &amp; Caleb, 2021: 548, figs 4446, 4951.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516CF75595E1FB04FAC4FADB" pageId="21" pageNumber="60" type="materials_examined">
<paragraph id="8BDED974516CF75595E1FB04FAC4FADB" blockId="21.[151,1437,1221,2040]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595E1FB04FE88FB6B" bold="true" box="[151,382,1221,1247]" pageId="21" pageNumber="60">Material examined.</emphasis>
<materialsCitation id="3B09D329516CF75594F4FB04FAD8FADB" collectingDate="2018-04-30" collectingDateMax="2018-05-15" collectingDateMin="2018-04-30" collectionCode="ZSI" collectorName="R. Tripathi &amp; A. K. Jangid" country="India" county="Jaisalmer" elevation="205" latitude="26.7525" location="Desert National Park Wildlife Sanctuary" longLatPrecision="20" longitude="71.42777" municipality="Thar Desert" pageId="21" pageNumber="60" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Rajasthan">
<emphasis id="B9150566516CF75594F4FB04FD97FB6B" bold="true" box="[386,609,1221,1248]" pageId="21" pageNumber="60">
<collectingCountry id="F37699E4516CF75594F4FB04FE2FFB6B" box="[386,473,1221,1247]" name="India" pageId="21" pageNumber="60">INDIA</collectingCountry>
:
<emphasis id="B9150566516CF7559494FB04FDAEFB54" bold="true" box="[482,600,1221,1248]" italics="true" pageId="21" pageNumber="60">
<collectingRegion id="49A51796516CF7559494FB04FDAEFB54" box="[482,600,1221,1248]" country="India" name="Rajasthan" pageId="21" pageNumber="60">Rajasthan</collectingRegion>
</emphasis>
:
</emphasis>
<collectingCounty id="62BFA1F8516CF7559710FB04FD24FB6B" box="[614,722,1221,1247]" pageId="21" pageNumber="60">Jaisalmer</collectingCounty>
,
<collectingMunicipality id="6BBA430E516CF75597AFFB04FCA8FB54" box="[729,862,1221,1248]" pageId="21" pageNumber="60">Thar Desert</collectingMunicipality>
,
<location id="8EBE8FAF516CF7559610FB07FAD7FB6B" LSID="urn:lsid:plazi:treatment:03C86862516CF75A95E1FC7BFB65FC7D:8EBE8FAF516CF7559610FB07FAD7FB6B" box="[870,1313,1221,1248]" country="India" county="Jaisalmer" latitude="26.7525" longLatPrecision="20" longitude="71.42777" municipality="Thar Desert" name="Desert National Park Wildlife Sanctuary" pageId="21" pageNumber="60" stateProvince="Rajasthan">
<location id="8EBE8FAF516CF7559610FB07FBBDFB6B" LSID="urn:lsid:plazi:treatment:03C86862516CF75A95E1FC7BFB65FC7D:8EBE8FAF516CF7559610FB07FBBDFB6B" box="[870,1099,1221,1248]" country="India" county="Jaisalmer" latitude="26.7525" longLatPrecision="20" longitude="71.42777" municipality="Thar Desert" name="Desert National Park" pageId="21" pageNumber="60" stateProvince="Rajasthan">Desert National Park</location>
Wildlife Sanctuary
</location>
,
<location id="8EBE8FAF516CF755905FFB04FF30FAB0" LSID="urn:lsid:plazi:treatment:03C86862516CF75A95E1FC7BFB65FC7D:8EBE8FAF516CF755905FFB04FF30FAB0" country="India" county="Jaisalmer" latitude="26.7525" longLatPrecision="20" longitude="71.42777" municipality="Thar Desert" name="Gajaimata area" pageId="21" pageNumber="60" stateProvince="Rajasthan">Gajaimata area</location>
(
<geoCoordinate id="EE55BFB3516CF75595A0FB28FE90FAB7" box="[214,358,1257,1283]" degrees="26" direction="north" minutes="40" orientation="latitude" pageId="21" pageNumber="60" precision="15" seconds="28" value="26.674444">26°4028N</geoCoordinate>
,
<geoCoordinate id="EE55BFB3516CF7559404FB28FE09FAB7" box="[370,511,1257,1283]" degrees="70" direction="east" minutes="34" orientation="longitude" pageId="21" pageNumber="60" precision="15" seconds="17" value="70.57139">70°3417E</geoCoordinate>
,
<elevation id="004C3E47516CF755977DFB28FD79FAB7" box="[523,655,1257,1284]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="21" pageNumber="60" unit="m" value="205.0">
<quantity id="4C997491516CF755977DFB28FDA5FAB0" box="[523,595,1257,1284]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="21" pageNumber="60" unit="m" value="205.0">205 m</quantity>
a.s.l.
</elevation>
),
<date id="FFDFFFB4516CF75597D0FB28FCD3FAB7" box="[678,805,1257,1283]" pageId="21" pageNumber="60" value="2018-04-30">
<collectingDate id="EF9B065C516CF75597D0FB28FCD3FAB7" box="[678,805,1257,1283]" pageId="21" pageNumber="60" value="2018-04-30">30.IV.2018</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2516CF7559617FB2BFC2DFAB7" box="[865,987,1257,1284]" pageId="21" pageNumber="60">R. Tripathi</collectorName>
&amp;
<collectorName id="2694BCA2516CF7559688FB2BFB73FAB7" box="[1022,1157,1257,1284]" pageId="21" pageNumber="60">A.K. Jangid</collectorName>
(from ground, by hand),
<specimenCount id="9D6712FD516CF75595E1FACCFF4CFA93" box="[151,186,1293,1319]" count="1" pageId="21" pageNumber="60" type="male">1♂</specimenCount>
(
<collectionCode id="ED7041B1516CF75595BCFACCFF03FA93" box="[202,245,1293,1319]" pageId="21" pageNumber="60">ZSI</collectionCode>
/WGRC/IR.-INV.27841); Same data as previous except
<location id="8EBE8FAF516CF7559602FACCFBFAFA9C" LSID="urn:lsid:plazi:treatment:03C86862516CF75A95E1FC7BFB65FC7D:8EBE8FAF516CF7559602FACCFBFAFA9C" box="[884,1036,1293,1320]" country="India" county="Jaisalmer" latitude="26.7525" longLatPrecision="20" longitude="71.42777" municipality="Thar Desert" name="Sudasari area" pageId="21" pageNumber="60" stateProvince="Rajasthan">Sudasari area</location>
(
<geoCoordinate id="EE55BFB3516CF755916AFACCFB5AFA93" box="[1052,1196,1293,1319]" degrees="26" direction="north" minutes="40" orientation="latitude" pageId="21" pageNumber="60" precision="15" seconds="07" value="26.66861">26°4007N</geoCoordinate>
,
<geoCoordinate id="EE55BFB3516CF75591CCFACCFAB0FA93" box="[1210,1350,1293,1319]" degrees="70" direction="east" minutes="36" orientation="longitude" pageId="21" pageNumber="60" precision="15" seconds="31" value="70.60861">70°3631E</geoCoordinate>
,
<elevation id="004C3E47516CF7559022FACCFF3AFAFF" metricMagnitude="2" metricUnit="m" metricValue="2.19" pageId="21" pageNumber="60" unit="m" value="219.0">
<quantity id="4C997491516CF7559022FACCFA6AFA9C" box="[1364,1436,1293,1320]" metricMagnitude="2" metricUnit="m" metricValue="2.19" pageId="21" pageNumber="60" unit="m" value="219.0">219 m</quantity>
a.s.l.
</elevation>
),
<date id="FFDFFFB4516CF7559597FAF0FEBFFAFF" box="[225,329,1329,1355]" pageId="21" pageNumber="60" value="2018-05-08">
<collectingDate id="EF9B065C516CF7559597FAF0FEBFFAFF" box="[225,329,1329,1355]" pageId="21" pageNumber="60" value="2018-05-08">8.V.2018</collectingDate>
</date>
,
<specimenCount id="9D6712FD516CF7559425FAF0FE83FAFF" box="[339,373,1329,1355]" count="2" pageId="21" pageNumber="60" type="male">2♂</specimenCount>
<specimenCount id="9D6712FD516CF755940AFAF0FE68FAFF" box="[380,414,1329,1355]" count="2" pageId="21" pageNumber="60" type="female">2♀</specimenCount>
(
<collectionCode id="ED7041B1516CF75594DAFAF0FE21FAFF" box="[428,471,1329,1355]" pageId="21" pageNumber="60">ZSI</collectionCode>
/WGRC/IR.-INV.27842, 27843, 27844 and 27845 respectively); Same data as previous except
<location id="8EBE8FAF516CF7559591FA94FEADFAC4" LSID="urn:lsid:plazi:treatment:03C86862516CF75A95E1FC7BFB65FC7D:8EBE8FAF516CF7559591FA94FEADFAC4" box="[231,347,1365,1392]" country="India" county="Jaisalmer" latitude="26.7525" longLatPrecision="20" longitude="71.42777" municipality="Thar Desert" name="Rasla area" pageId="21" pageNumber="60" stateProvince="Rajasthan">Rasla area</location>
(
<geoCoordinate id="EE55BFB3516CF755941DFA94FE0DFAC4" box="[363,507,1365,1392]" degrees="26" direction="north" minutes="45" orientation="latitude" pageId="21" pageNumber="60" precision="15" seconds="09" value="26.7525">26°4509N</geoCoordinate>
,
<geoCoordinate id="EE55BFB3516CF7559771FA94FD65FAC4" box="[519,659,1365,1392]" degrees="71" direction="east" minutes="25" orientation="longitude" pageId="21" pageNumber="60" precision="15" seconds="40" value="71.42777">71°2540E</geoCoordinate>
,
<elevation id="004C3E47516CF75597E9FA94FCD7FADB" box="[671,801,1365,1392]" metricMagnitude="2" metricUnit="m" metricValue="2.69" pageId="21" pageNumber="60" unit="m" value="269.0">
<quantity id="4C997491516CF75597E9FA94FD10FAC4" box="[671,742,1365,1392]" metricMagnitude="2" metricUnit="m" metricValue="2.69" pageId="21" pageNumber="60" unit="m" value="269.0">269 m</quantity>
a.s.l.
</elevation>
),
<date id="FFDFFFB4516CF755964EFA94FC58FAC4" box="[824,942,1365,1392]" pageId="21" pageNumber="60" value="2018-05-15">
<collectingDate id="EF9B065C516CF755964EFA94FC58FAC4" box="[824,942,1365,1392]" pageId="21" pageNumber="60" value="2018-05-15">15.V.2018</collectingDate>
</date>
,
<specimenCount id="9D6712FD516CF75596CFFA94FC2AFADB" box="[953,988,1365,1391]" count="1" pageId="21" pageNumber="60" type="female">1♀</specimenCount>
(
<collectionCode id="ED7041B1516CF755969CFA94FBE3FADB" box="[1002,1045,1365,1391]" pageId="21" pageNumber="60">ZSI</collectionCode>
/WGRC/IR.-INV.27846)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516CF75595B1FAB8FCA6F93B" pageId="21" pageNumber="60" type="diagnosis">
<paragraph id="8BDED974516CF75595B1FAB8FCA6F93B" blockId="21.[151,1437,1221,2040]" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595B1FAB8FEB4FA27" bold="true" box="[199,322,1401,1427]" pageId="21" pageNumber="60">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C61A2F7516CF75594CEFABBFD91FA27" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[440,615,1401,1428]" class="Arachnida" family="Gnaphosidae" genus="Megamyrmecion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B9150566516CF75594CEFABBFE3EFA27" box="[440,456,1402,1427]" italics="true" pageId="21" pageNumber="60">T</emphasis>
.
<emphasis id="B9150566516CF75594A1FAB8FD91FA27" box="[471,615,1401,1427]" italics="true" pageId="21" pageNumber="60">jodhpurensis</emphasis>
</taxonomicName>
<emphasis id="B9150566516CF7559706FAB8FD04FA27" bold="true" box="[624,754,1401,1427]" pageId="21" pageNumber="60">
<taxonomicNameLabel id="A226B81D516CF7559706FAB8FD04FA27" box="[624,754,1401,1427]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C61A2F7516CF7559689FABBFB9AFA27" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Sankaran &amp; Haddad &amp; Tripathi" baseAuthorityYear="2025" box="[1023,1132,1401,1427]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="furcata" status="comb. nov.">
<emphasis id="B9150566516CF7559689FABBFB9AFA27" box="[1023,1132,1401,1427]" italics="true" pageId="21" pageNumber="60">T. furcata</emphasis>
</taxonomicName>
<emphasis id="B9150566516CF7559103FAB8FB01FA27" bold="true" box="[1141,1271,1401,1427]" pageId="21" pageNumber="60">
<taxonomicNameLabel id="A226B81D516CF7559103FAB8FB01FA27" box="[1141,1271,1401,1427]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
in the general aspects of its palp, but can be distinguished from the latter by the short, stout prolateral and globular retrolateral branches of the RTA in retrolateral view (
<emphasis id="B9150566516CF7559709FA03FD60FA6F" box="[639,662,1474,1499]" italics="true" pageId="21" pageNumber="60">vs</emphasis>
. long, narrow prolateral and thumb-like retrolateral branches in
<taxonomicName id="4C61A2F7516CF75590F0FA03FF11FA4B" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Sankaran &amp; Haddad &amp; Tripathi" baseAuthorityYear="2025" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="furcata" status="comb. nov.">
<emphasis id="B9150566516CF75590F0FA03FA60FA6F" box="[1414,1430,1474,1499]" italics="true" pageId="21" pageNumber="60">T</emphasis>
.
<emphasis id="B9150566516CF75595E1FA24FF11FA4B" box="[151,231,1509,1535]" italics="true" pageId="21" pageNumber="60">furcata</emphasis>
</taxonomicName>
<emphasis id="B9150566516CF7559599FA24FE86FA4B" bold="true" box="[239,368,1509,1535]" pageId="21" pageNumber="60">
<taxonomicNameLabel id="A226B81D516CF7559599FA24FE86FA4B" box="[239,368,1509,1535]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
) (cf.
<figureCitation id="135AC5F1516CF75594D9FA24FDBCFA4B" box="[431,586,1509,1535]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="21" pageNumber="60">Figs 109112</figureCitation>
,
<figureCitation id="135AC5F1516CF7559721FA24FD40FA4B" box="[599,694,1509,1535]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="21" pageNumber="60">116117</figureCitation>
and
<bibRefCitation id="EFF0A485516CF7559799FA24FC52FA4B" author="Di Franco, F." box="[751,932,1509,1536]" pageId="21" pageNumber="60" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco 2001</bibRefCitation>
: figs 34). Females are similar to those of
<taxonomicName id="4C61A2F7516CF75590FEFA27FF1FF997" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Sankaran &amp; Haddad &amp; Tripathi" baseAuthorityYear="2025" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="pugnax" status="comb. nov.">
<emphasis id="B9150566516CF75590FEFA27FF1FF997" italics="true" pageId="21" pageNumber="60">T. pugnax</emphasis>
</taxonomicName>
<emphasis id="B9150566516CF7559584F9C8FE85F997" bold="true" box="[242,371,1545,1571]" pageId="21" pageNumber="60">
<taxonomicNameLabel id="A226B81D516CF7559584F9C8FE85F997" box="[242,371,1545,1571]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, as both have an epigynal posterior border line with a median notch and bean-shaped primary spermathecae, but can be distinguished from the latter by angular anterior part of the median extension (
<emphasis id="B9150566516CF755905CF9EFFAB7F9F3" box="[1322,1345,1582,1607]" italics="true" pageId="21" pageNumber="60">vs</emphasis>
. widely oval in
<taxonomicName id="4C61A2F7516CF755959BF993FEADF9DF" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Sankaran &amp; Haddad &amp; Tripathi" baseAuthorityYear="2025" box="[237,347,1618,1643]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="pugnax" status="comb. nov.">
<emphasis id="B9150566516CF755959BF993FEADF9DF" box="[237,347,1618,1643]" italics="true" pageId="21" pageNumber="60">T. pugnax</emphasis>
</taxonomicName>
<emphasis id="B9150566516CF7559412F990FE13F9DF" bold="true" box="[356,485,1617,1643]" pageId="21" pageNumber="60">
<taxonomicNameLabel id="A226B81D516CF7559412F990FE13F9DF" box="[356,485,1617,1643]" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
), and the secondary spermathecae with oval heads (
<emphasis id="B9150566516CF7559142F993FBBDF9DF" box="[1076,1099,1618,1643]" italics="true" pageId="21" pageNumber="60">vs</emphasis>
. globular in
<taxonomicName id="4C61A2F7516CF75591ABF993FABAF9DF" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Sankaran &amp; Haddad &amp; Tripathi" baseAuthorityYear="2025" box="[1245,1356,1618,1643]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="60" phylum="Arthropoda" rank="species" species="pugnax" status="comb. nov.">
<emphasis id="B9150566516CF75591ABF993FABAF9DF" box="[1245,1356,1618,1643]" italics="true" pageId="21" pageNumber="60">T. pugnax</emphasis>
</taxonomicName>
<emphasis id="B9150566516CF7559022F990FF3EF93B" bold="true" pageId="21" pageNumber="60">
<taxonomicNameLabel id="A226B81D516CF7559022F990FF3EF93B" pageId="21" pageNumber="60" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
) (cf.
<figureCitation id="135AC5F1516CF7559470F9B4FE56F924" box="[262,416,1653,1680]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="21" pageNumber="60">Figs 113115</figureCitation>
,
<figureCitation id="135AC5F1516CF75594DDF9B4FDFDF93B" box="[427,523,1653,1679]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="21" pageNumber="60">118119</figureCitation>
and
<bibRefCitation id="EFF0A485516CF7559737F9B7FD48F93B" author="Levy, G." box="[577,702,1653,1680]" pageId="21" pageNumber="60" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: figs 1314).
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516CF75A95B1F958FA9AFC11" lastPageId="26" lastPageNumber="65" pageId="21" pageNumber="60" type="description">
<paragraph id="8BDED974516CF75695B1F958FDE9FE1F" blockId="21.[151,1437,1221,2040]" lastBlockId="22.[151,1437,150,428]" lastPageId="22" lastPageNumber="61" pageId="21" pageNumber="60">
<emphasis id="B9150566516CF75595B1F958FEAFF907" bold="true" box="[199,345,1689,1715]" pageId="21" pageNumber="60">Description.</emphasis>
<emphasis id="B9150566516CF7559414F958FE6AF907" box="[354,412,1689,1715]" italics="true" pageId="21" pageNumber="60">Male</emphasis>
(Gajaimata area, ZSI/WGRC/IR.-INV.27841,
<figureCitation id="135AC5F1516CF75596DBF958FBC7F907" box="[941,1073,1689,1715]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Figs 9099</figureCitation>
). Carapace, clypeus, chelicerae, labium, endites, sternum, scutum and legs brown, eye field slightly darker than carapace, abdomen ash-grey with indistinct median longitudinal black stripe, spinnerets olive brown; carapace with faintly darker striae radiating from fovea, densely covered with white appressed hairs, cephalic part provided with few stiff black setae medially (
<figureCitation id="135AC5F1516CF75595E9F8E8FF00F8F7" box="[159,246,1833,1859]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 94</figureCitation>
). Fovea narrow, longitudinal, straight, red-brown (
<figureCitation id="135AC5F1516CF755964CF8E8FC67F8F7" box="[826,913,1833,1859]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 90</figureCitation>
). Cheliceral promargin with long black setae, with low keel and denticle (
<figureCitation id="135AC5F1516CF75594A1F88CFDDAF8DC" box="[471,556,1869,1896]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 95</figureCitation>
; arrow); retromargin without long setae, keel or denticle (
<figureCitation id="135AC5F1516CF75591B2F88CFAECF8D3" box="[1220,1306,1869,1895]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 96</figureCitation>
). Abdomen oval, hirsute (
<figureCitation id="135AC5F1516CF7559444F8B0FE71F83F" box="[306,391,1905,1931]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 90</figureCitation>
); dorsal abdomen anteriorly with short, inverted triangular-shaped scutum, with four distinct sigilla, two circular and remaining two rice grain-shaped (
<figureCitation id="135AC5F1516CF7559654F854FC81F81B" box="[802,887,1941,1967]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 90</figureCitation>
). Metatarsi III and all tarsi with well-developed scopulae, scopulae on tarsi complete, those on metatarsi restricted distally (
<figureCitation id="135AC5F1516CF7559681F878FB8BF867" box="[1015,1149,1977,2003]" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Figs 9899</figureCitation>
). Spinnerets hirsute (
<figureCitation id="135AC5F1516CF7559019F878FF41F843" captionStart="FIGURES 9099" captionStartId="22.[151,264,1875,1899]" captionTargetBox="[155,1432,451,1843]" captionTargetId="figure-197@22.[151,1436,446,1852]" captionTargetPageId="22" captionText="FIGURES 9099. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view.Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm." pageId="21" pageNumber="60">Fig. 97</figureCitation>
). Body length 3.72. Carapace 1.55 long, 1.11 wide. Abdomen 2.14 long, 1.23 wide. Eye sizes and interdistances: ALE 0.07, AME 0.12, PLE 0.08, PME 0.08; ALEPLE 0.05, AMEAME 0.05, AMEPME 0.10, PMEPLE 0.03, PMEPME 0.06. AME and ALE nearly contiguous. Clypeus height at AMEs 0.09, at ALEs 0.08. Chelicerae 0.44 long. Length of palp and legs: palp 1.28 [0.51, 0.18, 0.16, 0.43], I 3.18 [0.99, 0.48, 0.74, 0.54, 0.43], II 2.92 [0.92, 0.39, 0.68, 0.54, 0.39], III 3.03 [0.91, 0.40, 0.60, 0.66, 0.46], IV 3.94 [1.10, 0.50, 0.85, 1.01, 0.48]. Leg formula: 4132. Spination of palp: femur pld 1 do 2, patella pld 1, tibia rld 1, tarsus/cymbium pl 1 rl 2; legs: femur III pl 1 do 2, IIIIV pld 2 do 3 rld 2; patella III spineless, IIIIV pld 1 rld 1; tibia III plv 2 rlv 1, III pl 2 rl 3 plv 2 rlv 2, IV pl 1 rl 2 pld 1 rld 2 plv 3 rlv 3; metatarsus III plv 1 rlv 1, III pl 2 rl 1 pld 3 rld 2 plv 2 rlv 2, IV pl 1 rl 2 pld 3 rld 3 plv 3 rlv 3; tarsus IIV spineless.
</paragraph>
<caption id="DF1E89FC516FF75695E1F892FD7CF84D" pageId="22" pageNumber="61" startId="22.[151,264,1875,1899]" targetBox="[155,1432,451,1843]" targetPageId="22" targetType="figure">
<paragraph id="8BDED974516FF75695E1F892FD7CF84D" blockId="22.[151,1437,1875,2041]" pageId="22" pageNumber="61">
<emphasis id="B9150566516FF75695E1F892FEAEF8DF" bold="true" box="[151,344,1875,1899]" pageId="22" pageNumber="61">FIGURES 9099.</emphasis>
Somatic morphology of
<taxonomicName id="4C61A2F7516FF7569713F892FBF5F8DF" authority="(Gajbe, 1993)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[613,1027,1875,1899]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="61" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B9150566516FF7569713F892FC9EF8DF" box="[613,872,1875,1899]" italics="true" pageId="22" pageNumber="61">Trichothyse jodhpurensis</emphasis>
(
<bibRefCitation id="EFF0A485516FF756960FF892FC0AF8DF" author="Gajbe, U. A." box="[889,1020,1875,1899]" pageId="22" pageNumber="61" pagination="231 - 233" refId="ref19018" refString="Gajbe, U. A. (1993) A new Megamyrmecion spider from India (Araneae: Gnaphosidae). Records of the Zoological Survey of India, 91, 231-233. https://doi.org/10.26515/rzsi/v91/i2/1992/160945" type="journal article" year="1993">Gajbe, 1993</bibRefCitation>
)
</taxonomicName>
<emphasis id="B9150566516FF756917BF892FB73F8DF" bold="true" box="[1037,1157,1875,1899]" pageId="22" pageNumber="61">
<taxonomicNameLabel id="A226B81D516FF756917BF892FB73F8DF" box="[1037,1157,1875,1899]" pageId="22" pageNumber="61" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, male from Jaisalmer. 90. Habitus, dorsal view; 91. Same, retrolateral view; 92. Same, frontal view; 93. Eye region, dorsal view; 94. Carapace showing row of setae, dorsal view; 95. Left chelicera, prolateral view; 96. Same, retrolateral view; 97. Spinnerets, ventral view; 98. Leg I, retrolateral view; 99. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 9091 = 1.0 mm; 9296 = 0.1 mm; 97 = 0.2 mm; 9899 = 0.5 mm.
</paragraph>
</caption>
<caption id="DF1E89FC516EF75795E1F952FF30F88F" pageId="23" pageNumber="62" startId="23.[151,264,1683,1707]" targetBox="[155,1431,185,1654]" targetPageId="23" targetType="figure">
<paragraph id="8BDED974516EF75795E1F952FF30F88F" blockId="23.[151,1437,1683,1851]" pageId="23" pageNumber="62">
<emphasis id="B9150566516EF75795E1F952FE9BF91F" bold="true" box="[151,365,1683,1707]" pageId="23" pageNumber="62">FIGURES 100108.</emphasis>
Somatic morphology of
<taxonomicName id="4C61A2F7516EF757971CF952FC0DF91F" authority="(Gajbe, 1993)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[618,1019,1683,1707]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="62" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B9150566516EF757971CF952FC9FF91F" box="[618,873,1683,1707]" italics="true" pageId="23" pageNumber="62">Trichothyse jodhpurensis</emphasis>
(
<bibRefCitation id="EFF0A485516EF7579600F952FC02F91F" author="Gajbe, U. A." box="[886,1012,1683,1707]" pageId="23" pageNumber="62" pagination="231 - 233" refId="ref19018" refString="Gajbe, U. A. (1993) A new Megamyrmecion spider from India (Araneae: Gnaphosidae). Records of the Zoological Survey of India, 91, 231-233. https://doi.org/10.26515/rzsi/v91/i2/1992/160945" type="journal article" year="1993">Gajbe, 1993</bibRefCitation>
)
</taxonomicName>
<emphasis id="B9150566516EF7579174F952FB83F91F" bold="true" box="[1026,1141,1683,1707]" pageId="23" pageNumber="62">
<taxonomicNameLabel id="A226B81D516EF7579174F952FB83F91F" box="[1026,1141,1683,1707]" pageId="23" pageNumber="62" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, female from Jaisalmer. 100. Habitus, dorsal view; 101. Same, retrolateral view; 102. Same, frontal view; 103. Eye region, dorsal view; 104. Left chelicera, prolateral view; 105. Same, retrolateral view; 106. Spinnerets, ventral view; 107. Leg I, prolateral view; 108. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 100101 = 1.0 mm; 102105 = 0.1 mm, 106108 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDED974516EF75995B1F84CFBC2F8F3" blockId="23.[151,1436,1933,2032]" lastBlockId="25.[151,1437,1729,2008]" lastPageId="25" lastPageNumber="64" pageId="23" pageNumber="62">
<emphasis id="B9150566516EF75795B1F84CFF0AF813" box="[199,252,1933,1959]" italics="true" pageId="23" pageNumber="62">Palp</emphasis>
(
<figureCitation id="135AC5F1516EF757947FF84CFE55F81C" box="[265,419,1933,1960]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="23" pageNumber="62">Figs 109112</figureCitation>
,
<figureCitation id="135AC5F1516EF75794DAF84CFDE7F813" box="[428,529,1933,1959]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="23" pageNumber="62">116117</figureCitation>
): segments brown. RTA rod-shaped, nearly half the length of cymbium, with small, stout prolateral and roughly globular retrolateral branches (
<figureCitation id="135AC5F1516EF7579659F870FC3AF878" box="[815,972,1969,1996]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="23" pageNumber="62">Figs 109112</figureCitation>
,
<figureCitation id="135AC5F1516EF75796AEF870FBCAF87F" box="[984,1084,1969,1995]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="23" pageNumber="62">116117</figureCitation>
). Cymbial flange nearly half of length of cymbium, anteriorly with serration-like modification (
<figureCitation id="135AC5F1516EF7579603F814FC2BF844" box="[885,989,2005,2032]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="23" pageNumber="62">Figs 109</figureCitation>
,
<figureCitation id="135AC5F1516EF757969AF814FBEFF85B" box="[1004,1049,2005,2031]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="23" pageNumber="62">116</figureCitation>
). Tegulum elongate-oval, sac-like (
<figureCitation id="135AC5F15160F75995E9F900FEF5F96F" box="[159,259,1729,1755]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="25" pageNumber="64">Figs 109</figureCitation>
,
<figureCitation id="135AC5F15160F7599479F900FECAF96F" box="[271,316,1729,1755]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="25" pageNumber="64">116</figureCitation>
). Conductor membranous, with slight median curvature, flat distal part wrapping embolus, proximal part lying parallel to embolus, and smoothly rounded apex directed at 2-o clock ventrally (
<figureCitation id="135AC5F15160F75991F0F924FAD7F94B" box="[1158,1313,1765,1791]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="25" pageNumber="64">Figs 109110</figureCitation>
,
<figureCitation id="135AC5F15160F759905AF924FA66F94B" box="[1324,1424,1765,1791]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="25" pageNumber="64">116117</figureCitation>
). Embolus arising apically, narrow, claw-like, lying adjacent to conductor, slightly curved retrolaterally, with tip directed at 1-o clock ventrally, with broad embolic base (
<figureCitation id="135AC5F15160F759966AF8ECFC4EF8F3" box="[796,952,1837,1863]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="25" pageNumber="64">Figs 109110</figureCitation>
,
<figureCitation id="135AC5F15160F75996B5F8ECFBDEF8F3" box="[963,1064,1837,1863]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="25" pageNumber="64">116117</figureCitation>
).
</paragraph>
<caption id="DF1E89FC5161F75895E1F856FA6CF843" pageId="24" pageNumber="63" startId="24.[151,257,1943,1967]" targetBox="[190,1398,195,1913]" targetPageId="24" targetType="figure">
<paragraph id="8BDED9745161F75895E1F856FA6CF843" blockId="24.[151,1437,1943,2040]" pageId="24" pageNumber="63">
<emphasis id="B91505665161F75895E1F856FE94F81B" bold="true" box="[151,354,1943,1967]" pageId="24" pageNumber="63">FIGURES 109115.</emphasis>
Genitalic morphology of
<taxonomicName id="4C61A2F75161F7589711F856FC1DF804" authority="(Gajbe, 1993)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[615,1003,1943,1968]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B91505665161F7589711F856FCACF81B" box="[615,858,1943,1967]" italics="true" pageId="24" pageNumber="63">Trichothyse jodhpurensis</emphasis>
(
<bibRefCitation id="EFF0A4855161F758961CF856FC12F804" author="Gajbe, U. A." box="[874,996,1943,1968]" pageId="24" pageNumber="63" pagination="231 - 233" refId="ref19018" refString="Gajbe, U. A. (1993) A new Megamyrmecion spider from India (Araneae: Gnaphosidae). Records of the Zoological Survey of India, 91, 231-233. https://doi.org/10.26515/rzsi/v91/i2/1992/160945" type="journal article" year="1993">Gajbe, 1993</bibRefCitation>
)
</taxonomicName>
<emphasis id="B91505665161F7589682F859FB92F804" bold="true" box="[1012,1124,1944,1968]" pageId="24" pageNumber="63">
<taxonomicNameLabel id="A226B81D5161F7589682F859FB92F804" box="[1012,1124,1944,1968]" pageId="24" pageNumber="63" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="DF1E89FC5160F75995E1FA06FBEFF9DB" pageId="25" pageNumber="64" startId="25.[151,264,1479,1503]" targetBox="[161,1426,189,1443]" targetPageId="25" targetType="figure">
<paragraph id="8BDED9745160F75995E1FA06FBEFF9DB" blockId="25.[151,1437,1479,1647]" pageId="25" pageNumber="64">
<emphasis id="B91505665160F75995E1FA06FE9AFA6B" bold="true" box="[151,364,1479,1503]" pageId="25" pageNumber="64">FIGURES 116119.</emphasis>
Genitalic morphology of
<taxonomicName id="4C61A2F75160F759970CFA06FBE7FA6B" authority="(Gajbe, 1993)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[634,1041,1479,1503]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="64" phylum="Arthropoda" rank="species" species="jodhpurensis" status="comb. nov.">
<emphasis id="B91505665160F759970CFA06FC8DFA6B" box="[634,891,1479,1503]" italics="true" pageId="25" pageNumber="64">Trichothyse jodhpurensis</emphasis>
(
<bibRefCitation id="EFF0A4855160F75996FCFA06FBFCFA6B" author="Gajbe, U. A." box="[906,1034,1479,1503]" pageId="25" pageNumber="64" pagination="231 - 233" refId="ref19018" refString="Gajbe, U. A. (1993) A new Megamyrmecion spider from India (Araneae: Gnaphosidae). Records of the Zoological Survey of India, 91, 231-233. https://doi.org/10.26515/rzsi/v91/i2/1992/160945" type="journal article" year="1993">Gajbe, 1993</bibRefCitation>
)
</taxonomicName>
<emphasis id="B91505665160F759916FFA06FB79FA6B" bold="true" box="[1049,1167,1479,1503]" pageId="25" pageNumber="64">
<taxonomicNameLabel id="A226B81D5160F759916FFA06FB79FA6B" box="[1049,1167,1479,1503]" pageId="25" pageNumber="64" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BDED9745160F75A95B1F890FE4CFE3D" blockId="25.[151,1437,1729,2008]" lastBlockId="26.[151,1437,151,969]" lastPageId="26" lastPageNumber="65" pageId="25" pageNumber="64">
<emphasis id="B91505665160F75995B1F890FEEDF8DF" box="[199,283,1873,1899]" italics="true" pageId="25" pageNumber="64">Female</emphasis>
(Sudasari area, ZSI/WGRC/IR.-INV.27844,
<figureCitation id="135AC5F15160F7599653F890FC3EF8DF" box="[805,968,1873,1899]" captionStart="FIGURES 100108" captionStartId="23.[151,264,1683,1707]" captionTargetBox="[155,1431,185,1654]" captionTargetId="figure-17@23.[151,1436,179,1657]" captionTargetPageId="23" captionText="FIGURES 100108. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., female from Jaisalmer. 100. Habitus, dorsal view; 101. Same, retrolateral view; 102. Same, frontal view; 103. Eye region, dorsal view; 104. Left chelicera, prolateral view; 105. Same, retrolateral view; 106. Spinnerets, ventral view; 107. Leg I, prolateral view; 108. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 100101 = 1.0 mm; 102105 = 0.1 mm, 106108 = 0.5 mm." pageId="25" pageNumber="64">Figs 100108</figureCitation>
). General aspects essentially as in male except the following: habitus colour lighter than male. Carapace with randomly arranged black stiff setae. Dorsal abdomen without scutum, with six sigilla (
<figureCitation id="135AC5F15160F759970DF858FD29F807" box="[635,735,1945,1971]" captionStart="FIGURES 100108" captionStartId="23.[151,264,1683,1707]" captionTargetBox="[155,1431,185,1654]" captionTargetId="figure-17@23.[151,1436,179,1657]" captionTargetPageId="23" captionText="FIGURES 100108. Somatic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov., female from Jaisalmer. 100. Habitus, dorsal view; 101. Same, retrolateral view; 102. Same, frontal view; 103. Eye region, dorsal view; 104. Left chelicera, prolateral view; 105. Same, retrolateral view; 106. Spinnerets, ventral view; 107. Leg I, prolateral view; 108. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 100101 = 1.0 mm; 102105 = 0.1 mm, 106108 = 0.5 mm." pageId="25" pageNumber="64">Fig. 100</figureCitation>
). Body length 4.69. Carapace 2.09 long, 1.51 wide. Abdomen 2.57 long, 1.73 wide. Eye sizes and interdistances: ALE 0.08, AME 0.13, PLE 0.09, PME 0.09; ALEPLE 0.08, AMEAME 0.06, AMEPME 0.13, PMEPLE 0.06, PMEPME 0.09. Clypeus height at AMEs 0.10, at ALEs 0.09. Length of chelicerae 0.68. Length of palp and legs: palp 1.51 [0.50, 0.23, 0.30, 0.48], I 4.01 [1.28, 0.68, 0.87, 0.70, 0.48], II 3.88 [1.21, 0.55, 0.86, 0.74, 0.52], III 3.92 [1.21, 0.53, 0.78, 0.80, 0.60], IV 5.17 [1.50, 0.61, 1.13, 1.32, 0.61]. Leg formula: 4132. Spination of palp: femur do 2, tibia do 2 rld 1, tarsus pl 1 do 2 rlv 1; legs: femur I pl 1 do 3, II pl 2 do 3, III do 1 plv 2 rlv 2, IV do 3 pld 1 rld 1; patella III rld 1; tibia I plv 1 rlv 1, II plv 2, III pl 1 rl 2 pld 1 rld 2 plv 2 rlv 3, IV pl 1 rl 3 pld 1 rld 1 plv 3 rlv 3; metatarsus III plv 1, III pl 1 rl 2 pld 3 rld 2 plv 3 rlv 3, IV pl 2 rl 2 pld 2 rld 3 plv 3 rlv 3.
</paragraph>
<paragraph id="8BDED9745163F75A95B1FE52FBD3FDD5" blockId="26.[151,1437,151,969]" pageId="26" pageNumber="65">
<emphasis id="B91505665163F75A95B1FE52FEC7FE19" box="[199,305,403,429]" italics="true" pageId="26" pageNumber="65">Genitalia</emphasis>
(
<figureCitation id="135AC5F15163F75A9435FE52FE14FE19" box="[323,482,402,429]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Figs 113115</figureCitation>
,
<figureCitation id="135AC5F15163F75A9487FE52FDA0FE19" box="[497,598,403,429]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">118119</figureCitation>
): epigyne hirsute, less sclerotised, with weakly notched posterior border line, with median extension having broad angular posterior and narrow anterior parts, with large, median atrium (
<figureCitation id="135AC5F15163F75A95E9FE1AFECDFE41" box="[159,315,475,501]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Figs 113114</figureCitation>
,
<figureCitation id="135AC5F15163F75A943EFE1AFE83FE41" box="[328,373,475,501]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">118</figureCitation>
). Copulatory openings lie at lateral rims of atrium. Copulatory ducts slightly wavy (
<figureCitation id="135AC5F15163F75A9058FE1AFA66FE41" box="[1326,1424,475,501]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Fig. 119</figureCitation>
). Primary spermathecae bean-shaped, lying parallel to each other (
<figureCitation id="135AC5F15163F75A96F6FE3EFC11FDAD" box="[896,999,510,537]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Figs 115</figureCitation>
,
<figureCitation id="135AC5F15163F75A9683FE3EFBD4FDAD" box="[1013,1058,511,537]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">119</figureCitation>
). Paired secondary spermathecae short, parallel to each other, with oval heads, which were erroneously interpreted in Sankaran &amp; Caleb (2021) as copulatory ducts (
<figureCitation id="135AC5F15163F75A9416FD86FE32FDD5" box="[352,452,582,609]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Figs 115</figureCitation>
,
<figureCitation id="135AC5F15163F75A94A6FD86FE0BFDD5" box="[464,509,583,609]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">119</figureCitation>
). Fertilization ducts narrow, diverging (
<figureCitation id="135AC5F15163F75A96C1FD86FBEFFDD5" box="[951,1049,583,609]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Fig. 119</figureCitation>
).
</paragraph>
<paragraph id="8BDED9745163F75A95B1FDABFB52FD31" blockId="26.[151,1437,151,969]" box="[199,1188,618,645]" pageId="26" pageNumber="65">
<emphasis id="B91505665163F75A95B1FDABFEC9FD30" bold="true" box="[199,319,618,644]" pageId="26" pageNumber="65">Variation.</emphasis>
Male (n = 3): body length 3.323.72. Female (n = 3): body length 4.695.30.
</paragraph>
<paragraph id="8BDED9745163F75A95B1FD4FFA9AFC11" blockId="26.[151,1437,151,969]" pageId="26" pageNumber="65">
<emphasis id="B91505665163F75A95B1FD4FFE13FD1C" bold="true" box="[199,485,654,680]" pageId="26" pageNumber="65">Justification of transfer.</emphasis>
An examination of the
<typeStatus id="54DA67D65163F75A979AFD4EFCD1FD1D" box="[748,807,655,681]" pageId="26" pageNumber="65">types</typeStatus>
of
<taxonomicName id="4C61A2F75163F75A963AFD4EFBF7FD1D" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[844,1025,655,681]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="65" phylum="Arthropoda" rank="species" species="jodhpurensis">
<emphasis id="B91505665163F75A963AFD4EFC95FD1C" box="[844,867,655,680]" italics="true" pageId="26" pageNumber="65">M</emphasis>
.
<emphasis id="B91505665163F75A9607FD4EFBF7FD1D" box="[881,1025,655,681]" italics="true" pageId="26" pageNumber="65">jodhpurensis</emphasis>
</taxonomicName>
in a recent paper (Sankaran &amp; Caleb 2021) showed that it has all the diagnostic features of female
<taxonomicName id="4C61A2F75163F75A9642FD72FBE6FD79" box="[820,1040,691,717]" pageId="26" pageNumber="65">
<emphasis id="B91505665163F75A9642FD72FC41FD79" box="[820,951,691,717]" italics="true" pageId="26" pageNumber="65">Trichothyse</emphasis>
species
</taxonomicName>
, although at the time the association with this genus was unclear due to its geographical range. Recently collected male and female specimens of this species confirmed its similarity with
<taxonomicName id="4C61A2F75163F75A9745FD3AFCE7FCA1" box="[563,785,763,789]" pageId="26" pageNumber="65">
<emphasis id="B91505665163F75A9745FD3AFD40FCA1" box="[563,694,763,789]" italics="true" pageId="26" pageNumber="65">Trichothyse</emphasis>
species
</taxonomicName>
: distomedially originating broad conductor with prolateral fold, apically originating claw-like embolus with broad embolic base, medially contiguous sperm duct with single loop, robust RTA with apical bifurcation, epigyne with median extension, and vulva with short copulatory ducts, large, bean-shaped primary and small secondary spermathecae (cf.
<figureCitation id="135AC5F15163F75A9609FCA6FC19FC35" box="[895,1007,871,897]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 920" captionStartId-0="3.[151,264,1372,1396]" captionStartId-1="4.[151,264,1607,1631]" captionTargetBox-0="[152,1436,179,1346]" captionTargetBox-1="[151,1436,179,1581]" captionTargetId-0="figure-17@3.[151,1436,179,1346]" captionTargetId-1="figure-17@4.[151,1436,179,1581]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." captionText-1="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="26" pageNumber="65">Figs 120</figureCitation>
,
<figureCitation id="135AC5F15163F75A968CFCA7FBB4FC35" box="[1018,1090,870,897]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">5457</figureCitation>
,
<figureCitation id="135AC5F15163F75A913BFCA6FB12FC35" box="[1101,1252,871,897]" captionStart-0="FIGURES 109115" captionStart-1="FIGURES 116119" captionStartId-0="24.[151,257,1943,1967]" captionStartId-1="25.[151,264,1479,1503]" captionTargetBox-0="[190,1398,195,1913]" captionTargetBox-1="[161,1426,189,1443]" captionTargetId-0="figure-17@24.[179,1408,179,1917]" captionTargetId-1="figure-17@25.[151,1436,179,1453]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." captionText-1="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="26" pageNumber="65">Figs 109119</figureCitation>
and Sankaran &amp; Caleb 2021: figs 4951). Based on these observations, we propose transferring
<taxonomicName id="4C61A2F75163F75A9166FC4AFB49FC11" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Gajbe" baseAuthorityYear="1993" box="[1040,1215,907,933]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="65" phylum="Arthropoda" rank="species" species="jodhpurensis">
<emphasis id="B91505665163F75A9166FC4AFBD7FC10" box="[1040,1057,907,932]" italics="true" pageId="26" pageNumber="65">P</emphasis>
.
<emphasis id="B91505665163F75A9159FC4AFB49FC11" box="[1071,1215,907,933]" italics="true" pageId="26" pageNumber="65">jodhpurensis</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75163F75A9195FC4AFA90FC11" authorityName="Tucker" authorityYear="1923" box="[1251,1382,907,933]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665163F75A9195FC4AFA90FC11" box="[1251,1382,907,933]" italics="true" pageId="26" pageNumber="65">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5163F75A95B1FC6FFB65FC7D" box="[199,1171,942,969]" pageId="26" pageNumber="65" type="distribution">
<paragraph id="8BDED9745163F75A95B1FC6FFB65FC7D" blockId="26.[151,1437,151,969]" box="[199,1171,942,969]" pageId="26" pageNumber="65">
<emphasis id="B91505665163F75A95B1FC6FFE97FC7C" bold="true" box="[199,353,942,968]" pageId="26" pageNumber="65">Distribution.</emphasis>
Known from the Jaisalmer and Jodhpur districts of
<collectingRegion id="49A517965163F75A96D1FC6EFBE3FC7D" box="[935,1045,943,969]" country="India" name="Rajasthan" pageId="26" pageNumber="65">Rajasthan</collectingRegion>
(
<figureCitation id="135AC5F15163F75A9153FC6EFB70FC7D" box="[1061,1158,943,969]" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="26" pageNumber="65">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,186 @@
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<mods:title id="1FD5C90D11F02D2760534A0E0BCFD359">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:namePart id="0BE08D75F45F7D3590B15416824BCF03">Haddad, Charles R.</mods:namePart>
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<mods:affiliation id="FC5E2313C0078DF757881B7C8FA743CD">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="BA0E6E3BDD75263109E7499265890FF8" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:namePart id="4C4325F103E5C66139E73E7E303D42B0">Tripathi, Rishikesh</mods:namePart>
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<mods:affiliation id="B3190EC8BC99F13329FC58B11E70C29B">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Thrissur, Kerala 680 125, India.</mods:affiliation>
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<paragraph id="8BDED974516DF75495E1FF1EFD31FF4D" blockId="20.[151,711,223,250]" box="[151,711,223,250]" pageId="20" pageNumber="59">
<heading id="D0966E18516DF75495E1FF1EFD31FF4D" bold="true" box="[151,711,223,250]" fontSize="11" level="1" pageId="20" pageNumber="59" reason="1">
<emphasis id="B9150566516DF75495E1FF1EFD31FF4D" bold="true" box="[151,711,223,250]" pageId="20" pageNumber="59">
<taxonomicName id="4C61A2F7516DF75495E1FF1EFDCFFF4D" authority="(Simon, 1914)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Simon" baseAuthorityYear="1914" box="[151,569,223,250]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="furcata" status="comb. nov.">
<emphasis id="B9150566516DF75495E1FF1EFE73FF4D" bold="true" box="[151,389,223,250]" italics="true" pageId="20" pageNumber="59">Trichothyse furcata</emphasis>
(
<bibRefCitation id="EFF0A485516DF75494E2FF1EFDC6FF4D" author="Simon, E." box="[404,560,223,249]" pageId="20" pageNumber="59" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon, 1914</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A226B81D516DF7549736FF1EFD31FF4D" box="[576,711,223,249]" pageId="20" pageNumber="59" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37B8AFF516DF75495E1FEE5FC4CFEEE" pageId="20" pageNumber="59" type="reference_group">
<paragraph id="8BDED974516DF75495E1FEE5FC4CFEEE" blockId="20.[151,1436,291,346]" pageId="20" pageNumber="59">
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<treatmentCitation id="0AC0FF65516DF75495E1FEE5FDDAFE88" author="Simon, E." box="[151,556,292,316]" page="184" pageId="20" pageNumber="59" year="1914">
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<emphasis id="B9150566516DF75495E1FEE5FE9BFE88" box="[151,365,292,316]" italics="true" pageId="20" pageNumber="59">Poecilochroa furcata</emphasis>
<bibRefCitation id="EFF0A485516DF7549402FEE5FDDAFE88" author="Simon, E." box="[372,556,292,316]" pageId="20" pageNumber="59" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon, 1914: 184</bibRefCitation>
</taxonomicName>
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, 221, figs 393394;
<treatmentCitation id="0AC0FF65516DF754978BFEE5FC2FFE88" author="Di Franco, F." box="[765,985,292,316]" page="203" pageId="20" pageNumber="59" year="2001">
<bibRefCitation id="EFF0A485516DF754978BFEE5FC2FFE88" author="Di Franco, F." box="[765,985,292,316]" pageId="20" pageNumber="59" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco, 2001: 203</bibRefCitation>
</treatmentCitation>
, figs 34 (
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<specimenCount id="9D6712FD516DF75491D3FEE5FB4DFE8F" box="[1189,1211,292,315]" count="1" pageId="20" pageNumber="59" type="male"></specimenCount>
:
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<collectingCountry id="F37699E4516DF75491B3FEE2FAC4FE8F" box="[1221,1330,291,315]" name="France" pageId="20" pageNumber="59">FRANCE</collectingCountry>
:
</emphasis>
Provence [
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,
<geoCoordinate id="EE55BFB3516DF7549442FE83FE7AFEEE" box="[308,396,322,346]" degrees="06" direction="east" minutes="12" orientation="longitude" pageId="20" pageNumber="59" precision="925" value="6.2">06°12E</geoCoordinate>
], date and collector unknown, MNHN—not examined.
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<emphasis id="B9150566516DF75495E1FE49FEDFFE16" bold="true" box="[151,297,392,418]" pageId="20" pageNumber="59">Description.</emphasis>
<emphasis id="B9150566516DF7549446FE49FE9CFE16" box="[304,362,392,418]" italics="true" pageId="20" pageNumber="59">Male</emphasis>
. For redescription of the male, see
<bibRefCitation id="EFF0A485516DF7549783FE49FC4CFE17" author="Di Franco, F." box="[757,954,392,419]" pageId="20" pageNumber="59" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco (2001)</bibRefCitation>
.
</paragraph>
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<emphasis id="B9150566516DF75495B1FE6DFEEDFE72" box="[199,283,428,454]" italics="true" pageId="20" pageNumber="59">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BDED974516DF75495B1FE11FBC3FD77" blockId="20.[151,1437,392,779]" pageId="20" pageNumber="59">
<emphasis id="B9150566516DF75495B1FE11FE12FE5E" bold="true" box="[199,484,464,490]" pageId="20" pageNumber="59">Justification of transfer.</emphasis>
The illustrations of the palp provided in
<bibRefCitation id="EFF0A485516DF75496DFFE11FBC8FE5F" author="Simon, E." box="[937,1086,464,491]" pageId="20" pageNumber="59" refId="ref20190" refString="Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book" year="1914">Simon (1914</bibRefCitation>
: fig. 393) and
<bibRefCitation id="EFF0A485516DF75491ABFE11FA6EFE5F" author="Di Franco, F." box="[1245,1432,464,491]" pageId="20" pageNumber="59" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco (2001</bibRefCitation>
: figs 34) are diagnostic and were used for comparison. Even though we did not examine the
<typeStatus id="54DA67D6516DF75491D6FE35FAF4FDBA" box="[1184,1282,500,526]" pageId="20" pageNumber="59" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C61A2F7516DF7549051FE34FA63FDBB" authorityName="Simon" authorityYear="1914" box="[1319,1429,501,527]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="furcata">
<emphasis id="B9150566516DF7549051FE34FACEFDBA" box="[1319,1336,501,526]" italics="true" pageId="20" pageNumber="59">P</emphasis>
.
<emphasis id="B9150566516DF7549033FE34FA63FDBB" box="[1349,1429,501,527]" italics="true" pageId="20" pageNumber="59">furcata</emphasis>
</taxonomicName>
, the original illustrations of the palp of this species are consistent with the diagnostic features of male
<taxonomicName id="4C61A2F7516DF754906CFDD9FF1DFDE2" pageId="20" pageNumber="59">
<emphasis id="B9150566516DF754906CFDD9FA6BFD86" box="[1306,1437,536,562]" italics="true" pageId="20" pageNumber="59">Trichothyse</emphasis>
species
</taxonomicName>
, in contrast to
<emphasis id="B9150566516DF75494ECFDFDFD3FFDE2" box="[410,713,572,598]" italics="true" pageId="20" pageNumber="59">
<taxonomicName id="4C61A2F7516DF75494ECFDFDFDC6FDE2" authorityName="Westring" authorityYear="1874" box="[410,560,572,598]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="59" phylum="Arthropoda" rank="genus" sensu="stricto">Poecilochroa</taxonomicName>
<taxonomicNameLabel id="A226B81D516DF754974FFDFCFD3FFDE2" box="[569,713,573,598]" pageId="20" pageNumber="59" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C61A2F7516DF7549672FDFDFC5DFDE2" authorityName="Wunderlich" authorityYear="2011" box="[772,939,572,598]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B9150566516DF7549672FDFDFC5DFDE2" box="[772,939,572,598]" italics="true" pageId="20" pageNumber="59">Macarophaeus</emphasis>
</taxonomicName>
: distomedially originating broad conductor with prolateral fold, apically originating claw-like embolus with broad embolic base, medially contiguous sperm duct with single loop and robust RTA with apical bifurcation (cf.
<bibRefCitation id="EFF0A485516DF75496F5FD45FBCCFD2B" author="Di Franco, F." box="[899,1082,644,671]" pageId="20" pageNumber="59" pagination="195 - 215" refId="ref18761" refString="Di Franco, F. (2001) Gnaphosidae (Arachnida, Araneae) della Riserva Naturale Orientata &quot; Oasi del Simeto &quot; (Catania). Naturalista Siciliano, 25, 195-215." type="journal article" year="2001">Di Franco 2001</bibRefCitation>
: figs 34 and
<figureCitation id="135AC5F1516DF75491ABFD45FAB4FD2B" box="[1245,1346,644,671]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="20" pageNumber="59">Figs 18</figureCitation>
,
<figureCitation id="135AC5F1516DF7549039FD45FA6EFD2B" box="[1359,1432,644,671]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="20" pageNumber="59">5455</figureCitation>
,
<figureCitation id="135AC5F1516DF75495E1FD68FF0DFD77" box="[151,251,681,707]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="20" pageNumber="59">109112</figureCitation>
,
<figureCitation id="135AC5F1516DF7549470FD69FE9DFD76" box="[262,363,680,706]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="20" pageNumber="59">116117</figureCitation>
). We therefore propose the transfer of
<taxonomicName id="4C61A2F7516DF754966EFD68FC71FD77" authorityName="Simon" authorityYear="1914" box="[792,903,681,707]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="59" phylum="Arthropoda" rank="species" species="furcata">
<emphasis id="B9150566516DF754966EFD68FCDFFD76" box="[792,809,681,706]" italics="true" pageId="20" pageNumber="59">P</emphasis>
.
<emphasis id="B9150566516DF7549641FD68FC71FD77" box="[823,903,681,707]" italics="true" pageId="20" pageNumber="59">furcata</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F7516DF75496DDFD69FBD8FD76" authorityName="Tucker" authorityYear="1923" box="[939,1070,680,706]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B9150566516DF75496DDFD69FBD8FD76" box="[939,1070,680,706]" italics="true" pageId="20" pageNumber="59">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF516DF75495B1FD0DFF24FCBF" pageId="20" pageNumber="59" type="distribution">
<paragraph id="8BDED974516DF75495B1FD0DFF24FCBF" blockId="20.[151,1437,392,779]" pageId="20" pageNumber="59">
<emphasis id="B9150566516DF75495B1FD0DFE97FD52" bold="true" box="[199,353,716,742]" pageId="20" pageNumber="59">Distribution.</emphasis>
Recorded from three countries in Mediterranean Europe, i.e.
<collectingCountry id="F37699E4516DF7549140FD0CFB73FD53" box="[1078,1157,717,743]" name="France" pageId="20" pageNumber="59">France</collectingCountry>
,
<collectingCountry id="F37699E4516DF75491E2FD0CFB10FD53" box="[1172,1254,717,743]" name="Greece" pageId="20" pageNumber="59">Greece</collectingCountry>
, and
<collectingCountry id="F37699E4516DF754905EFD0DFAADFD52" box="[1320,1371,716,742]" name="Italy" pageId="20" pageNumber="59">Italy</collectingCountry>
(
<figureCitation id="135AC5F1516DF7549019FD0DFF32FCBF" captionStart="FIGURE 121" captionStartId="27.[151,250,1113,1137]" captionTargetBox="[152,1435,181,1086]" captionTargetId="figure-17@27.[151,1436,179,1088]" captionTargetPageId="27" captionText="FIGURE 121. Distribution records of the known Palaearctic Trichothyse species." pageId="20" pageNumber="59">Fig. 121</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,406 @@
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<mods:title id="B19B5CE1E70F940DC03C429C703C497C">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:namePart id="1025F137225DBE49A6E7989B9EEF2684">Sankaran, Pradeep M.</mods:namePart>
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<mods:namePart id="E26AB3DAAF4AEBD9B6860F9334C72A37">Haddad, Charles R.</mods:namePart>
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<mods:affiliation id="CB89739415C41D0C40BAD430041117A9">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="046658919A61841DAE0AB464BF9A6BB2" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:affiliation id="AF413E075C8875A3A7D50BFF3E7081B5">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Thrissur, Kerala 680 125, India.</mods:affiliation>
<mods:nameIdentifier id="177046D369704010A33A6D40E38E88CD" type="email">rishikesh.tripathi14@gmail.com</mods:nameIdentifier>
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<subSubSection id="C37B8AFF5170F74995E1FC9FFD10FCCD" box="[151,742,862,889]" pageId="9" pageNumber="48" type="nomenclature">
<paragraph id="8BDED9745170F74995E1FC9FFD10FCCD" blockId="9.[151,742,862,925]" box="[151,742,862,889]" pageId="9" pageNumber="48">
<heading id="D0966E185170F74995E1FC9FFD10FCCD" bold="true" box="[151,742,862,889]" fontSize="11" level="1" pageId="9" pageNumber="48" reason="1">
<emphasis id="B91505665170F74995E1FC9FFD10FCCD" bold="true" box="[151,742,862,889]" pageId="9" pageNumber="48">
<taxonomicName id="4C61A2F75170F74995E1FC9FFD76FCCC" authority="Haddad &amp; Sankaran" authorityName="Haddad &amp; Sankaran" authorityYear="2025" box="[151,640,862,889]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B91505665170F74995E1FC9FFE85FCCD" bold="true" box="[151,371,862,889]" italics="true" pageId="9" pageNumber="48">Trichothyse karoo</emphasis>
Haddad &amp; Sankaran
</taxonomicName>
<taxonomicNameLabel id="A226B81D5170F74997FEFC9EFD10FCCD" box="[648,742,863,889]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5170F74995E1FC42FE69FC29" box="[151,415,898,925]" pageId="9" pageNumber="48" type="description">
<paragraph id="8BDED9745170F74995E1FC42FE69FC29" blockId="9.[151,742,862,925]" box="[151,415,898,925]" pageId="9" pageNumber="48">
<figureCitation id="135AC5F15170F74995E1FC42FF0CFC29" box="[151,250,899,925]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="9" pageNumber="48">Figs 3, 7</figureCitation>
,
<figureCitation id="135AC5F15170F7499470FC42FEBBFC29" box="[262,333,899,925]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="9" pageNumber="48">11, 17</figureCitation>
,
<figureCitation id="135AC5F15170F749942FFC43FE69FC29" box="[345,415,898,925]" captionStart-0="FIGURES 4247" captionStart-1="FIGURES 4853" captionStart-2="FIGURES 5457" captionStartId-0="10.[151,264,1511,1535]" captionStartId-1="11.[151,264,1906,1930]" captionStartId-2="12.[151,264,1679,1703]" captionTargetBox-0="[151,1436,414,1485]" captionTargetBox-1="[151,1436,810,1880]" captionTargetBox-2="[162,1425,190,1642]" captionTargetId-0="figure-191@10.[151,1436,414,1485]" captionTargetId-1="figure-357@11.[151,1436,810,1880]" captionTargetId-2="figure-17@12.[151,1435,179,1653]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." captionText-1="FIGURES 4853. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., paratype female from Jagerskraal: 48. Habitus, dorsal view; 49. Eye region, anterior view; 50. Same, dorsal view; 51. Chelicerae, posteroventral view; 52. Leg I, retrolateral view; 53. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 48 = 1 mm; 4953 = 0.5 mm." captionText-2="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="9" pageNumber="48">4257</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5170F74995E1FC0AFA61FB55" pageId="9" pageNumber="48" type="materials_examined">
<paragraph id="8BDED9745170F74995E1FC0AFAF7FBBD" blockId="9.[151,1437,970,2041]" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995E1FC0AFEB3FC50" bold="true" box="[151,325,970,997]" pageId="9" pageNumber="48">Type material.</emphasis>
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:
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:
<emphasis id="B91505665170F74997B0FC0AFC90FC51" bold="true" box="[710,870,971,997]" italics="true" pageId="9" pageNumber="48">
<collectingRegion id="49A517965170F74997B0FC0AFC90FC51" box="[710,870,971,997]" country="South Africa" name="Western Cape" pageId="9" pageNumber="48">Western Cape</collectingRegion>
</emphasis>
:
</emphasis>
<collectingMunicipality id="6BBA430E5170F7499601FC0AFBEBFC51" box="[887,1053,971,997]" pageId="9" pageNumber="48">Matjiesfontein</collectingMunicipality>
,
<location id="8EBE8FAF5170F749915FFC0AFB1AFC51" LSID="urn:lsid:plazi:treatment:03C868625170F74B95E1FC9FFB2BFCA1:8EBE8FAF5170F749915FFC0AFB1AFC51" box="[1065,1260,971,997]" country="South Africa" latitude="-33.22211" longLatPrecision="1" longitude="20.409945" municipality="Matjiesfontein" name="Farm Jagerskraal" pageId="9" pageNumber="48" stateProvince="Western Cape">Farm Jagerskraal</location>
,
<geoCoordinate id="EE55BFB35170F749918EFC0AFA61FC51" box="[1272,1431,971,997]" degrees="33" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="19.6" value="-33.22211">33°1319.6S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35170F74995E1FC2FFECEFBBD" box="[151,312,1006,1033]" degrees="20" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="48" precision="1" seconds="35.8" value="20.409945">20°2435.8E</geoCoordinate>
,
<elevation id="004C3E475170F7499432FC2FFE3EFBBD" box="[324,456,1006,1033]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="9" pageNumber="48" unit="m" value="975.0">
<quantity id="4C9974915170F7499432FC2FFE7DFBBD" box="[324,395,1006,1033]" metricMagnitude="2" metricUnit="m" metricValue="9.75" pageId="9" pageNumber="48" unit="m" value="975.0">975 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45170F74994A2FC2FFDBBFBBD" box="[468,589,1006,1033]" pageId="9" pageNumber="48" value="2015-10-18">
<collectingDate id="EF9B065C5170F74994A2FC2FFDBBFBBD" box="[468,589,1006,1033]" pageId="9" pageNumber="48" value="2015-10-18">18.X.2015</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25170F74997FFFC2EFD2BFBBD" box="[649,733,1007,1033]" pageId="9" pageNumber="48">Z. Mbo</collectorName>
(hand collecting, karoo veld) (
<specimenCode id="DBC7710F5170F7499143FC2EFB44FBBD" box="[1077,1202,1007,1033]" collectionCode="NCA" pageId="9" pageNumber="48">NCA 2016</specimenCode>
/2262).
</materialsCitation>
</paragraph>
<paragraph id="8BDED9745170F74995B1FBD2FA61FB55" blockId="9.[151,1437,970,2041]" pageId="9" pageNumber="48">
<materialsCitation id="3B09D3295170F74995B1FBD2FA65FB55" collectingDate="2015-09-08" collectingDateMax="2016-12-31" collectingDateMin="2015-09-08" collectionCode="NCA" collectorName="Z. Mbo &amp; W. Jubber" country="South Africa" county="Laingsburg" elevation="660" latitude="-33.81525" location="Les Hauts de Montagu" longLatPrecision="1" longitude="20.151249" municipality="Anysberg Nature Reserve" pageId="9" pageNumber="48" specimenCode="NCA 2016, NCA 2017" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="54DA67D65170F74995B1FBD2FECFFB99" box="[199,313,1043,1069]" pageId="9" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B91505665170F7499426FBD3FCF3FB99" bold="true" box="[336,773,1042,1069]" pageId="9" pageNumber="48">
<collectingCountry id="F37699E45170F7499426FBD3FDC1FB98" box="[336,567,1042,1068]" name="South Africa" pageId="9" pageNumber="48">SOUTH AFRICA</collectingCountry>
:
<emphasis id="B91505665170F7499739FBD2FD0AFB99" bold="true" box="[591,764,1043,1069]" italics="true" pageId="9" pageNumber="48">
<collectingRegion id="49A517965170F7499739FBD2FD0AFB99" box="[591,764,1043,1069]" country="South Africa" name="Western Cape" pageId="9" pageNumber="48">Western Cape</collectingRegion>
</emphasis>
:
</emphasis>
<collectingCounty id="62BFA1F85170F749966EFBD2FC6DFB99" box="[792,923,1043,1069]" pageId="9" pageNumber="48">Laingsburg</collectingCounty>
,
<collectingMunicipality id="6BBA430E5170F74996C7FBD2FB19FB99" box="[945,1263,1043,1069]" pageId="9" pageNumber="48">Anysberg Nature Reserve</collectingMunicipality>
,
<geoCoordinate id="EE55BFB35170F7499071FBD3FA6EFB99" box="[1287,1432,1042,1069]" degrees="33" direction="south" minutes="28.454" orientation="latitude" pageId="9" pageNumber="48" precision="1" value="-33.47423">33°28.454S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35170F74995E1FBF6FEDCFBE5" box="[151,298,1079,1105]" degrees="20" direction="east" minutes="40.696" orientation="longitude" pageId="9" pageNumber="48" precision="1" value="20.678267">20°40.696E</geoCoordinate>
,
<elevation id="004C3E475170F7499441FBF6FE48FBE5" box="[311,446,1079,1105]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="48" unit="m" value="660.0">
<quantity id="4C9974915170F7499441FBF6FE89FBE5" box="[311,383,1079,1105]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="48" unit="m" value="660.0">660 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45170F74994BAFBF7FD8FFBE5" box="[460,633,1078,1105]" pageId="9" pageNumber="48" value="2015-09-08" valueMax="2015-10-08" valueMin="2015-09-08">
<collectingDate id="EF9B065C5170F74994BAFBF7FD8FFBE5" box="[460,633,1078,1105]" pageId="9" pageNumber="48" value="2015-09-08" valueMax="2015-10-08" valueMin="2015-09-08">8.IX8.X.2015</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25170F74997CEFBF6FCFBFBE5" box="[696,781,1079,1105]" pageId="9" pageNumber="48">Z. Mbo</collectorName>
(
<collectingMethod id="5220A1635170F7499668FBF6FC53FBE5" box="[798,933,1079,1105]" pageId="9" pageNumber="48">pitfall traps</collectingMethod>
, riparian woodland),
<specimenCount id="9D6712FD5170F74991ECFBF6FB4BFBE5" box="[1178,1213,1079,1105]" count="1" pageId="9" pageNumber="48" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F5170F74991BBFBF6FABBFBE5" box="[1229,1357,1079,1105]" collectionCode="NCA" pageId="9" pageNumber="48">NCA 2016</specimenCode>
/2493); Matjiesfontein, Farm Jagerskraal,
<geoCoordinate id="EE55BFB35170F749975DFB9AFD3AFBC1" box="[555,716,1115,1141]" degrees="33" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="28.6" value="-33.224613">33°1328.6S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35170F74997A8FB9AFC76FBC1" box="[734,896,1115,1141]" degrees="20" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="48" precision="1" seconds="29.8" value="20.408278">20°2429.8E</geoCoordinate>
,
<elevation id="004C3E475170F74996E4FB9BFBD4FBC1" box="[914,1058,1114,1141]" metricMagnitude="2" metricUnit="m" metricValue="9.95" pageId="9" pageNumber="48" unit="m" value="995.0">
<quantity id="4C9974915170F74996E4FB9BFC29FBC1" box="[914,991,1114,1141]" metricMagnitude="2" metricUnit="m" metricValue="9.95" pageId="9" pageNumber="48" unit="m" value="995.0">995 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45170F7499142FB9BFB58FBC1" box="[1076,1198,1114,1141]" pageId="9" pageNumber="48" value="2015-10-16">
<collectingDate id="EF9B065C5170F7499142FB9BFB58FBC1" box="[1076,1198,1114,1141]" pageId="9" pageNumber="48" value="2015-10-16">16.X.2015</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25170F7499183FB9AFAB9FBC1" box="[1269,1359,1115,1141]" pageId="9" pageNumber="48">Z. Mbo</collectorName>
(hand collecting, karoo veld),
<specimenCount id="9D6712FD5170F74994D2FBBEFE31FB2D" box="[420,455,1151,1177]" count="1" pageId="9" pageNumber="48" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5170F74994A0FBBEFDA2FB2D" box="[470,596,1151,1177]" collectionCode="NCA" pageId="9" pageNumber="48">NCA 2016</specimenCode>
/2243); Same locality,
<geoCoordinate id="EE55BFB35170F7499627FBBEFC05FB2D" box="[849,1011,1151,1177]" degrees="33" direction="south" minutes="13" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="48.1" value="-33.23003">33°1348.1S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35170F7499689FBBFFB54FB2D" box="[1023,1186,1150,1177]" degrees="20" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="48" precision="1" seconds="54.4" value="20.415112">20°2454.4E</geoCoordinate>
,
<elevation id="004C3E475170F74991D8FBBFFAB6FB2D" box="[1198,1344,1150,1177]" metricMagnitude="3" metricUnit="m" metricValue="1.045" pageId="9" pageNumber="48" unit="m" value="1045.0">
<quantity id="4C9974915170F74991D8FBBFFAF5FB2D" box="[1198,1283,1150,1177]" metricMagnitude="3" metricUnit="m" metricValue="1.045" pageId="9" pageNumber="48" unit="m" value="1045.0">1045 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45170F749903BFBBEFEE7FB09" pageId="9" pageNumber="48" value="2015-09-10" valueMax="2015-10-15" valueMin="2015-09-10">
<collectingDate id="EF9B065C5170F749903BFBBEFEE7FB09" pageId="9" pageNumber="48" value="2015-09-10" valueMax="2015-10-15" valueMin="2015-09-10">10.IX 15.X.2015</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25170F7499422FB62FE5AFB09" box="[340,428,1187,1213]" pageId="9" pageNumber="48">Z. Mbo</collectorName>
(
<collectingMethod id="5220A1635170F74994C8FB62FDBEFB09" box="[446,584,1187,1213]" pageId="9" pageNumber="48">pitfall traps</collectingMethod>
, open ground),
<specimenCount id="9D6712FD5170F7499675FB62FCD0FB09" box="[771,806,1187,1213]" count="1" pageId="9" pageNumber="48" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F5170F749964EFB62FC4FFB09" box="[824,953,1187,1213]" collectionCode="NCA" pageId="9" pageNumber="48">NCA 2016</specimenCode>
/2224); Montagu,
<location id="8EBE8FAF5170F74991FCFB62FA6EFB09" LSID="urn:lsid:plazi:treatment:03C868625170F74B95E1FC9FFB2BFCA1:8EBE8FAF5170F74991FCFB62FA6EFB09" box="[1162,1432,1187,1213]" country="South Africa" county="Laingsburg" latitude="-33.81525" longLatPrecision="1" longitude="20.151249" municipality="Anysberg Nature Reserve" name="Les Hauts de Montagu" pageId="9" pageNumber="48" stateProvince="Western Cape">Les Hauts de Montagu</location>
,
<geoCoordinate id="EE55BFB35170F74995E1FB07FECEFB55" box="[151,312,1222,1249]" degrees="33" direction="south" minutes="48" orientation="latitude" pageId="9" pageNumber="48" precision="1" seconds="54.9" value="-33.81525">33°4854.9S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35170F7499433FB07FE11FB55" box="[325,487,1222,1249]" degrees="20" direction="east" minutes="09" orientation="longitude" pageId="9" pageNumber="48" precision="1" seconds="04.5" value="20.151249">20°0904.5E</geoCoordinate>
,
<elevation id="004C3E475170F7499482FB06FD8EFB55" box="[500,632,1223,1249]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="48" unit="m" value="360.0">
<quantity id="4C9974915170F7499482FB06FDCDFB55" box="[500,571,1223,1249]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="48" unit="m" value="360.0">360 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45170F74997F2FB06FCD5FB55" box="[644,803,1223,1249]" pageId="9" pageNumber="48" value="2016-07" valueMax="2016-12-31" valueMin="2016-07-00">
<collectingDate id="EF9B065C5170F74997F2FB06FCD5FB55" box="[644,803,1223,1249]" pageId="9" pageNumber="48" value="2016-07" valueMax="2016-12-31" valueMin="2016-07-00">VIIXII.2016</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25170F7499629FB06FC26FB55" box="[863,976,1223,1249]" pageId="9" pageNumber="48">W. Jubber</collectorName>
(hand collecting),
<specimenCount id="9D6712FD5170F74991D0FB06FB3FFB55" box="[1190,1225,1223,1249]" count="1" pageId="9" pageNumber="48" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F5170F74991AEFB06FAA1FB55" box="[1240,1367,1222,1249]" collectionCode="NCA" pageId="9" pageNumber="48">NCA 2017</specimenCode>
/521)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5170F74995B1FB2AFDC2FA9D" pageId="9" pageNumber="48" type="etymology">
<paragraph id="8BDED9745170F74995B1FB2AFDC2FA9D" blockId="9.[151,1437,970,2041]" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995B1FB2AFEB8FAB1" bold="true" box="[199,334,1259,1285]" pageId="9" pageNumber="48">Etymology.</emphasis>
The specific epithet is a noun in apposition referring to the habitat in which it was collected, the Nama Karoo Biome of
<collectingCountry id="F37699E45170F74994E8FACEFDC6FA9D" box="[414,560,1295,1321]" name="South Africa" pageId="9" pageNumber="48">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5170F74995B1FAF3FDE9F991" pageId="9" pageNumber="48" type="diagnosis">
<paragraph id="8BDED9745170F74995B1FAF3FDE9F991" blockId="9.[151,1437,970,2041]" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995B1FAF3FEB4FAF8" bold="true" box="[199,322,1330,1356]" pageId="9" pageNumber="48">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C61A2F75170F74994C0FAF2FDE3FAF9" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[438,533,1331,1357]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B91505665170F74994C0FAF2FE30FAF8" box="[438,454,1331,1356]" italics="true" pageId="9" pageNumber="48">T</emphasis>
.
<emphasis id="B91505665170F74994A3FAF2FDE3FAF9" box="[469,533,1331,1357]" italics="true" pageId="9" pageNumber="48">karoo</emphasis>
</taxonomicName>
<emphasis id="B91505665170F749976BFAF2FD8EFAF9" bold="true" box="[541,632,1331,1357]" pageId="9" pageNumber="48">
<taxonomicNameLabel id="A226B81D5170F749976BFAF2FD8EFAF9" box="[541,632,1331,1357]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C61A2F75170F74996F4FAF2FBF3FAF9" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[898,1029,1331,1357]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B91505665170F74996F4FAF2FBF3FAF9" box="[898,1029,1331,1357]" italics="true" pageId="9" pageNumber="48">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B91505665170F749917BFAF2FB79FAF9" bold="true" box="[1037,1167,1331,1357]" pageId="9" pageNumber="48">
<taxonomicNameLabel id="A226B81D5170F749917BFAF2FB79FAF9" box="[1037,1167,1331,1357]" pageId="9" pageNumber="48" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
in the general structure of the palp but can be distinguished by the different passage of the sperm duct, the more gradual curvature of the embolus (
<figureCitation id="135AC5F15170F7499472FABAFE9FFA21" box="[260,361,1403,1429]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="9" pageNumber="48">Figs 34</figureCitation>
,
<figureCitation id="135AC5F15170F7499402FABBFE65FA21" box="[372,403,1402,1429]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="9" pageNumber="48">54</figureCitation>
), and the shorter basal section and the longer/sharper dorsal branch of the RTA in retrolateral view (
<figureCitation id="135AC5F15170F74995A8FA5EFEB3FA0D" box="[222,325,1439,1465]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="9" pageNumber="48">Figs 78</figureCitation>
,
<figureCitation id="135AC5F15170F7499424FA5FFE84FA0D" box="[338,370,1438,1465]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="9" pageNumber="48">55</figureCitation>
). Females of
<taxonomicName id="4C61A2F75170F749977BFA5EFD9CFA0D" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[525,618,1439,1465]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B91505665170F749977BFA5EFD9CFA0D" box="[525,618,1439,1465]" italics="true" pageId="9" pageNumber="48">T. karoo</emphasis>
</taxonomicName>
<emphasis id="B91505665170F7499705FA5EFD38FA0D" bold="true" box="[627,718,1439,1465]" pageId="9" pageNumber="48">
<taxonomicNameLabel id="A226B81D5170F7499705FA5EFD38FA0D" box="[627,718,1439,1465]" pageId="9" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share with
<taxonomicName id="4C61A2F75170F7499623FA5EFC20FA0D" authorityName="Lawrence" authorityYear="1928" box="[853,982,1439,1465]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B91505665170F7499623FA5EFC20FA0D" box="[853,982,1439,1465]" italics="true" pageId="9" pageNumber="48">T. fontensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C61A2F75170F7499166FA5EFB62FA0D" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[1040,1172,1439,1465]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B91505665170F7499166FA5EFB62FA0D" box="[1040,1172,1439,1465]" italics="true" pageId="9" pageNumber="48">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B91505665170F74991EAFA5EFAE8FA0D" bold="true" box="[1180,1310,1439,1465]" pageId="9" pageNumber="48">
<taxonomicNameLabel id="A226B81D5170F74991EAFA5EFAE8FA0D" box="[1180,1310,1439,1465]" pageId="9" pageNumber="48" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
the almost parallel mesal margins of the primary spermathecae but can be distinguished from all their congeners by the presence of a sclerotised posterior mesal ridge that terminates in a small teardrop-shaped atrium centrally in the epigyne (cf.
<figureCitation id="135AC5F15170F74995E1F9CAFEACF991" box="[151,346,1546,1573]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="9" pageNumber="48">Figs 9, 13, 15, 19</figureCitation>
and
<figureCitation id="135AC5F15170F74994E7F9CAFDE5F991" box="[401,531,1546,1573]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="9" pageNumber="48">Figs 5657</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5170F74B95B1F9EFFC9EFD45" lastPageId="11" lastPageNumber="50" pageId="9" pageNumber="48" type="description">
<paragraph id="8BDED9745170F74A95B1F9EFFB15FE3D" blockId="9.[151,1437,970,2041]" lastBlockId="10.[151,1437,150,393]" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995B1F9EFFEAFF9FC" bold="true" box="[199,345,1582,1608]" pageId="9" pageNumber="48">Description.</emphasis>
<emphasis id="B91505665170F7499415F9EEFE6BF9FD" box="[355,413,1583,1609]" italics="true" pageId="9" pageNumber="48">Male</emphasis>
(
<typeStatus id="54DA67D65170F74994D9F9EEFDE3F9FD" box="[431,533,1583,1609]" pageId="9" pageNumber="48" type="holotype">holotype</typeStatus>
, Matjiesfontein, NCA 2016/2262,
<figureCitation id="135AC5F15170F74996D1F9EEFBDAF9FD" box="[935,1068,1583,1609]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Figs 4247</figureCitation>
). Carapace, clypeus, chelicerae, labium, endites and sternum deep orange-brown (
<figureCitation id="135AC5F15170F74997CDF992FCCBF9D9" box="[699,829,1618,1645]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Figs 4245</figureCitation>
); legs deep yellow-brown, with black mottling, femora slightly darker than other segments (
<figureCitation id="135AC5F15170F7499735F9B6FD3CF925" box="[579,714,1655,1681]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Figs 4647</figureCitation>
); eye field slightly darker than rest of carapace (
<figureCitation id="135AC5F15170F749907EF9B6FA66F925" box="[1288,1424,1655,1681]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Figs 4344</figureCitation>
); abdomen dark grey dorsally, almost black laterally; spinnerets dark grey; venter creamy-grey. Carapace with faintly darker striae radiating from fovea, densely covered with white appressed hairs, cephalic part, and posterior slope with several stiff, erect black setae. Eye field as in genus description (
<figureCitation id="135AC5F15170F74996B0F922FBBAF949" box="[966,1100,1763,1789]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Figs 4344</figureCitation>
). Fovea narrow, longitudinal, straight, dark brown. Anterior surface of cheliceral paturon with numerous long black setae; promargin with low keel ending in small denticle, with additional distal tiny denticle (
<figureCitation id="135AC5F15170F7499611F8EAFC4FF8F1" box="[871,953,1834,1861]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Fig. 45</figureCitation>
; arrow); keel continuing along entire length of promargin; retromargin without keel or denticle. Abdomen oval, hirsute; dorsum anteriorly with narrow, shiny brown scutum extending slightly less than ½ abdominal length, with three pairs of distinct oval sigilla, two pairs close together near posterior of scutum, at ⅓ abdomen length, third pair just behind midpoint (
<figureCitation id="135AC5F15170F74991BFF856FAE8F805" box="[1225,1310,1943,1969]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Fig. 42</figureCitation>
). Metatarsi III and all tarsi with well-developed complete scopulae (
<figureCitation id="135AC5F15170F749965CF87AFC59F861" box="[810,943,1979,2005]" captionStart="FIGURES 4247" captionStartId="10.[151,264,1511,1535]" captionTargetBox="[151,1436,414,1485]" captionTargetId="figure-191@10.[151,1436,414,1485]" captionTargetPageId="10" captionText="FIGURES 4247. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm." pageId="9" pageNumber="48">Figs 4647</figureCitation>
), on metatarsi III and IV restricted distally. Spinnerets hirsute. Body length 5.40. Carapace 2.60 long, 1.76 wide. Abdomen 3.22 long, 1.80 wide. Eye sizes and interdistances: ALE 0.14, AME 0.17, PLE 0.14, PME 0.13; ALEPLE 0.08, AMEAME 0.05, AMEPME 0.16, PMEPLE 0.09, PMEPME 0.10. AME and ALE nearly contiguous. Clypeus height at AMEs 0.12, at ALEs 0.10. Chelicerae 0.83 long. Length of palp and legs: palp 2.10 [0.73, 0.36, 0.23, 0.78], I 5.77 [1.70, 1.05, 1.23, 1.09, 0.70], II 5.53 [1.62, 0.98, 1.19, 1.08, 0.66], III 5.26 [1.56, 0.82, 1.12, 1.13, 0.63], IV 7.01 [1.90, 1.05, 1.43, 1.85, 0.78]. Leg formula: 4123. Spination of palp: femur pl 1 do 2, patella pl 1 do 1, tibia pl 1, tarsus plv 1 rlv 1; legs: femur III pl 1 do 3, IIIIV pl 2 do 3 rl 2; patella I, II, IV spineless, III rl 1; tibia III vt 2, III pl 4 rl 2 plv 1 rlv 1 vt 3, IV pl 3 rl 2 plv 2 rlv 1 vt 2; metatarsus III plv 1 rlv 1, III pl 4 rl 2 plv 1 rlv 1 vt 3, IV pl 4 rl 4 plv 2 rlv 2 vt 3.
</paragraph>
<caption id="DF1E89FC5173F74A95E1FA26FEB0F9DF" pageId="10" pageNumber="49" startId="10.[151,264,1511,1535]" targetBox="[151,1436,414,1485]" targetPageId="10" targetType="figure">
<paragraph id="8BDED9745173F74A95E1FA26FEB0F9DF" blockId="10.[151,1437,1511,1643]" pageId="10" pageNumber="49">
<emphasis id="B91505665173F74A95E1FA26FEA0FA4B" bold="true" box="[151,342,1511,1535]" pageId="10" pageNumber="49">FIGURES 4247.</emphasis>
Somatic morphology of
<taxonomicName id="4C61A2F75173F74A972EFA26FC1EFA4B" authority="Haddad &amp; Sankaran" authorityName="Haddad &amp; Sankaran" authorityYear="2025" box="[600,1000,1511,1535]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B91505665173F74A972EFA26FCE6FA4B" box="[600,784,1511,1535]" italics="true" pageId="10" pageNumber="49">Trichothyse karoo</emphasis>
Haddad &amp; Sankaran
</taxonomicName>
<emphasis id="B91505665173F74A9686FA26FBB4FA4B" bold="true" box="[1008,1090,1511,1535]" pageId="10" pageNumber="49">
<taxonomicNameLabel id="A226B81D5173F74A9686FA26FBB4FA4B" box="[1008,1090,1511,1535]" pageId="10" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male from Jagerskraal: 42. Habitus, dorsal view; 43. Eye region, anterior view; 44. Same, dorsal view; 45. Chelicerae, posteroventral view; 46. Leg I, retrolateral view; 47. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 42 = 1.0 mm; 4347 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDED9745173F74A95B1F97CFB5CF838" blockId="10.[151,1437,1725,2042]" pageId="10" pageNumber="49">
<emphasis id="B91505665173F74A95B1F97CFF0AF963" box="[199,252,1725,1751]" italics="true" pageId="10" pageNumber="49">Palp</emphasis>
(
<figureCitation id="135AC5F15173F74A947CF97CFE9AF96C" box="[266,364,1725,1752]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="10" pageNumber="49">Figs 3, 7</figureCitation>
,
<figureCitation id="135AC5F15173F74A9401F97CFE35F96C" box="[375,451,1725,1752]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="10" pageNumber="49">5455</figureCitation>
): segments yellow-brown, cymbium darker (
<figureCitation id="135AC5F15173F74A96C4F97CFBE2F963" box="[946,1044,1725,1751]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="10" pageNumber="49">Figs 3, 7</figureCitation>
). RTA with stout base, with slender finger-like ventral branch and slightly shorter sharply pointed triangular dorsal branch (
<figureCitation id="135AC5F15173F74A911BF920FB24F948" box="[1133,1234,1761,1788]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="10" pageNumber="49">Figs 3, 7</figureCitation>
,
<figureCitation id="135AC5F15173F74A91A8F920FADFF948" box="[1246,1321,1761,1788]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="10" pageNumber="49">5455</figureCitation>
). Cymbial flange indistinct, approximately half the length of cymbium, smooth along most of its length. Tegulum elongate-oval, sac-like (
<figureCitation id="135AC5F15173F74A944CF8E8FE77F8F7" box="[314,385,1833,1859]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="10" pageNumber="49">Figs 3</figureCitation>
,
<figureCitation id="135AC5F15173F74A94FBF8E8FE5BF8F0" box="[397,429,1833,1860]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="10" pageNumber="49">54</figureCitation>
). Conductor membranous, passing behind embolus, with proximal part parallel to embolus and smoothly rounded apex directed retrolaterally (
<figureCitation id="135AC5F15173F74A9795F88CFCBBF8DC" box="[739,845,1869,1896]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="10" pageNumber="49">Figs 3, 7</figureCitation>
,
<figureCitation id="135AC5F15173F74A962DF88CFC50F8DC" box="[859,934,1869,1896]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="10" pageNumber="49">5455</figureCitation>
). Embolus arising prolaterally, then curving retrolaterally at distal part to form a claw-shaped tip, directed at 3-oclock (
<figureCitation id="135AC5F15173F74A9695F8B0FBBEF838" box="[995,1096,1905,1932]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="10" pageNumber="49">Figs 3, 7</figureCitation>
,
<figureCitation id="135AC5F15173F74A9122F8B0FB68F838" box="[1108,1182,1905,1932]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="10" pageNumber="49">5455</figureCitation>
).
</paragraph>
<paragraph id="8BDED9745173F74B95B1F854FD8CFE65" blockId="10.[151,1437,1725,2042]" lastBlockId="11.[151,1437,151,789]" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49">
<emphasis id="B91505665173F74A95B1F854FEEDF81B" box="[199,283,1941,1967]" italics="true" pageId="10" pageNumber="49">Female</emphasis>
(
<typeStatus id="54DA67D65173F74A945CF854FE79F81B" box="[298,399,1941,1967]" pageId="10" pageNumber="49" type="paratype">paratype</typeStatus>
, Matjiesfontein, NCA 2016/2243,
<figureCitation id="135AC5F15173F74A9667F854FC62F804" box="[785,916,1941,1968]" captionStart="FIGURES 4853" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[151,1436,810,1880]" captionTargetId="figure-357@11.[151,1436,810,1880]" captionTargetPageId="11" captionText="FIGURES 4853. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., paratype female from Jagerskraal: 48. Habitus, dorsal view; 49. Eye region, anterior view; 50. Same, dorsal view; 51. Chelicerae, posteroventral view; 52. Leg I, retrolateral view; 53. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 48 = 1 mm; 4953 = 0.5 mm." pageId="10" pageNumber="49">Figs 4853</figureCitation>
). General aspects essentially as in male except the following: habitus colour lighter than male, legs yellow-brown, with metatarsi and tarsi slightly darker (
<figureCitation id="135AC5F15173F74A901CF878FEFBF84C" captionStart="FIGURES 4853" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[151,1436,810,1880]" captionTargetId="figure-357@11.[151,1436,810,1880]" captionTargetPageId="11" captionText="FIGURES 4853. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., paratype female from Jagerskraal: 48. Habitus, dorsal view; 49. Eye region, anterior view; 50. Same, dorsal view; 51. Chelicerae, posteroventral view; 52. Leg I, retrolateral view; 53. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 48 = 1 mm; 4953 = 0.5 mm." pageId="10" pageNumber="49">Figs 48, 5253</figureCitation>
). Abdomen without dorsal scutum, with three pairs of distinct oval sigilla, first two pairs at ¼ abdomen length and third pair at ½ abdomen length (
<figureCitation id="135AC5F15172F74B97F1FF56FD2BFF05" box="[647,733,151,177]" captionStart="FIGURES 4853" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[151,1436,810,1880]" captionTargetId="figure-357@11.[151,1436,810,1880]" captionTargetPageId="11" captionText="FIGURES 4853. Somatic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov., paratype female from Jagerskraal: 48. Habitus, dorsal view; 49. Eye region, anterior view; 50. Same, dorsal view; 51. Chelicerae, posteroventral view; 52. Leg I, retrolateral view; 53. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 48 = 1 mm; 4953 = 0.5 mm." pageId="11" pageNumber="50">Fig. 48</figureCitation>
). Body length 7.12. Carapace 2.86 long, 1.80 wide. Abdomen 3.91 long, 2.16 wide. Eye sizes and interdistances: ALE 0.16, AME 0.17, PLE 0.15, PME 0.14; ALEPLE 0.13, AMEAME 0.09, AMEPME 0.15, PMEPLE 0.10, PMEPME 0.11. Clypeus height at AMEs 0.11, at ALEs 0.10. Chelicerae 0.95 long. Length of palp and legs: palp 2.24 [0.81, 0.35, 0.36, 0.72], I 6.08 [1.80, 1.10, 1.28, 1.12, 0.78], II 6.03 [1.79, 1.11, 1.28, 1.17, 0.68], III 5.72 [1.65, 0.92, 1.12, 1.23, 0.80], IV 7.41 [2.04, 1.10, 1.57, 1.88, 0.82]. Leg formula: 4123. Spination of palp: femur pl 1 do 2, patella pl 1, tibia pl 2 do 1 rl 1 plv 2, tarsus pl 1 rl 1 plv 2 rlv 2 vt 2; legs: femur I pl 1 do 3, II pl 2 do 3, III pl 2 do 3 rl 2, IV pl 1 do 2 rl 1; patella I, II and IV spineless, III pl 1 rl 1; tibia III vt 1, III pl 4 rl 2 plv 2 rlv 1 vt 2, IV pl 3 rl 2 plv 2 rlv 1 vt 2; metatarsus I plv 1, II plv 1 rlv 1, III pl 4 rl 3 plv 1 rlv 1 vt 3, IV pl 4 rl 4 plv 2 rlv 2 vt 4.
</paragraph>
<paragraph id="8BDED9745172F74B95B1FE1AFED8FD79" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="50">
<emphasis id="B91505665172F74B95B1FE1AFEC7FE41" box="[199,305,475,501]" italics="true" pageId="11" pageNumber="50">Genitalia</emphasis>
(
<figureCitation id="135AC5F15172F74B9432FE1AFE3AFE41" box="[324,460,475,501]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="11" pageNumber="50">Figs 11, 17</figureCitation>
,
<figureCitation id="135AC5F15172F74B94AAFE1BFDD1FE41" box="[476,551,474,501]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="11" pageNumber="50">5657</figureCitation>
): epigyne originally hirsute, hairs lost during cleaning, with narrow median hourglass-shaped ridge in posterior half formed by posterior lobes, terminating anteriorly in small teardrop-shaped atrium in centre of epigyne (
<figureCitation id="135AC5F15172F74B9490FDE2FDB6FD89" box="[486,576,547,573]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="11" pageNumber="50">Figs 11</figureCitation>
,
<figureCitation id="135AC5F15172F74B9739FDE3FD98FD89" box="[591,622,546,573]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="11" pageNumber="50">56</figureCitation>
; arrow). Copulatory openings indistinct, at posterior edge of posterior lobes. Copulatory ducts very short, partly obscured by primary spermathecae (
<figureCitation id="135AC5F15172F74B917CFD86FB94FDD5" box="[1034,1122,583,609]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="11" pageNumber="50">Figs 17</figureCitation>
,
<figureCitation id="135AC5F15172F74B9118FD87FB78FDD5" box="[1134,1166,582,609]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="11" pageNumber="50">57</figureCitation>
). Primary spermathecae parallel-sided, separated by slightly less than half their width (
<figureCitation id="135AC5F15172F74B962CFDAAFC44FD31" box="[858,946,619,645]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="11" pageNumber="50">Figs 17</figureCitation>
,
<figureCitation id="135AC5F15172F74B96C9FDABFC29FD31" box="[959,991,618,645]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="11" pageNumber="50">57</figureCitation>
). Paired secondary spermathecae short, globular, partly obscured by primary spermathecae (
<figureCitation id="135AC5F15172F74B979CFD4EFCB4FD1D" box="[746,834,655,681]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="11" pageNumber="50">Figs 17</figureCitation>
,
<figureCitation id="135AC5F15172F74B9626FD4FFC86FD1D" box="[848,880,654,681]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="11" pageNumber="50">57</figureCitation>
). Fertilization ducts broad, undulating, diverging (
<figureCitation id="135AC5F15172F74B95E9FD72FF00FD79" box="[159,246,691,717]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="11" pageNumber="50">Figs 17</figureCitation>
,
<figureCitation id="135AC5F15172F74B9474FD73FED4FD79" box="[258,290,690,717]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="11" pageNumber="50">57</figureCitation>
).
</paragraph>
<paragraph id="8BDED9745172F74B95B1FD17FC9EFD45" blockId="11.[151,1437,151,789]" box="[199,872,726,753]" pageId="11" pageNumber="50">
<emphasis id="B91505665172F74B95B1FD17FEC9FD44" bold="true" box="[199,319,726,752]" pageId="11" pageNumber="50">Variation.</emphasis>
Male (n = 4): body length 4.606.05, mean 5.28.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5172F74B95B1FD3BFB2BFCA1" box="[199,1245,762,789]" pageId="11" pageNumber="50" type="distribution">
<paragraph id="8BDED9745172F74B95B1FD3BFB2BFCA1" blockId="11.[151,1437,151,789]" box="[199,1245,762,789]" pageId="11" pageNumber="50">
<emphasis id="B91505665172F74B95B1FD3BFE97FCA0" bold="true" box="[199,353,762,788]" pageId="11" pageNumber="50">Distribution.</emphasis>
Known only from the semi-arid south-western parts of
<collectingCountry id="F37699E45172F74B96A7FD3AFB96FCA1" box="[977,1120,763,789]" name="South Africa" pageId="11" pageNumber="50">South Africa</collectingCountry>
(
<figureCitation id="135AC5F15172F74B9118FD3AFB27FCA1" box="[1134,1233,763,789]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="11" pageNumber="50">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,211 @@
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<mods:title id="C7FA6F1DF896DCB8AEFDC4E63F82EA2D">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:identifier id="308CB32790959A17602D22A55650F6D7" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<subSubSection id="C37B8AFF5174F74D95E1FCF1FD8EFCFF" box="[151,632,816,843]" pageId="13" pageNumber="52" type="nomenclature">
<paragraph id="8BDED9745174F74D95E1FCF1FD8EFCFF" blockId="13.[151,632,816,880]" box="[151,632,816,843]" pageId="13" pageNumber="52">
<heading id="D0966E185174F74D95E1FCF1FD8EFCFF" bold="true" box="[151,632,816,843]" fontSize="11" level="1" pageId="13" pageNumber="52" reason="1">
<taxonomicName id="4C61A2F75174F74D95E1FCF1FD8EFCFF" ID-CoL="7CT7J" authority="Lawrence, 1927" authorityName="Lawrence" authorityYear="1927" box="[151,632,816,843]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="subtropica">
<emphasis id="B91505665174F74D95E1FCF1FD8EFCFF" bold="true" box="[151,632,816,843]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95E1FCF1FE5AFCFF" bold="true" box="[151,428,816,843]" italics="true" pageId="13" pageNumber="52">Trichothyse subtropica</emphasis>
<bibRefCitation id="EFF0A4855174F74D94C5FCF0FD8EFCFF" author="Lawrence, R. F." box="[435,632,817,843]" pageId="13" pageNumber="52" pagination="1 - 75" refId="ref19265" refString="Lawrence, R. F. (1927) Contributions to a knowledge of the fauna of South-West Africa V. Arachnida. Annals of the South African Museum, 25, 1-75." type="journal article" year="1927">Lawrence, 1927</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74D95E1FC94FE9EFCC4" box="[151,360,853,880]" pageId="13" pageNumber="52" type="description">
<paragraph id="8BDED9745174F74D95E1FC94FE9EFCC4" blockId="13.[151,632,816,880]" box="[151,360,853,880]" pageId="13" pageNumber="52">
<figureCitation id="135AC5F15174F74D95E1FC94FEE0FCDB" box="[151,278,853,879]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Figs 12, 18</figureCitation>
,
<figureCitation id="135AC5F15174F74D9454FC94FE9EFCC4" box="[290,360,853,880]" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">5863</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74D95E1FC58FB80FC64" pageId="13" pageNumber="52" type="reference_group">
<paragraph id="8BDED9745174F74D95E1FC58FB80FC64" blockId="13.[151,1437,921,976]" pageId="13" pageNumber="52">
<treatmentCitationGroup id="AB71FE5A5174F74D95E1FC58FB80FC64" pageId="13" pageNumber="52">
<treatmentCitation id="0AC0FF655174F74D95E1FC58FDAFFC05" author="Lawrence, R. F." box="[151,601,921,945]" page="15" pageId="13" pageNumber="52" year="1927">
<taxonomicName id="4C61A2F75174F74D95E1FC58FDAFFC05" ID-CoL="7CT7J" authority="Lawrence, 1927: 15" authorityName="Lawrence" authorityPageNumber="15" authorityYear="1927" box="[151,601,921,945]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="subtropica">
<emphasis id="B91505665174F74D95E1FC58FE77FC05" box="[151,385,921,945]" italics="true" pageId="13" pageNumber="52">Trichothyse subtropica</emphasis>
<materialsCitation id="3B09D3295174F74D94FFFC5BFDAFFC05" box="[393,601,921,945]" collectingDate="1923-03" collectorName="R. F. Lawrence" country="Namibia" latitude="-19.066668" location="River" longLatPrecision="1273" longitude="14.216666" pageId="13" pageNumber="52" specimenCount="2" specimenCount-female="2" stateProvince="Kunene" typeStatus="lectotype">
<bibRefCitation id="EFF0A4855174F74D94FFFC5BFDAFFC05" author="Lawrence, R. F." box="[393,601,921,945]" pageId="13" pageNumber="52" pagination="1 - 75" refId="ref19265" refString="Lawrence, R. F. (1927) Contributions to a knowledge of the fauna of South-West Africa V. Arachnida. Annals of the South African Museum, 25, 1-75." type="journal article" year="1927">Lawrence, 1927: 15</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, plate I, fig. 15 (
<typeStatus id="54DA67D65174F74D967FFC58FC9EFC05" box="[777,872,921,945]" pageId="13" pageNumber="52" type="lectotype">lectotype</typeStatus>
<specimenCount id="9D6712FD5174F74D9606FC5BFC75FC05" box="[880,899,922,945]" count="1" pageId="13" pageNumber="52" type="female"></specimenCount>
, here designated, and
<typeStatus id="54DA67D65174F74D911BFC58FB03FC05" box="[1133,1269,921,945]" pageId="13" pageNumber="52" type="paralectotype">paralectotype</typeStatus>
<specimenCount id="9D6712FD5174F74D9188FC5BFAE5FC05" box="[1278,1299,922,945]" count="1" pageId="13" pageNumber="52" type="female"></specimenCount>
:
<emphasis id="B91505665174F74D9056FC58FA6BFC05" bold="true" box="[1312,1437,921,945]" pageId="13" pageNumber="52">
<collectingCountry id="F37699E45174F74D9056FC58FA61FC05" box="[1312,1431,921,945]" name="Namibia" pageId="13" pageNumber="52">NAMIBIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49A517965174F74D95B1FC79FEE0FC7B" box="[199,278,952,975]" country="Namibia" name="Kunene" pageId="13" pageNumber="52">Kunene</collectingRegion>
<location id="8EBE8FAF5174F74D946AFC79FEA2FC64" LSID="urn:lsid:plazi:treatment:03C868625174F74E95E1FCF1FEF8FEA9:8EBE8FAF5174F74D946AFC79FEA2FC64" box="[284,340,952,976]" country="Namibia" latitude="-19.066668" longLatPrecision="1273" longitude="14.216666" name="River" pageId="13" pageNumber="52" stateProvince="Kunene">River</location>
[
<geoCoordinate id="EE55BFB35174F74D9414FC79FE4FFC64" box="[354,441,952,976]" degrees="19" direction="south" minutes="04" orientation="latitude" pageId="13" pageNumber="52" precision="925" value="-19.066668">19°04S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35174F74D94B5FC79FDEDFC64" box="[451,539,952,976]" degrees="14" direction="east" minutes="13" orientation="longitude" pageId="13" pageNumber="52" precision="925" value="14.216666">14°13E</geoCoordinate>
],
<date id="FFDFFFB45174F74D975BFC79FD77FC64" box="[557,641,952,976]" pageId="13" pageNumber="52" value="1923-03">
<collectingDate id="EF9B065C5174F74D975BFC79FD77FC64" box="[557,641,952,976]" pageId="13" pageNumber="52" value="1923-03">III.1923</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25174F74D97CEFC79FCB9FC7B" box="[696,847,952,975]" pageId="13" pageNumber="52">R.F. Lawrence</collectorName>
, SAMC B6143—examined).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74D95E1FC3FFCC4FBD4" pageId="13" pageNumber="52" type="discussion">
<paragraph id="8BDED9745174F74D95E1FC3FFCC4FBD4" blockId="13.[151,1437,1022,2021]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95E1FC3FFEF6FBAC" bold="true" box="[151,256,1022,1048]" pageId="13" pageNumber="52">Remark.</emphasis>
The
<typeStatus id="54DA67D65174F74D9434FC3FFE5FFBAC" box="[322,425,1022,1048]" pageId="13" pageNumber="52" type="lectotype">lectotype</typeStatus>
designated here is the larger of the
<specimenCount id="9D6712FD5174F74D962CFC3EFC10FBAC" box="[858,998,1022,1049]" count="2" pageId="13" pageNumber="52" type="female">two females</specimenCount>
in the type repository. The types are quite obviously faded in colour, and the true colouration of this species is probably consistent with the congeners (re)described in this paper from recently collected material.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74D95B1FBABFAC3FB78" pageId="13" pageNumber="52" type="diagnosis">
<paragraph id="8BDED9745174F74D95B1FBABFAC3FB78" blockId="13.[151,1437,1022,2021]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95B1FBABFEB4FB30" bold="true" box="[199,322,1130,1156]" pageId="13" pageNumber="52">Diagnosis.</emphasis>
Among the Afrotropical species, the females of
<taxonomicName id="4C61A2F75174F74D961AFBAAFBF6FB30" authorityName="Lawrence" authorityYear="1927" box="[876,1024,1130,1156]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="subtropica">
<emphasis id="B91505665174F74D961AFBAAFBF6FB30" box="[876,1024,1130,1156]" italics="true" pageId="13" pageNumber="52">T. subtropica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C61A2F75174F74D914FFBAAFB49FB30" authorityName="Tucker" authorityYear="1923" box="[1081,1215,1130,1157]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="hortensis">
<emphasis id="B91505665174F74D914FFBAAFBBFFB30" box="[1081,1097,1131,1156]" italics="true" pageId="13" pageNumber="52">T</emphasis>
.
<emphasis id="B91505665174F74D912EFBABFB49FB30" box="[1112,1215,1130,1156]" italics="true" pageId="13" pageNumber="52">hortensis</emphasis>
</taxonomicName>
share the convexly shaped primary spermathecae, but
<taxonomicName id="4C61A2F75174F74D9763FB4EFD50FB1C" authorityName="Lawrence" authorityYear="1927" box="[533,678,1166,1192]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="subtropica">
<emphasis id="B91505665174F74D9763FB4EFD50FB1C" box="[533,678,1166,1192]" italics="true" pageId="13" pageNumber="52">T. subtropica</emphasis>
</taxonomicName>
can be recognised by the less strongly curved primary spermathecae, while separated by more than 1.5 times the width in
<taxonomicName id="4C61A2F75174F74D9794FB72FC93FB78" authorityName="Tucker" authorityYear="1923" box="[738,869,1202,1228]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="hortensis">
<emphasis id="B91505665174F74D9794FB72FC93FB78" box="[738,869,1202,1228]" italics="true" pageId="13" pageNumber="52">T. hortensis</emphasis>
</taxonomicName>
(cf.
<figureCitation id="135AC5F15174F74D96EFFB73FC1EFB78" box="[921,1000,1202,1228]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Fig. 18</figureCitation>
and
<figureCitation id="135AC5F15174F74D9168FB73FB87FB78" box="[1054,1137,1202,1228]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Fig. 16</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74E95B1FB17FF35FF61" lastPageId="14" lastPageNumber="53" pageId="13" pageNumber="52" type="description">
<paragraph id="8BDED9745174F74D95B1FB17FD49F8E0" blockId="13.[151,1437,1022,2021]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95B1FB17FEAFFB44" bold="true" box="[199,345,1238,1264]" pageId="13" pageNumber="52">Description.</emphasis>
<emphasis id="B91505665174F74D9415FB17FE41FB44" box="[355,439,1238,1264]" italics="true" pageId="13" pageNumber="52">Female</emphasis>
(
<typeStatus id="54DA67D65174F74D94BFFB17FDC2FB44" box="[457,564,1238,1264]" pageId="13" pageNumber="52" type="lectotype">lectotype</typeStatus>
,
<collectingRegion id="49A517965174F74D9735FB16FD6CFB45" box="[579,666,1239,1265]" country="Namibia" name="Kunene" pageId="13" pageNumber="52">Kunene</collectingRegion>
River, SAMC B6143,
<figureCitation id="135AC5F15174F74D96DEFB17FBDBFB45" box="[936,1069,1238,1265]" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">Figs 5863</figureCitation>
). Carapace, clypeus, chelicerae, labium, endites, sternum, scutum and legs orange-brown (
<figureCitation id="135AC5F15174F74D965EFB3BFC5BFAA1" box="[808,941,1274,1301]" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">Figs 5861</figureCitation>
). Abdomen uniform grey (
<figureCitation id="135AC5F15174F74D91ADFB3BFAC6FAA1" box="[1243,1328,1274,1301]" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">Fig. 58</figureCitation>
), anterior lateral spinnerets yellow-brown, posterior, median and lateral spinnerets creamy-grey; carapace with faintly darker striae radiating from fovea, densely covered with white appressed hairs; clypeus and posterior slope provided with numerous erect stiff black setae. Fovea narrow, longitudinal, straight, dark brown. Anterior surface of cheliceral paturon with long black setae, promargin with low keel ending in distal tooth, with small denticle beyond it (
<figureCitation id="135AC5F15174F74D9019FA4BFF40FA7C" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">Fig. 61</figureCitation>
; arrow); retromargin with single tiny basal denticle. Abdomen oval, hirsute; dorsum without scutum, with three pairs of distinct oval sigilla, two pairs at ¼ abdomen length and third pair at midpoint (
<figureCitation id="135AC5F15174F74D911FFA13FB48FA59" box="[1129,1214,1490,1517]" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">Fig. 58</figureCitation>
). Metatarsi III and all tarsi with well-developed complete scopulae (
<figureCitation id="135AC5F15174F74D97B2FA37FCB1F9A4" box="[708,839,1526,1552]" captionStart="FIGURES 5863" captionStartId="14.[151,264,1409,1433]" captionTargetBox="[151,1436,306,1383]" captionTargetId="figure-52@14.[151,1436,306,1383]" captionTargetPageId="14" captionText="FIGURES 5863. Somatic morphology of Trichothyse subtropica Lawrence, 1927, lectotype female: 58. Habitus, dorsal view; 59. Eye region, anterior view; 60. Same, dorsal view; 61. Chelicerae, posteroventral view; 62. Legs I &amp; II, retrolateral view; 63. Leg IV, retrolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 58 = 1.0 mm; 5963 = 0.5 mm." pageId="13" pageNumber="52">Figs 6263</figureCitation>
), scopulae on metatarsi III and IV restricted distally. Spinnerets hirsute. Body length 12.10. Carapace 4.40 long, 3.14 wide.Abdomen 7.62 long, 4.39 wide. Eye sizes and interdistances: ALE 0.26, AME 0.23, PLE 0.19, PME 0.17; ALEPLE 0.16, AMEAME 0.10, AMEPME 0.25, PMEPLE 0.17, PMEPME 0.21. AME and ALE nearly contiguous. Clypeus height at AMEs 0.19, at ALEs 0.17. Chelicerae 1.48 long. Length of palp and legs: palp 2.93 [1.11, 0.54, 0.50, 0.78], I 8.94 [2.65, 1.70, 1.93, 1.71, 0.95], II 8.92 [2.59, 1.73, 1.89, 1.70, 1.01], III 8.54 [2.55, 1.35, 1.72, 1.93, 0.99], IV 10.66 [2.98, 1.58, 2.28, 2.81, 1.01]. Leg formula: 4123. Spination of palp: femur pl 2 do 2, patella pl 1, tibia pl 2 rl 1 rlv 1, tarsus plv 2 rlv 2 vt 2; legs: femur I pl 1 do 3, II pl 2 do 3, III pl 2 do 3 rl 2, IV pl 1 do 3 rl 1; patella III spineless, III pl 1 rl 1, IV rl 1; tibia III vt 1, III pl 3 rl 2 plv 2 rlv 1 vt 2, IV pl 2 rl 2 plv 2 rlv 2 vt 2; metatarsus III plv 1 rlv 1, III pl 3 rl 3 plv 1 rlv 1 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3; tarsus IIV spineless.
</paragraph>
<paragraph id="8BDED9745174F74E95B1F89FFF35FF61" blockId="13.[151,1437,1022,2021]" lastBlockId="14.[151,1437,151,285]" lastPageId="14" lastPageNumber="53" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95B1F89FFEC7F8CC" box="[199,305,1886,1912]" italics="true" pageId="13" pageNumber="52">Genitalia</emphasis>
(
<figureCitation id="135AC5F15174F74D9437F89FFE30F8CC" box="[321,454,1886,1912]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Figs 12, 18</figureCitation>
): epigyne hirsute, with paired S-shaped margins forming a broad shallow atrium (
<figureCitation id="135AC5F15174F74D9019F89FFF40F828" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Fig. 12</figureCitation>
; arrow). Copulatory openings indistinct, hidden behind anterior edge of posterior lobes, visible when epigyne is rotated on its longitudinal axis. Copulatory ducts short, and secondary spermathecae entirely obscured by primary spermathecae (
<figureCitation id="135AC5F15174F74D9436F80BFE62F850" box="[320,404,1994,2020]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Fig. 18</figureCitation>
). Primary spermathecae convex, forming arrowhead-shaped space between them, separated by less than the width of each spermatheca (
<figureCitation id="135AC5F15177F74E97F4FF56FD23FF05" box="[642,725,151,177]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="14" pageNumber="53">Fig. 18</figureCitation>
). Fertilization ducts narrow, slightly undulating, diverging (
<figureCitation id="135AC5F15177F74E9019FF56FF40FF61" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="14" pageNumber="53">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5177F74E95B1FF1FFEF8FEA9" pageId="14" pageNumber="53" type="distribution">
<paragraph id="8BDED9745177F74E95B1FF1FFEF8FEA9" blockId="14.[151,1437,151,285]" pageId="14" pageNumber="53">
<emphasis id="B91505665177F74E95B1FF1FFE97FF4C" bold="true" box="[199,353,222,248]" pageId="14" pageNumber="53">Distribution.</emphasis>
Known only from the
<typeStatus id="54DA67D65177F74E971CFF1EFD6CFF4D" box="[618,666,223,249]" pageId="14" pageNumber="53">type</typeStatus>
locality, on the north-western border between
<collectingCountry id="F37699E45177F74E91C7FF1EFAE7FF4D" box="[1201,1297,223,249]" name="Namibia" pageId="14" pageNumber="53">Namibia</collectingCountry>
and
<collectingCountry id="F37699E45177F74E903CFF1EFA6AFF4D" box="[1354,1436,223,249]" name="Angola" pageId="14" pageNumber="53">Angola</collectingCountry>
(
<figureCitation id="135AC5F15177F74E95E9FEC2FEF4FEA9" box="[159,258,259,285]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="14" pageNumber="53">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,237 @@
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<mods:title id="5C64C5AFCE3AFAD226EB1C1881089906">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<heading id="D0966E185175F74C95E1F8BAFC6CF822" bold="true" box="[151,922,1915,1942]" fontSize="11" level="1" pageId="12" pageNumber="51" reason="1">
<emphasis id="B91505665175F74C95E1F8BAFC6CF822" bold="true" box="[151,922,1915,1942]" pageId="12" pageNumber="51">
<taxonomicName id="4C61A2F75175F74C95E1F8BAFCFBF822" authority="(O. Pickard-Cambridge, 1874)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1874" box="[151,781,1915,1942]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="pugnax" status="comb. nov.">
<emphasis id="B91505665175F74C95E1F8BAFE7EF822" bold="true" box="[151,392,1915,1942]" italics="true" pageId="12" pageNumber="51">Trichothyse pugnax</emphasis>
(O.
<bibRefCitation id="EFF0A4855175F74C94C9F8BDFCF2F822" author="Pickard-Cambridge, O." box="[447,772,1916,1942]" pageId="12" pageNumber="51" pagination="370 - 419" refId="ref19799" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42, 370-419, pls. LI - LII. https://doi.org/10.1111/j.1096-3642.1874.tb02495.x" type="journal article" year="1874">Pickard-Cambridge, 1874</bibRefCitation>
)
</taxonomicName>
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</emphasis>
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<treatmentCitation id="0AC0FF655175F74C95E1F800FD53F86D" author="Pickard-Cambridge, O." box="[151,677,1984,2009]" page="339" pageId="12" pageNumber="51" year="1874">
<taxonomicName id="4C61A2F75175F74C95E1F800FD53F86D" authority="O. Pickard-Cambridge, 1874: 339" authorityName="O. Pickard-Cambridge" authorityPageNumber="339" authorityYear="1874" box="[151,677,1984,2009]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="51" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B91505665175F74C95E1F800FECBF86C" box="[151,317,1985,2008]" italics="true" pageId="12" pageNumber="51">Drassus pugnax</emphasis>
<materialsCitation id="3B09D3295175F74C9433F800FD53F86D" box="[325,677,1984,2009]" collectionCode="OUMNH" collectorName="O. Pickard-Cambridge" country="Egypt" latitude="30.033333" location="Cairo" longLatPrecision="1224" longitude="31.233334" pageId="12" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Al Qahirah" typeStatus="holotype">
O.
<bibRefCitation id="EFF0A4855175F74C9410F801FD53F86D" author="Pickard-Cambridge, O." box="[358,677,1984,2009]" pageId="12" pageNumber="51" pagination="370 - 419" refId="ref19799" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42, 370-419, pls. LI - LII. https://doi.org/10.1111/j.1096-3642.1874.tb02495.x" type="journal article" year="1874">Pickard-Cambridge, 1874: 339</bibRefCitation>
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, plate LII, fig. 25 (
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<specimenCount id="9D6712FD5175F74C96B9F800FC13F86C" box="[975,997,1985,2008]" count="1" pageId="12" pageNumber="51" type="male"></specimenCount>
:
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<collectingCountry id="F37699E45175F74C9687F801FBBCF86C" box="[1009,1098,1984,2008]" name="Egypt" pageId="12" pageNumber="51">EGYPT</collectingCountry>
:
</emphasis>
<collectingRegion id="49A517965175F74C9121F801FB79F86C" box="[1111,1167,1984,2008]" country="Egypt" name="Al Qahirah" pageId="12" pageNumber="51">Cairo</collectingRegion>
[
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,
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], date unknown, leg.
<collectorName id="2694BCA25175F74C9428F81EFDB0F843" box="[350,582,2015,2039]" pageId="12" pageNumber="51">O. Pickard-Cambridge</collectorName>
, repository OUMNH (B. 254)—not examined).
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<treatmentCitationGroup id="AB71FE5A5174F74D95E1FF56FD5BFF1B" box="[151,685,151,175]" pageId="13" pageNumber="52">
<treatmentCitation id="0AC0FF655174F74D95E1FF56FDECFF1B" author="Karsch, F." box="[151,538,151,175]" page="12" pageId="13" pageNumber="52" year="1881">
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<emphasis id="B91505665174F74D95E1FF56FE92FF1B" box="[151,356,151,175]" italics="true" pageId="13" pageNumber="52">Drassus sockniensis</emphasis>
<bibRefCitation id="EFF0A4855174F74D941CFF56FDECFF1B" author="Karsch, F." box="[362,538,151,175]" pageId="13" pageNumber="52" pagination="1 - 14" refId="ref19073" refString="Karsch, F. (1881) Verzeichniss der wahrend der Rohlfs'schen Afrikanischen Expedition erbeuteten Myriopoden und Arachniden. Archiv fur Naturgeschichte, 47, 1-14. https://doi.org/10.5962/bhl.part.13207" type="journal article" year="1881">Karsch, 1881: 12</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, plate I, fig. 9.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745174F74D95E1FF74FB04FF79" blockId="13.[151,1266,151,267]" box="[151,1266,181,206]" pageId="13" pageNumber="52">
<treatmentCitationGroup id="AB71FE5A5174F74D95E1FF74FB04FF79" box="[151,1266,181,206]" pageId="13" pageNumber="52">
<treatmentCitation id="0AC0FF655174F74D95E1FF74FDD8FF79" author="Simon, E." box="[151,558,181,205]" page="423" pageId="13" pageNumber="52" year="1908">
<taxonomicName id="4C61A2F75174F74D95E1FF74FDD8FF79" ID-CoL="4KTCB" authority="Simon, 1908: 423" authorityName="Simon" authorityPageNumber="423" authorityYear="1908" box="[151,558,181,205]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B91505665174F74D95E1FF74FE86FF79" box="[151,368,181,205]" italics="true" pageId="13" pageNumber="52">Poecilochroa pugnax</emphasis>
<bibRefCitation id="EFF0A4855174F74D9400FF74FDD8FF79" author="Simon, E." box="[374,558,181,205]" pageId="13" pageNumber="52" pagination="419 - 438" refId="ref20149" refString="Simon, E. (1908) Etude sur les arachnides recueillis par M. le Dr. Klaptocz en Tripolitaine. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere, 26, 419-438." type="journal article" year="1908">Simon, 1908: 423</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AC0FF655174F74D974EFF77FD29FF79" author="Levy, G." box="[568,735,181,206]" page="433" pageId="13" pageNumber="52" year="1999">
<bibRefCitation id="EFF0A4855174F74D974EFF77FD29FF79" author="Levy, G." box="[568,735,181,206]" pageId="13" pageNumber="52" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy, 1999: 433</bibRefCitation>
</treatmentCitation>
, figs 1014;
<treatmentCitation id="0AC0FF655174F74D9615FF77FB71FF79" author="Bosmans, R. &amp; Herve, C." box="[867,1159,181,205]" page="60" pageId="13" pageNumber="52" year="2021">
<bibRefCitation id="EFF0A4855174F74D9615FF77FB71FF79" author="Bosmans, R. &amp; Herve, C." box="[867,1159,181,205]" pageId="13" pageNumber="52" pagination="58 - 64" refId="ref18430" refString="Bosmans, R. &amp; Herve, C. (2021) Less is more: eight new synonyms in Mediterranean spiders (Araneae), with a new Pelecopsis species from Tunisia (Linyphiidae). Arachnologische Mitteilungen, 61, 58-64. https://doi.org/10.30963/aramit6109" type="journal article" year="2021">Bosmans &amp; Hervé, 2021: 60</bibRefCitation>
</treatmentCitation>
, fig. 2de.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745174F74D95E1FF15FDB7FF58" blockId="13.[151,1266,151,267]" box="[151,577,212,236]" pageId="13" pageNumber="52">
<treatmentCitationGroup id="AB71FE5A5174F74D95E1FF15FDB7FF58" box="[151,577,212,236]" pageId="13" pageNumber="52">
<treatmentCitation id="0AC0FF655174F74D95E1FF15FDCBFF58" author="Strand, E." box="[151,573,212,236]" page="62" pageId="13" pageNumber="52" year="1908">
<taxonomicName id="4C61A2F75174F74D95E1FF15FDCBFF58" authority="Strand, 1908: 62" authorityName="Strand" authorityPageNumber="62" authorityYear="1908" box="[151,573,212,236]" class="Arachnida" family="Gnaphosidae" genus="Scotophaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="subpugnax">
<emphasis id="B91505665174F74D95E1FF15FE7AFF58" box="[151,396,212,236]" italics="true" pageId="13" pageNumber="52">Scotophaeus subpugnax</emphasis>
<bibRefCitation id="EFF0A4855174F74D94E4FF15FDCBFF58" author="Strand, E." box="[402,573,212,236]" pageId="13" pageNumber="52" pagination="11 - 101" refId="ref20228" refString="Strand, E. (1908) Nordafrikanische, hauptsachlich von Carlo Freiherr von Erlanger gesammelte Aviculariidae, Drassidae und Theridiidae. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg, 64, 11-101. https://doi.org/10.5962/bhl.part.27907" type="journal article" year="1908">Strand, 1908: 62</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDED9745174F74D95E1FF32FD10FEBF" blockId="13.[151,1266,151,267]" box="[151,742,243,267]" pageId="13" pageNumber="52">
<treatmentCitationGroup id="AB71FE5A5174F74D95E1FF32FD10FEBF" box="[151,742,243,267]" pageId="13" pageNumber="52">
<treatmentCitation id="0AC0FF655174F74D95E1FF32FDEEFEBF" author="Denis, J." box="[151,536,243,267]" page="62" pageId="13" pageNumber="52" year="1947">
<taxonomicName id="4C61A2F75174F74D95E1FF32FDEEFEBF" ID-CoL="4KTC3" authority="Denis, 1947: 62" authorityName="Denis" authorityPageNumber="62" authorityYear="1947" box="[151,536,243,267]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="lesserti">
<emphasis id="B91505665174F74D95E1FF32FE98FEBF" box="[151,366,243,267]" italics="true" pageId="13" pageNumber="52">Poecilochroa lesserti</emphasis>
<bibRefCitation id="EFF0A4855174F74D9403FF32FDEEFEBF" author="Denis, J." box="[373,536,243,267]" pageId="13" pageNumber="52" pagination="17 - 103" refId="ref18678" refString="Denis, J. (1947) Results of the Armstrong College expedition to Siwa Oasis (Libyan desert), 1935. Spiders [Araneae]. Bulletin de la Societe Fouad 1 er d'Entomologie, 31, 17-103, pls. I - VI." type="journal article" year="1947">Denis, 1947: 62</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, plate III, figs 810.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74D95E1FEF8FB31FD0F" pageId="13" pageNumber="52" type="description">
<paragraph id="8BDED9745174F74D95E1FEF8FB83FEE7" blockId="13.[151,1437,313,735]" box="[151,1141,313,339]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95E1FEF8FEDFFEE7" bold="true" box="[151,297,313,339]" pageId="13" pageNumber="52">Description.</emphasis>
<emphasis id="B91505665174F74D9446FEF8FE9CFEE7" box="[304,362,313,339]" italics="true" pageId="13" pageNumber="52">Male</emphasis>
and
<emphasis id="B91505665174F74D94D6FEF8FE1DFEE7" box="[416,491,313,339]" italics="true" pageId="13" pageNumber="52">female</emphasis>
. For description of the male and female, see
<bibRefCitation id="EFF0A4855174F74D9695FEF8FB87FEE7" author="Levy, G." box="[995,1137,313,339]" pageId="13" pageNumber="52" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy (1999)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED9745174F74D95B1FE9CFB31FD0F" blockId="13.[151,1437,313,735]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95B1FE9CFE1EFEC3" bold="true" box="[199,488,349,375]" pageId="13" pageNumber="52">Justification of transfer.</emphasis>
The illustrations of male and female genitalia provided in
<bibRefCitation id="EFF0A4855174F74D91F1FE9CFAF8FEC3" author="Levy, G." box="[1159,1294,349,375]" pageId="13" pageNumber="52" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy (1999</bibRefCitation>
: figs 1014) are diagnostic and were used for comparison. Although we did not examine the
<typeStatus id="54DA67D65174F74D9167FE40FBBAFE2F" box="[1041,1100,385,411]" pageId="13" pageNumber="52">types</typeStatus>
of
<taxonomicName id="4C61A2F75174F74D9106FE43FB17FE2F" authorityName="Simon" authorityYear="1908" box="[1136,1249,385,411]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B91505665174F74D9106FE43FB77FE2F" box="[1136,1153,386,411]" italics="true" pageId="13" pageNumber="52">P</emphasis>
.
<emphasis id="B91505665174F74D91F9FE43FB17FE2F" box="[1167,1249,386,411]" italics="true" pageId="13" pageNumber="52">pugnax</emphasis>
</taxonomicName>
, the illustrations of the genitalia provided for this species are consistent with the diagnostic features of male and female
<taxonomicName id="4C61A2F75174F74D906CFE64FF1DFE57" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D906CFE64FA6BFE0B" box="[1306,1437,421,447]" italics="true" pageId="13" pageNumber="52">Trichothyse</emphasis>
species
</taxonomicName>
, in contrast to
<emphasis id="B91505665174F74D94ECFE08FD3FFE57" box="[410,713,457,483]" italics="true" pageId="13" pageNumber="52">
<taxonomicName id="4C61A2F75174F74D94ECFE08FDC6FE57" authorityName="Westring" authorityYear="1874" box="[410,560,457,483]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="genus" sensu="stricto">Poecilochroa</taxonomicName>
<taxonomicNameLabel id="A226B81D5174F74D974FFE0BFD3FFE57" box="[569,713,458,483]" pageId="13" pageNumber="52" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C61A2F75174F74D9672FE08FC5DFE57" authorityName="Wunderlich" authorityYear="2011" box="[772,939,457,483]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665174F74D9672FE08FC5DFE57" box="[772,939,457,483]" italics="true" pageId="13" pageNumber="52">Macarophaeus</emphasis>
</taxonomicName>
: distomedially originating broad conductor with prolateral bend, apically originating claw-like embolus with broad embolic base, medially contiguous sperm duct with single loop, robust RTA with apical bifurcation, epigyne with a median extension of anterior margin and vulva with short copulatory ducts, large, bean-shaped primary and small secondary spermathecae (cf.
<figureCitation id="135AC5F15174F74D9052FDF4FA6EFDFB" box="[1316,1432,565,591]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 920" captionStartId-0="3.[151,264,1372,1396]" captionStartId-1="4.[151,264,1607,1631]" captionTargetBox-0="[152,1436,179,1346]" captionTargetBox-1="[151,1436,179,1581]" captionTargetId-0="figure-17@3.[151,1436,179,1346]" captionTargetId-1="figure-17@4.[151,1436,179,1581]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." captionText-1="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="13" pageNumber="52">Figs 120</figureCitation>
,
<figureCitation id="135AC5F15174F74D95E1FD98FF16FDC0" box="[151,224,601,628]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="13" pageNumber="52">5457</figureCitation>
,
<figureCitation id="135AC5F15174F74D959AFD98FEBBFDC7" box="[236,333,601,627]" captionStart-0="FIGURES 109115" captionStart-1="FIGURES 116119" captionStartId-0="24.[151,257,1943,1967]" captionStartId-1="25.[151,264,1479,1503]" captionTargetBox-0="[190,1398,195,1913]" captionTargetBox-1="[161,1426,189,1443]" captionTargetId-0="figure-17@24.[179,1408,179,1917]" captionTargetId-1="figure-17@25.[151,1436,179,1453]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." captionText-1="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="13" pageNumber="52">109119</figureCitation>
and
<bibRefCitation id="EFF0A4855174F74D94F3FD98FDF4FDC7" author="Levy, G." box="[389,514,601,627]" pageId="13" pageNumber="52" pagination="427 - 452" refId="ref19371" refString="Levy, G. (1999) Spiders of six uncommon drassodine genera (Araneae: Gnaphosidae) from Israel. Israel Journal of Zoology, 45, 427-452." type="journal article" year="1999">Levy 1999</bibRefCitation>
: figs 1014). Moreover, in the original description, O.
<bibRefCitation id="EFF0A4855174F74D9118FD98FA6AFDC7" author="Pickard-Cambridge, O." box="[1134,1436,601,627]" pageId="13" pageNumber="52" pagination="370 - 419" refId="ref19799" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42, 370-419, pls. LI - LII. https://doi.org/10.1111/j.1096-3642.1874.tb02495.x" type="journal article" year="1874">Pickard-Cambridge (1874)</bibRefCitation>
clearly mentioned that this species has very large AMEs, also indicating its incorrect placement in
<emphasis id="B91505665174F74D9070FDBCFED2FD0F" italics="true" pageId="13" pageNumber="52">
<taxonomicName id="4C61A2F75174F74D9070FDBCFA6AFD23" authorityName="Westring" authorityYear="1874" box="[1286,1436,637,663]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="genus" sensu="stricto">Poecilochroa</taxonomicName>
<taxonomicNameLabel id="A226B81D5174F74D95E1FD63FED2FD0F" box="[151,292,674,699]" pageId="13" pageNumber="52" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
. Based on these observations, we propose the transfer of
<taxonomicName id="4C61A2F75174F74D96DEFD63FBEFFD0F" authorityName="Simon" authorityYear="1908" box="[936,1049,673,699]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B91505665174F74D96DEFD63FC4FFD0F" box="[936,953,674,699]" italics="true" pageId="13" pageNumber="52">P</emphasis>
.
<emphasis id="B91505665174F74D96B1FD63FBEFFD0F" box="[967,1049,674,699]" italics="true" pageId="13" pageNumber="52">pugnax</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75174F74D914BFD60FB36FD0F" authorityName="Tucker" authorityYear="1923" box="[1085,1216,673,699]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665174F74D914BFD60FB36FD0F" box="[1085,1216,673,699]" italics="true" pageId="13" pageNumber="52">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5174F74D95B1FD04FC42FD6B" box="[199,948,709,735]" pageId="13" pageNumber="52" type="distribution">
<paragraph id="8BDED9745174F74D95B1FD04FC42FD6B" blockId="13.[151,1437,313,735]" box="[199,948,709,735]" pageId="13" pageNumber="52">
<emphasis id="B91505665174F74D95B1FD04FE97FD6B" bold="true" box="[199,353,709,735]" pageId="13" pageNumber="52">Distribution.</emphasis>
Mediterranean North Africa and
<collectingCountry id="F37699E45174F74D97A0FD04FCC3FD6B" box="[726,821,709,735]" name="Ethiopia" pageId="13" pageNumber="52">Ethiopia</collectingCountry>
(
<figureCitation id="135AC5F15174F74D9633FD04FC51FD6B" box="[837,935,709,735]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="13" pageNumber="52">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,563 @@
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<mods:title id="387AC2D8C7A5CD1B1DC6AEF5C8D2056C">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
</mods:titleInfo>
<mods:name id="CFEDE0FECD61A69B1A6E0B5522616E74" type="personal">
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<mods:namePart id="476FFD6A8A021E96FEA6471BB2CF144E">Sankaran, Pradeep M.</mods:namePart>
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<mods:affiliation id="2056769BA3413FDB3AF401773220AFF9">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
<mods:nameIdentifier id="C62BAF4A5C7DCCA9E056A2B8E0550936" type="email">pradeepmspala@rediffmail.com</mods:nameIdentifier>
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<mods:namePart id="B818B52E8636DD9B568EB5E2D91418A3">Haddad, Charles R.</mods:namePart>
<mods:nameIdentifier id="C6E87CCBA9290B35299B75A05B4AFE58" type="ORCID">0000-0002-2317-7760</mods:nameIdentifier>
<mods:affiliation id="8964C637A865FFC3BFED39183D40BA03">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="75123A2A3467AA790A4CF20EE0B9EB9B" type="email">haddadcr@ufs.ac.za</mods:nameIdentifier>
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<mods:namePart id="083B81E8EE6B4EB88C091D1F317BB552">Tripathi, Rishikesh</mods:namePart>
<mods:nameIdentifier id="99DF6D2F22F699B55E3E7776AF2B4DAF" type="ORCID">0000-0002-9192-4609</mods:nameIdentifier>
<mods:affiliation id="F308107A00D9401D54716ADA1D55324F">Centre for Animal Taxonomy and Ecology, Department of Zoology, Christ College, Irinjalakuda, Thrissur, Kerala 680 125, India.</mods:affiliation>
<mods:nameIdentifier id="7222C5611CC3610954D103F1EF6EF35C" type="email">rishikesh.tripathi14@gmail.com</mods:nameIdentifier>
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<mods:titleInfo id="E9D961C0127DE4453256DAABA80259F5">
<mods:title id="6071FF35EC15642569C7858446FC646C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EA5A26BE0BB0AB91B42BE5E9E258BE66">
<mods:date id="C2E61B60CAA3972A29876E0829CCAE37">2025</mods:date>
<mods:detail id="A8933FFD26BEA30DCBACA5298C34D1B6" type="pubDate">
<mods:number id="380421A2D9D1C4FCB20E721883529BAA">2025-01-31</mods:number>
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<mods:detail id="FABAEC0A55B132D51B01644FE60650C0" type="volume">
<mods:number id="DAA5731BF3017FDDBC1FB2F1F8E2A7DC">5583</mods:number>
</mods:detail>
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<mods:number id="BA4A872803498BCF0E0BCB370F659CCD">1</mods:number>
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<mods:start id="2B3F51205FE23A3366DBB3DE46D25D93">39</mods:start>
<mods:end id="409D4DCCEF283D9D0219F2C1B6E81B71">70</mods:end>
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<mods:url id="26FB18963898503E5769197E8990E9FF">https://doi.org/10.11646/zootaxa.5583.1.2</mods:url>
</mods:location>
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<mods:identifier id="1D01D75797C345FA53E182EEB7F9E223" type="DOI">10.11646/zootaxa.5583.1.2</mods:identifier>
<mods:identifier id="62BE59A848DA594840193EE60A3DD4CA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C868625177F75195E1F9F2FB91FF4D" ID-DOI="http://doi.org/10.5281/zenodo.14797902" ID-Zenodo-Dep="14797902" LSID="urn:lsid:plazi:treatment:03C868625177F75195E1F9F2FB91FF4D" httpUri="http://treatment.plazi.org/id/03C868625177F75195E1F9F2FB91FF4D" lastPageId="17" lastPageNumber="56" pageId="14" pageNumber="53">
<subSubSection id="C37B8AFF5177F74E95E1F9F2FCFCF9FA" box="[151,778,1587,1614]" pageId="14" pageNumber="53" type="nomenclature">
<paragraph id="8BDED9745177F74E95E1F9F2FCFCF9FA" blockId="14.[151,778,1587,1651]" box="[151,778,1587,1614]" pageId="14" pageNumber="53">
<heading id="D0966E185177F74E95E1F9F2FCFCF9FA" bold="true" box="[151,778,1587,1614]" fontSize="11" level="1" pageId="14" pageNumber="53" reason="1">
<emphasis id="B91505665177F74E95E1F9F2FCFCF9FA" bold="true" box="[151,778,1587,1614]" pageId="14" pageNumber="53">
<taxonomicName id="4C61A2F75177F74E95E1F9F2FD8BF9F9" authority="(Lawrence, 1938)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[151,637,1587,1614]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="53" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B91505665177F74E95E1F9F2FE6BF9FA" bold="true" box="[151,413,1587,1614]" italics="true" pageId="14" pageNumber="53">Trichothyse zuluensis</emphasis>
(
<bibRefCitation id="EFF0A4855177F74E94DBF9F2FD82F9F9" author="Lawrence, R. F." box="[429,628,1587,1613]" pageId="14" pageNumber="53" pagination="455 - 524" refId="ref19341" refString="Lawrence, R. F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455-524." type="journal article" year="1938">Lawrence, 1938</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A226B81D5177F74E97F2F9F5FCFCF9FA" box="[644,778,1588,1614]" pageId="14" pageNumber="53" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5177F74E95E1F999FE56F9C7" box="[151,416,1624,1651]" pageId="14" pageNumber="53" type="description">
<paragraph id="8BDED9745177F74E95E1F999FE56F9C7" blockId="14.[151,778,1587,1651]" box="[151,416,1624,1651]" pageId="14" pageNumber="53">
<figureCitation id="135AC5F15177F74E95E1F999FF0FF9C6" box="[151,249,1624,1650]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="14" pageNumber="53">Figs 4, 8</figureCitation>
,
<figureCitation id="135AC5F15177F74E9470F999FEB8F9C6" box="[262,334,1624,1650]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="14" pageNumber="53">13, 19</figureCitation>
,
<figureCitation id="135AC5F15177F74E942CF999FE56F9C7" box="[346,416,1624,1651]" captionStart-0="FIGURES 6469" captionStart-1="FIGURES 7075" captionStartId-0="15.[151,264,1835,1859]" captionStartId-1="17.[151,264,1367,1391]" captionTargetBox-0="[151,1436,738,1808]" captionTargetBox-1="[151,1436,270,1341]" captionTargetId-0="figure-279@15.[151,1436,738,1809]" captionTargetId-1="figure-50@17.[151,1436,270,1341]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionText-0="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." captionText-1="FIGURES 7075. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., female from Ndumo: 70. Habitus, dorsal view; 71. Eye region, anterior view; 72. Same, dorsal view; 73. Chelicerae, posteroventral view; 74. Legs I &amp; II, retrolateral view; 75. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 70 = 1.0 mm; 7175 = 0.5 mm." pageId="14" pageNumber="53">6475</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5177F74E95E1F95DFEA2F946" pageId="14" pageNumber="53" type="reference_group">
<paragraph id="8BDED9745177F74E95E1F95DFEA2F946" blockId="14.[151,1437,1692,1778]" pageId="14" pageNumber="53">
<treatmentCitationGroup id="AB71FE5A5177F74E95E1F95DFEA2F946" pageId="14" pageNumber="53">
<treatmentCitation id="0AC0FF655177F74E95E1F95DFDA8F900" author="Lawrence, R. F." box="[151,606,1692,1716]" page="479" pageId="14" pageNumber="53" year="1938">
<taxonomicName id="4C61A2F75177F74E95E1F95DFDA8F900" ID-CoL="5CD6M" authority="Lawrence, 1938: 479" authorityName="Lawrence" authorityPageNumber="479" authorityYear="1938" box="[151,606,1692,1716]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="53" phylum="Arthropoda" rank="species" species="zuluensis">
<emphasis id="B91505665177F74E95E1F95DFE80F900" box="[151,374,1692,1716]" italics="true" pageId="14" pageNumber="53">Xerophaeus zuluensis</emphasis>
<bibRefCitation id="EFF0A4855177F74E9409F95CFDA8F900" author="Lawrence, R. F." box="[383,606,1692,1716]" pageId="14" pageNumber="53" pagination="455 - 524" refId="ref19341" refString="Lawrence, R. F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455-524." type="journal article" year="1938">Lawrence, 1938: 479</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 14;
<treatmentCitation id="0AC0FF655177F74E97B7F95DFBBFF900" author="Dippenaar-Schoeman, A. S. &amp; Haddad, C. R. &amp; Foord, S. H. &amp; Lotz, L. N." box="[705,1097,1692,1716]" page="63" pageId="14" pageNumber="53" year="2021">
<bibRefCitation id="EFF0A4855177F74E97B7F95DFBBFF900" author="Dippenaar-Schoeman, A. S. &amp; Haddad, C. R. &amp; Foord, S. H. &amp; Lotz, L. N." box="[705,1097,1692,1716]" pageId="14" pageNumber="53" refId="ref18797" refString="Dippenaar-Schoeman, A. S., Haddad, C. R., Foord, S. H. &amp; Lotz, L. N. (2021) The Gnaphosidae of South Africa. Part 3 (T - X). Version 1. South African National Survey of Arachnida Photo Identification Guide, Irene, 65 pp. https://doi.org/10.5281/zenodo.7197741" type="book" year="2021">
Dippenaar-Schoeman
<emphasis id="B91505665177F74E96D3F95CFC2AF900" box="[933,988,1692,1716]" italics="true" pageId="14" pageNumber="53">et al.</emphasis>
, 2021: 63
</bibRefCitation>
</treatmentCitation>
, figs 14 (
<typeStatus id="54DA67D65177F74E91CAF95DFAE0F900" box="[1212,1302,1692,1716]" pageId="14" pageNumber="53" type="holotype">holotype</typeStatus>
<specimenCount id="9D6712FD5177F74E9056F95CFAC3F900" box="[1312,1333,1693,1716]" count="1" pageId="14" pageNumber="53" type="female"></specimenCount>
:
<emphasis id="B91505665177F74E9032F95DFEC5F967" bold="true" pageId="14" pageNumber="53">
<collectingCountry id="F37699E45177F74E9032F95DFEDBF967" name="South Africa" pageId="14" pageNumber="53">SOUTH AFRICA</collectingCountry>
:
</emphasis>
<collectingRegion id="49A517965177F74E944BF97AFE16F967" box="[317,480,1723,1747]" country="South Africa" name="KwaZulu-Natal" pageId="14" pageNumber="53">KwaZulu-Natal</collectingRegion>
: Nkandhla [
<geoCoordinate id="EE55BFB35177F74E9717F97AFD41F967" box="[609,695,1723,1747]" degrees="28" direction="south" minutes="37" orientation="latitude" pageId="14" pageNumber="53" precision="925" value="-28.616667">28°37S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35177F74E97B1F97AFCD6F967" box="[711,800,1723,1747]" degrees="31" direction="east" minutes="05" orientation="longitude" pageId="14" pageNumber="53" precision="925" value="31.083334">31°05E</geoCoordinate>
],
<date id="FFDFFFB45177F74E9643F97AFC8EF967" box="[821,888,1723,1747]" pageId="14" pageNumber="53" value="1937-01">I.1937</date>
, leg. R.F. Lawrence, NMSA 1400—examined from photographs).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5177F74F95E1F8DEFB49FE65" lastPageId="15" lastPageNumber="54" pageId="14" pageNumber="53" type="materials_examined">
<paragraph id="8BDED9745177F74F95E1F8DEFB49FE65" blockId="14.[151,1436,1823,2031]" lastBlockId="15.[151,1437,150,681]" lastPageId="15" lastPageNumber="54" pageId="14" pageNumber="53">
<emphasis id="B91505665177F74E95E1F8DEFE3AF88D" bold="true" box="[151,460,1823,1849]" pageId="14" pageNumber="53">Other material examined.</emphasis>
<materialsCitation id="3B09D3295177F74F94A5F8DEFB4DFE65" collectingDate="2004-07-10" collectingDateMax="2020-10-17" collectingDateMin="2004-07-10" collectionCode="NCA" collectorName="C. Haddad &amp; Haddad, R &amp; J. Neethling &amp; R. Booysen &amp; R. Steenkamp &amp; Same &amp; Shokwe Pan" country="South Africa" county="Ndumo Game Reserve" elevation="35" lastPageId="15" lastPageNumber="54" latitude="-26.875" location="Pongola River" longLatPrecision="20" longitude="32.20667" municipality="Banzi Pan" pageId="14" pageNumber="53" specimenCode="NCA 2019, NCA 2014, NCA 2016, NCA 2020" specimenCount="35" specimenCount-female="27" specimenCount-male="8" stateProvince="KwaZulu-Natal">
<emphasis id="B91505665177F74E94A5F8DEFC80F88E" bold="true" box="[467,886,1823,1850]" pageId="14" pageNumber="53">
<collectingCountry id="F37699E45177F74E94A5F8DEFD5AF88D" box="[467,684,1823,1849]" name="South Africa" pageId="14" pageNumber="53">SOUTH AFRICA</collectingCountry>
:
<emphasis id="B91505665177F74E97CFF8DEFC9BF88E" bold="true" box="[697,877,1823,1850]" italics="true" pageId="14" pageNumber="53">
<collectingRegion id="49A517965177F74E97CFF8DEFC9BF88E" box="[697,877,1823,1850]" country="South Africa" name="KwaZulu-Natal" pageId="14" pageNumber="53">KwaZulu-Natal</collectingRegion>
</emphasis>
:
</emphasis>
<collectingCounty id="62BFA1F85177F74E9608F8E1FB76F88E" box="[894,1152,1824,1850]" pageId="14" pageNumber="53">Ndumo Game Reserve</collectingCounty>
,
<collectingMunicipality id="6BBA430E5177F74E91FAF8E1FAF7F88E" box="[1164,1281,1824,1850]" pageId="14" pageNumber="53">Banzi Pan</collectingMunicipality>
,
<geoCoordinate id="EE55BFB35177F74E907BF8E1FA6EF88F" box="[1293,1432,1824,1851]" degrees="26" direction="south" minutes="53" orientation="latitude" pageId="14" pageNumber="53" precision="15" seconds="03" value="-26.884167">26°5303S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35177F74E95E1F885FED2F8EA" box="[151,292,1860,1886]" degrees="32" direction="east" minutes="17" orientation="longitude" pageId="14" pageNumber="53" precision="15" seconds="13" value="32.286945">32°1713E</geoCoordinate>
,
<elevation id="004C3E475177F74E9444F885FE5AF8EA" box="[306,428,1860,1887]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="53" unit="m" value="35.0">
<quantity id="4C9974915177F74E9444F885FE9BF8EA" box="[306,365,1860,1887]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="53" unit="m" value="35.0">35 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45177F74E94CDF885FDCEF8EA" box="[443,568,1860,1886]" pageId="14" pageNumber="53" value="2018-12-07">
<collectingDate id="EF9B065C5177F74E94CDF885FDCEF8EA" box="[443,568,1860,1886]" pageId="14" pageNumber="53" value="2018-12-07">7.XII.2018</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25177F74E970EF885FD03F8EA" box="[632,757,1860,1886]" pageId="14" pageNumber="53">C. Haddad</collectorName>
, R. Booysen &amp;
<collectorName id="2694BCA25177F74E96C6F885FBCFF8EA" box="[944,1081,1860,1886]" pageId="14" pageNumber="53">J. Neethling</collectorName>
(canopy fogging,
<taxonomicName id="4C61A2F75177F74E9078F885FF00F836" class="Magnoliopsida" family="Euphorbiaceae" genus="Spirostachys" kingdom="Plantae" order="Malpighiales" pageId="14" pageNumber="53" phylum="Tracheophyta" rank="species" species="africana">
<emphasis id="B91505665177F74E9078F885FF00F836" italics="true" pageId="14" pageNumber="53">Spirostachys africana</emphasis>
</taxonomicName>
),
<specimenCount id="9D6712FD5177F74E9466F8A9FEC5F836" box="[272,307,1896,1922]" count="1" pageId="14" pageNumber="53" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F5177F74E9432F8A9FE32F837" box="[324,452,1896,1923]" collectionCode="NCA" pageId="14" pageNumber="53">NCA 2019</specimenCode>
/215); Same data but leg.
<collectorName id="2694BCA25177F74E979BF8A9FC91F836" box="[749,871,1896,1922]" pageId="14" pageNumber="53">C. Haddad</collectorName>
&amp;
<collectorName id="2694BCA25177F74E96F9F8A9FBE2F836" box="[911,1044,1896,1922]" pageId="14" pageNumber="53">R. Booysen</collectorName>
(under bark of
<taxonomicName id="4C61A2F75177F74E91BAF8A8FA7AF836" box="[1228,1420,1897,1922]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="53" phylum="Mollusca" rank="species" species="sycomorus">
<emphasis id="B91505665177F74E91BAF8A8FA7AF836" box="[1228,1420,1897,1922]" italics="true" pageId="14" pageNumber="53">Ficus sycomorus</emphasis>
</taxonomicName>
),
<specimenCount id="9D6712FD5177F74E95E1F84DFF4CF812" box="[151,186,1932,1958]" count="1" pageId="14" pageNumber="53" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5177F74E95B8F84DFEA5F813" box="[206,339,1932,1959]" collectionCode="NCA" pageId="14" pageNumber="53">NCA 2019</specimenCode>
/175); Same locality, SW shore of Banzi Pan,
<geoCoordinate id="EE55BFB35177F74E9609F84DFBFCF813" box="[895,1034,1932,1959]" degrees="26" direction="south" minutes="53" orientation="latitude" pageId="14" pageNumber="53" precision="15" seconds="07" value="-26.885277">26°5307S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35177F74E916DF84DFB51F813" box="[1051,1191,1932,1959]" degrees="32" direction="east" minutes="16" orientation="longitude" pageId="14" pageNumber="53" precision="15" seconds="55" value="32.281944">32°1655E</geoCoordinate>
,
<date id="FFDFFFB45177F74E91CFF84DFACDF813" box="[1209,1339,1932,1959]" pageId="14" pageNumber="53" value="2005-06-17">
<collectingDate id="EF9B065C5177F74E91CFF84DFACDF813" box="[1209,1339,1932,1959]" pageId="14" pageNumber="53" value="2005-06-17">17.VI.2005</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25177F74E90F5F84DFF18F87E" pageId="14" pageNumber="53">C. Haddad</collectorName>
(
<taxonomicName id="4C61A2F75177F74E9588F871FDF3F87E" box="[254,517,1968,1994]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="53" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B91505665177F74E9588F871FDF3F87E" box="[254,517,1968,1994]" italics="true" pageId="14" pageNumber="53">Vachellia xanthophloea</emphasis>
</taxonomicName>
bark),
<specimenCount id="9D6712FD5177F74E9723F871FD8EF87E" box="[597,632,1968,1994]" count="1" pageId="14" pageNumber="53" type="male">1♂</specimenCount>
<specimenCount id="9D6712FD5177F74E9709F871FD54F87E" box="[639,674,1968,1994]" count="1" pageId="14" pageNumber="53" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5177F74E97C6F871FCD8F87F" box="[688,814,1968,1995]" collectionCode="NCA" pageId="14" pageNumber="53">NCA 2014</specimenCode>
/1255); Same data but
<date id="FFDFFFB45177F74E9151F871FB62F87E" box="[1063,1172,1968,1994]" pageId="14" pageNumber="53" value="2006-01-23">
<collectingDate id="EF9B065C5177F74E9151F871FB62F87E" box="[1063,1172,1968,1994]" pageId="14" pageNumber="53" value="2006-01-23">23.I.2006</collectingDate>
</date>
,
<specimenCount id="9D6712FD5177F74E91D6F871FB35F87E" box="[1184,1219,1968,1994]" count="1" pageId="14" pageNumber="53" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5177F74E91A7F871FAB8F87E" box="[1233,1358,1968,1994]" collectionCode="NCA" pageId="14" pageNumber="53">NCA 2014</specimenCode>
/1274); Same locality, Crocodile Farm,
<location id="8EBE8FAF5177F74E9771F815FD5CF85A" LSID="urn:lsid:plazi:treatment:03C868625177F75195E1F9F2FB91FF4D:8EBE8FAF5177F74E9771F815FD5CF85A" box="[519,682,2004,2030]" country="South Africa" county="Ndumo Game Reserve" latitude="-26.875" longLatPrecision="20" longitude="32.20667" municipality="Banzi Pan" name="Pongola River" pageId="14" pageNumber="53" stateProvince="KwaZulu-Natal">
Pongola
<location id="8EBE8FAF5177F74E971AF815FD5CF85A" LSID="urn:lsid:plazi:treatment:03C868625177F75195E1F9F2FB91FF4D:8EBE8FAF5177F74E971AF815FD5CF85A" box="[620,682,2004,2030]" country="South Africa" county="Ndumo Game Reserve" latitude="-26.875" longLatPrecision="20" longitude="32.20667" municipality="Banzi Pan" name="River Floodplain" pageId="14" pageNumber="53" stateProvince="KwaZulu-Natal">River</location>
</location>
Floodplain,
<geoCoordinate id="EE55BFB35177F74E9649F815FC3CF85B" box="[831,970,2004,2031]" degrees="26" direction="south" minutes="54" orientation="latitude" pageId="14" pageNumber="53" precision="15" seconds="25" value="-26.906944">26°5425S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35177F74E96AEF815FB92F85A" box="[984,1124,2004,2030]" degrees="32" direction="east" minutes="19" orientation="longitude" pageId="14" pageNumber="53" precision="15" seconds="11" value="32.31972">32°1911E</geoCoordinate>
,
<date id="FFDFFFB45177F74E9104F815FB16F85A" box="[1138,1248,2004,2030]" pageId="14" pageNumber="53" value="2006-01-24">
<collectingDate id="EF9B065C5177F74E9104F815FB16F85A" box="[1138,1248,2004,2030]" pageId="14" pageNumber="53" value="2006-01-24">24.I.2006</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25177F74E9054F815FA6AF85A" box="[1314,1436,2004,2030]" pageId="14" pageNumber="53">C. Haddad</collectorName>
(
<taxonomicName id="4C61A2F75176F74F95D7FF56FEA1FF05" box="[161,343,151,177]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="54" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B91505665176F74F95D7FF56FEA1FF05" box="[161,343,151,177]" italics="true" pageId="15" pageNumber="54">V. xanthophloea</emphasis>
</taxonomicName>
bark),
<specimenCount id="9D6712FD5176F74F94DAFF56FE39FF05" box="[428,463,151,177]" count="2" pageId="15" pageNumber="54" type="female">2♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F9496FF56FD97FF05" box="[480,609,151,177]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1478); Same locality, S shore of Hotwe Pan,
<geoCoordinate id="EE55BFB35176F74F9104FF57FB0BFF05" box="[1138,1277,150,177]" degrees="26" direction="south" minutes="52" orientation="latitude" pageId="15" pageNumber="54" precision="15" seconds="43" value="-26.87861">26°5243S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35176F74F907DFF56FA61FF05" box="[1291,1431,151,177]" degrees="32" direction="east" minutes="18" orientation="longitude" pageId="15" pageNumber="54" precision="15" seconds="27" value="32.3075">32°1827E</geoCoordinate>
,
<date id="FFDFFFB45176F74F95E1FF7AFED4FF61" box="[151,290,187,213]" pageId="15" pageNumber="54" value="2004-07-10">
<collectingDate id="EF9B065C5176F74F95E1FF7AFED4FF61" box="[151,290,187,213]" pageId="15" pageNumber="54" value="2004-07-10">10.VII.2004</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25176F74F9429FF7AFE21FF61" box="[351,471,187,213]" pageId="15" pageNumber="54">C. Haddad</collectorName>
(
<taxonomicName id="4C61A2F75176F74F949EFF7AFD6AFF61" box="[488,668,187,213]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="54" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B91505665176F74F949EFF7AFD6AFF61" box="[488,668,187,213]" italics="true" pageId="15" pageNumber="54">V. xanthophloea</emphasis>
</taxonomicName>
bark),
<specimenCount id="9D6712FD5176F74F9798FF7AFCE6FF61" box="[750,784,187,213]" count="2" pageId="15" pageNumber="54" type="male">2♂</specimenCount>
(
<specimenCode id="DBC7710F5176F74F9669FF7AFC68FF61" box="[799,926,187,213]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1293); Same data but
<date id="FFDFFFB45176F74F91ECFF7BFAF5FF61" box="[1178,1283,186,213]" pageId="15" pageNumber="54" value="2005-02-07">
<collectingDate id="EF9B065C5176F74F91ECFF7BFAF5FF61" box="[1178,1283,186,213]" pageId="15" pageNumber="54" value="2005-02-07">7.II.2005</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F9079FF7AFAC2FF61" box="[1295,1332,187,213]" count="4" pageId="15" pageNumber="54" type="female">4♀</specimenCount>
, together with
<specimenCount id="9D6712FD5176F74F95B9FF1EFF04FF4D" box="[207,242,223,249]" count="1" pageId="15" pageNumber="54" type="male">1♂</specimenCount>
<taxonomicName id="4C61A2F75176F74F958EFF1EFE81FF4D" authorityName="Lawrence" authorityYear="1928" box="[248,375,223,249]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B91505665176F74F958EFF1EFE81FF4D" box="[248,375,223,249]" italics="true" pageId="15" pageNumber="54">T. fontensis</emphasis>
</taxonomicName>
(
<specimenCode id="DBC7710F5176F74F94F2FF1EFDF7FF4D" box="[388,513,223,249]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1317); Same data but
<date id="FFDFFFB45176F74F978EFF1FFC8DFF4D" box="[760,891,222,249]" pageId="15" pageNumber="54" value="2005-06-22">
<collectingDate id="EF9B065C5176F74F978EFF1FFC8DFF4D" box="[760,891,222,249]" pageId="15" pageNumber="54" value="2005-06-22">22.VI.2005</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F96F3FF1EFC5EFF4D" box="[901,936,223,249]" count="1" pageId="15" pageNumber="54" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F96C0FF1EFBC4FF4D" box="[950,1074,223,249]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1334); Same data but
<date id="FFDFFFB45176F74F905CFF1EFA6EFF4D" box="[1322,1432,223,249]" pageId="15" pageNumber="54" value="2006-01-27">
<collectingDate id="EF9B065C5176F74F905CFF1EFA6EFF4D" box="[1322,1432,223,249]" pageId="15" pageNumber="54" value="2006-01-27">27.I.2006</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F95E1FEC3FF4CFEA9" box="[151,186,258,285]" count="5" pageId="15" pageNumber="54" type="female">5♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F95BAFEC2FEBBFEA9" box="[204,333,259,285]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1306); Same locality, W shore of Nyamiti Pan,
<geoCoordinate id="EE55BFB35176F74F9609FEC3FBFCFEA9" box="[895,1034,258,285]" degrees="26" direction="south" minutes="53" orientation="latitude" pageId="15" pageNumber="54" precision="15" seconds="46" value="-26.89611">26°5346S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35176F74F916FFEC2FB53FEA9" box="[1049,1189,259,285]" degrees="32" direction="east" minutes="16" orientation="longitude" pageId="15" pageNumber="54" precision="15" seconds="33" value="32.275833">32°1633E</geoCoordinate>
,
<date id="FFDFFFB45176F74F91C3FEC2FAB6FEA9" box="[1205,1344,259,285]" pageId="15" pageNumber="54" value="2004-07-10">
<collectingDate id="EF9B065C5176F74F91C3FEC2FAB6FEA9" box="[1205,1344,259,285]" pageId="15" pageNumber="54" value="2004-07-10">10.VII.2004</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25176F74F90F5FEC2FF18FEF5" pageId="15" pageNumber="54">C. Haddad</collectorName>
(
<taxonomicName id="4C61A2F75176F74F9476FEE6FE42FEF5" box="[256,436,295,321]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="54" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B91505665176F74F9476FEE6FE42FEF5" box="[256,436,295,321]" italics="true" pageId="15" pageNumber="54">V. xanthophloea</emphasis>
</taxonomicName>
bark),
<specimenCount id="9D6712FD5176F74F9771FEE6FDDCFEF5" box="[519,554,295,321]" count="2" pageId="15" pageNumber="54" type="male">2♂</specimenCount>
<specimenCount id="9D6712FD5176F74F9744FEE6FDA3FEF5" box="[562,597,295,321]" count="2" pageId="15" pageNumber="54" type="female">2♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F9713FEE6FD12FEF5" box="[613,740,295,321]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1149); Same data but
<date id="FFDFFFB45176F74F9694FEE7FBBDFEF5" box="[994,1099,294,321]" pageId="15" pageNumber="54" value="2005-02-06">
<collectingDate id="EF9B065C5176F74F9694FEE7FBBDFEF5" box="[994,1099,294,321]" pageId="15" pageNumber="54" value="2005-02-06">6.II.2005</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F912EFEE6FB8DFEF5" box="[1112,1147,295,321]" count="1" pageId="15" pageNumber="54" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F91FCFEE6FAFFFEF5" box="[1162,1289,295,321]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1167); Same data but
<date id="FFDFFFB45176F74F9582FE8BFE83FED1" box="[244,373,330,357]" pageId="15" pageNumber="54" value="2005-06-19">
<collectingDate id="EF9B065C5176F74F9582FE8BFE83FED1" box="[244,373,330,357]" pageId="15" pageNumber="54" value="2005-06-19">19.VI.2005</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F9409FE8AFE54FED1" box="[383,418,331,357]" count="2" pageId="15" pageNumber="54" type="female">2♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F94D8FE8AFDDCFED1" box="[430,554,331,357]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1183); Same data but
<date id="FFDFFFB45176F74F966DFE8BFC6FFED1" box="[795,921,330,357]" pageId="15" pageNumber="54" value="2015-12-04">
<collectingDate id="EF9B065C5176F74F966DFE8BFC6FFED1" box="[795,921,330,357]" pageId="15" pageNumber="54" value="2015-12-04">4.XII.2015</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F96D4FE8BFC33FED1" box="[930,965,330,357]" count="5" pageId="15" pageNumber="54" type="female">5♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F96A4FE8AFBBAFED1" box="[978,1100,331,357]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2016</specimenCode>
/2117); Same locality, Nyamiti Pan,
<geoCoordinate id="EE55BFB35176F74F95A4FEAFFEABFE3D" box="[210,349,366,393]" degrees="26" direction="south" minutes="53" orientation="latitude" pageId="15" pageNumber="54" precision="15" seconds="21" value="-26.889166">26°5321S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35176F74F941AFEAEFE0EFE3D" box="[364,504,367,393]" degrees="32" direction="east" minutes="17" orientation="longitude" pageId="15" pageNumber="54" precision="15" seconds="42" value="32.295">32°1742E</geoCoordinate>
,
<elevation id="004C3E475176F74F9771FEAFFD75FE3D" box="[519,643,366,393]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="15" pageNumber="54" unit="m" value="53.0">
<quantity id="4C9974915176F74F9771FEAFFDB5FE3D" box="[519,579,366,393]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="15" pageNumber="54" unit="m" value="53.0">53 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB45176F74F97E4FEAEFCFCFE3D" box="[658,778,367,393]" pageId="15" pageNumber="54" value="2020-10-17">
<collectingDate id="EF9B065C5176F74F97E4FEAEFCFCFE3D" box="[658,778,367,393]" pageId="15" pageNumber="54" value="2020-10-17">17.X.2020</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25176F74F963BFEAEFC25FE3D" box="[845,979,367,393]" pageId="15" pageNumber="54">R. Booysen</collectorName>
&amp;
<collectorName id="2694BCA25176F74F968AFEAEFB6AFE3D" box="[1020,1180,367,393]" pageId="15" pageNumber="54">R. Steenkamp</collectorName>
(hand collecting),
<specimenCount id="9D6712FD5176F74F900FFEAEFA6AFE3D" box="[1401,1436,367,393]" count="1" pageId="15" pageNumber="54" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F95E8FE52FEEBFE19" box="[158,285,402,429]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2020</specimenCode>
/857);
<collectorName id="2694BCA25176F74F9410FE52FE52FE19" box="[358,420,403,429]" pageId="15" pageNumber="54">Same</collectorName>
locality, E shore of
<collectorName id="2694BCA25176F74F97FAFE52FCEDFE19" box="[652,795,403,429]" pageId="15" pageNumber="54">Shokwe Pan</collectorName>
,
<geoCoordinate id="EE55BFB35176F74F965EFE53FC44FE19" box="[808,946,402,429]" degrees="26" direction="south" minutes="52" orientation="latitude" pageId="15" pageNumber="54" precision="15" seconds="30" value="-26.875">26°5230S</geoCoordinate>
,
<geoCoordinate id="EE55BFB35176F74F96B6FE52FBBAFE19" box="[960,1100,403,429]" degrees="32" direction="east" minutes="12" orientation="longitude" pageId="15" pageNumber="54" precision="15" seconds="24" value="32.20667">32°1224E</geoCoordinate>
,
<date id="FFDFFFB45176F74F912FFE52FB13FE19" box="[1113,1253,403,429]" pageId="15" pageNumber="54" value="2004-07-13">
<collectingDate id="EF9B065C5176F74F912FFE52FB13FE19" box="[1113,1253,403,429]" pageId="15" pageNumber="54" value="2004-07-13">13.VII.2004</collectingDate>
</date>
, leg.
<collectorName id="2694BCA25176F74F9055FE52FA6AFE19" box="[1315,1436,403,429]" pageId="15" pageNumber="54">C. Haddad</collectorName>
(
<taxonomicName id="4C61A2F75176F74F95D7FE76FEA2FE65" box="[161,340,439,465]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="54" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B91505665176F74F95D7FE76FEA2FE65" box="[161,340,439,465]" italics="true" pageId="15" pageNumber="54">V. xanthophloea</emphasis>
</taxonomicName>
bark),
<specimenCount id="9D6712FD5176F74F94D2FE76FE31FE65" box="[420,455,439,465]" count="1" pageId="15" pageNumber="54" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F5176F74F94A0FE76FDA5FE65" box="[470,595,438,465]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1350);
<collectorName id="2694BCA25176F74F97DEFE76FD10FE65" box="[680,742,439,465]" pageId="15" pageNumber="54">Same</collectorName>
data but
<date id="FFDFFFB45176F74F9638FE77FC41FE65" box="[846,951,438,465]" pageId="15" pageNumber="54" value="2005-02-06">
<collectingDate id="EF9B065C5176F74F9638FE77FC41FE65" box="[846,951,438,465]" pageId="15" pageNumber="54" value="2005-02-06">6.II.2005</collectingDate>
</date>
,
<specimenCount id="9D6712FD5176F74F96B5FE76FC13FE65" box="[963,997,439,465]" count="1" pageId="15" pageNumber="54" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F5176F74F9682FE76FB87FE65" box="[1012,1137,439,465]" collectionCode="NCA" pageId="15" pageNumber="54">NCA 2014</specimenCode>
/1370)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5176F74F95B1FE1BFB28FD1D" pageId="15" pageNumber="54" type="diagnosis">
<paragraph id="8BDED9745176F74F95B1FE1BFB28FD1D" blockId="15.[151,1437,150,681]" pageId="15" pageNumber="54">
<emphasis id="B91505665176F74F95B1FE1BFEB4FE40" bold="true" box="[199,322,474,500]" pageId="15" pageNumber="54">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C61A2F75176F74F94DBFE1AFDD8FE41" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[429,558,475,501]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B91505665176F74F94DBFE1AFDD8FE41" box="[429,558,475,501]" italics="true" pageId="15" pageNumber="54">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B91505665176F74F9745FE1AFD44FE41" bold="true" box="[563,690,475,501]" pageId="15" pageNumber="54">
<taxonomicNameLabel id="A226B81D5176F74F9745FE1AFD44FE41" box="[563,690,475,501]" pageId="15" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
closely resemble those of
<taxonomicName id="4C61A2F75176F74F96A5FE1AFBD9FE41" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[979,1071,475,501]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B91505665176F74F96A5FE1AFC15FE40" box="[979,995,475,500]" italics="true" pageId="15" pageNumber="54">T</emphasis>
.
<emphasis id="B91505665176F74F9699FE1AFBD9FE41" box="[1007,1071,475,501]" italics="true" pageId="15" pageNumber="54">karoo</emphasis>
</taxonomicName>
<emphasis id="B91505665176F74F9143FE1AFB7BFE41" bold="true" box="[1077,1165,475,501]" pageId="15" pageNumber="54">
<taxonomicNameLabel id="A226B81D5176F74F9143FE1AFB7BFE41" box="[1077,1165,475,501]" pageId="15" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but can be distinguished by the different passage of the sperm duct, the sharper bend of the embolus (
<figureCitation id="135AC5F15176F74F968EFE3EFBA8FDAD" box="[1016,1118,511,537]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="15" pageNumber="54">Figs 34</figureCitation>
), the longer basal section of the RTA (approximately the same length as the dorsal branch), and the shorter, stouter dorsal branch of the RTA in retrolateral view (
<figureCitation id="135AC5F15176F74F9428FD86FE32FDD5" box="[350,452,583,609]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="15" pageNumber="54">Figs 78</figureCitation>
). Females of
<taxonomicName id="4C61A2F75176F74F972DFD86FD2BFDD5" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[603,733,583,609]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B91505665176F74F972DFD86FD2BFDD5" box="[603,733,583,609]" italics="true" pageId="15" pageNumber="54">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B91505665176F74F9792FD86FC92FDD5" bold="true" box="[740,868,583,609]" pageId="15" pageNumber="54">
<taxonomicNameLabel id="A226B81D5176F74F9792FD86FC92FDD5" box="[740,868,583,609]" pageId="15" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
have almost parallel mesal margins of the primary spermathecae, as do
<taxonomicName id="4C61A2F75176F74F94F7FDAAFE2BFD31" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[385,477,619,645]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B91505665176F74F94F7FDAAFE2BFD31" box="[385,477,619,645]" italics="true" pageId="15" pageNumber="54">T. karoo</emphasis>
</taxonomicName>
<emphasis id="B91505665176F74F9493FDAAFDC9FD31" bold="true" box="[485,575,619,645]" pageId="15" pageNumber="54">
<taxonomicNameLabel id="A226B81D5176F74F9493FDAAFDC9FD31" box="[485,575,619,645]" pageId="15" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but can be distinguished from it by the heart-shaped atrium being placed at posterior middle of the epigyne (cf.
<figureCitation id="135AC5F15176F74F975DFD4EFD5EFD1D" box="[555,680,655,681]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="15" pageNumber="54">Figs 13, 19</figureCitation>
and
<figureCitation id="135AC5F15176F74F97A8FD4EFCA8FD1D" box="[734,862,655,681]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="15" pageNumber="54">Figs 11, 17</figureCitation>
; also see
<bibRefCitation id="EFF0A4855176F74F96BEFD4EFB8CFD1D" author="Lawrence, R. F." box="[968,1146,655,681]" pageId="15" pageNumber="54" pagination="455 - 524" refId="ref19341" refString="Lawrence, R. F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455-524." type="journal article" year="1938">Lawrence 1938</bibRefCitation>
: fig. 14).
</paragraph>
</subSubSection>
<caption id="DF1E89FC5176F74F95E1F8EAFF30F81A" pageId="15" pageNumber="54" startId="15.[151,264,1835,1859]" targetBox="[151,1436,738,1808]" targetPageId="15" targetType="figure">
<paragraph id="8BDED9745176F74F95E1F8EAFF30F81A" blockId="15.[151,1436,1835,1966]" pageId="15" pageNumber="54">
<emphasis id="B91505665176F74F95E1F8EAFEA5F8F7" bold="true" box="[151,339,1835,1859]" pageId="15" pageNumber="54">FIGURES 6469.</emphasis>
Somatic morphology of
<taxonomicName id="4C61A2F75176F74F973DF8EAFC2CF8F7" authority="(Lawrence, 1938)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[587,986,1835,1859]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="54" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B91505665176F74F973DF8EAFCD5F8F7" box="[587,803,1835,1859]" italics="true" pageId="15" pageNumber="54">Trichothyse zuluensis</emphasis>
(
<bibRefCitation id="EFF0A4855176F74F9659F8EAFC25F8F7" author="Lawrence, R. F." box="[815,979,1835,1859]" pageId="15" pageNumber="54" pagination="455 - 524" refId="ref19341" refString="Lawrence, R. F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455-524." type="journal article" year="1938">Lawrence, 1938</bibRefCitation>
)
</taxonomicName>
<emphasis id="B91505665176F74F9696F8EAFBA4F8F7" bold="true" box="[992,1106,1835,1859]" pageId="15" pageNumber="54">
<taxonomicNameLabel id="A226B81D5176F74F9696F8EAFBA4F8F7" box="[992,1106,1835,1859]" pageId="15" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C37B8AFF5169F75195B1FF57FD95FF61" lastPageId="17" lastPageNumber="56" pageId="16" pageNumber="55" type="description">
<paragraph id="8BDED9745169F75095B1FF57FE31FC8D" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="55">
<emphasis id="B91505665169F75095B1FF57FEAFFF04" bold="true" box="[199,345,150,176]" pageId="16" pageNumber="55">Description.</emphasis>
<emphasis id="B91505665169F7509413FF56FE69FF05" box="[357,415,151,177]" italics="true" pageId="16" pageNumber="55">Male</emphasis>
(Ndumo, NCA 2021/1255,
<figureCitation id="135AC5F15169F750979DFF56FC87FF05" box="[747,881,151,177]" captionStart="FIGURES 6469" captionStartId="15.[151,264,1835,1859]" captionTargetBox="[151,1436,738,1808]" captionTargetId="figure-279@15.[151,1436,738,1809]" captionTargetPageId="15" captionText="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." pageId="16" pageNumber="55">Figs 6469</figureCitation>
). Carapace and chelicerae dark brown, labium, endites, sternum, scutum and legs orange-brown (
<figureCitation id="135AC5F15169F75097A7FF7AFC75FF61" box="[721,899,187,213]" captionStart="FIGURES 6469" captionStartId="15.[151,264,1835,1859]" captionTargetBox="[151,1436,738,1808]" captionTargetId="figure-279@15.[151,1436,738,1809]" captionTargetPageId="15" captionText="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." pageId="16" pageNumber="55">Figs 64, 6769</figureCitation>
); legs with faint black mottling, femora darker and tarsi paler than patellae, tibiae and metatarsi. Abdomen dark grey, spinnerets dark brown; carapace with faintly darker striae radiating from fovea, densely covered with white appressed hairs, with stiff erect black setae in eye region, along midline and on posterior slope. Fovea narrow, longitudinal, straight, dark brown. Anterior surface of cheliceral paturon with long black setae, promargin with low undulating keel ending in blunt distal tooth (
<figureCitation id="135AC5F15169F7509019FE8AFF40FE3D" captionStart="FIGURES 6469" captionStartId="15.[151,264,1835,1859]" captionTargetBox="[151,1436,738,1808]" captionTargetId="figure-279@15.[151,1436,738,1809]" captionTargetPageId="15" captionText="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." pageId="16" pageNumber="55">Fig. 67</figureCitation>
; arrow); retromargin without denticle. Abdomen oval, hirsute, with deep orange-brown oval dorsal scutum extending slightly more than ⅓ abdomen length (
<figureCitation id="135AC5F15169F75097B4FE52FCE0FE19" box="[706,790,403,429]" captionStart="FIGURES 6469" captionStartId="15.[151,264,1835,1859]" captionTargetBox="[151,1436,738,1808]" captionTargetId="figure-279@15.[151,1436,738,1809]" captionTargetPageId="15" captionText="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." pageId="16" pageNumber="55">Fig. 64</figureCitation>
); three pairs of distinct sigilla, first and second pairs at ⅓ abdomen length, on posterior margin of scutum, third pair slightly beyond midpoint (
<figureCitation id="135AC5F15169F7509112FE76FB4CFE65" box="[1124,1210,439,465]" captionStart="FIGURES 6469" captionStartId="15.[151,264,1835,1859]" captionTargetBox="[151,1436,738,1808]" captionTargetId="figure-279@15.[151,1436,738,1809]" captionTargetPageId="15" captionText="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." pageId="16" pageNumber="55">Fig. 64</figureCitation>
). Metatarsi III and all tarsi with well-developed complete scopulae (
<figureCitation id="135AC5F15169F75097B2FE1AFCB1FE41" box="[708,839,475,501]" captionStart="FIGURES 6469" captionStartId="15.[151,264,1835,1859]" captionTargetBox="[151,1436,738,1808]" captionTargetId="figure-279@15.[151,1436,738,1809]" captionTargetPageId="15" captionText="FIGURES 6469. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., male from Ndumo: 64. Habitus, dorsal view; 65. Eye region, anterior view; 66. Same, dorsal view; 67. Chelicerae, posteroventral view; 68. Leg I, retrolateral view; 69. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 64 = 1.0 mm; 6569 = 0.5 mm." pageId="16" pageNumber="55">Figs 6869</figureCitation>
), scopulae on metatarsi III and IV restricted distally. Spinnerets hirsute. Body length 8.20. Carapace 3.75 long, 2.55 wide. Abdomen 4.60 long, 2.53 wide. Eye sizes and interdistances: ALE 0.17, AME 0.21, PLE 0.17, PME 0.16; ALEPLE 0.13, AMEAME 0.17, AMEPME 0.17, PMEPLE 0.10, PMEPME 0.15. AME and ALE nearly contiguous. Clypeus height at AMEs 0.14, at ALEs 0.16. Chelicerae 1.32 long. Length of palp and legs: palp 3.37 [1.18, 0.52, 0.45, 1.22], I 8.45 [2.55, 1.39, 1.90, 1.73, 0.88], II 8.43 [2.32, 1.62, 1.87, 1.70, 0.92], III 8.06 [2.20, 1.30, 1.70, 1.88, 0.98], IV 9.84 [2.33, 1.52, 2.23, 2.76, 1.00]. Leg formula: 4123. Spination of palp: femur pl 2 do 2, patella and tibia spineless, tarsus plv 1 rlv 2; legs: femur I pl 1 do 3, II pl 2 do 3, III pl 2 do 3 rl 2, IV pl 1 do 3 rl 2; patella I, II and IV spineless, III pl 1 rl 1; tibia III plv 1 vt 1, III pl 4 rl 2 plv 2 rlv 1 vt 2, IV pl 3 rl 2 plv 2 rlv 1 vt 1; metatarsus III plv 1 rlv 1, III pl 4 rl 2 plv 1 rlv 1 vt 4, IV pl 4 rl 2 plv 2 rlv 2 vt 3.
</paragraph>
<paragraph id="8BDED9745169F75095B1FC82FF43FB81" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="55">
<emphasis id="B91505665169F75095B1FC82FF0AFCE9" box="[199,252,835,861]" italics="true" pageId="16" pageNumber="55">Palp</emphasis>
(
<figureCitation id="135AC5F15169F7509478FC82FE8CFCE9" box="[270,378,835,861]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="16" pageNumber="55">Figs 4, 8</figureCitation>
): segments brown, cymbium darker (
<figureCitation id="135AC5F15169F7509658FC82FC81FCE9" box="[814,887,835,861]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="16" pageNumber="55">Fig. 4</figureCitation>
). RTA with stout base, with straight finger-like ventral branch directed almost vertically and markedly shorter, sharply pointed triangular dorsal branch (
<figureCitation id="135AC5F15169F750905CFCA6FA78FC35" box="[1322,1422,871,897]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="16" pageNumber="55">Figs 4, 8</figureCitation>
). Cymbial flange indistinct, approximately half the length of cymbium, smooth along most of its length. Tegulum oval, sac-like (
<figureCitation id="135AC5F15169F750944CFC6EFE89FC7D" box="[314,383,943,969]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="16" pageNumber="55">Fig. 4</figureCitation>
). Conductor membranous, passing behind embolus, with proximal section parallel to embolus, bending at right angle, apex smoothly rounded and directed at 3-oclock (
<figureCitation id="135AC5F15169F75096B8FC12FBC5FC59" box="[974,1075,979,1005]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="16" pageNumber="55">Figs 4, 8</figureCitation>
). Embolus arising apically, base parallel-sided and initially directed at 10-oclock, with sharp bend to spike-like tip, directed at 1-oclock (
<figureCitation id="135AC5F15169F750903BFC36FF51FB81" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="16" pageNumber="55">Figs 4, 8</figureCitation>
).
</paragraph>
<paragraph id="8BDED9745169F75095B1FBFEFC42F999" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="55">
<emphasis id="B91505665169F75095B1FBFEFEEDFBED" box="[199,283,1087,1113]" italics="true" pageId="16" pageNumber="55">Female</emphasis>
(Ndumo, NCA 2020/857,
<figureCitation id="135AC5F15169F750973CFBFEFD3BFBED" box="[586,717,1086,1113]" captionStart="FIGURES 7075" captionStartId="17.[151,264,1367,1391]" captionTargetBox="[151,1436,270,1341]" captionTargetId="figure-50@17.[151,1436,270,1341]" captionTargetPageId="17" captionText="FIGURES 7075. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., female from Ndumo: 70. Habitus, dorsal view; 71. Eye region, anterior view; 72. Same, dorsal view; 73. Chelicerae, posteroventral view; 74. Legs I &amp; II, retrolateral view; 75. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 70 = 1.0 mm; 7175 = 0.5 mm." pageId="16" pageNumber="55">Figs 7075</figureCitation>
). Similar to male except the following: carapace and chelicerae brown, labium, endites, sternum and legs yellow-brown (
<figureCitation id="135AC5F15169F750966AFBA2FC3DFBC9" box="[796,971,1122,1149]" captionStart="FIGURES 7075" captionStartId="17.[151,264,1367,1391]" captionTargetBox="[151,1436,270,1341]" captionTargetId="figure-50@17.[151,1436,270,1341]" captionTargetPageId="17" captionText="FIGURES 7075. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., female from Ndumo: 70. Habitus, dorsal view; 71. Eye region, anterior view; 72. Same, dorsal view; 73. Chelicerae, posteroventral view; 74. Legs I &amp; II, retrolateral view; 75. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 70 = 1.0 mm; 7175 = 0.5 mm." pageId="16" pageNumber="55">Figs 70, 7375</figureCitation>
); legs covered in faint black mottling, all metatarsi and tarsi I and II slightly darker than other segments; palps similar but tarsi darker. Abdomen mottled grey, spinnerets yellow-brown. Cheliceral promargin with low keel ending in distal tooth, with small denticle beyond it (
<figureCitation id="135AC5F15169F75095E9FB0EFF03FB5D" box="[159,245,1231,1257]" captionStart="FIGURES 7075" captionStartId="17.[151,264,1367,1391]" captionTargetBox="[151,1436,270,1341]" captionTargetId="figure-50@17.[151,1436,270,1341]" captionTargetPageId="17" captionText="FIGURES 7075. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., female from Ndumo: 70. Habitus, dorsal view; 71. Eye region, anterior view; 72. Same, dorsal view; 73. Chelicerae, posteroventral view; 74. Legs I &amp; II, retrolateral view; 75. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 70 = 1.0 mm; 7175 = 0.5 mm." pageId="16" pageNumber="55">Fig. 73</figureCitation>
; arrow); retromargin without denticle. Abdomen oval, hirsute (
<figureCitation id="135AC5F15169F75096BEFB0EFBEBFB5D" box="[968,1053,1231,1257]" captionStart="FIGURES 7075" captionStartId="17.[151,264,1367,1391]" captionTargetBox="[151,1436,270,1341]" captionTargetId="figure-50@17.[151,1436,270,1341]" captionTargetPageId="17" captionText="FIGURES 7075. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., female from Ndumo: 70. Habitus, dorsal view; 71. Eye region, anterior view; 72. Same, dorsal view; 73. Chelicerae, posteroventral view; 74. Legs I &amp; II, retrolateral view; 75. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 70 = 1.0 mm; 7175 = 0.5 mm." pageId="16" pageNumber="55">Fig. 70</figureCitation>
), with three pairs of distinct oval sigilla, two pairs at ⅓ abdomen length and third pair just beyond midpoint (
<figureCitation id="135AC5F15169F7509680FB32FBBAFAB9" box="[1014,1100,1267,1293]" captionStart="FIGURES 7075" captionStartId="17.[151,264,1367,1391]" captionTargetBox="[151,1436,270,1341]" captionTargetId="figure-50@17.[151,1436,270,1341]" captionTargetPageId="17" captionText="FIGURES 7075. Somatic morphology of Trichothyse zuluensis (Lawrence, 1938) comb. nov., female from Ndumo: 70. Habitus, dorsal view; 71. Eye region, anterior view; 72. Same, dorsal view; 73. Chelicerae, posteroventral view; 74. Legs I &amp; II, retrolateral view; 75. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 70 = 1.0 mm; 7175 = 0.5 mm." pageId="16" pageNumber="55">Fig. 70</figureCitation>
). Body length 8.10. Carapace 2.84 long, 1.99 wide. Abdomen 5.20 long, 2.83 wide. Eye sizes and interdistances: ALE 0.16, AME 0.19, PLE 0.14, PME 0.12; ALEPLE 0.10, AMEAME 0.05, AMEPME 0.13, PMEPLE 0.10, PMEPME 0.14. Clypeus height at AMEs 0.10, at ALEs 0.09. Chelicerae 1.25 long. Length of palp and legs: palp 2.11 [0.65, 0.37, 0.36, 0.73], I 6.00 [1.88, 1.08, 1.23, 1.13, 0.68], II 5.97 [1.80, 1.05, 1.24, 1.17, 0.71], III 5.71 [1.67, 0.90, 1.12, 1.30, 0.72], IV 7.59 [2.15, 1.13, 1.59, 1.94, 0.78]. Leg formula: 4123. Spination of palp: femur pl 1 do 2, patella pl 1 do 1, tibia pl 2 rl 2 plv 1, tarsus pl 1 rl 1 plv 2 rlv 2 vt 2; legs: femur I pl 1 do 3, II pl 2 do 3, III pl 2 do 3 rl 2, IV pl 1 do 3 rl 1; patella I, II and IV spineless, III pl 1 rl 1; tibia III plv 1 vt 1, III pl 4 rl 2 vt 2, IV pl 2 rl 2 plv 2 rlv 1 vt 2; metatarsus I plv 1, II plv 1 rlv 1, III pl 2 rl 2 plv 1 rlv 1 vt 3, IV pl 5 rl 4 plv 2 rlv 2 vt 3.
</paragraph>
<paragraph id="8BDED9745169F75095B1F9F6FC9CF955" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="55">
<emphasis id="B91505665169F75095B1F9F6FEC7F9E5" box="[199,305,1591,1617]" italics="true" pageId="16" pageNumber="55">Genitalia</emphasis>
(
<figureCitation id="135AC5F15169F7509434F9F6FE3FF9E5" box="[322,457,1591,1617]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="16" pageNumber="55">Figs 13, 19</figureCitation>
): epigyne hirsute, with paired comma-shaped ridges posteriorly, forming small heart-shaped atrium (
<figureCitation id="135AC5F15169F7509430F99AFE6CF9C1" box="[326,410,1627,1653]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="16" pageNumber="55">Fig. 13</figureCitation>
; arrow). Copulatory openings indistinct, hidden in lateral edge of atrium. Copulatory ducts short, hidden behind primary spermathecae (
<figureCitation id="135AC5F15169F75097E6F9BEFD13F92D" box="[656,741,1663,1689]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="16" pageNumber="55">Fig. 19</figureCitation>
). Paired secondary spermathecae globose, partly obscured by primary spermathecae (
<figureCitation id="135AC5F15169F75094EAF962FE18F909" box="[412,494,1699,1725]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="16" pageNumber="55">Fig. 19</figureCitation>
). Primary spermathecae parallel-sided, close together, with angular anteromesal ends (
<figureCitation id="135AC5F15169F75095E9F906FF05F955" box="[159,243,1735,1761]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="16" pageNumber="55">Fig. 19</figureCitation>
). Fertilization ducts narrow, curved, diverging (
<figureCitation id="135AC5F15169F750967FF906FCAAF955" box="[777,860,1735,1761]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="16" pageNumber="55">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BDED9745169F75095B1F92BFF26F89D" blockId="16.[151,1437,150,2013]" pageId="16" pageNumber="55">
<emphasis id="B91505665169F75095B1F92BFEC9F8B0" bold="true" box="[199,319,1770,1796]" pageId="16" pageNumber="55">Variation.</emphasis>
Male (n = 7): body length 7.509.70, mean 8.33. Female (n = 27): body length 5.7010.20, mean 7.95.
</paragraph>
<paragraph id="8BDED9745169F75195B1F8F3FD95FF61" blockId="16.[151,1437,150,2013]" lastBlockId="17.[151,1436,151,249]" lastPageId="17" lastPageNumber="56" pageId="16" pageNumber="55">
<emphasis id="B91505665169F75095B1F8F3FE1DF8F8" bold="true" box="[199,491,1842,1868]" pageId="16" pageNumber="55">Justification of transfer.</emphasis>
The
<typeStatus id="54DA67D65169F750975CF8F2FDACF8F9" box="[554,602,1843,1869]" pageId="16" pageNumber="55">type</typeStatus>
and fresh material of
<taxonomicName id="4C61A2F75169F7509616F8F2FC1FF8F9" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[864,1001,1843,1869]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="55" phylum="Arthropoda" rank="species" species="zuluensis">
<emphasis id="B91505665169F7509616F8F2FC1FF8F9" box="[864,1001,1843,1869]" italics="true" pageId="16" pageNumber="55">X. zuluensis</emphasis>
</taxonomicName>
is consistent in somatic and genitalic morphology with the other species of
<taxonomicName id="4C61A2F75169F750974AF896FD49F8C5" authorityName="Tucker" authorityYear="1923" box="[572,703,1879,1905]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665169F750974AF896FD49F8C5" box="[572,703,1879,1905]" italics="true" pageId="16" pageNumber="55">Trichothyse</emphasis>
</taxonomicName>
(re)described in this paper, and shows considerable differences to
<taxonomicName id="4C61A2F75169F75095E1F8BAFE4AF821" authority="Purcell, 1907" authorityName="Purcell" authorityYear="1907" box="[151,444,1915,1941]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665169F75095E1F8BAFEEAF821" box="[151,284,1915,1941]" italics="true" pageId="16" pageNumber="55">Xerophaeus</emphasis>
Purcell, 1907
</taxonomicName>
, an entirely African genus with large gnaphosids generally&gt;
<quantity id="4C9974915169F750911EF8BAFB5EF821" box="[1128,1192,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="55" unit="mm" value="8.0">8 mm</quantity>
in length (see
<bibRefCitation id="EFF0A4855169F7509038F8BAFF39F80D" author="Tucker, R. W. E." pageId="16" pageNumber="55" pagination="251 - 437" refId="ref20373" refString="Tucker, R. W. E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251-437." type="journal article" year="1923">Tucker 1923</bibRefCitation>
and
<bibRefCitation id="EFF0A4855169F7509473F85EFE55F80D" author="Murphy, J." box="[261,419,1951,1977]" pageId="16" pageNumber="55" pagination="1 - 92" refId="ref19543" refString="Murphy, J. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambridgeshire, 1 &amp; 2, i - xii + 1-92 &amp; i - ii + 93 - 605." type="journal article" year="2007">Murphy 2007</bibRefCitation>
). Most
<taxonomicName id="4C61A2F75169F7509483F85EFD8CF80D" authorityName="Purcell" authorityYear="1907" box="[501,634,1951,1977]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665169F7509483F85EFD8CF80D" box="[501,634,1951,1977]" italics="true" pageId="16" pageNumber="55">Xerophaeus</emphasis>
</taxonomicName>
have shorter, stouter legs, are grey in colour, have a simple RTA, a long slender conductor and embolus, and an epigyne with a pronounced anterior hood or tongue-like scape (see
<bibRefCitation id="EFF0A4855169F7519038F802FF39FF05" author="Tucker, R. W. E." lastPageId="17" lastPageNumber="56" pageId="16" pageNumber="55" pagination="251 - 437" refId="ref20373" refString="Tucker, R. W. E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251-437." type="journal article" year="1923">Tucker 1923</bibRefCitation>
and
<bibRefCitation id="EFF0A4855168F7519473FF56FD8FFF05" author="Dippenaar-Schoeman, A. S. &amp; Haddad, C. R. &amp; Foord, S. H. &amp; Lotz, L. N." box="[261,633,151,177]" pageId="17" pageNumber="56" refId="ref18797" refString="Dippenaar-Schoeman, A. S., Haddad, C. R., Foord, S. H. &amp; Lotz, L. N. (2021) The Gnaphosidae of South Africa. Part 3 (T - X). Version 1. South African National Survey of Arachnida Photo Identification Guide, Irene, 65 pp. https://doi.org/10.5281/zenodo.7197741" type="book" year="2021">
Dippenaar-Schoeman
<emphasis id="B91505665168F7519488FF56FDC0FF05" box="[510,566,151,177]" italics="true" pageId="17" pageNumber="56">et al.</emphasis>
2021
</bibRefCitation>
), traits inconsistent with the species examined here. We thus propose its transfer from
<taxonomicName id="4C61A2F75168F7519447FF7AFE40FF61" authorityName="Purcell" authorityYear="1907" box="[305,438,187,213]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665168F7519447FF7AFE40FF61" box="[305,438,187,213]" italics="true" pageId="17" pageNumber="56">Xerophaeus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F75168F75194ACFF7AFDABFF61" authorityName="Tucker" authorityYear="1923" box="[474,605,187,213]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="B91505665168F75194ACFF7AFDABFF61" box="[474,605,187,213]" italics="true" pageId="17" pageNumber="56">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5168F75195B1FF1FFB91FF4D" box="[199,1127,222,249]" pageId="17" pageNumber="56" type="distribution">
<paragraph id="8BDED9745168F75195B1FF1FFB91FF4D" blockId="17.[151,1436,151,249]" box="[199,1127,222,249]" pageId="17" pageNumber="56">
<emphasis id="B91505665168F75195B1FF1FFE97FF4C" bold="true" box="[199,353,222,248]" pageId="17" pageNumber="56">Distribution.</emphasis>
Known only from northern
<collectingRegion id="49A517965168F75197EBFF1EFCA6FF4D" box="[669,848,223,249]" country="South Africa" name="KwaZulu-Natal" pageId="17" pageNumber="56">KwaZulu-Natal</collectingRegion>
,
<collectingCountry id="F37699E45168F751962DFF1EFC1FFF4D" box="[859,1001,223,249]" name="South Africa" pageId="17" pageNumber="56">South Africa</collectingCountry>
(
<figureCitation id="135AC5F15168F751968EFF1EFBADFF4D" box="[1016,1115,223,249]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="17" pageNumber="56">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,215 @@
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<mods:title id="EADF3857C86DDE42E43AAC232A660BC9">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<mods:affiliation id="8FE75BFCDB52A5A8170E3E76508A016C">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:identifier id="63C422D160849699466E19273CC58AC0" type="ZooBank">77CA9A8E-3717-406E-9B1E-04B1FEDD0D85</mods:identifier>
</mods:mods>
<treatment id="03C86862517BF74395E1F8BBFC9CF8D8" ID-DOI="http://doi.org/10.5281/zenodo.14797888" ID-Zenodo-Dep="14797888" LSID="urn:lsid:plazi:treatment:03C86862517BF74395E1F8BBFC9CF8D8" httpUri="http://treatment.plazi.org/id/03C86862517BF74395E1F8BBFC9CF8D8" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41">
<subSubSection id="C37B8AFF517BF74295E1F8BBFD35F821" box="[151,707,1914,1941]" pageId="2" pageNumber="41" type="nomenclature">
<paragraph id="8BDED974517BF74295E1F8BBFD35F821" blockId="2.[151,707,1914,1941]" box="[151,707,1914,1941]" pageId="2" pageNumber="41">
<heading id="D0966E18517BF74295E1F8BBFD35F821" bold="true" box="[151,707,1914,1941]" fontSize="11" level="1" pageId="2" pageNumber="41" reason="1">
<emphasis id="B9150566517BF74295E1F8BBFD35F821" bold="true" box="[151,707,1914,1941]" pageId="2" pageNumber="41">
<taxonomicName id="4C61A2F7517BF74295E1F8BBFDC3F820" authority="(Fage, 1929)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Fage" baseAuthorityYear="1929" box="[151,565,1914,1941]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="antineae" status="comb. nov.">
<emphasis id="B9150566517BF74295E1F8BBFE63F821" bold="true" box="[151,405,1914,1941]" italics="true" pageId="2" pageNumber="41">Trichothyse antineae</emphasis>
(
<bibRefCitation id="EFF0A485517BF74294D3F8BBFDDBF820" author="Fage, L." box="[421,557,1914,1940]" pageId="2" pageNumber="41" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage, 1929</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A226B81D517BF742974BF8BAFD35F821" box="[573,707,1915,1941]" pageId="2" pageNumber="41" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517BF74295E1F87EFD11F842" pageId="2" pageNumber="41" type="reference_group">
<paragraph id="8BDED974517BF74295E1F87EFD11F842" blockId="2.[151,1437,1983,2038]" pageId="2" pageNumber="41">
<treatmentCitationGroup id="AB71FE5A517BF74295E1F87EFD11F842" pageId="2" pageNumber="41">
<treatmentCitation id="0AC0FF65517BF74295E1F87EFDD1F863" author="Fage, L." box="[151,551,1983,2008]" page="248" pageId="2" pageNumber="41" year="1929">
<taxonomicName id="4C61A2F7517BF74295E1F87EFDD1F863" ID-CoL="4KTBF" authority="Fage, 1929: 248" authorityName="Fage" authorityPageNumber="248" authorityYear="1929" box="[151,551,1983,2008]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="antineae">
<emphasis id="B9150566517BF74295E1F87EFE8AF863" box="[151,380,1983,2007]" italics="true" pageId="2" pageNumber="41">Poecilochroa antineae</emphasis>
<bibRefCitation id="EFF0A485517BF74294F4F801FDD1F863" author="Fage, L." box="[386,551,1983,2008]" pageId="2" pageNumber="41" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage, 1929: 248</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1 (
<typeStatus id="54DA67D6517BF7429702F87EFD38F863" box="[628,718,1983,2007]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
<specimenCount id="9D6712FD517BF74297A2F801FD1CF863" box="[724,746,1984,2007]" count="1" pageId="2" pageNumber="41" type="male"></specimenCount>
:
<emphasis id="B9150566517BF7429782F87EFCB4F863" bold="true" box="[756,834,1983,2007]" pageId="2" pageNumber="41">MALI:</emphasis>
Asselar [Asler] [
<geoCoordinate id="EE55BFB3517BF742969AF87EFBBEF863" box="[1004,1096,1983,2007]" degrees="18" direction="north" minutes="52" orientation="latitude" pageId="2" pageNumber="41" precision="925" value="18.866667">18°52N</geoCoordinate>
,
<geoCoordinate id="EE55BFB3517BF7429124F87EFB5DF863" box="[1106,1195,1983,2007]" degrees="00" direction="east" minutes="15" orientation="longitude" pageId="2" pageNumber="41" precision="925" value="0.25">00°15E</geoCoordinate>
],
<date id="FFDFFFB4517BF74291CDF87EFACFF863" box="[1211,1337,1983,2007]" pageId="2" pageNumber="41" value="1927-12-21">21.XII.1927</date>
, collector unknown, #596, repository unknown—not examined.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF1E89FC517AF74395E1FA9DFB48FA08" pageId="3" pageNumber="42" startId="3.[151,264,1372,1396]" targetBox="[152,1436,179,1346]" targetPageId="3" targetType="figure">
<paragraph id="8BDED974517AF74395E1FA9DFB48FA08" blockId="3.[151,1436,1372,1468]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF74395E1FA9DFECCFAC0" bold="true" box="[151,314,1372,1396]" pageId="3" pageNumber="42">FIGURES 18.</emphasis>
Palpal morphology of male Afrotropical
<taxonomicName id="4C61A2F7517AF74397AAFA9DFC54FAC0" box="[732,930,1372,1396]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF74397AAFA9DFCA4FAC0" box="[732,850,1372,1396]" italics="true" pageId="3" pageNumber="42">Trichothyse</emphasis>
species
</taxonomicName>
in ventral (14) and retrolateral (58) views. 1, 5.
<taxonomicName id="4C61A2F7517AF74395E1FA40FE4FFA2C" authority="Lawrence, 1928" authorityName="Lawrence" authorityYear="1928" box="[151,441,1408,1432]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517AF74395E1FA40FEFDFA2C" box="[151,267,1408,1432]" italics="true" pageId="3" pageNumber="42">T. fontensis</emphasis>
<bibRefCitation id="EFF0A485517AF7439464FA40FE4FFA2C" author="Lawrence, R. F." box="[274,441,1408,1432]" pageId="3" pageNumber="42" pagination="217 - 312" refId="ref19301" refString="Lawrence, R. F. (1928) Contributions to a knowledge of the fauna of South-West Africa VII. Arachnida (Part 2). Annals of the South African Museum, 25, 217-312." type="journal article" year="1928">Lawrence, 1928</bibRefCitation>
</taxonomicName>
, holotype from Sesfontein; 2, 6.
<taxonomicName id="4C61A2F7517AF743967CFA40FC88FA2C" authorityName="Lawrence" authorityYear="1928" box="[778,894,1408,1432]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517AF743967CFA40FC88FA2C" box="[778,894,1408,1432]" italics="true" pageId="3" pageNumber="42">T. fontensis</emphasis>
</taxonomicName>
from Ndumo; 3, 7.
<taxonomicName id="4C61A2F7517AF743913AFA40FA80FA2C" authority="Haddad &amp; Sankaran" authorityName="Haddad &amp; Sankaran" authorityYear="2025" box="[1100,1398,1408,1432]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B9150566517AF743913AFA40FB56FA2C" box="[1100,1184,1408,1432]" italics="true" pageId="3" pageNumber="42">T. karoo</emphasis>
Haddad &amp; Sankaran
</taxonomicName>
<emphasis id="B9150566517AF7439008FA41FF35FA08" bold="true" pageId="3" pageNumber="42">
<taxonomicNameLabel id="A226B81D517AF7439008FA41FF35FA08" pageId="3" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Jagerskraal; 4, 8.
<taxonomicName id="4C61A2F7517AF74394C5FA64FD15FA08" authority="(Lawrence, 1938)" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[435,739,1444,1468]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B9150566517AF74394C5FA64FDDFFA08" box="[435,553,1444,1468]" italics="true" pageId="3" pageNumber="42">T. zuluensis</emphasis>
(
<bibRefCitation id="EFF0A485517AF7439740FA65FD2AFA08" author="Lawrence, R. F." box="[566,732,1444,1468]" pageId="3" pageNumber="42" pagination="455 - 524" refId="ref19341" refString="Lawrence, R. F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455-524." type="journal article" year="1938">Lawrence, 1938</bibRefCitation>
)
</taxonomicName>
<emphasis id="B9150566517AF743979CFA65FCA8FA08" bold="true" box="[746,862,1444,1468]" pageId="3" pageNumber="42">
<taxonomicNameLabel id="A226B81D517AF743979CFA65FCA8FA08" box="[746,862,1444,1468]" pageId="3" pageNumber="42" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
from Ndumo. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C37B8AFF517AF74395E1F9CFFDB8F8FD" pageId="3" pageNumber="42" type="description">
<paragraph id="8BDED974517AF74395E1F9CFFC99F99C" blockId="3.[151,1437,1550,1901]" box="[151,879,1550,1577]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF74395E1F9CFFEDFF99C" bold="true" box="[151,297,1550,1576]" pageId="3" pageNumber="42">Description.</emphasis>
<emphasis id="B9150566517AF7439446F9CFFE9CF99C" box="[304,362,1550,1576]" italics="true" pageId="3" pageNumber="42">Male</emphasis>
. For description of the male, see
<bibRefCitation id="EFF0A485517AF74397A9F9CEFC9CF99C" author="Fage, L." box="[735,874,1550,1577]" pageId="3" pageNumber="42" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage (1929)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDED974517AF74395B1F9F3FE69F9F8" blockId="3.[151,1437,1550,1901]" box="[199,415,1586,1613]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF74395B1F9F3FEEDF9F8" box="[199,283,1586,1612]" italics="true" pageId="3" pageNumber="42">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BDED974517AF74395B1F997FDB8F8FD" blockId="3.[151,1437,1550,1901]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF74395B1F997FE11F9C4" bold="true" box="[199,487,1622,1648]" pageId="3" pageNumber="42">Justification of transfer.</emphasis>
The illustrations of the
<typeStatus id="54DA67D6517AF743978EF997FCACF9C4" box="[760,858,1622,1648]" pageId="3" pageNumber="42" type="holotype">holotype</typeStatus>
palp provided in
<bibRefCitation id="EFF0A485517AF7439155F996FB50F9C4" author="Fage, L." box="[1059,1190,1622,1649]" pageId="3" pageNumber="42" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage (1929</bibRefCitation>
: fig. 1) are diagnostic and were used for comparison. Even though we did not examine the
<typeStatus id="54DA67D6517AF74396F3F9BBFC11F920" box="[901,999,1658,1684]" pageId="3" pageNumber="42" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C61A2F7517AF743917FF9BAFB71F920" authorityName="Fage" authorityYear="1929" box="[1033,1159,1659,1685]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="antineae">
<emphasis id="B9150566517AF743917FF9BAFBECF920" box="[1033,1050,1659,1684]" italics="true" pageId="3" pageNumber="42">P</emphasis>
.
<emphasis id="B9150566517AF7439151F9BAFB71F920" box="[1063,1159,1659,1684]" italics="true" pageId="3" pageNumber="42">antineae</emphasis>
</taxonomicName>
, the original illustrations of the palp have all of the typical features of male
<taxonomicName id="4C61A2F7517AF7439794F95FFC34F90C" box="[738,962,1694,1720]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF7439794F95FFC93F90C" box="[738,869,1694,1720]" italics="true" pageId="3" pageNumber="42">Trichothyse</emphasis>
species
</taxonomicName>
, in contrast to
<emphasis id="B9150566517AF7439118F95FFA6BF90C" box="[1134,1437,1694,1720]" italics="true" pageId="3" pageNumber="42">
<taxonomicName id="4C61A2F7517AF7439118F95FFAF2F90C" authorityName="Westring" authorityYear="1874" box="[1134,1284,1694,1720]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus" sensu="stricto">Poecilochroa</taxonomicName>
<taxonomicNameLabel id="A226B81D517AF743907BF95EFA6BF90C" box="[1293,1437,1695,1720]" pageId="3" pageNumber="42" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C61A2F7517AF74395BEF903FE99F968" authorityName="Wunderlich" authorityYear="2011" box="[200,367,1730,1756]" class="Arachnida" family="Gnaphosidae" genus="Macarophaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9150566517AF74395BEF903FE99F968" box="[200,367,1730,1756]" italics="true" pageId="3" pageNumber="42">Macarophaeus</emphasis>
</taxonomicName>
: distomedially originating broad conductor with prolateral fold, apically originating claw-like embolus with broad embolic base, medially contiguous sperm duct with single loop, and a robust RTA with apical bifurcation (cf.
<figureCitation id="135AC5F1517AF7439432F8CBFE50F890" box="[324,422,1802,1828]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="3" pageNumber="42">Figs 18</figureCitation>
,
<figureCitation id="135AC5F1517AF74394C7F8CBFE0FF891" box="[433,505,1802,1829]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="3" pageNumber="42">5455</figureCitation>
,
<figureCitation id="135AC5F1517AF7439775F8CBFD91F890" box="[515,615,1802,1828]" captionStart="FIGURES 109115" captionStartId="24.[151,257,1943,1967]" captionTargetBox="[190,1398,195,1913]" captionTargetId="figure-17@24.[179,1408,179,1917]" captionTargetPageId="24" captionText="FIGURES 109115. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 109112. Male left palp: 109. Ventral view; 110. Same, retrolateral view; 111. Same, dorsal view; 112. RTA enlarged, retrolateral view. 113115. Female genitalia: 113. Intact epigyne, ventral view; 114. Epigyne after clearing in 10% KOH, ventral view; 115. Vulva, dorsal view. Scale bars: 0.1 mm." pageId="3" pageNumber="42">109112</figureCitation>
,
<figureCitation id="135AC5F1517AF7439707F8CBFD27F890" box="[625,721,1802,1828]" captionStart="FIGURES 116119" captionStartId="25.[151,264,1479,1503]" captionTargetBox="[161,1426,189,1443]" captionTargetId="figure-17@25.[151,1436,179,1453]" captionTargetPageId="25" captionText="FIGURES 116119. Genitalic morphology of Trichothyse jodhpurensis (Gajbe, 1993) comb. nov.: 116117. Male left palp: 116. Ventral view; 117. Same, retrolateral view. 118119. Female genitalia: 118. Epigyne, ventral view; 119. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; CyF, cymbial flange; E, embolus; EB, embolic base; FD, fertilisation duct; ME, median extension of anterior epigynal margin; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="3" pageNumber="42">116117</figureCitation>
and
<bibRefCitation id="EFF0A485517AF7439673F8CAFC88F890" author="Fage, L." box="[773,894,1802,1829]" pageId="3" pageNumber="42" pagination="248 - 254" refId="ref18925" refString="Fage, L. (1929) Mission saharienne Augieras-Draper 1927 - 1928. Araignees nouvelles. Bulletin du Museum National d'Histoire Naturelle de Paris, 1, 248-254." type="journal article" year="1929">Fage 1929</bibRefCitation>
: fig. 1). Based on these observations, we propose transferring
<taxonomicName id="4C61A2F7517AF7439457F8EEFE56F8FC" authorityName="Fage" authorityYear="1929" box="[289,416,1839,1865]" class="Arachnida" family="Gnaphosidae" genus="Poecilochroa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="antineae">
<emphasis id="B9150566517AF7439457F8EEFEC4F8FC" box="[289,306,1839,1864]" italics="true" pageId="3" pageNumber="42">P</emphasis>
.
<emphasis id="B9150566517AF7439436F8EEFE56F8FC" box="[320,416,1839,1864]" italics="true" pageId="3" pageNumber="42">antineae</emphasis>
</taxonomicName>
to
<taxonomicName id="4C61A2F7517AF74394B2F8EFFDB1F8FC" authorityName="Tucker" authorityYear="1923" box="[452,583,1838,1864]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B9150566517AF74394B2F8EFFDB1F8FC" box="[452,583,1838,1864]" italics="true" pageId="3" pageNumber="42">Trichothyse</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517AF74395B1F893FC9CF8D8" box="[199,874,1874,1901]" pageId="3" pageNumber="42" type="distribution">
<paragraph id="8BDED974517AF74395B1F893FC9CF8D8" blockId="3.[151,1437,1550,1901]" box="[199,874,1874,1901]" pageId="3" pageNumber="42">
<emphasis id="B9150566517AF74395B1F893FE97F8D8" bold="true" box="[199,353,1874,1900]" pageId="3" pageNumber="42">Distribution.</emphasis>
Known only from the
<typeStatus id="54DA67D6517AF7439716F892FD66F8D9" box="[608,656,1875,1901]" pageId="3" pageNumber="42">type</typeStatus>
locality (
<figureCitation id="135AC5F1517AF743978DF893FCABF8D8" box="[763,861,1874,1900]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="3" pageNumber="42">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="98872B47120C0551E142EF86B593FCCC">A review of the ground spider genus Trichothyse Tucker, 1923 (Araneae, Gnaphosidae)</mods:title>
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<emphasis id="B9150566517DF74495E1F880FE6CF8EF" bold="true" box="[151,410,1857,1884]" italics="true" pageId="4" pageNumber="43">Trichothyse fontensis</emphasis>
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<figureCitation id="135AC5F1517DF74495E1F8A4FEC4F834" box="[151,306,1893,1920]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="4" pageNumber="43">Figs 12, 56</figureCitation>
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<figureCitation id="135AC5F1517DF7449448F8A7FE8EF834" box="[318,376,1893,1920]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="4" pageNumber="43">9, 15</figureCitation>
,
<figureCitation id="135AC5F1517DF74494F2F8A7FE3DF834" box="[388,459,1894,1920]" captionStart-0="FIGURES 2127" captionStart-1="FIGURES 2833" captionStartId-0="5.[151,264,1193,1217]" captionStartId-1="6.[151,264,1393,1417]" captionTargetBox-0="[151,1436,179,1167]" captionTargetBox-1="[151,1436,179,1367]" captionTargetId-0="figure-17@5.[151,1436,179,1167]" captionTargetId-1="figure-17@6.[151,1436,179,1368]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." captionText-1="FIGURES 2833. Somatic morphology of Trichothyse fontensis Lawrence, 1928, female from Ndumo: 28. Habitus, dorsal view; 29. Eye region, anterior view; 30. Same, dorsal view; 31. Chelicerae, posteroventral view; 32. Legs I &amp; II, retrolateral view; 33. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 28 = 1.0 mm; 2933 = 0.5 mm." pageId="4" pageNumber="43">2133</figureCitation>
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<emphasis id="B9150566517CF74595E1FB68FEA1FB75" bold="true" box="[151,343,1193,1217]" pageId="5" pageNumber="44">FIGURES 2127.</emphasis>
Somatic morphology of
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, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm.
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<emphasis id="B9150566517CF74595E1FAB6FE3DFA25" bold="true" box="[151,459,1399,1425]" pageId="5" pageNumber="44">Other material examined.</emphasis>
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</taxonomicName>
bark),
<specimenCount id="9D6712FD517CF74591EDFA5DFB48FA02" box="[1179,1214,1436,1462]" count="1" pageId="5" pageNumber="44" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F517CF74591B9FA5DFAB8FA02" box="[1231,1358,1436,1462]" collectionCode="NCA" pageId="5" pageNumber="44">NCA 2014</specimenCode>
/1417);
<location id="8EBE8FAF517CF74595E1FA01FF23FA6E" LSID="urn:lsid:plazi:treatment:03C86862517DF74795E1F880FAEEFAB9:8EBE8FAF517CF74595E1FA01FF23FA6E" box="[151,213,1472,1498]" country="South Africa" county="Ndumo Game Reserve" latitude="-26.87861" longLatPrecision="20" longitude="32.3075" municipality="Eastern" name="Same" pageId="5" pageNumber="44" stateProvince="KwaZulu-Natal">Same</location>
locality,
<location id="8EBE8FAF517CF745944DFA7EFDC0FA6E" LSID="urn:lsid:plazi:treatment:03C86862517DF74795E1F880FAEEFAB9:8EBE8FAF517CF745944DFA7EFDC0FA6E" box="[315,566,1471,1498]" country="South Africa" county="Ndumo Game Reserve" latitude="-26.87861" longLatPrecision="20" longitude="32.3075" municipality="Eastern" name="Environmental Centre" pageId="5" pageNumber="44" stateProvince="KwaZulu-Natal">Environmental Centre</location>
,
<geoCoordinate id="EE55BFB3517CF7459736FA7EFD3DFA6E" box="[576,715,1471,1498]" degrees="26" direction="south" minutes="55" orientation="latitude" pageId="5" pageNumber="44" precision="15" seconds="24" value="-26.923332">26°5524S</geoCoordinate>
,
<geoCoordinate id="EE55BFB3517CF74597A0FA01FC94FA6E" box="[726,866,1472,1498]" degrees="32" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="44" precision="15" seconds="08" value="32.302223">32°1808E</geoCoordinate>
,
<elevation id="004C3E47517CF7459618FA01FC06FA6D" box="[878,1008,1471,1498]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="5" pageNumber="44" unit="m" value="122.0">
<quantity id="4C997491517CF7459618FA01FC42FA6E" box="[878,948,1472,1498]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="5" pageNumber="44" unit="m" value="122.0">122 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB4517CF745968AFA01FB88FA6E" box="[1020,1150,1472,1498]" pageId="5" pageNumber="44" value="2009-11-29">
<collectingDate id="EF9B065C517CF745968AFA01FB88FA6E" box="[1020,1150,1472,1498]" pageId="5" pageNumber="44" value="2009-11-29">29.XI.2009</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2517CF74591CFFA01FAC5FA6D" box="[1209,1331,1471,1498]" pageId="5" pageNumber="44">C. Haddad</collectorName>
, V. Swart &amp;
<collectorName id="2694BCA2517CF74595C0FA25FE99FA49" box="[182,367,1507,1534]" pageId="5" pageNumber="44">A. Kirk-Spriggs</collectorName>
(broadleaf woodland, canopy fogging 23,
<emphasis id="B9150566517CF7459612FA22FBC5FA49" box="[868,1075,1507,1533]" italics="true" pageId="5" pageNumber="44">Strynchos spinosa</emphasis>
),
<specimenCount id="9D6712FD517CF7459138FA25FB87FA4A" box="[1102,1137,1508,1534]" count="1" pageId="5" pageNumber="44" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F517CF74591F2FA25FAF2FA4A" box="[1156,1284,1507,1534]" collectionCode="NCA" pageId="5" pageNumber="44">NCA 2021</specimenCode>
/1255); Same locality, Near Abattoir,
<geoCoordinate id="EE55BFB3517CF74594D2F9C6FDD8F996" box="[420,558,1543,1570]" degrees="26" direction="south" minutes="54" orientation="latitude" pageId="5" pageNumber="44" precision="15" seconds="44" value="-26.912222">26°5444S</geoCoordinate>
,
<geoCoordinate id="EE55BFB3517CF745974AF9C9FD3FF996" box="[572,713,1544,1570]" degrees="32" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="44" precision="15" seconds="20" value="32.305553">32°1820E</geoCoordinate>
,
<elevation id="004C3E47517CF74597A1F9C9FCABF995" box="[727,861,1543,1570]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="5" pageNumber="44" unit="m" value="114.0">
<quantity id="4C997491517CF74597A1F9C9FCE9F996" box="[727,799,1544,1570]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="5" pageNumber="44" unit="m" value="114.0">114 m</quantity>
a.s.l.
</elevation>
,
<date id="FFDFFFB4517CF745961AF9C9FC1CF996" box="[876,1002,1544,1570]" pageId="5" pageNumber="44" value="2009-12-09">
<collectingDate id="EF9B065C517CF745961AF9C9FC1CF996" box="[876,1002,1544,1570]" pageId="5" pageNumber="44" value="2009-12-09">9.XII.2009</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2517CF745915FF9C9FB50F995" box="[1065,1190,1543,1570]" pageId="5" pageNumber="44">C. Haddad</collectorName>
, V. Swart &amp;
<collectorName id="2694BCA2517CF745904DF9C9FF1BF9F1" pageId="5" pageNumber="44">A. Kirk-Spriggs</collectorName>
(broadleaf woodland, canopy fogging 47,
<taxonomicName id="4C61A2F7517CF7459796F9EDFC27F9F1" box="[736,977,1579,1606]" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="44" phylum="Tracheophyta" rank="species" species="livingstonei">
<emphasis id="B9150566517CF7459796F9EDFC27F9F1" box="[736,977,1579,1606]" italics="true" pageId="5" pageNumber="44">Garcinia livingstonei</emphasis>
</taxonomicName>
),
<specimenCount id="9D6712FD517CF745969AF9EDFBF9F9F2" box="[1004,1039,1580,1606]" count="1" pageId="5" pageNumber="44" type="male">1♂</specimenCount>
(
<specimenCode id="DBC7710F517CF7459157F9EDFB57F9F2" box="[1057,1185,1579,1606]" collectionCode="NCA" pageId="5" pageNumber="44">NCA 2021</specimenCode>
/1256);
<collectorName id="2694BCA2517CF745918CF9EDFACEF9F2" box="[1274,1336,1580,1606]" pageId="5" pageNumber="44">Same</collectorName>
locality,
<collectorName id="2694BCA2517CF74595E1F98EFF0AF9DD" box="[151,252,1615,1641]" pageId="5" pageNumber="44">Southern</collectorName>
shore of
<collectorName id="2694BCA2517CF7459413F991FE12F9DE" box="[357,484,1616,1642]" pageId="5" pageNumber="44">Hotwe Pan</collectorName>
,
<geoCoordinate id="EE55BFB3517CF7459499F98EFD8DF9DE" box="[495,635,1615,1642]" degrees="26" direction="south" minutes="52" orientation="latitude" pageId="5" pageNumber="44" precision="15" seconds="43" value="-26.87861">26°5243S</geoCoordinate>
,
<geoCoordinate id="EE55BFB3517CF74597F0F991FCE4F9DE" box="[646,786,1616,1642]" degrees="32" direction="east" minutes="18" orientation="longitude" pageId="5" pageNumber="44" precision="15" seconds="27" value="32.3075">32°1827E</geoCoordinate>
,
<date id="FFDFFFB4517CF7459668F98EFC71F9DE" box="[798,903,1615,1642]" pageId="5" pageNumber="44" value="2005-02-07">
<collectingDate id="EF9B065C517CF7459668F98EFC71F9DE" box="[798,903,1615,1642]" pageId="5" pageNumber="44" value="2005-02-07">7.II.2005</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2517CF74596B5F991FBCDF9DD" box="[963,1083,1615,1642]" pageId="5" pageNumber="44">C. Haddad</collectorName>
(
<taxonomicName id="4C61A2F7517CF745913DF98EFAA5F9DD" box="[1099,1363,1615,1641]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="44" phylum="Tracheophyta" rank="species" species="xanthophloea">
<emphasis id="B9150566517CF745913DF98EFAA5F9DD" box="[1099,1363,1615,1641]" italics="true" pageId="5" pageNumber="44">Vachellia xanthophloea</emphasis>
</taxonomicName>
bark),
<specimenCount id="9D6712FD517CF74595E1F9B5FF4AF93A" box="[151,188,1652,1678]" count="1" pageId="5" pageNumber="44" type="male">1♂</specimenCount>
, together with
<specimenCount id="9D6712FD517CF7459412F9B5FE71F93A" box="[356,391,1652,1678]" count="4" pageId="5" pageNumber="44" type="female">4♀</specimenCount>
<taxonomicName id="4C61A2F7517CF74594F8F9B5FDE6F939" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[398,528,1651,1677]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B9150566517CF74594F8F9B5FDE6F939" box="[398,528,1651,1677]" italics="true" pageId="5" pageNumber="44">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF7459761F9B5FD61F93A" bold="true" box="[535,663,1652,1678]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF7459761F9B5FD61F93A" box="[535,663,1652,1678]" pageId="5" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<specimenCode id="DBC7710F517CF74597D0F9B5FCD2F93A" box="[678,804,1652,1678]" collectionCode="NCA" pageId="5" pageNumber="44">NCA 2014</specimenCode>
/1317)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517CF74595B1F956FE49F842" pageId="5" pageNumber="44" type="diagnosis">
<paragraph id="8BDED974517CF74595B1F956FE49F842" blockId="5.[151,1437,1399,2038]" pageId="5" pageNumber="44">
<emphasis id="B9150566517CF74595B1F956FEB4F905" bold="true" box="[199,322,1687,1713]" pageId="5" pageNumber="44">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C61A2F7517CF74594C5F959FDC3F905" authorityName="Lawrence" authorityYear="1928" box="[435,565,1687,1714]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517CF74594C5F959FE35F905" box="[435,451,1688,1713]" italics="true" pageId="5" pageNumber="44">T</emphasis>
.
<emphasis id="B9150566517CF74594A7F956FDC3F905" box="[465,565,1687,1713]" italics="true" pageId="5" pageNumber="44">fontensis</emphasis>
</taxonomicName>
are similar to the males of
<taxonomicName id="4C61A2F7517CF745961FF959FC1AF905" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[873,1004,1687,1713]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B9150566517CF745961FF959FC1AF905" box="[873,1004,1687,1713]" italics="true" pageId="5" pageNumber="44">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF7459685F959FB82F906" bold="true" box="[1011,1140,1688,1714]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF7459685F959FB82F906" box="[1011,1140,1688,1714]" pageId="5" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
in the general structure of the palp, particularly the rounded end of the conductor and the similar shape of the sperm duct (
<figureCitation id="135AC5F1517CF7459192F97AFA91F962" box="[1252,1383,1723,1750]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="5" pageNumber="44">Figs 12, 4</figureCitation>
) but can be distinguished by the different shape of the embolus, with the tip directed apically in
<taxonomicName id="4C61A2F7517CF74591E5F921FAE5F94D" authorityName="Lawrence" authorityYear="1928" box="[1171,1299,1759,1785]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517CF74591E5F921FAE5F94D" box="[1171,1299,1759,1785]" italics="true" pageId="5" pageNumber="44">T. fontensis</emphasis>
</taxonomicName>
(
<figureCitation id="135AC5F1517CF7459057F91EFA93F94E" box="[1313,1381,1759,1786]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="5" pageNumber="44">Fig. 2</figureCitation>
) and retrolaterally in
<taxonomicName id="4C61A2F7517CF745943CF8C5FE3AF8A9" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[330,460,1795,1821]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B9150566517CF745943CF8C5FE3AF8A9" box="[330,460,1795,1821]" italics="true" pageId="5" pageNumber="44">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF74594A4F8C5FDA7F8AA" bold="true" box="[466,593,1796,1822]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF74594A4F8C5FDA7F8AA" box="[466,593,1796,1822]" pageId="5" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135AC5F1517CF7459716F8C2FD55F8AA" box="[608,675,1795,1822]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="5" pageNumber="44">Fig. 4</figureCitation>
), and the lack of a strong bifurcation of the RTA in
<taxonomicName id="4C61A2F7517CF74591ABF8C5FAADF8A9" authorityName="Lawrence" authorityYear="1928" box="[1245,1371,1795,1821]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517CF74591ABF8C5FAADF8A9" box="[1245,1371,1795,1821]" italics="true" pageId="5" pageNumber="44">T. fontensis</emphasis>
</taxonomicName>
(only a slight protuberance;
<figureCitation id="135AC5F1517CF74594E4F8E6FE20F8F5" box="[402,470,1831,1857]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="5" pageNumber="44">Fig. 6</figureCitation>
). Females of
<taxonomicName id="4C61A2F7517CF7459706F8E9FD19F8F5" authorityName="Lawrence" authorityYear="1928" box="[624,751,1831,1857]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517CF7459706F8E9FD19F8F5" box="[624,751,1831,1857]" italics="true" pageId="5" pageNumber="44">T. fontensis</emphasis>
</taxonomicName>
share with
<taxonomicName id="4C61A2F7517CF7459604F8E9FC38F8F5" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[882,974,1831,1857]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B9150566517CF7459604F8E9FC38F8F5" box="[882,974,1831,1857]" italics="true" pageId="5" pageNumber="44">T. karoo</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF74596A0F8E9FBC6F8F6" bold="true" box="[982,1072,1832,1858]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF74596A0F8E9FBC6F8F6" box="[982,1072,1832,1858]" pageId="5" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C61A2F7517CF745911EF8E9FB1DF8F5" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[1128,1259,1831,1857]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B9150566517CF745911EF8E9FB1DF8F5" box="[1128,1259,1831,1857]" italics="true" pageId="5" pageNumber="44">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF7459184F8E9FA85F8F6" bold="true" box="[1266,1395,1832,1858]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF7459184F8E9FA85F8F6" box="[1266,1395,1832,1858]" pageId="5" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
the almost parallel mesal margins of the primary spermathecae, but can be distinguished by the angular anterolateral edge of the primary spermathecae (as opposed to the anteromesal edge of the latter two species), and the small hood-shaped anterior atrium compared to the small central teardrop-shaped atrium (as in
<taxonomicName id="4C61A2F7517CF74591FCF855FB10F819" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" box="[1162,1254,1939,1965]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="B9150566517CF74591FCF855FB10F819" box="[1162,1254,1939,1965]" italics="true" pageId="5" pageNumber="44">T. karoo</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF7459199F855FABFF81A" bold="true" box="[1263,1353,1940,1966]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF7459199F855FABFF81A" box="[1263,1353,1940,1966]" pageId="5" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) or the heart-shaped posterior atrium in the epigyne as in
<taxonomicName id="4C61A2F7517CF74597BFF879FCBDF865" authorityName="Sankaran &amp; Haddad &amp; Tripathi" authorityYear="2025" baseAuthorityName="Lawrence" baseAuthorityYear="1938" box="[713,843,1975,2001]" class="Arachnida" family="Gnaphosidae" genus="Xerophaeus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="zuluensis" status="comb. nov.">
<emphasis id="B9150566517CF74597BFF879FCBDF865" box="[713,843,1975,2001]" italics="true" pageId="5" pageNumber="44">T. zuluensis</emphasis>
</taxonomicName>
<emphasis id="B9150566517CF7459624F879FC24F866" bold="true" box="[850,978,1976,2002]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="A226B81D517CF7459624F879FC24F866" box="[850,978,1976,2002]" pageId="5" pageNumber="44" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(see
<bibRefCitation id="EFF0A485517CF745917BF879FB49F866" author="Lawrence, R. F." box="[1037,1215,1976,2002]" pageId="5" pageNumber="44" pagination="455 - 524" refId="ref19341" refString="Lawrence, R. F. (1938) A collection of spiders from Natal and Zululand. Annals of the Natal Museum, 8, 455-524." type="journal article" year="1938">Lawrence 1938</bibRefCitation>
: fig. 14) (cf.
<figureCitation id="135AC5F1517CF7459026F876FEAAF842" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="5" pageNumber="44">Figs 9, 11, 13, 15, 17, 19</figureCitation>
,
<figureCitation id="135AC5F1517CF745941EF81AFE45F842" box="[360,435,2011,2038]" captionStart="FIGURES 5457" captionStartId="12.[151,264,1679,1703]" captionTargetBox="[162,1425,190,1642]" captionTargetId="figure-17@12.[151,1435,179,1653]" captionTargetPageId="12" captionText="FIGURES 5457. Genitalic morphology of Trichothyse karoo Haddad &amp; Sankaran sp. nov.: 5455. Holotype male left palp: 54. Ventral view; 55. Retrolateral view. 5657. Paratype female: 56. Epigyne, ventral view; 57. Vulva, dorsal view. Abbreviations: A, atrium; C, conductor; CD, copulatory duct; E, embolus; EB, embolic base; FD, fertilisation duct; PL, posterior lobe; PS, primary spermatheca; RTA, retrolateral tibial apophysis; SD, sperm duct; SS, secondary spermatheca; T, tegulum. Scale bars: 0.1 mm." pageId="5" pageNumber="44">5657</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF1E89FC517FF74695E1FAB0FF30FA41" pageId="6" pageNumber="45" startId="6.[151,264,1393,1417]" targetBox="[151,1436,179,1367]" targetPageId="6" targetType="figure">
<paragraph id="8BDED974517FF74695E1FAB0FF30FA41" blockId="6.[151,1436,1393,1525]" pageId="6" pageNumber="45">
<emphasis id="B9150566517FF74695E1FAB0FEA1FA3D" bold="true" box="[151,343,1393,1417]" pageId="6" pageNumber="45">FIGURES 2833.</emphasis>
Somatic morphology of
<taxonomicName id="4C61A2F7517FF7469728FAB0FC11FA3E" authority="Lawrence, 1928" authorityName="Lawrence" authorityYear="1928" box="[606,999,1393,1418]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="fontensis">
<emphasis id="B9150566517FF7469728FAB0FCC1FA3E" box="[606,823,1393,1418]" italics="true" pageId="6" pageNumber="45">Trichothyse fontensis</emphasis>
<bibRefCitation id="EFF0A485517FF7469649FAB3FC11FA3E" author="Lawrence, R. F." box="[831,999,1394,1418]" pageId="6" pageNumber="45" pagination="217 - 312" refId="ref19301" refString="Lawrence, R. F. (1928) Contributions to a knowledge of the fauna of South-West Africa VII. Arachnida (Part 2). Annals of the South African Museum, 25, 217-312." type="journal article" year="1928">Lawrence, 1928</bibRefCitation>
</taxonomicName>
, female from Ndumo: 28. Habitus, dorsal view; 29. Eye region, anterior view; 30. Same, dorsal view; 31. Chelicerae, posteroventral view; 32. Legs I &amp; II, retrolateral view; 33. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 28 = 1.0 mm; 2933 = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C37B8AFF517FF74795B1F986FC9EFB5D" lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45" type="description">
<paragraph id="8BDED974517FF74795B1F986FA6BFE3D" blockId="6.[151,1437,1607,2030]" lastBlockId="7.[151,1437,151,1293]" lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45">
<emphasis id="B9150566517FF74695B1F986FEAFF9D5" bold="true" box="[199,345,1607,1633]" pageId="6" pageNumber="45">Description.</emphasis>
<emphasis id="B9150566517FF7469412F989FE68F9D6" box="[356,414,1608,1634]" italics="true" pageId="6" pageNumber="45">Male</emphasis>
(Ndumo, NCA 2021/1255,
<figureCitation id="135AC5F1517FF746979FF989FC86F9D6" box="[745,880,1608,1634]" captionStart="FIGURES 2127" captionStartId="5.[151,264,1193,1217]" captionTargetBox="[151,1436,179,1167]" captionTargetId="figure-17@5.[151,1436,179,1167]" captionTargetPageId="5" captionText="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." pageId="6" pageNumber="45">Figs 2227</figureCitation>
). Carapace deep red-brown, chelicerae, labium, endites and legs dark brown, sternum orange-brown, eye field slightly darker than carapace (
<figureCitation id="135AC5F1517FF74691B5F9ADFAECF932" box="[1219,1306,1644,1670]" captionStart="FIGURES 2127" captionStartId="5.[151,264,1193,1217]" captionTargetBox="[151,1436,179,1167]" captionTargetId="figure-17@5.[151,1436,179,1167]" captionTargetPageId="5" captionText="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." pageId="6" pageNumber="45">Fig. 22</figureCitation>
). Abdomen dark grey dorsally, spinnerets dark brown; carapace with faintly darker striae radiating from fovea, densely covered with white appressed hairs, cephalic part, and posterior slope with several stiff erect black setae. Fovea narrow, longitudinal, straight, dark brown. Anterior surface of cheliceral paturon with long black setae, promargin with low serrated keel and distal denticle, with keel extending along entire length of promargin (
<figureCitation id="135AC5F1517FF7469117F93DFB42F8A2" box="[1121,1204,1788,1814]" captionStart="FIGURES 2127" captionStartId="5.[151,264,1193,1217]" captionTargetBox="[151,1436,179,1167]" captionTargetId="figure-17@5.[151,1436,179,1167]" captionTargetPageId="5" captionText="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." pageId="6" pageNumber="45">Fig. 27</figureCitation>
; arrow); retromargin without keel or denticle (
<figureCitation id="135AC5F1517FF74694C6F8E1FDF2F88E" box="[432,516,1824,1850]" captionStart="FIGURES 2127" captionStartId="5.[151,264,1193,1217]" captionTargetBox="[151,1436,179,1167]" captionTargetId="figure-17@5.[151,1436,179,1167]" captionTargetPageId="5" captionText="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." pageId="6" pageNumber="45">Fig. 27</figureCitation>
). Abdomen oval, hirsute, anterior surface with dense anterior dorsal strong curved setae; dorsal abdomen anteriorly with deep red oval scutum extending to ½ abdominal length, with three pairs of distinct oval sigilla, first two pairs close together, within posterior margin of dorsal scutum, third pair at ⅗ abdominal length (
<figureCitation id="135AC5F1517FF746959BF84DFEB5F812" box="[237,323,1932,1958]" captionStart="FIGURES 2127" captionStartId="5.[151,264,1193,1217]" captionTargetBox="[151,1436,179,1167]" captionTargetId="figure-17@5.[151,1436,179,1167]" captionTargetPageId="5" captionText="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." pageId="6" pageNumber="45">Fig. 22</figureCitation>
). Metatarsi III and all tarsi with well-developed complete scopulae, scopulae on metatarsi III and IV restricted distally (
<figureCitation id="135AC5F1517FF74694FAF871FDE6F87E" box="[396,528,1967,1994]" captionStart="FIGURES 2127" captionStartId="5.[151,264,1193,1217]" captionTargetBox="[151,1436,179,1167]" captionTargetId="figure-17@5.[151,1436,179,1167]" captionTargetPageId="5" captionText="FIGURES 2127. Somatic morphology of Trichothyse fontensis Lawrence, 1928, holotype male from Sesfontein (21) and non-type male from Ndumo (2227): 21, 22. Habitus, dorsal view; 23. Eye region, anterior view; 24. Same, dorsal view; 25. Leg I, retrolateral view; 26. Leg IV, prolateral view; 27. Chelicerae, posteroventral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 2122 = 1.0 mm; 2327 = 0.5 mm." pageId="6" pageNumber="45">Figs 2526</figureCitation>
). Spinnerets hirsute. Body length 6.45. Carapace 3.05 long, 2.16 wide. Abdomen 3.57 long, 2.20 wide. Eye sizes and interdistances: ALE 0.16, AME 0.21, PLE 0.15, PME 0.14; ALEPLE 0.10, AMEAME 0.04, AMEPME 0.17, PMEPLE 0.10, PMEPME 0.14. AME and ALE nearly contiguous. Clypeus height at AMEs 0.13, at ALEs 0.12. Chelicerae 1.30 long. Length of palp and legs: palp 2.78 [0.90, 0.46, 0.32, 1.10], I 6.99 [2.11, 1.30, 1.53, 1.32, 0.73], II 7.17 [2.17, 1.31, 1.50, 1.40, 0.79], III 6.89 [2.05, 1.13, 1.35, 1.55, 0.81], IV 8.72 [2.45, 1.30, 1.85, 2.28, 0.84]. Leg formula: 4213. Spination of palp: femur pl 1 do 2, patella pl 1 do 1, tibia spineless, tarsus plv 1 rlv 1; legs: femur I pl 1 do 3, II pl 2 do 3, III pl 2 do 3 rl 2, IV pl 1 do 3 rl 1; patella I, II and IV spineless, III pl 1 rl 1; tibia III plv 1 vt 2, III pl 4 rl 2 plv 2 rlv 1 vt 2, IV pl 3 rl 2 plv 2 rlv 1 vt 2; metatarsus III plv 1 rlv 1, III pl 2 do 1 rl 2 plv 1 rlv 1 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 4; tarsus IIV spineless.
</paragraph>
<paragraph id="8BDED974517EF74795B1FE52FE84FD31" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1FE52FF0AFE19" box="[199,252,403,429]" italics="true" pageId="7" pageNumber="46">Palp</emphasis>
(
<figureCitation id="135AC5F1517EF7479478FE52FE44FE19" box="[270,434,402,429]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="7" pageNumber="46">Figs 12, 56</figureCitation>
): segments orange-brown. RTA more than half the length of cymbium, with prominent ventral branch at approximately ⅔ its length, but inconspicuous dorsal “lobe” and restricted to a dorsal angle (
<figureCitation id="135AC5F1517EF747901CFE76FF30FE41" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="7" pageNumber="46">Figs 2, 6</figureCitation>
). Cymbial flange nearly half of the length of cymbium, slightly concave along most of its length (
<figureCitation id="135AC5F1517EF7479030FE1AFA79FE41" box="[1350,1423,475,501]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="7" pageNumber="46">Fig. 2</figureCitation>
). Tegulum long-oval, sac-like (
<figureCitation id="135AC5F1517EF7479497FE3EFDD0FDAD" box="[481,550,511,537]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="7" pageNumber="46">Fig. 2</figureCitation>
). Conductor membranous, with almost right-angled bend in distal half, passing behind embolus, with proximal part parallel to embolus and smoothly rounded apex directed retrolaterally (
<figureCitation id="135AC5F1517EF747903CFDE2FA79FD89" box="[1354,1423,547,573]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="7" pageNumber="46">Fig. 2</figureCitation>
). Embolus arising apically, narrow, initially straight, looping prolaterally then distally, with spike-like tip directed at 12-oclock (
<figureCitation id="135AC5F1517EF7479456FDAAFE93FD31" box="[288,357,619,645]" captionStart="FIGURES 18" captionStartId="3.[151,264,1372,1396]" captionTargetBox="[152,1436,179,1346]" captionTargetId="figure-17@3.[151,1436,179,1346]" captionTargetPageId="3" captionText="FIGURES 18. Palpal morphology of male Afrotropical Trichothyse species in ventral (14) and retrolateral (58) views. 1, 5. T. fontensis Lawrence, 1928, holotype from Sesfontein; 2, 6. T. fontensis from Ndumo; 3, 7. T. karoo Haddad &amp; Sankaran sp. nov. from Jagerskraal; 4, 8. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo. Scale bars: 0.5 mm." pageId="7" pageNumber="46">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BDED974517EF74795B1FD4EFCB0FC59" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1FD4EFEEDFD1D" box="[199,283,655,681]" italics="true" pageId="7" pageNumber="46">Female</emphasis>
(Ndumo, NCA 2014/1417,
<figureCitation id="135AC5F1517EF7479728FD4EFD15FD1D" box="[606,739,655,681]" captionStart="FIGURES 2833" captionStartId="6.[151,264,1393,1417]" captionTargetBox="[151,1436,179,1367]" captionTargetId="figure-17@6.[151,1436,179,1368]" captionTargetPageId="6" captionText="FIGURES 2833. Somatic morphology of Trichothyse fontensis Lawrence, 1928, female from Ndumo: 28. Habitus, dorsal view; 29. Eye region, anterior view; 30. Same, dorsal view; 31. Chelicerae, posteroventral view; 32. Legs I &amp; II, retrolateral view; 33. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 28 = 1.0 mm; 2933 = 0.5 mm." pageId="7" pageNumber="46">Figs 2833</figureCitation>
). As in male except the following: habitus generally orange-brown, abdomen dark grey (
<figureCitation id="135AC5F1517EF74794A5FD72FDD1FD79" box="[467,551,691,717]" captionStart="FIGURES 2833" captionStartId="6.[151,264,1393,1417]" captionTargetBox="[151,1436,179,1367]" captionTargetId="figure-17@6.[151,1436,179,1368]" captionTargetPageId="6" captionText="FIGURES 2833. Somatic morphology of Trichothyse fontensis Lawrence, 1928, female from Ndumo: 28. Habitus, dorsal view; 29. Eye region, anterior view; 30. Same, dorsal view; 31. Chelicerae, posteroventral view; 32. Legs I &amp; II, retrolateral view; 33. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 28 = 1.0 mm; 2933 = 0.5 mm." pageId="7" pageNumber="46">Fig. 28</figureCitation>
). Abdomen with three pairs of oval sigilla, first two pairs at ⅓ abdomen length, third pair just beyond midpoint (
<figureCitation id="135AC5F1517EF747977DFD16FD96FD45" box="[523,608,727,753]" captionStart="FIGURES 2833" captionStartId="6.[151,264,1393,1417]" captionTargetBox="[151,1436,179,1367]" captionTargetId="figure-17@6.[151,1436,179,1368]" captionTargetPageId="6" captionText="FIGURES 2833. Somatic morphology of Trichothyse fontensis Lawrence, 1928, female from Ndumo: 28. Habitus, dorsal view; 29. Eye region, anterior view; 30. Same, dorsal view; 31. Chelicerae, posteroventral view; 32. Legs I &amp; II, retrolateral view; 33. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 28 = 1.0 mm; 2933 = 0.5 mm." pageId="7" pageNumber="46">Fig. 28</figureCitation>
). Body length 7.45. Carapace 3.80 long, 2.40 wide. Abdomen 3.60 long, 2.45 wide. Eye sizes and interdistances: ALE 0.22, AME 0.24, PLE 0.19, PME 0.16; ALEPLE 0.11, AMEAME 0.10, AMEPME 0.19, PMEPLE 0.14, PMEPME 0.18. Clypeus height at AMEs 0.14, at ALEs 0.11. Chelicerae 1.17 long. Length of palp and legs: palp 2.97 [1.06, 0.55, 0.49, 0.87], I 7.52 [2.16, 1.42, 1.60, 1.45, 0.89], II 7.33 [2.12, 1.32, 1.58, 1.45, 0.86], III 8.01 [2.10, 1.23, 1.41, 1.59, 1.68], IV missing. Spination of palp: femur pl 2 do 2, patella pl 1 do 1, tibia pl 2 rl 2 plv 2 rlv 1 vt 1, tarsus pl 1 rl 1 plv 2 rlv 2 vt 2; legs (both leg IV missing): femur I pl 1 do 3, II pl 2 do 3, III pl 2 do 3 rl 2; patella I and II spineless, III pl 1 rl 1; tibia III plv 1 vt 1, III pl 2 rl 2 plv 2 rlv 1 vt 2; metatarsus III plv 1 rlv 1, III pl 4 rl 2 plv 1 rlv 2 vt 4.
</paragraph>
<paragraph id="8BDED974517EF74795B1FC36FE51FB71" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1FC36FEC7FBA5" box="[199,305,1015,1041]" italics="true" pageId="7" pageNumber="46">Genitalia</emphasis>
(
<figureCitation id="135AC5F1517EF7479435FC36FE48FBA5" box="[323,446,1014,1041]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Figs 9, 15</figureCitation>
): epigyne hirsute, well sclerotised, with broadly notched posterior border, with small hood-shaped anterior atrium (
<figureCitation id="135AC5F1517EF747949AFBDAFDC2FB81" box="[492,564,1051,1077]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Fig. 9</figureCitation>
; arrow) and paired semi-circular lobes posteriorly, forming shallow median depression. Copulatory openings indistinct, possibly at the edge of posterior lobes. Copulatory ducts very short, mostly hidden by primary spermathecae (
<figureCitation id="135AC5F1517EF7479714FBA2FD42FBC9" box="[610,692,1122,1149]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Fig. 15</figureCitation>
). Primary spermathecae parallel, anteriorly forming diverging ends (
<figureCitation id="135AC5F1517EF74795E9FB46FF03FB15" box="[159,245,1158,1185]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Fig. 15</figureCitation>
). Paired secondary spermathecae short, globular, partly obscured by primary spermathecae. Fertilization ducts narrow, diverging.
</paragraph>
<paragraph id="8BDED974517EF74795B1FB0FFC9EFB5D" blockId="7.[151,1437,151,1293]" box="[199,872,1230,1257]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1FB0FFEC9FB5C" bold="true" box="[199,319,1230,1256]" pageId="7" pageNumber="46">Variation.</emphasis>
Male (n = 4): body length 6.306.75, mean 6.51.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517EF74795B1FB33FAEEFAB9" box="[199,1304,1266,1293]" pageId="7" pageNumber="46" type="distribution">
<paragraph id="8BDED974517EF74795B1FB33FAEEFAB9" blockId="7.[151,1437,151,1293]" box="[199,1304,1266,1293]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1FB33FE97FAB8" bold="true" box="[199,353,1266,1292]" pageId="7" pageNumber="46">Distribution.</emphasis>
Known only from north-western
<collectingCountry id="F37699E4517EF74797ACFB32FCCCFAB9" box="[730,826,1267,1293]" name="Namibia" pageId="7" pageNumber="46">Namibia</collectingCountry>
and north-eastern
<collectingCountry id="F37699E4517EF747917DFB32FB6CFAB9" box="[1035,1178,1267,1293]" name="South Africa" pageId="7" pageNumber="46">South Africa</collectingCountry>
(
<figureCitation id="135AC5F1517EF74791DFFB32FAFDFAB9" box="[1193,1291,1267,1293]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="7" pageNumber="46">Fig. 120</figureCitation>
).
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,254 @@
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<taxonomicName id="4C61A2F7517EF74795E1FA9FFDB4FACC" ID-CoL="7CSVJ" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[151,578,1374,1401]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="hortensis">
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<emphasis id="B9150566517EF74795E1FA9FFE6AFACD" bold="true" box="[151,412,1374,1401]" italics="true" pageId="7" pageNumber="46">Trichothyse hortensis</emphasis>
<bibRefCitation id="EFF0A485517EF74794D4FA9EFDB4FACC" author="Tucker, R. W. E." box="[418,578,1374,1401]" pageId="7" pageNumber="46" pagination="251 - 437" refId="ref20373" refString="Tucker, R. W. E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251-437." type="journal article" year="1923">Tucker, 1923</bibRefCitation>
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<paragraph id="8BDED974517EF74795E1FA42FE9FFA29" blockId="7.[151,578,1374,1437]" box="[151,361,1411,1437]" pageId="7" pageNumber="46">
<figureCitation id="135AC5F1517EF74795E1FA42FEE0FA29" box="[151,278,1411,1437]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Figs 10, 16</figureCitation>
,
<figureCitation id="135AC5F1517EF7479454FA42FE9FFA29" box="[290,361,1411,1437]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="7" pageNumber="46">3441</figureCitation>
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<subSubSection id="C37B8AFF517EF74795E1FA06FD78F9A8" pageId="7" pageNumber="46" type="reference_group">
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<bibRefCitation id="EFF0A485517EF7479401FA06FDC4FA6B" author="Tucker, R. W. E." box="[375,562,1479,1503]" pageId="7" pageNumber="46" pagination="251 - 437" refId="ref20373" refString="Tucker, R. W. E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251-437." type="journal article" year="1923">Tucker, 1923: 332</bibRefCitation>
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, plate IX, fig. 53;
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<bibRefCitation id="EFF0A485517EF747979AFA06FC53FA6B" author="Murphy, J." box="[748,933,1479,1503]" pageId="7" pageNumber="46" pagination="1 - 92" refId="ref19543" refString="Murphy, J. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambridgeshire, 1 &amp; 2, i - xii + 1-92 &amp; i - ii + 93 - 605." type="journal article" year="2007">Murphy, 2007: 50</bibRefCitation>
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, figs 382383;
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Dippenaar-Schoeman
<emphasis id="B9150566517EF7479055FA09FAA0FA6B" box="[1315,1366,1479,1503]" italics="true" pageId="7" pageNumber="46">et al.</emphasis>
, 2021: 28
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, figs 17 (
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<collectingCountry id="F37699E4517EF7479621FA27FC38FA4A" box="[855,974,1510,1534]" name="Namibia" pageId="7" pageNumber="46">NAMIBIA</collectingCountry>
:
</emphasis>
Waterberg [ca.
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,
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], leg. R.W. Tucker,
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<emphasis id="B9150566517EF74795E1F98BFE2FF9D0" bold="true" box="[151,473,1610,1636]" pageId="7" pageNumber="46">Other material examined.</emphasis>
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:
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,
<date id="FFDFFFB4517EF7479459F9AEFE6BF93D" box="[303,413,1647,1673]" pageId="7" pageNumber="46" value="2014-01-28">
<collectingDate id="EF9B065C517EF7479459F9AEFE6BF93D" box="[303,413,1647,1673]" pageId="7" pageNumber="46" value="2014-01-28">28.I.2014</collectingDate>
</date>
, leg.
<collectorName id="2694BCA2517EF74794AFF9AEFDBCF93D" box="[473,586,1647,1673]" pageId="7" pageNumber="46">N. Josling</collectorName>
(in garden amongst plants),
<specimenCount id="9D6712FD517EF74796FDF9AEFC58F93D" box="[907,942,1647,1673]" count="1" pageId="7" pageNumber="46" type="female">1♀</specimenCount>
(
<specimenCode id="DBC7710F517EF74796CAF9AEFBCCF93D" box="[956,1082,1646,1673]" collectionCode="NCA" pageId="7" pageNumber="46">NCA 2015</specimenCode>
/1903)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517EF74795B1F952FE83F965" pageId="7" pageNumber="46" type="discussion">
<paragraph id="8BDED974517EF74795B1F952FE83F965" blockId="7.[151,1437,1610,2033]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1F952FEC6F919" bold="true" box="[199,304,1683,1709]" pageId="7" pageNumber="46">Remark.</emphasis>
The
<typeStatus id="54DA67D6517EF7479418F952FE26F919" box="[366,464,1683,1709]" pageId="7" pageNumber="46" type="holotype">holotype</typeStatus>
females colouration is faded (
<figureCitation id="135AC5F1517EF7479640F952FC7AF919" box="[822,908,1683,1709]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="7" pageNumber="46">Fig. 34</figureCitation>
), so the redescription is based on the recently collected specimen.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517EF74795B1F91BFCD0F889" pageId="7" pageNumber="46" type="diagnosis">
<paragraph id="8BDED974517EF74795B1F91BFCD0F889" blockId="7.[151,1437,1610,2033]" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1F91BFEB4F940" bold="true" box="[199,322,1754,1780]" pageId="7" pageNumber="46">Diagnosis.</emphasis>
The females of
<taxonomicName id="4C61A2F7517EF7479481F91AFD8DF941" authorityName="Tucker" authorityYear="1923" box="[503,635,1755,1781]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="hortensis">
<emphasis id="B9150566517EF7479481F91AFDF1F940" box="[503,519,1755,1780]" italics="true" pageId="7" pageNumber="46">T</emphasis>
.
<emphasis id="B9150566517EF7479762F91AFD8DF941" box="[532,635,1755,1781]" italics="true" pageId="7" pageNumber="46">hortensis</emphasis>
</taxonomicName>
share with
<taxonomicName id="4C61A2F7517EF747978BF91AFC79F941" authorityName="Lawrence" authorityYear="1927" box="[765,911,1755,1781]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="subtropica">
<emphasis id="B9150566517EF747978BF91AFC79F941" box="[765,911,1755,1781]" italics="true" pageId="7" pageNumber="46">T. subtropica</emphasis>
</taxonomicName>
the convex shape of the primary spermathecae but can be distinguished from it by the far stronger curvature of the same structure, being separated by at least 1.5 times their width (cf.
<figureCitation id="135AC5F1517EF74794FFF8E2FE2EF889" box="[393,472,1827,1853]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Fig. 16</figureCitation>
and
<figureCitation id="135AC5F1517EF7479779F8E2FD97F889" box="[527,609,1827,1853]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="7" pageNumber="46">Fig. 18</figureCitation>
). Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF517EF74995B1F887FCD5FD45" lastPageId="9" lastPageNumber="48" pageId="7" pageNumber="46" type="description">
<paragraph id="8BDED974517EF74995B1F887FC56FE41" blockId="7.[151,1437,1610,2033]" lastBlockId="9.[151,1437,151,789]" lastPageId="9" lastPageNumber="48" pageId="7" pageNumber="46">
<emphasis id="B9150566517EF74795B1F887FEAFF8D4" bold="true" box="[199,345,1862,1888]" pageId="7" pageNumber="46">Description.</emphasis>
<emphasis id="B9150566517EF7479410F886FE4CF8D5" box="[358,442,1863,1889]" italics="true" pageId="7" pageNumber="46">Female</emphasis>
(Kalkfontein, NCA 2015/1903,
<figureCitation id="135AC5F1517EF747964AF886FC32F8D5" box="[828,964,1862,1889]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="7" pageNumber="46">Figs 3541</figureCitation>
). Carapace, clypeus, chelicerae, labium, endites, sternum, scutum and legs yellow-brown, eye field slightly darker than carapace, carapace with faint black mottling (
<figureCitation id="135AC5F1517EF7479470F84EFEACF81D" box="[262,346,1934,1961]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="7" pageNumber="46">Fig. 35</figureCitation>
). Abdomen dark grey, mottled creamy-grey dorsally in anterior half (
<figureCitation id="135AC5F1517EF7479113F84EFB4FF81D" box="[1125,1209,1934,1961]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="7" pageNumber="46">Fig. 35</figureCitation>
), spinnerets yellowgrey; carapace with faintly darker striae radiating from fovea, densely covered with grey appressed hairs; cephalic part, midline and posterior slope with numerous stiff erect brown setae (
<figureCitation id="135AC5F1517EF7479698F816FBB0F845" box="[1006,1094,2006,2033]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="7" pageNumber="46">Fig. 35</figureCitation>
). Fovea narrow, longitudinal, straight, brown. Anterior surface of cheliceral paturon with long brown setae, with low serrated keel ending in distinct denticle, and additional sharp denticle beyond it (
<figureCitation id="135AC5F15171F7489679FF7AFC97FF61" box="[783,865,187,213]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="8" pageNumber="47">Fig. 38</figureCitation>
; arrow); retromargin without keel, with two weakly developed, indistinct denticles. Abdomen oval, hirsute, dorsum with three pairs of distinct oval sigilla, first two pairs at ¼ abdomen length and third pair at midpoint (
<figureCitation id="135AC5F15171F7489679FEC2FC93FEA9" box="[783,869,258,285]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="8" pageNumber="47">Fig. 35</figureCitation>
). Metatarsi III and all tarsi with well-developed scopulae (
<figureCitation id="135AC5F15170F749947EFF56FE7CFF05" box="[264,394,151,177]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="9" pageNumber="48">Figs 4041</figureCitation>
), scopulae on metatarsi I and II complete while that on metatarsi III and IV restricted distally. Spinnerets hirsute (
<figureCitation id="135AC5F15170F7499406FF7AFE32FF61" box="[368,452,187,213]" captionStart="FIGURES 3441" captionStartId="8.[151,264,1882,1906]" captionTargetBox="[151,1436,342,1856]" captionTargetId="figure-86@8.[151,1436,342,1856]" captionTargetPageId="8" captionText="FIGURES 3441. Somatic morphology of Trichothyse hortensis Tucker, 1923, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm." pageId="9" pageNumber="48">Fig. 39</figureCitation>
). Body length 5.90. Carapace 2.55 long, 1.38 wide. Abdomen 3.42 long, 2.23 wide. Eye sizes and interdistances:ALE 0.15, AME 0.17, PLE 0.13, PME 0.12; ALEPLE 0.07, AMEAME 0.05, AMEPME 0.13, PMEPLE 0.10, PMEPME 0.12. AME and ALE nearly contiguous. Clypeus height at AMEs 0.13, at ALEs 0.11. Chelicerae 0.84 long. Length of palp and legs: palp 2.21 [0.78, 0.38, 0.35, 0.70], I 5.59 [1.59, 1.05, 1.15, 1.08, 0.72], II 5.53 [1.54, 1.01, 1.16, 1.10, 0.72], III 5.33 [1.52, 0.90, 1.07, 1.06, 0.78], IV 6.10 [1.82, 1.05, 1.43, 1.08, 0.72]. Leg formula: 4123. Spination of palp: femur pl 1 do 2, patella pl 1 do 2, tibia pl 2 rl 2 plv 1, tarsus pl 1 rl 1 plv 2 rlv 2 vt 2; legs: femur III pl 1 do 3, III pl 2 do 3 rl 2, IV pl 1 do 3 rl 1; patella III spineless, IIIIV pl 1 rl 1; tibia I vt 1, II plv 1 vt 1, III pl 2 rl 2 plv 2 vt 2, IV pl 2 rl 2 plv 2 rlv 1 vt 2; metatarsus I plv 1, II plv 1 rlv 1, III pl 2 rl 2 plv 1 rlv 1 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3; tarsus IIV spineless.
</paragraph>
<caption id="DF1E89FC5171F74895E1F89BFB9BF86A" pageId="8" pageNumber="47" startId="8.[151,264,1882,1906]" targetBox="[151,1436,342,1856]" targetPageId="8" targetType="figure">
<paragraph id="8BDED9745171F74895E1F89BFB9BF86A" blockId="8.[151,1437,1882,2015]" pageId="8" pageNumber="47">
<emphasis id="B91505665171F74895E1F89BFEA2F8C6" bold="true" box="[151,340,1882,1906]" pageId="8" pageNumber="47">FIGURES 3441.</emphasis>
Somatic morphology of
<taxonomicName id="4C61A2F75171F7489727F89BFC4EF8C6" authority="Tucker, 1923" authorityName="Tucker" authorityYear="1923" box="[593,952,1882,1906]" class="Arachnida" family="Gnaphosidae" genus="Trichothyse" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="hortensis">
<emphasis id="B91505665171F7489727F89BFCDDF8C6" box="[593,811,1882,1906]" italics="true" pageId="8" pageNumber="47">Trichothyse hortensis</emphasis>
<bibRefCitation id="EFF0A4855171F7489646F89BFC4EF8C6" author="Tucker, R. W. E." box="[816,952,1882,1906]" pageId="8" pageNumber="47" pagination="251 - 437" refId="ref20373" refString="Tucker, R. W. E. (1923) The Drassidae of South Africa. Annals of the South African Museum, 19, 251-437." type="journal article" year="1923">Tucker, 1923</bibRefCitation>
</taxonomicName>
, holotype female from Waterberg (34) and nontype female from Kalkfontein Dam (3541): 34, 35. Habitus, dorsal view; 36. Eye region, anterior view; 37. Same, dorsal view; 38. Chelicerae, posteroventral view; 39. Spinnerets, posteroventral view; 40. Leg I, retrolateral view; 41. Leg IV, prolateral view. Arrow indicates keel on the cheliceral promargin. Scale bars: 3435 = 1.0 mm; 3641 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDED9745170F74995B1FE3EFDDDFD79" blockId="9.[151,1437,151,789]" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995B1FE3EFEC7FDAD" box="[199,305,511,537]" italics="true" pageId="9" pageNumber="48">Genitalia</emphasis>
(
<figureCitation id="135AC5F15170F7499432FE3EFE26FDAD" box="[324,464,511,537]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="9" pageNumber="48">Figs 10, 16</figureCitation>
): epigyne originally hirsute, hairs lost during cleaning, with paired S-shaped lobes posteriorly, forming posterior border and broad shallow atrium (
<figureCitation id="135AC5F15170F749960EFDE2FC38FD89" box="[888,974,547,573]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="9" pageNumber="48">Fig. 10</figureCitation>
; arrow). Copulatory openings indistinct, probably at anterior edge of posterior lobes. Copulatory ducts very short, partly obscured by primary spermathecae (
<figureCitation id="135AC5F15170F74995E9FDAAFF05FD31" box="[159,243,619,645]" captionStart="FIGURES 920" captionStartId="4.[151,264,1607,1631]" captionTargetBox="[151,1436,179,1581]" captionTargetId="figure-17@4.[151,1436,179,1581]" captionTargetPageId="4" captionText="FIGURES 920. Epigyne morphology of Afrotropical Trichothyse species in ventral (914) and dorsal (1520) views. 9, 15. T. fontensis Lawrence, 1928 from Ndumo; 10, 16. T. hortensis Tucker, 1923, lectotype from Waterberg; 11, 17. T. karoo Haddad &amp; Sankaran sp. nov., paratype from Jagerskraal; 12, 18. T. subtropica Lawrence, 1927, lectotype from Kunene River; 13, 19. T. zuluensis (Lawrence, 1938) comb. nov. from Ndumo; 14, 20. T. africana (Tucker, 1923) from Dereham. Arrows indicate epigynal atrium. Scale bars: 0.25 mm." pageId="9" pageNumber="48">Fig. 16</figureCitation>
). Primary spermathecae strongly convex, forming diamond-shaped space between them, at least 1.5 times their width. Paired secondary spermathecae short, globular, partly obscured by primary spermathecae. Fertilization ducts narrow, finger-like, diverging.
</paragraph>
<paragraph id="8BDED9745170F74995B1FD17FCD5FD45" blockId="9.[151,1437,151,789]" box="[199,803,726,753]" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995B1FD17FEC9FD44" bold="true" box="[199,319,726,752]" pageId="9" pageNumber="48">Variation.</emphasis>
The
<typeStatus id="54DA67D65170F749940EFD16FE2CFD45" box="[376,474,727,753]" pageId="9" pageNumber="48" type="holotype">holotype</typeStatus>
female is
<quantity id="4C9974915170F7499738FD16FD44FD45" box="[590,690,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="9.02" pageId="9" pageNumber="48" unit="mm" value="9.02">9.02 mm</quantity>
in length.
</paragraph>
</subSubSection>
<subSubSection id="C37B8AFF5170F74995B1FD3BFA9CFCA1" box="[199,1386,762,789]" pageId="9" pageNumber="48" type="distribution">
<paragraph id="8BDED9745170F74995B1FD3BFA9CFCA1" blockId="9.[151,1437,151,789]" box="[199,1386,762,789]" pageId="9" pageNumber="48">
<emphasis id="B91505665170F74995B1FD3BFE97FCA0" bold="true" box="[199,353,762,788]" pageId="9" pageNumber="48">Distribution.</emphasis>
Known only from northern
<collectingCountry id="F37699E45170F74997EBFD3AFD0BFCA1" box="[669,765,763,789]" name="Namibia" pageId="9" pageNumber="48">Namibia</collectingCountry>
and the
<collectingRegion id="49A517965170F749962BFD3AFBCFFCA1" box="[861,1081,763,789]" country="South Africa" name="Free State" pageId="9" pageNumber="48">Free State Province</collectingRegion>
of
<collectingCountry id="F37699E45170F7499128FD3AFB1AFCA1" box="[1118,1260,763,789]" name="South Africa" pageId="9" pageNumber="48">South Africa</collectingCountry>
(
<figureCitation id="135AC5F15170F749918DFD3AFAA8FCA1" box="[1275,1374,763,789]" captionStart="FIGURE 120" captionStartId="26.[151,250,1961,1985]" captionTargetBox="[151,1429,1029,1932]" captionTargetId="figure-441@26.[151,1436,1026,1935]" captionTargetPageId="26" captionText="FIGURE 120. Distribution records of the known Afrotropical Trichothyse species." pageId="9" pageNumber="48">Fig. 120</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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Kunt, Danışman and Yağmur
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,
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asl
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,
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,
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.
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(
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),
<collectingCountry id="F3717298FFEAF162FDBA0DD4FDE3F9C4" box="[530,620,1530,1555]" name="Turkey" pageId="1" pageNumber="398">TURKEY</collectingCountry>
:
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,
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,
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north of
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(
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,
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), c
</materialsCitation>
.
<quantity id="4C9E9FEDFFEAF162FD070E33FD79F9E1" box="[687,758,1565,1590]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="1" pageNumber="398" unit="m" value="922.0">922 m</quantity>
asl,
<date id="FFD814C8FFEAF162FC810E33FB81F9E1" box="[809,1038,1565,1590]" pageId="1" pageNumber="398" value="2015-04-05" valueMax="2015-05-05" valueMin="2015-04-05">05 April-5 May 2015</date>
, MSS trap, Leg. E.A.
<materialsCitation id="3B0E3855FFEAF162FEA20E11FDCCF9AC" collectionCode="HZH" country="Turkey" county="Yagmur" latitude="38.353058" location="Kuslar (Dagmarmara) Village" longLatPrecision="1" longitude="-27.839167" municipality="Turgutlu District" pageId="1" pageNumber="398" specimenCount="1" specimenCount-male="1" stateProvince="Manisa" typeStatus="paratype">
<collectingCounty id="62B84A84FFEAF162FEA20E11FEEDF98F" box="[266,354,1599,1624]" pageId="1" pageNumber="398">Yağmur</collectingCounty>
<specimenCount id="9D60F981FFEAF162FE2A0E11FE22F980" box="[386,429,1599,1624]" count="1" pageId="1" pageNumber="398" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED77AACDFFEAF162FE140E11FE7CF98F" box="[444,499,1599,1624]" pageId="1" pageNumber="398">HZH</collectionCode>
),
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,
<collectingMunicipality id="6BBDA872FFEAF162FD660E11FC06F98F" box="[718,905,1599,1624]" pageId="1" pageNumber="398">Turgutlu District</collectingMunicipality>
,
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(
<geoCoordinate id="EE5254CFFFEAF162FF5E0E4CFE07F9AC" box="[246,392,1634,1659]" degrees="38.353056" direction="north" orientation="latitude" pageId="1" pageNumber="398" precision="1" value="38.353058">38.353056°N</geoCoordinate>
,
<geoCoordinate id="EE5254CFFFEAF162FE380E4CFDA9F9AC" box="[400,550,1634,1659]" degrees="27.839167" direction="west" orientation="longitude" pageId="1" pageNumber="398" precision="1" value="-27.839167">27.839167°W</geoCoordinate>
), c
</materialsCitation>
.
<quantity id="4C9E9FEDFFEAF162FDE40E4CFD1EF9AC" box="[588,657,1634,1659]" metricMagnitude="2" metricUnit="m" metricValue="9.62" pageId="1" pageNumber="398" unit="m" value="962.0">962 m</quantity>
asl,
<date id="FFD814C8FFEAF162FD170E4CFC24F9AC" box="[703,939,1634,1659]" pageId="1" pageNumber="398" value="2016-05-10" valueMax="2016-06-26" valueMin="2016-05-10">10 May-26 June 2016</date>
, pitfall traps, Leg. E. A.
<materialsCitation id="3B0E3855FFEAF162FF150EABFE72F917" collectingDate="2017-11-26" collectingDateMax="2018-03-02" collectingDateMin="2017-11-26" collectionCode="AZMM, KUAM" collectorName="Leg" country="Turkey" location="Yagmur - Same" pageId="1" pageNumber="398" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<location id="8EB964D3FFEAF162FF150EABFEFFF949" LSID="urn:lsid:plazi:treatment:03CF831EFFEAF164FEBA0D21FCD8FBA3:8EB964D3FFEAF162FF150EABFEFFF949" box="[189,368,1669,1694]" country="Turkey" name="Yagmur - Same" pageId="1" pageNumber="398">Yağmur Same</location>
place but
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(
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),
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(
<collectionCode id="ED77AACDFFEAF162FD1E0EABFC89F949" box="[694,774,1669,1694]" pageId="1" pageNumber="398">AZMM</collectionCode>
),
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(
<collectionCode id="ED77AACDFFEAF162FCE50EABFC19F949" box="[845,918,1669,1694]" pageId="1" pageNumber="398">KUAM</collectionCode>
)
<date id="FFD814C8FFEAF162FC0C0EABFBF2F949" box="[932,1149,1669,1694]" pageId="1" pageNumber="398" value="2017-11-26">
<collectingDate id="EF9CED20FFEAF162FC0C0EABFBF2F949" box="[932,1149,1669,1694]" pageId="1" pageNumber="398" value="2017-11-26">26 November 2017</collectingDate>
</date>
-
<date id="FFD814C8FFEAF162FF080E89FEB4F917" box="[160,315,1703,1728]" pageId="1" pageNumber="398" value="2018-03-02">
<collectingDate id="EF9CED20FFEAF162FF080E89FEB4F917" box="[160,315,1703,1728]" pageId="1" pageNumber="398" value="2018-03-02">2 March 2018</collectingDate>
</date>
,
<collectingMethod id="52274A1FFFEAF162FEEE0E89FE47F917" box="[326,456,1703,1728]" pageId="1" pageNumber="398">pitfall traps</collectingMethod>
,
<collectorName id="269357DEFFEAF162FE7A0E89FE72F917" box="[466,509,1703,1728]" pageId="1" pageNumber="398">Leg</collectorName>
</materialsCitation>
. E.A.
<materialsCitation id="3B0E3855FFEAF162FD900E89FD1CF917" box="[568,659,1703,1728]" country="Turkey" location="Yagmur" pageId="1" pageNumber="398" specimenCount="1" typeStatus="paratype">
<location id="8EB964D3FFEAF162FD900E89FD1CF917" LSID="urn:lsid:plazi:treatment:03CF831EFFEAF164FEBA0D21FCD8FBA3:8EB964D3FFEAF162FD900E89FD1CF917" box="[568,659,1703,1728]" country="Turkey" name="Yagmur" pageId="1" pageNumber="398">Yağmur</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF196280FFE9F161FF080E8DFB0BF923" pageId="2" pageNumber="399" startId="2.[160,225,1699,1722]" targetBox="[166,1141,147,1654]" targetPageId="2" targetType="figure">
<paragraph id="8BD93208FFE9F161FF080E8DFB0BF923" blockId="2.[160,1157,1699,1780]" pageId="2" pageNumber="399">
<emphasis id="B912EE1AFFE9F161FF080E8DFF75F96D" bold="true" box="[160,250,1699,1722]" pageId="2" pageNumber="399">Figure 1.</emphasis>
<taxonomicName id="4C66498BFFE9F161FEA90E8DFE77F96D" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[257,504,1699,1722]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. nov.">
<emphasis id="B912EE1AFFE9F161FEA90E8DFE77F96D" box="[257,504,1699,1722]" italics="true" pageId="2" pageNumber="399">Harpactea bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE9F161FE560E8DFDB7F96D" bold="true" box="[510,568,1699,1722]" pageId="2" pageNumber="399">
<taxonomicNameLabel id="A2215361FFE9F161FE560E8DFDB7F96D" box="[510,568,1699,1722]" pageId="2" pageNumber="399" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus, T tegulum.
</paragraph>
</caption>
<caption id="DF196280FFE8F160FF080C46FE10FB23" pageId="3" pageNumber="400" startId="3.[160,225,1128,1151]" targetBox="[163,1153,142,1096]" targetPageId="3" targetType="figure">
<paragraph id="8BD93208FFE8F160FF080C46FE10FB23" blockId="3.[160,1157,1128,1268]" pageId="3" pageNumber="400">
<emphasis id="B912EE1AFFE8F160FF080C46FF76FBA8" bold="true" box="[160,249,1128,1151]" pageId="3" pageNumber="400">Figure 2.</emphasis>
<taxonomicName id="4C66498BFFE8F160FEA80C46FE78FBA8" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[256,503,1128,1151]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. nov.">
<emphasis id="B912EE1AFFE8F160FEA80C46FE78FBA8" box="[256,503,1128,1151]" italics="true" pageId="3" pageNumber="400">Harpactea bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE8F160FE540C46FDBAFBA8" bold="true" box="[508,565,1128,1151]" pageId="3" pageNumber="400">
<taxonomicNameLabel id="A2215361FFE8F160FE540C46FDBAFBA8" box="[508,565,1128,1151]" pageId="3" pageNumber="400" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(AC) Vulva, dorsal view from different angles; in Figure 2C, the outlines of the posterior diverticulum are drawn and highlighted for visibility. (D) Same, ventral view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, PD posterior diverticulum, RS roundish structures, S spermatheca, SB sclerotised belt, SK spermathecal keel, TB transversal bar.
</paragraph>
</caption>
<subSubSection id="C37C6183FFE8F160FF080D12FBFBFA4C" pageId="3" pageNumber="400" type="etymology">
<paragraph id="8BD93208FFE8F160FF080D12FE92FA81" blockId="3.[160,1155,1340,1435]" box="[160,285,1340,1366]" pageId="3" pageNumber="400">
<emphasis id="B912EE1AFFE8F160FF080D12FE92FA81" bold="true" box="[160,285,1340,1366]" italics="true" pageId="3" pageNumber="400">Etymology</emphasis>
</paragraph>
<paragraph id="8BD93208FFE8F160FF080D71FBFBFA4C" blockId="3.[160,1155,1340,1435]" pageId="3" pageNumber="400">
<materialsCitation id="3B0E3855FFE8F160FF080D71FBFFFA4C" country="Turkey" location="The" municipality="Bozdaglar'" pageId="3" pageNumber="400" specimenCount="1" stateProvince="Izmir">
<location id="8EB964D3FFE8F160FF080D71FF46FAAF" LSID="urn:lsid:plazi:treatment:03CF831EFFEAF164FEBA0D21FCD8FBA3:8EB964D3FFE8F160FF080D71FF46FAAF" box="[160,201,1375,1400]" country="Turkey" municipality="Bozdaglar'" name="The" pageId="3" pageNumber="400" stateProvince="Izmir">The</location>
specific epithet is a toponym for the
<collectingMunicipality id="6BBDA872FFE8F160FD2D0D71FD70FAAF" box="[645,767,1375,1400]" pageId="3" pageNumber="400">Bozdağlar</collectingMunicipality>
, a mountain range located in the provinces of
<collectingRegion id="49A2FCEAFFE8F160FE9A0DACFEE6FA4C" box="[306,361,1410,1435]" country="Turkey" name="Izmir" pageId="3" pageNumber="400">İzmir</collectingRegion>
and
<collectingRegion id="49A2FCEAFFE8F160FE0A0DACFE7EFA4C" box="[418,497,1410,1435]" country="Turkey" name="Manisa" pageId="3" pageNumber="400">Manisa</collectingRegion>
in
<collectingCountry id="F3717298FFE8F160FDBD0DACFDECFA4C" box="[533,611,1410,1435]" name="Turkey" pageId="3" pageNumber="400">Turkey</collectingCountry>
, including the
<typeStatus id="54DD8CAAFFE8F160FCA20DACFCB3FA4C" box="[778,828,1410,1435]" pageId="3" pageNumber="400">type</typeStatus>
locality of the new species
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFE8F167FF080DF8FBAEFA4C" lastPageId="4" lastPageNumber="401" pageId="3" pageNumber="400" type="diagnosis">
<paragraph id="8BD93208FFE8F160FF080DF8FE9BFA27" blockId="3.[160,1155,1494,1728]" box="[160,276,1494,1520]" pageId="3" pageNumber="400">
<emphasis id="B912EE1AFFE8F160FF080DF8FE9BFA27" bold="true" box="[160,276,1494,1520]" italics="true" pageId="3" pageNumber="400">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD93208FFE8F167FF080DD4FF6CFB7F" blockId="3.[160,1155,1494,1728]" lastBlockId="4.[160,1156,1098,1435]" lastPageId="4" lastPageNumber="401" pageId="3" pageNumber="400">
The male copulatory organ of
<taxonomicName id="4C66498BFFE8F160FE540DD4FD36F9C4" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[508,697,1530,1555]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFE8F160FE540DD4FD36F9C4" box="[508,697,1530,1555]" italics="true" pageId="3" pageNumber="400">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE8F160FD6B0DD4FC8AF9C4" bold="true" box="[707,773,1530,1555]" pageId="3" pageNumber="400">
<taxonomicNameLabel id="A2215361FFE8F160FD6B0DD4FC8AF9C4" box="[707,773,1530,1555]" pageId="3" pageNumber="400" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
shows affinity with
<taxonomicName id="4C66498BFFE8F160FC450DD4FEB9F9E1" authority="Lazarov, 2006" authorityName="Lazarov" authorityYear="2006" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="B912EE1AFFE8F160FC450DD4FB0CF9C4" box="[1005,1155,1530,1555]" italics="true" pageId="3" pageNumber="400">H. alexandrae</emphasis>
<bibRefCitation id="EFF74FF9FFE8F160FF080E33FEB9F9E1" author="Lazarov S" box="[160,310,1565,1590]" pageId="3" pageNumber="400" pagination="13 - 16" refId="ref5882" refString="Lazarov S. 2006. A New Spider Species from Bulgaria, Harpactea alexandrae sp. n. (Araneae: Dysderidae). Acta Zool. Bulg. 58: 13-16." type="journal article" year="2006">Lazarov, 2006</bibRefCitation>
</taxonomicName>
, whose
<typeStatus id="54DD8CAAFFE8F160FE380E33FE4EF9E1" box="[400,449,1565,1590]" pageId="3" pageNumber="400">type</typeStatus>
locality is
<collectingCountry id="F3717298FFE8F160FD9B0E33FD00F9E1" box="[563,655,1565,1590]" name="Bulgaria" pageId="3" pageNumber="400">Bulgaria</collectingCountry>
. However, in
<taxonomicName id="4C66498BFFE8F160FC890E33FC3BF9E1" authorityName="Lazarov" authorityYear="2006" box="[801,948,1565,1590]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="B912EE1AFFE8F160FC890E33FC3BF9E1" box="[801,948,1565,1590]" italics="true" pageId="3" pageNumber="400">H. alexandrae</emphasis>
</taxonomicName>
, the embolus (E) is longer than the conductor (C) (see
<bibRefCitation id="EFF74FF9FFE8F160FD870E11FD4DF98F" author="Lazarov S" box="[559,706,1599,1624]" pageId="3" pageNumber="400" pagination="13 - 16" refId="ref5882" refString="Lazarov S. 2006. A New Spider Species from Bulgaria, Harpactea alexandrae sp. n. (Araneae: Dysderidae). Acta Zool. Bulg. 58: 13-16." type="journal article" year="2006">Lazarov 2006</bibRefCitation>
, figs. 2, 3). Also, the angle between the embolus and the conductor is much narrower in
<taxonomicName id="4C66498BFFE8F160FCAA0E4CFC46F9AC" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[770,969,1634,1659]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFE8F160FCAA0E4CFC46F9AC" box="[770,969,1634,1659]" italics="true" pageId="3" pageNumber="400">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE8F160FC7F0E4CFB91F9AC" bold="true" box="[983,1054,1634,1659]" pageId="3" pageNumber="400">
<taxonomicNameLabel id="A2215361FFE8F160FC7F0E4CFB91F9AC" box="[983,1054,1634,1659]" pageId="3" pageNumber="400" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C66498BFFE8F160FF080EABFEBDF949" authorityName="Lazarov" authorityYear="2006" box="[160,306,1669,1694]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="B912EE1AFFE8F160FF080EABFEBDF949" box="[160,306,1669,1694]" italics="true" pageId="3" pageNumber="400">H. alexandrae</emphasis>
</taxonomicName>
(
<figureCitation id="135D2E8DFFE8F160FE970EABFE5FF949" box="[319,464,1669,1694]" captionStart="Figure 2" captionStartId="3.[160,225,1128,1151]" captionTargetBox="[163,1153,142,1096]" captionTargetId="figure-131@3.[163,1153,142,1096]" captionTargetPageId="3" captionText="Figure 2. Harpactea bozdaglarensis sp. n. (AC) Vulva, dorsal view from different angles; in Figure 2C, the outlines of the posterior diverticulum are drawn and highlighted for visibility. (D) Same, ventral view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, PD posterior diverticulum, RS roundish structures, S spermatheca, SB sclerotised belt, SK spermathecal keel, TB transversal bar." pageId="3" pageNumber="400">Figures 2BC</figureCitation>
,
<figureCitation id="135D2E8DFFE8F160FE700EABFD9CF949" box="[472,531,1669,1694]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="3" pageNumber="400">4AC</figureCitation>
). The vulva of the females of these two species is entirely different. Whereas in
<taxonomicName id="4C66498BFFE8F160FE300E89FDD7F917" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[408,600,1703,1728]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFE8F160FE300E89FDD7F917" box="[408,600,1703,1728]" italics="true" pageId="3" pageNumber="400">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE8F160FDCB0E89FD29F917" bold="true" box="[611,678,1703,1728]" pageId="3" pageNumber="400">
<taxonomicNameLabel id="A2215361FFE8F160FDCB0E89FD29F917" box="[611,678,1703,1728]" pageId="3" pageNumber="400" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, the distal end of the spermatheca (S) is concave or straight, the middle is bulging, in
<taxonomicName id="4C66498BFFEFF167FD180C64FCC9FBB4" authorityName="Lazarov" authorityYear="2006" box="[688,838,1098,1123]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="B912EE1AFFEFF167FD180C64FCC9FBB4" box="[688,838,1098,1123]" italics="true" pageId="4" pageNumber="401">H. alexandrae</emphasis>
</taxonomicName>
, the spermatheca is almost straight, and the tip is narrow and convex (see
<figureCitation id="135D2E8DFFEFF167FD310C43FCAEFB51" box="[665,801,1133,1158]" captionStart="Figure 2" captionStartId="3.[160,225,1128,1151]" captionTargetBox="[163,1153,142,1096]" captionTargetId="figure-131@3.[163,1153,142,1096]" captionTargetPageId="3" captionText="Figure 2. Harpactea bozdaglarensis sp. n. (AC) Vulva, dorsal view from different angles; in Figure 2C, the outlines of the posterior diverticulum are drawn and highlighted for visibility. (D) Same, ventral view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, PD posterior diverticulum, RS roundish structures, S spermatheca, SB sclerotised belt, SK spermathecal keel, TB transversal bar." pageId="4" pageNumber="401">Figure 2AD</figureCitation>
;
<bibRefCitation id="EFF74FF9FFEFF167FC830C43FBF1FB51" author="Spiegelaere WD &amp; Bosmans R" box="[811,1150,1133,1158]" pageId="4" pageNumber="401" pagination="7 - 16" refId="ref6003" refString="Spiegelaere WD, Bosmans R. 2009. Spider fauna in a grass dune remnant at the Black Sea coast (Romania), presenting seven new species for the Romanian fauna including the first description of the female Harpactea alexandrae Lazarov, 2006 (Dysderidae). Analele Stiintifice ale Universitatii &quot; Al. I. Cuza &quot; Iasi, s. Biologie animala 55: 7-16." type="journal article" year="2009">Spiegelaere and Bosmans 2009</bibRefCitation>
, fig. 2).
</paragraph>
<caption id="DF196280FFEFF167FF080B56FF75FBD2" pageId="4" pageNumber="401" startId="4.[160,225,888,911]" targetBox="[172,1149,145,852]" targetPageId="4" targetType="figure">
<paragraph id="8BD93208FFEFF167FF080B56FF75FBD2" blockId="4.[160,1155,888,1029]" pageId="4" pageNumber="401">
<emphasis id="B912EE1AFFEFF167FF080B56FF76FC58" bold="true" box="[160,249,888,911]" pageId="4" pageNumber="401">Figure 3.</emphasis>
<taxonomicName id="4C66498BFFEFF167FF570B56FE4DFC58" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[255,450,888,911]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. nov.">
<emphasis id="B912EE1AFFEFF167FF570B56FE4DFC58" box="[255,450,888,911]" italics="true" pageId="4" pageNumber="401">Harpactea dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEFF167FE600B56FD8EFC58" bold="true" box="[456,513,888,911]" pageId="4" pageNumber="401">
<taxonomicNameLabel id="A2215361FFEFF167FE600B56FD8EFC58" box="[456,513,888,911]" pageId="4" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum.
</paragraph>
</caption>
<paragraph id="8BD93208FFEFF167FF680C9CFBAEFA4C" blockId="4.[160,1156,1098,1435]" pageId="4" pageNumber="401">
Another species resembling
<taxonomicName id="4C66498BFFEFF167FDA50C9CFD5EFB1C" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[525,721,1202,1227]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFEFF167FDA50C9CFD5EFB1C" box="[525,721,1202,1227]" italics="true" pageId="4" pageNumber="401">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEFF167FD740C9CFCAFFB1C" bold="true" box="[732,800,1202,1227]" pageId="4" pageNumber="401">
<taxonomicNameLabel id="A2215361FFEFF167FD740C9CFCAFFB1C" box="[732,800,1202,1227]" pageId="4" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is
<taxonomicName id="4C66498BFFEFF167FCE00C9CFBF1FB1C" authority="Bosmans, 2009" authorityName="Bosmans" authorityYear="2009" box="[840,1150,1202,1227]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="clementi">
<emphasis id="B912EE1AFFEFF167FCE00C9CFC49FB1C" box="[840,966,1202,1227]" italics="true" pageId="4" pageNumber="401">H. clementi</emphasis>
Bosmans, 2009
</taxonomicName>
. However, while the tegulum is cylindrical in
<taxonomicName id="4C66498BFFEFF167FD3E0CFBFCD9FB39" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[662,854,1237,1262]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFEFF167FD3E0CFBFCD9FB39" box="[662,854,1237,1262]" italics="true" pageId="4" pageNumber="401">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEFF167FCF50CFBFC11FB39" bold="true" box="[861,926,1237,1262]" pageId="4" pageNumber="401">
<taxonomicNameLabel id="A2215361FFEFF167FCF50CFBFC11FB39" box="[861,926,1237,1262]" pageId="4" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, it is pear-shaped in
<taxonomicName id="4C66498BFFEFF167FF080CD9FE93FAC7" authorityName="Bosmans" authorityYear="2009" box="[160,284,1271,1296]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="clementi">
<emphasis id="B912EE1AFFEFF167FF080CD9FE93FAC7" box="[160,284,1271,1296]" italics="true" pageId="4" pageNumber="401">H. clementi</emphasis>
</taxonomicName>
. In addition, the embolus, which is cylindrical in
<taxonomicName id="4C66498BFFEFF167FCF10CD9FC5AFAC7" authorityName="Bosmans" authorityYear="2009" box="[857,981,1271,1296]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="clementi">
<emphasis id="B912EE1AFFEFF167FCF10CD9FC5AFAC7" box="[857,981,1271,1296]" italics="true" pageId="4" pageNumber="401">H. clementi</emphasis>
</taxonomicName>
, is flattened in
<taxonomicName id="4C66498BFFEFF167FF080D34FEEAFAE4" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[160,357,1306,1331]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFEFF167FF080D34FEEAFAE4" box="[160,357,1306,1331]" italics="true" pageId="4" pageNumber="401">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEFF167FED90D34FE39FAE4" bold="true" box="[369,438,1306,1331]" pageId="4" pageNumber="401">
<taxonomicNameLabel id="A2215361FFEFF167FED90D34FE39FAE4" box="[369,438,1306,1331]" pageId="4" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
When the females of these two species are compared, it is characteristic that the anchor-like chitinous parts of the vulva of
<taxonomicName id="4C66498BFFEFF167FC3C0D13FBD8FA81" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[916,1111,1341,1366]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFEFF167FC3C0D13FBD8FA81" box="[916,1111,1341,1366]" italics="true" pageId="4" pageNumber="401">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEFF167FBC90D13FF39FAAF" bold="true" pageId="4" pageNumber="401">
<taxonomicNameLabel id="A2215361FFEFF167FBC90D13FF39FAAF" pageId="4" pageNumber="401" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are wider than those of
<taxonomicName id="4C66498BFFEFF167FE600D71FDCEFAAF" authorityName="Bosmans" authorityYear="2009" box="[456,577,1375,1400]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="401" phylum="Arthropoda" rank="species" species="clementi">
<emphasis id="B912EE1AFFEFF167FE600D71FDCEFAAF" box="[456,577,1375,1400]" italics="true" pageId="4" pageNumber="401">H. clementi</emphasis>
</taxonomicName>
and that the vulva of the new species has a distinct sclerotised belt (see
<figureCitation id="135D2E8DFFEFF167FE2E0DACFD97FA4C" box="[390,536,1410,1435]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="4" pageNumber="401">Figures 1BC</figureCitation>
and
<figureCitation id="135D2E8DFFEFF167FDFA0DACFD00FA4C" box="[594,655,1410,1435]" captionStart="Figure 2" captionStartId="3.[160,225,1128,1151]" captionTargetBox="[163,1153,142,1096]" captionTargetId="figure-131@3.[163,1153,142,1096]" captionTargetPageId="3" captionText="Figure 2. Harpactea bozdaglarensis sp. n. (AC) Vulva, dorsal view from different angles; in Figure 2C, the outlines of the posterior diverticulum are drawn and highlighted for visibility. (D) Same, ventral view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, PD posterior diverticulum, RS roundish structures, S spermatheca, SB sclerotised belt, SK spermathecal keel, TB transversal bar." pageId="4" pageNumber="401">2AD</figureCitation>
;
<bibRefCitation id="EFF74FF9FFEFF167FD340DACFC25FA4C" author="Keer JV &amp; Bosmans R" box="[668,938,1410,1435]" pageId="4" pageNumber="401" pagination="277 - 285" refId="ref5846" refString="Keer JV, Bosmans R. 2009. On some new Harpactea and Stalagtia species from Lesbos, Greece (Araneae: Dysderidae). Acta Zool. Bulg. 61: 277-285." type="journal article" year="2009">Keer and Bosmans 2009</bibRefCitation>
, figs. 16).
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFEFF164FF080DF8FB0CFD04" lastPageId="7" lastPageNumber="404" pageId="4" pageNumber="401" type="description">
<paragraph id="8BD93208FFEFF167FF080DF8FEA9FA27" blockId="4.[160,1155,1494,1728]" box="[160,294,1494,1520]" pageId="4" pageNumber="401">
<emphasis id="B912EE1AFFEFF167FF080DF8FEA9FA27" bold="true" box="[160,294,1494,1520]" italics="true" pageId="4" pageNumber="401">Description</emphasis>
</paragraph>
<paragraph id="8BD93208FFEFF167FF080DD7FC9FF9AC" blockId="4.[160,1155,1494,1728]" pageId="4" pageNumber="401">
<emphasis id="B912EE1AFFEFF167FF080DD7FEDAF9C4" bold="true" box="[160,341,1529,1555]" italics="true" pageId="4" pageNumber="401">Measurements.</emphasis>
<emphasis id="B912EE1AFFEFF167FECC0DD4FE53F9C4" bold="true" box="[356,476,1530,1555]" pageId="4" pageNumber="401">
[
<typeStatus id="54DD8CAAFFEFF167FEC30DD4FE53F9C4" box="[363,476,1530,1555]" pageId="4" pageNumber="401" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFEFF167FE4F0DD4FE74F9C6" box="[487,507,1530,1553]" count="1" pageId="4" pageNumber="401" type="male"></specimenCount>
<emphasis id="B912EE1AFFEFF167FE530DD4FDF2F9C4" bold="true" box="[507,637,1530,1555]" pageId="4" pageNumber="401">
/
<typeStatus id="54DD8CAAFFEFF167FDAB0DD4FDF2F9C4" box="[515,637,1530,1555]" pageId="4" pageNumber="401" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFEFF167FD200DD4FD13F9C6" box="[648,668,1530,1553]" count="1" pageId="4" pageNumber="401" type="female"></specimenCount>
<emphasis id="B912EE1AFFEFF167FD340DD4FD2BF9C4" bold="true" box="[668,676,1530,1555]" pageId="4" pageNumber="401">]</emphasis>
TL 5.60/5.706.65; AL 2.91/3.103.55; CL 2.69/2.603.10;
<subScript id="17E2304DFFEFF167FEFF0E33FE2EF9E0" attach="left" box="[343,417,1565,1591]" fontSize="7" pageId="4" pageNumber="401">CWmax</subScript>
2.02/2.052.40;
<subScript id="17E2304DFFEFF167FDCC0E33FD23F9E0" attach="left" box="[612,684,1565,1591]" fontSize="7" pageId="4" pageNumber="401">CWmin</subScript>
0.87/0.901.05; Clh 0.08/0.060.07; Aed 0.12/0.110.11; iAE 0.08/0.060.08; PLEd 0.11/0.100.12; PMEd 0.10/0.080.10; ChF 0.61/0.530.54; ChG 0.31/0.230.25; ChL 1.24/1.001.25.
</paragraph>
<paragraph id="8BD93208FFEFF167FF680EABFCAEF917" blockId="4.[160,1155,1494,1728]" pageId="4" pageNumber="401">
<emphasis id="B912EE1AFFEFF167FF680EABFEBBF949" bold="true" box="[192,308,1669,1694]" pageId="4" pageNumber="401">
[
<typeStatus id="54DD8CAAFFEFF167FF6E0EABFEBBF949" box="[198,308,1669,1694]" pageId="4" pageNumber="401" type="paratype">Paratype</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFEFF167FE930EABFEC0F94B" box="[315,335,1669,1692]" count="1" pageId="4" pageNumber="401" type="male"></specimenCount>
<emphasis id="B912EE1AFFEFF167FEE70EABFED8F949" bold="true" box="[335,343,1669,1694]" pageId="4" pageNumber="401">]</emphasis>
TL 5.58; AL 2.94; CL 2.64;
<subScript id="17E2304DFFEFF167FDDF0EABFD4EF948" attach="left" box="[631,705,1669,1695]" fontSize="7" pageId="4" pageNumber="401">CWmax</subScript>
1.92;
<subScript id="17E2304DFFEFF167FCAA0EABFCC7F948" attach="left" box="[770,840,1669,1695]" fontSize="7" pageId="4" pageNumber="401">CWmin</subScript>
0.77; Clh 0.06; Aed 0.12; iAE 0.09; PLEd 0.12; PMEd 0.11; ChF 0.61; ChG 0.32; ChL 1.16.
</paragraph>
<caption id="DF196280FFEEF166FF080C8AFE48FB21" pageId="5" pageNumber="402" startId="5.[160,225,1188,1211]" targetBox="[163,1153,142,1157]" targetPageId="5" targetType="figure">
<paragraph id="8BD93208FFEEF166FF080C8AFE48FB21" blockId="5.[160,1156,1188,1270]" pageId="5" pageNumber="402">
<emphasis id="B912EE1AFFEEF166FF080C8AFF75FB6C" bold="true" box="[160,250,1188,1211]" pageId="5" pageNumber="402">Figure 4.</emphasis>
Scanning electron microscopy images of the male palp. (AC)
<taxonomicName id="4C66498BFFEEF166FC2A0C8AFB0BFB6C" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[898,1156,1188,1211]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="402" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. nov.">
<emphasis id="B912EE1AFFEEF166FC2A0C8AFB0BFB6C" box="[898,1156,1188,1211]" italics="true" pageId="5" pageNumber="402">Harpactea bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEEF166FF080CECFF53FB0E" bold="true" box="[160,220,1218,1241]" pageId="5" pageNumber="402">
<taxonomicNameLabel id="A2215361FFEEF166FF080CECFF53FB0E" box="[160,220,1218,1241]" pageId="5" pageNumber="402" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(DE)
<taxonomicName id="4C66498BFFEEF166FE8A0CECFE61FB0E" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[290,494,1218,1241]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="402" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. nov.">
<emphasis id="B912EE1AFFEEF166FE8A0CECFE61FB0E" box="[290,494,1218,1241]" italics="true" pageId="5" pageNumber="402">Harpactea dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEEF166FE5D0CECFDBEFB0E" bold="true" box="[501,561,1218,1241]" pageId="5" pageNumber="402">
<taxonomicNameLabel id="A2215361FFEEF166FE5D0CECFDBEFB0E" box="[501,561,1218,1241]" pageId="5" pageNumber="402" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet.
</paragraph>
</caption>
<paragraph id="8BD93208FFEEF166FF680D63FC93F9C4" blockId="5.[160,1156,1357,1728]" pageId="5" pageNumber="402">Middle-sized harpacteine spider. Carapace reddish brown with smooth surface. Eye region and margins of thoracic region darker than general. Fovea longitudinal, blackish, distinct. Surface of carapace with sparse yellowish short setae. Setae around eyes relatively long compared to others. Anterior margin of eye region convex, while posterior margin of thoracic region straight. Eyes silver-coloured, arranged annularly. AE&gt;PLE&gt;PME. AE seperated; AE-PLE 0.01, PLE-PME 0.03, PME-PME 0.02.</paragraph>
<paragraph id="8BD93208FFEEF165FF680E33FD30FF1D" blockId="5.[160,1156,1357,1728]" lastBlockId="6.[160,1156,142,445]" lastPageId="6" lastPageNumber="403" pageId="5" pageNumber="402">
Sternum glabrous, orangish-red; length 1.54, width 1.19. Surface with brownish hairs. Labium and gnathocoxae reddish. Labium length 0.39, width 0.30. Gnathocoxae length 0.69, width 0.55. Chelicerae reddish brown. Cheliceral groove with four triangular teeth (
<figureCitation id="135D2E8DFFEEF166FF000EABFE92F949" box="[168,285,1669,1694]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="5" pageNumber="402">Figure 1A</figureCitation>
). Promarginal teeth more strongly developed than retromarginal teeth. Promarginal teeth almost same size. Of retromarginal teeth, one in centre of cheliceral groove about twice size of one at base. There is a distance of 0.02 between the promarginal and 0.08 between the retromarginal teeth.
</paragraph>
<paragraph id="8BD93208FFEDF165FF6808FAFEA0FE6A" blockId="6.[160,1156,142,445]" pageId="6" pageNumber="403">
Abdomen greyish. Surface covered with brownish setae. Legs yellowish light brown. Anterior legs slightly darker than posterior legs. Tarsi with three claws. Paired claws toothed. Paired claws of legs I and II with 7 teeth; legs III and IV with 5 teeth. Anterior legs without scopulae. Tarsi three and four and first distal quarter of metatarsi three and four with scopulae. Anterior femora and third and fourth coxae with spine. Leg formula: VI, I, II, III. Leg spination and measurements are given in
<tableCitation id="C6E407B3FFEDF165FC6009AFFBA3FE4D" box="[968,1068,385,410]" captionStart="Table 1" captionStartId="6.[239,295,901,924]" captionText="Table 1. Leg spination of H. bozdaglarensis sp. n. (Holotype ♂/Paratype ♀)." pageId="6" pageNumber="403">Tables 1</tableCitation>
and
<tableCitation id="C6E407B3FFEDF165FBD809AFFBF1FE4D" box="[1136,1150,385,410]" captionStart="Table 2" captionStartId="6.[182,238,1320,1343]" captionText="Table 2. Leg measurements of H. bozdaglarensis sp. n. (Holotype ♂/Paratypes ♂–♀)." pageId="6" pageNumber="403">2</tableCitation>
, respectively.
</paragraph>
<paragraph id="8BD93208FFEDF165FF0809D6FC82FC9D" blockId="6.[160,1156,504,842]" pageId="6" pageNumber="403">
<emphasis id="B912EE1AFFEDF165FF0809D6FE7DFDC5" bold="true" box="[160,498,504,530]" pageId="6" pageNumber="403">
<emphasis id="B912EE1AFFEDF165FF0809D6FF5BFDC5" bold="true" box="[160,212,504,530]" italics="true" pageId="6" pageNumber="403">Palp</emphasis>
. (
<figureCitation id="135D2E8DFFEDF165FF4609D6FE1BFDC5" box="[238,404,504,530]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="6" pageNumber="403">Figures 1BD</figureCitation>
,
<figureCitation id="135D2E8DFFEDF165FE0C09D6FE6BFDC5" box="[420,484,504,530]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="6" pageNumber="403">4AC</figureCitation>
).
</emphasis>
Palpal tarsus cylindrical, length 0.63. Palpal tibia longer than tarsus (
<figureCitation id="135D2E8DFFEDF165FE980A32FE13FDE2" box="[304,412,540,565]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="6" pageNumber="403">Figure 1B</figureCitation>
), length 0.65. Ratio Ti/Ta 0.97. Tegulum (T) cylindrical, brownish; length 0.55, width 0.31. Embolus (E) strongly sclerotised in the proximal half and weakly sclerotised in distal half, with a membranous appearance (
<figureCitation id="135D2E8DFFEDF165FC990A4FFC26FDAD" box="[817,937,609,634]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="6" pageNumber="403">Figures 1C</figureCitation>
,
<figureCitation id="135D2E8DFFEDF165FC1C0A4FFC5DFDAD" box="[948,978,609,634]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="6" pageNumber="403">4A</figureCitation>
). Conductor (C) massive, strongly sclerotised. Embolus and conductor almost same size (
<figureCitation id="135D2E8DFFEDF165FC7B0AAAFBC2FD4A" box="[979,1101,644,669]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="6" pageNumber="403">Figures 1D</figureCitation>
,
<figureCitation id="135D2E8DFFEDF165FBF10AAAFBF9FD4A" box="[1113,1142,644,669]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="6" pageNumber="403">4C</figureCitation>
). Embolus length 0.27, conductor 0.28. Embolus originated from distal centre of tegulum, whereas conductor originated from distal prolateral margin. Embolus flattens and narrows towards tip (
<figureCitation id="135D2E8DFFEDF165FED30AC2FE77FCD2" box="[379,504,748,773]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="6" pageNumber="403">Figures 1C</figureCitation>
,
<figureCitation id="135D2E8DFFEDF165FDA00AC2FDCCFCD2" box="[520,579,748,773]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="6" pageNumber="403">4AC</figureCitation>
). Conductor cylindrical, spiniform with a sharply tapering end. As it distances from distal part of tegulum, tips of embolus and conductor become more distant from each other (
<figureCitation id="135D2E8DFFEDF165FDF60B1FFD57FC9D" box="[606,728,817,842]" captionStart="Figure 1" captionStartId="2.[160,225,1699,1722]" captionTargetBox="[166,1141,147,1654]" captionTargetId="figure-3@2.[163,1153,142,1667]" captionTargetPageId="2" captionText="Figure 1. Harpactea bozdaglarensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, right, prolateral view. (C) Same, horizontally rotated. (D) Male palp, right, retrolateral view. Abbreviations: C conductor, E embolus,T tegulum." pageId="6" pageNumber="403">Figures 1D</figureCitation>
,
<figureCitation id="135D2E8DFFEDF165FD4B0B1FFC8FFC9D" box="[739,768,817,842]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="6" pageNumber="403">4C</figureCitation>
).
</paragraph>
<caption id="DF196280FFEDF165FF470BABFC50FC4B" box="[239,991,901,924]" pageId="6" pageNumber="403" startId="6.[239,295,901,924]" targetType="table">
<paragraph id="8BD93208FFEDF165FF470BABFC50FC4B" blockId="6.[239,991,901,924]" box="[239,991,901,924]" pageId="6" pageNumber="403">
<emphasis id="B912EE1AFFEDF165FF470BABFECFFC4B" bold="true" box="[239,320,901,924]" pageId="6" pageNumber="403">Table 1.</emphasis>
Leg spination of
<taxonomicName id="4C66498BFFEDF165FE470BABFD17FC4B" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[495,664,901,924]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFEDF165FE470BABFD17FC4B" box="[495,664,901,924]" italics="true" pageId="6" pageNumber="403">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEDF165FD360BABFD57FC4B" bold="true" box="[670,728,901,924]" pageId="6" pageNumber="403">
<taxonomicNameLabel id="A2215361FFEDF165FD360BABFD57FC4B" box="[670,728,901,924]" pageId="6" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Holotype ♂/Paratype ♀).
</paragraph>
</caption>
<paragraph id="8BD93208FFEDF165FF510B87FBA2FB33" pageId="6" pageNumber="403">
<table id="F966C0A8FFED0E9CFF510B87FBA2FB33" box="[249,1069,937,1252]" gridcols="5" gridrows="12" pageId="6" pageNumber="403">
<tr id="3556304AFFED0E9CFF510B87FBA2FC6A" box="[249,1069,937,957]" gridrow="0" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510B87FE90FC6A" box="[249,287,937,957]" gridcol="0" gridrow="0" pageId="6" pageNumber="403">Legs</th>
<th id="76875936FFED0E9CFE3C0B87FE46FC6A" box="[404,457,937,957]" gridcol="1" gridrow="0" pageId="6" pageNumber="403">I</th>
<th id="76875936FFED0E9CFD810B87FDFDFC6A" box="[553,626,937,957]" gridcol="2" gridrow="0" pageId="6" pageNumber="403">II</th>
<th id="76875936FFED0E9CFD790B87FCE2FC6A" box="[721,877,937,957]" gridcol="3" gridrow="0" pageId="6" pageNumber="403">III</th>
<th id="76875936FFED0E9CFC650B87FBA2FC6A" box="[973,1069,937,957]" gridcol="4" gridrow="0" pageId="6" pageNumber="403">IV</th>
</tr>
<tr id="3556304AFFED0E9CFF510BE8FBA2FC0B" box="[249,1069,966,988]" gridrow="1" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510BE8FBA2FC0B" box="[249,1069,966,988]" colspan="5" colspanRight="4" gridcol="0" gridrow="1" pageId="6" pageNumber="403"></th>
</tr>
<tr id="3556304AFFED0E9CFF510BCEFBA2FC23" box="[249,1069,992,1012]" gridrow="2" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510BCEFE90FC23" box="[249,287,992,1012]" gridcol="0" gridrow="2" pageId="6" pageNumber="403">Cx</th>
<td id="76875936FFED0E9CFE3C0BCEFE46FC23" box="[404,457,992,1012]" gridcol="1" gridrow="2" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810BCEFDFDFC23" box="[553,626,992,1012]" gridcol="2" gridrow="2" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790BCEFCE2FC23" box="[721,877,992,1012]" gridcol="3" gridrow="2" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFC650BCEFBA2FC23" box="[973,1069,992,1012]" gridcol="4" gridrow="2" pageId="6" pageNumber="403">4 pl</td>
</tr>
<tr id="3556304AFFED0E9CFF510BD6FBA2FBDB" box="[249,1069,1016,1036]" gridrow="3" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510BD6FE90FBDB" box="[249,287,1016,1036]" gridcol="0" gridrow="3" pageId="6" pageNumber="403">Fe</th>
<td id="76875936FFED0E9CFE3C0BD6FE46FBDB" box="[404,457,1016,1036]" gridcol="1" gridrow="3" pageId="6" pageNumber="403">1, 2 pl</td>
<td id="76875936FFED0E9CFD810BD6FDFDFBDB" box="[553,626,1016,1036]" gridcol="2" gridrow="3" pageId="6" pageNumber="403">1, 1, 2 pl</td>
<td id="76875936FFED0E9CFD790BD6FCE2FBDB" box="[721,877,1016,1036]" gridcol="3" gridrow="3" pageId="6" pageNumber="403">8 d (4 pd 4 rd)</td>
<td id="76875936FFED0E9CFC650BD6FBA2FBDB" box="[973,1069,1016,1036]" gridcol="4" gridrow="3" pageId="6" pageNumber="403">8 d</td>
</tr>
<tr id="3556304AFFED0E9CFF510C3EFBA2FBF3" box="[249,1069,1040,1060]" gridrow="4" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510C3EFE90FBF3" box="[249,287,1040,1060]" gridcol="0" gridrow="4" pageId="6" pageNumber="403">Pa</th>
<td id="76875936FFED0E9CFE3C0C3EFE46FBF3" box="[404,457,1040,1060]" gridcol="1" gridrow="4" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810C3EFDFDFBF3" box="[553,626,1040,1060]" gridcol="2" gridrow="4" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790C3EFCE2FBF3" box="[721,877,1040,1060]" gridcol="3" gridrow="4" pageId="6" pageNumber="403">1 rl</td>
<td id="76875936FFED0E9CFC650C3EFBA2FBF3" box="[973,1069,1040,1060]" gridcol="4" gridrow="4" pageId="6" pageNumber="403">1 rl</td>
</tr>
<tr id="3556304AFFED0E9CFF510C06FBA2FBEB" box="[249,1069,1064,1084]" gridrow="5" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510C06FE90FBEB" box="[249,287,1064,1084]" gridcol="0" gridrow="5" pageId="6" pageNumber="403">Ti</th>
<td id="76875936FFED0E9CFE3C0C06FE46FBEB" box="[404,457,1064,1084]" gridcol="1" gridrow="5" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810C06FDFDFBEB" box="[553,626,1064,1084]" gridcol="2" gridrow="5" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790C06FCE2FBEB" box="[721,877,1064,1084]" gridcol="3" gridrow="5" pageId="6" pageNumber="403">3 rl 2 pl 6 v</td>
<td id="76875936FFED0E9CFC650C06FBA2FBEB" box="[973,1069,1064,1084]" gridcol="4" gridrow="5" pageId="6" pageNumber="403">3 rl 3 pl 6 v</td>
</tr>
<tr id="3556304AFFED0E9CFF510C6EFBA2FBBB" box="[249,1069,1088,1132]" gridrow="6" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510C6EFE90FBBB" box="[249,287,1088,1132]" gridcol="0" gridrow="6" pageId="6" pageNumber="403">Me ♀</th>
<td id="76875936FFED0E9CFE3C0C6EFE46FBBB" box="[404,457,1088,1132]" gridcol="1" gridrow="6" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810C6EFDFDFBBB" box="[553,626,1088,1132]" gridcol="2" gridrow="6" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790C6EFCE2FBBB" box="[721,877,1088,1132]" gridcol="3" gridrow="6" pageId="6" pageNumber="403">3 rl 2 pl 4 v</td>
<td id="76875936FFED0E9CFC650C6EFBA2FBBB" box="[973,1069,1088,1132]" gridcol="4" gridrow="6" pageId="6" pageNumber="403">3 rl 4 pl 5 v</td>
</tr>
<tr id="3556304AFFED0E9CFF510C5EFBA2FB53" box="[249,1069,1136,1156]" gridrow="7" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510C5EFE90FB53" box="[249,287,1136,1156]" gridcol="0" gridrow="7" pageId="6" pageNumber="403">Cx</th>
<td id="76875936FFED0E9CFE3C0C5EFE46FB53" box="[404,457,1136,1156]" gridcol="1" gridrow="7" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810C5EFDFDFB53" box="[553,626,1136,1156]" gridcol="2" gridrow="7" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790C5EFCE2FB53" box="[721,877,1136,1156]" gridcol="3" gridrow="7" pageId="6" pageNumber="403">0 (1) pl</td>
<td id="76875936FFED0E9CFC650C5EFBA2FB53" box="[973,1069,1136,1156]" gridcol="4" gridrow="7" pageId="6" pageNumber="403">3 pl</td>
</tr>
<tr id="3556304AFFED0E9CFF510CA6FBA2FB4B" box="[249,1069,1160,1180]" gridrow="8" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510CA6FE90FB4B" box="[249,287,1160,1180]" gridcol="0" gridrow="8" pageId="6" pageNumber="403">Fe</th>
<td id="76875936FFED0E9CFE3C0CA6FE46FB4B" box="[404,457,1160,1180]" gridcol="1" gridrow="8" pageId="6" pageNumber="403">1, 2 pl</td>
<td id="76875936FFED0E9CFD810CA6FDFDFB4B" box="[553,626,1160,1180]" gridcol="2" gridrow="8" pageId="6" pageNumber="403">1, 1, 2 pl</td>
<td id="76875936FFED0E9CFD790CA6FCE2FB4B" box="[721,877,1160,1180]" gridcol="3" gridrow="8" pageId="6" pageNumber="403">7 d (3 pd 1 d 3 rd)</td>
<td id="76875936FFED0E9CFC650CA6FBA2FB4B" box="[973,1069,1160,1180]" gridcol="4" gridrow="8" pageId="6" pageNumber="403">7 d</td>
</tr>
<tr id="3556304AFFED0E9CFF510C8EFBA2FB63" box="[249,1069,1184,1204]" gridrow="9" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510C8EFE90FB63" box="[249,287,1184,1204]" gridcol="0" gridrow="9" pageId="6" pageNumber="403">Pa</th>
<td id="76875936FFED0E9CFE3C0C8EFE46FB63" box="[404,457,1184,1204]" gridcol="1" gridrow="9" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810C8EFDFDFB63" box="[553,626,1184,1204]" gridcol="2" gridrow="9" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790C8EFCE2FB63" box="[721,877,1184,1204]" gridcol="3" gridrow="9" pageId="6" pageNumber="403">1 rl</td>
<td id="76875936FFED0E9CFC650C8EFBA2FB63" box="[973,1069,1184,1204]" gridcol="4" gridrow="9" pageId="6" pageNumber="403">1 rl</td>
</tr>
<tr id="3556304AFFED0E9CFF510C96FBA2FB1B" box="[249,1069,1208,1228]" gridrow="10" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510C96FE90FB1B" box="[249,287,1208,1228]" gridcol="0" gridrow="10" pageId="6" pageNumber="403">Ti</th>
<td id="76875936FFED0E9CFE3C0C96FE46FB1B" box="[404,457,1208,1228]" gridcol="1" gridrow="10" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810C96FDFDFB1B" box="[553,626,1208,1228]" gridcol="2" gridrow="10" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790C96FCE2FB1B" box="[721,877,1208,1228]" gridcol="3" gridrow="10" pageId="6" pageNumber="403">3 rl 2 pl 6 v</td>
<td id="76875936FFED0E9CFC650C96FBA2FB1B" box="[973,1069,1208,1228]" gridcol="4" gridrow="10" pageId="6" pageNumber="403">3 rl 3 pl 6 v</td>
</tr>
<tr id="3556304AFFED0E9CFF510CFEFBA2FB33" box="[249,1069,1232,1252]" gridrow="11" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF510CFEFE90FB33" box="[249,287,1232,1252]" gridcol="0" gridrow="11" pageId="6" pageNumber="403">Me</th>
<td id="76875936FFED0E9CFE3C0CFEFE46FB33" box="[404,457,1232,1252]" gridcol="1" gridrow="11" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD810CFEFDFDFB33" box="[553,626,1232,1252]" gridcol="2" gridrow="11" pageId="6" pageNumber="403">0</td>
<td id="76875936FFED0E9CFD790CFEFCE2FB33" box="[721,877,1232,1252]" gridcol="3" gridrow="11" pageId="6" pageNumber="403">3 rl 2 pl 5 v</td>
<td id="76875936FFED0E9CFC650CFEFBA2FB33" box="[973,1069,1232,1252]" gridcol="4" gridrow="11" pageId="6" pageNumber="403">3 rl 4 pl 6 v</td>
</tr>
</table>
</paragraph>
<caption id="DF196280FFEDF165FF1E0D06FB8AFAE8" box="[182,1029,1320,1343]" pageId="6" pageNumber="403" startId="6.[182,238,1320,1343]" targetType="table">
<paragraph id="8BD93208FFEDF165FF1E0D06FB8AFAE8" blockId="6.[182,1029,1320,1343]" box="[182,1029,1320,1343]" pageId="6" pageNumber="403">
<emphasis id="B912EE1AFFEDF165FF1E0D06FE88FAE8" bold="true" box="[182,263,1320,1343]" pageId="6" pageNumber="403">Table 2.</emphasis>
Leg measurements of
<taxonomicName id="4C66498BFFEDF165FE440D06FD1AFAE8" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[492,661,1320,1343]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="403" phylum="Arthropoda" rank="species" species="bozdaglarensis" status="sp. n.">
<emphasis id="B912EE1AFFEDF165FE440D06FD1AFAE8" box="[492,661,1320,1343]" italics="true" pageId="6" pageNumber="403">H. bozdaglarensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFEDF165FD330D06FD5AFAE8" bold="true" box="[667,725,1320,1343]" pageId="6" pageNumber="403">
<taxonomicNameLabel id="A2215361FFEDF165FD330D06FD5AFAE8" box="[667,725,1320,1343]" pageId="6" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Holotype ♂/Paratypes ♂–♀).
</paragraph>
</caption>
<paragraph id="8BD93208FFEDF165FF680D62FBECF960" pageId="6" pageNumber="403">
<table id="F966C0A8FFED0E9CFF680D62FBECF960" box="[192,1123,1356,1719]" gridcols="5" gridrows="14" pageId="6" pageNumber="403">
<tr id="3556304AFFED0E9CFF680D62FBECFAB7" box="[192,1123,1356,1376]" gridrow="0" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680D62FF7DFAB7" box="[192,242,1356,1376]" gridcol="0" gridrow="0" pageId="6" pageNumber="403">Legs</th>
<th id="76875936FFED0E9CFEF40D62FE59FAB7" box="[348,470,1356,1376]" gridcol="1" gridrow="0" pageId="6" pageNumber="403">I</th>
<th id="76875936FFED0E9CFD9D0D62FD20FAB7" box="[565,687,1356,1376]" gridcol="2" gridrow="0" pageId="6" pageNumber="403">II</th>
<th id="76875936FFED0E9CFCA70D62FC06FAB7" box="[783,905,1356,1376]" gridcol="3" gridrow="0" pageId="6" pageNumber="403">III</th>
<th id="76875936FFED0E9CFC410D62FBECFAB7" box="[1001,1123,1356,1376]" gridcol="4" gridrow="0" pageId="6" pageNumber="403">IV</th>
</tr>
<tr id="3556304AFFED0E9CFF680D47FBECFAA8" box="[192,1123,1385,1407]" gridrow="1" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680D47FBECFAA8" box="[192,1123,1385,1407]" colspan="5" colspanRight="4" gridcol="0" gridrow="1" pageId="6" pageNumber="403"></th>
</tr>
<tr id="3556304AFFED0E9CFF680DADFBECFA40" box="[192,1123,1411,1431]" gridrow="2" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680DADFF7DFA40" box="[192,242,1411,1431]" gridcol="0" gridrow="2" pageId="6" pageNumber="403">Fe</th>
<td id="76875936FFED0E9CFEF40DADFE59FA40" box="[348,470,1411,1431]" gridcol="1" gridrow="2" pageId="6" pageNumber="403">2.81</td>
<td id="76875936FFED0E9CFD9D0DADFD20FA40" box="[565,687,1411,1431]" gridcol="2" gridrow="2" pageId="6" pageNumber="403">2.46</td>
<td id="76875936FFED0E9CFCA70DADFC06FA40" box="[783,905,1411,1431]" gridcol="3" gridrow="2" pageId="6" pageNumber="403">2.06</td>
<td id="76875936FFED0E9CFC410DADFBECFA40" box="[1001,1123,1411,1431]" gridcol="4" gridrow="2" pageId="6" pageNumber="403">2.88</td>
</tr>
<tr id="3556304AFFED0E9CFF680DB5FBECFA78" box="[192,1123,1435,1455]" gridrow="3" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680DB5FF7DFA78" box="[192,242,1435,1455]" gridcol="0" gridrow="3" pageId="6" pageNumber="403">Pa</th>
<td id="76875936FFED0E9CFEF40DB5FE59FA78" box="[348,470,1435,1455]" gridcol="1" gridrow="3" pageId="6" pageNumber="403">1.65</td>
<td id="76875936FFED0E9CFD9D0DB5FD20FA78" box="[565,687,1435,1455]" gridcol="2" gridrow="3" pageId="6" pageNumber="403">1.54</td>
<td id="76875936FFED0E9CFCA70DB5FC06FA78" box="[783,905,1435,1455]" gridcol="3" gridrow="3" pageId="6" pageNumber="403">0.94</td>
<td id="76875936FFED0E9CFC410DB5FBECFA78" box="[1001,1123,1435,1455]" gridcol="4" gridrow="3" pageId="6" pageNumber="403">1.07</td>
</tr>
<tr id="3556304AFFED0E9CFF680D9DFBECFA10" box="[192,1123,1459,1479]" gridrow="4" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680D9DFF7DFA10" box="[192,242,1459,1479]" gridcol="0" gridrow="4" pageId="6" pageNumber="403">Ti</th>
<td id="76875936FFED0E9CFEF40D9DFE59FA10" box="[348,470,1459,1479]" gridcol="1" gridrow="4" pageId="6" pageNumber="403">2.15</td>
<td id="76875936FFED0E9CFD9D0D9DFD20FA10" box="[565,687,1459,1479]" gridcol="2" gridrow="4" pageId="6" pageNumber="403">2.20</td>
<td id="76875936FFED0E9CFCA70D9DFC06FA10" box="[783,905,1459,1479]" gridcol="3" gridrow="4" pageId="6" pageNumber="403">1.69</td>
<td id="76875936FFED0E9CFC410D9DFBECFA10" box="[1001,1123,1459,1479]" gridcol="4" gridrow="4" pageId="6" pageNumber="403">2.40</td>
</tr>
<tr id="3556304AFFED0E9CFF680DE5FBECFA08" box="[192,1123,1483,1503]" gridrow="5" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680DE5FF7DFA08" box="[192,242,1483,1503]" gridcol="0" gridrow="5" pageId="6" pageNumber="403">Me</th>
<td id="76875936FFED0E9CFEF40DE5FE59FA08" box="[348,470,1483,1503]" gridcol="1" gridrow="5" pageId="6" pageNumber="403">2.34</td>
<td id="76875936FFED0E9CFD9D0DE5FD20FA08" box="[565,687,1483,1503]" gridcol="2" gridrow="5" pageId="6" pageNumber="403">2.08</td>
<td id="76875936FFED0E9CFCA70DE5FC06FA08" box="[783,905,1483,1503]" gridcol="3" gridrow="5" pageId="6" pageNumber="403">2.04</td>
<td id="76875936FFED0E9CFC410DE5FBECFA08" box="[1001,1123,1483,1503]" gridcol="4" gridrow="5" pageId="6" pageNumber="403">2.59</td>
</tr>
<tr id="3556304AFFED0E9CFF680DCDFBECFA20" box="[192,1123,1507,1527]" gridrow="6" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680DCDFF7DFA20" box="[192,242,1507,1527]" gridcol="0" gridrow="6" pageId="6" pageNumber="403">Ta</th>
<td id="76875936FFED0E9CFEF40DCDFE59FA20" box="[348,470,1507,1527]" gridcol="1" gridrow="6" pageId="6" pageNumber="403">0.61</td>
<td id="76875936FFED0E9CFD9D0DCDFD20FA20" box="[565,687,1507,1527]" gridcol="2" gridrow="6" pageId="6" pageNumber="403">0.60</td>
<td id="76875936FFED0E9CFCA70DCDFC06FA20" box="[783,905,1507,1527]" gridcol="3" gridrow="6" pageId="6" pageNumber="403">0.60</td>
<td id="76875936FFED0E9CFC410DCDFBECFA20" box="[1001,1123,1507,1527]" gridcol="4" gridrow="6" pageId="6" pageNumber="403">0.63</td>
</tr>
<tr id="3556304AFFED0E9CFF680DD5FBECF9F0" box="[192,1123,1531,1575]" gridrow="7" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680DD5FF7DF9F0" box="[192,242,1531,1575]" gridcol="0" gridrow="7" pageId="6" pageNumber="403">Total ♂/♀</th>
<td id="76875936FFED0E9CFEF40DD5FE59F9F0" box="[348,470,1531,1575]" gridcol="1" gridrow="7" pageId="6" pageNumber="403">9.56</td>
<td id="76875936FFED0E9CFD9D0DD5FD20F9F0" box="[565,687,1531,1575]" gridcol="2" gridrow="7" pageId="6" pageNumber="403">8.88</td>
<td id="76875936FFED0E9CFCA70DD5FC06F9F0" box="[783,905,1531,1575]" gridcol="3" gridrow="7" pageId="6" pageNumber="403">7.33</td>
<td id="76875936FFED0E9CFC410DD5FBECF9F0" box="[1001,1123,1531,1575]" gridcol="4" gridrow="7" pageId="6" pageNumber="403">9.57</td>
</tr>
<tr id="3556304AFFED0E9CFF680E05FBECF9E8" box="[192,1123,1579,1599]" gridrow="8" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680E05FF7DF9E8" box="[192,242,1579,1599]" gridcol="0" gridrow="8" pageId="6" pageNumber="403">Fe</th>
<td id="76875936FFED0E9CFEF40E05FE59F9E8" box="[348,470,1579,1599]" gridcol="1" gridrow="8" pageId="6" pageNumber="403">2.49/2.302.85</td>
<td id="76875936FFED0E9CFD9D0E05FD20F9E8" box="[565,687,1579,1599]" gridcol="2" gridrow="8" pageId="6" pageNumber="403">2.45/2.252.40</td>
<td id="76875936FFED0E9CFCA70E05FC06F9E8" box="[783,905,1579,1599]" gridcol="3" gridrow="8" pageId="6" pageNumber="403">1.71/1.722.00</td>
<td id="76875936FFED0E9CFC410E05FBECF9E8" box="[1001,1123,1579,1599]" gridcol="4" gridrow="8" pageId="6" pageNumber="403">2.61/2.652.75</td>
</tr>
<tr id="3556304AFFED0E9CFF680E6DFBECF980" box="[192,1123,1603,1623]" gridrow="9" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680E6DFF7DF980" box="[192,242,1603,1623]" gridcol="0" gridrow="9" pageId="6" pageNumber="403">Pa</th>
<td id="76875936FFED0E9CFEF40E6DFE59F980" box="[348,470,1603,1623]" gridcol="1" gridrow="9" pageId="6" pageNumber="403">1.68/1.501.75</td>
<td id="76875936FFED0E9CFD9D0E6DFD20F980" box="[565,687,1603,1623]" gridcol="2" gridrow="9" pageId="6" pageNumber="403">1.53/1.401.50</td>
<td id="76875936FFED0E9CFCA70E6DFC06F980" box="[783,905,1603,1623]" gridcol="3" gridrow="9" pageId="6" pageNumber="403">0.82/0.801.10</td>
<td id="76875936FFED0E9CFC410E6DFBECF980" box="[1001,1123,1603,1623]" gridcol="4" gridrow="9" pageId="6" pageNumber="403">1.21/1.151.25</td>
</tr>
<tr id="3556304AFFED0E9CFF680E75FBECF9B8" box="[192,1123,1627,1647]" gridrow="10" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680E75FF7DF9B8" box="[192,242,1627,1647]" gridcol="0" gridrow="10" pageId="6" pageNumber="403">Ti</th>
<td id="76875936FFED0E9CFEF40E75FE59F9B8" box="[348,470,1627,1647]" gridcol="1" gridrow="10" pageId="6" pageNumber="403">2.33/2.152.30</td>
<td id="76875936FFED0E9CFD9D0E75FD20F9B8" box="[565,687,1627,1647]" gridcol="2" gridrow="10" pageId="6" pageNumber="403">2.21/2.002.20</td>
<td id="76875936FFED0E9CFCA70E75FC06F9B8" box="[783,905,1627,1647]" gridcol="3" gridrow="10" pageId="6" pageNumber="403">1.40/1.161.50</td>
<td id="76875936FFED0E9CFC410E75FBECF9B8" box="[1001,1123,1627,1647]" gridcol="4" gridrow="10" pageId="6" pageNumber="403">2.21/2.402.50</td>
</tr>
<tr id="3556304AFFED0E9CFF680E5DFBECF950" box="[192,1123,1651,1671]" gridrow="11" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680E5DFF7DF950" box="[192,242,1651,1671]" gridcol="0" gridrow="11" pageId="6" pageNumber="403">Me</th>
<td id="76875936FFED0E9CFEF40E5DFE59F950" box="[348,470,1651,1671]" gridcol="1" gridrow="11" pageId="6" pageNumber="403">2.03/1.952.05</td>
<td id="76875936FFED0E9CFD9D0E5DFD20F950" box="[565,687,1651,1671]" gridcol="2" gridrow="11" pageId="6" pageNumber="403">2.07/1.902.20</td>
<td id="76875936FFED0E9CFCA70E5DFC06F950" box="[783,905,1651,1671]" gridcol="3" gridrow="11" pageId="6" pageNumber="403">1.85/1.682.00</td>
<td id="76875936FFED0E9CFC410E5DFBECF950" box="[1001,1123,1651,1671]" gridcol="4" gridrow="11" pageId="6" pageNumber="403">2.58/2.602.75</td>
</tr>
<tr id="3556304AFFED0E9CFF680EA5FBECF948" box="[192,1123,1675,1695]" gridrow="12" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680EA5FF7DF948" box="[192,242,1675,1695]" gridcol="0" gridrow="12" pageId="6" pageNumber="403">Ta</th>
<td id="76875936FFED0E9CFEF40EA5FE59F948" box="[348,470,1675,1695]" gridcol="1" gridrow="12" pageId="6" pageNumber="403">0.52/0.500.55</td>
<td id="76875936FFED0E9CFD9D0EA5FD20F948" box="[565,687,1675,1695]" gridcol="2" gridrow="12" pageId="6" pageNumber="403">0.60/0.550.60</td>
<td id="76875936FFED0E9CFCA70EA5FC06F948" box="[783,905,1675,1695]" gridcol="3" gridrow="12" pageId="6" pageNumber="403">0.57/0.500.55</td>
<td id="76875936FFED0E9CFC410EA5FBECF948" box="[1001,1123,1675,1695]" gridcol="4" gridrow="12" pageId="6" pageNumber="403">0.66/0.600.60</td>
</tr>
<tr id="3556304AFFED0E9CFF680E8DFBECF960" box="[192,1123,1699,1719]" gridrow="13" pageId="6" pageNumber="403">
<th id="76875936FFED0E9CFF680E8DFF7DF960" box="[192,242,1699,1719]" gridcol="0" gridrow="13" pageId="6" pageNumber="403">Total</th>
<td id="76875936FFED0E9CFEF40E8DFE59F960" box="[348,470,1699,1719]" gridcol="1" gridrow="13" pageId="6" pageNumber="403">9.05/8.409.50</td>
<td id="76875936FFED0E9CFD9D0E8DFD20F960" box="[565,687,1699,1719]" gridcol="2" gridrow="13" pageId="6" pageNumber="403">8.86/8.108.90</td>
<td id="76875936FFED0E9CFCA70E8DFC06F960" box="[783,905,1699,1719]" gridcol="3" gridrow="13" pageId="6" pageNumber="403">6.36/5.867.15</td>
<td id="76875936FFED0E9CFC410E8DFBECF960" box="[1001,1123,1699,1719]" gridcol="4" gridrow="13" pageId="6" pageNumber="403">9.27/9.409.85</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD93208FFECF164FF6808A0FE63FF3A" blockId="7.[160,1156,142,237]" pageId="7" pageNumber="404">Although there are size differences between the somatic parts of males and females, they are generally similar. Palpal tarsus cylindrical, length 0.750.90. Palpal tibia shorter than tarsus, length 0.600.65.</paragraph>
<paragraph id="8BD93208FFECF164FF080937FD1CFE37" blockId="7.[160,1156,281,723]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080937FEE1FEE4" bold="true" box="[160,366,281,307]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080937FF6CFEE4" bold="true" box="[160,227,281,307]" italics="true" pageId="7" pageNumber="404">Vulva</emphasis>
. (
<figureCitation id="135D2E8DFFECF164FF520937FED0FEE4" box="[250,351,281,307]" captionStart="Figure 2" captionStartId="3.[160,225,1128,1151]" captionTargetBox="[163,1153,142,1096]" captionTargetId="figure-131@3.[163,1153,142,1096]" captionTargetPageId="3" captionText="Figure 2. Harpactea bozdaglarensis sp. n. (AC) Vulva, dorsal view from different angles; in Figure 2C, the outlines of the posterior diverticulum are drawn and highlighted for visibility. (D) Same, ventral view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, PD posterior diverticulum, RS roundish structures, S spermatheca, SB sclerotised belt, SK spermathecal keel, TB transversal bar." pageId="7" pageNumber="404">Figure 2</figureCitation>
).
</emphasis>
Anterior arc (AC) wider than long. Spermathecal keel (SK) concave in some individuals and relatively straight in others. In all cases, lateral edges pointed (
<figureCitation id="135D2E8DFFECF164FF000971FEB8FEAF" box="[168,311,351,376]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4AD</figureCitation>
). Approximate middle part of spermatheca (S) slightly recurved to sides (
<figureCitation id="135D2E8DFFECF164FF0009ACFEB9FE4C" box="[168,310,386,411]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4AD</figureCitation>
). Anterior part of this region longer than part posterior to it. In addition, posterior part sclerotised at same level as anterior margin of the anterior arc (AA) and more strongly sclerotised than anterior part.
</paragraph>
<paragraph id="8BD93208FFECF164FF6809C4FB0CFD04" blockId="7.[160,1156,281,723]" pageId="7" pageNumber="404">
Lateral margins of anterior margin of the anterior arc are widened laterally (
<figureCitation id="135D2E8DFFECF164FBA409C4FBF9FDD4" box="[1036,1142,490,515]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4B</figureCitation>
). These parts more weakly sclerotised than central regions. Round structures (
<collectingRegion id="49A2FCEAFFECF164FB870A22FBC0FDF2" box="[1071,1103,524,549]" country="Bulgaria" name="Ruse" pageId="7" pageNumber="404">RS</collectingRegion>
) are symmetrical and distinct as conical tubercles (
<figureCitation id="135D2E8DFFECF164FD0F0A01FCA0FD9F" box="[679,815,559,584]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4AB</figureCitation>
). Width of transversal bar (TB) approximately equal to anterior margin of the anterior arc (
<figureCitation id="135D2E8DFFECF164FCEE0A7CFC3DFDBC" box="[838,946,594,619]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4C</figureCitation>
). Posterior diverticulum membranous, with indistinct borders, attached to the transversal bar (
<figureCitation id="135D2E8DFFECF164FBA20A5AFBF9FD5A" box="[1034,1142,628,653]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4C</figureCitation>
). Between anterior margin of the anterior arc and transversal bar there is a sclerotised belt whose width is approximately twice that of anterior margin of anterior arc (
<figureCitation id="135D2E8DFFECF164FC460A94FBF9FD04" box="[1006,1142,698,723]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">Figure 4AC</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFECF164FF080AD1FCD8FBA3" pageId="7" pageNumber="404" type="distribution">
<paragraph id="8BD93208FFECF164FF080AD1FEA3FCCE" blockId="7.[160,524,767,828]" box="[160,300,767,793]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080AD1FEA3FCCE" bold="true" box="[160,300,767,793]" italics="true" pageId="7" pageNumber="404">Distribution</emphasis>
</paragraph>
<paragraph id="8BD93208FFECF164FF080B0DFD83FCEB" blockId="7.[160,524,767,828]" box="[160,524,803,828]" pageId="7" pageNumber="404">
Known only from
<typeStatus id="54DD8CAAFFECF164FEC20B0DFE13FCEB" box="[362,412,803,828]" pageId="7" pageNumber="404">type</typeStatus>
localities.
</paragraph>
<paragraph id="8BD93208FFECF164FF080B46FF76FC55" blockId="7.[160,1156,872,1140]" box="[160,249,872,898]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080B46FF76FC55" bold="true" box="[160,249,872,898]" italics="true" pageId="7" pageNumber="404">Habitat</emphasis>
</paragraph>
<paragraph id="8BD93208FFECF164FF080BA2FCD8FBA3" blockId="7.[160,1156,872,1140]" pageId="7" pageNumber="404">
The collection site, situated
<quantity id="4C9E9FEDFFECF164FE7D0BA2FD83FC72" box="[469,524,908,933]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="404" unit="km" value="2.0">2 km</quantity>
south of Allahdiyen Village, comprises dense oak bushes and scattered red pine trees. This habitat features a thick layer of leaf litter and is a serene and humid area located on the outskirts of a large valley. The MSS traps were positioned among accumulated, fragmented stone pieces in a small, cool, and humid valley. The area also encompasses plane trees and accumulated stones blanketed with leaf litter composed of dried plane leaves. Mixed red pine trees and oak bushes distinguish the Kuşlar site. It is a cool and humid environment with dense leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,580 @@
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<mods:title id="C1D0ACD1B2998A4BE8CEE6AD22B458AA">Two new species of Harpactea Bristowe, 1939 from Turkey (Araneae: Dysderidae)</mods:title>
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<mods:namePart id="EB48ECBC548E364562F42A29DA08B678">Kunt, Kadir BoğaÇ</mods:namePart>
<mods:affiliation id="1764FFA6F66D5D31C2B176D00B121631">Graduate School of Natural and Applied Sciences, Department of Biology, Kırıkkale University, Yahşihan, Turkey; &amp; Cyprus Wildlife Research Institute, Taşkent, Cyprus;</mods:affiliation>
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<mods:namePart id="861315B80035D374E0CCA62C495C62BB">Danışman, Tarık</mods:namePart>
<mods:affiliation id="CAA04F6706B09AB1577B60100025B117">Faculty of Engineering and Natural Sciences, Department of Biology, Kırıkkale University, Yahşihan, Turkey;</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="4E9FBD19857459B1AF95446F7A007F0E">Yağmur, Ersen Aydın</mods:namePart>
<mods:affiliation id="8C4B994A06B569C5C853F07491C1DB45">Alaşehir Vocational School, Manisa Celal Bayar University, Alaşehir, Turkey</mods:affiliation>
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<mods:date id="7A4A09B30412DD5FB940D546109083BC">2025</mods:date>
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<mods:number id="66ACBE24E390627DA1E570DB8BE21F50">5 - 8</mods:number>
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<treatment id="03CF831EFFECF168FE9B0C9DFE37FDB0" LSID="urn:lsid:plazi:treatment:03CF831EFFECF168FE9B0C9DFE37FDB0" httpUri="http://treatment.plazi.org/id/03CF831EFFECF168FE9B0C9DFE37FDB0" lastPageId="11" lastPageNumber="408" pageId="7" pageNumber="404">
<subSubSection id="C37C6183FFECF164FE9B0C9DFC7EFB1A" box="[307,1009,1203,1229]" pageId="7" pageNumber="404" type="nomenclature">
<paragraph id="8BD93208FFECF164FE9B0C9DFC7EFB1A" blockId="7.[307,1009,1203,1258]" box="[307,1009,1203,1229]" pageId="7" pageNumber="404">
<heading id="D0918564FFECF164FE9B0C9DFC7EFB1A" bold="true" box="[307,1009,1203,1229]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="404" reason="3">
<emphasis id="B912EE1AFFECF164FE9B0C9DFC7EFB1A" bold="true" box="[307,1009,1203,1229]" pageId="7" pageNumber="404">
<taxonomicName id="4C66498BFFECF164FE9B0C9DFC2AFB1A" authority="Kunt, Danisman and Yagmur" authorityName="Kunt, Danisman and Yagmur" authorityYear="2025" box="[307,933,1203,1229]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. nov.">
<emphasis id="B912EE1AFFECF164FE9B0C9DFDBDFB1A" bold="true" box="[307,562,1203,1229]" italics="true" pageId="7" pageNumber="404">Harpactea dilekensis</emphasis>
Kunt, Danışman and Yağmur
</taxonomicName>
<taxonomicNameLabel id="A2215361FFECF164FC050C9DFC7EFB1A" box="[941,1009,1203,1229]" pageId="7" pageNumber="404" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFECF164FD890CFEFC8CFB3D" box="[545,771,1232,1258]" pageId="7" pageNumber="404" type="description">
<paragraph id="8BD93208FFECF164FD890CFEFC8CFB3D" blockId="7.[307,1009,1203,1258]" box="[545,771,1232,1258]" pageId="7" pageNumber="404">
(
<figureCitation id="135D2E8DFFECF164FD810CFEFD1BFB3D" box="[553,660,1232,1258]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="7" pageNumber="404">Figures 3</figureCitation>
,
<figureCitation id="135D2E8DFFECF164FD0A0CFEFD50FB3D" box="[674,735,1232,1258]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="7" pageNumber="404">4DE</figureCitation>
,
<figureCitation id="135D2E8DFFECF164FD440CFEFD75FB3D" box="[748,762,1232,1258]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="7" pageNumber="404">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFECF164FF080D09FEA5FA1B" pageId="7" pageNumber="404" type="materials_examined">
<paragraph id="8BD93208FFECF164FF080D09FEF0FA96" blockId="7.[160,1156,1319,1484]" box="[160,383,1319,1345]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080D09FEF0FA96" bold="true" box="[160,383,1319,1345]" italics="true" pageId="7" pageNumber="404">Material examined</emphasis>
</paragraph>
<paragraph id="8BD93208FFECF164FF080D65FEA5FA1B" blockId="7.[160,1156,1319,1484]" pageId="7" pageNumber="404">
<materialsCitation id="3B0E3855FFECF164FF080D65FED2FA7E" collectingDate="2017-05-12" collectionCode="KUAM" collectorName="Dilek Peninsula NP &amp; Litter &amp; Leg. E. A. Yagmur" country="Turkey" elevation="163" latitude="37.690556" location="South of GuzelCamli Village" longLatPrecision="1" longitude="-27.237223" pageId="7" pageNumber="404" specimenCount="1" specimenCount-male="1" stateProvince="Aydin" typeStatus="holotype">
<emphasis id="B912EE1AFFECF164FF080D65FEF3FAB3" bold="true" box="[160,380,1355,1380]" pageId="7" pageNumber="404">
<typeStatus id="54DD8CAAFFECF164FF080D65FE9EFAB3" box="[160,273,1355,1380]" pageId="7" pageNumber="404" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED77AACDFFECF164FE8E0D65FEFAFAB3" box="[294,373,1355,1380]" pageId="7" pageNumber="404">KUAM</collectionCode>
)
</emphasis>
<specimenCount id="9D60F981FFECF164FE210D65FE12FAB5" box="[393,413,1355,1378]" count="1" pageId="7" pageNumber="404" type="male"></specimenCount>
<collectingCountry id="F3717298FFECF164FE020D65FD89FAB3" box="[426,518,1355,1380]" name="Turkey" pageId="7" pageNumber="404">TURKEY</collectingCountry>
:
<collectingRegion id="49A2FCEAFFECF164FDBD0D65FD43FAB3" box="[533,716,1355,1380]" country="Turkey" name="Aydin" pageId="7" pageNumber="404">Aydın Province</collectingRegion>
,
<collectorName id="269357DEFFECF164FD740D65FC4EFAB3" box="[732,961,1355,1380]" pageId="7" pageNumber="404">Dilek Peninsula NP</collectorName>
,
<quantity id="4C9E9FEDFFECF164FC790D65FB80FAB3" box="[977,1039,1355,1380]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="404" unit="km" value="4.0">4 km</quantity>
<location id="8EB964D3FFECF164FBB40D65FEE1FA50" LSID="urn:lsid:plazi:treatment:03CF831EFFECF168FE9B0C9DFE37FDB0:8EB964D3FFECF164FBB40D65FEE1FA50" country="Turkey" latitude="37.690556" longLatPrecision="1" longitude="-27.237223" name="South of GuzelCamli Village" pageId="7" pageNumber="404" stateProvince="Aydin">South of GüzelÇamlı Village</location>
(
<geoCoordinate id="EE5254CFFFECF164FED40D40FD82FA50" box="[380,525,1390,1415]" degrees="37.690556" direction="north" orientation="latitude" pageId="7" pageNumber="404" precision="1" value="37.690556">37.690556°N</geoCoordinate>
,
<geoCoordinate id="EE5254CFFFECF164FDB10D40FD20FA50" box="[537,687,1390,1415]" degrees="27.237222" direction="west" orientation="longitude" pageId="7" pageNumber="404" precision="1" value="-27.237223">27.237222°W</geoCoordinate>
), c.
<elevation id="004BD53BFFECF164FD720D40FCC5FA50" box="[730,842,1390,1415]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="404" unit="m" value="163.0">
<quantity id="4C9E9FEDFFECF164FD720D40FCAEFA50" box="[730,801,1390,1415]" metricMagnitude="2" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="404" unit="m" value="163.0">163 m</quantity>
asl
</elevation>
,
<date id="FFD814C8FFECF164FCFE0D40FC65FA50" box="[854,1002,1390,1415]" pageId="7" pageNumber="404" value="2017-05-12">
<collectingDate id="EF9CED20FFECF164FCFE0D40FC65FA50" box="[854,1002,1390,1415]" pageId="7" pageNumber="404" value="2017-05-12">12 May 2017</collectingDate>
</date>
,
<collectorName id="269357DEFFECF164FC5D0D40FBBFFA50" box="[1013,1072,1390,1415]" pageId="7" pageNumber="404">Litter</collectorName>
sifting,
<collectorName id="269357DEFFECF164FF080DBEFED2FA7E" box="[160,349,1424,1449]" pageId="7" pageNumber="404">Leg. E.A. Yağmur</collectorName>
</materialsCitation>
.
<emphasis id="B912EE1AFFECF164FECD0DBEFE53FA7E" bold="true" box="[357,476,1424,1449]" pageId="7" pageNumber="404">
<typeStatus id="54DD8CAAFFECF164FECD0DBEFE53FA7E" box="[357,476,1424,1449]" pageId="7" pageNumber="404" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFECF164FE4A0DBEFD86FA70" box="[482,521,1424,1449]" count="3" pageId="7" pageNumber="404" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D60F981FFECF164FDBD0DBEFDB3FA70" box="[533,572,1424,1449]" count="2" pageId="7" pageNumber="404" type="female">2 ♀</specimenCount>
(
<collectionCode id="ED77AACDFFECF164FDE00DBEFD1CFA7E" box="[584,659,1424,1449]" pageId="7" pageNumber="404">KUAM</collectionCode>
),
<specimenCount id="9D60F981FFECF164FD0A0DBEFD46FA70" box="[674,713,1424,1449]" count="1" pageId="7" pageNumber="404" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D60F981FFECF164FD7C0DBEFD74FA70" box="[724,763,1424,1449]" count="1" pageId="7" pageNumber="404" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED77AACDFFECF164FCAF0DBEFCD8FA7E" box="[775,855,1424,1449]" pageId="7" pageNumber="404">AZMM</collectionCode>
),
<specimenCount id="9D60F981FFECF164FCCD0DBEFC02FA70" box="[869,909,1424,1449]" count="1" pageId="7" pageNumber="404" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D60F981FFECF164FC300DBEFC30FA70" box="[920,959,1424,1449]" count="1" pageId="7" pageNumber="404" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED77AACDFFECF164FC630DBEFB8DFA7E" box="[971,1026,1424,1449]" pageId="7" pageNumber="404">HZH</collectionCode>
), same data as
<typeStatus id="54DD8CAAFFECF164FF170D9DFEA9FA1B" box="[191,294,1459,1484]" pageId="7" pageNumber="404" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFECF164FF080DD6FBB2F9E2" box="[160,1085,1528,1589]" pageId="7" pageNumber="404" type="etymology">
<paragraph id="8BD93208FFECF164FF080DD6FE92F9C5" blockId="7.[160,1085,1528,1589]" box="[160,285,1528,1554]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080DD6FE92F9C5" bold="true" box="[160,285,1528,1554]" italics="true" pageId="7" pageNumber="404">Etymology</emphasis>
</paragraph>
<paragraph id="8BD93208FFECF164FF080E32FBB2F9E2" blockId="7.[160,1085,1528,1589]" box="[160,1085,1564,1589]" pageId="7" pageNumber="404">
The specific epithet is a toponym for the Dilek Peninsula,
<collectingRegion id="49A2FCEAFFECF164FC870E32FC54F9E2" box="[815,987,1564,1589]" country="Turkey" name="Aydin" pageId="7" pageNumber="404">Aydın Province</collectingRegion>
, Turkey.
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFECF16AFF080E4FFDEAFE82" lastPageId="9" lastPageNumber="406" pageId="7" pageNumber="404" type="diagnosis">
<paragraph id="8BD93208FFECF164FF080E4FFE9BF9AC" blockId="7.[160,1156,1633,1728]" box="[160,276,1633,1659]" pageId="7" pageNumber="404">
<emphasis id="B912EE1AFFECF164FF080E4FFE9BF9AC" bold="true" box="[160,276,1633,1659]" italics="true" pageId="7" pageNumber="404">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD93208FFECF16BFF080EABFCE4F96E" blockId="7.[160,1156,1633,1728]" lastBlockId="8.[160,1156,1592,1721]" lastPageId="8" lastPageNumber="405" pageId="7" pageNumber="404">
The male copulatory organ of
<taxonomicName id="4C66498BFFECF164FDBD0EABFD2DF949" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[533,674,1669,1694]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFECF164FDBD0EABFD2DF949" box="[533,674,1669,1694]" italics="true" pageId="7" pageNumber="404">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFECF164FD060EABFD7BF949" bold="true" box="[686,756,1669,1694]" pageId="7" pageNumber="404">
<taxonomicNameLabel id="A2215361FFECF164FD060EABFD7BF949" box="[686,756,1669,1694]" pageId="7" pageNumber="404" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles that of
<taxonomicName id="4C66498BFFECF164FC770EABFEC7F917" authority="Bosmans, 2023" authorityName="Bosmans" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="chaniaensis">
<emphasis id="B912EE1AFFECF164FC770EABFB0BF949" box="[991,1156,1669,1694]" italics="true" pageId="7" pageNumber="404">H. chaniaensis</emphasis>
Bosmans, 2023
</taxonomicName>
(
<collectingRegion id="49A2FCEAFFECF164FEFD0E89FE1BF917" box="[341,404,1703,1728]" country="Greece" name="Kriti" pageId="7" pageNumber="404">Crete</collectingRegion>
) and
<taxonomicName id="4C66498BFFECF164FE7C0E89FCB6F917" authority="Bosmans, 2023" authorityName="Bosmans" authorityYear="2023" box="[468,825,1703,1728]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="404" phylum="Arthropoda" rank="species" species="walterdebucki">
<emphasis id="B912EE1AFFECF164FE7C0E89FD06F917" box="[468,649,1703,1728]" italics="true" pageId="7" pageNumber="404">H. walterdebucki</emphasis>
Bosmans, 2023
</taxonomicName>
(Cyclades, Iraklia) distributed in the
<collectingRegion id="49A2FCEAFFE3F16BFF420E16FECFF986" box="[234,320,1592,1617]" country="Greece" name="Voreio Aigaio" pageId="8" pageNumber="405">Aegean</collectingRegion>
islands. However,
<taxonomicName id="4C66498BFFE3F16BFDBB0E16FD16F986" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[531,665,1592,1617]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFE3F16BFDBB0E16FD16F986" box="[531,665,1592,1617]" italics="true" pageId="8" pageNumber="405">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE3F16BFD090E16FD6EF986" bold="true" box="[673,737,1592,1617]" pageId="8" pageNumber="405">
<taxonomicNameLabel id="A2215361FFE3F16BFD090E16FD6EF986" box="[673,737,1592,1617]" pageId="8" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from the latter two species by the robust, thick and comma-shaped embolus (E). Indeed, the embolus is straight in
<taxonomicName id="4C66498BFFE3F16BFE8E0E50FE45F940" authorityName="Bosmans" authorityYear="2023" box="[294,458,1662,1687]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="chaniaensis">
<emphasis id="B912EE1AFFE3F16BFE8E0E50FE45F940" box="[294,458,1662,1687]" italics="true" pageId="8" pageNumber="405">H. chaniaensis</emphasis>
</taxonomicName>
and slightly curved in
<taxonomicName id="4C66498BFFE3F16BFD410E50FC28F940" authorityName="Bosmans" authorityYear="2023" box="[745,935,1662,1687]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="walterdebucki">
<emphasis id="B912EE1AFFE3F16BFD410E50FC28F940" box="[745,935,1662,1687]" italics="true" pageId="8" pageNumber="405">H. walterdebucki</emphasis>
</taxonomicName>
(see
<figureCitation id="135D2E8DFFE3F16BFC450E50FBF1F940" box="[1005,1150,1662,1687]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="8" pageNumber="405">Figure 3BG</figureCitation>
; Bosmans 2023, fig. 7bh;
<bibRefCitation id="EFF74FF9FFE3F16BFE170E8EFD7EF96E" author="Bosmans R &amp; Gavalas I" box="[447,753,1696,1721]" pageId="8" pageNumber="405" pagination="1 - 91" refId="ref5650" refString="Bosmans R, Gavalas I. 2023. The spiders (Araneae) of the tiny Greek island Iraklia (Kiklades), with the descriptions of 5 new Harpactea species (Araneae: Dysderidae), 3 species new to Europe and 8 new to Greece. Parnassiana Arch. 11 (Supplementum 1): 1-91." type="journal article" year="2023">Bosmans and Gavalas 2023</bibRefCitation>
, fig. 7cg).
</paragraph>
<caption id="DF196280FFE3F16BFF080DB4FC48F9DF" pageId="8" pageNumber="405" startId="8.[160,225,1434,1457]" targetBox="[163,1153,142,1401]" targetPageId="8" targetType="figure">
<paragraph id="8BD93208FFE3F16BFF080DB4FC48F9DF" blockId="8.[160,1157,1434,1544]" pageId="8" pageNumber="405">
<emphasis id="B912EE1AFFE3F16BFF080DB4FF74FA66" bold="true" box="[160,251,1434,1457]" pageId="8" pageNumber="405">Figure 5.</emphasis>
<taxonomicName id="4C66498BFFE3F16BFEAB0DB4FE48FA66" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[259,455,1434,1457]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="405" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. nov.">
<emphasis id="B912EE1AFFE3F16BFEAB0DB4FE48FA66" box="[259,455,1434,1457]" italics="true" pageId="8" pageNumber="405">Harpactea dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE3F16BFE670DB4FD85FA66" bold="true" box="[463,522,1434,1457]" pageId="8" pageNumber="405">
<taxonomicNameLabel id="A2215361FFE3F16BFE670DB4FD85FA66" box="[463,522,1434,1457]" pageId="8" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar.
</paragraph>
</caption>
<paragraph id="8BD93208FFE2F16AFF6808A0FDEAFE82" blockId="9.[160,1156,142,341]" pageId="9" pageNumber="406">
In the female of
<taxonomicName id="4C66498BFFE2F16AFED208A0FE70FF70" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[378,511,142,167]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFE2F16AFED208A0FE70FF70" box="[378,511,142,167]" italics="true" pageId="9" pageNumber="406">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE2F16AFDAE08A0FDC9FF70" bold="true" box="[518,582,142,167]" pageId="9" pageNumber="406">
<taxonomicNameLabel id="A2215361FFE2F16AFDAE08A0FDC9FF70" box="[518,582,142,167]" pageId="9" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the vulva is similar to that of
<taxonomicName id="4C66498BFFE2F16AFC3008A0FBC3FF70" authorityName="Bosmans" authorityYear="2023" box="[920,1100,142,167]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="walterdebucki">
<emphasis id="B912EE1AFFE2F16AFC3008A0FBC3FF70" box="[920,1100,142,167]" italics="true" pageId="9" pageNumber="406">H. walterdebucki</emphasis>
</taxonomicName>
. The anterior arcs (AC) of the two species are similar in terms of their general morphological features. The most important difference is that the spermathecal keel (SK) is narrower in
<taxonomicName id="4C66498BFFE2F16AFF0808D8FEACFED8" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[160,291,246,271]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFE2F16AFF0808D8FEACFED8" box="[160,291,246,271]" italics="true" pageId="9" pageNumber="406">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE2F16AFE8108D8FEE8FED8" bold="true" box="[297,359,246,271]" pageId="9" pageNumber="406">
<taxonomicNameLabel id="A2215361FFE2F16AFE8108D8FEE8FED8" box="[297,359,246,271]" pageId="9" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="4C66498BFFE2F16AFE6908D8FDFBFED8" authorityName="Bosmans" authorityYear="2023" box="[449,628,246,271]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="walterdebucki">
<emphasis id="B912EE1AFFE2F16AFE6908D8FDFBFED8" box="[449,628,246,271]" italics="true" pageId="9" pageNumber="406">H. walterdebucki</emphasis>
</taxonomicName>
. There is also a prominent sclerotised plate (SP) posterior to the spermatheca in
<taxonomicName id="4C66498BFFE2F16AFDA70937FD1BFEE5" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[527,660,281,306]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFE2F16AFDA70937FD1BFEE5" box="[527,660,281,306]" italics="true" pageId="9" pageNumber="406">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE2F16AFD340937FD53FEE5" bold="true" box="[668,732,281,306]" pageId="9" pageNumber="406">
<taxonomicNameLabel id="A2215361FFE2F16AFD340937FD53FEE5" box="[668,732,281,306]" pageId="9" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, whereas in
<taxonomicName id="4C66498BFFE2F16AFCC40937FBAEFEE5" authorityName="Bosmans" authorityYear="2023" box="[876,1057,281,306]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="406" phylum="Arthropoda" rank="species" species="walterdebucki">
<emphasis id="B912EE1AFFE2F16AFCC40937FBAEFEE5" box="[876,1057,281,306]" italics="true" pageId="9" pageNumber="406">H. walterdebucki</emphasis>
</taxonomicName>
, there is a gently developed sclerotised belt (SB).
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFE2F168FF0809AEFD14FE82" lastPageId="11" lastPageNumber="408" pageId="9" pageNumber="406" type="description">
<paragraph id="8BD93208FFE2F16AFF0809AEFEA9FE4D" blockId="9.[160,1157,384,1485]" box="[160,294,384,410]" pageId="9" pageNumber="406">
<emphasis id="B912EE1AFFE2F16AFF0809AEFEA9FE4D" bold="true" box="[160,294,384,410]" italics="true" pageId="9" pageNumber="406">Description</emphasis>
</paragraph>
<paragraph id="8BD93208FFE2F16AFF08098DFC9EFDF2" blockId="9.[160,1157,384,1485]" pageId="9" pageNumber="406">
<emphasis id="B912EE1AFFE2F16AFF08098DFEDAFE6A" bold="true" box="[160,341,419,445]" italics="true" pageId="9" pageNumber="406">Measurements.</emphasis>
<emphasis id="B912EE1AFFE2F16AFECC098AFE53FE6A" bold="true" box="[356,476,420,445]" pageId="9" pageNumber="406">
[
<typeStatus id="54DD8CAAFFE2F16AFEC3098AFE53FE6A" box="[363,476,420,445]" pageId="9" pageNumber="406" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFE2F16AFE4F098AFE74FE6C" box="[487,507,420,443]" count="1" pageId="9" pageNumber="406" type="male"></specimenCount>
<emphasis id="B912EE1AFFE2F16AFE53098AFDF2FE6A" bold="true" box="[507,637,420,445]" pageId="9" pageNumber="406">
/
<typeStatus id="54DD8CAAFFE2F16AFDAB098AFDF2FE6A" box="[515,637,420,445]" pageId="9" pageNumber="406" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFE2F16AFD20098AFD13FE6C" box="[648,668,420,443]" count="1" pageId="9" pageNumber="406" type="female"></specimenCount>
<emphasis id="B912EE1AFFE2F16AFD34098AFD2BFE6A" bold="true" box="[668,676,420,445]" pageId="9" pageNumber="406">]</emphasis>
TL 2.94/4.084.19; AL 1.71/2.552.71; CL 1.23/1.481.53;
<subScript id="17E2304DFFE2F16AFEFF09E9FE2EFE36" attach="left" box="[343,417,455,481]" fontSize="7" pageId="9" pageNumber="406">CWmax</subScript>
0.99/1.111.14;
<subScript id="17E2304DFFE2F16AFDCC09E9FD23FE36" attach="left" box="[612,684,455,481]" fontSize="7" pageId="9" pageNumber="406">CWmin</subScript>
0.47/0.490.51; Clh 0.04/0.040.04; Aed 0.07/0.070.08; iAE 0.04/0.040.04; PLEd 0.06/0.060.08; PMEd 0.05/0.050.06; ChF 0.27/0.320.33; ChG 0.13/0.140.15; ChL 0.44/0.620.63.
</paragraph>
<paragraph id="8BD93208FFE2F16AFF680A01FCAEFDBD" blockId="9.[160,1157,384,1485]" pageId="9" pageNumber="406">
<emphasis id="B912EE1AFFE2F16AFF680A01FEBBFD9F" bold="true" box="[192,308,559,584]" pageId="9" pageNumber="406">
[
<typeStatus id="54DD8CAAFFE2F16AFF6E0A01FEBBFD9F" box="[198,308,559,584]" pageId="9" pageNumber="406" type="paratype">Paratype</typeStatus>
</emphasis>
<specimenCount id="9D60F981FFE2F16AFE930A01FEC0FD91" box="[315,335,559,582]" count="1" pageId="9" pageNumber="406" type="male"></specimenCount>
<emphasis id="B912EE1AFFE2F16AFEE70A01FED8FD9F" bold="true" box="[335,343,559,584]" pageId="9" pageNumber="406">]</emphasis>
TL 3.12; AL 1.73; CL 1.39;
<subScript id="17E2304DFFE2F16AFDDF0A01FD4EFD9E" attach="left" box="[631,705,559,585]" fontSize="7" pageId="9" pageNumber="406">CWmax</subScript>
1.07;
<subScript id="17E2304DFFE2F16AFCAA0A01FCC7FD9E" attach="left" box="[770,840,559,585]" fontSize="7" pageId="9" pageNumber="406">CWmin</subScript>
0.46; Clh 0.04; Aed 0.07; iAE 0.05; PLEd 0.06; PMEd 0.05; ChF 0.28; ChG 0.14; ChL 0.46.
</paragraph>
<paragraph id="8BD93208FFE2F16AFF680A5AFB0BFCCF" blockId="9.[160,1157,384,1485]" pageId="9" pageNumber="406">Small-sized harpacteine spider. Carapace brownish with smooth surface. Eye region of carapace darker than thoracic region. Fovea longitudinal, blackish-brown, distinct. Surface of carapace with sparse blackish short setae. There is a single line of hairs between fovea and PME. Eye region slightly higher than thoracic region. They are almost in alignment. Anterior margin of eyes region convex, while posterior margin of thoracic region straight.</paragraph>
<paragraph id="8BD93208FFE2F16AFF680B0FFE77FA1A" blockId="9.[160,1157,384,1485]" pageId="9" pageNumber="406">
Eyes silvery honey-coloured, arranged annularly. AE&gt;PLE = PME. AE seperated; AE-PLE 0.01, PLE-PME 0.01, PME-PME 0.01. Sternum yellowish brown; length 0.85, width 0.68. Surface with brownish setae. Setae in front of 3
<superScript id="7C139F40FFE2F16AFC2B0B4FFC1AFCA6" attach="left" box="[899,917,865,881]" fontSize="7" pageId="9" pageNumber="406">rd</superScript>
and 4
<superScript id="7C139F40FFE2F16AFC410B4FFC74FCA6" attach="left" box="[1001,1019,865,881]" fontSize="7" pageId="9" pageNumber="406">th</superScript>
coxae and posterior margin of sternum longer. Labium and gnathocoxae brownish. Labium length 0.26, width 0.19. Gnathocoxae length 0.48, width 0.25. Chelicerae brownish. Cheliceral groove with 4 teeth (
<figureCitation id="135D2E8DFFE2F16AFD910BE1FD3EFC3F" box="[569,689,975,1000]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="9" pageNumber="406">Figure 3A</figureCitation>
). Promarginal teeth triangular, retromarginal teeth conical. Promarginal teeth more strongly developed than retromarginal teeth. They are several times larger than retromarginal teeth. While retromarginal teeth are equal in length to each other, promarginal tooth at the base of the cheliceral groove slightly larger than one following it. There is a distance of 0.01 between the promarginal and 0.04 between the retromarginal teeth. Abdomen brownish greyish. Surface covered with brownish setae. Legs yellowish light brown. Anterior legs slightly darker than posterior legs. Tarsi with 3 claws. Paired claws toothed. Paired claws of legs I and II with 67 teeth; legs III and IV with 5 teeth. Anterior legs without scopulae. Tarsi three and four and first distal quarter of metatarsi three and four with weak scopulae. First and second femora with 2 spines prolaterally. First femoral spines arranged side by side, and second femoral spines arranged back to back. Third and fourth coxae also with spine. Leg formula: VI, I, II, III. Leg spination and measurements are given in
<tableCitation id="C6E407B3FFE2F16AFF080D9AFE8BFA1A" box="[160,260,1460,1485]" captionStart="Table 3" captionStartId="10.[243,299,142,165]" captionText="Table 3. Leg spination of H. dilekensis sp. n. (Holotype ♂/Paratype ♀)." pageId="9" pageNumber="406">Tables 3</tableCitation>
and
<tableCitation id="C6E407B3FFE2F16AFEEF0D9AFEDAFA1A" box="[327,341,1460,1485]" captionStart="Table 4" captionStartId="10.[182,238,561,584]" captionText="Table 4. Leg measurements of H. dilekensis sp. n. (Holotype ♂/Paratypes ♂–♀)." pageId="9" pageNumber="406">4</tableCitation>
, respectively.
</paragraph>
<paragraph id="8BD93208FFE2F169FF080DD7FEA7FB3A" blockId="9.[160,1156,1529,1728]" lastBlockId="10.[160,1157,1028,1400]" lastPageId="10" lastPageNumber="407" pageId="9" pageNumber="406">
<emphasis id="B912EE1AFFE2F16AFF080DD7FE78F9C4" bold="true" box="[160,503,1529,1555]" pageId="9" pageNumber="406">
<emphasis id="B912EE1AFFE2F16AFF080DD7FF5AF9C4" bold="true" box="[160,213,1529,1555]" italics="true" pageId="9" pageNumber="406">Palp</emphasis>
. (
<figureCitation id="135D2E8DFFE2F16AFF580DD7FE16F9C4" box="[240,409,1529,1555]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="9" pageNumber="406">Figures 3BG</figureCitation>
,
<figureCitation id="135D2E8DFFE2F16AFE010DD7FE66F9C4" box="[425,489,1529,1555]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="9" pageNumber="406">4DE</figureCitation>
).
</emphasis>
Palpal tarsus triangular, length 0.36 (
<figureCitation id="135D2E8DFFE2F16AFC770DD4FBF9F9C4" box="[991,1142,1530,1555]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="9" pageNumber="406">Figure 3DE</figureCitation>
). Palpal tibia shorter than tarsus, length 0.28. Ratio Ti/Ta 0.77. Tegulum (T) yellowish brown, spherical, gradually narrowing distally; length 0.34, width 0.32 (
<figureCitation id="135D2E8DFFE2F16AFBA20E11FF20F9AC" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="9" pageNumber="406">Figure 3B, C</figureCitation>
). Embolus (E), conductor (C) and median apophysis (MA) black, strongly sclerotised. Embolus longer than conductor and median apophysis (
<figureCitation id="135D2E8DFFE2F16AFC740EABFBFAF949" box="[988,1141,1669,1694]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="9" pageNumber="406">Figure 3CF</figureCitation>
). Embolus length 0.21, conductor 0.08. Embolar base wide. It is on same axis as tegulum. Conductor and median apophysis aligned and positioned opposite each other. Median apophysis shaped like a small tubercle, while the conductor relatively larger (
<figureCitation id="135D2E8DFFE1F169FE960C64FE6FFBB4" box="[318,480,1098,1123]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="10" pageNumber="407">Figures 3D, G</figureCitation>
,
<figureCitation id="135D2E8DFFE1F169FE450C64FDA7FBB4" box="[493,552,1098,1123]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="10" pageNumber="407">4DE</figureCitation>
). Conductor on a convex-shaped sclerotised plate. Its shape like a triangular hook, and its tip orientated at three oclock. Embolus comma-shaped and gradually tapers from base to tip (
<figureCitation id="135D2E8DFFE1F169FC9E0CA1FC4AFB7F" box="[822,965,1167,1192]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="10" pageNumber="407">Figure 4DE</figureCitation>
). Base on same axis as tegulum and tip curved ventrally at an angle of about 45° to this axis (
<figureCitation id="135D2E8DFFE1F169FF000CFAFE94FB3A" box="[168,283,1236,1261]" captionStart="Figure 3" captionStartId="4.[160,225,888,911]" captionTargetBox="[172,1149,145,852]" captionTargetId="figure-211@4.[163,1153,142,857]" captionTargetPageId="4" captionText="Figure 3. Harpactea dilekensis sp. n. (A) Chelicerae, ventral view (red dots indicate promarginal; white dots indicate retromarginal teeth). (B) Male palp, left, nearly retrolateral. (C) Same, retrolateral. (D) Same, nearly retrolateral view. (E) Same, nearly retrolateral view. (F) Same, nearly ventral view. (G) Same, prolateral view. Abbreviations: C conductor, E embolus, MA median apophysis, T tegulum." pageId="10" pageNumber="407">Figure 3C</figureCitation>
).
</paragraph>
<caption id="DF196280FFE1F169FF5B08A0FC20FF72" box="[243,943,142,165]" pageId="10" pageNumber="407" startId="10.[243,299,142,165]" targetType="table">
<paragraph id="8BD93208FFE1F169FF5B08A0FC20FF72" blockId="10.[243,943,142,165]" box="[243,943,142,165]" pageId="10" pageNumber="407">
<emphasis id="B912EE1AFFE1F169FF5B08A0FECBFF72" bold="true" box="[243,324,142,165]" pageId="10" pageNumber="407">Table 3.</emphasis>
Leg spination of
<taxonomicName id="4C66498BFFE1F169FE5B08A0FDE7FF72" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[499,616,142,165]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFE1F169FE5B08A0FDE7FF72" box="[499,616,142,165]" italics="true" pageId="10" pageNumber="407">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE1F169FDC708A0FD26FF72" bold="true" box="[623,681,142,165]" pageId="10" pageNumber="407">
<taxonomicNameLabel id="A2215361FFE1F169FDC708A0FD26FF72" box="[623,681,142,165]" pageId="10" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Holotype ♂/Paratype ♀).
</paragraph>
</caption>
<paragraph id="8BD93208FFE1F169FF55089CFBA9FE39" pageId="10" pageNumber="407">
<table id="F966C0A8FFE10E9CFF55089CFBA9FE39" box="[253,1062,178,494]" gridcols="5" gridrows="12" pageId="10" pageNumber="407">
<tr id="3556304AFFE10E9CFF55089CFBA9FF11" box="[253,1062,178,198]" gridrow="0" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF55089CFEACFF11" box="[253,291,178,198]" gridcol="0" gridrow="0" pageId="10" pageNumber="407">Legs</th>
<th id="76875936FFE10E9CFE30089CFE36FF11" box="[408,441,178,198]" gridcol="1" gridrow="0" pageId="10" pageNumber="407">I</th>
<th id="76875936FFE10E9CFDB0089CFDC2FF11" box="[536,589,178,198]" gridcol="2" gridrow="0" pageId="10" pageNumber="407">II</th>
<th id="76875936FFE10E9CFD05089CFCB9FF11" box="[685,822,178,198]" gridcol="3" gridrow="0" pageId="10" pageNumber="407">III</th>
<th id="76875936FFE10E9CFC3E089CFBA9FF11" box="[918,1062,178,198]" gridcol="4" gridrow="0" pageId="10" pageNumber="407">IV</th>
</tr>
<tr id="3556304AFFE10E9CFF5508E1FBA9FF32" box="[253,1062,207,229]" gridrow="1" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF5508E1FBA9FF32" box="[253,1062,207,229]" colspan="5" colspanRight="4" gridcol="0" gridrow="1" pageId="10" pageNumber="407"></th>
</tr>
<tr id="3556304AFFE10E9CFF5508C4FBA9FF29" box="[253,1062,234,254]" gridrow="2" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF5508C4FEACFF29" box="[253,291,234,254]" gridcol="0" gridrow="2" pageId="10" pageNumber="407">Cx</th>
<td id="76875936FFE10E9CFE3008C4FE36FF29" box="[408,441,234,254]" gridcol="1" gridrow="2" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB008C4FDC2FF29" box="[536,589,234,254]" gridcol="2" gridrow="2" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD0508C4FCB9FF29" box="[685,822,234,254]" gridcol="3" gridrow="2" pageId="10" pageNumber="407">1 pl</td>
<td id="76875936FFE10E9CFC3E08C4FBA9FF29" box="[918,1062,234,254]" gridcol="4" gridrow="2" pageId="10" pageNumber="407">2 pl</td>
</tr>
<tr id="3556304AFFE10E9CFF55092CFBA9FEC1" box="[253,1062,258,278]" gridrow="3" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF55092CFEACFEC1" box="[253,291,258,278]" gridcol="0" gridrow="3" pageId="10" pageNumber="407">Fe</th>
<td id="76875936FFE10E9CFE30092CFE36FEC1" box="[408,441,258,278]" gridcol="1" gridrow="3" pageId="10" pageNumber="407">2 pl</td>
<td id="76875936FFE10E9CFDB0092CFDC2FEC1" box="[536,589,258,278]" gridcol="2" gridrow="3" pageId="10" pageNumber="407">1, 1 pl</td>
<td id="76875936FFE10E9CFD05092CFCB9FEC1" box="[685,822,258,278]" gridcol="3" gridrow="3" pageId="10" pageNumber="407">2(3) rd 2 pd</td>
<td id="76875936FFE10E9CFC3E092CFBA9FEC1" box="[918,1062,258,278]" gridcol="4" gridrow="3" pageId="10" pageNumber="407">1 d 2 pl</td>
</tr>
<tr id="3556304AFFE10E9CFF550934FBA9FEF9" box="[253,1062,282,302]" gridrow="4" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF550934FEACFEF9" box="[253,291,282,302]" gridcol="0" gridrow="4" pageId="10" pageNumber="407">Pa</th>
<td id="76875936FFE10E9CFE300934FE36FEF9" box="[408,441,282,302]" gridcol="1" gridrow="4" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB00934FDC2FEF9" box="[536,589,282,302]" gridcol="2" gridrow="4" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD050934FCB9FEF9" box="[685,822,282,302]" gridcol="3" gridrow="4" pageId="10" pageNumber="407">1 rd</td>
<td id="76875936FFE10E9CFC3E0934FBA9FEF9" box="[918,1062,282,302]" gridcol="4" gridrow="4" pageId="10" pageNumber="407">0</td>
</tr>
<tr id="3556304AFFE10E9CFF55091CFBA9FE91" box="[253,1062,306,326]" gridrow="5" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF55091CFEACFE91" box="[253,291,306,326]" gridcol="0" gridrow="5" pageId="10" pageNumber="407">Ti</th>
<td id="76875936FFE10E9CFE30091CFE36FE91" box="[408,441,306,326]" gridcol="1" gridrow="5" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB0091CFDC2FE91" box="[536,589,306,326]" gridcol="2" gridrow="5" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD05091CFCB9FE91" box="[685,822,306,326]" gridcol="3" gridrow="5" pageId="10" pageNumber="407">3 rl 2 pl 1, 1, 2 v</td>
<td id="76875936FFE10E9CFC3E091CFBA9FE91" box="[918,1062,306,326]" gridcol="4" gridrow="5" pageId="10" pageNumber="407">3 rl 3 pl 1, 1, 2 v</td>
</tr>
<tr id="3556304AFFE10E9CFF550964FBA9FEA2" box="[253,1062,330,373]" gridrow="6" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF550964FEACFEA2" box="[253,291,330,373]" gridcol="0" gridrow="6" pageId="10" pageNumber="407">Me ♀</th>
<td id="76875936FFE10E9CFE300964FE36FEA2" box="[408,441,330,373]" gridcol="1" gridrow="6" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB00964FDC2FEA2" box="[536,589,330,373]" gridcol="2" gridrow="6" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD050964FCB9FEA2" box="[685,822,330,373]" gridcol="3" gridrow="6" pageId="10" pageNumber="407">2 pl 1, 1, 2 v</td>
<td id="76875936FFE10E9CFC3E0964FBA9FEA2" box="[918,1062,330,373]" gridcol="4" gridrow="6" pageId="10" pageNumber="407">3 rl 3 pd 1, 2 v</td>
</tr>
<tr id="3556304AFFE10E9CFF550954FBA9FE59" box="[253,1062,378,398]" gridrow="7" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF550954FEACFE59" box="[253,291,378,398]" gridcol="0" gridrow="7" pageId="10" pageNumber="407">Cx</th>
<td id="76875936FFE10E9CFE300954FE36FE59" box="[408,441,378,398]" gridcol="1" gridrow="7" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB00954FDC2FE59" box="[536,589,378,398]" gridcol="2" gridrow="7" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD050954FCB9FE59" box="[685,822,378,398]" gridcol="3" gridrow="7" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFC3E0954FBA9FE59" box="[918,1062,378,398]" gridcol="4" gridrow="7" pageId="10" pageNumber="407">1 pl</td>
</tr>
<tr id="3556304AFFE10E9CFF5509BCFBA9FE71" box="[253,1062,402,422]" gridrow="8" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF5509BCFEACFE71" box="[253,291,402,422]" gridcol="0" gridrow="8" pageId="10" pageNumber="407">Fe</th>
<td id="76875936FFE10E9CFE3009BCFE36FE71" box="[408,441,402,422]" gridcol="1" gridrow="8" pageId="10" pageNumber="407">2 pl</td>
<td id="76875936FFE10E9CFDB009BCFDC2FE71" box="[536,589,402,422]" gridcol="2" gridrow="8" pageId="10" pageNumber="407">1, 1 pl</td>
<td id="76875936FFE10E9CFD0509BCFCB9FE71" box="[685,822,402,422]" gridcol="3" gridrow="8" pageId="10" pageNumber="407">3 rd 2 pd</td>
<td id="76875936FFE10E9CFC3E09BCFBA9FE71" box="[918,1062,402,422]" gridcol="4" gridrow="8" pageId="10" pageNumber="407">1 d 2 pl</td>
</tr>
<tr id="3556304AFFE10E9CFF550984FBA9FE69" box="[253,1062,426,446]" gridrow="9" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF550984FEACFE69" box="[253,291,426,446]" gridcol="0" gridrow="9" pageId="10" pageNumber="407">Pa</th>
<td id="76875936FFE10E9CFE300984FE36FE69" box="[408,441,426,446]" gridcol="1" gridrow="9" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB00984FDC2FE69" box="[536,589,426,446]" gridcol="2" gridrow="9" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD050984FCB9FE69" box="[685,822,426,446]" gridcol="3" gridrow="9" pageId="10" pageNumber="407">1 rd</td>
<td id="76875936FFE10E9CFC3E0984FBA9FE69" box="[918,1062,426,446]" gridcol="4" gridrow="9" pageId="10" pageNumber="407">0</td>
</tr>
<tr id="3556304AFFE10E9CFF5509ECFBA9FE01" box="[253,1062,450,470]" gridrow="10" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF5509ECFEACFE01" box="[253,291,450,470]" gridcol="0" gridrow="10" pageId="10" pageNumber="407">Ti</th>
<td id="76875936FFE10E9CFE3009ECFE36FE01" box="[408,441,450,470]" gridcol="1" gridrow="10" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB009ECFDC2FE01" box="[536,589,450,470]" gridcol="2" gridrow="10" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD0509ECFCB9FE01" box="[685,822,450,470]" gridcol="3" gridrow="10" pageId="10" pageNumber="407">3 rl 2 pl 1, 1, 2 v</td>
<td id="76875936FFE10E9CFC3E09ECFBA9FE01" box="[918,1062,450,470]" gridcol="4" gridrow="10" pageId="10" pageNumber="407">3 rl 3 pl 1, 1, 2 v</td>
</tr>
<tr id="3556304AFFE10E9CFF5509F4FBA9FE39" box="[253,1062,474,494]" gridrow="11" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF5509F4FEACFE39" box="[253,291,474,494]" gridcol="0" gridrow="11" pageId="10" pageNumber="407">Me</th>
<td id="76875936FFE10E9CFE3009F4FE36FE39" box="[408,441,474,494]" gridcol="1" gridrow="11" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFDB009F4FDC2FE39" box="[536,589,474,494]" gridcol="2" gridrow="11" pageId="10" pageNumber="407">0</td>
<td id="76875936FFE10E9CFD0509F4FCB9FE39" box="[685,822,474,494]" gridcol="3" gridrow="11" pageId="10" pageNumber="407">2 rl 2 pl 1, 1, 2 v</td>
<td id="76875936FFE10E9CFC3E09F4FBA9FE39" box="[918,1062,474,494]" gridcol="4" gridrow="11" pageId="10" pageNumber="407">3 rl 3 pd 1, 1, 2 v</td>
</tr>
</table>
</paragraph>
<caption id="DF196280FFE1F169FF1E0A1FFC5EFD9F" box="[182,977,561,584]" pageId="10" pageNumber="407" startId="10.[182,238,561,584]" targetType="table">
<paragraph id="8BD93208FFE1F169FF1E0A1FFC5EFD9F" blockId="10.[182,977,561,584]" box="[182,977,561,584]" pageId="10" pageNumber="407">
<emphasis id="B912EE1AFFE1F169FF1E0A1FFE88FD9F" bold="true" box="[182,263,561,584]" pageId="10" pageNumber="407">Table 4.</emphasis>
Leg measurements of
<taxonomicName id="4C66498BFFE1F169FE440A1FFDEEFD9F" authorityName="Kunt &amp; Danışman &amp; Yağmur" authorityYear="2025" box="[492,609,561,584]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="407" phylum="Arthropoda" rank="species" species="dilekensis" status="sp. n.">
<emphasis id="B912EE1AFFE1F169FE440A1FFDEEFD9F" box="[492,609,561,584]" italics="true" pageId="10" pageNumber="407">H. dilekensis</emphasis>
</taxonomicName>
<emphasis id="B912EE1AFFE1F169FDC00A1FFD2DFD9F" bold="true" box="[616,674,561,584]" pageId="10" pageNumber="407">
<taxonomicNameLabel id="A2215361FFE1F169FDC00A1FFD2DFD9F" box="[616,674,561,584]" pageId="10" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Holotype ♂/Paratypes ♂–♀).
</paragraph>
</caption>
<paragraph id="8BD93208FFE1F169FF680A7BFBECFC16" pageId="10" pageNumber="407">
<table id="F966C0A8FFE10E9CFF680A7BFBECFC16" box="[192,1123,597,961]" gridcols="5" gridrows="14" pageId="10" pageNumber="407">
<tr id="3556304AFFE10E9CFF680A7BFBECFDBE" box="[192,1123,597,617]" gridrow="0" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680A7BFF7DFDBE" box="[192,242,597,617]" gridcol="0" gridrow="0" pageId="10" pageNumber="407">Legs</th>
<th id="76875936FFE10E9CFEF40A7BFE59FDBE" box="[348,470,597,617]" gridcol="1" gridrow="0" pageId="10" pageNumber="407">I</th>
<th id="76875936FFE10E9CFD9D0A7BFD3FFDBE" box="[565,688,597,617]" gridcol="2" gridrow="0" pageId="10" pageNumber="407">II</th>
<th id="76875936FFE10E9CFCA70A7BFC06FDBE" box="[783,905,597,617]" gridcol="3" gridrow="0" pageId="10" pageNumber="407">III</th>
<th id="76875936FFE10E9CFC410A7BFBECFDBE" box="[1001,1123,597,617]" gridcol="4" gridrow="0" pageId="10" pageNumber="407">IV</th>
</tr>
<tr id="3556304AFFE10E9CFF680A5CFBECFD5F" box="[192,1123,626,648]" gridrow="1" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680A5CFBECFD5F" box="[192,1123,626,648]" colspan="5" colspanRight="4" gridcol="0" gridrow="1" pageId="10" pageNumber="407"></th>
</tr>
<tr id="3556304AFFE10E9CFF680AA3FBECFD76" box="[192,1123,653,673]" gridrow="2" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680AA3FF7DFD76" box="[192,242,653,673]" gridcol="0" gridrow="2" pageId="10" pageNumber="407">Fe</th>
<td id="76875936FFE10E9CFEF40AA3FE59FD76" box="[348,470,653,673]" gridcol="1" gridrow="2" pageId="10" pageNumber="407">0.94</td>
<td id="76875936FFE10E9CFD9D0AA3FD3FFD76" box="[565,688,653,673]" gridcol="2" gridrow="2" pageId="10" pageNumber="407">1.01</td>
<td id="76875936FFE10E9CFCA70AA3FC06FD76" box="[783,905,653,673]" gridcol="3" gridrow="2" pageId="10" pageNumber="407">0.80</td>
<td id="76875936FFE10E9CFC410AA3FBECFD76" box="[1001,1123,653,673]" gridcol="4" gridrow="2" pageId="10" pageNumber="407">1.14</td>
</tr>
<tr id="3556304AFFE10E9CFF680A8BFBECFD6E" box="[192,1123,677,697]" gridrow="3" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680A8BFF7DFD6E" box="[192,242,677,697]" gridcol="0" gridrow="3" pageId="10" pageNumber="407">Pa</th>
<td id="76875936FFE10E9CFEF40A8BFE59FD6E" box="[348,470,677,697]" gridcol="1" gridrow="3" pageId="10" pageNumber="407">0.72</td>
<td id="76875936FFE10E9CFD9D0A8BFD3FFD6E" box="[565,688,677,697]" gridcol="2" gridrow="3" pageId="10" pageNumber="407">0.63</td>
<td id="76875936FFE10E9CFCA70A8BFC06FD6E" box="[783,905,677,697]" gridcol="3" gridrow="3" pageId="10" pageNumber="407">0.40</td>
<td id="76875936FFE10E9CFC410A8BFBECFD6E" box="[1001,1123,677,697]" gridcol="4" gridrow="3" pageId="10" pageNumber="407">0.54</td>
</tr>
<tr id="3556304AFFE10E9CFF680A93FBECFD06" box="[192,1123,701,721]" gridrow="4" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680A93FF7DFD06" box="[192,242,701,721]" gridcol="0" gridrow="4" pageId="10" pageNumber="407">Ti</th>
<td id="76875936FFE10E9CFEF40A93FE59FD06" box="[348,470,701,721]" gridcol="1" gridrow="4" pageId="10" pageNumber="407">0.87</td>
<td id="76875936FFE10E9CFD9D0A93FD3FFD06" box="[565,688,701,721]" gridcol="2" gridrow="4" pageId="10" pageNumber="407">0.82</td>
<td id="76875936FFE10E9CFCA70A93FC06FD06" box="[783,905,701,721]" gridcol="3" gridrow="4" pageId="10" pageNumber="407">0.61</td>
<td id="76875936FFE10E9CFC410A93FBECFD06" box="[1001,1123,701,721]" gridcol="4" gridrow="4" pageId="10" pageNumber="407">0.89</td>
</tr>
<tr id="3556304AFFE10E9CFF680AFBFBECFD3E" box="[192,1123,725,745]" gridrow="5" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680AFBFF7DFD3E" box="[192,242,725,745]" gridcol="0" gridrow="5" pageId="10" pageNumber="407">Me</th>
<td id="76875936FFE10E9CFEF40AFBFE59FD3E" box="[348,470,725,745]" gridcol="1" gridrow="5" pageId="10" pageNumber="407">0.80</td>
<td id="76875936FFE10E9CFD9D0AFBFD3FFD3E" box="[565,688,725,745]" gridcol="2" gridrow="5" pageId="10" pageNumber="407">0.74</td>
<td id="76875936FFE10E9CFCA70AFBFC06FD3E" box="[783,905,725,745]" gridcol="3" gridrow="5" pageId="10" pageNumber="407">0.75</td>
<td id="76875936FFE10E9CFC410AFBFBECFD3E" box="[1001,1123,725,745]" gridcol="4" gridrow="5" pageId="10" pageNumber="407">1.17</td>
</tr>
<tr id="3556304AFFE10E9CFF680AC3FBECFCD6" box="[192,1123,749,769]" gridrow="6" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680AC3FF7DFCD6" box="[192,242,749,769]" gridcol="0" gridrow="6" pageId="10" pageNumber="407">Ta</th>
<td id="76875936FFE10E9CFEF40AC3FE59FCD6" box="[348,470,749,769]" gridcol="1" gridrow="6" pageId="10" pageNumber="407">0.29</td>
<td id="76875936FFE10E9CFD9D0AC3FD3FFCD6" box="[565,688,749,769]" gridcol="2" gridrow="6" pageId="10" pageNumber="407">0.29</td>
<td id="76875936FFE10E9CFCA70AC3FC06FCD6" box="[783,905,749,769]" gridcol="3" gridrow="6" pageId="10" pageNumber="407">0.29</td>
<td id="76875936FFE10E9CFC410AC3FBECFCD6" box="[1001,1123,749,769]" gridcol="4" gridrow="6" pageId="10" pageNumber="407">0.30</td>
</tr>
<tr id="3556304AFFE10E9CFF680B2BFBECFCE6" box="[192,1123,773,817]" gridrow="7" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680B2BFF7DFCE6" box="[192,242,773,817]" gridcol="0" gridrow="7" pageId="10" pageNumber="407">Total ♂/♀</th>
<td id="76875936FFE10E9CFEF40B2BFE59FCE6" box="[348,470,773,817]" gridcol="1" gridrow="7" pageId="10" pageNumber="407">3.62</td>
<td id="76875936FFE10E9CFD9D0B2BFD3FFCE6" box="[565,688,773,817]" gridcol="2" gridrow="7" pageId="10" pageNumber="407">3.49</td>
<td id="76875936FFE10E9CFCA70B2BFC06FCE6" box="[783,905,773,817]" gridcol="3" gridrow="7" pageId="10" pageNumber="407">2.85</td>
<td id="76875936FFE10E9CFC410B2BFBECFCE6" box="[1001,1123,773,817]" gridcol="4" gridrow="7" pageId="10" pageNumber="407">4.04</td>
</tr>
<tr id="3556304AFFE10E9CFF680B1BFBECFC9E" box="[192,1123,821,841]" gridrow="8" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680B1BFF7DFC9E" box="[192,242,821,841]" gridcol="0" gridrow="8" pageId="10" pageNumber="407">Fe</th>
<td id="76875936FFE10E9CFEF40B1BFE59FC9E" box="[348,470,821,841]" gridcol="1" gridrow="8" pageId="10" pageNumber="407">1.01/1.011.14</td>
<td id="76875936FFE10E9CFD9D0B1BFD3FFC9E" box="[565,688,821,841]" gridcol="2" gridrow="8" pageId="10" pageNumber="407">0.97/1.031.09</td>
<td id="76875936FFE10E9CFCA70B1BFC06FC9E" box="[783,905,821,841]" gridcol="3" gridrow="8" pageId="10" pageNumber="407">0.81/0.860.98</td>
<td id="76875936FFE10E9CFC410B1BFBECFC9E" box="[1001,1123,821,841]" gridcol="4" gridrow="8" pageId="10" pageNumber="407">1.17/1.221.24</td>
</tr>
<tr id="3556304AFFE10E9CFF680B63FBECFCB6" box="[192,1123,845,865]" gridrow="9" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680B63FF7DFCB6" box="[192,242,845,865]" gridcol="0" gridrow="9" pageId="10" pageNumber="407">Pa</th>
<td id="76875936FFE10E9CFEF40B63FE59FCB6" box="[348,470,845,865]" gridcol="1" gridrow="9" pageId="10" pageNumber="407">0.69/0.690.71</td>
<td id="76875936FFE10E9CFD9D0B63FD3FFCB6" box="[565,688,845,865]" gridcol="2" gridrow="9" pageId="10" pageNumber="407">0.64/0.530.62</td>
<td id="76875936FFE10E9CFCA70B63FC06FCB6" box="[783,905,845,865]" gridcol="3" gridrow="9" pageId="10" pageNumber="407">0.44/0.390.44</td>
<td id="76875936FFE10E9CFC410B63FBECFCB6" box="[1001,1123,845,865]" gridcol="4" gridrow="9" pageId="10" pageNumber="407">0.57/0.590.64</td>
</tr>
<tr id="3556304AFFE10E9CFF680B4BFBECFCAE" box="[192,1123,869,889]" gridrow="10" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680B4BFF7DFCAE" box="[192,242,869,889]" gridcol="0" gridrow="10" pageId="10" pageNumber="407">Ti</th>
<td id="76875936FFE10E9CFEF40B4BFE59FCAE" box="[348,470,869,889]" gridcol="1" gridrow="10" pageId="10" pageNumber="407">0.91/0.590.81</td>
<td id="76875936FFE10E9CFD9D0B4BFD3FFCAE" box="[565,688,869,889]" gridcol="2" gridrow="10" pageId="10" pageNumber="407">0.83/0.780.81</td>
<td id="76875936FFE10E9CFCA70B4BFC06FCAE" box="[783,905,869,889]" gridcol="3" gridrow="10" pageId="10" pageNumber="407">0.59/0.630.67</td>
<td id="76875936FFE10E9CFC410B4BFBECFCAE" box="[1001,1123,869,889]" gridcol="4" gridrow="10" pageId="10" pageNumber="407">0.98/1.011.04</td>
</tr>
<tr id="3556304AFFE10E9CFF680B53FBECFC46" box="[192,1123,893,913]" gridrow="11" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680B53FF7DFC46" box="[192,242,893,913]" gridcol="0" gridrow="11" pageId="10" pageNumber="407">Me</th>
<td id="76875936FFE10E9CFEF40B53FE59FC46" box="[348,470,893,913]" gridcol="1" gridrow="11" pageId="10" pageNumber="407">0.79/0.580.62</td>
<td id="76875936FFE10E9CFD9D0B53FD3FFC46" box="[565,688,893,913]" gridcol="2" gridrow="11" pageId="10" pageNumber="407">0.75/0.740.79</td>
<td id="76875936FFE10E9CFCA70B53FC06FC46" box="[783,905,893,913]" gridcol="3" gridrow="11" pageId="10" pageNumber="407">0.69/0.710.85</td>
<td id="76875936FFE10E9CFC410B53FBECFC46" box="[1001,1123,893,913]" gridcol="4" gridrow="11" pageId="10" pageNumber="407">1.09/1.131.22</td>
</tr>
<tr id="3556304AFFE10E9CFF680BBBFBECFC7E" box="[192,1123,917,937]" gridrow="12" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680BBBFF7DFC7E" box="[192,242,917,937]" gridcol="0" gridrow="12" pageId="10" pageNumber="407">Ta</th>
<td id="76875936FFE10E9CFEF40BBBFE59FC7E" box="[348,470,917,937]" gridcol="1" gridrow="12" pageId="10" pageNumber="407">0.32/0.300.30</td>
<td id="76875936FFE10E9CFD9D0BBBFD3FFC7E" box="[565,688,917,937]" gridcol="2" gridrow="12" pageId="10" pageNumber="407">0.31/0.300.31</td>
<td id="76875936FFE10E9CFCA70BBBFC06FC7E" box="[783,905,917,937]" gridcol="3" gridrow="12" pageId="10" pageNumber="407">0.31/0.320.35</td>
<td id="76875936FFE10E9CFC410BBBFBECFC7E" box="[1001,1123,917,937]" gridcol="4" gridrow="12" pageId="10" pageNumber="407">0.34/0.320.34</td>
</tr>
<tr id="3556304AFFE10E9CFF680B83FBECFC16" box="[192,1123,941,961]" gridrow="13" pageId="10" pageNumber="407">
<th id="76875936FFE10E9CFF680B83FF7DFC16" box="[192,242,941,961]" gridcol="0" gridrow="13" pageId="10" pageNumber="407">Total</th>
<td id="76875936FFE10E9CFEF40B83FE59FC16" box="[348,470,941,961]" gridcol="1" gridrow="13" pageId="10" pageNumber="407">3.72/3.173.58</td>
<td id="76875936FFE10E9CFD9D0B83FD3FFC16" box="[565,688,941,961]" gridcol="2" gridrow="13" pageId="10" pageNumber="407">3.50/3.383.62</td>
<td id="76875936FFE10E9CFCA70B83FC06FC16" box="[783,905,941,961]" gridcol="3" gridrow="13" pageId="10" pageNumber="407">2.84/2.913.29</td>
<td id="76875936FFE10E9CFC410B83FBECFC16" box="[1001,1123,941,961]" gridcol="4" gridrow="13" pageId="10" pageNumber="407">4.15/4.274.48</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD93208FFE1F169FF680CD9FCD7FAAF" blockId="10.[160,1157,1028,1400]" pageId="10" pageNumber="407">Although there are size differences between the somatic parts of males and females, they are generally similar. Palpal tarsus cylindrical, length 0.350.37. Palpal tibia shorter than tarsus, length 0.260.28. Female individuals have slightly developed epigastral scutum in gastral area. In females, third coxae without spines.</paragraph>
<paragraph id="8BD93208FFE1F169FF080D9DFE5DF9C4" blockId="10.[160,1156,1459,1728]" pageId="10" pageNumber="407">
<emphasis id="B912EE1AFFE1F169FF080D9DFEF6FA1A" bold="true" box="[160,377,1459,1485]" pageId="10" pageNumber="407">
<emphasis id="B912EE1AFFE1F169FF080D9DFF6BFA1A" bold="true" box="[160,228,1459,1485]" italics="true" pageId="10" pageNumber="407">Vulva</emphasis>
. (
<figureCitation id="135D2E8DFFE1F169FF570D9DFEE6FA1A" box="[255,361,1459,1485]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="10" pageNumber="407">Figure 5</figureCitation>
).
</emphasis>
General parts of vulva strongly sclerotised at almost same level. Sclerotised plate (SP), posterior and central region of spermatheca (S) less sclerotised than in general (
<figureCitation id="135D2E8DFFE1F169FEF30DD4FE4AF9C4" box="[347,453,1530,1555]" captionStart="Figure 4" captionStartId="5.[160,225,1188,1211]" captionTargetBox="[163,1153,142,1157]" captionTargetId="figure-127@5.[163,1153,142,1157]" captionTargetPageId="5" captionText="Figure 4. Scanning electron microscopy images of the male palp. (AC) Harpactea bozdaglarensis sp. n. (DE) Harpactea dilekensis sp. n. Abbreviations: C conductor, E embolus, MA median apophysis, o sperm outlet." pageId="10" pageNumber="407">Figure 4B</figureCitation>
).
</paragraph>
<paragraph id="8BD93208FFE1F168FF680E32FD14FE82" blockId="10.[160,1156,1459,1728]" lastBlockId="11.[160,1156,142,341]" lastPageId="11" lastPageNumber="408" pageId="10" pageNumber="407">
Anterior arc (AC) longer in width than length. Central region of spermatheca bulging, ovoid. Posterior and anterior half almost equal in size. However, quadrangular posterior half wider than anterior half. Spermathecal keel (SK) blunt (
<figureCitation id="135D2E8DFFE1F169FC9D0E4CFC32F9AC" box="[821,957,1634,1659]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="10" pageNumber="407">Figure 5BD</figureCitation>
). Anterior margin of anterior arc (AA) thick and robust. Lateral margins initially oriented anteriorly in a halfmoon shape and then bent and oriented laterally. Distinctive roundish structures (R
<emphasis id="B912EE1AFFE1F169FBC20E89FBF9F917" box="[1130,1142,1703,1728]" italics="true" pageId="10" pageNumber="407">S</emphasis>
), nearly half height of anterior arcs anterior margin (
<figureCitation id="135D2E8DFFE0F168FD5108A0FC0AFF70" box="[761,901,142,167]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="11" pageNumber="408">Figure 5BD</figureCitation>
). Transversal bar (TB), same length as anterior margin of anterior arc (
<figureCitation id="135D2E8DFFE0F168FD6D089FFCC1FF1D" box="[709,846,177,202]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="11" pageNumber="408">Figure 5BC</figureCitation>
). A planar, oval sclerotised plate located posterior to transversal bar. Lateral margins of this plate slightly curved dorsally (
<figureCitation id="135D2E8DFFE0F168FEA108D8FEF8FED8" box="[265,375,246,271]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="11" pageNumber="408">Figure 5C</figureCitation>
). Behind this plate, transparent, membranous folds associated with posterior diverticulum (
<figureCitation id="135D2E8DFFE0F168FE060937FD90FEE5" box="[430,543,281,306]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="11" pageNumber="408">Figure 5D</figureCitation>
). Posterior diverticulum (PD) roundish, membranous, and equal in width and length (
<figureCitation id="135D2E8DFFE0F168FDAF0912FD01FE82" box="[519,654,316,341]" captionStart="Figure 5" captionStartId="8.[160,225,1434,1457]" captionTargetBox="[163,1153,142,1401]" captionTargetId="figure-58@8.[163,1153,142,1402]" captionTargetPageId="8" captionText="Figure 5. Harpactea dilekensis sp. n. (A) Vulva, ventral view. (B) Same, dorsal view. (C) Same, nearly dorsal view. (D) Same, posterior dorsal view. Abbreviations: AA anterior margin of the anterior arch, AC anterior arc, ES epigastral scutum, PD posterior diverticulum, RS roundish structures, S spermatheca, SK spermathecal keel, SP sclerotised plate, TB transversal bar." pageId="11" pageNumber="408">Figure 5A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C6183FFE0F168FF0809BEFE37FDB0" pageId="11" pageNumber="408" type="distribution">
<paragraph id="8BD93208FFE0F168FF0809BEFEA3FE7D" blockId="11.[160,524,400,461]" box="[160,300,400,426]" pageId="11" pageNumber="408">
<emphasis id="B912EE1AFFE0F168FF0809BEFEA3FE7D" bold="true" box="[160,300,400,426]" italics="true" pageId="11" pageNumber="408">Distribution</emphasis>
</paragraph>
<paragraph id="8BD93208FFE0F168FF08099AFD83FE1A" blockId="11.[160,524,400,461]" box="[160,524,436,461]" pageId="11" pageNumber="408">
Known only from
<typeStatus id="54DD8CAAFFE0F168FEC2099AFE13FE1A" box="[362,412,436,461]" pageId="11" pageNumber="408">type</typeStatus>
localities.
</paragraph>
<paragraph id="8BD93208FFE0F168FF080A26FF76FDF5" blockId="11.[160,1155,520,615]" box="[160,249,520,546]" pageId="11" pageNumber="408">
<emphasis id="B912EE1AFFE0F168FF080A26FF76FDF5" bold="true" box="[160,249,520,546]" italics="true" pageId="11" pageNumber="408">Habitat</emphasis>
</paragraph>
<paragraph id="8BD93208FFE0F168FF080A02FE37FDB0" blockId="11.[160,1155,520,615]" pageId="11" pageNumber="408">The collection locality consists of red pine trees and oak bushes. It is a cool and humid area with thick leaf litter.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,84 +1,85 @@
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<taxonomicName id="4C7E4D757F4FFD3209C2F98687FAF9CC" authority="J. Maza &amp; R. G. Maza, 1989" authorityName="J. Maza &amp; R. G. Maza" authorityYear="1989" box="[151,762,1585,1610]" class="Insecta" family="Nymphalidae" genus="Opoptera" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="subSpecies" species="staudingeri" subSpecies="mexicana">
@ -213,11 +214,11 @@ m. 16-VIII-86 Col. J. de la Maza/
J. de la Maza E.
</taxonomicName>
/ (
<figureCitation id="13452A737F4FFD320886F87E8728F863" box="[467,552,1994,2021]" captionStart="FIGURE 34" captionStartId="43.[151,250,1217,1241]" captionTargetBox="[223,1363,191,1193]" captionTargetId="figure-20@43.[223,1363,181,1193]" captionTargetPageId="43" captionText="FIGURE 34. Opoptera mexicana J. Maza &amp; R. G. Maza, 1989 stat. nov.: Female holotype of Opoptera staudingeri mexicana J. Maza &amp; R. G. Maza, 1989: a. dorsal view, b. ventral view. Scale bar: 1 cm. Photos: J. de la Maza &amp; R. G. de la Maza." pageId="42" pageNumber="43">Fig. 34</figureCitation>
<figureCitation id="13452A737F4FFD320886F87E8728F863" box="[467,552,1994,2021]" captionStart="FIGURE 34" captionStartId="43.[151,250,1217,1241]" captionTargetBox="[223,1363,191,1193]" captionTargetId="figure-20@43.[223,1363,181,1193]" captionTargetPageId="43" captionText="FIGURE 34. Opoptera mexicana J. Maza &amp; R. G. Maza, 1989 stat. nov.: Female holotype of Opoptera staudingeri mexicana J. Maza &amp; R. G. Maza, 1989: a. dorsal view, b. ventral view. Scale bar: 1 cm. Photos: J. de la Maza &amp; R. G. de la Maza." figureDoi="http://doi.org/10.5281/zenodo.14797400" httpUri="https://zenodo.org/record/14797400/files/figure.png" pageId="42" pageNumber="43">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF01667E7F4EFD3309C2FB75805AFB7B" pageId="43" pageNumber="44" startId="43.[151,250,1217,1241]" targetBox="[223,1363,191,1193]" targetPageId="43" targetType="figure">
<caption id="DF01667E7F4EFD3309C2FB75805AFB7B" ID-DOI="http://doi.org/10.5281/zenodo.14797400" ID-Zenodo-Dep="14797400" httpUri="https://zenodo.org/record/14797400/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1217,1241]" targetBox="[223,1363,191,1193]" targetPageId="43" targetType="figure">
<paragraph id="8BC136F67F4EFD3309C2FB75805AFB7B" blockId="43.[151,1436,1217,1278]" pageId="43" pageNumber="44">
<emphasis id="B90AEAE47F4EFD3309C2FB758420FB5F" bold="true" box="[151,288,1217,1241]" pageId="43" pageNumber="44">FIGURE 34.</emphasis>
<taxonomicName id="4C7E4D757F4EFD330873FB76861AFB5C" authority="J. Maza &amp; R. G. Maza, 1989" authorityName="J. Maza &amp; R. G. Maza" authorityYear="1989" baseAuthorityName="J. Maza &amp; R. G. Maza" baseAuthorityYear="1989" box="[294,794,1218,1242]" class="Insecta" family="Nymphalidae" genus="Opoptera" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="mexicana" status="stat. nov.">
@ -251,7 +252,7 @@ by not having the orange band on DFW and by having a submarginal band between M
CuA
<subScript id="17FA34B37F4EFD3308C8FA1E84A5FA3E" attach="left" box="[413,421,1450,1464]" fontSize="6" pageId="43" pageNumber="44">2</subScript>
on VFW (
<figureCitation id="13452A737F4EFD330B49FA2C8770FA34" box="[540,624,1432,1458]" captionStart="FIGURE 34" captionStartId="43.[151,250,1217,1241]" captionTargetBox="[223,1363,191,1193]" captionTargetId="figure-20@43.[223,1363,181,1193]" captionTargetPageId="43" captionText="FIGURE 34. Opoptera mexicana J. Maza &amp; R. G. Maza, 1989 stat. nov.: Female holotype of Opoptera staudingeri mexicana J. Maza &amp; R. G. Maza, 1989: a. dorsal view, b. ventral view. Scale bar: 1 cm. Photos: J. de la Maza &amp; R. G. de la Maza." pageId="43" pageNumber="44">Fig. 34</figureCitation>
<figureCitation id="13452A737F4EFD330B49FA2C8770FA34" box="[540,624,1432,1458]" captionStart="FIGURE 34" captionStartId="43.[151,250,1217,1241]" captionTargetBox="[223,1363,191,1193]" captionTargetId="figure-20@43.[223,1363,181,1193]" captionTargetPageId="43" captionText="FIGURE 34. Opoptera mexicana J. Maza &amp; R. G. Maza, 1989 stat. nov.: Female holotype of Opoptera staudingeri mexicana J. Maza &amp; R. G. Maza, 1989: a. dorsal view, b. ventral view. Scale bar: 1 cm. Photos: J. de la Maza &amp; R. G. de la Maza." figureDoi="http://doi.org/10.5281/zenodo.14797400" httpUri="https://zenodo.org/record/14797400/files/figure.png" pageId="43" pageNumber="44">Fig. 34</figureCitation>
).
</paragraph>
<paragraph id="8BC136F67F4EFD330992FA088725FA50" blockId="43.[151,1437,1359,1962]" box="[199,549,1467,1494]" pageId="43" pageNumber="44">
@ -315,7 +316,7 @@ Geographic distribution.
(
<bibRefCitation id="EFEF4B077F4EFD330D87F8B48057F89C" author="Wolfe, K. V." box="[1234,1367,1792,1818]" pageId="43" pageNumber="44" pagination="28 - 29" refId="ref40516" refString="Wolfe, K. V. (2017) Probable last-instar larvae of Opoptera staudingeri mexicana (Nymphalidae, Satyrinae, Brassolini). News of The Lepidopterists' Society, 59 (1), 28-29." type="journal article" year="2017">Wolfe 2017</bibRefCitation>
) (
<figureCitation id="13452A737F4EFD330C3AF8B485B7F8B8" captionStart="FIGURE 60" captionStartId="69.[151,250,1107,1131]" captionTargetBox="[151,1436,181,1084]" captionTargetId="figure-20@69.[151,1436,181,1084]" captionTargetPageId="69" captionText="FIGURE 60. Geographical distribution of Opoptera species. The asterisk (*) inside the localities symbols indicates that the distribution data were taken originate from the literature." pageId="43" pageNumber="44">Fig. 60</figureCitation>
<figureCitation id="13452A737F4EFD330C3AF8B485B7F8B8" captionStart="FIGURE 60" captionStartId="69.[151,250,1107,1131]" captionTargetBox="[151,1436,181,1084]" captionTargetId="figure-20@69.[151,1436,181,1084]" captionTargetPageId="69" captionText="FIGURE 60. Geographical distribution of Opoptera species. The asterisk (*) inside the localities symbols indicates that the distribution data were taken originate from the literature." figureDoi="http://doi.org/10.5281/zenodo.14797467" httpUri="https://zenodo.org/record/14797467/files/figure.png" pageId="43" pageNumber="44">Fig. 60</figureCitation>
). Elevation of
<quantity id="4C869B137F4EFD33080EF89084B9F8B8" box="[347,441,1828,1854]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="43" pageNumber="44" unit="m" value="1800.0">1,800 m</quantity>
, according to the single elevation record.

View file

@ -1,51 +1,52 @@
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View file

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<figureCitation id="1360098EFFA3560BFF22688E38CAF83B" box="[159,241,1926,1952]" captionStart="FIGURES 1316" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[155,1435,188,1124]" captionTargetId="figure-34@12.[151,1436,181,1125]" captionTargetPageId="12" captionText="FIGURES 1316. Habitus of: (13) Ophiomyia arabica (Deeming, 2006), Male (SAUDI ARABIA:Jazan, Abu Aresh, Al- Mahdag Village, 520.vi.2011, Malaise trap, H.A. Dawah); (14). Ophiomyia beckeri (Hendel, 1923), Male (SAUDI ARABIA: Asir, Abha, Hay Al-Nusub (Abha Farm Centre), 3.vi.2001, Malaise trap, H.A. Dawah; (1516) Ophiomyia centrosematis (de Meijere, 1940): (15) Female, lateral view (SAUDI ARABIA: Asir, Abha, Madinate Al-Ameer Sultan, 6.xi.2003, Malaise trap, H.A. Dawah); (16) Male, lateral view (SAUDI ARABIA: Asir, Abha, Madinate Al-Ameer Sultan, 627.viii.2013, Malaise trap, H.A. Dawah)." figureDoi="http://doi.org/10.5281/zenodo.14797555" httpUri="https://zenodo.org/record/14797555/files/figure.png" pageId="10" pageNumber="11">Fig. 14</figureCitation>
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<figureCitation id="1360098EFFAF5607FF226D633938FD1E" box="[159,259,619,645]" captionStart="FIGURES 78" captionStartId="6.[151,264,1820,1844]" captionTargetBox="[158,1435,1254,1795]" captionTargetId="figure-337@6.[151,1436,1203,1796]" captionTargetPageId="6" captionText="FIGURES 78. Habitus of: Agromyza panici de Meijere, 1934: (7) Male, lateral view (SAUDI ARABIA: Asir, Keratha, Al- Ethrebany Fruit Farm, 125.v.2013, Malaise trap, H.A. Dawah); (8) Female (YEMEN: Sug Bani Manour, 28.vii26.ix.201, A. van Harten)." figureDoi="http://doi.org/10.5281/zenodo.14797543" httpUri="https://zenodo.org/record/14797543/files/figure.png" pageId="6" pageNumber="7">Figs 78</figureCitation>
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<figureCitation id="1360098EFFB3561BFF226D4F38CAFDFA" box="[159,241,583,609]" captionStart="FIGURES 3233" captionStartId="26.[151,264,1871,1895]" captionTargetBox="[156,1434,1138,1847]" captionTargetId="figure-317@26.[151,1436,1131,1848]" captionTargetPageId="26" captionText="FIGURES 3233. Habitus of: (32) Pseudonapomyza acanthacearum Spencer, 1959, Male, lateral view (SAUDI ARABIA: Jazan, Fifa (Al-Tatweer Centre), 526.i.2014, Malaise trap, H.A. Dawah); (33) Pseudonapomyza asiatica Spencer, 1961, Female (GAMBIA, Central River Division, Sokuto, Naini District, sorghum field, 24.viii.1997, D.J. Mann)." pageId="26" pageNumber="27">Fig. 33</figureCitation>
<figureCitation id="1360098EFFB3561BFF226D4F38CAFDFA" box="[159,241,583,609]" captionStart="FIGURES 3233" captionStartId="26.[151,264,1871,1895]" captionTargetBox="[156,1434,1138,1847]" captionTargetId="figure-317@26.[151,1436,1131,1848]" captionTargetPageId="26" captionText="FIGURES 3233. Habitus of: (32) Pseudonapomyza acanthacearum Spencer, 1959, Male, lateral view (SAUDI ARABIA: Jazan, Fifa (Al-Tatweer Centre), 526.i.2014, Malaise trap, H.A. Dawah); (33) Pseudonapomyza asiatica Spencer, 1961, Female (GAMBIA, Central River Division, Sokuto, Naini District, sorghum field, 24.viii.1997, D.J. Mann)." figureDoi="http://doi.org/10.5281/zenodo.14797578" httpUri="https://zenodo.org/record/14797578/files/figure.png" pageId="26" pageNumber="27">Fig. 33</figureCitation>
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.
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<emphasis id="B92FC919FFB3561BFF2A6847396CF8F3" bold="true" box="[151,343,1871,1896]" pageId="26" pageNumber="27">FIGURES 3233.</emphasis>
Habitus of:

View file

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@ -488,7 +489,7 @@ and
; Černý 2023). It is widespread on Pacific Islands.
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<emphasis id="B92FC919FFB4561CFF2A6B6D39BEFBE6" bold="true" box="[151,389,1125,1149]" pageId="29" pageNumber="30">FIGURES 3436. (34)</emphasis>
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Habitus of:

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leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF0FF8FFF6B2B30FA3BFD64" pageId="12" pageNumber="166" type="discussion">
<paragraph id="8BEC36C9FFF0FF8FFF6B2B30FDABFD94" blockId="12.[151,1437,259,718]" pageId="12" pageNumber="166">
<emphasis id="B927EADBFFF0FF8FFF6B2B30FE8AFE20" bold="true" box="[199,315,367,393]" pageId="12" pageNumber="166">Remarks.</emphasis>
Probably not a very common European species, which was originally described from
<collectingCountry id="F3447659FFF0FF8FFB7A2B2FFA81FE23" box="[1238,1328,368,394]" name="Norway" pageId="12" pageNumber="166">Norway</collectingCountry>
(northern
<collectingRegion id="4997F82BFFF0FF8FFF3B2BCCFEB8FE04" box="[151,265,403,429]" country="Norway" name="Finnmark" pageId="12" pageNumber="166">Finnmark</collectingRegion>
), where it inhabits sandy coastal meadows and dunes. It was also recorded in southern
<collectingCountry id="F3447659FFF0FF8FFB5C2BCBFAFAFE07" box="[1264,1355,404,430]" name="Norway" pageId="12" pageNumber="166">Norway</collectingCountry>
, where it occupies quite a different
<typeStatus id="54E8886BFFF0FF8FFE7A2BE7FDB7FE7B" box="[470,518,440,466]" pageId="12" pageNumber="166">type</typeStatus>
of habitat:
<emphasis id="B927EADBFFF0FF8FFD262BE8FEBDFE5C" italics="true" pageId="12" pageNumber="166">eroded fine-grained sediments of glacio-lacustrine origin with sparse vegetation</emphasis>
(
<bibRefCitation id="EFC24B38FFF0FF8FFEB62B84FE72FE5C" author="Fjellberg, A." box="[282,451,475,501]" pageId="12" pageNumber="166" refId="ref12541" refString="Fjellberg, A. (1998) The Collembola of Fennoscandinavia and Denmark. Part I. Poduromorpha. Fauna Entomologica Scandinavica. Vol. 35. Brill, Leiden, 184 pp. https://doi.org/10.1163/9789004273634" type="book" year="1998">Fjellberg 1998</bibRefCitation>
, p. 96). In our materials,
<taxonomicName id="4C534D4AFFF0FF8FFD732B84FCD8FE5C" box="[735,873,475,501]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="12" pageNumber="166" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B927EADBFFF0FF8FFD732B84FCD8FE5C" box="[735,873,475,501]" italics="true" pageId="12" pageNumber="166">A. echinatus</emphasis>
</taxonomicName>
was found only in floodplain communities of one of the sites of the “Privolzhskaya Lesostep” Nature Reserve, this significantly increasing not only its distribution, but also its ecological preferences.
</paragraph>
<paragraph id="8BEC36C9FFF0FF8FFF6B2818FA3BFD64" blockId="12.[151,1437,259,718]" pageId="12" pageNumber="166">
As the species has been described in detail (
<bibRefCitation id="EFC24B38FFF0FF8FFD6C2818FCDBFDC8" author="Fjellberg, A." box="[704,874,583,609]" pageId="12" pageNumber="166" pagination="35 - 41" refId="ref12462" refString="Fjellberg, A. (1988) Six new species of Collembola from north Norway (Hypogastruridae, Odontellidae, Onychiuridae, Isotomidae). Fauna Norvegica, Series B, 35 (1), 35-41." type="journal article" year="1988">Fjellberg 1988</bibRefCitation>
,
<bibRefCitation id="EFC24B38FFF0FF8FFCDB2818FC02FDC8" author="Fjellberg, A." box="[887,947,583,609]" pageId="12" pageNumber="166" refId="ref12541" refString="Fjellberg, A. (1998) The Collembola of Fennoscandinavia and Denmark. Part I. Poduromorpha. Fauna Entomologica Scandinavica. Vol. 35. Brill, Leiden, 184 pp. https://doi.org/10.1163/9789004273634" type="book" year="1998">1998</bibRefCitation>
), only a few minor additions can be made. Thus, our specimens from the Volga region always have only two prelabral setae (medial pair absent), lacking the basal pair of setae on the manubrial field and having only 3+3 setae on its main part (
<figureCitation id="13682A4CFFF0FF8FFB9928D0FB36FD03" box="[1077,1159,655,682]" captionStart="FIGURES 1015" captionStartId="3.[151,264,1349,1373]" captionTargetBox="[160,1424,185,1319]" captionTargetId="figure-19@3.[151,1436,181,1325]" captionTargetPageId="3" captionText="FIGURES 1015. Axenyllodes ghilarovi (1012), A. bayeri (13), A. nematodes (14) and A. echinatus (15): 10, dorsal chaetotaxy of head and thorax, holotype; 11, abdominal tip, holotype; 1215, chaetotaxy of manubrial field: m—setae on main part, b— basal setae and bl—basolateral setae. Scales: 1011, 0.1 mm, 1215, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797990" httpUri="https://zenodo.org/record/14797990/files/figure.png" pageId="12" pageNumber="166">Fig. 15</figureCitation>
). The latter character, i.e. the chaetotaxy of the manubrial field was confirmed by A. Fjellberg for Scandinavian populations (pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,297 @@
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<mods:title id="C4A5E6709556276F61B03B601FF9C883">Two new and two little-known species of the genus Axenyllodes Stach, 1949 from Russia and Kazakhstan</mods:title>
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<taxonomicName id="4C534D4AFFF5FF8AFF3B2CBFFE29F952" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[151,408,1760,1787]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFF3B2CBFFE29F952" bold="true" box="[151,408,1760,1787]" italics="true" pageId="9" pageNumber="163">Axenyllodes potapovi</emphasis>
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<paragraph id="8BEC36C9FFF5FF8AFF3B2D5AFEA4F8B6" blockId="9.[151,509,1760,1823]" box="[151,277,1797,1823]" pageId="9" pageNumber="163">
<figureCitation id="13682A4CFFF5FF8AFF3B2D5AFEA4F8B6" box="[151,277,1797,1823]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="9" pageNumber="163">Figs 3339</figureCitation>
</paragraph>
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<paragraph id="8BEC36C9FFF5FF8AFF3B2D12FE58F806" blockId="9.[151,1437,1869,2039]" pageId="9" pageNumber="163">
<emphasis id="B927EADBFFF5FF8AFF3B2D12FE8EF8CE" bold="true" box="[151,319,1869,1895]" pageId="9" pageNumber="163">Type material</emphasis>
.
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<typeStatus id="54E8886BFFF5FF8AFEE22D12FE08F8CE" box="[334,441,1869,1895]" pageId="9" pageNumber="163" type="holotype">Holotype</typeStatus>
, juvenile,
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,
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,
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, north of
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,
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42.6090
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N
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47.7898
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E
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, coast of
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, ~
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alt., sand, zone of weak vegetation (flotation),
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.
<collectorName id="26A6531FFFF5FF8AFB6A2D2EFAC1F822" box="[1222,1392,1905,1931]" pageId="9" pageNumber="163">N. Kuznetsova</collectorName>
,
<collectorName id="26A6531FFFF5FF8AFAD02D2EFF42F806" pageId="9" pageNumber="163">M. Potapov</collectorName>
&amp;
<collectorName id="26A6531FFFF5FF8AFEB92DCAFE09F806" box="[277,440,1941,1967]" pageId="9" pageNumber="163">A. Kremenitsa</collectorName>
leg.
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<emphasis id="B927EADBFFF5FF8AFF6B2DE6FE8BF87A" bold="true" box="[199,314,1977,2003]" pageId="9" pageNumber="163">Diagnosis</emphasis>
. A blind species of the genus with the complete absence of dark pigment, characterized by: a more or less distinct pattern of cuticular granulation on thorax and in the anterior part of the abdomen; the presence of 7 clearly differentiated dorsal sensilla and a rather large microsensillum on Ant. IV; star-shaped PAO with three lobes, not sunken into the cuticle; 2+2 postlabial setae; Th. IIIII with setae m1 present and lateral microsensilla absent, lateral sensilla on the thorax slightly thickened; the basic
<typeStatus id="54E8886BFFF6FF89FCBF2A80FCF2FF50" box="[787,835,223,249]" pageId="10" pageNumber="164">type</typeStatus>
of VT, tenaculum and furca; manubrial field with 8+8 setae; the tibiotarsi with 777 subequal setae and 7 setae on each femur; and the absence of
<collectingCountry id="F3447659FFF6FF89FB042B5CFB7EFEB4" box="[1192,1231,259,285]" name="American Samoa" pageId="10" pageNumber="164">AS</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF6FF89FF6B2B79FC94FAFC" pageId="10" pageNumber="164" type="description">
<paragraph id="8BEC36C9FFF6FF89FF6B2B79FEFFFE78" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
<emphasis id="B927EADBFFF6FF89FF6B2B79FEE8FEE9" bold="true" box="[199,345,294,320]" pageId="10" pageNumber="164">Description.</emphasis>
Length (without antennae) of the only known specimen,
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. Habitus typical of the genus: body oblong, appendices short. Colour white without any traces of dark pigment. Tegument granulation rather coarse, secondary granules usually spinous at tip. Th. IIAbd. II with rather long and narrow longitudinal fields of finer granulations located symmetrically with respect to medial line, only traces of such fields developed on Abd. IIIIV (
<figureCitation id="13682A4CFFF6FF89FF422BE8FEF3FE78" box="[238,322,439,465]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2B84FD64FD2C" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
Antennae about as long as head. Ant. IV with a large simple vesicle apically, with 7 clearly differentiated sensilla (four dorso-internal [S1S4] and three dorso-external [S7S9, S8 longer and thinner, S9 located higher than usual], ms about 0.4 times as long as nearest sensillum, subapical organite present, ventral side with few ordinary setae (
<figureCitation id="13682A4CFFF6FF89FF712818FEEEFDC8" box="[221,351,583,609]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Figs 3334</figureCitation>
). Antennal organ of Ant. III typical, inner sensilla small, sgv &amp; sgd larger and subequal to ms. Ant. IIII with 7, 10 and 17 ordinary setae, respectively.
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B28D0FC1AFCF4" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
Ocelli absent. PAO star-shaped, with three lobes, located superficially in a small depression (
<figureCitation id="13682A4CFFF6FF89FB4928D0FA88FD00" box="[1253,1337,654,681]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 35</figureCitation>
). Buccal cone short, typical of the genus (
<figureCitation id="13682A4CFFF6FF89FDA028ECFDD3FD64" box="[524,610,691,717]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 37</figureCitation>
). Labrum with 8 labral setae, medial pair of which in anterior row much stronger and with their tips curved laterally, as usual; prelabral setae invisible. Maxillary palp simple. Labium with several small apical spinules and three common setae; basomedial and basolateral fields of labium with 4+3 setae, respectively. Maxilla typical of the family with a strong and triangular head, mandibles invisible. Ventral side of head in
<typeStatus id="54E8886BFFF6FF89FF5C291CFEE3FCF4" box="[240,338,835,861]" pageId="10" pageNumber="164" type="holotype">holotype</typeStatus>
with 2+2 postlabial setae along ventral line (
<figureCitation id="13682A4CFFF6FF89FCE7291CFC2EFCF4" box="[843,927,835,861]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2938FD49FBB8" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
Ordinary setae on dorsal side of body short, smooth and acuminate, those on last abdominal terga only slightly longer, dorsal sensilla undifferentiated, but lateral ones on Th. IIIII clearly thickened (
<figureCitation id="13682A4CFFF6FF89FBFA29D4FB18FC0C" box="[1110,1193,907,933]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 36</figureCitation>
). Main characteristics of dorsal chaetotaxy: setae m1 present on Th. IIIII and absent from Abd. IV; Th. IIIII with only three setae in a-row (a1, a3 and a5), three m-setae (including S=m6), and five setae in p-row as usual, lateral microsensilla absent; Abd. IIII with four setae in a-row (a1, a2, a3 and a5).
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2E44FF57FB40" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
Thoracic sterna without setae. Ventral tube with 3+3 distal setae, one seta also present each side of VT on sternum of Abd. I, Abd. IIIII with 3 and 5 ventral setae each side, respectively (
<figureCitation id="13682A4CFFF6FF89FBA22E60FBD0FBF0" box="[1038,1121,1087,1113]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 38</figureCitation>
). Tenaculum with 2+2 teeth, as usual. Furca small (
<figureCitation id="13682A4CFFF6FF89FE342E3CFE56FBD4" box="[408,487,1123,1149]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig 38</figureCitation>
), manubrial field with 4+4 setae on main part, 1+1 basal and 3+3 basolateral setae (8+8 setae totally); dorsal side of dens with two setae and rather fine granulation. Mucro about as long as or slightly longer than dens, strongly hooked, lateral lamella present. Anal opening located terminally; anal spines completely absent.
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2EACFC94FAFC" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
Chaetotaxy of Legs IIII as following: 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 4, 4 setae on coxae, 5, 5, 45 on trochanters, 7, 7, 7 setae on femora, and seven subequal setae also present on each tibiotarsi. Unguis toothless, unguiculus invisible (
<figureCitation id="13682A4CFFF6FF89FD682F64FCA9FAFC" box="[708,792,1339,1365]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF6FF89FF6B2F00FBBFF874" pageId="10" pageNumber="164" type="etymology">
<paragraph id="8BEC36C9FFF6FF89FF6B2F00FD92FA34" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
<emphasis id="B927EADBFFF6FF89FF6B2F00FEF9FAD0" bold="true" box="[199,328,1375,1401]" pageId="10" pageNumber="164">Etymology</emphasis>
. The new species is to honour our friend and colleague, Mikhail B. Potapov, whose contribution to springtail taxonomy is outstanding.
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2FF9FDDFF9F8" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
<emphasis id="B927EADBFFF6FF89FF6B2FF9FE84FA69" bold="true" box="[199,309,1446,1472]" pageId="10" pageNumber="164">Affinities</emphasis>
. Currently, only four completely blind species of the genus which lack anal spines are known:
<taxonomicName id="4C534D4AFFF6FF89FA902FF8FE8DFA4C" authority="Thibaud, 1995" authorityName="Thibaud" authorityYear="1995" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="clevai">
<emphasis id="B927EADBFFF6FF89FA902FF8FA2DFA68" box="[1340,1436,1447,1473]" italics="true" pageId="10" pageNumber="164">A. clevai</emphasis>
<bibRefCitation id="EFC24B38FFF6FF89FF3B2F94FE8DFA4C" author="Thibaud, J. - M." box="[151,316,1482,1509]" pageId="10" pageNumber="164" pagination="31 - 37" refId="ref13962" refString="Thibaud, J. - M. (1995) Recherche sur la faune interstitielle aerienne des sables fins: les Collemboles littoraux de l'Herault. Annales de la Societe Entomologique de France, 31, 31-37. https://doi.org/10.1080/21686351.1995.12277718" type="journal article" year="1995">Thibaud, 1995</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F3447659FFF6FF89FEE02F94FE2FFA4C" box="[332,414,1483,1509]" name="France" pageId="10" pageNumber="164">France</collectingCountry>
),
<taxonomicName id="4C534D4AFFF6FF89FE1F2F94FD4EFA4C" authority="Fjellberg, 1995" authorityName="Fjellberg" authorityYear="1995" box="[435,767,1482,1509]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="nematodes">
<emphasis id="B927EADBFFF6FF89FE1F2F94FDFAFA4C" box="[435,587,1483,1509]" italics="true" pageId="10" pageNumber="164">A. nematodes</emphasis>
<bibRefCitation id="EFC24B38FFF6FF89FDFF2F94FD4EFA4C" author="Fjellberg, A." box="[595,767,1482,1509]" pageId="10" pageNumber="164" pagination="153 - 158" refId="ref12502" refString="Fjellberg, A. (1995) Collembola of the Canary Islands. II. Family Odontellidae. Entomologica Scandinavica, 26, 153-158. https://doi.org/10.1163/187631295X00152" type="journal article" year="1995">Fjellberg, 1995</bibRefCitation>
</taxonomicName>
(Canary Islands),
<taxonomicName id="4C534D4AFFF6FF89FC7F2F94FBD1FA4D" box="[979,1120,1483,1509]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B927EADBFFF6FF89FC7F2F94FBD1FA4D" box="[979,1120,1483,1509]" italics="true" pageId="10" pageNumber="164">A. japonicus</emphasis>
</taxonomicName>
(
<collectingCountry id="F3447659FFF6FF89FBDC2F94FB02FA4C" box="[1136,1203,1483,1509]" name="Japan" pageId="10" pageNumber="164">Japan</collectingCountry>
) and
<taxonomicName id="4C534D4AFFF6FF89FB582F94FA2DFA4D" authorityName="Thibaud" authorityYear="2011" box="[1268,1436,1483,1509]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="comoriensis">
<emphasis id="B927EADBFFF6FF89FB582F94FA2DFA4D" box="[1268,1436,1483,1509]" italics="true" pageId="10" pageNumber="164">A. comoriensis</emphasis>
</taxonomicName>
(
<collectingCountry id="F3447659FFF6FF89FF332FB0FEE9F9A0" box="[159,344,1519,1545]" name="Comoros" pageId="10" pageNumber="164">Comoro Islands</collectingCountry>
). In addition,
<taxonomicName id="4C534D4AFFF6FF89FE582FB0FB91F9A0" authority="Vazquez &amp; Palacios-Vargas, 1989" authorityName="Vazquez &amp; Palacios-Vargas" authorityYear="1989" box="[500,1056,1519,1545]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B927EADBFFF6FF89FE582FB0FD26F9A1" box="[500,663,1519,1545]" italics="true" pageId="10" pageNumber="164">A. americanus</emphasis>
<bibRefCitation id="EFC24B38FFF6FF89FD312FB0FB91F9A0" author="Vazquez, M. M. &amp; Palacios-Vargas, J. G." box="[669,1056,1519,1545]" pageId="10" pageNumber="164" pagination="37 - 41" refId="ref14178" refString="Vazquez, M. M. &amp; Palacios-Vargas, J. G. (1989) Une nouvelle espece mexicaine de Axenyllodes et biogeographie de la lignee Xenyllodienne (Collembola, Odontellidae). Revue francaise d'Entomologie, New Series, 11 (1), 37-41." type="journal article" year="1989">Vázquez &amp; Palacios-Vargas, 1989</bibRefCitation>
</taxonomicName>
, the only South American species of the genus, sometimes also has no anal spines, but it also lacks a mucro (a unique feature for the genus) and is hardly comparable to
<taxonomicName id="4C534D4AFFF6FF89FE202C68FDBCF9F9" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[396,525,1591,1617]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF6FF89FE202C68FE2CF9F8" box="[396,413,1591,1617]" italics="true" pageId="10" pageNumber="164">A</emphasis>
.
<emphasis id="B927EADBFFF6FF89FE072C68FDBCF9F9" box="[427,525,1591,1616]" italics="true" pageId="10" pageNumber="164">potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF6FF89FDB82C68FDDFF9F8" bold="true" box="[532,622,1591,1617]" pageId="10" pageNumber="164">
<taxonomicNameLabel id="A21457A0FFF6FF89FDB82C68FDDFF9F8" box="[532,622,1591,1617]" pageId="10" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2C04FE0EF8D8" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
Three of the four species listed above are fairly easily distinguished from
<taxonomicName id="4C534D4AFFF6FF89FBBD2C04FB25F9DD" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[1041,1172,1627,1653]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF6FF89FBBD2C04FB93F9DC" box="[1041,1058,1627,1653]" italics="true" pageId="10" pageNumber="164">A</emphasis>
.
<emphasis id="B927EADBFFF6FF89FB9E2C04FB25F9DD" box="[1074,1172,1627,1652]" italics="true" pageId="10" pageNumber="164">potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF6FF89FB302C04FB46F9DC" bold="true" box="[1180,1271,1627,1653]" pageId="10" pageNumber="164">
<taxonomicNameLabel id="A21457A0FFF6FF89FB302C04FB46F9DC" box="[1180,1271,1627,1653]" pageId="10" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
because all of them have lateral microsensilla on Th. IIIII (for other differences, see
<tableCitation id="C6D10372FFF6FF89FC092C20FC47F930" box="[933,1014,1663,1689]" captionStart="TABLE 1" captionStartId="13.[151,243,474,500]" captionText="TABLE 1. Main diagnostic characters of the known species of the genus Axenyllodes Stach, 1949." pageId="10" pageNumber="164">Table 1</tableCitation>
). The only exception is
<taxonomicName id="4C534D4AFFF6FF89FAAC2C20FA24F930" authorityName="Fjellberg" authorityYear="1995" box="[1280,1429,1663,1689]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="nematodes">
<emphasis id="B927EADBFFF6FF89FAAC2C20FA24F930" box="[1280,1429,1663,1689]" italics="true" pageId="10" pageNumber="164">A. nematodes</emphasis>
</taxonomicName>
, which, however, unlike
<taxonomicName id="4C534D4AFFF6FF89FE002CFCFD81F915" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[428,560,1699,1725]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF6FF89FE002CFCFE0CF914" box="[428,445,1699,1725]" italics="true" pageId="10" pageNumber="164">A</emphasis>
.
<emphasis id="B927EADBFFF6FF89FE622CFCFD81F915" box="[462,560,1699,1724]" italics="true" pageId="10" pageNumber="164">potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF6FF89FD962CFCFD27F914" bold="true" box="[570,662,1699,1725]" pageId="10" pageNumber="164">
<taxonomicNameLabel id="A21457A0FFF6FF89FD962CFCFD27F914" box="[570,662,1699,1725]" pageId="10" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is characterized by the presence 6 and 6 setae on each femur and tibiotarsus, respectively (
<emphasis id="B927EADBFFF6FF89FE182C98FE7AF949" box="[436,459,1735,1760]" italics="true" pageId="10" pageNumber="164">vs</emphasis>
7 and 7 setae in
<taxonomicName id="4C534D4AFFF6FF89FD3C2C98FCA3F949" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[656,786,1735,1761]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF6FF89FD3C2C98FD10F948" box="[656,673,1735,1761]" italics="true" pageId="10" pageNumber="164">A</emphasis>
.
<emphasis id="B927EADBFFF6FF89FD1C2C98FCA3F949" box="[688,786,1735,1760]" italics="true" pageId="10" pageNumber="164">potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF6FF89FCB72C98FCC7F948" bold="true" box="[795,886,1735,1761]" pageId="10" pageNumber="164">
<taxonomicNameLabel id="A21457A0FFF6FF89FCB72C98FCC7F948" box="[795,886,1735,1761]" pageId="10" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and 7+7 setae on the manubrial field, i.e. basal pair of setae absent (
<figureCitation id="13682A4CFFF6FF89FE2D2CB4FE62F8AC" box="[385,467,1771,1797]" captionStart="FIGURES 1015" captionStartId="3.[151,264,1349,1373]" captionTargetBox="[160,1424,185,1319]" captionTargetId="figure-19@3.[151,1436,181,1325]" captionTargetPageId="3" captionText="FIGURES 1015. Axenyllodes ghilarovi (1012), A. bayeri (13), A. nematodes (14) and A. echinatus (15): 10, dorsal chaetotaxy of head and thorax, holotype; 11, abdominal tip, holotype; 1215, chaetotaxy of manubrial field: m—setae on main part, b— basal setae and bl—basolateral setae. Scales: 1011, 0.1 mm, 1215, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797990" httpUri="https://zenodo.org/record/14797990/files/figure.png" pageId="10" pageNumber="164">Fig. 14</figureCitation>
) (
<emphasis id="B927EADBFFF6FF89FE402CB4FDB2F8AD" box="[492,515,1771,1796]" italics="true" pageId="10" pageNumber="164">vs</emphasis>
8+8 setae including a basal pair in
<taxonomicName id="4C534D4AFFF6FF89FC3A2CB4FBA9F8AD" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[918,1048,1771,1797]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF6FF89FC3A2CB4FC16F8AC" box="[918,935,1771,1797]" italics="true" pageId="10" pageNumber="164">A</emphasis>
.
<emphasis id="B927EADBFFF6FF89FC1A2CB4FBA9F8AD" box="[950,1048,1771,1796]" italics="true" pageId="10" pageNumber="164">potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF6FF89FBB32CB4FBCBF8AC" bold="true" box="[1055,1146,1771,1797]" pageId="10" pageNumber="164">
<taxonomicNameLabel id="A21457A0FFF6FF89FBB32CB4FBCBF8AC" box="[1055,1146,1771,1797]" pageId="10" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<figureCitation id="13682A4CFFF6FF89FB242CB4FB6DF8AC" box="[1160,1244,1771,1797]" captionStart="FIGURES 3339" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[173,1414,195,1905]" captionTargetId="figure-19@11.[159,1428,181,1919]" captionTargetPageId="11" captionText="FIGURES 3339. Axenyllodes potapovi sp. nov.: 33, chaetotaxy of antenna, ventral view; 34, ibid., dorsal view; 35, PAO; 36, cuticular granulation and dorsal chaetotaxy; 37, ventral chaetotaxy of head; 38, ventral chaetotaxy of abdomen; 39, chaetotaxy of leg II. Scales: 3334, 3638, 0.05 mm, 35, 39, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14798000" httpUri="https://zenodo.org/record/14798000/files/figure.png" pageId="10" pageNumber="164">Fig. 38</figureCitation>
). In other words, the
<typeStatus id="54E8886BFFF6FF89FF6C2D50FE93F880" box="[192,290,1807,1833]" pageId="10" pageNumber="164" type="holotype">holotype</typeStatus>
of the new species is clearly characterized by a larger number of setae on the legs and in the manubrial region than
<taxonomicName id="4C534D4AFFF6FF89FEB22D6CFE04F8E4" authorityName="Fjellberg" authorityYear="1995" box="[286,437,1843,1869]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="nematodes">
<emphasis id="B927EADBFFF6FF89FEB22D6CFE04F8E4" box="[286,437,1843,1869]" italics="true" pageId="10" pageNumber="164">A. nematodes</emphasis>
</taxonomicName>
, not fewer, as could be expected based on its juvenile state. In our opinion, this supports its status as a new species.
</paragraph>
<paragraph id="8BEC36C9FFF6FF89FF6B2D24FBBFF874" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="164">
There is one more character that could probably be used to distinguish the two species (3+3 postlabial setae ventrally on the head in
<taxonomicName id="4C534D4AFFF6FF89FE1C2DC0FDF8F810" authorityName="Fjellberg" authorityYear="1995" box="[432,585,1951,1977]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="nematodes">
<emphasis id="B927EADBFFF6FF89FE1C2DC0FDF8F810" box="[432,585,1951,1977]" italics="true" pageId="10" pageNumber="164">A. nematodes</emphasis>
</taxonomicName>
,
<emphasis id="B927EADBFFF6FF89FDF52DC0FDC1F811" box="[601,624,1951,1976]" italics="true" pageId="10" pageNumber="164">vs</emphasis>
2+2 such setae in
<taxonomicName id="4C534D4AFFF6FF89FCE62DC0FC7CF811" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[842,973,1951,1977]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="10" pageNumber="164" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF6FF89FCE62DC0FC7CF811" box="[842,973,1951,1977]" italics="true" pageId="10" pageNumber="164">A. potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF6FF89FC7A2DC0FB80F810" bold="true" box="[982,1073,1951,1977]" pageId="10" pageNumber="164">
<taxonomicNameLabel id="A21457A0FFF6FF89FC7A2DC0FB80F810" box="[982,1073,1951,1977]" pageId="10" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), but unfortunately this may be related to the immature status of the only known specimen of the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,432 @@
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<mods:title id="CF319AEAC4F0382BE725B58EC697DDD2">Two new and two little-known species of the genus Axenyllodes Stach, 1949 from Russia and Kazakhstan</mods:title>
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<mods:affiliation id="AC11B60136CAEAC7A7B0C41DD8A8A775">State Natural Reserve « Privolzhskaya Lesostep », Penza 440031, Russia.</mods:affiliation>
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<subSubSection id="C3496542FFF8FF87FF3B2F95FDD5FA4D" box="[151,612,1482,1509]" pageId="4" pageNumber="158" type="nomenclature">
<paragraph id="8BEC36C9FFF8FF87FF3B2F95FDD5FA4D" blockId="4.[151,652,1482,1575]" box="[151,612,1482,1509]" pageId="4" pageNumber="158">
<heading id="D0A481A5FFF8FF87FF3B2F95FDD5FA4D" bold="true" box="[151,612,1482,1509]" fontSize="11" level="1" pageId="4" pageNumber="158" reason="1">
<taxonomicName id="4C534D4AFFF8FF87FF3B2F95FDD5FA4D" ID-CoL="K3LC" authority="(Kseneman, 1935)" baseAuthorityName="Kseneman" baseAuthorityYear="1935" box="[151,612,1482,1509]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="158" phylum="Arthropoda" rank="species" species="bayeri">
<emphasis id="B927EADBFFF8FF87FF3B2F95FDD5FA4D" bold="true" box="[151,612,1482,1509]" pageId="4" pageNumber="158">
<emphasis id="B927EADBFFF8FF87FF3B2F95FECDFA4C" bold="true" box="[151,380,1482,1509]" italics="true" pageId="4" pageNumber="158">Axenyllodes bayeri</emphasis>
(
<bibRefCitation id="EFC24B38FFF8FF87FE272F95FDEDFA4D" author="Kseneman, M." box="[395,604,1482,1508]" pageId="4" pageNumber="158" pagination="39 - 65" refId="ref12941" refString="Kseneman, M. (1935). Apterygota z uzemi rybniku Lednickych. II. Easopis narodniho Musea, oddil pOiro-dovedny, Praha, 109, 39-65." type="journal article" year="1935">Kseneman, 1935</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF8FF87FF3B2FB4FE8FF98E" pageId="4" pageNumber="158" type="reference_group">
<paragraph id="8BEC36C9FFF8FF87FF3B2FB4FD3DF9AA" blockId="4.[151,652,1482,1575]" box="[151,652,1515,1539]" pageId="4" pageNumber="158">
Basionym:
<taxonomicName id="4C534D4AFFF8FF87FEA72FB4FD3DF9AA" authority="Kseneman, 1935: 3" authorityName="Kseneman" authorityPageNumber="3" authorityYear="1935" box="[267,652,1515,1539]" class="Collembola" family="Odontellidae" genus="Xenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="158" phylum="Arthropoda" rank="species" species="bayeri">
<emphasis id="B927EADBFFF8FF87FEA72FB4FE73F9AA" box="[267,450,1515,1539]" italics="true" pageId="4" pageNumber="158">Xenyllodes bayeri</emphasis>
<bibRefCitation id="EFC24B38FFF8FF87FE642FB3FD3DF9AA" author="Kseneman, M." box="[456,652,1515,1539]" pageId="4" pageNumber="158" pagination="39 - 65" refId="ref12941" refString="Kseneman, M. (1935). Apterygota z uzemi rybniku Lednickych. II. Easopis narodniho Musea, oddil pOiro-dovedny, Praha, 109, 39-65." type="journal article" year="1935">Kseneman, 1935: 3</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BEC36C9FFF8FF87FF3B2C52FE8FF98E" blockId="4.[151,652,1482,1575]" box="[151,318,1549,1575]" pageId="4" pageNumber="158">
<figureCitation id="13682A4CFFF8FF87FF3B2C52FF5DF98E" box="[151,236,1549,1575]" captionStart="FIGURES 1015" captionStartId="3.[151,264,1349,1373]" captionTargetBox="[160,1424,185,1319]" captionTargetId="figure-19@3.[151,1436,181,1325]" captionTargetPageId="3" captionText="FIGURES 1015. Axenyllodes ghilarovi (1012), A. bayeri (13), A. nematodes (14) and A. echinatus (15): 10, dorsal chaetotaxy of head and thorax, holotype; 11, abdominal tip, holotype; 1215, chaetotaxy of manubrial field: m—setae on main part, b— basal setae and bl—basolateral setae. Scales: 1011, 0.1 mm, 1215, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797990" httpUri="https://zenodo.org/record/14797990/files/figure.png" pageId="4" pageNumber="158">Figs 13</figureCitation>
,
<figureCitation id="13682A4CFFF8FF87FF542C52FE8FF98E" box="[248,318,1549,1575]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="4" pageNumber="158">1623</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF8FF86FF3B2C0AFEBFFECC" lastPageId="5" lastPageNumber="159" pageId="4" pageNumber="158" type="materials_examined">
<paragraph id="8BEC36C9FFF8FF87FF3B2C0AFD85F8EE" blockId="4.[151,1436,1621,1972]" pageId="4" pageNumber="158">
<emphasis id="B927EADBFFF8FF87FF3B2C0AFED3F9C6" bold="true" box="[151,354,1621,1647]" pageId="4" pageNumber="158">Studied material</emphasis>
.
<collectingCountry id="F3447659FFF8FF87FED82C0AFE72F9C6" box="[372,451,1621,1647]" name="Russia" pageId="4" pageNumber="158">Russia</collectingCountry>
:
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<emphasis id="B927EADBFFF8FF87FE7D2C0AFE6EF9C6" bold="true" box="[465,479,1621,1647]" pageId="4" pageNumber="158">8</emphasis>
females
</specimenCount>
and
<specimenCount id="9D55FD40FFF8FF87FDD22C0AFD42F9C6" box="[638,755,1621,1647]" count="1" pageId="4" pageNumber="158" type="juvenile">
<emphasis id="B927EADBFFF8FF87FDD22C0AFD3DF9C6" bold="true" box="[638,652,1621,1647]" pageId="4" pageNumber="158">1</emphasis>
juvenile
</specimenCount>
, Middle Volga River Basin,
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, “Privolzhskaya Lesostep” State Nature Reserve, xerophytic steppe with fescue on sandy soil (
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<emphasis id="B927EADBFFF8FF87FC562C25FB07F93A" box="[1018,1206,1657,1683]" italics="true" pageId="4" pageNumber="158">Festuca polesica</emphasis>
</taxonomicName>
,
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<emphasis id="B927EADBFFF8FF87FB682C25FA24F93A" box="[1220,1429,1657,1684]" italics="true" pageId="4" pageNumber="158">Stipa borysthenica</emphasis>
</taxonomicName>
;
<taxonomicName id="4C534D4AFFF8FF87FF3B2CC2FEC0F91E" box="[151,369,1693,1719]" class="Magnoliopsida" family="Rosaceae" genus="Potentilla" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="158" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B927EADBFFF8FF87FF3B2CC2FEC0F91E" box="[151,369,1693,1719]" italics="true" pageId="4" pageNumber="158">Potentilla argentea</emphasis>
</taxonomicName>
),
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52.4816
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N
</geoCoordinate>
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46.2027
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E
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,
<date id="FFED1009FFF8FF87FD312CC2FCCFF91E" box="[669,894,1693,1719]" pageId="4" pageNumber="158" value="2017-09-22">22 September 2017</date>
;
<specimenCount id="9D55FD40FFF8FF87FC212CC2FBB9F91E" box="[909,1032,1693,1719]" count="11" pageId="4" pageNumber="158" type="female">
<emphasis id="B927EADBFFF8FF87FC212CC2FC16F91E" bold="true" box="[909,935,1693,1719]" pageId="4" pageNumber="158">11</emphasis>
females
</specimenCount>
and
<specimenCount id="9D55FD40FFF8FF87FBE82CC2FB72F91E" box="[1092,1219,1693,1719]" count="7" pageId="4" pageNumber="158" type="juvenile">
<emphasis id="B927EADBFFF8FF87FBE82CC2FBE3F91E" bold="true" box="[1092,1106,1693,1719]" pageId="4" pageNumber="158">7</emphasis>
juveniles
</specimenCount>
, same biotope, but
<date id="FFED1009FFF8FF87FF3B2C9EFEC4F972" box="[151,373,1729,1755]" pageId="4" pageNumber="158" value="2018-09-27">27 September 2018</date>
;
<specimenCount id="9D55FD40FFF8FF87FE2D2C9EFE4DF972" box="[385,508,1729,1755]" count="30" pageId="4" pageNumber="158" type="female">
<emphasis id="B927EADBFFF8FF87FE2D2C9EFE2CF972" bold="true" box="[385,413,1729,1755]" pageId="4" pageNumber="158">30</emphasis>
females
</specimenCount>
and
<specimenCount id="9D55FD40FFF8FF87FD9F2C9EFD14F972" box="[563,677,1729,1755]" count="1" pageId="4" pageNumber="158" type="juvenile">
<emphasis id="B927EADBFFF8FF87FD9F2C9EFDF0F972" bold="true" box="[563,577,1729,1755]" pageId="4" pageNumber="158">1</emphasis>
juvenile
</specimenCount>
, same habitat, but
<date id="FFED1009FFF8FF87FCD42C9EFB89F972" box="[888,1080,1729,1755]" pageId="4" pageNumber="158" value="2019-10-01">01 October 2019</date>
;
<specimenCount id="9D55FD40FFF8FF87FBE82C9EFB05F972" box="[1092,1204,1729,1755]" count="2" pageId="4" pageNumber="158" type="female">
<emphasis id="B927EADBFFF8FF87FBE82C9EFBE3F972" bold="true" box="[1092,1106,1729,1755]" pageId="4" pageNumber="158">2</emphasis>
females
</specimenCount>
, same area, meadow steppe with
<taxonomicName id="4C534D4AFFF8FF87FEB72CBAFD9EF956" box="[283,559,1765,1791]" class="Liliopsida" family="Poaceae" genus="Brachypodium" kingdom="Plantae" order="Poales" pageId="4" pageNumber="158" phylum="Tracheophyta" rank="species" species="pinnatum">
<emphasis id="B927EADBFFF8FF87FEB72CBAFD9EF956" box="[283,559,1765,1791]" italics="true" pageId="4" pageNumber="158">Brachypodium pinnatum</emphasis>
</taxonomicName>
,
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52.4858
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N
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46.2113
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,
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.All Yu. Shveenkova leg.;
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<emphasis id="B927EADBFFF8FF87FA9A2CBAFAF5F956" bold="true" box="[1334,1348,1765,1791]" pageId="4" pageNumber="158">1</emphasis>
female
</specimenCount>
,
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, Pinega State Nature Reserve [
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64.56
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N
</geoCoordinate>
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43.26
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E
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], entrance to Golubinsky proval cave,
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<emphasis id="B927EADBFFF8FF87FAEE2D55FADEF88A" box="[1346,1391,1802,1827]" italics="true" pageId="4" pageNumber="158">Poa</emphasis>
sp.
</taxonomicName>
,
<date id="FFED1009FFF8FF87FF3B2D72FEC4F8EE" box="[151,373,1837,1863]" pageId="4" pageNumber="158" value="2004-09-21">21 September 2004</date>
. A. Babenko leg.
</paragraph>
<paragraph id="8BEC36C9FFF8FF87FF6B2D0EFE6BF81A" blockId="4.[151,1436,1621,1972]" pageId="4" pageNumber="158">
<collectingCountry id="F3447659FFF8FF87FF6B2D0EFEFFF8C2" box="[199,334,1873,1899]" name="Kazakhstan" pageId="4" pageNumber="158">Kazakhstan</collectingCountry>
:
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<emphasis id="B927EADBFFF8FF87FEF62D0EFEC7F8C2" bold="true" box="[346,374,1873,1899]" pageId="4" pageNumber="158">12</emphasis>
females
</specimenCount>
, Pavlodar Region, Motogul [
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53.39
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N
</geoCoordinate>
,
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75.87
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E
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], small birch grove, nest of
<taxonomicName id="4C534D4AFFF8FF87FA902D0DFEB5F826" authorityName="Foerster" authorityYear="1850" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="158" phylum="Arthropoda" rank="species" species="polyctena">
<emphasis id="B927EADBFFF8FF87FA902D0DFEB5F826" italics="true" pageId="4" pageNumber="158">Formica polyctena</emphasis>
</taxonomicName>
,
<date id="FFED1009FFF8FF87FEB82D2AFD91F826" box="[276,544,1909,1935]" pageId="4" pageNumber="158" value="1977-09-08" valueMax="1977-09-09" valueMin="1977-09-08">0809 September 1977</date>
. S.K. Stebaeva leg.;
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<emphasis id="B927EADBFFF8FF87FCBD2D2AFCAEF826" bold="true" box="[785,799,1909,1935]" pageId="4" pageNumber="158">5</emphasis>
females
</specimenCount>
, same area, date and collector, but steppe with
<taxonomicName id="4C534D4AFFF8FF87FF3B2DC5FEB7F81D" box="[151,262,1946,1972]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="4" pageNumber="158" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B927EADBFFF8FF87FF3B2DC5FF60F81D" box="[151,209,1946,1972]" italics="true" pageId="4" pageNumber="158">Stipa</emphasis>
spp.
</taxonomicName>
and
<emphasis id="B927EADBFFF8FF87FE902DC6FE6BF81A" box="[316,474,1945,1971]" italics="true" pageId="4" pageNumber="158">
<taxonomicName id="4C534D4AFFF8FF87FE902DC6FE67F81A" box="[316,470,1945,1971]" class="Magnoliopsida" family="Apiaceae" genus="Silaum" kingdom="Plantae" order="Apiales" pageId="4" pageNumber="158" phylum="Tracheophyta" rank="species" species="silaus">Silaum silaus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BEC36C9FFF9FF86FF6B2AC8FEBFFECC" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
<materialsCitation id="3B3B3C94FFF9FF86FF6B2AC8FD1CFF7C" collectingDate="2004-01-20" collectionCode="MNCN" collectorName="C. Gutierrez Martin" country="Spain" latitude="42.5003" location="Leiva" longLatPrecision="6" longitude="-3.0497" municipality="Ent" pageId="5" pageNumber="159" specimenCount="2" specimenCount-female="2" stateProvince="La Rioja">
<collectingCountry id="F3447659FFF9FF86FF6B2AC8FEBAFF18" box="[199,267,151,177]" name="Spain" pageId="5" pageNumber="159">Spain</collectingCountry>
:
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<emphasis id="B927EADBFFF9FF86FEBA2AC9FE95FF19" bold="true" box="[278,292,150,176]" pageId="5" pageNumber="159">2</emphasis>
females
</specimenCount>
(
<collectionCode id="ED42AE0CFFF9FF86FE3E2AC8FE56FF18" box="[402,487,151,177]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="159" type="Museum">MNCN</collectionCode>
_
<collectingMunicipality id="6B88ACB3FFF9FF86FE5A2AC8FDACFF18" box="[502,541,151,177]" pageId="5" pageNumber="159">Ent</collectingMunicipality>
45863 and 75086),
<location id="8E8C6012FFF9FF86FCAD2AC8FCF5FF18" LSID="urn:lsid:plazi:treatment:03FA87DFFFF8FF86FF3B2F95FCFDF8E4:8E8C6012FFF9FF86FCAD2AC8FCF5FF18" box="[769,836,151,177]" country="Spain" latitude="42.5003" longLatPrecision="6" longitude="-3.0497" municipality="Ent" name="Leiva" pageId="5" pageNumber="159" stateProvince="La Rioja">Leiva</location>
,
<collectingRegion id="4997F82BFFF9FF86FCE32AC8FBAEFF18" box="[847,1055,151,177]" country="Spain" name="La Rioja" pageId="5" pageNumber="159">La Rioja Province</collectingRegion>
, fruit culture,
<geoCoordinate id="EE67500EFFF9FF86FB132AC9FA8EFF18" box="[1215,1343,150,177]" degrees="42.5003" direction="north" orientation="latitude" pageId="5" pageNumber="159" precision="5" value="42.5003">42.5003º N</geoCoordinate>
<geoCoordinate id="EE67500EFFF9FF86FAEB2AC8FF03FF7C" degrees="3.0497" direction="west" orientation="longitude" pageId="5" pageNumber="159" precision="5" value="-3.0497">3.0497º W</geoCoordinate>
,
<date id="FFED1009FFF9FF86FF6C2AE4FE32FF7C" box="[192,387,187,213]" pageId="5" pageNumber="159" value="2004-01-20">
<collectingDate id="EFA9E9E1FFF9FF86FF6C2AE4FE32FF7C" box="[192,387,187,213]" pageId="5" pageNumber="159" value="2004-01-20">20 January 2004</collectingDate>
</date>
,
<collectorName id="26A6531FFFF9FF86FE3E2AE4FDC7FF7C" box="[402,630,187,213]" pageId="5" pageNumber="159">C. Gutiérrez Martín</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B3B3C94FFF9FF86FD102AE5FA94FF50" collectingDate="1954-04-04" collectionCode="MNCN" collectorName="W. Steiner" country="Spain" latitude="40.3589" location="Vallecas" longLatPrecision="6" longitude="-3.6322" municipality="Ent" pageId="5" pageNumber="159" specimenCount="3" specimenCount-female="3" stateProvince="Madrid">
<specimenCount id="9D55FD40FFF9FF86FD102AE5FC9DFF7C" box="[700,812,186,213]" count="3" pageId="5" pageNumber="159" type="female">
<emphasis id="B927EADBFFF9FF86FD102AE5FD7BFF7D" bold="true" box="[700,714,186,212]" pageId="5" pageNumber="159">3</emphasis>
females
</specimenCount>
(
<collectionCode id="ED42AE0CFFF9FF86FC922AE4FC22FF7C" box="[830,915,187,213]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="159" type="Museum">MNCN</collectionCode>
_
<collectingMunicipality id="6B88ACB3FFF9FF86FC0D2AE4FC79FF7C" box="[929,968,187,213]" pageId="5" pageNumber="159">Ent</collectingMunicipality>
109949, 109950 and 109951),
<location id="8E8C6012FFF9FF86FA942AE4FA29FF7C" LSID="urn:lsid:plazi:treatment:03FA87DFFFF8FF86FF3B2F95FCFDF8E4:8E8C6012FFF9FF86FA942AE4FA29FF7C" box="[1336,1432,187,213]" country="Spain" latitude="40.3589" longLatPrecision="6" longitude="-3.6322" municipality="Ent" name="Vallecas" pageId="5" pageNumber="159" stateProvince="Madrid">Vallecas</location>
,
<collectingRegion id="4997F82BFFF9FF86FF3B2A80FEE6FF50" box="[151,343,223,249]" country="Spain" name="Madrid" pageId="5" pageNumber="159">Madrid Province</collectingRegion>
, grass on gypsum soil (sierozem),
<geoCoordinate id="EE67500EFFF9FF86FD762A81FCE8FF50" box="[730,857,222,249]" degrees="40.3589" direction="north" orientation="latitude" pageId="5" pageNumber="159" precision="5" value="40.3589">40.3589º N</geoCoordinate>
<geoCoordinate id="EE67500EFFF9FF86FCCC2A80FC66FF50" box="[864,983,223,249]" degrees="3.6322" direction="west" orientation="longitude" pageId="5" pageNumber="159" precision="5" value="-3.6322">3.6322º W</geoCoordinate>
,
<date id="FFED1009FFF9FF86FC4E2A80FBC3FF50" box="[994,1138,222,249]" pageId="5" pageNumber="159" value="1954-04-04">
<collectingDate id="EFA9E9E1FFF9FF86FC4E2A80FBC3FF50" box="[994,1138,222,249]" pageId="5" pageNumber="159" value="1954-04-04">4 April 1954</collectingDate>
</date>
,
<collectorName id="26A6531FFFF9FF86FBD02A80FB40FF50" box="[1148,1265,223,249]" pageId="5" pageNumber="159">W. Steiner</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B3B3C94FFF9FF86FA9C2A81FDC6FEE8" collectionCode="MNCN" collectorName="A. Signoret" country="Spain" county="Ent" latitude="41.7961" location="Cabreja Range" longLatPrecision="6" longitude="-2.7878" municipality="Abejar" pageId="5" pageNumber="159" specimenCount="2" specimenCount-female="2" stateProvince="Soria Province">
<specimenCount id="9D55FD40FFF9FF86FA9C2A81FA2DFF50" box="[1328,1436,222,249]" count="2" pageId="5" pageNumber="159" type="female">
<emphasis id="B927EADBFFF9FF86FA9C2A81FA8FFF51" bold="true" box="[1328,1342,222,248]" pageId="5" pageNumber="159">2</emphasis>
females
</specimenCount>
(
<collectionCode id="ED42AE0CFFF9FF86FF332B5CFF45FEB4" box="[159,244,259,285]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="159" type="Museum">MNCN</collectionCode>
_
<collectingCounty id="628D4E45FFF9FF86FEAE2B5CFE98FEB4" box="[258,297,259,285]" pageId="5" pageNumber="159">Ent</collectingCounty>
45366 and 45373),
<collectingMunicipality id="6B88ACB3FFF9FF86FDBE2B5CFDD1FEB4" box="[530,608,259,285]" pageId="5" pageNumber="159">Abejar</collectingMunicipality>
,
<location id="8E8C6012FFF9FF86FDC12B5CFCA6FEB4" LSID="urn:lsid:plazi:treatment:03FA87DFFFF8FF86FF3B2F95FCFDF8E4:8E8C6012FFF9FF86FDC12B5CFCA6FEB4" box="[621,791,259,285]" country="Spain" county="Ent" latitude="41.7961" longLatPrecision="6" longitude="-2.7878" municipality="Abejar" name="Cabreja Range" pageId="5" pageNumber="159" stateProvince="Soria Province">Cabreja Range</location>
,
<collectingRegion id="4997F82BFFF9FF86FC892B5CFC61FEB4" box="[805,976,259,285]" inferredBySuffix="true" pageId="5" pageNumber="159">Soria Province</collectingRegion>
, juniper leaf litter (
<taxonomicName id="4C534D4AFFF9FF86FB1D2B5CFA3DFEB4" box="[1201,1420,259,285]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="159" phylum="Tracheophyta" rank="species" species="thurifera">
<emphasis id="B927EADBFFF9FF86FB1D2B5CFA3DFEB4" box="[1201,1420,259,285]" italics="true" pageId="5" pageNumber="159">Juniperus thurifera</emphasis>
</taxonomicName>
),
<geoCoordinate id="EE67500EFFF9FF86FF3B2B78FEA5FEE8" box="[151,276,295,321]" degrees="41.7961" direction="north" orientation="latitude" pageId="5" pageNumber="159" precision="5" value="41.7961">41.7961º N</geoCoordinate>
<geoCoordinate id="EE67500EFFF9FF86FEB62B78FE21FEE8" box="[282,400,295,321]" degrees="2.7878" direction="west" orientation="longitude" pageId="5" pageNumber="159" precision="5" value="-2.7878">2.7878º W</geoCoordinate>
,
<collectorName id="26A6531FFFF9FF86FE342B78FDA6FEE8" box="[408,535,295,321]" pageId="5" pageNumber="159">A. Signoret</collectorName>
leg. The
</materialsCitation>
<materialsCitation id="3B3B3C94FFF9FF86FDD02B78FEBBFECC" collectionCode="MNCN" country="Spain" location="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="159" specimenCount="1" stateProvince="Madrid">
material is stored in the
<location id="8E8C6012FFF9FF86FC2E2B78FA81FEE8" LSID="urn:lsid:plazi:treatment:03FA87DFFFF8FF86FF3B2F95FCFDF8E4:8E8C6012FFF9FF86FC2E2B78FA81FEE8" box="[898,1328,295,321]" country="Spain" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="159" stateProvince="Madrid">Museo Nacional de Ciencias Naturales</location>
(
<collectionCode id="ED42AE0CFFF9FF86FA912B78FA24FEE8" box="[1341,1429,295,321]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="5" pageNumber="159" type="Museum">MNCN</collectionCode>
) in
<collectingRegion id="4997F82BFFF9FF86FF182B14FEBBFECC" box="[180,266,331,357]" country="Spain" name="Madrid" pageId="5" pageNumber="159">Madrid</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF9FF86FF6B2B31FE7BFE04" pageId="5" pageNumber="159" type="diagnosis">
<paragraph id="8BEC36C9FFF9FF86FF6B2B31FE7BFE04" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
<emphasis id="B927EADBFFF9FF86FF6B2B31FE8BFE21" bold="true" box="[199,314,366,392]" pageId="5" pageNumber="159">Diagnosis</emphasis>
. A species of the genus distinguished from all other known congeners by its large PAO that usually having four irregular lobes.
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF9FF86FF6B2BE9FAC9FA68" pageId="5" pageNumber="159" type="description">
<paragraph id="8BEC36C9FFF9FF86FF6B2BE9FCD6FD94" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
<emphasis id="B927EADBFFF9FF86FF6B2BE9FCEEFE79" bold="true" box="[199,863,438,465]" pageId="5" pageNumber="159">
Description based on specimens from
<collectingCountry id="F3447659FFF9FF86FD252BE9FD69FE79" box="[649,728,438,464]" name="Russia" pageId="5" pageNumber="159">Russia</collectingCountry>
and
<collectingCountry id="F3447659FFF9FF86FCBF2BE9FCEBFE79" box="[787,858,438,464]" name="Spain" pageId="5" pageNumber="159">Spain</collectingCountry>
.
</emphasis>
Length (without antennae)
<quantity id="4CAB9B2CFFF9FF86FB392BE9FA8AFE78" box="[1173,1339,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.3" metricValueMin="4.3" pageId="5" pageNumber="159" unit="mm" value="0.48" valueMax="0.53" valueMin="0.43">0.430.53 mm</quantity>
. Habitus typical of the genus. Colour white, without any traces of dark pigment. Tegument granulation coarse, secondary granules rounded at tip. Head and all terga from Th. I to Abd. V with rounded fields with finer granulations located more or less symmetrically with respect to medial line (
<figureCitation id="13682A4CFFF9FF86FCAB287CFCEBFD94" box="[775,858,547,573]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BEC36C9FFF9FF86FF6B2818FBA1FD58" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
Antennae about as long as head. Ant. IV with a large simple vesicle apically, six rather thin, but clearly differentiated sensilla with pointed apices (three dorso-external [S7, S8, S9] and three dorso-internal [S1, S2, S4]), ms rather large and broadened at apex, notably shorter in length than sensilla, subapical organite present (
<figureCitation id="13682A4CFFF9FF86FA9628D0FA3FFD00" box="[1338,1422,655,681]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 19</figureCitation>
); ventral side of Ant. IV with few ordinary setae (
<figureCitation id="13682A4CFFF9FF86FD1B28ECFCBAFD64" box="[695,779,691,717]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 18</figureCitation>
). Antennal organ of Ant. III typical, all its sensilla and ms subequal (
<figureCitation id="13682A4CFFF9FF86FEA62888FEECFD58" box="[266,349,727,753]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 17</figureCitation>
). Ant. IIII with 7, 10 and 1517 ordinary setae, respectively.
</paragraph>
<paragraph id="8BEC36C9FFF9FF86FF6B28A4FC9EFBB8" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
Head with 2+2 ocelli, their diameter equal to 0.380.50 of PAO length, distance between them usually longer than their diameter (1.01.6: 1). PAO large (913 μm in length) and irregularly lobed, located in a depression (
<figureCitation id="13682A4CFFF9FF86FF33291CFF44FCF4" box="[159,245,835,861]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 20</figureCitation>
), sometimes partly hidden under cuticular folds. Buccal cone short, typical of the genus. Labrum with 8 usual setae, medial pair of setae in anterior row stronger, with tips curved laterally, only two prelabral setae always present. Maxillary palp simple. Labium with five apical spinules and three common setae subequal to them in length. Basomedial and basolateral fields of labium with 4+4 setae, respectively. Maxilla typical of the family with strong, triangular head, mandibles invisible. Ventral side of head in adults with 3+3 postlabial setae along ventral line (
<figureCitation id="13682A4CFFF9FF86FF7C29A8FE92FBB8" box="[208,291,1015,1041]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 21</figureCitation>
), but juveniles often with 2+2 postlabial setae.
</paragraph>
<paragraph id="8BEC36C9FFF9FF86FF6B2E44FB01FB08" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
Ordinary setae on dorsal side of body smooth and acuminate, slightly longer on last abdominal terga, sensilla undifferentiated (
<figureCitation id="13682A4CFFF9FF86FEF72E60FE00FBF0" box="[347,433,1087,1113]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 16</figureCitation>
). Main characteristics of dorsal chaetotaxy: Th. IIIII with three setae present in a-row (a1, a3 and a5), two setae and sensilla in m-row (m1, m4 and S [=m6]) and five setae in p-row, lateral microsensilla invisible; Abd. IIII with 45 setae in a-row (seta a4 often absent); Abd. IV without setae m1.
</paragraph>
<paragraph id="8BEC36C9FFF9FF86FF6B2EF4FAFEFAFC" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
Thoracic sterna without setae. Ventral tube with 3+3 distal setae, one seta also present on each side of VT on sternum of Abd. I, Abd. IIIII with 4 and 5 ventral setae on each side, respectively (
<figureCitation id="13682A4CFFF9FF86FBE72E90FB11FB40" box="[1099,1184,1231,1257]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 22</figureCitation>
). Tenaculum with 2+2 teeth. Furca small (
<figureCitation id="13682A4CFFF9FF86FEC12EACFE75FAA4" box="[365,452,1267,1293]" captionStart="FIGURES 1015" captionStartId="3.[151,264,1349,1373]" captionTargetBox="[160,1424,185,1319]" captionTargetId="figure-19@3.[151,1436,181,1325]" captionTargetPageId="3" captionText="FIGURES 1015. Axenyllodes ghilarovi (1012), A. bayeri (13), A. nematodes (14) and A. echinatus (15): 10, dorsal chaetotaxy of head and thorax, holotype; 11, abdominal tip, holotype; 1215, chaetotaxy of manubrial field: m—setae on main part, b— basal setae and bl—basolateral setae. Scales: 1011, 0.1 mm, 1215, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797990" httpUri="https://zenodo.org/record/14797990/files/figure.png" pageId="5" pageNumber="159">Figs 13</figureCitation>
,
<figureCitation id="13682A4CFFF9FF86FE7C2EACFE5EFAA4" box="[464,495,1267,1293]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">22</figureCitation>
), manubrium with 4+4 setae on main part, 1+1 basal setae and 3+3 basolateral setae (8+8 setae in total); dorsal side of dens with two setae and rather fine granulation. Mucro about as long as dens, strongly curved, lateral lamella not especially high. Anal spines strong and curved, set on subequal papillae.
</paragraph>
<paragraph id="8BEC36C9FFF9FF86FF6B2F00FAC9FA68" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
Legs IIII with more or less stable sets of setae: 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 4, 4, 4 setae on coxae, 5, (4)5, 45 on trochanters, 8, 8, 8 setae on femora, and 10109(10) setae on each tibiotarsus, the latter setae clearly differing in length. Unguis toothless, unguiculus needle-shaped and rather long (
<figureCitation id="13682A4CFFF9FF86FAB42FF8FADDFA68" box="[1304,1388,1447,1473]" captionStart="FIGURES 1623" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[167,1420,191,1909]" captionTargetId="figure-17@6.[153,1434,181,1919]" captionTargetPageId="6" captionText="FIGURES 1623. Axenyllodes bayeri: 16, cuticular granulation and dorsal chaetotaxy; 17, AOIII; 18, chaetotaxy of antenna, ventral view; 19, ibid., dorsal view; 20, PAO and ocelli; 21, ventral chaetotaxy of head; 22, ventral chaetotaxy of abdomen; 23, tibiotarsi I. Scale: 16, 2122, 0.05 mm, 1819, 0.025 mm, 17, 20, 23, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797992" httpUri="https://zenodo.org/record/14797992/files/figure.png" pageId="5" pageNumber="159">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFF9FF86FF6B2F94FCFDF8E4" pageId="5" pageNumber="159" type="discussion">
<paragraph id="8BEC36C9FFF9FF86FF6B2F94FCFDF8E4" blockId="5.[151,1437,150,1869]" pageId="5" pageNumber="159">
<emphasis id="B927EADBFFF9FF86FF6B2F94FE85FA4C" bold="true" box="[199,308,1483,1509]" pageId="5" pageNumber="159">Remarks</emphasis>
.
<taxonomicName id="4C534D4AFFF9FF86FEEC2F94FDA4FA4C" box="[320,533,1483,1509]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="5" pageNumber="159" phylum="Arthropoda" rank="species" species="bayeri">
<emphasis id="B927EADBFFF9FF86FEEC2F94FDA4FA4C" box="[320,533,1483,1509]" italics="true" pageId="5" pageNumber="159">Axenyllodes bayeri</emphasis>
</taxonomicName>
, the
<typeStatus id="54E8886BFFF9FF86FDE42F94FDC9FA4C" box="[584,632,1483,1509]" pageId="5" pageNumber="159">type</typeStatus>
species of the genus, differs quite radically from all other representatives of the genus by its unique shape of the PAO. It is widespread in the Palaearctic and, in addition to the original description from the
<collectingCountry id="F3447659FFF9FF86FED22C4CFD83F984" box="[382,562,1555,1581]" name="Czech Republic" pageId="5" pageNumber="159">Czech Republic</collectingCountry>
, there are several more or less detailed redescriptions (
<bibRefCitation id="EFC24B38FFF9FF86FB222C4CFABEF984" author="Stach, J." box="[1166,1295,1555,1581]" pageId="5" pageNumber="159" refId="ref13426" refString="Stach, J. (1949) The Apterygotan Fauna of Poland in Relation to the World-Fauna of this Group of Insects. Families: Neogastruridae and Brachystomellidae. Acta Monographica Musei Historiae Naturalis, Krakow, 341 pp." type="book" year="1949">Stach 1949</bibRefCitation>
;
<bibRefCitation id="EFC24B38FFF9FF86FAB62C4CFA26F984" author="Gisin, H." box="[1306,1431,1555,1581]" pageId="5" pageNumber="159" refId="ref12624" refString="Gisin, H. (1960) Collembolenfauna Europas. Museum d'Histoire Naturelle, Geneve, 312 pp." type="book" year="1960">Gisin 1960</bibRefCitation>
; Pallissa 1964;
<bibRefCitation id="EFC24B38FFF9FF86FE972C68FDA5F9F8" author="Jordana, R. &amp; Arbea, J. I. &amp; Simon, C. &amp; Lucianez, M. J." box="[315,532,1591,1617]" pageId="5" pageNumber="159" pagination="1 - 807" refId="ref12702" refString="Jordana, R., Arbea, J. I., Simon, C. &amp; Lucianez, M. J. (1997) Collembola, Poduromorpha. In: Ramos, M. A. (Ed.), Fauna Iberica. Vol. 8. Museo Nacional de Ciencas Naturales, CSIC, Madrid, pp 1-807." type="book chapter" year="1997">
Jordana
<emphasis id="B927EADBFFF9FF86FE362C68FE7AF9F8" box="[410,459,1591,1617]" italics="true" pageId="5" pageNumber="159">et al</emphasis>
. 1997
</bibRefCitation>
;
<bibRefCitation id="EFC24B38FFF9FF86FD8C2C68FD7FF9F8" author="Fjellberg, A." box="[544,718,1590,1617]" pageId="5" pageNumber="159" pagination="153 - 158" refId="ref12502" refString="Fjellberg, A. (1995) Collembola of the Canary Islands. II. Family Odontellidae. Entomologica Scandinavica, 26, 153-158. https://doi.org/10.1163/187631295X00152" type="journal article" year="1995">Fjellberg, 1995</bibRefCitation>
), between which and the above description, being mainly based on material from the Volga region, there are but few discrepancies. The most important of these is the presence or absence of lateral microsensilla on the thoracic segments.According to fig.
<quantity id="4CAB9B2CFFF9FF86FC722C20FB92F930" box="[990,1059,1663,1689]" metricMagnitude="0" metricUnit="m" metricValue="2.9718" pageId="5" pageNumber="159" unit="in" value="117.0">117 in</quantity>
<bibRefCitation id="EFC24B38FFF9FF86FB852C20FAA1F930" author="Jordana, R. &amp; Arbea, J. I. &amp; Simon, C. &amp; Lucianez, M. J." box="[1065,1296,1663,1689]" pageId="5" pageNumber="159" pagination="1 - 807" refId="ref12702" refString="Jordana, R., Arbea, J. I., Simon, C. &amp; Lucianez, M. J. (1997) Collembola, Poduromorpha. In: Ramos, M. A. (Ed.), Fauna Iberica. Vol. 8. Museo Nacional de Ciencas Naturales, CSIC, Madrid, pp 1-807." type="book chapter" year="1997">
Jordana
<emphasis id="B927EADBFFF9FF86FB2A2C20FB06F930" box="[1158,1207,1663,1689]" italics="true" pageId="5" pageNumber="159">et al</emphasis>
. (1997)
</bibRefCitation>
, the Spanish specimens have microsensilla on both thoracic segments. It appears that Thibauds statement (2006, p. 114) of the presence of lateral sensilla is based on this figure:
<emphasis id="B927EADBFFF9FF86FD6F2C98FA81F949" box="[707,1328,1735,1761]" italics="true" pageId="5" pageNumber="159">thorax II et III avec chacun 1 paire de microsensilles ms</emphasis>
. We have revised the specimens from
<collectingCountry id="F3447659FFF9FF86FE7F2CB4FDA2F8AC" box="[467,531,1771,1797]" name="Spain" pageId="5" pageNumber="159">Spain</collectingCountry>
(
<collectingRegion id="4997F82BFFF9FF86FD8E2CB4FD38F8AC" box="[546,649,1771,1797]" country="Spain" name="La Rioja" pageId="5" pageNumber="159">La Rioja</collectingRegion>
, Soria,
<collectingRegion id="4997F82BFFF9FF86FD722CB4FC83F8AC" box="[734,818,1771,1797]" country="Spain" name="Madrid" pageId="5" pageNumber="159">Madrid</collectingRegion>
) which served for the description in the Ibérian Fauna, but did not find the ms on Th. IIIII. Thus, we can confidently say that the material described above from the Central Palaearctic belongs to the same species that occurs in Europe.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="D0A481A5FFFBFF84FF3B2AC8FE52FF18" bold="true" box="[151,483,151,178]" fontSize="11" level="1" pageId="7" pageNumber="161" reason="1">
<emphasis id="B927EADBFFFBFF84FF3B2AC8FE52FF18" bold="true" box="[151,483,151,178]" pageId="7" pageNumber="161">
<taxonomicName id="4C534D4AFFFBFF84FF3B2AC8FECFFF1B" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[151,382,151,178]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFFBFF84FF3B2AC8FECFFF1B" bold="true" box="[151,382,151,178]" italics="true" pageId="7" pageNumber="161">Axenyllodes chinki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457A0FFFBFF84FE292AC8FE52FF18" box="[389,483,151,177]" pageId="7" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFFBFF84FF3B2AE4FEA5FF7C" box="[151,276,187,213]" pageId="7" pageNumber="161" type="description">
<paragraph id="8BEC36C9FFFBFF84FF3B2AE4FEA5FF7C" blockId="7.[151,483,151,213]" box="[151,276,187,213]" pageId="7" pageNumber="161">
<figureCitation id="13682A4CFFFBFF84FF3B2AE4FEA5FF7C" box="[151,276,187,213]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Figs 2431</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFFBFF84FF3B2B5CFBDAFECC" pageId="7" pageNumber="161" type="materials_examined">
<paragraph id="8BEC36C9FFFBFF84FF3B2B5CFBDAFECC" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
<emphasis id="B927EADBFFFBFF84FF3B2B5CFEF3FEB4" bold="true" box="[151,322,259,285]" pageId="7" pageNumber="161">Type material</emphasis>
.
<materialsCitation id="3B3B3C94FFFBFF84FEF82B5CFE71FECC" collectingDate="2023-10-14" collectingDateMax="2023-10-16" collectingDateMin="2023-10-14" collectorName="L. Kim" country="Kazakhstan" county="Syr Darya River" elevation="230" location="Turkistan Region" municipality="Kyzylkum" pageId="7" pageNumber="161" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E8886BFFFBFF84FEF82B5CFE70FEB4" box="[340,449,259,285]" pageId="7" pageNumber="161" type="holotype">Holotype</typeStatus>
: female,
<collectingCountry id="F3447659FFFBFF84FD822B5CFD05FEB4" box="[558,692,259,285]" name="Kazakhstan" pageId="7" pageNumber="161">Kazakhstan</collectingCountry>
,
<location id="8E8C6012FFFBFF84FD6E2B5CFC3AFEB4" LSID="urn:lsid:plazi:treatment:03FA87DFFFFBFF8AFF3B2AC8FEB0F933:8E8C6012FFFBFF84FD6E2B5CFC3AFEB4" box="[706,907,259,285]" country="Kazakhstan" county="Syr Darya River" municipality="Kyzylkum" name="Turkistan Region" pageId="7" pageNumber="161">Turkistan Region</location>
,
<collectingMunicipality id="6B88ACB3FFFBFF84FC362B5CFBA1FEB4" box="[922,1040,259,285]" pageId="7" pageNumber="161">Kyzylkum</collectingMunicipality>
sandy desert, basin of
<collectingCounty id="628D4E45FFFBFF84FA8A2B5CFF66FEE8" pageId="7" pageNumber="161">Syr Darya River</collectingCounty>
,
<locationDeviation id="9A5E2FEBFFFBFF84FF4F2B78FE18FEE8" box="[227,425,295,322]" location="Arys" pageId="7" pageNumber="161">
<quantity id="4CAB9B2CFFFBFF84FF4F2B78FE9BFEE8" box="[227,298,295,322]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="161" unit="km" value="85.0">85 km</quantity>
W of Arys
</locationDeviation>
, 42.2325
<superScript id="7C269B81FFFBFF84FDBC2B78FD9DFEEB" attach="right" box="[528,556,295,322]" fontSize="6" pageId="7" pageNumber="161">oN</superScript>
67.8128
<superScript id="7C269B81FFFBFF84FD232B78FD1BFEEB" attach="right" box="[655,682,295,322]" fontSize="6" pageId="7" pageNumber="161">oE</superScript>
,
<quantity id="4CAB9B2CFFFBFF84FD1A2B78FD4FFEEB" box="[694,766,295,322]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="161" unit="m" value="230.0">
<elevation id="007ED1FAFFFBFF84FD1A2B78FD4FFEEB" box="[694,766,295,322]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="161" unit="m" value="230.0">230 m</elevation>
</quantity>
alt., bottom of SE slope of “Small Chink”, pit-traps,
<date id="FFED1009FFFBFF84FAFA2B78FE85FECC" pageId="7" pageNumber="161" value="2023-10-14" valueMax="2023-10-16" valueMin="2023-10-14">
<collectingDate id="EFA9E9E1FFFBFF84FAFA2B78FE85FECC" pageId="7" pageNumber="161" value="2023-10-14" valueMax="2023-10-16" valueMin="2023-10-14">1416 October 2023</collectingDate>
</date>
.
<collectorName id="26A6531FFFFBFF84FE922B13FE3EFECC" box="[318,399,331,358]" pageId="7" pageNumber="161">L. Kim</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B3B3C94FFFBFF84FE6A2B13FBD6FECC" box="[454,1127,331,358]" collectingDate="2023-10-14" collectingDateMax="2023-10-16" collectingDateMin="2023-10-14" collectorName="L. Kim" country="Kazakhstan" county="Syr Darya River" elevation="230" location="Turkistan Region" municipality="Kyzylkum" pageId="7" pageNumber="161" specimenCount="10" specimenCount-female="6" specimenCount-juvenile="4" typeStatus="paratype">
<typeStatus id="54E8886BFFFBFF84FE6A2B13FD86FECF" box="[454,567,332,358]" pageId="7" pageNumber="161" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D55FD40FFFBFF84FDEE2B14FD1FFECC" box="[578,686,331,357]" count="6" pageId="7" pageNumber="161" type="female">
<emphasis id="B927EADBFFFBFF84FDEE2B14FDE1FECC" bold="true" box="[578,592,331,357]" pageId="7" pageNumber="161">6</emphasis>
females
</specimenCount>
and
<specimenCount id="9D55FD40FFFBFF84FD492B14FCD3FECC" box="[741,866,331,357]" count="4" pageId="7" pageNumber="161" type="juvenile">
<emphasis id="B927EADBFFFBFF84FD492B14FD42FECC" bold="true" box="[741,755,331,357]" pageId="7" pageNumber="161">4</emphasis>
juveniles
</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFFBFF84FF6B2B30FAB1FDB0" pageId="7" pageNumber="161" type="diagnosis">
<paragraph id="8BEC36C9FFFBFF84FF6B2B30FAB1FDB0" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
<emphasis id="B927EADBFFFBFF84FF6B2B30FE8BFE20" bold="true" box="[199,314,367,393]" pageId="7" pageNumber="161">Diagnosis</emphasis>
.
<materialsCitation id="3B3B3C94FFFBFF84FEE42B30FB4DFDB0" collectorName="Ant. IV &amp; Th. II" country="American Samoa" location="AS" pageId="7" pageNumber="161" specimenCount="1">
A species of the genus, characterized by the presence of (6)7 clearly differentiated dorsal sensilla and a rather large microsensillum on
<collectorName id="26A6531FFFFBFF84FD9B2BCCFD3EFE07" box="[567,655,403,430]" pageId="7" pageNumber="161">Ant. IV</collectorName>
; 2+2 ocelli with traces of dark pigment under each of them; a distinct pattern of cuticular granulation on the terga; a specific form of PAO, almost completely hidden under the cuticular fold; the absence of lateral microsensilla and the presence of setae m1 on
<collectorName id="26A6531FFFFBFF84FC692B84FBB5FE5F" box="[965,1028,475,502]" pageId="7" pageNumber="161">Th. II</collectorName>
III; the basic
<typeStatus id="54E8886BFFFBFF84FB082B83FB65FE5F" box="[1188,1236,476,502]" pageId="7" pageNumber="161">type</typeStatus>
of VT, tenaculum and furca; the tibiotarsi with 777 setae, two of which are distinctly clavate; and the absence of
<collectingCountry id="F3447659FFFBFF84FB792BA0FB4DFDB0" box="[1237,1276,511,537]" name="American Samoa" pageId="7" pageNumber="161">AS</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFFBFF84FF6B287CFD8DF914" pageId="7" pageNumber="161" type="description">
<paragraph id="8BEC36C9FFFBFF84FF6B287CFF75FD5B" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
<emphasis id="B927EADBFFFBFF84FF6B287CFEE8FD94" bold="true" box="[199,345,547,573]" pageId="7" pageNumber="161">Description.</emphasis>
Length (without antennae)
<quantity id="4CAB9B2CFFFBFF84FD3B287CFC8FFD97" box="[663,830,547,574]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" metricValueMax="8.299999999999999" metricValueMin="6.8999999999999995" pageId="7" pageNumber="161" unit="mm" value="0.76" valueMax="0.83" valueMin="0.69">0.690.83 mm</quantity>
. Habitus typical of the genus: body slim and oblong, appendices short (
<figureCitation id="13682A4CFFFBFF84FEF22818FE01FDC8" box="[350,432,583,609]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 24</figureCitation>
). Colour white with few granules of dark pigment under ocelli. Tegument granulation rather coarse, secondary granules usually spinous at tip. Head with fields of fine granulations in central and posterolateral areas, forming a characteristic pattern of longitudinal stripes; similar granulations present each side of thoracic segments. Abd. IV with two oval fields of fine granulations located symmetrically with respect to medial line (
<figureCitation id="13682A4CFFFBFF84FAC328ECFF06FD5B" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="8BEC36C9FFFBFF84FF6B28A4FEDBFC60" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
Antennae about as long as head. Ant. IV with a large simple vesicle apically, with 7 clearly differentiated sensilla with pointed apices (four dorso-internal [S1S4] and three dorso-external [S7S9]; S9 often absent from immature specimens), ms large, clearly broadened at apex, only slightly shorter in length than sensilla, subapical organite present (
<figureCitation id="13682A4CFFFBFF84FEF22938FE02FC28" box="[350,435,871,897]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 27</figureCitation>
), ventral side with few ordinary setae. Antennal organ of Ant. III typical, inner sensilla small, sgv &amp; sgd only slightly larger, ms and about 1618 ordinary setae present on Ant. III. Ant. III with 7 and 10 setae, respectively.
</paragraph>
<paragraph id="8BEC36C9FFFBFF84FF6B298CFE13FB6C" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
Head with 2+2 subequal ocelli, their diameter equal to 0.240.36 of PAO length. PAO sunken in a deep cavity and hardly visible from above, broadly oval in shape, edges pointed (
<figureCitation id="13682A4CFFFBFF84FC1D29A8FB8EFBB8" box="[945,1087,1015,1041]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 26, 26a</figureCitation>
). Buccal cone short, typical of the genus (
<figureCitation id="13682A4CFFFBFF84FEB82E44FEDBFB9C" box="[276,362,1051,1077]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 28</figureCitation>
). Maxillary palp simple. Labrum with 8 usual setae, medial pair of setae in anterior row much stronger, with tips curved laterally; four prelabral setae also present. Labium with five small apical spinules, two tiny organites and three common setae. Basomedial and basolateral fields of labium with 4+4 setae each. Maxilla typical of the family, with a strong and triangular head, mandibles invisible. Ventral side of head with 3+3 postlabial setae along ventral line.
</paragraph>
<paragraph id="8BEC36C9FFFBFF84FF6B2E90FE24FAD3" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
Ordinary setae on dorsal side of body short, smooth and acuminate, distinctly longer on last abdominal terga, sensilla undifferentiated (
<figureCitation id="13682A4CFFFBFF84FE152EACFDA1FAA7" box="[441,528,1267,1294]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 25</figureCitation>
). Main characteristics of dorsal chaetotaxy: setae m1 present on Th. IIIII and absent from Abd. IV; Th. IIIII with four setae present in a-row (a1, a3, a4 and a5), m-row with two ordinary setae (m1 and m4) and lateral sensillum in m6 position, lateral microsensilla invisible; Abd. IIII with five setae in a-row (a1, a2, a3, a4 and a5).
</paragraph>
<paragraph id="8BEC36C9FFFBFF84FF6B2FDCFBA0F984" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
Thoracic sterna without setae. Ventral tube with 3+3 distal setae, one seta also present each side of VT on sternum of Abd. I, Abd. IIIII with 45 ventral setae each side (
<figureCitation id="13682A4CFFFBFF84FCF12FF8FC00FA68" box="[861,945,1447,1473]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 29</figureCitation>
). Tenaculum with 2+2 teeth, as usual. Furca small (
<figureCitation id="13682A4CFFFBFF84FF482F94FE87FA4C" box="[228,310,1483,1509]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 31</figureCitation>
), manubrium with 4+4 setae on main part, 1+1 basal and 3+3 basolateral setae (8+
<quantity id="4CAB9B2CFFFBFF84FB702F94FAB7FA4C" box="[1244,1286,1483,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="7" pageNumber="161" unit="in" value="8.0">8 in</quantity>
total); dorsal side of dens with two setae and rather fine granulations. Mucro about as long as dens, strongly hooked and with clear lateral lamella. Anal opening located terminally; anal spines completely absent.
</paragraph>
<paragraph id="8BEC36C9FFFBFF84FF6B2C67FD8DF914" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
Legs IIII with a stable set of setae: 1, 2, 2 setae on upper subcoxae, 0, 3, 3 setae on lower subcoxae, 34, 34, 34 setae on coxae, 5, 5, 4(5) on trochanters, 7, 7, 7 setae on femora, inner one thinner and longer. Seven setae also present on each tibiotarsi including two longer and clearly clavate at apex (
<figureCitation id="13682A4CFFFBFF84FC402C20FBF0F930" box="[1004,1089,1663,1689]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 30</figureCitation>
). Unguis toothless, unguiculus rather long, needle-shaped (
<figureCitation id="13682A4CFFFBFF84FE632CFCFD9EF914" box="[463,559,1699,1725]" captionStart="FIGURES 2431" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[191,1396,195,1905]" captionTargetId="figure-17@8.[177,1410,181,1919]" captionTargetPageId="8" captionText="FIGURES 2431. Axenyllodes chinki sp. nov.: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="7" pageNumber="161">Fig. 30a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3496542FFFBFF8AFF6B2C98FEB0F933" lastPageId="9" lastPageNumber="163" pageId="7" pageNumber="161" type="etymology">
<paragraph id="8BEC36C9FFFBFF84FF6B2C98FDDBF880" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="161">
<emphasis id="B927EADBFFFBFF84FF6B2C98FEF9F948" bold="true" box="[199,328,1735,1761]" pageId="7" pageNumber="161">Etymology</emphasis>
. Named after “chink”, a regional term used, according to Wikipedia &lt;https://en.m.wikipedia.org/ wiki/Chink_(geology)&gt;, “
<emphasis id="B927EADBFFFBFF84FE162CB3FE5CF880" italics="true" pageId="7" pageNumber="161">
in Central Asia for steep chalk and limestone escarpments and cliffs heights up to
<quantity id="4CAB9B2CFFFBFF84FAFC2CB4FA29F8AC" box="[1360,1432,1771,1797]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="161" unit="m" value="350.0">350 m</quantity>
, often around flat-top elevation
</emphasis>
” (
<figureCitation id="13682A4CFFFBFF84FDA52D50FDECF880" box="[521,605,1807,1833]" captionStart="FIGURES 32" captionStartId="9.[151,264,981,1005]" captionTargetBox="[227,1359,181,957]" captionTargetId="figure-19@9.[227,1359,181,957]" captionTargetPageId="9" captionText="FIGURES 32. The type locality of Axenyllodes chinki sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14797998" httpUri="https://zenodo.org/record/14797998/files/figure.png" pageId="7" pageNumber="161">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="8BEC36C9FFFBFF8AFF6B2D6CFEC3FB6C" blockId="7.[151,1437,259,2014]" lastBlockId="9.[151,1437,1087,1690]" lastPageId="9" lastPageNumber="163" pageId="7" pageNumber="161">
<emphasis id="B927EADBFFFBFF84FF6B2D6CFE84F8E4" bold="true" box="[199,309,1843,1869]" pageId="7" pageNumber="161">Affinities</emphasis>
. Currently, there are only five known species in the genus characterized by such a small number of chaetae (7 or 6) on the tibiotarsi as in
<taxonomicName id="4C534D4AFFFBFF84FDEA2D08FD1BF8D8" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[582,682,1879,1905]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFFBFF84FDEA2D08FD1BF8D8" box="[582,682,1879,1905]" italics="true" pageId="7" pageNumber="161">A. chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFFBFF84FD1E2D08FCBDF8D8" bold="true" box="[690,780,1879,1905]" pageId="7" pageNumber="161">
<taxonomicNameLabel id="A21457A0FFFBFF84FD1E2D08FCBDF8D8" box="[690,780,1879,1905]" pageId="7" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<taxonomicName id="4C534D4AFFFBFF84FCB02D08FC16F8D8" box="[796,935,1879,1905]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="echinatus">
<emphasis id="B927EADBFFFBFF84FCB02D08FC16F8D8" box="[796,935,1879,1905]" italics="true" pageId="7" pageNumber="161">A. echinatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C534D4AFFFBFF84FC1A2D08FAB5F8DB" authority="Fjellberg, 1995" authorityName="Fjellberg" authorityYear="1995" box="[950,1284,1879,1906]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="nematodes">
<emphasis id="B927EADBFFFBFF84FC1A2D08FBFFF8D8" box="[950,1102,1879,1905]" italics="true" pageId="7" pageNumber="161">A. nematodes</emphasis>
<bibRefCitation id="EFC24B38FFFBFF84FBF92D08FAB5F8DB" author="Fjellberg, A." box="[1109,1284,1879,1906]" pageId="7" pageNumber="161" pagination="153 - 158" refId="ref12502" refString="Fjellberg, A. (1995) Collembola of the Canary Islands. II. Family Odontellidae. Entomologica Scandinavica, 26, 153-158. https://doi.org/10.1163/187631295X00152" type="journal article" year="1995">Fjellberg, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C534D4AFFFBFF84FAA32D08FF5DF83F" authority="Tamura" authorityName="Tamura" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B927EADBFFFBFF84FAA32D08FA2DF8D8" box="[1295,1436,1879,1905]" italics="true" pageId="7" pageNumber="161">A. japonicus</emphasis>
Tamura
</taxonomicName>
in
<bibRefCitation id="EFC24B38FFFBFF84FEA12D23FE47F83C" author="Tamura, H. &amp; Yue, Q." box="[269,502,1915,1942]" pageId="7" pageNumber="161" pagination="47 - 53" refId="ref13749" refString="Tamura, H. &amp; Yue, Q. (1999) Two new species of the genus Axenyllodes from Japan and China (Collembola: Pseudachorutidae). Edaphologia, 62, 47-53." type="journal article" year="1999">Tamura &amp; Yue, 1999</bibRefCitation>
,
<taxonomicName id="4C534D4AFFFBFF84FE532D24FDC7F83C" box="[511,630,1915,1941]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B927EADBFFFBFF84FE532D24FDC7F83C" box="[511,630,1915,1941]" italics="true" pageId="7" pageNumber="161">A. sinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D4AFFFBFF84FD062D24FC49F83C" authority="Thibaud, 2011" authorityName="Thibaud" authorityYear="2011" box="[682,1016,1915,1941]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="comoriensis">
<emphasis id="B927EADBFFFBFF84FD062D24FCFFF83C" box="[682,846,1915,1941]" italics="true" pageId="7" pageNumber="161">A. comoriensis</emphasis>
<bibRefCitation id="EFC24B38FFFBFF84FCF82D24FC49F83C" author="Thibaud, J. - M." box="[852,1016,1915,1941]" pageId="7" pageNumber="161" pagination="5 - 12" refId="ref14050" refString="Thibaud, J. - M. (2011) Les Collemboles des sables littoraux de l'Ile de la Grande Comore. Revue francaise d'Entomologie, New Series, 33 (1 - 4), 5-12." type="journal article" year="2011">Thibaud, 2011</bibRefCitation>
</taxonomicName>
. Most of them have a reduced number of ocelli (01). The only exception is
<taxonomicName id="4C534D4AFFFBFF84FDEF2DC0FD0DF810" box="[579,700,1951,1977]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B927EADBFFFBFF84FDEF2DC0FD0DF810" box="[579,700,1951,1977]" italics="true" pageId="7" pageNumber="161">A. sinensis</emphasis>
</taxonomicName>
, described from the mountainous regions of southwestern
<collectingCountry id="F3447659FFFBFF84FAFE2DC0FA29F810" box="[1362,1432,1951,1977]" name="China" pageId="7" pageNumber="161">China</collectingCountry>
, which, like
<taxonomicName id="4C534D4AFFFBFF84FEB32D9CFE35F874" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[287,388,1987,2013]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="7" pageNumber="161" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFFBFF84FEB32D9CFE35F874" box="[287,388,1987,2013]" italics="true" pageId="7" pageNumber="161">A. chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFFBFF84FE222D9CFE5BF874" bold="true" box="[398,490,1987,2013]" pageId="7" pageNumber="161">
<taxonomicNameLabel id="A21457A0FFFBFF84FE222D9CFE5BF874" box="[398,490,1987,2013]" pageId="7" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has 2+2 ocelli, almost identical dorsal and ventral chaetotaxies, as well as similar pattern of the integument granulation. These two species can easily be distinguished by the shape of the PAO (a single triangular lobe in
<taxonomicName id="4C534D4AFFF5FF8AFE022E3BFD99FBD4" box="[430,552,1124,1150]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B927EADBFFF5FF8AFE022E3BFD99FBD4" box="[430,552,1124,1150]" italics="true" pageId="9" pageNumber="163">A. sinensis</emphasis>
</taxonomicName>
,
<emphasis id="B927EADBFFF5FF8AFD9A2E3BFDFCFBD4" box="[566,589,1124,1149]" italics="true" pageId="9" pageNumber="163">vs</emphasis>
broadly oval PAO with pointed edges, sunken in a deep cavity and hardly visible from above in
<taxonomicName id="4C534D4AFFF5FF8AFE3E2ED7FE46FB08" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[402,503,1159,1186]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFE3E2ED7FE46FB08" box="[402,503,1159,1186]" italics="true" pageId="9" pageNumber="163">A. chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF5FF8AFE532ED7FDEBFB0B" bold="true" box="[511,602,1160,1186]" pageId="9" pageNumber="163">
<taxonomicNameLabel id="A21457A0FFF5FF8AFE532ED7FDEBFB0B" box="[511,602,1160,1186]" pageId="9" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), as well as the presence of anal spines and the absence of clavate tenent setae in
<emphasis id="B927EADBFFF5FF8AFF5F2EF3FEC3FB6C" box="[243,370,1196,1222]" italics="true" pageId="9" pageNumber="163">
<taxonomicName id="4C534D4AFFF5FF8AFF5F2EF3FEDFFB6C" box="[243,366,1196,1222]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="sinensis">A. sinensis</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="DF2C6641FFF4FF8BFF3B2DC8FB4BF85E" ID-DOI="http://doi.org/10.5281/zenodo.14797996" ID-Zenodo-Dep="14797996" httpUri="https://zenodo.org/record/14797996/files/figure.png" pageId="8" pageNumber="162" startId="8.[151,264,1943,1967]" targetBox="[191,1396,195,1905]" targetPageId="8" targetType="figure">
<paragraph id="8BEC36C9FFF4FF8BFF3B2DC8FB4BF85E" blockId="8.[151,1436,1943,2040]" pageId="8" pageNumber="162">
<emphasis id="B927EADBFFF4FF8BFF3B2DC8FEFCF806" bold="true" box="[151,333,1943,1967]" pageId="8" pageNumber="162">FIGURES 2431</emphasis>
.
<taxonomicName id="4C534D4AFFF4FF8BFEF52DC7FDA6F819" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[345,535,1944,1968]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="8" pageNumber="162" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFF4FF8BFEF52DC7FDA6F819" box="[345,535,1944,1968]" italics="true" pageId="8" pageNumber="162">Axenyllodes chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF4FF8BFDB02DC7FDDDF819" bold="true" box="[540,620,1944,1968]" pageId="8" pageNumber="162">
<taxonomicNameLabel id="A21457A0FFF4FF8BFDB02DC7FDDDF819" box="[540,620,1944,1968]" pageId="8" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 24, general habitus; 25, cuticular granulation and dorsal chaetotaxy; 26 and 26a, PAO and ocelli; 27, chaetotaxy of antenna, dorsal view; 28, ventral chaetotaxy of head; 29, ventral chaetotaxy of abdomen; 30, tibiotarsi III; 30a, tip of leg III; 31, furca, lateral view. Scales: 2425, 2829, 0.1 mm, 2627, 3031, 0.01 mm.
</paragraph>
</caption>
<caption id="DF2C6641FFF5FF8AFF3B298AFCA6FC44" ID-DOI="http://doi.org/10.5281/zenodo.14797998" ID-Zenodo-Dep="14797998" box="[151,791,981,1006]" httpUri="https://zenodo.org/record/14797998/files/figure.png" pageId="9" pageNumber="163" startId="9.[151,264,981,1005]" targetBox="[227,1359,181,957]" targetPageId="9" targetType="figure">
<paragraph id="8BEC36C9FFF5FF8AFF3B298AFCA6FC44" blockId="9.[151,791,981,1006]" box="[151,791,981,1006]" pageId="9" pageNumber="163">
<emphasis id="B927EADBFFF5FF8AFF3B298AFE96FC44" bold="true" box="[151,295,981,1005]" pageId="9" pageNumber="163">FIGURES 32</emphasis>
. The type locality of
<taxonomicName id="4C534D4AFFF5FF8AFDAD298AFD71FC44" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[513,704,981,1005]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFDAD298AFD71FC44" box="[513,704,981,1005]" italics="true" pageId="9" pageNumber="163">Axenyllodes chinki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457A0FFF5FF8AFD6A298AFCA6FC44" box="[710,791,981,1005]" pageId="9" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BEC36C9FFF5FF8AFF6B2E8FFE1FFA9B" blockId="9.[151,1437,1087,1690]" pageId="9" pageNumber="163">
One more species,
<taxonomicName id="4C534D4AFFF5FF8AFE0A2E8FFD9BFB40" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[422,554,1232,1258]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="potapovi" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFE0A2E8FFD9BFB40" box="[422,554,1232,1258]" italics="true" pageId="9" pageNumber="163">A. potapovi</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF5FF8AFD982E8FFD21FB43" bold="true" box="[564,656,1232,1258]" pageId="9" pageNumber="163">
<taxonomicNameLabel id="A21457A0FFF5FF8AFD982E8FFD21FB43" box="[564,656,1232,1258]" pageId="9" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, also characterized by a strong reduction of the tibiotarsal setae, is described below. It can easily be distinguished from
<taxonomicName id="4C534D4AFFF5FF8AFD472EABFCFEFAA4" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[747,847,1267,1294]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFD472EABFCFEFAA4" box="[747,847,1267,1294]" italics="true" pageId="9" pageNumber="163">A. chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF5FF8AFCF42EABFC03FAA7" bold="true" box="[856,946,1268,1294]" pageId="9" pageNumber="163">
<taxonomicNameLabel id="A21457A0FFF5FF8AFCF42EABFC03FAA7" box="[856,946,1268,1294]" pageId="9" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the complete absence of eyes (for other differences, see
<tableCitation id="C6D10372FFF5FF8AFEE72F48FE2CFA9B" box="[331,413,1303,1330]" captionStart="TABLE 1" captionStartId="13.[151,243,474,500]" captionText="TABLE 1. Main diagnostic characters of the known species of the genus Axenyllodes Stach, 1949." pageId="9" pageNumber="163">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BEC36C9FFF5FF8AFF6B2F64FD65FA4C" blockId="9.[151,1437,1087,1690]" pageId="9" pageNumber="163">
Unfortunately, the number of tibiotarsal setae was not specified in two known species of the genus, i.e.
<taxonomicName id="4C534D4AFFF5FF8AFA9E2F63FA2DFAFC" baseAuthorityName="Gisin" baseAuthorityYear="1952" box="[1330,1436,1340,1366]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="B927EADBFFF5FF8AFA9E2F63FA2DFAFC" box="[1330,1436,1340,1366]" italics="true" pageId="9" pageNumber="163">A. caecus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C534D4AFFF5FF8AFF6A2F3FFD1CFAD3" authority="(Jordana &amp; Ardanaz, 1981)" baseAuthorityName="Jordana &amp; Ardanaz" baseAuthorityYear="1981" box="[198,685,1375,1402]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="monoculatus">
<emphasis id="B927EADBFFF5FF8AFF6A2F3FFEC5FAD0" box="[198,372,1375,1402]" italics="true" pageId="9" pageNumber="163">A. monoculatus</emphasis>
(
<bibRefCitation id="EFC24B38FFF5FF8AFE2E2F00FD15FAD3" author="Jordana, R. &amp; Ardanaz, A." box="[386,676,1375,1402]" pageId="9" pageNumber="163" pagination="33 - 45" refId="ref12643" refString="Jordana, R., Ardanaz, A. (1981) Xenyllodes monoculatus, n. sp., Uzelia setifera clavata n. ssp. y Folsomia sexoculata trisetata n. ssp. (Collembola) de Navarra. Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica, 6, 33-45." type="journal article" year="1981">Jordana &amp; Ardanaz, 1981</bibRefCitation>
)
</taxonomicName>
, in the original descriptions or subsequently, although the Canarian specimen identified by A. Fjellberg as
<taxonomicName id="4C534D4AFFF5FF8AFDE12FDBFC91FA34" baseAuthorityName="Jordana &amp; Ardanaz" baseAuthorityYear="1981" box="[589,800,1411,1438]" class="Collembola" family="Odontellidae" genus="Axenyllodes" isUncertain="true" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="monoculatus">
<emphasis id="B927EADBFFF5FF8AFDE12FDBFDEFFA37" box="[589,606,1412,1438]" italics="true" pageId="9" pageNumber="163">A</emphasis>
. cf.
<emphasis id="B927EADBFFF5FF8AFD3D2FDCFC91FA34" box="[657,800,1411,1437]" italics="true" pageId="9" pageNumber="163">monoculatus</emphasis>
</taxonomicName>
was noted to have 10109 setae on the tibiotarsi and 7 femoral setae (
<bibRefCitation id="EFC24B38FFF5FF8AFE902FF8FE5DFA6B" author="Fjellberg, A." box="[316,492,1447,1474]" pageId="9" pageNumber="163" pagination="153 - 158" refId="ref12502" refString="Fjellberg, A. (1995) Collembola of the Canary Islands. II. Family Odontellidae. Entomologica Scandinavica, 26, 153-158. https://doi.org/10.1163/187631295X00152" type="journal article" year="1995">Fjellberg, 1995</bibRefCitation>
, p. 156). Both of them have a reduced number of ocelli (less than 2+2):
<taxonomicName id="4C534D4AFFF5FF8AFABB2FF7FA33FA68" baseAuthorityName="Gisin" baseAuthorityYear="1952" box="[1303,1410,1448,1474]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="B927EADBFFF5FF8AFABB2FF7FA33FA68" box="[1303,1410,1448,1474]" italics="true" pageId="9" pageNumber="163">A. caecus</emphasis>
</taxonomicName>
is blind, whereas
<taxonomicName id="4C534D4AFFF5FF8AFEED2F93FE41FA4C" baseAuthorityName="Jordana &amp; Ardanaz" baseAuthorityYear="1981" box="[321,496,1483,1510]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="monoculatus">
<emphasis id="B927EADBFFF5FF8AFEED2F93FE41FA4C" box="[321,496,1483,1510]" italics="true" pageId="9" pageNumber="163">A. monoculatus</emphasis>
</taxonomicName>
has only 1+1 ocelli.
</paragraph>
<paragraph id="8BEC36C9FFF5FF8AFF6B2FB0FEB0F933" blockId="9.[151,1437,1087,1690]" pageId="9" pageNumber="163">
Using the most recent key to species of the genus (
<bibRefCitation id="EFC24B38FFF5FF8AFD582FB0FC24F9A0" author="Thibaud, J. - M." box="[756,917,1519,1545]" pageId="9" pageNumber="163" pagination="111 - 116" refId="ref14012" refString="Thibaud, J. - M. (2006) Etude des Collemboles des sables littoraux de Bretagne (France). Revue francaise d'Entomologie, New Series, 28 (3), 111-116." type="journal article" year="2006">Thibaud 2006</bibRefCitation>
), the new species can be identified as
<taxonomicName id="4C534D4AFFF5FF8AFA9B2FAFFA24F9A0" authorityName="Thibaud &amp; Peja" authorityYear="1996" box="[1335,1429,1520,1546]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="marci">
<emphasis id="B927EADBFFF5FF8AFA9B2FAFFA24F9A0" box="[1335,1429,1520,1546]" italics="true" pageId="9" pageNumber="163">A. marci</emphasis>
</taxonomicName>
, described from a sandy beach in
<collectingCountry id="F3447659FFF5FF8AFDA82C4CFDD1F984" box="[516,608,1555,1581]" name="Albania" pageId="9" pageNumber="163">Albania</collectingCountry>
, which, like
<taxonomicName id="4C534D4AFFF5FF8AFD412C4BFCE1F984" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[749,848,1555,1582]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFD412C4BFCE1F984" box="[749,848,1555,1582]" italics="true" pageId="9" pageNumber="163">A. chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF5FF8AFCFB2C4BFC00F987" bold="true" box="[855,945,1556,1582]" pageId="9" pageNumber="163">
<taxonomicNameLabel id="A21457A0FFF5FF8AFCFB2C4BFC00F987" box="[855,945,1556,1582]" pageId="9" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, has 2+2 ocelli and lacks anal spines. These two species clearly differ in the chaetotaxy of the thoracic terga (
<taxonomicName id="4C534D4AFFF5FF8AFCDB2C67FC66F9F8" authorityName="Thibaud &amp; Peja" authorityYear="1996" box="[887,983,1592,1618]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="marci">
<emphasis id="B927EADBFFF5FF8AFCDB2C67FC66F9F8" box="[887,983,1592,1618]" italics="true" pageId="9" pageNumber="163">A. marci</emphasis>
</taxonomicName>
is characterized by the absence of setae m1 and the presence of lateral ms on Th. IIIII) and the number of tibiotarsal setae (
<quantity id="4CAB9B2CFFF5FF8AFB922C03FBC6F9DC" box="[1086,1143,1627,1654]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="163" unit="in" value="10.0">10 in</quantity>
<taxonomicName id="4C534D4AFFF5FF8AFBD22C03FB6DF9DC" authorityName="Thibaud &amp; Peja" authorityYear="1996" box="[1150,1244,1628,1654]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="marci">
<emphasis id="B927EADBFFF5FF8AFBD22C03FB6DF9DC" box="[1150,1244,1628,1654]" italics="true" pageId="9" pageNumber="163">A. marci</emphasis>
</taxonomicName>
,
<emphasis id="B927EADBFFF5FF8AFB462C03FAB0F9DC" box="[1258,1281,1628,1653]" italics="true" pageId="9" pageNumber="163">vs</emphasis>
<quantity id="4CAB9B2CFFF5FF8AFAA42C03FA82F9DC" box="[1288,1331,1627,1654]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="9" pageNumber="163" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="4C534D4AFFF5FF8AFA952C03FA2DF9DC" authorityName="Babenko &amp; Shveenkova &amp; Arbea" authorityYear="2025" box="[1337,1436,1627,1654]" class="Collembola" family="Odontellidae" genus="Axenyllodes" kingdom="Animalia" order="Poduromorpha" pageId="9" pageNumber="163" phylum="Arthropoda" rank="species" species="chinki" status="sp. nov.">
<emphasis id="B927EADBFFF5FF8AFA952C03FA2DF9DC" box="[1337,1436,1627,1654]" italics="true" pageId="9" pageNumber="163">A. chinki</emphasis>
</taxonomicName>
<emphasis id="B927EADBFFF5FF8AFF3B2CDFFF40F933" bold="true" box="[151,241,1664,1690]" pageId="9" pageNumber="163">
<taxonomicNameLabel id="A21457A0FFF5FF8AFF3B2CDFFF40F933" box="[151,241,1664,1690]" pageId="9" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,330 @@
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<mods:title id="D776C88527FDB7DFA746816BA1F1CC12">Lost for more than 85 years-rediscovery of Channa amphibeus (McClelland, 1845), the worlds most elusive snakehead species (Teleostei, Labyrinthici, Channidae)</mods:title>
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<mods:namePart id="31691B2B93198407ABF3A6CADE1A3F38">Moulitharan, Nallathambi</mods:namePart>
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<treatment id="2C0087A0FFE74C68FF76FBD4776695F9" ID-DOI="http://doi.org/10.5281/zenodo.14797724" ID-Zenodo-Dep="14797724" LSID="urn:lsid:plazi:treatment:2C0087A0FFE74C68FF76FBD4776695F9" httpUri="http://treatment.plazi.org/id/2C0087A0FFE74C68FF76FBD4776695F9" lastPageId="6" lastPageNumber="93" pageId="2" pageNumber="89" scope_family="Channidae">
<subSubSection id="ECB3653DFFE74C6CFF76FBD4746F932C" box="[151,635,1122,1149]" pageId="2" pageNumber="89" type="nomenclature">
<paragraph id="A41636B6FFE74C6CFF76FBD4746F932C" blockId="2.[151,635,1122,1184]" box="[151,635,1122,1149]" pageId="2" pageNumber="89">
<heading id="FF5E81DAFFE74C6CFF76FBD4746F932C" bold="true" box="[151,635,1122,1149]" fontSize="11" level="1" pageId="2" pageNumber="89" reason="1">
<taxonomicName id="63A94D35FFE74C6CFF76FBD4746F932C" ID-CoL="TM2G" authority="(McClelland, 1845)" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[151,635,1122,1149]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="89" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE74C6CFF76FBD4746F932C" bold="true" box="[151,635,1122,1149]" pageId="2" pageNumber="89">
<emphasis id="96DDEAA4FFE74C6CFF76FBD47790932D" bold="true" box="[151,388,1122,1149]" italics="true" pageId="2" pageNumber="89">Channa amphibeus</emphasis>
(
<bibRefCitation id="C0384B47FFE74C6CFE75FBD47467932C" author="McClelland, J." box="[404,627,1122,1148]" pageId="2" pageNumber="89" pagination="274 - 282" refId="ref8002" refString="McClelland, J. (1845) Description of four species of fishes from the rivers at the foot of Boutan Mountains. Journal of Natural History, Calcutta, 5 (18), 274-282." type="journal article" year="1845">McClelland, 1845</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECB3653DFFE74C6CFF76FB30777493F0" box="[151,352,1158,1184]" pageId="2" pageNumber="89" type="description">
<paragraph id="A41636B6FFE74C6CFF76FB30777493F0" blockId="2.[151,635,1122,1184]" box="[151,352,1158,1184]" pageId="2" pageNumber="89">
(
<figureCitation id="3C922A33FFE74C6CFF7EFB3076F993F0" box="[159,237,1158,1184]" captionStart="FIGURE 1" captionStartId="3.[151,250,1184,1208]" captionTargetBox="[353,1232,349,1152]" captionTargetId="figure-100@3.[347,1240,342,1158]" captionTargetPageId="3" captionText="FIGURE 1. Channa amphibeus illustrations. (A) Drawing of a specimen misidentified as Ophicephalus barca in McClelland 1840, plate xi, fig. 3, (B) Coloured illustration of C. amphibeus reproduced from Shaw &amp; Shebbeare (1938), plate 1. (C) Monochrome photograph of C. amphibeus reproduced from Shaw &amp; Shebbeare (1938), plate 6 (Image donated by: Ralf Britz)." figureDoi="http://doi.org/10.5281/zenodo.14798018" httpUri="https://zenodo.org/record/14798018/files/figure.png" pageId="2" pageNumber="89">Figs. 1</figureCitation>
,
<figureCitation id="3C922A33FFE74C6CFF1AFB30771F93F0" box="[251,267,1158,1184]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="2" pageNumber="89">2</figureCitation>
,
<figureCitation id="3C922A33FFE74C6CFEF6FB30773193F0" box="[279,293,1158,1184]" captionStart="FIGURE 3" captionStartId="5.[152,255,1006,1030]" captionTargetBox="[340,1247,181,982]" captionTargetId="figure-19@5.[340,1247,181,982]" captionTargetPageId="5" captionText="FIGURE 3. Channa amphibeus topotypes, A. lateral, dorsal, and ventral view, KUFOS-AN-2024.12, 205.0 mm SL, colouration after preservation; B. BNHS FWF 1186, 420 mm SL, colouration immediately after exposure to formalin." figureDoi="http://doi.org/10.5281/zenodo.14798022" httpUri="https://zenodo.org/record/14798022/files/figure.png" pageId="2" pageNumber="89">3</figureCitation>
&amp;
<figureCitation id="3C922A33FFE74C6CFEA8FB30774C93F0" box="[329,344,1158,1184]" captionStart="FIGURE 7" captionStartId="8.[152,255,846,870]" captionTargetBox="[340,1247,181,822]" captionTargetId="figure-21@8.[340,1247,181,822]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of head, (A &amp; B) Channa amphibeus, ca. 270 and 500 mm SL, showing neon-blue portion (square region) and maroon blotches; C &amp; D) Channa barca, ca. 200 &amp; 500 mm SL, showing black blotches and absence of neon-blue portion (Photo credits, C &amp; D: Heok Hui Tan and Dominik Niemeier)." figureDoi="http://doi.org/10.5281/zenodo.14798030" httpUri="https://zenodo.org/record/14798030/files/figure.png" pageId="2" pageNumber="89">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ECB3653DFFE74C6CFF76FB7873979204" pageId="2" pageNumber="89" type="materials_examined">
<paragraph id="A41636B6FFE74C6CFF76FB78727E9260" blockId="2.[151,1437,1230,2013]" pageId="2" pageNumber="89">
<emphasis id="96DDEAA4FFE74C6CFF76FB78779593B8" bold="true" box="[151,385,1230,1256]" pageId="2" pageNumber="89">Material examined.</emphasis>
<taxonomicName id="63A94D35FFE74C6CFE68FB78747693B8" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[393,610,1230,1256]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="89" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE74C6CFE68FB78747693B8" box="[393,610,1230,1256]" italics="true" pageId="2" pageNumber="89">Channa amphibeus</emphasis>
</taxonomicName>
,
<materialsCitation id="14C13CEBFFE74C6CFD90FB7877789260" collectingDate="2024-09-26" collectionCode="BNHS" collectorName="J. Praveenraj" country="India" location="Chel River" municipality="Kalimpong District" pageId="2" pageNumber="89" specimenCode="FWF 1185-86, 180" specimenCount="1" stateProvince="West Bengal">
KUFOS-AN-2024.12, 205.0 mm SL, collected from a local fisher at the
<location id="A176606DFFE74C6CFF76FB447701925C" LSID="urn:lsid:plazi:treatment:2C0087A0FFE74C68FF76FBD4776695F9:A176606DFFE74C6CFF76FB447701925C" box="[151,277,1266,1292]" country="India" municipality="Kalimpong District" name="Chel River" pageId="2" pageNumber="89" stateProvince="West Bengal">Chel River</location>
, near Gorubathan,
<collectingMunicipality id="4472ACCCFFE74C6CFE0EFB4474D9925C" box="[495,717,1266,1292]" pageId="2" pageNumber="89">Kalimpong District</collectingMunicipality>
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,
<collectingCountry id="DCBE7626FFE74C6CFC98FB4475A1925C" box="[889,949,1266,1292]" name="India" pageId="2" pageNumber="89">India</collectingCountry>
,
<collectorName id="095C5360FFE74C6CFC23FB457241925C" box="[962,1109,1266,1293]" pageId="2" pageNumber="89">J. Praveenraj</collectorName>
&amp; team,
<date id="D0171076FFE74C6CFB22FB447351925C" box="[1219,1349,1266,1292]" pageId="2" pageNumber="89" value="2024-09-26">
<collectingDate id="C053E99EFFE74C6CFB22FB447351925C" box="[1219,1349,1266,1292]" pageId="2" pageNumber="89" value="2024-09-26">26.09.2024</collectingDate>
</date>
;
<collectionCode id="C2B8AE73FFE74C6CFAB3FB447389925C" box="[1362,1437,1266,1292]" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="2" pageNumber="89">BNHS</collectionCode>
<specimenCode id="F40F9ECDFFE74C6CFF76FAA177239261" box="[151,311,1302,1329]" collectionCode="FWF" pageId="2" pageNumber="89">FWF 1185-86</specimenCode>
,
<specimenCode id="F40F9ECDFFE74C6CFEA3FAA077789260" box="[322,364,1302,1328]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="2" pageNumber="89">180</specimenCode>
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&amp;
<quantity id="63519B53FFE74C6CFE71FAA077F99261" box="[400,493,1302,1329]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="89" unit="mm" value="420.0">420 mm</quantity>
SL, 2 ex, collection location same as above,
<date id="D0171076FFE74C6CFC07FAA072729260" box="[998,1126,1302,1328]" pageId="2" pageNumber="89" value="2024-11-26">26.11.2024</date>
.
</paragraph>
<paragraph id="A41636B6FFE74C6CFF26FA8C73979204" blockId="2.[151,1437,1230,2013]" box="[199,1411,1338,1365]" pageId="2" pageNumber="89">
<emphasis id="96DDEAA4FFE74C6CFF26FA8C77CE9204" bold="true" box="[199,474,1338,1364]" pageId="2" pageNumber="89">Photographs examined</emphasis>
:
<taxonomicName id="63A94D35FFE74C6CFE08FA8C74D59204" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[489,705,1338,1364]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="89" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE74C6CFE08FA8C74D59204" box="[489,705,1338,1364]" italics="true" pageId="2" pageNumber="89">Channa amphibeus</emphasis>
</taxonomicName>
, ca. 270, 370, and
<quantity id="63519B53FFE74C6CFC77FA8C75E79205" box="[918,1011,1338,1365]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="89" unit="mm" value="500.0">500 mm</quantity>
SL, location details same as above.
</paragraph>
</subSubSection>
<subSubSection id="ECB3653DFFE74C68FF26FAE877F39785" lastPageId="6" lastPageNumber="93" pageId="2" pageNumber="89" type="description">
<paragraph id="A41636B6FFE74C6DFF26FAE877C99611" blockId="2.[151,1437,1230,2013]" lastBlockId="3.[151,1437,151,321]" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
<emphasis id="96DDEAA4FFE74C6CFF26FAE8774D9228" bold="true" box="[199,345,1374,1400]" pageId="2" pageNumber="89">Description.</emphasis>
See
<figureCitation id="3C922A33FFE74C6CFE79FAE877FF9228" box="[408,491,1374,1400]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="2" pageNumber="89">Figs. 2</figureCitation>
,
<figureCitation id="3C922A33FFE74C6CFE1AFAE8741E9228" box="[507,522,1374,1400]" captionStart="FIGURE 3" captionStartId="5.[152,255,1006,1030]" captionTargetBox="[340,1247,181,982]" captionTargetId="figure-19@5.[340,1247,181,982]" captionTargetPageId="5" captionText="FIGURE 3. Channa amphibeus topotypes, A. lateral, dorsal, and ventral view, KUFOS-AN-2024.12, 205.0 mm SL, colouration after preservation; B. BNHS FWF 1186, 420 mm SL, colouration immediately after exposure to formalin." figureDoi="http://doi.org/10.5281/zenodo.14798022" httpUri="https://zenodo.org/record/14798022/files/figure.png" pageId="2" pageNumber="89">3</figureCitation>
,
<figureCitation id="3C922A33FFE74C6CFDFDFAE874689229" box="[540,636,1374,1401]" captionStart="FIGURE 7" captionStartId="8.[152,255,846,870]" captionTargetBox="[340,1247,181,822]" captionTargetId="figure-21@8.[340,1247,181,822]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of head, (A &amp; B) Channa amphibeus, ca. 270 and 500 mm SL, showing neon-blue portion (square region) and maroon blotches; C &amp; D) Channa barca, ca. 200 &amp; 500 mm SL, showing black blotches and absence of neon-blue portion (Photo credits, C &amp; D: Heok Hui Tan and Dominik Niemeier)." figureDoi="http://doi.org/10.5281/zenodo.14798030" httpUri="https://zenodo.org/record/14798030/files/figure.png" pageId="2" pageNumber="89">7A &amp; B</figureCitation>
for general appearance and
<tableCitation id="E92B030DFFE74C6CFC2FFAE872319228" box="[974,1061,1374,1400]" captionStart="TABLE 1" captionStartId="9.[151,239,151,175]" captionText="TABLE 1. Morphometric data of Channa amphibeus (n=3) (BNHS FWF 1185-86 &amp; KUFOS-AN-2024.12; topotypes from Chel River)." pageId="2" pageNumber="89">Table 1</tableCitation>
for morphometric data. A large snakehead species reaching at least
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SL (present study), but reported by local fishers to reach
<quantity id="63519B53FFE74C6CFAEEFA34737992CD" box="[1295,1389,1410,1437]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="89" unit="mm" value="800.0">800 mm</quantity>
TL. Body elongate, round in cross section anteriorly, laterally compressed at caudal peduncle, body depth 5.9 times SL. Dorsal profile convex, rising evenly from tip of snout to dorsal-fin origin, then almost straight to base of caudal fin. Ventral contour descending gently until below opercle, almost straight to base of caudal fin. Head large, length 3.5 times SL. Head widest, halfway between posterior margin of eye and opercle. Eyes located on anterior half of head, closer to snout. Interorbital region somewhat bulging. Mouth large, posterior end of maxilla and premaxilla extending beyond posterior margin of eye. Lower jaw slightly protruding beyond upper jaw, lips fleshy. Pectoral-fin insertion almost at vertical through origin of dorsal fin. Body depth and width greatest at dorsal-fin origin. Dorsal- and anal-fin bases long, 69.9% and 46.7% SL, respectively. Premaxilla with multiple rows of small, sharp, pointed teeth. Dentary with two rows of teeth, inner row with medium- to large canines; outer row with mediumsized, villiform teeth terminating in three rows at jaw symphysis. Vomer with an inner series of large conical teeth and outer series of much smaller teeth. Palatine with external series of smaller teeth and internal row of up to nine much larger canines. Dorsal-fin rays 4852, anal-fin rays 3336. Caudal fin with 67+67 principal rays, and 3 dorsal and 4 ventral procurrent rays. Pectoral-fin rays 15, pelvic-fin rays 6. Lateral line with 7593 pored scales extending from shoulder girdle in horizontal line up to caudal peduncle, continuous, not dropping a scale row after pre-drop, plus 34 non-pored scales on caudal-fin base; transverse scale rows in dorsoventral direction 7.5/1/13.5 14.5, circumpeduncular scales 3034, cheek scales 8, prepelvic scales 1728, predorsal scales 1619. Vertebrae 53 (
<emphasis id="96DDEAA4FFE64C6DFF40FF2176BB97E0" box="[161,175,151,176]" italics="true" pageId="3" pageNumber="90">n</emphasis>
=1) with 17 abdominal and 36 caudal vertebrae. Adult male (
<figureCitation id="3C922A33FFE64C6DFCB4FF2175A297E1" box="[853,950,151,177]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="3" pageNumber="90">Figs. 2D</figureCitation>
&amp;
<figureCitation id="3C922A33FFE64C6DFC38FF2175E997E1" box="[985,1021,151,177]" captionStart="FIGURE 7" captionStartId="8.[152,255,846,870]" captionTargetBox="[340,1247,181,822]" captionTargetId="figure-21@8.[340,1247,181,822]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of head, (A &amp; B) Channa amphibeus, ca. 270 and 500 mm SL, showing neon-blue portion (square region) and maroon blotches; C &amp; D) Channa barca, ca. 200 &amp; 500 mm SL, showing black blotches and absence of neon-blue portion (Photo credits, C &amp; D: Heok Hui Tan and Dominik Niemeier)." figureDoi="http://doi.org/10.5281/zenodo.14798030" httpUri="https://zenodo.org/record/14798030/files/figure.png" pageId="3" pageNumber="90">7B</figureCitation>
): Head wedge-shaped in dorsal view. Cheeks bulging laterally. Predorsal profile ascending very steeply in straight line from tip of snout to nape, gently convex more posteriorly up to dorsal-fin origin, and thereafter almost straight to base of caudal fin. Ventral contour convex from tip of snout to almost vertical through nape, then gently curved till pelvic-fin origin, thereafter almost straight to base of caudal fin.
</paragraph>
<caption id="F0D6663EFFE64C6DFF76FB1676CD9274" ID-DOI="http://doi.org/10.5281/zenodo.14798018" ID-Zenodo-Dep="14798018" httpUri="https://zenodo.org/record/14798018/files/figure.png" pageId="3" pageNumber="90" startId="3.[151,250,1184,1208]" targetBox="[353,1232,349,1152]" targetPageId="3" targetType="figure">
<paragraph id="A41636B6FFE64C6DFF76FB1676CD9274" blockId="3.[151,1437,1184,1316]" pageId="3" pageNumber="90">
<emphasis id="96DDEAA4FFE64C6DFF76FB16770093E8" bold="true" box="[151,276,1184,1208]" pageId="3" pageNumber="90">FIGURE 1.</emphasis>
<taxonomicName id="63A94D35FFE64C6DFEFAFB1677F493E8" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[283,480,1184,1208]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="90" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE64C6DFEFAFB1677F493E8" box="[283,480,1184,1208]" italics="true" pageId="3" pageNumber="90">Channa amphibeus</emphasis>
</taxonomicName>
illustrations. (A) Drawing of a specimen misidentified as
<taxonomicName id="63A94D35FFE64C6DFBD4FB16731793E8" authorityName="Hamilton" authorityYear="1822" box="[1077,1283,1184,1208]" family="Channidae" genus="Ophicephalus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="90" phylum="Chordata" rank="species" species="barca">
<emphasis id="96DDEAA4FFE64C6DFBD4FB16731793E8" box="[1077,1283,1184,1208]" italics="true" pageId="3" pageNumber="90">Ophicephalus barca</emphasis>
</taxonomicName>
in
<bibRefCitation id="C0384B47FFE64C6DFAC4FB1676D8938D" author="McClelland, J." pageId="3" pageNumber="90" pagination="427 - 429" refId="ref7967" refString="McClelland, J. (1840) Remarks on Miscellaneous Subjects. Journal of Natural History, Calcutta, 1 (3), 427-429, pl. xi, fig 3." type="journal article" year="1840">McClelland 1840</bibRefCitation>
, plate xi, fig. 3, (B) Coloured illustration of
<taxonomicName id="63A94D35FFE64C6DFD4DFB73752F938C" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[684,827,1220,1244]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="90" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE64C6DFD4DFB73752F938C" box="[684,827,1220,1244]" italics="true" pageId="3" pageNumber="90">C. amphibeus</emphasis>
</taxonomicName>
reproduced from
<bibRefCitation id="C0384B47FFE64C6DFC1DFB727301938D" author="Shaw, G. E. &amp; Shebbeare, E. O." box="[1020,1301,1220,1245]" pageId="3" pageNumber="90" pagination="1 - 124" refId="ref8898" refString="Shaw, G. E. &amp; Shebbeare, E. O. (1938) Fishes of Northern Bengal. Journal Royal Asiatic Society of Bengal, 3, 1-124." type="journal article" year="1938">Shaw &amp; Shebbeare (1938)</bibRefCitation>
, plate 1. (C) Monochrome photograph of
<taxonomicName id="63A94D35FFE64C6DFE29FB5F744C9250" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[456,600,1256,1280]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="90" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE64C6DFE29FB5F744C9250" box="[456,600,1256,1280]" italics="true" pageId="3" pageNumber="90">C. amphibeus</emphasis>
</taxonomicName>
reproduced from
<bibRefCitation id="C0384B47FFE64C6DFCFAFB5E72239251" author="Shaw, G. E. &amp; Shebbeare, E. O." box="[795,1079,1256,1281]" pageId="3" pageNumber="90" pagination="1 - 124" refId="ref8898" refString="Shaw, G. E. &amp; Shebbeare, E. O. (1938) Fishes of Northern Bengal. Journal Royal Asiatic Society of Bengal, 3, 1-124." type="journal article" year="1938">Shaw &amp; Shebbeare (1938)</bibRefCitation>
, plate 6 (Image donated by: Ralf Britz).
</paragraph>
</caption>
<paragraph id="A41636B6FFE64C6DFF26FAC073389035" blockId="3.[151,1437,1398,2037]" pageId="3" pageNumber="90">
<emphasis id="96DDEAA4FFE64C6DFF26FAC077B392C0" bold="true" box="[199,423,1398,1424]" pageId="3" pageNumber="90">Colouration in life.</emphasis>
Specimens of
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SL (
<figureCitation id="3C922A33FFE64C6DFCEEFAC175BB92C1" box="[783,943,1399,1425]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="3" pageNumber="90">Figs. 2A, B, C</figureCitation>
&amp;
<figureCitation id="3C922A33FFE64C6DFC2EFAC175E292C1" box="[975,1014,1399,1425]" captionStart="FIGURE 7" captionStartId="8.[152,255,846,870]" captionTargetBox="[340,1247,181,822]" captionTargetId="figure-21@8.[340,1247,181,822]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of head, (A &amp; B) Channa amphibeus, ca. 270 and 500 mm SL, showing neon-blue portion (square region) and maroon blotches; C &amp; D) Channa barca, ca. 200 &amp; 500 mm SL, showing black blotches and absence of neon-blue portion (Photo credits, C &amp; D: Heok Hui Tan and Dominik Niemeier)." figureDoi="http://doi.org/10.5281/zenodo.14798030" httpUri="https://zenodo.org/record/14798030/files/figure.png" pageId="3" pageNumber="90">7A</figureCitation>
): base colour of body iridescent-green with numerous black spots, and sides with 1112 irregular, orange or chrome- to ochre-yellow bars originating from dorsal-fin base and extending beyond lateral-line, but not reaching base of anal fin. The bars equal in width to iridescent-green interbars between them. In some specimens, at mid to posterior portion of body, bars not matching up, incomplete or bifurcating (
<figureCitation id="3C922A33FFE64C6DFE11F9B174479171" box="[496,595,1543,1569]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="3" pageNumber="90">Figs. 2B</figureCitation>
, &amp; C). Dorsal surface of head greenish with numerous maroon to chrome-yellow, elongated spots forming reticulated pattern; side of head also with numerous elongated, chrome-yellow spots. Lips, snout, and postorbital region, neon-blue with rich-brown to maroon, elongated blotches. Gular region bluish. Series of 13, large, irregular, black spots above and posterior to opercular opening at lateral-line origin. Dorsal fin with basal half yellowish to orange, green and black subdistal bands, and narrow, white distal margin; dorsal-fin membrane with numerous black spots, arranged as five horizontal rows along entire fin length. Anal fin iridescent-green, becoming darker towards dorsal margin. Caudal fin with ochre-yellow fin rays, greenish interradial membranes, and narrow white to orange distal margin. Pectoral fin bright-orange with bluish base, and 34 narrow, semi-circular light brown bands; in some specimens a series of orange spots arranged in loose semi-circular lines on fin rays. Pelvic fin with pale-grey rays and greenish interradial membrane becoming darker towards tips.
</paragraph>
<paragraph id="A41636B6FFE64C6AFF26F8D9778B964D" blockId="3.[151,1437,1398,2037]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="91" pageId="3" pageNumber="90">
Specimens of
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SL (
<figureCitation id="3C922A33FFE64C6DFDC9F8D9749E90D9" box="[552,650,1903,1929]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="3" pageNumber="90">Figs. 2D</figureCitation>
,
<figureCitation id="3C922A33FFE64C6DFD75F8D974A390D9" box="[660,695,1903,1929]" captionStart="FIGURE 3" captionStartId="5.[152,255,1006,1030]" captionTargetBox="[340,1247,181,982]" captionTargetId="figure-19@5.[340,1247,181,982]" captionTargetPageId="5" captionText="FIGURE 3. Channa amphibeus topotypes, A. lateral, dorsal, and ventral view, KUFOS-AN-2024.12, 205.0 mm SL, colouration after preservation; B. BNHS FWF 1186, 420 mm SL, colouration immediately after exposure to formalin." figureDoi="http://doi.org/10.5281/zenodo.14798022" httpUri="https://zenodo.org/record/14798022/files/figure.png" pageId="3" pageNumber="90">3B</figureCitation>
,
<figureCitation id="3C922A33FFE64C6DFD20F8D974F190D9" box="[705,741,1903,1929]" captionStart="FIGURE 7" captionStartId="8.[152,255,846,870]" captionTargetBox="[340,1247,181,822]" captionTargetId="figure-21@8.[340,1247,181,822]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of head, (A &amp; B) Channa amphibeus, ca. 270 and 500 mm SL, showing neon-blue portion (square region) and maroon blotches; C &amp; D) Channa barca, ca. 200 &amp; 500 mm SL, showing black blotches and absence of neon-blue portion (Photo credits, C &amp; D: Heok Hui Tan and Dominik Niemeier)." figureDoi="http://doi.org/10.5281/zenodo.14798030" httpUri="https://zenodo.org/record/14798030/files/figure.png" pageId="3" pageNumber="90">7B</figureCitation>
): Body bluish-green with numerous black spots, sides with 16 18, narrow, maroon bars. In some specimens bars completely absent, or barely visible as highly reduced, irregular, broken blotches (
<figureCitation id="3C922A33FFE64C6DFEB8F80177A79081" box="[345,435,1975,2001]" captionStart="FIGURE 2" captionStartId="4.[152,255,1476,1500]" captionTargetBox="[340,1247,309,1453]" captionTargetId="figure-75@4.[340,1247,309,1453]" captionTargetPageId="4" captionText="FIGURE 2. Channa amphibeus, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued)." figureDoi="http://doi.org/10.5281/zenodo.14798020" httpUri="https://zenodo.org/record/14798020/files/figure.png" pageId="3" pageNumber="90">Fig. 2D</figureCitation>
). Head bluish-green with numerous, rounded to elongated maroon spots, more prominent on lips. Snout, lips, and portion below eye neon-blue. Dorsal and caudal fin olive-green with numerous black spots along entire fin length, subdistal margin black, and distal margin narrow yellow and white. Anal fin bluish-green with olive-green to brown fin rays, becoming darker towards margins, and narrow white distal margin; numerous black spots along entire fin length. Pectoral fin brown with orange spots arranged transversely. Pelvic fin bluish-green with greyish tips.
</paragraph>
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<paragraph id="A41636B6FFE14C6AFF79FA7272DC9150" blockId="4.[151,1436,1476,1536]" pageId="4" pageNumber="91">
<emphasis id="96DDEAA4FFE14C6AFF79FA72770D928C" bold="true" box="[152,281,1476,1500]" pageId="4" pageNumber="91">FIGURE 2.</emphasis>
<taxonomicName id="63A94D35FFE14C6AFEFEFA7277F0928C" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[287,484,1476,1500]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE14C6AFEFEFA7277F0928C" box="[287,484,1476,1500]" italics="true" pageId="4" pageNumber="91">Channa amphibeus</emphasis>
</taxonomicName>
, showing live colouration (A) KUFOS-AN-2024.12, 205.0 mm SL, female (B) ca. 270.0 mm SL, male (C) ca. 350 mm SL, male (D) ca. 500 mm SL, adult male (B, C, &amp; D, not fixed and catalogued).
</paragraph>
</caption>
<paragraph id="A41636B6FFE14C6AFF26F9E475E990A8" blockId="4.[151,1437,1618,2041]" pageId="4" pageNumber="91">
<emphasis id="96DDEAA4FFE14C6AFF26F9E4756A913C" bold="true" box="[199,894,1618,1644]" pageId="4" pageNumber="91">
Colour details according to
<bibRefCitation id="C0384B47FFE14C6AFDEEF9E47573913C" author="Shaw, G. E. &amp; Shebbeare, E. O." box="[527,871,1618,1644]" pageId="4" pageNumber="91" pagination="1 - 124" refId="ref8898" refString="Shaw, G. E. &amp; Shebbeare, E. O. (1938) Fishes of Northern Bengal. Journal Royal Asiatic Society of Bengal, 3, 1-124." type="journal article" year="1938">Shaw &amp; Shebbeare (1938: 12</bibRefCitation>
l):
</emphasis>
“A gorgeously coloured fish. The ground-colour is blue when viewed obliquely and iridescent green when viewed at right angles to the surface. On the body this colour is sprinkled with dark spots, uniform in size but irregular in shape. These spots are absent from the belly, sparse below the lateral line and increasingly plentiful towards the back where they coalesce. On the head, the spots are larger and rounded, rich brown below the level of the eye and becoming darker and more plentiful towards the top of the head where they coalesce. The brightest blue (green) and richest brown are in the region of the upper lip. Along the body 1316 more or less irregular vertical bands about equal in width to the interspaces between them and extending from the dorsal to below the lateral line. These are bright orange bordered with brown and merging into brown on the back and are free from black spots. The dorsal has the basal half brown or orange, the outer half blue (green), darkening outwards but having a narrow pale blue or white edge. Pectoral deep orange. Pelvics blue. Anal iridescent blue (green) with a narrower dark border. Caudal brown at the base, then iridescent blue (green) with dark rays, then blackish with a narrow white or bluish-white border” (see
<figureCitation id="3C922A33FFE14C6AFC78F86875FB90A8" box="[921,1007,2014,2040]" captionStart="FIGURE 1" captionStartId="3.[151,250,1184,1208]" captionTargetBox="[353,1232,349,1152]" captionTargetId="figure-100@3.[347,1240,342,1158]" captionTargetPageId="3" captionText="FIGURE 1. Channa amphibeus illustrations. (A) Drawing of a specimen misidentified as Ophicephalus barca in McClelland 1840, plate xi, fig. 3, (B) Coloured illustration of C. amphibeus reproduced from Shaw &amp; Shebbeare (1938), plate 1. (C) Monochrome photograph of C. amphibeus reproduced from Shaw &amp; Shebbeare (1938), plate 6 (Image donated by: Ralf Britz)." figureDoi="http://doi.org/10.5281/zenodo.14798018" httpUri="https://zenodo.org/record/14798018/files/figure.png" pageId="4" pageNumber="91">Fig. 1B</figureCitation>
).
</paragraph>
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<paragraph id="A41636B6FFE04C6BFF79FC5872DD937A" blockId="5.[151,1436,1006,1066]" pageId="5" pageNumber="92">
<emphasis id="96DDEAA4FFE04C6BFF79FC5877039356" bold="true" box="[152,279,1006,1030]" pageId="5" pageNumber="92">FIGURE 3.</emphasis>
<taxonomicName id="63A94D35FFE04C6BFEFDFC5877CB9356" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[284,479,1006,1030]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE04C6BFEFDFC5877CB9356" box="[284,479,1006,1030]" italics="true" pageId="5" pageNumber="92">Channa amphibeus</emphasis>
</taxonomicName>
topotype
<emphasis id="96DDEAA4FFE04C6BFDDDFC5974529356" box="[572,582,1007,1030]" italics="true" pageId="5" pageNumber="92">s</emphasis>
,
<emphasis id="96DDEAA4FFE04C6BFDB0FC58747D9356" bold="true" box="[593,617,1006,1030]" pageId="5" pageNumber="92">A.</emphasis>
lateral, dorsal, and ventral view, KUFOS-AN-2024.12, 205.0 mm SL, colouration after preservation;
<emphasis id="96DDEAA4FFE04C6BFEB7FBA4777A937A" bold="true" box="[342,366,1042,1066]" pageId="5" pageNumber="92">B.</emphasis>
BNHS FWF 1186, 420 mm SL, colouration immediately after exposure to formalin.
</paragraph>
</caption>
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<paragraph id="A41636B6FFE04C6BFF79F91572339054" blockId="5.[151,1436,1699,1796]" pageId="5" pageNumber="92">
<emphasis id="96DDEAA4FFE04C6BFF79F915770D91EB" bold="true" box="[152,281,1699,1723]" pageId="5" pageNumber="92">FIGURE 4.</emphasis>
Map of Northeast India, showing distribution of
<taxonomicName id="63A94D35FFE04C6BFCF2F91575CC91EB" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[787,984,1699,1723]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE04C6BFCF2F91575CC91EB" box="[787,984,1699,1723]" italics="true" pageId="5" pageNumber="92">Channa amphibeus</emphasis>
</taxonomicName>
,
<taxonomicName id="63A94D35FFE04C6BFC04F91272B991EB" box="[997,1197,1699,1723]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="aurantimaculata">
<emphasis id="96DDEAA4FFE04C6BFC04F91272B991EB" box="[997,1197,1699,1723]" italics="true" pageId="5" pageNumber="92">C. aurantimaculata</emphasis>
</taxonomicName>
, and
<emphasis id="96DDEAA4FFE04C6BFB07F912735291EB" box="[1254,1350,1699,1723]" italics="true" pageId="5" pageNumber="92">
<taxonomicName id="63A94D35FFE04C6BFB07F912735591EB" authorityName="Hamilton" authorityYear="1822" box="[1254,1345,1699,1723]" family="Channidae" genus="Ophicephalus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="barca">C. barca</taxonomicName>
.
</emphasis>
(Circles indicate type locality, squares and triangles indicate additional records; for
<taxonomicName id="63A94D35FFE04C6BFC68F97E7200918F" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[905,1044,1735,1759]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="92" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE04C6BFC68F97E7200918F" box="[905,1044,1735,1759]" italics="true" pageId="5" pageNumber="92">C. amphibeus</emphasis>
</taxonomicName>
the type locality and current collection locality are likely to be the same; star indicate record based on
<bibRefCitation id="C0384B47FFE04C6BFCF7F95D72089054" author="Shaw, G. E. &amp; Shebbeare, E. O." box="[790,1052,1771,1796]" pageId="5" pageNumber="92" pagination="1 - 124" refId="ref8898" refString="Shaw, G. E. &amp; Shebbeare, E. O. (1938) Fishes of Northern Bengal. Journal Royal Asiatic Society of Bengal, 3, 1-124." type="journal article" year="1938">Shaw &amp; Shebbeare, 1938</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="A41636B6FFE04C68FF26F8E377F39785" blockId="5.[151,1436,1877,2012]" lastBlockId="6.[151,1437,151,681]" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
<emphasis id="96DDEAA4FFE04C6BFF26F8E3740D903F" bold="true" box="[199,537,1877,1903]" pageId="5" pageNumber="92">Colouration in preservative.</emphasis>
See
<figureCitation id="3C922A33FFE04C6BFDB0F8E074BF9020" box="[593,683,1878,1904]" captionStart="FIGURE 3" captionStartId="5.[152,255,1006,1030]" captionTargetBox="[340,1247,181,982]" captionTargetId="figure-19@5.[340,1247,181,982]" captionTargetPageId="5" captionText="FIGURE 3. Channa amphibeus topotypes, A. lateral, dorsal, and ventral view, KUFOS-AN-2024.12, 205.0 mm SL, colouration after preservation; B. BNHS FWF 1186, 420 mm SL, colouration immediately after exposure to formalin." figureDoi="http://doi.org/10.5281/zenodo.14798022" httpUri="https://zenodo.org/record/14798022/files/figure.png" pageId="5" pageNumber="92">Fig. 3A</figureCitation>
. In 10% formalin: Dorsum, sides of head and body dark-grey with black spots. Ventral surface of body pale-grey to creamy-white. Bars on sides beige. Spots on head pale-grey. Pectoral fin greyish, with 3 narrow, semi-circular grey bands. Dorsal fin dark-grey with narrow white distal margin. Anal fin pale grey with 23 horizontal rows of black spots. Caudal fin pale-grey with black fin rays and whitish narrow margin. Pelvic fin pale-grey with dark-grey tips. Lower jaw ventral surface and belly creamy-white, branchiostegal membrane dark grey to black.
</paragraph>
</subSubSection>
<subSubSection id="ECB3653DFFE34C68FF26FF68776695F9" pageId="6" pageNumber="93" type="distribution">
<paragraph id="A41636B6FFE34C68FF26FF68728F9549" blockId="6.[151,1437,151,681]" pageId="6" pageNumber="93">
<emphasis id="96DDEAA4FFE34C68FF26FF6874AA97A9" bold="true" box="[199,702,222,249]" pageId="6" pageNumber="93">Distribution, habitat, and ecological notes.</emphasis>
Currently,
<taxonomicName id="63A94D35FFE34C68FCDAFF69720797A9" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[827,1043,223,249]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE34C68FCDAFF69720797A9" box="[827,1043,223,249]" italics="true" pageId="6" pageNumber="93">Channa amphibeus</emphasis>
</taxonomicName>
is known only from the Chel River system (
<figureCitation id="3C922A33FFE34C68FF14FEB5772F964D" box="[245,315,259,285]" captionStart="FIGURE 4" captionStartId="5.[152,255,1699,1723]" captionTargetBox="[355,1232,1131,1671]" captionTargetId="figure-56@5.[351,1237,1127,1676]" captionTargetPageId="5" captionText="FIGURE 4. Map of Northeast India, showing distribution of Channa amphibeus, C. aurantimaculata, and C. barca. (Circles indicate type locality, squares and triangles indicate additional records; for C. amphibeus the type locality and current collection locality are likely to be the same; star indicate record based on Shaw &amp; Shebbeare, 1938)." figureDoi="http://doi.org/10.5281/zenodo.14798024" httpUri="https://zenodo.org/record/14798024/files/figure.png" pageId="6" pageNumber="93">Fig. 4</figureCitation>
). The locality details mentioned in
<bibRefCitation id="C0384B47FFE34C68FD28FEB57560964D" author="Pearson, J. T." box="[713,884,259,285]" pageId="6" pageNumber="93" pagination="551 - 552" refId="ref8375" refString="Pearson, J. T. (1839) On the Bora Chung, or the Ground Fish of Bootan. In: The journal of the Asiatic Society of Bengal. 8 (91). Bishop's College Press, Calcutta, pp. 551-552. [https://www.biodiversitylibrary.org/page/40036648]" type="book chapter" year="1839">Pearson (1839)</bibRefCitation>
and
<bibRefCitation id="C0384B47FFE34C68FC4DFEB57296964D" author="McClelland, J." box="[940,1154,258,285]" pageId="6" pageNumber="93" pagination="274 - 282" refId="ref8002" refString="McClelland, J. (1845) Description of four species of fishes from the rivers at the foot of Boutan Mountains. Journal of Natural History, Calcutta, 5 (18), 274-282." type="journal article" year="1845">McClelland (1845)</bibRefCitation>
literature reveal that this species was collected on the borders of the Chel River, not immediately from the main river, but from the dry region which possess grass jungle. The dry region with grass jungle actually indicates the Terai grasslands, and the specimens might have been collected during the summer or late winter. We presume that the locality from where we obtained our specimens, and the original
<typeStatus id="7B128814FFE34C68FD78FE2574DD96FD" box="[665,713,403,429]" pageId="6" pageNumber="93">type</typeStatus>
locality may be in close proximity as the fish described in this paper were collected from a muddy, slow-flowing, shallow stream with riparian vegetation amidst Terai grasses. From interviews with local fishers, we infer that this species grows to a length of ~
<quantity id="63519B53FFE34C68FBCBFE6D728D96A5" box="[1066,1177,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="93" unit="cm" value="70.0" valueMax="80.0" valueMin="60.0">6080 cm</quantity>
TL; adults live as pairs during the dry season in submerged holes originally created (and later abandoned) by crabs.
</paragraph>
<paragraph id="A41636B6FFE34C68FF26FD94776695F9" blockId="6.[151,1437,151,681]" pageId="6" pageNumber="93">
<emphasis id="96DDEAA4FFE34C68FF26FD947785956C" bold="true" box="[199,401,546,572]" pageId="6" pageNumber="93">Genetic analysis.</emphasis>
In the maximum likelihood phylogenetic tree based on mitochondrial co1 gene sequences,
<taxonomicName id="63A94D35FFE34C68FF76FDF177259531" baseAuthorityName="McClelland" baseAuthorityYear="1845" box="[151,305,583,609]" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE34C68FF76FDF177259531" box="[151,305,583,609]" italics="true" pageId="6" pageNumber="93">C. amphibeus</emphasis>
</taxonomicName>
was recovered as the sister species of
<taxonomicName id="63A94D35FFE34C68FD3FFDF1752B9531" authorityName="Hamilton" authorityYear="1822" box="[734,831,583,609]" family="Channidae" genus="Ophicephalus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="barca">
<emphasis id="96DDEAA4FFE34C68FD3FFDF1752B9531" box="[734,831,583,609]" italics="true" pageId="6" pageNumber="93">C. barca</emphasis>
</taxonomicName>
with a divergence of 3.13.2%. Additionally,
<taxonomicName id="63A94D35FFE34C68FAA5FDF1770495D5" baseAuthorityName="McClelland" baseAuthorityYear="1845" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="amphibeus">
<emphasis id="96DDEAA4FFE34C68FAA5FDF1770495D5" italics="true" pageId="6" pageNumber="93">Channa amphibeus</emphasis>
</taxonomicName>
differs from all species in the Gachua group (sensu
<bibRefCitation id="C0384B47FFE34C68FCB8FDDD75C095D5" author="Britz, R." box="[857,980,619,645]" pageId="6" pageNumber="93" pagination="335 - 344" refId="ref6527" refString="Britz, R. (2008) Channa ornatipinnis and C. pulchra, two new species of dwarf snakeheads from Myanmar (Teleostei: Channidae). Ichthyological Exploration of Freshwaters, 18, 335-344." type="journal article" year="2008">Britz 2008</bibRefCitation>
), by a raw genetic distance ranging from 3.119.0% (
<figureCitation id="3C922A33FFE34C68FEC1FD39777095F9" box="[288,356,654,681]" captionStart="FIGURE 5" captionStartId="6.[152,255,1456,1480]" captionTargetBox="[362,1229,709,1424]" captionTargetId="figure-255@6.[354,1234,705,1433]" captionTargetPageId="6" captionText="FIGURE 5. Maximum likelihood phylogenetic tree based on the co1 gene showing relationship of Channa amphibeus with members of the Channa gachua group." figureDoi="http://doi.org/10.5281/zenodo.14798026" httpUri="https://zenodo.org/record/14798026/files/figure.png" pageId="6" pageNumber="93">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="844233F92CC9BD72E6C45FCEBB6291D6" type="CLB-Dataset">307960</mods:identifier>
<mods:identifier id="2C2D13E6B4608849A9D0738F2F7D61E1" type="DOI">10.11646/zootaxa.5583.1.6</mods:identifier>
<mods:identifier id="83603A21D21C405B1CEA7D64012D765B" type="GBIF-Dataset">d6cbd30a-bd65-4bf8-a4af-56814f369760</mods:identifier>
<mods:identifier id="79B04C05C9562CF962A7B095657FA90F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="99837EFCD5293ED1590E564FADE8ED85" type="Zenodo-Dep">14797933</mods:identifier>
<mods:identifier id="13EEED8333231BABDF6C7CEECE246DD8" type="ZooBank">4B646824-BD55-4BCA-92DD-94634E2FABED</mods:identifier>
</mods:mods>
<treatment id="673087CAFFE5FFCFFF610609EA7CFC5E" ID-DOI="http://doi.org/10.5281/zenodo.14797922" ID-Zenodo-Dep="14797922" LSID="urn:lsid:plazi:treatment:673087CAFFE5FFCFFF610609EA7CFC5E" httpUri="http://treatment.plazi.org/id/673087CAFFE5FFCFFF610609EA7CFC5E" lastPageId="9" lastPageNumber="122" pageId="5" pageNumber="118" scope_family="Dugesiidae" scope_order="Tricladida" scope_phylum="Platyhelminthes" scope_subOrder="Continenticola">
<subSubSection id="A7836557FFE5FFC3FF610609EC88FBE4" box="[151,730,1057,1084]" pageId="5" pageNumber="118" type="nomenclature">
<paragraph id="EF2636DCFFE5FFC3FF610609EC88FBE4" blockId="5.[151,730,1057,1084]" box="[151,730,1057,1084]" pageId="5" pageNumber="118">
<heading id="B46E81B0FFE5FFC3FF610609EC88FBE4" bold="true" box="[151,730,1057,1084]" fontSize="11" level="1" pageId="5" pageNumber="118" reason="1">
<emphasis id="DDEDEACEFFE5FFC3FF610609EC88FBE4" bold="true" box="[151,730,1057,1084]" pageId="5" pageNumber="118">
<taxonomicName id="28994D5FFFE5FFC3FF610609EC3DFBE4" authority="Harrath &amp; Sluys" authorityName="Harrath &amp; Sluys" authorityYear="2025" box="[151,623,1057,1084]" family="Dugesiidae" genus="Dugesia" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="118" phylum="Platyhelminthes" rank="species" species="bursagrossa" status="sp. nov.">
<emphasis id="DDEDEACEFFE5FFC3FF610609EFC7FBE3" bold="true" box="[151,405,1057,1084]" italics="true" pageId="5" pageNumber="118">Dugesia bursagrossa</emphasis>
Harrath &amp; Sluys
</taxonomicName>
,
<taxonomicNameLabel id="C6DE57B5FFE5FFC3FD8B0609EC88FBE4" box="[637,730,1057,1083]" pageId="5" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7836557FFE5FFC3FF610641EFD4FACC" pageId="5" pageNumber="118" type="materials_examined">
<paragraph id="EF2636DCFFE5FFC3FF610641EFD4FACC" blockId="5.[151,1437,1129,2020]" pageId="5" pageNumber="118">
<emphasis id="DDEDEACEFFE5FFC3FF610641EFD6FB5C" bold="true" box="[151,388,1129,1155]" pageId="5" pageNumber="118">Material examined.</emphasis>
<materialsCitation id="5FF13C81FFE5FFC3FE790641EBCAFB78" collectingDate="2020-08-16" collectionCode="RMNH" collectorName="Waleed Aldahmash &amp; Abdel Halim Harrath" country="Saudi Arabia" latitude="18.279" location="Ain Dhiba" longLatPrecision="1" longitude="42.3885" municipality="Abha province" pageId="5" pageNumber="118" specimenCount="1" typeStatus="holotype">
<typeStatus id="3022887EFFE5FFC3FE790641EFAEFB5C" box="[399,508,1129,1155]" pageId="5" pageNumber="118" type="holotype">Holotype</typeStatus>
:
<collectionCode id="8988AE19FFE5FFC3FDFC0642EC32FB5B" box="[522,608,1130,1156]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="118" type="Museum">RMNH</collectionCode>
.VER.21535.1,
<location id="EA466007FFE5FFC3FCF90641EDD9FB5C" LSID="urn:lsid:plazi:treatment:673087CAFFE5FFCFFF610609EA7CFC5E:EA466007FFE5FFC3FCF90641EDD9FB5C" box="[783,907,1129,1155]" country="Saudi Arabia" latitude="18.279" longLatPrecision="1" longitude="42.3885" municipality="Abha province" name="Ain Dhiba" pageId="5" pageNumber="118">Ain Dhiba</location>
,
<collectingMunicipality id="0F42ACA6FFE5FFC3FC6F0641EA14FB5C" box="[921,1094,1129,1155]" pageId="5" pageNumber="118">Abha province</collectingMunicipality>
,
<collectingCountry id="978E764CFFE5FFC3FBA30641EAB8FB5C" box="[1109,1258,1129,1155]" name="Saudi Arabia" pageId="5" pageNumber="118">Saudi Arabia</collectingCountry>
(
<geoCoordinate id="8AAD501BFFE5FFC3FB0B0641EBCEFB5C" box="[1277,1436,1129,1155]" degrees="18" direction="north" minutes="16" orientation="latitude" pageId="5" pageNumber="118" precision="1" seconds="44.4" value="18.279">18°1644.4”N</geoCoordinate>
<geoCoordinate id="8AAD501BFFE5FFC3FF6106A5EF67FB78" box="[151,309,1165,1191]" degrees="42" direction="east" minutes="23" orientation="longitude" pageId="5" pageNumber="118" precision="1" seconds="18.6" value="42.3885">42°2318.6”E</geoCoordinate>
),
<date id="9B27101CFFE5FFC3FEBC06A5EC59FB78" box="[330,523,1165,1191]" pageId="5" pageNumber="118" value="2020-08-16">
<collectingDate id="8B63E9F4FFE5FFC3FEBC06A5EC59FB78" box="[330,523,1165,1191]" pageId="5" pageNumber="118" value="2020-08-16">
16
<superScript id="18EC9B94FFE5FFC3FE9006A5EF21FB44" attach="left" box="[358,371,1165,1179]" fontSize="6" pageId="5" pageNumber="118">th</superScript>
August 2020
</collectingDate>
</date>
, coll.
<collectorName id="426C530AFFE5FFC3FDBB06A5ED74FB78" box="[589,806,1165,1191]" pageId="5" pageNumber="118">Waleed Aldahmash</collectorName>
and
<collectorName id="426C530AFFE5FFC3FCAC06A5EA19FB78" box="[858,1099,1165,1191]" pageId="5" pageNumber="118">Abdel Halim Harrath</collectorName>
, sagittal sections on 42 slides
</materialsCitation>
.
<materialsCitation id="5FF13C81FFE5FFC3FF61069AEDD7FB14" box="[151,901,1201,1228]" collectingDate="2020-08-16" collectionCode="RMNH" collectorName="Waleed Aldahmash &amp; Abdel Halim Harrath" country="Saudi Arabia" latitude="18.279" location="Ain Dhiba" longLatPrecision="1" longitude="42.3885" municipality="Abha province" pageId="5" pageNumber="118" specimenCount="1" typeStatus="paratype">
<typeStatus id="3022887EFFE5FFC3FF61069AEF5AFB13" box="[151,264,1202,1228]" pageId="5" pageNumber="118" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="8988AE19FFE5FFC3FEE6069AEF34FB13" box="[272,358,1202,1228]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="118" type="Museum">RMNH</collectionCode>
.VER.21535.2, ibid., sagittal sections on 44 slides
</materialsCitation>
;
<materialsCitation id="5FF13C81FFE5FFC3FC78069AEF74FB30" collectingDate="2020-08-16" collectionCode="RMNH" collectorName="Waleed Aldahmash &amp; Abdel Halim Harrath" country="Saudi Arabia" latitude="18.279" location="Ain Dhiba" longLatPrecision="1" longitude="42.3885" municipality="Abha province" pageId="5" pageNumber="118" specimenCount="1" typeStatus="paratype">
<collectionCode id="8988AE19FFE5FFC3FC78069AEDB6FB13" box="[910,996,1202,1228]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="118" type="Museum">RMNH</collectionCode>
.VER.21535.3, ibid., horizontal sections on 42 slides
</materialsCitation>
,
<materialsCitation id="5FF13C81FFE5FFC3FEC006FEED83FB30" box="[310,977,1237,1264]" collectingDate="2020-08-16" collectionCode="RMNH" collectorName="Waleed Aldahmash &amp; Abdel Halim Harrath" country="Saudi Arabia" latitude="18.279" location="Ain Dhiba" longLatPrecision="1" longitude="42.3885" municipality="Abha province" pageId="5" pageNumber="118" specimenCount="1" typeStatus="paratype">
<collectionCode id="8988AE19FFE5FFC3FEC006FEEFDEFB2F" box="[310,396,1238,1264]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="118" type="Museum">RMNH</collectionCode>
.VER.21535.4, ibid., sagittal sections on 61 slides
</materialsCitation>
;
<materialsCitation id="5FF13C81FFE5FFC3FC1606FEEFD3FACC" collectingDate="2020-08-16" collectionCode="RMNH" collectorName="Waleed Aldahmash &amp; Abdel Halim Harrath" country="Saudi Arabia" latitude="18.279" location="Ain Dhiba" longLatPrecision="1" longitude="42.3885" municipality="Abha province" pageId="5" pageNumber="118" specimenCount="1" typeStatus="paratype">
<collectionCode id="8988AE19FFE5FFC3FC1606FEEA64FB2F" box="[992,1078,1238,1264]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="118" type="Museum">RMNH</collectionCode>
.VER.21535.5, ibid., horizontal sections on 27 slides
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A7836557FFE5FFC3FF310735EEA2FAA0" pageId="5" pageNumber="118" type="etymology">
<paragraph id="EF2636DCFFE5FFC3FF310735EEA2FAA0" blockId="5.[151,1437,1129,2020]" pageId="5" pageNumber="118">
<emphasis id="DDEDEACEFFE5FFC3FF310735EF1CFAE8" bold="true" box="[199,334,1309,1335]" pageId="5" pageNumber="118">Etymology.</emphasis>
The specific epithet is based on the Latin noun
<emphasis id="DDEDEACEFFE5FFC3FCA10735EDC5FAE8" box="[855,919,1309,1335]" italics="true" pageId="5" pageNumber="118">bursa</emphasis>
, alluding to the copulatory bursa in planarians, and the Latin adjective
<emphasis id="DDEDEACEFFE5FFC3FE5D076AEC52FA84" box="[427,512,1346,1371]" italics="true" pageId="5" pageNumber="118">grossus</emphasis>
, big, thick; thus, the name refers to the very large copulatory bursa in the new species.
</paragraph>
</subSubSection>
<subSubSection id="A7836557FFE5FFCFFF3107A1EA7CFC5E" lastPageId="9" lastPageNumber="122" pageId="5" pageNumber="118" type="diagnosis">
<paragraph id="EF2636DCFFE5FFC3FF3107A1EC5BF8D4" blockId="5.[151,1437,1129,2020]" pageId="5" pageNumber="118">
<emphasis id="DDEDEACEFFE5FFC3FF3107A1EF10FA7C" bold="true" box="[199,322,1417,1443]" pageId="5" pageNumber="118">Diagnosis.</emphasis>
<taxonomicName id="28994D5FFFE5FFC3FEB307A2EC62FA7C" authorityName="Harrath &amp; Sluys" authorityYear="2025" box="[325,560,1417,1443]" family="Dugesiidae" genus="Dugesia" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="118" phylum="Platyhelminthes" rank="species" species="bursagrossa">
<emphasis id="DDEDEACEFFE5FFC3FEB307A2EC62FA7C" box="[325,560,1417,1443]" italics="true" pageId="5" pageNumber="118">Dugesia bursagrossa</emphasis>
</taxonomicName>
is characterized by the following anatomical features:dorsal testes; symmetrical and highly glandular, barrel-shaped penis papilla; vasa deferentia opening separately into the antero-dorsal portion of a small intrabulbar seminal vesicle; ejaculatory duct opening subterminally through the ventral side of the penis papilla; muscular penis bulb composed of intermingled longitudinal and circular muscles; two ovaries positioned ventrally at approximately one-third of the distance between the brain and the root of the pharynx; oviducts opening separately and at different levels into the vaginal portion of the bursal canal; shell glands opening into the proximal, vaginal portion of the bursal canal; very large copulatory bursa, occupying almost the entire ample space between the posterior wall of the pharyngeal pocket and the penis bulb; bursal canal lined with a nucleated epithelium and covered by subepithelial layers of longitudinal muscles, followed by layers of circular muscles; mixoploid karyotype, with diploid complements of 2n = 18 and triploid complements of 3n = 27 + 1B chromosomes, with all chromosomes being metacentric.
</paragraph>
<paragraph id="EF2636DCFFE5FFC3FF31053DEAB5F83C" blockId="5.[151,1437,1129,2020]" pageId="5" pageNumber="118">
<emphasis id="DDEDEACEFFE5FFC3FF31053DEC58F8F0" bold="true" box="[199,522,1813,1839]" pageId="5" pageNumber="118">Morphological description.</emphasis>
Live animals measuring up to
<quantity id="28619B39FFE5FFC3FCA8053DEDFEF8EF" box="[862,940,1813,1840]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="118" unit="mm" value="20.0">20 mm</quantity>
in length and
<quantity id="28619B39FFE5FFC3FBBE053DEADAF8EF" box="[1096,1160,1813,1840]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="118" unit="mm" value="3.0">3 mm</quantity>
in width. Low triangular head provided with two blunt auricles and two eyes set in pigment-free patches (
<figureCitation id="77A22A59FFE5FFC3FBE40511EA3EF88C" box="[1042,1132,1849,1875]" captionStart="FIGURE 4" captionStartId="6.[151,250,1467,1491]" captionTargetBox="[154,1433,159,1443]" captionTargetId="figure-35@6.[154,1433,159,1443]" captionTargetPageId="6" captionText="FIGURE 4. Dugesia bursagrossa. A. Photograph of live specimen in dorsal view. B. Paratype RMNH.VER.21535.4. Sagittal section showing the pharynx musculature. C. Holotype RMNH.VER.21535.1. Transverse section of the pharynx, showing its musculature. D. Paratype RMNH.VER.21535.4. Detail at higher magnification of the rectangular inset in panel B. Scale bars: A: 1 mm; B: 500 µm; C: 300 µm; D: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14797941" httpUri="https://zenodo.org/record/14797941/files/figure.png" pageId="5" pageNumber="118">Fig. 4A</figureCitation>
). Dorsal surface pigmented light brown, while the ventral surface is paler. The pharynx is situated in the middle of the body and measures about 1/6 of the body length (
<figureCitation id="77A22A59FFE5FFC3FE1A05A9EC3BF844" box="[492,617,1921,1947]" captionStart="FIGURE 4" captionStartId="6.[151,250,1467,1491]" captionTargetBox="[154,1433,159,1443]" captionTargetId="figure-35@6.[154,1433,159,1443]" captionTargetPageId="6" captionText="FIGURE 4. Dugesia bursagrossa. A. Photograph of live specimen in dorsal view. B. Paratype RMNH.VER.21535.4. Sagittal section showing the pharynx musculature. C. Holotype RMNH.VER.21535.1. Transverse section of the pharynx, showing its musculature. D. Paratype RMNH.VER.21535.4. Detail at higher magnification of the rectangular inset in panel B. Scale bars: A: 1 mm; B: 500 µm; C: 300 µm; D: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14797941" httpUri="https://zenodo.org/record/14797941/files/figure.png" pageId="5" pageNumber="118">Fig. 4A, B</figureCitation>
). It is covered with an epithelium, which is underlain by a subepithelial layer of longitudinal muscle followed by layers of circular muscles. The inner pharynx epithelium is underlain by subepithelial layers of circular muscles, followed by a layer of longitudinal muscle (
<figureCitation id="77A22A59FFE5FFC3FBB305E1EA8DF83C" box="[1093,1247,1993,2019]" captionStart="FIGURE 4" captionStartId="6.[151,250,1467,1491]" captionTargetBox="[154,1433,159,1443]" captionTargetId="figure-35@6.[154,1433,159,1443]" captionTargetPageId="6" captionText="FIGURE 4. Dugesia bursagrossa. A. Photograph of live specimen in dorsal view. B. Paratype RMNH.VER.21535.4. Sagittal section showing the pharynx musculature. C. Holotype RMNH.VER.21535.1. Transverse section of the pharynx, showing its musculature. D. Paratype RMNH.VER.21535.4. Detail at higher magnification of the rectangular inset in panel B. Scale bars: A: 1 mm; B: 500 µm; C: 300 µm; D: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.14797941" httpUri="https://zenodo.org/record/14797941/files/figure.png" pageId="5" pageNumber="118">Fig. 4B, CD</figureCitation>
)
</paragraph>
<caption id="BBE66654FFE6FFC0FF610793EC3AF99F" ID-DOI="http://doi.org/10.5281/zenodo.14797941" ID-Zenodo-Dep="14797941" httpUri="https://zenodo.org/record/14797941/files/figure.png" pageId="6" pageNumber="119" startId="6.[151,250,1467,1491]" targetBox="[154,1433,159,1443]" targetPageId="6" targetType="figure">
<paragraph id="EF2636DCFFE6FFC0FF610793EC3AF99F" blockId="6.[151,1437,1467,1600]" pageId="6" pageNumber="119">
<emphasis id="DDEDEACEFFE6FFC0FF610793EF41FA0C" bold="true" box="[151,275,1467,1491]" pageId="6" pageNumber="119">FIGURE 4.</emphasis>
<taxonomicName id="28994D5FFFE6FFC0FEEC0794EFA0FA0C" authorityName="Harrath &amp; Sluys" authorityYear="2025" box="[282,498,1467,1491]" family="Dugesiidae" genus="Dugesia" kingdom="Animalia" order="Tricladida" pageId="6" pageNumber="119" phylum="Platyhelminthes" rank="species" species="bursagrossa">
<emphasis id="DDEDEACEFFE6FFC0FEEC0794EFA0FA0C" box="[282,498,1467,1491]" italics="true" pageId="6" pageNumber="119">Dugesia bursagrossa</emphasis>
</taxonomicName>
.
<emphasis id="DDEDEACEFFE6FFC0FE090793EC4AFA0C" bold="true" box="[511,536,1467,1491]" pageId="6" pageNumber="119">A.</emphasis>
Photograph of live specimen in dorsal view.
<emphasis id="DDEDEACEFFE6FFC0FC100794EDAFFA0B" bold="true" box="[998,1021,1468,1492]" pageId="6" pageNumber="119">B.</emphasis>
Paratype RMNH.VER.21535.4. Sagittal section showing the pharynx musculature.
<emphasis id="DDEDEACEFFE6FFC0FDB807F7EC34FA28" bold="true" box="[590,614,1503,1527]" pageId="6" pageNumber="119">C.</emphasis>
Holotype RMNH.VER.21535.1. Transverse section of the pharynx, showing its musculature.
<emphasis id="DDEDEACEFFE6FFC0FED6042CEF6BF9C3" bold="true" box="[288,313,1540,1564]" pageId="6" pageNumber="119">D.</emphasis>
Paratype RMNH.VER.21535.4. Detail at higher magnification of the rectangular inset in panel B. Scale bars: A: 1 mm; B: 500 µm; C: 300 µm; D: 100 µm.
</paragraph>
</caption>
<paragraph id="EF2636DCFFE6FFC0FF310446EEB7F82F" blockId="6.[151,1437,1646,2032]" pageId="6" pageNumber="119">
The numerous testes are located dorsally, extending from the posterior level of the ovaries into the posterior part of the body (
<figureCitation id="77A22A59FFE6FFC0FEDC04BAEFD4F973" box="[298,390,1681,1708]" captionStart="FIGURE 5" captionStartId="7.[151,250,1750,1774]" captionTargetBox="[202,1385,156,1731]" captionTargetId="figure-22@7.[198,1389,151,1738]" captionTargetPageId="7" captionText="FIGURE 5. Male reproductive system of D. bursagrossa A. Paratype RMNH.VER.21535.4. Sagittal section showing the small, dorsally situated testes (indicated by arrows). B. Paratype RMNH.VER.21535.4. Testicular follicles showing the characteristics of normal spermatogenesis. C. Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the vas deferens (vd) that recurves before opening into the seminal vesicle. D. Paratype RMNH.VER.21535.5. Horizontal section, showing the small diaphragm between the seminal vesicle and the ejaculatoryduct. E. Paratype RMNH.VER.21535.5.Horizontal section of the copulatory apparatus showing the spermiducal vesicle narrowing to form the portion of the sperm duct that opens into the seminal vesicle. F. Paratype RMNH.VER.21535.3. Horizontal section, showing the large copulatory bursa, containing remnants of a spermatophore, and the symmetrical openings of the sperm ducts into the seminal vesicle. Scale bars: A: 500 µm; B: 100 µm; C, E and F: 300 µm; D: 250 µm. (Arrowhead indicates the anterior side of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797943" httpUri="https://zenodo.org/record/14797943/files/figure.png" pageId="6" pageNumber="119">Fig. 5A</figureCitation>
); the follicles show various stages of spermatogenesis (
<figureCitation id="77A22A59FFE6FFC0FC0B04BAEA05F973" box="[1021,1111,1681,1708]" captionStart="FIGURE 5" captionStartId="7.[151,250,1750,1774]" captionTargetBox="[202,1385,156,1731]" captionTargetId="figure-22@7.[198,1389,151,1738]" captionTargetPageId="7" captionText="FIGURE 5. Male reproductive system of D. bursagrossa A. Paratype RMNH.VER.21535.4. Sagittal section showing the small, dorsally situated testes (indicated by arrows). B. Paratype RMNH.VER.21535.4. Testicular follicles showing the characteristics of normal spermatogenesis. C. Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the vas deferens (vd) that recurves before opening into the seminal vesicle. D. Paratype RMNH.VER.21535.5. Horizontal section, showing the small diaphragm between the seminal vesicle and the ejaculatoryduct. E. Paratype RMNH.VER.21535.5.Horizontal section of the copulatory apparatus showing the spermiducal vesicle narrowing to form the portion of the sperm duct that opens into the seminal vesicle. F. Paratype RMNH.VER.21535.3. Horizontal section, showing the large copulatory bursa, containing remnants of a spermatophore, and the symmetrical openings of the sperm ducts into the seminal vesicle. Scale bars: A: 500 µm; B: 100 µm; C, E and F: 300 µm; D: 250 µm. (Arrowhead indicates the anterior side of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797943" httpUri="https://zenodo.org/record/14797943/files/figure.png" pageId="6" pageNumber="119">Fig. 5B</figureCitation>
). On either side of the body, a large sperm duct or vas deferens runs ventrally towards the posterior end of the body. At the level of the penis papilla, the sperm ducts turn dorso-medially and then narrow before individually, but symmetrically, opening into the intrabulbar seminal vesicle (
<figureCitation id="77A22A59FFE6FFC0FE0904D6EC24F8C7" box="[511,630,1789,1816]" captionStart="FIGURE 5" captionStartId="7.[151,250,1750,1774]" captionTargetBox="[202,1385,156,1731]" captionTargetId="figure-22@7.[198,1389,151,1738]" captionTargetPageId="7" captionText="FIGURE 5. Male reproductive system of D. bursagrossa A. Paratype RMNH.VER.21535.4. Sagittal section showing the small, dorsally situated testes (indicated by arrows). B. Paratype RMNH.VER.21535.4. Testicular follicles showing the characteristics of normal spermatogenesis. C. Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the vas deferens (vd) that recurves before opening into the seminal vesicle. D. Paratype RMNH.VER.21535.5. Horizontal section, showing the small diaphragm between the seminal vesicle and the ejaculatoryduct. E. Paratype RMNH.VER.21535.5.Horizontal section of the copulatory apparatus showing the spermiducal vesicle narrowing to form the portion of the sperm duct that opens into the seminal vesicle. F. Paratype RMNH.VER.21535.3. Horizontal section, showing the large copulatory bursa, containing remnants of a spermatophore, and the symmetrical openings of the sperm ducts into the seminal vesicle. Scale bars: A: 500 µm; B: 100 µm; C, E and F: 300 µm; D: 250 µm. (Arrowhead indicates the anterior side of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797943" httpUri="https://zenodo.org/record/14797943/files/figure.png" pageId="6" pageNumber="119">Fig. 5CF</figureCitation>
). This elongated seminal vesicle is lined with a columnar and nucleated epithelium and is surrounded by a well-developed coat of intermingled muscles. A small diaphragm is situated at the distal, posterior portion of the seminal vesicle and opens into the ejaculatory duct (
<figureCitation id="77A22A59FFE6FFC0FB91056EEA93F8BF" box="[1127,1217,1861,1888]" captionStart="FIGURE 5" captionStartId="7.[151,250,1750,1774]" captionTargetBox="[202,1385,156,1731]" captionTargetId="figure-22@7.[198,1389,151,1738]" captionTargetPageId="7" captionText="FIGURE 5. Male reproductive system of D. bursagrossa A. Paratype RMNH.VER.21535.4. Sagittal section showing the small, dorsally situated testes (indicated by arrows). B. Paratype RMNH.VER.21535.4. Testicular follicles showing the characteristics of normal spermatogenesis. C. Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the vas deferens (vd) that recurves before opening into the seminal vesicle. D. Paratype RMNH.VER.21535.5. Horizontal section, showing the small diaphragm between the seminal vesicle and the ejaculatoryduct. E. Paratype RMNH.VER.21535.5.Horizontal section of the copulatory apparatus showing the spermiducal vesicle narrowing to form the portion of the sperm duct that opens into the seminal vesicle. F. Paratype RMNH.VER.21535.3. Horizontal section, showing the large copulatory bursa, containing remnants of a spermatophore, and the symmetrical openings of the sperm ducts into the seminal vesicle. Scale bars: A: 500 µm; B: 100 µm; C, E and F: 300 µm; D: 250 µm. (Arrowhead indicates the anterior side of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797943" httpUri="https://zenodo.org/record/14797943/files/figure.png" pageId="6" pageNumber="119">Fig. 5D</figureCitation>
). The latter is lined with a columnar and nucleated epithelium, but any surrounding musculature was not observed. From its point of origin near the diaphragm, the ejaculatory duct at first follows a central course through the penis papilla for a short distance, but then turns towards the ventral side of the papilla where it opens subterminally to the exterior (
<figureCitation id="77A22A59FFE6FFC0FAC8059AEBCAF813" box="[1342,1432,1969,1996]" captionStart="FIGURE 5" captionStartId="7.[151,250,1750,1774]" captionTargetBox="[202,1385,156,1731]" captionTargetId="figure-22@7.[198,1389,151,1738]" captionTargetPageId="7" captionText="FIGURE 5. Male reproductive system of D. bursagrossa A. Paratype RMNH.VER.21535.4. Sagittal section showing the small, dorsally situated testes (indicated by arrows). B. Paratype RMNH.VER.21535.4. Testicular follicles showing the characteristics of normal spermatogenesis. C. Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the vas deferens (vd) that recurves before opening into the seminal vesicle. D. Paratype RMNH.VER.21535.5. Horizontal section, showing the small diaphragm between the seminal vesicle and the ejaculatoryduct. E. Paratype RMNH.VER.21535.5.Horizontal section of the copulatory apparatus showing the spermiducal vesicle narrowing to form the portion of the sperm duct that opens into the seminal vesicle. F. Paratype RMNH.VER.21535.3. Horizontal section, showing the large copulatory bursa, containing remnants of a spermatophore, and the symmetrical openings of the sperm ducts into the seminal vesicle. Scale bars: A: 500 µm; B: 100 µm; C, E and F: 300 µm; D: 250 µm. (Arrowhead indicates the anterior side of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797943" httpUri="https://zenodo.org/record/14797943/files/figure.png" pageId="6" pageNumber="119">Figs 5E</figureCitation>
,
<figureCitation id="77A22A59FFE6FFC0FF6105FEEEEEF82F" box="[151,188,2006,2032]" captionStart="FIGURE 6" captionStartId="8.[151,250,1707,1731]" captionTargetBox="[161,1426,187,1683]" captionTargetId="figure-35@8.[157,1430,181,1688]" captionTargetPageId="8" captionText="FIGURE 6. Dugesia bursagrossa. Photomicrographs. A. Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct. B. Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore. C. Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes. D. Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries. E. Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis. F. Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa. G. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="6" pageNumber="119">6A</figureCitation>
,
<figureCitation id="77A22A59FFE6FFC0FF3105FEEE85F82F" box="[199,215,2006,2032]" captionStart="FIGURE 7" captionStartId="9.[151,250,1362,1386]" captionTargetBox="[159,1433,928,1334]" captionTargetId="figure-391@9.[154,1433,921,1339]" captionTargetPageId="9" captionText="FIGURE 7. Dugesia bursagrossa, Sagittal reconstruction of the copulatory apparatus of the holotype. Scale bar: 100 µm. Anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.14797949" httpUri="https://zenodo.org/record/14797949/files/figure.png" pageId="6" pageNumber="119">7</figureCitation>
).
</paragraph>
<caption id="BBE66654FFE7FFC1FF6104FEEAD0F830" ID-DOI="http://doi.org/10.5281/zenodo.14797943" ID-Zenodo-Dep="14797943" httpUri="https://zenodo.org/record/14797943/files/figure.png" pageId="7" pageNumber="120" startId="7.[151,250,1750,1774]" targetBox="[202,1385,156,1731]" targetPageId="7" targetType="figure">
<paragraph id="EF2636DCFFE7FFC1FF6104FEEAD0F830" blockId="7.[151,1436,1750,2031]" pageId="7" pageNumber="120">
<emphasis id="DDEDEACEFFE7FFC1FF6104FEEF40F930" bold="true" box="[151,274,1750,1775]" pageId="7" pageNumber="120">FIGURE 5.</emphasis>
Male reproductive system of
<taxonomicName id="28994D5FFFE7FFC1FDCA04FFEC84F930" authorityName="Harrath &amp; Sluys" authorityYear="2025" box="[572,726,1751,1775]" family="Dugesiidae" genus="Dugesia" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="120" phylum="Platyhelminthes" rank="species" species="bursagrossa">
<emphasis id="DDEDEACEFFE7FFC1FDCA04FFEC84F930" box="[572,726,1751,1775]" italics="true" pageId="7" pageNumber="120">D. bursagrossa</emphasis>
</taxonomicName>
<emphasis id="DDEDEACEFFE7FFC1FD2D04FEECA6F931" bold="true" box="[731,756,1750,1774]" pageId="7" pageNumber="120">A.</emphasis>
Paratype RMNH.VER.21535.4. Sagittal section showing the small, dorsally situated testes (indicated by arrows).
<emphasis id="DDEDEACEFFE7FFC1FD9304DFEC2EF8D0" bold="true" box="[613,636,1783,1807]" pageId="7" pageNumber="120">B.</emphasis>
Paratype RMNH.VER.21535.4. Testicular follicles showing the characteristics of normal spermatogenesis.
<emphasis id="DDEDEACEFFE7FFC1FE48053EEF85F8F1" bold="true" box="[446,471,1814,1838]" pageId="7" pageNumber="120">C.</emphasis>
Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the vas deferens (vd) that recurves before opening into the seminal vesicle.
<emphasis id="DDEDEACEFFE7FFC1FC84051FEDD9F890" bold="true" box="[882,907,1847,1871]" pageId="7" pageNumber="120">D.</emphasis>
Paratype RMNH.VER.21535.5. Horizontal section, showing the small diaphragm between the seminal vesicle and the ejaculatory duct.
<emphasis id="DDEDEACEFFE7FFC1FC26057FEDB5F8B0" bold="true" box="[976,999,1879,1903]" pageId="7" pageNumber="120">E.</emphasis>
Paratype RMNH.VER.21535.5. Horizontal section of the copulatory apparatus showing the spermiducal vesicle narrowing to form the portion of the sperm duct that opens into the seminal vesicle.
<emphasis id="DDEDEACEFFE7FFC1FE6205BFEFF5F870" bold="true" box="[404,423,1943,1967]" pageId="7" pageNumber="120">F.</emphasis>
Paratype RMNH.VER.21535.3. Horizontal section, showing the large copulatory bursa, containing remnants of a spermatophore, and the symmetrical openings of the sperm ducts into the seminal vesicle. Scale bars: A: 500 µm; B: 100 µm; C, E and F: 300 µm; D: 250 µm. (Arrowhead indicates the anterior side of the animal).
</paragraph>
</caption>
<caption id="BBE66654FFE8FFCEFF610483EEBEF837" ID-DOI="http://doi.org/10.5281/zenodo.14797945" ID-Zenodo-Dep="14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="8" pageNumber="121" startId="8.[151,250,1707,1731]" targetBox="[161,1426,187,1683]" targetPageId="8" targetType="figure">
<paragraph id="EF2636DCFFE8FFCEFF610483EEBEF837" blockId="8.[151,1437,1707,2024]" pageId="8" pageNumber="121">
<emphasis id="DDEDEACEFFE8FFCEFF610483EF41F91C" bold="true" box="[151,275,1707,1731]" pageId="8" pageNumber="121">FIGURE 6.</emphasis>
<taxonomicName id="28994D5FFFE8FFCEFEEC0484EFA3F91B" authorityName="Harrath &amp; Sluys" authorityYear="2025" box="[282,497,1708,1732]" family="Dugesiidae" genus="Dugesia" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="121" phylum="Platyhelminthes" rank="species" species="bursagrossa">
<emphasis id="DDEDEACEFFE8FFCEFEEC0484EFA3F91B" box="[282,497,1708,1732]" italics="true" pageId="8" pageNumber="121">Dugesia bursagrossa</emphasis>
</taxonomicName>
. Photomicrographs.
<emphasis id="DDEDEACEFFE8FFCEFD350483EC8EF91C" bold="true" box="[707,732,1707,1731]" pageId="8" pageNumber="121">A.</emphasis>
Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct.
<emphasis id="DDEDEACEFFE8FFCEFDD404C4EC68F8DB" bold="true" box="[546,570,1772,1796]" pageId="8" pageNumber="121">B.</emphasis>
Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore.
<emphasis id="DDEDEACEFFE8FFCEFBCD0523EA06F8FC" bold="true" box="[1083,1108,1803,1827]" pageId="8" pageNumber="121">C.</emphasis>
Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes.
<emphasis id="DDEDEACEFFE8FFCEFB330504EA8CF89B" bold="true" box="[1221,1246,1836,1860]" pageId="8" pageNumber="121">D.</emphasis>
Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries.
<emphasis id="DDEDEACEFFE8FFCEFCFF0564ED72F8BB" bold="true" box="[777,800,1868,1892]" pageId="8" pageNumber="121">E.</emphasis>
Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis.
<emphasis id="DDEDEACEFFE8FFCEFD560544ECE1F85B" bold="true" box="[672,691,1900,1924]" pageId="8" pageNumber="121">F.</emphasis>
Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa.
<emphasis id="DDEDEACEFFE8FFCEFE3605A3EF86F87C" bold="true" box="[448,468,1931,1955]" pageId="8" pageNumber="121">G</emphasis>
. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal).
</paragraph>
</caption>
<paragraph id="EF2636DCFFE9FFCFFF3102BFED40FE56" blockId="9.[151,1437,151,897]" pageId="9" pageNumber="122">
The highly glandular, more or less barrel-shaped penis papilla is situated within the male atrium, which communicates with a small common atrium via a constriction (
<figureCitation id="77A22A59FFE9FFCFFC830293ED83FF0A" box="[885,977,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1707,1731]" captionTargetBox="[161,1426,187,1683]" captionTargetId="figure-35@8.[157,1430,181,1688]" captionTargetPageId="8" captionText="FIGURE 6. Dugesia bursagrossa. Photomicrographs. A. Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct. B. Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore. C. Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes. D. Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries. E. Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis. F. Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa. G. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="9" pageNumber="122">Fig. 6B</figureCitation>
). The papilla is covered by a columnar, nucleated epithelium (
<figureCitation id="77A22A59FFE9FFCFFE6302F7EFA3FF26" box="[405,497,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1707,1731]" captionTargetBox="[161,1426,187,1683]" captionTargetId="figure-35@8.[157,1430,181,1688]" captionTargetPageId="8" captionText="FIGURE 6. Dugesia bursagrossa. Photomicrographs. A. Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct. B. Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore. C. Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes. D. Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries. E. Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis. F. Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa. G. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="9" pageNumber="122">Fig. 6A</figureCitation>
), under which lies a layer of circular muscles along most of its extent. Numerous ducts from abundant erythrophilic bulbar and extra-bulbar glands traverse a wide zone within the penis papilla, eventually releasing their secretions into the posterior section of the ejaculatory duct (
<figureCitation id="77A22A59FFE9FFCFFB83030FEAA1FE9E" box="[1141,1267,295,321]" captionStart="FIGURE 6" captionStartId="8.[151,250,1707,1731]" captionTargetBox="[161,1426,187,1683]" captionTargetId="figure-35@8.[157,1430,181,1688]" captionTargetPageId="8" captionText="FIGURE 6. Dugesia bursagrossa. Photomicrographs. A. Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct. B. Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore. C. Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes. D. Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries. E. Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis. F. Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa. G. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="9" pageNumber="122">Fig. 6A, B</figureCitation>
). Additionally, some other penial glands are located in the parenchymal region of the penis papilla and discharge their secretions mainly through the epithelium near the tip of the papilla.
</paragraph>
<paragraph id="EF2636DCFFE9FFCFFF3103BBEDAFFD5A" blockId="9.[151,1437,151,897]" pageId="9" pageNumber="122">
The ovaries are situated ventrally, immediately above the nerve cords at approximately one-third of the distance between the brain and the root of the pharynx and about 120 μm behind the eyes (
<figureCitation id="77A22A59FFE9FFCFFBE9039FEA24FE0E" box="[1055,1142,439,465]" captionStart="FIGURE 6" captionStartId="8.[151,250,1707,1731]" captionTargetBox="[161,1426,187,1683]" captionTargetId="figure-35@8.[157,1430,181,1688]" captionTargetPageId="8" captionText="FIGURE 6. Dugesia bursagrossa. Photomicrographs. A. Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct. B. Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore. C. Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes. D. Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries. E. Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis. F. Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa. G. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="9" pageNumber="122">Fig. 6C</figureCitation>
); the gonads show various stages of normal oogenesis (
<figureCitation id="77A22A59FFE9FFCFFE2103F3EC00FE2A" box="[471,594,475,501]" captionStart="FIGURE 6" captionStartId="8.[151,250,1707,1731]" captionTargetBox="[161,1426,187,1683]" captionTargetId="figure-35@8.[157,1430,181,1688]" captionTargetPageId="8" captionText="FIGURE 6. Dugesia bursagrossa. Photomicrographs. A. Holotype RMNH.VER.21535.1. Sagittal section of the penis papilla, showing the subterminal opening of the ejaculatory duct. Note the abundance of penial glands, discharging their secretion (arrowheads) into the ejaculatory duct. B. Holotype RMNH.VER.21535.1. Sagittal section of the copulatory apparatus showing the penis papilla, the constriction in the male atrium, the common atrium, and the gonopore. C. Paratype RMNH.VER.21535.4. Sagittal section showing the form and position of ovary in relation to the anterior end and the eyes. D. Paratype RMNH. VER.21535.5. Horizontal section showing the two ovaries. E. Paratype RMNH.VER.21535.5. Horizontal section of ovary, showing the characteristics of normal oogenesis. F. Holotype RMNH.VER.21535.1. Sagittal section of the very large and elongated copulatory bursa. G. Paratype RMNH.VER.21535.4. Sagittal section of the very large and elongated copulatory bursa. Scale bars: A, B and D: 200 µm; C: 400 µm; E: 150 µm; F and G: 300 µm (Arrowhead indicates the anterior end of the animal)." figureDoi="http://doi.org/10.5281/zenodo.14797945" httpUri="https://zenodo.org/record/14797945/files/figure.png" pageId="9" pageNumber="122">Fig. 6D, E</figureCitation>
). The oviducts run dorsally to the ventral nerve cords until arriving shortly posterior to the gonopore, where they turn dorso-medially to open separately into the vaginal section of the bursal canal. The oviducal openings are asymmetrical in that the left oviduct opens into the ventral portion of the bursal canal, near its communication with the atrium, whereas the right oviduct opens much higher into the bursal canal. The shell glands discharge their secretion into this portion of the bursal canal.
</paragraph>
<paragraph id="EF2636DCFFE9FFCFFF3100A7EA7CFC5E" blockId="9.[151,1437,151,897]" pageId="9" pageNumber="122">
From its vaginal region, the bursal canal curves forwards, meanwhile diminishing in diameter. The canal is lined with a nucleated epithelium composed of cells of variable height, while it is coated with a subepithelial layer of longitudinal muscles, followed by a thick layer of circular muscles. At a level approximately at the anterior portion of the penis bulb, the bursal canal opens through the postero-dorsal wall of the copulatory bursa. This copulatory bursa is a very large and particularly elongated, sac-shaped structure that is located immediately anterior to the penis bulb and occupies almost the entire ample space between the posterior wall of the pharyngeal pocket and the penis bulb (
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,
<figureCitation id="77A22A59FFE9FFCFFEAA014FEF3EFC5E" box="[348,364,871,897]" captionStart="FIGURE 7" captionStartId="9.[151,250,1362,1386]" captionTargetBox="[159,1433,928,1334]" captionTargetId="figure-391@9.[154,1433,921,1339]" captionTargetPageId="9" captionText="FIGURE 7. Dugesia bursagrossa, Sagittal reconstruction of the copulatory apparatus of the holotype. Scale bar: 100 µm. Anterior to the left." figureDoi="http://doi.org/10.5281/zenodo.14797949" httpUri="https://zenodo.org/record/14797949/files/figure.png" pageId="9" pageNumber="122">7</figureCitation>
); a muscle layer around the copulatory bursa was not detected.
</paragraph>
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</treatment>
</document>

View file

@ -1,71 +1,72 @@
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@ -160,7 +161,7 @@ Externally the new species is similar to
<emphasis id="3D95EAD7FF94FFA29D2DF8B4EF7A5A34" box="[777,1075,1811,1837]" italics="true" pageId="3" pageNumber="189">Eucosma gonzalezalvarezi</emphasis>
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but differs in the lighter ground colour of the forewing that attenuates the contrast with the costal strigulae and the ill-defined markings (
<figureCitation id="97DA2A40FF94FFA29B37F897EEC75A48" box="[1299,1422,1847,1873]" captionStart="FIGURE 2" captionStartId="2.[152,255,1435,1459]" captionTargetBox="[340,1247,345,1412]" captionTargetId="figure-79@2.[340,1247,345,1412]" captionTargetPageId="2" captionText="FIGURE 2. Adult specimens. A. Holotype Eucosma callei sp. nov. ♂; B. Eucosma gonzalezalvarezi Agenjo, 1970 ♂. Photograpy by Rubio and Ortiz." pageId="3" pageNumber="189">Figs 2A, B</figureCitation>
<figureCitation id="97DA2A40FF94FFA29B37F897EEC75A48" box="[1299,1422,1847,1873]" captionStart="FIGURE 2" captionStartId="2.[152,255,1435,1459]" captionTargetBox="[340,1247,345,1412]" captionTargetId="figure-79@2.[340,1247,345,1412]" captionTargetPageId="2" captionText="FIGURE 2. Adult specimens. A. Holotype Eucosma callei sp. nov. ♂; B. Eucosma gonzalezalvarezi Agenjo, 1970 ♂. Photograpy by Rubio and Ortiz." figureDoi="http://doi.org/10.5281/zenodo.14797520" httpUri="https://zenodo.org/record/14797520/files/figure.png" pageId="3" pageNumber="189">Figs 2A, B</figureCitation>
). The male genitalia of
<taxonomicName id="C8E14D46FF94FFA29FB4F8FCEAA45A6C" authorityName="Girdley, Garre, Rubio, &amp; Ortiz" authorityYear="2025" box="[400,493,1883,1909]" class="Insecta" family="Tortricidae" genus="Eucosma" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="189" phylum="Arthropoda" rank="species" species="callei">
<emphasis id="3D95EAD7FF94FFA29FB4F8FCEAA45A6C" box="[400,493,1883,1909]" italics="true" pageId="3" pageNumber="189">E. callei</emphasis>
@ -170,7 +171,7 @@ differ from those of
<emphasis id="3D95EAD7FF94FFA29CFEF8FCE8FC5A6C" box="[730,949,1883,1909]" italics="true" pageId="3" pageNumber="189">E. gonzalezalvarezi</emphasis>
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in the position of ventral edge of sacculus, the angle of sacculus, the length of neck of valve, the depth of the ventral incision of valva and the size of uncus (
<figureCitation id="97DA2A40FF94FFA29EBBF803EA575AA4" box="[159,286,1955,1981]" captionStart="FIGURE 3" captionStartId="3.[152,255,1345,1369]" captionTargetBox="[340,1247,181,1321]" captionTargetId="figure-21@3.[340,1247,181,1321]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia. A. Eucosma callei sp. nov., Genitalia slide JG-412; B. Eucosma gonzalezalvarezi Agenjo, 1970, Genitalia slide MG-1407. Photograp by Rubio and Ortiz." pageId="3" pageNumber="189">Figs 3A, B</figureCitation>
<figureCitation id="97DA2A40FF94FFA29EBBF803EA575AA4" box="[159,286,1955,1981]" captionStart="FIGURE 3" captionStartId="3.[152,255,1345,1369]" captionTargetBox="[340,1247,181,1321]" captionTargetId="figure-21@3.[340,1247,181,1321]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia. A. Eucosma callei sp. nov., Genitalia slide JG-412; B. Eucosma gonzalezalvarezi Agenjo, 1970, Genitalia slide MG-1407. Photograp by Rubio and Ortiz." figureDoi="http://doi.org/10.5281/zenodo.14797522" httpUri="https://zenodo.org/record/14797522/files/figure.png" pageId="3" pageNumber="189">Figs 3A, B</figureCitation>
). Finally, these two and related species are separated by DNA barcode sequences, except
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<emphasis id="3D95EAD7FF94FFA29B2AF804EED55AA4" box="[1294,1436,1955,1981]" italics="true" pageId="3" pageNumber="189">E. aemulana</emphasis>
@ -180,7 +181,7 @@ and
<emphasis id="3D95EAD7FF94FFA29EE3F868EA1F5AF8" box="[199,342,1991,2017]" italics="true" pageId="3" pageNumber="189">E. tripoliana</emphasis>
</taxonomicName>
which have identical barcodes (
<figureCitation id="97DA2A40FF94FFA29C9AF867E84D5AF8" box="[702,772,1991,2017]" captionStart="FIGURE 4" captionStartId="4.[152,255,1625,1649]" captionTargetBox="[356,1232,801,1597]" captionTargetId="figure-189@4.[343,1245,796,1602]" captionTargetPageId="4" captionText="FIGURE 4. Maximum Likelihood tree of Eucosma species, obtained from 70 nucleotide COI sequences. The depth of each branch shows divergence within lineages. Bootstrap values are provided at major nodes. The scale bar represents 0.01 genetic difference." pageId="3" pageNumber="189">Fig. 4</figureCitation>
<figureCitation id="97DA2A40FF94FFA29C9AF867E84D5AF8" box="[702,772,1991,2017]" captionStart="FIGURE 4" captionStartId="4.[152,255,1625,1649]" captionTargetBox="[356,1232,801,1597]" captionTargetId="figure-189@4.[343,1245,796,1602]" captionTargetPageId="4" captionText="FIGURE 4. Maximum Likelihood tree of Eucosma species, obtained from 70 nucleotide COI sequences. The depth of each branch shows divergence within lineages. Bootstrap values are provided at major nodes. The scale bar represents 0.01 genetic difference." figureDoi="http://doi.org/10.5281/zenodo.14797526" httpUri="https://zenodo.org/record/14797526/files/figure.png" pageId="3" pageNumber="189">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
@ -334,7 +335,7 @@ Taxon names, BOLD accession numbers for the specimens used in distance estimatio
</tr>
</table>
</paragraph>
<caption id="5B9E664DFF93FFA59EBCF9F9EA4C5BA0" pageId="4" pageNumber="190" startId="4.[152,255,1625,1649]" targetBox="[356,1232,801,1597]" targetPageId="4" targetType="figure">
<caption id="5B9E664DFF93FFA59EBCF9F9EA4C5BA0" ID-DOI="http://doi.org/10.5281/zenodo.14797526" ID-Zenodo-Dep="14797526" httpUri="https://zenodo.org/record/14797526/files/figure.png" pageId="4" pageNumber="190" startId="4.[152,255,1625,1649]" targetBox="[356,1232,801,1597]" targetPageId="4" targetType="figure">
<paragraph id="0F5E36C5FF93FFA59EBCF9F9EA4C5BA0" blockId="4.[151,1436,1625,1721]" pageId="4" pageNumber="190">
<emphasis id="3D95EAD7FF93FFA59EBCF9F9EA535B68" bold="true" box="[152,282,1625,1649]" pageId="4" pageNumber="190">FIGURE 4.</emphasis>
Maximum Likelihood tree of
@ -349,7 +350,7 @@ species
<paragraph id="0F5E36C5FF93FFA49EE3F8ABE8305C58" blockId="4.[151,1436,1803,2010]" lastBlockId="5.[151,1437,151,969]" lastPageId="5" lastPageNumber="191" pageId="4" pageNumber="190">
<emphasis id="3D95EAD7FF93FFA59EE3F8ABEA105A3C" bold="true" box="[199,345,1803,1829]" pageId="4" pageNumber="190">Description.</emphasis>
Male. Adult (
<figureCitation id="97DA2A40FF93FFA59C27F8ACE9155A3F" box="[515,604,1804,1830]" captionStart="FIGURE 2" captionStartId="2.[152,255,1435,1459]" captionTargetBox="[340,1247,345,1412]" captionTargetId="figure-79@2.[340,1247,345,1412]" captionTargetPageId="2" captionText="FIGURE 2. Adult specimens. A. Holotype Eucosma callei sp. nov. ♂; B. Eucosma gonzalezalvarezi Agenjo, 1970 ♂. Photograpy by Rubio and Ortiz." pageId="4" pageNumber="190">Fig 2A</figureCitation>
<figureCitation id="97DA2A40FF93FFA59C27F8ACE9155A3F" box="[515,604,1804,1830]" captionStart="FIGURE 2" captionStartId="2.[152,255,1435,1459]" captionTargetBox="[340,1247,345,1412]" captionTargetId="figure-79@2.[340,1247,345,1412]" captionTargetPageId="2" captionText="FIGURE 2. Adult specimens. A. Holotype Eucosma callei sp. nov. ♂; B. Eucosma gonzalezalvarezi Agenjo, 1970 ♂. Photograpy by Rubio and Ortiz." figureDoi="http://doi.org/10.5281/zenodo.14797520" httpUri="https://zenodo.org/record/14797520/files/figure.png" pageId="4" pageNumber="190">Fig 2A</figureCitation>
). Head: frons whitish; vertex whitish; antenna filiform with scape and pedicellus whitish; labial palpus whitish somewhat ochreous on sides, porrect. Thorax: patagium pale brown; mesonotum ochreous; tegula ochreous; metanotum grey; thorax ventrally whitish; forewing length
<quantity id="C8199B20FF93FFA59AD2F8F3EE065A77" box="[1270,1359,1875,1902]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="4" pageNumber="190" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
(mean
@ -360,7 +361,7 @@ length of forewing; upperside ground colour creamy with brown admixture; nine pa
</paragraph>
<paragraph id="0F5E36C5FF92FFA49EE3FEEBEAAC5F00" blockId="5.[151,1437,151,969]" pageId="5" pageNumber="191">
Male genitalia based on 5 preparations (
<figureCitation id="97DA2A40FF92FFA49C5DFEEBE9985C7C" box="[633,721,331,357]" captionStart="FIGURE 3" captionStartId="3.[152,255,1345,1369]" captionTargetBox="[340,1247,181,1321]" captionTargetId="figure-21@3.[340,1247,181,1321]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia. A. Eucosma callei sp. nov., Genitalia slide JG-412; B. Eucosma gonzalezalvarezi Agenjo, 1970, Genitalia slide MG-1407. Photograp by Rubio and Ortiz." pageId="5" pageNumber="191">Fig. 3A</figureCitation>
<figureCitation id="97DA2A40FF92FFA49C5DFEEBE9985C7C" box="[633,721,331,357]" captionStart="FIGURE 3" captionStartId="3.[152,255,1345,1369]" captionTargetBox="[340,1247,181,1321]" captionTargetId="figure-21@3.[340,1247,181,1321]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia. A. Eucosma callei sp. nov., Genitalia slide JG-412; B. Eucosma gonzalezalvarezi Agenjo, 1970, Genitalia slide MG-1407. Photograp by Rubio and Ortiz." figureDoi="http://doi.org/10.5281/zenodo.14797522" httpUri="https://zenodo.org/record/14797522/files/figure.png" pageId="5" pageNumber="191">Fig. 3A</figureCitation>
). Uncus reduced, rounded apically, slightly hairy. Socii drooping, thick and convex on the external side. Tegumen simple, large, straight-edged and tapering distally. Gnathos and anal cone slightly sclerotized. Basal part of valva relatively narrow. Sacculus slightly convex ventrally with weakly rounded caudal angle; neck of valva slender; ventral incision of valva distinct, but rather shallow. Cucullus large, with rounded dorsal angle and distinct ventral lobe, densely setous. Phallus short, tapering terminally, which contains a clump of deciduous cornuti.
</paragraph>
<paragraph id="0F5E36C5FF92FFA49EE3FD83EAEB5F24" blockId="5.[151,1437,151,969]" box="[199,418,547,573]" pageId="5" pageNumber="191">
@ -439,11 +440,11 @@ Asso
after
<bibRefCitation id="6B704B34FF92FFA49C3AFC0FE98B5ED0" author="Agenjo, R." box="[542,706,943,969]" pageId="5" pageNumber="191" pagination="1 - 4" refId="ref4087" refString="Agenjo, R. (1970) Una nueva Eucosma Hb., 1826, madrilena, dedicada al Excmo. Sr. Prof. Dr. D. Angel Gonzalez Alvarez, Secretario General del Consejo Superior de Investigaciones Cientificas (Lep. Tortricidae). Eos Revista espanola de entomologia, 45 (1-4), 7 - 12, pl. I." type="journal article" year="1970">Agenjo (1970)</bibRefCitation>
(
<figureCitation id="97DA2A40FF92FFA49CF6FC0FE85F5ED0" box="[722,790,942,969]" captionStart="FIGURE 5" captionStartId="5.[152,255,1696,1720]" captionTargetBox="[340,1247,993,1673]" captionTargetId="figure-362@5.[340,1247,993,1673]" captionTargetPageId="5" captionText="FIGURE 5. Habitat in the Fuente de los Almeces, Moratalla Mountains, Murcia. Photograph by J. Girdley." pageId="5" pageNumber="191">Fig. 5</figureCitation>
<figureCitation id="97DA2A40FF92FFA49CF6FC0FE85F5ED0" box="[722,790,942,969]" captionStart="FIGURE 5" captionStartId="5.[152,255,1696,1720]" captionTargetBox="[340,1247,993,1673]" captionTargetId="figure-362@5.[340,1247,993,1673]" captionTargetPageId="5" captionText="FIGURE 5. Habitat in the Fuente de los Almeces, Moratalla Mountains, Murcia. Photograph by J. Girdley." figureDoi="http://doi.org/10.5281/zenodo.14797528" httpUri="https://zenodo.org/record/14797528/files/figure.png" pageId="5" pageNumber="191">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="5B9E664DFF92FFA49EBCF900EF955BA0" box="[152,1244,1696,1721]" pageId="5" pageNumber="191" startId="5.[152,255,1696,1720]" targetBox="[340,1247,993,1673]" targetPageId="5" targetType="figure">
<caption id="5B9E664DFF92FFA49EBCF900EF955BA0" ID-DOI="http://doi.org/10.5281/zenodo.14797528" ID-Zenodo-Dep="14797528" box="[152,1244,1696,1721]" httpUri="https://zenodo.org/record/14797528/files/figure.png" pageId="5" pageNumber="191" startId="5.[152,255,1696,1720]" targetBox="[340,1247,993,1673]" targetPageId="5" targetType="figure">
<paragraph id="0F5E36C5FF92FFA49EBCF900EF955BA0" blockId="5.[152,1244,1696,1721]" box="[152,1244,1696,1721]" pageId="5" pageNumber="191">
<emphasis id="3D95EAD7FF92FFA49EBCF900EA515BA0" bold="true" box="[152,280,1696,1721]" pageId="5" pageNumber="191">FIGURE 5.</emphasis>
Habitat in the Fuente de los Almeces, Moratalla Mountains, Murcia. Photograph by J. Girdley.

View file

@ -0,0 +1,434 @@
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<paragraph id="13679254292B6A78FF7AE279FDB2FBA5" blockId="9.[151,941,1086,1185]" box="[151,519,1086,1113]" pageId="9" pageNumber="80">
<heading id="482F2538292B6A78FF7AE279FDB2FBA5" bold="true" box="[151,519,1086,1113]" fontSize="11" level="1" pageId="9" pageNumber="80" reason="1">
<emphasis id="21AC4E46292B6A78FF7AE279FDB2FBA5" bold="true" box="[151,519,1086,1113]" pageId="9" pageNumber="80">
<taxonomicName id="D4D8E9D7292B6A78FF7AE279FE14FBA5" authorityName="Sassi" authorityYear="2025" box="[151,417,1086,1113]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="tommasoi" status="sp. nov.">
<emphasis id="21AC4E46292B6A78FF7AE279FE14FBA5" bold="true" box="[151,417,1086,1113]" italics="true" pageId="9" pageNumber="80">Metallactus tommasoi</emphasis>
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<taxonomicNameLabel id="3A9FF33D292B6A78FE44E278FDB2FBA5" box="[425,519,1087,1113]" pageId="9" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<subSubSection id="5BC2C1DF292B6A78FF7AE2C0FCCCF94A" pageId="9" pageNumber="80" type="description">
<paragraph id="13679254292B6A78FF7AE2C0FE32FB5D" blockId="9.[151,941,1086,1185]" box="[151,391,1158,1185]" pageId="9" pageNumber="80">
(
<figureCitation id="8BE38ED1292B6A78FF72E2C0FF5DFB5D" box="[159,232,1159,1185]" captionStart="FIGURE 3" captionStartId="10.[151,250,1600,1624]" captionTargetBox="[167,1420,181,1566]" captionTargetId="figure-33@10.[151,1436,181,1577]" captionTargetPageId="10" captionText="FIGURE 3. Metallactus tommasoi. ab: habitus, dorsal and lateral view (HT); c: frontal view, male (HT); d: frontal view, female (PT, Felipe Solá, Buenos Aires, Argentina); eh: colour variations (e: female, PT, Felipe Solá, Buenos Aires, Argentina; f: male, PT, Felipe Solá, Buenos Aires, Argentina; g: male, PT, Tandil, Buenos Aires, Argentina; h: male, PT, Felipe Solá, Buenos Aires, Argentina). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.14798008" httpUri="https://zenodo.org/record/14798008/files/figure.png" pageId="9" pageNumber="80">Figs 3</figureCitation>
;
<figureCitation id="8BE38ED1292B6A78FF18E2C0FE9BFB5D" box="[245,302,1159,1185]" captionStart="FIGURE 4" captionStartId="11.[151,250,1594,1618]" captionTargetBox="[171,1416,309,1554]" captionTargetId="figure-80@11.[171,1416,309,1570]" captionTargetPageId="11" captionText="FIGURE 4. ad: Metallactus minax, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg: M. tommasoi, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="9" pageNumber="80">4eg</figureCitation>
;
<figureCitation id="8BE38ED1292B6A78FED7E2C1FEE2FB5D" box="[314,343,1158,1185]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="9" pageNumber="80">5c</figureCitation>
;
<figureCitation id="8BE38ED1292B6A78FE8EE2C0FECBFB5D" box="[355,382,1159,1185]" captionStart="FIGURE 6" captionStartId="13.[151,250,948,972]" captionTargetBox="[151,1436,273,910]" captionTargetId="figure-48@13.[151,1436,273,924]" captionTargetPageId="13" captionText="FIGURE 6. Distribution maps for the Metallactus minax species-group. a: M. minax; b: M. abstersus; c: M. tommasoi. Circles: data from specimens analysed during the study. Star in fig. b: the available information allows to point out the presence in the state but without more precise data on the location. Star in fig. c: Identification uncertain based on females only, therefore excluded from the type series. In fig. a: record of the species for the state of Pará, indicated on the label of one of the paratypes (see text), is not shown on the map because this data is considered to require further confirmation." figureDoi="http://doi.org/10.5281/zenodo.14798014" httpUri="https://zenodo.org/record/14798014/files/figure.png" pageId="9" pageNumber="80">6c</figureCitation>
)
</paragraph>
<paragraph id="13679254292B6A78FF7AE28FFCCCF94A" blockId="9.[151,1437,1224,2043]" pageId="9" pageNumber="80">
<emphasis id="21AC4E46292B6A78FF7AE28FFF55FB1E" bold="true" box="[151,224,1224,1250]" italics="true" pageId="9" pageNumber="80">Types.</emphasis>
<materialsCitation id="A3B09809292B6A78FF04E28FFF5BFAB2" collectorName="Monros En. &amp; F. Monros Collection" country="Argentina" location="Bahia Blanca" pageId="9" pageNumber="80" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
<typeStatus id="CC632CF6292B6A78FF04E28FFEEAFB1E" box="[233,351,1224,1250]" pageId="9" pageNumber="80" type="holotype">HOLOTYPE</typeStatus>
:
<emphasis id="21AC4E46292B6A78FE9DE28FFD60FB1E" bold="true" box="[368,725,1224,1250]" pageId="9" pageNumber="80">
<collectingCountry id="6BCFD2C4292B6A78FE9DE28FFD95FB1E" box="[368,544,1224,1250]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
:
<collectingRegion id="D11C5CB6292B6A78FDC3E28FFD65FB1E" box="[558,720,1224,1250]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="9" pageNumber="80">Buenos Aires</collectingRegion>
:
</emphasis>
<specimenCount id="05DE59DD292B6A78FD32E28EFD43FB1F" box="[735,758,1225,1251]" count="1" pageId="9" pageNumber="80" type="male"></specimenCount>
, gLued, detached abdomen gLued on the same card, median lobe of aedeagus glued on a separate card, // “
<collectingCountry id="6BCFD2C4292B6A78FD70E2ABFCF7FAFA" box="[669,834,1260,1286]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
Bs. AIRES, Felipe Sola (
<location id="1607C48F292B6A78FB8EE2ABFB48FAFA" LSID="urn:lsid:plazi:treatment:9B712342292B6A7FFF7AE279FC20FD0D:1607C48F292B6A78FB8EE2ABFB48FAFA" box="[1123,1277,1260,1286]" country="Argentina" name="Bahia Blanca" pageId="9" pageNumber="80" stateProvince="Ciudad Autonoma de Buenos Aires">Bahia Blanca</location>
) coll.
<collectorName id="BE2DF782292B6A78FAA8E2ABFF0BFAD7" pageId="9" pageNumber="80">Monrós En.</collectorName>
942” [white label, handwritten] // “
<collectorName id="BE2DF782292B6A78FDA7E357FC8CFAD6" box="[586,825,1296,1323]" pageId="9" pageNumber="80">F. Monros Collection</collectorName>
1959” [White label, printed] // “
<taxonomicName id="D4D8E9D7292B6A78FB48E357FA29FAD6" authorityName="Sassi" authorityYear="2025" box="[1189,1436,1296,1322]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="tommasoi" status="sp. nov.">
<emphasis id="21AC4E46292B6A78FB48E357FA29FAD6" box="[1189,1436,1296,1322]" italics="true" pageId="9" pageNumber="80">Metallactus tommasoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3A9FF33D292B6A78FF7AE373FF5BFAB2" box="[151,238,1332,1358]" pageId="9" pageNumber="80" rank="species">
<emphasis id="21AC4E46292B6A78FF7AE373FF5BFAB2" bold="true" box="[151,238,1332,1358]" pageId="9" pageNumber="80">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="21AC4E46292B6A78FF02E373FF41FAB2" bold="true" box="[239,244,1332,1358]" pageId="9" pageNumber="80">.</emphasis>
<materialsCitation id="A3B09809292B6A78FF10E373FED5F94A" collectingDate="1938-12" collectionCode="USNMNH, DSPC, ZSM" collectorName="D. Sassi &amp; Felipe Sola &amp; Monros Feb. &amp; F. Monros Collection &amp; F. Monros &amp; Tandil &amp; Monros En. &amp; Manuel J. Viana" country="Argentina" county="Bahia" location="Dep. de Calamuchita El Sauce" municipality="FeLipe SoLa" pageId="9" pageNumber="80" specimenCount="24" specimenCount-female="12" specimenCount-male="12" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
<typeStatus id="CC632CF6292B6A78FF10E373FEC8FAB2" box="[253,381,1332,1358]" pageId="9" pageNumber="80" type="holotype">HOLOTYPUS</typeStatus>
<collectorName id="BE2DF782292B6A78FE6AE372FE53FAB2" box="[391,486,1332,1359]" pageId="9" pageNumber="80">D. Sassi</collectorName>
des.” [red label, printed] // (
<collectionCode id="75C90A91292B6A78FCDEE373FC06FAB2" box="[819,947,1332,1358]" pageId="9" pageNumber="80">USNMNH</collectionCode>
).
<typeStatus id="CC632CF6292B6A78FC25E373FB88FAB2" box="[968,1085,1332,1358]" pageId="9" pageNumber="80" type="paratype">PARATYPES</typeStatus>
(24):
<emphasis id="21AC4E46292B6A78FB6AE373FF55FA8E" bold="true" pageId="9" pageNumber="80">
<collectingCountry id="6BCFD2C4292B6A78FB6AE373FA82FAB2" box="[1159,1335,1332,1358]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
:
<collectingRegion id="D11C5CB6292B6A78FAA8E373FF6FFA8E" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="9" pageNumber="80">Buenos Aires</collectingRegion>
:
</emphasis>
<specimenCount id="05DE59DD292B6A78FF04E31FFEA1FA8E" box="[233,276,1368,1394]" count="2" pageId="9" pageNumber="80" type="male">2♂</specimenCount>
<specimenCount id="05DE59DD292B6A78FEF9E31FFE85FA8E" box="[276,304,1368,1394]" count="1" pageId="9" pageNumber="80" type="female">1♀</specimenCount>
, same data as the hoLotype (
<collectionCode id="75C90A91292B6A78FD9AE31FFD47FA8E" box="[631,754,1368,1394]" pageId="9" pageNumber="80">USNMNH</collectionCode>
&amp;
<collectionCode id="75C90A91292B6A78FCF6E31FFCD3FA8E" box="[795,870,1368,1394]" pageId="9" pageNumber="80">DSPC</collectionCode>
);
<specimenCount id="05DE59DD292B6A78FC96E31FFC2BFA8E" box="[891,926,1368,1394]" count="4" pageId="9" pageNumber="80" type="female">4♀</specimenCount>
// “
<collectingCountry id="6BCFD2C4292B6A78FC24E31FFBDBFA8E" box="[969,1134,1368,1394]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
Bs. AIRES,
<collectingCounty id="FA06EAD8292B6A78FAE9E31FFAF0FA8E" box="[1284,1349,1368,1394]" pageId="9" pageNumber="80">Bahia</collectingCounty>
BLanca
<collectorName id="BE2DF782292B6A78FF7AE33BFEA3FA6A" box="[151,278,1404,1430]" pageId="9" pageNumber="80">Felipe Sola</collectorName>
coll.
<collectorName id="BE2DF782292B6A78FEBEE33BFE57FA6A" box="[339,482,1404,1430]" pageId="9" pageNumber="80">Monrós Feb.</collectorName>
942” [white label, handwritten] // “
<collectorName id="BE2DF782292B6A78FC80E33BFBEEFA6A" box="[877,1115,1404,1431]" pageId="9" pageNumber="80">F. Monros Collection</collectorName>
1959” [White label, printed] // (
<collectionCode id="75C90A91292B6A78FF58E3E7FE80FA46" box="[181,309,1440,1466]" pageId="9" pageNumber="80">USNMNH</collectionCode>
);
<specimenCount id="05DE59DD292B6A78FEA5E3E7FEDAFA46" box="[328,367,1440,1466]" count="7" pageId="9" pageNumber="80" type="male">7♂</specimenCount>
<specimenCount id="05DE59DD292B6A78FE82E3E7FE3AFA46" box="[367,399,1440,1466]" count="2" pageId="9" pageNumber="80" type="female">2♀</specimenCount>
// “
<collectingCountry id="6BCFD2C4292B6A78FE58E3E7FDEFFA46" box="[437,602,1440,1466]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
Bs. AIRES,
<collectingMunicipality id="F303082E292B6A78FD07E3E7FCDCFA46" box="[746,873,1440,1466]" pageId="9" pageNumber="80">FeLipe SoLá</collectingMunicipality>
coLL. Monrós Feb. 942” [white LabeL, handwritten] // “
<collectorName id="BE2DF782292B6A78FF57E383FE9AFA23" box="[186,303,1476,1503]" pageId="9" pageNumber="80">F. Monros</collectorName>
CoLLection 1959” [White LabeL, printed] // (
<collectionCode id="75C90A91292B6A78FCF3E383FC2CFA22" box="[798,921,1476,1502]" pageId="9" pageNumber="80">USNMNH</collectionCode>
&amp;
<collectionCode id="75C90A91292B6A78FC2DE383FBBEFA22" box="[960,1035,1476,1502]" pageId="9" pageNumber="80">DSPC</collectionCode>
);
<specimenCount id="05DE59DD292B6A78FBF2E383FBF6FA22" box="[1055,1091,1476,1502]" count="3" pageId="9" pageNumber="80" type="male">3♂</specimenCount>
// “
<collectingCountry id="6BCFD2C4292B6A78FB81E383FAA4FA22" box="[1132,1297,1476,1502]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
Bs. AIRES,
<collectorName id="BE2DF782292B6A78FF7AE3AFFF6BF9FE" box="[151,222,1512,1538]" pageId="9" pageNumber="80">Tandil</collectorName>
coll.
<collectorName id="BE2DF782292B6A78FECEE3AFFE1FF9FF" box="[291,426,1512,1539]" pageId="9" pageNumber="80">Monrós En.</collectorName>
1941” [white label, handwritten] // “
<collectorName id="BE2DF782292B6A78FCB5E3AFFBF8F9FE" box="[856,1101,1512,1539]" pageId="9" pageNumber="80">F. Monros Collection</collectorName>
1959” [White label, printed] // (
<collectionCode id="75C90A91292B6A78FF55E04BFE86F9DA" box="[184,307,1548,1574]" pageId="9" pageNumber="80">USNMNH</collectionCode>
&amp;
<collectionCode id="75C90A91292B6A78FEB1E04BFE12F9DA" box="[348,423,1548,1574]" pageId="9" pageNumber="80">DSPC</collectionCode>
);
<specimenCount id="05DE59DD292B6A78FE50E04BFE55F9DA" box="[445,480,1548,1574]" count="1" pageId="9" pageNumber="80" type="female">1♀</specimenCount>
// “ARGENTINIEN Prov.
<collectingRegion id="D11C5CB6292B6A78FCF6E04AFC01F9DA" box="[795,948,1548,1575]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="9" pageNumber="80">Buenos Aires</collectingRegion>
Sierra de La Ventana 20.3.54 [white LabeL, partly handwritten] // “Zool. Staatslg. München” [White label, printed] // (
<collectionCode id="75C90A91292B6A78FC1FE077FB84F9B6" box="[1010,1073,1584,1610]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="80" type="Museum">ZSM</collectionCode>
).
<emphasis id="21AC4E46292B6A78FBAAE068FB04F9B6" bold="true" box="[1095,1201,1583,1610]" pageId="9" pageNumber="80">
<collectingRegion id="D11C5CB6292B6A78FBAAE068FB04F9B6" box="[1095,1201,1583,1610]" country="Argentina" name="Cordoba" pageId="9" pageNumber="80">Córdoba</collectingRegion>
</emphasis>
:
<specimenCount id="05DE59DD292B6A78FB2FE077FB50F9B6" box="[1218,1253,1584,1610]" count="4" pageId="9" pageNumber="80" type="female">4♀</specimenCount>
// “
<collectingRegion id="D11C5CB6292B6A78FAFFE077FA29F9B6" box="[1298,1436,1584,1610]" country="Argentina" name="Cordoba" pageId="9" pageNumber="80">CORDOBA</collectingRegion>
-
<collectingCountry id="6BCFD2C4292B6A78FF47E013FEF9F992" box="[170,332,1620,1646]" name="Argentina" pageId="9" pageNumber="80">ARGENTINA</collectingCountry>
<location id="1607C48F292B6A78FEBBE012FD0EF992" LSID="urn:lsid:plazi:treatment:9B712342292B6A7FFF7AE279FC20FD0D:1607C48F292B6A78FEBBE012FD0EF992" box="[342,699,1620,1647]" country="Argentina" county="Bahia" municipality="FeLipe SoLa" name="Dep. de Calamuchita El Sauce" pageId="9" pageNumber="80" stateProvince="Ciudad Autonoma de Buenos Aires">Dep. de Calamuchita El Sauce</location>
<date id="6766B494292B6A78FD2BE013FC9BF992" box="[710,814,1620,1646]" pageId="9" pageNumber="80" value="1938-12">
<collectingDate id="77224D7C292B6A78FD2BE013FC9BF992" box="[710,814,1620,1646]" pageId="9" pageNumber="80" value="1938-12">XII-1938</collectingDate>
</date>
<collectorName id="BE2DF782292B6A78FCD4E013FC43F992" box="[825,1014,1620,1647]" pageId="9" pageNumber="80">Manuel J. Viana</collectorName>
[white label, printed] // “
<collectorName id="BE2DF782292B6A78FAC8E013FEB9F96E" pageId="9" pageNumber="80">F. Monros Collection</collectorName>
1959” [White label, printed] // (
<collectionCode id="75C90A91292B6A78FD7BE03FFCA4F96E" box="[662,785,1656,1682]" pageId="9" pageNumber="80">USNMNH</collectionCode>
&amp;
<collectionCode id="75C90A91292B6A78FCADE03FFC3EF96E" box="[832,907,1656,1682]" pageId="9" pageNumber="80">DSPC</collectionCode>
). All
<typeStatus id="CC632CF6292B6A78FC3CE03EFB89F96F" box="[977,1084,1657,1683]" pageId="9" pageNumber="80" type="paratype">paratypes</typeStatus>
also labelled: // “
<taxonomicName id="D4D8E9D7292B6A78FAF5E03FFEB1F94A" authorityName="Sassi" authorityYear="2025" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="tommasoi" status="sp. nov.">
<emphasis id="21AC4E46292B6A78FAF5E03FFEB1F94A" italics="true" pageId="9" pageNumber="80">Metallactus tommasoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3A9FF33D292B6A78FEE6E0DBFED5F94A" box="[267,352,1692,1718]" pageId="9" pageNumber="80" rank="species">
<emphasis id="21AC4E46292B6A78FEE6E0DBFED5F94A" bold="true" box="[267,352,1692,1718]" pageId="9" pageNumber="80">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="21AC4E46292B6A78FE8DE0DBFED0F94A" bold="true" box="[352,357,1692,1718]" pageId="9" pageNumber="80">.</emphasis>
<typeStatus id="CC632CF6292B6A78FE81E0DBFE54F94A" box="[364,481,1692,1718]" pageId="9" pageNumber="80" type="paratype">PARATYPUS</typeStatus>
D. Sassi des.” [red label, printed] //.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF292B6A78FF2AE0F8FCADF84E" pageId="9" pageNumber="80" type="materials_examined">
<paragraph id="13679254292B6A78FF2AE0F8FB89F872" blockId="9.[151,1437,1224,2043]" pageId="9" pageNumber="80">
<materialsCitation id="A3B09809292B6A78FF2AE0F8FB82F872" collectingDate="2013-12-12" collectingDateMax="2013-12-15" collectingDateMin="2013-12-12" collectionCode="UDPC" country="Paraguay" location="Morombi" pageId="9" pageNumber="80" specimenCount="2" stateProvince="Caaguazu">
<emphasis id="21AC4E46292B6A78FF2AE0F8FDACF926" bold="true" box="[199,537,1727,1754]" italics="true" pageId="9" pageNumber="80">
<typeStatus id="CC632CF6292B6A78FF2AE0F8FDACF926" box="[199,537,1727,1754]" isOtherTypeStatus="true" pageId="9" pageNumber="80">Additional material examined</typeStatus>
</emphasis>
.A single female was examined, bearing the following information:
<emphasis id="21AC4E46292B6A78FB14E087FA20F926" bold="true" box="[1273,1429,1728,1754]" pageId="9" pageNumber="80">
<collectingCountry id="6BCFD2C4292B6A78FB14E087FA20F926" box="[1273,1429,1728,1754]" name="Paraguay" pageId="9" pageNumber="80">PARAGUAY</collectingCountry>
</emphasis>
:
<emphasis id="21AC4E46292B6A78FF7AE0A3FEBAF902" bold="true" box="[151,271,1764,1790]" pageId="9" pageNumber="80">
<collectingRegion id="D11C5CB6292B6A78FF7AE0A3FEBAF902" box="[151,271,1764,1790]" country="Paraguay" name="Caaguazu" pageId="9" pageNumber="80">Caaguazu</collectingRegion>
</emphasis>
:
<location id="1607C48F292B6A78FEF0E0A3FE32F902" LSID="urn:lsid:plazi:treatment:9B712342292B6A7FFF7AE279FC20FD0D:1607C48F292B6A78FEF0E0A3FE32F902" box="[285,391,1764,1790]" country="Paraguay" name="Morombi" pageId="9" pageNumber="80" stateProvince="Caaguazu">Morombi</location>
<date id="6766B494292B6A78FE63E0A3FDF4F903" box="[398,577,1764,1791]" pageId="9" pageNumber="80" value="2013-12-12" valueMax="2013-12-15" valueMin="2013-12-12">
<collectingDate id="77224D7C292B6A78FE63E0A3FDF4F903" box="[398,577,1764,1791]" pageId="9" pageNumber="80" value="2013-12-12" valueMax="2013-12-15" valueMin="2013-12-12">1215.XII.2013</collectingDate>
</date>
(
<collectionCode id="75C90A91292B6A78FDA2E0A3FD15F902" box="[591,672,1764,1790]" pageId="9" pageNumber="80">UDPC</collectionCode>
). The specimen is slightly larger than the recorded range for females of the new species. Additionally, its coloration is partly different, with the elytra being completely yellow except for a small, rounded spot at the apex. Despite matching the rest of the observed characteristics, it was deemed more prudent not to consider this specimen as part of the
<typeStatus id="CC632CF6292B6A78FCF1E116FCF9F897" box="[796,844,1873,1899]" pageId="9" pageNumber="80">type</typeStatus>
series of the new species. Therefore, the presence of
<taxonomicName id="D4D8E9D7292B6A78FF7AE133FE3AF872" authorityName="Sassi" authorityYear="2025" box="[151,399,1908,1934]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="tommasoi">
<emphasis id="21AC4E46292B6A78FF7AE133FE3AF872" box="[151,399,1908,1934]" italics="true" pageId="9" pageNumber="80">Metallactus tommasoi</emphasis>
</taxonomicName>
in
<collectingRegion id="D11C5CB6292B6A78FE5EE132FDAEF873" box="[435,539,1909,1935]" country="Paraguay" name="Asuncion" pageId="9" pageNumber="80">Paraguay</collectingRegion>
remains doubtful, pending further confirmation
</materialsCitation>
.
</paragraph>
<paragraph id="13679254292B6A78FF2AE1DFFCADF84E" blockId="9.[151,1437,1224,2043]" box="[199,792,1943,1970]" pageId="9" pageNumber="80">
<materialsCitation id="A3B09809292B6A78FF2AE1DFFCA0F84E" box="[199,789,1943,1970]" country="Argentina" location="Felipe Sola" pageId="9" pageNumber="80" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
<emphasis id="21AC4E46292B6A78FF2AE1DFFEEAF84E" bold="true" box="[199,351,1943,1970]" italics="true" pageId="9" pageNumber="80">
<typeStatus id="CC632CF6292B6A78FF2AE1DFFF4BF84E" box="[199,254,1944,1970]" pageId="9" pageNumber="80">Type</typeStatus>
locality.
</emphasis>
<location id="1607C48F292B6A78FE8BE1DFFE50F84E" LSID="urn:lsid:plazi:treatment:9B712342292B6A7FFF7AE279FC20FD0D:1607C48F292B6A78FE8BE1DFFE50F84E" box="[358,485,1944,1970]" country="Argentina" name="Felipe Sola" pageId="9" pageNumber="80" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
(
<collectingRegion id="D11C5CB6292B6A78FE1EE1DFFD3BF84E" box="[499,654,1944,1970]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="9" pageNumber="80">Buenos Aires</collectingRegion>
,
<collectingCountry id="6BCFD2C4292B6A78FD75E1DFFCB8F84E" box="[664,781,1944,1970]" name="Argentina" pageId="9" pageNumber="80">Argentina</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF292B6A78FF2AE1FCFDE6F806" pageId="9" pageNumber="80" type="etymology">
<paragraph id="13679254292B6A78FF2AE1FCFDE6F806" blockId="9.[151,1437,1224,2043]" pageId="9" pageNumber="80">
<emphasis id="21AC4E46292B6A78FF2AE1FCFEFDF82A" bold="true" box="[199,328,1979,2006]" italics="true" pageId="9" pageNumber="80">Etymology.</emphasis>
The new species is dedicated to my son Tommaso, a keen naturalist and skilled biologist, specialized in a field different from biosystematics.
</paragraph>
</subSubSection>
<caption id="47A7C2DC29286A7BFF7AE007FDDBF938" ID-DOI="http://doi.org/10.5281/zenodo.14798008" ID-Zenodo-Dep="14798008" httpUri="https://zenodo.org/record/14798008/files/figure.png" pageId="10" pageNumber="81" startId="10.[151,250,1600,1624]" targetBox="[167,1420,181,1566]" targetPageId="10" targetType="figure">
<paragraph id="1367925429286A7BFF7AE007FDDBF938" blockId="10.[151,1436,1600,1733]" pageId="10" pageNumber="81">
<emphasis id="21AC4E4629286A7BFF7AE007FEA0F9A4" bold="true" box="[151,277,1600,1624]" pageId="10" pageNumber="81">FIGURE 3.</emphasis>
<emphasis id="21AC4E4629286A7BFEF2E007FDBCF9A4" box="[287,521,1600,1624]" italics="true" pageId="10" pageNumber="81">
<taxonomicName id="D4D8E9D729286A7BFEF2E007FDB3F9A4" authorityName="Sassi" authorityYear="2025" box="[287,518,1600,1624]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="tommasoi">Metallactus tommasoi</taxonomicName>
.
</emphasis>
ab: habitus, dorsal and lateral view (HT); c: frontal view, male (HT); d: frontal view, female (PT, Felipe Solá, Buenos Aires, Argentina); eh: colour variations (e: female, PT, Felipe Solá, Buenos Aires, Argentina; f: male, PT, Felipe Solá, Buenos Aires, Argentina; g: male, PT, Tandil, Buenos Aires, Argentina; h: male, PT, Felipe Solá, Buenos Aires, Argentina). Images not to scale.
</paragraph>
</caption>
<subSubSection id="5BC2C1DF29286A7AFF2AE151FBFBFF29" lastPageId="11" lastPageNumber="82" pageId="10" pageNumber="81" type="distribution">
<paragraph id="1367925429286A7BFF2AE151FDC7F8CD" blockId="10.[151,1437,1814,2021]" box="[199,626,1814,1841]" pageId="10" pageNumber="81">
<emphasis id="21AC4E4629286A7BFF2AE151FEE6F8CD" bold="true" box="[199,339,1814,1841]" italics="true" pageId="10" pageNumber="81">Distribution</emphasis>
.
<collectingCountry id="6BCFD2C429286A7BFE8DE150FE67F8CD" box="[352,466,1815,1841]" name="Argentina" pageId="10" pageNumber="81">Argentina</collectingCountry>
,
<collectingCountry id="6BCFD2C429286A7BFE30E150FDF0F8CD" box="[477,581,1815,1841]" name="Paraguay" pageId="10" pageNumber="81">Paraguay</collectingCountry>
(?).
</paragraph>
<paragraph id="1367925429286A7AFF2AE17CFBFBFF29" blockId="10.[151,1437,1814,2021]" lastBlockId="11.[151,1437,151,285]" lastPageId="11" lastPageNumber="82" pageId="10" pageNumber="81">
<emphasis id="21AC4E4629286A7BFF2AE17CFE10F8A9" bold="true" box="[199,421,1851,1877]" italics="true" pageId="10" pageNumber="81">Comparative notes</emphasis>
. The new species chromatically differs from both
<taxonomicName id="D4D8E9D729286A7BFBE8E17BFBC1F8A9" authorityName="Suffrian" authorityYear="1866" box="[1029,1140,1852,1877]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629286A7BFBE8E17BFBC1F8A9" box="[1029,1140,1852,1877]" italics="true" pageId="10" pageNumber="81">M. minax</emphasis>
</taxonomicName>
and
<taxonomicName id="D4D8E9D729286A7BFB5BE17BFAFEF8A9" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1206,1355,1851,1877]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E4629286A7BFB5BE17BFAFEF8A9" box="[1206,1355,1851,1877]" italics="true" pageId="10" pageNumber="81">M. abstersus</emphasis>
</taxonomicName>
in the coloration of the head, which is predominantly yellow on frons and clypeus rather than totally or almost totally black. Additionally, the elytral colour pattern appears quite characteristic, with the suture generally broadly banded in black, and the presence of spots in the middle of the elytron predominantly oriented longitudinally. Moreover, elytral punctation is usually less dense than in the other species, and arranged in almost regular rows. The most evident difference, however, lies in the shape of the median lobe, where the median bulge on the ventral surface is significantly reduced compared to what is observed in the other species of the group.
</paragraph>
</subSubSection>
<subSubSection id="5BC2C1DF29296A7FFF2AE698FC20FD0D" lastPageId="14" lastPageNumber="85" pageId="11" pageNumber="82" type="description">
<paragraph id="1367925429296A7AFF2AE698FCB1FEE1" blockId="11.[151,1437,151,285]" pageId="11" pageNumber="82">
<emphasis id="21AC4E4629296A7AFF2AE698FE04FF05" bold="true" box="[199,433,222,249]" italics="true" pageId="11" pageNumber="82">Description of male.</emphasis>
Habitus in figs 3ab (HT). BL =
<quantity id="D4203FB129296A7AFCC6E699FC01FF05" box="[811,948,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.3" metricValueMin="5.2" pageId="11" pageNumber="82" unit="mm" value="5.25" valueMax="5.3" valueMin="5.2">5.25.3 mm</quantity>
, BW = 3.0
<quantity id="D4203FB129296A7AFBD0E698FB21FF05" box="[1085,1172,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="82" unit="mm" value="3.1">3.1 mm</quantity>
, PL =
<quantity id="D4203FB129296A7AFB33E698FAD2FF05" box="[1246,1383,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="11" pageNumber="82" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
, PW =
<quantity id="D4203FB129296A7AFF43E745FE8DFEE1" box="[174,312,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="11" pageNumber="82" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
. Interocular distance 11.411.6 % of BL.
</paragraph>
<caption id="47A7C2DC29296A7AFF7AE07DFE5AF942" ID-DOI="http://doi.org/10.5281/zenodo.14798010" ID-Zenodo-Dep="14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="11" pageNumber="82" startId="11.[151,250,1594,1618]" targetBox="[171,1416,309,1554]" targetPageId="11" targetType="figure">
<paragraph id="1367925429296A7AFF7AE07DFE5AF942" blockId="11.[151,1437,1594,1726]" pageId="11" pageNumber="82">
<emphasis id="21AC4E4629296A7AFF7AE07DFEA6F9AD" bold="true" box="[151,275,1594,1618]" pageId="11" pageNumber="82">FIGURE 4.</emphasis>
ad:
<taxonomicName id="D4D8E9D729296A7AFEA2E07DFDB9F9AE" authorityName="Suffrian" authorityYear="1866" box="[335,524,1594,1618]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E4629296A7AFEA2E07DFDB9F9AE" box="[335,524,1594,1618]" italics="true" pageId="11" pageNumber="82">Metallactus minax</emphasis>
</taxonomicName>
, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg:
<taxonomicName id="D4D8E9D729296A7AFC59E018FB8FF98A" authorityName="Sassi" authorityYear="2025" box="[948,1082,1631,1654]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="tommasoi">
<emphasis id="21AC4E4629296A7AFC59E018FB8FF98A" box="[948,1082,1631,1654]" italics="true" pageId="11" pageNumber="82">M. tommasoi</emphasis>
</taxonomicName>
, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment.
</paragraph>
</caption>
<paragraph id="1367925429296A7DFF2AE157FEEEFEE1" blockId="11.[151,1436,1808,2015]" lastBlockId="12.[151,1437,151,285]" lastPageId="12" lastPageNumber="83" pageId="11" pageNumber="82">
<emphasis id="21AC4E4629296A7AFF2AE157FEB1F8D6" box="[199,260,1808,1834]" italics="true" pageId="11" pageNumber="82">Head</emphasis>
: usually black, with clypeus and lower part of frons yellow. Usually, yellow pattern extended upwards in form of two curved spots along inner border of eyes, leaving a black line in middle. At times such spots reduced to small round markings between upper lobes of eyes. Labrum light yellow as well. Surface of upper frons and vertex quite convex, head punctation sparse, moderately impressed on vertex, denser and stronger on clypeus and along inner rim of eyes. Setosity almost missing on vertex and frons but, again, thick and adpressed along lower part of inner eye margins and along median line between eyes. Mid-cranial suture usually scarcely visible. Upper lobes of eyes well separated along median lines. Ocular lines shallow, strictly adhering to ocular rim, marked by rows of small punctures. Ocular canthus rounded, scarcely setose. Antennae rather short, slender, blackish with first five antennomeres partly yellowish, 35 bright, subcylindrical, the subsequent ones dull, moderately flattened and more diffusedly setose.
</paragraph>
<caption id="47A7C2DC292E6A7DFF7AE091FA28F8EF" ID-DOI="http://doi.org/10.5281/zenodo.14798012" ID-Zenodo-Dep="14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="12" pageNumber="83" startId="12.[151,250,1750,1774]" targetBox="[159,1425,313,1713]" targetPageId="12" targetType="figure">
<paragraph id="13679254292E6A7DFF7AE091FA28F8EF" blockId="12.[151,1437,1750,1811]" pageId="12" pageNumber="83">
<emphasis id="21AC4E46292E6A7DFF7AE091FEA6F912" bold="true" box="[151,275,1750,1774]" pageId="12" pageNumber="83">FIGURE 5.</emphasis>
a:
<taxonomicName id="D4D8E9D7292E6A7DFEDFE091FE5BF912" authorityName="Suffrian" authorityYear="1866" box="[306,494,1750,1774]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="83" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E46292E6A7DFEDFE091FE5BF912" box="[306,494,1750,1774]" italics="true" pageId="12" pageNumber="83">Metallactus minax</emphasis>
</taxonomicName>
, endophallus [0.3 mm]; bd: spermatheca. b:
<taxonomicName id="D4D8E9D7292E6A7DFC2CE090FB94F912" authorityName="Suffrian" authorityYear="1866" box="[961,1057,1751,1774]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="83" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E46292E6A7DFC2CE090FB94F912" box="[961,1057,1751,1774]" italics="true" pageId="12" pageNumber="83">M. minax</emphasis>
</taxonomicName>
[0.6 mm]; c:
<taxonomicName id="D4D8E9D7292E6A7DFB41E090FA84F912" authorityName="Sassi" authorityYear="2025" box="[1196,1329,1751,1774]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="83" phylum="Arthropoda" rank="species" species="tommasoi">
<emphasis id="21AC4E46292E6A7DFB41E090FA84F912" box="[1196,1329,1751,1774]" italics="true" pageId="12" pageNumber="83">M. tommasoi</emphasis>
</taxonomicName>
[0.6 mm]; d:
<taxonomicName id="D4D8E9D7292E6A7DFF5CE0BCFE86F8EE" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[177,307,1786,1810]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="83" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E46292E6A7DFF5CE0BCFE86F8EE" box="[177,307,1786,1810]" italics="true" pageId="12" pageNumber="83">M. abstersus</emphasis>
</taxonomicName>
[0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment.
</paragraph>
</caption>
<paragraph id="13679254292E6A7CFF2AE122FEA8FF05" blockId="12.[151,1437,1892,2027]" lastBlockId="13.[151,1436,151,249]" lastPageId="13" lastPageNumber="84" pageId="12" pageNumber="83">
<emphasis id="21AC4E46292E6A7DFF2AE122FE83F882" box="[199,310,1893,1918]" italics="true" pageId="12" pageNumber="83">Pronotum</emphasis>
: black with variable yellow pattern. Usually lateral and anterior margins are yellow, as well as outer part of posterior one. Besides, a longitudinal yellow line present at center of disc, starting from anterior margin and often reaching posterior one. In this case such yellow line broadening backwards and containing small black fleck just in front of scutellum. Pronotal outline weakly transverse, regularly convex. Lateral margins narrow, regularly curved with maximum width just behind half of length. Posterolateral impression short, shallow. Surface moderately lustrous with scattered, well-impressed, often double-sized punctation, denser on sides, almost obliterated on central part of disc.
</paragraph>
<caption id="47A7C2DC292F6A7CFF7AE5F3FBCFFBA0" ID-DOI="http://doi.org/10.5281/zenodo.14798014" ID-Zenodo-Dep="14798014" httpUri="https://zenodo.org/record/14798014/files/figure.png" pageId="13" pageNumber="84" startId="13.[151,250,948,972]" targetBox="[151,1436,273,910]" targetPageId="13" targetType="figure">
<paragraph id="13679254292F6A7CFF7AE5F3FBCFFBA0" blockId="13.[151,1437,948,1116]" pageId="13" pageNumber="84">
<emphasis id="21AC4E46292F6A7CFF7AE5F3FEA6FC30" bold="true" box="[151,275,948,972]" pageId="13" pageNumber="84">FIGURE 6.</emphasis>
Distribution maps for the
<taxonomicName id="D4D8E9D7292F6A7CFDF0E5F3FD6CFC30" authorityName="Suffrian" authorityYear="1866" box="[541,729,948,972]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E46292F6A7CFDF0E5F3FD6CFC30" box="[541,729,948,972]" italics="true" pageId="13" pageNumber="84">Metallactus minax</emphasis>
</taxonomicName>
species-group. a:
<taxonomicName id="D4D8E9D7292F6A7CFC7DE5F2FC45FC30" authorityName="Suffrian" authorityYear="1866" box="[912,1008,949,972]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E46292F6A7CFC7DE5F2FC45FC30" box="[912,1008,949,972]" italics="true" pageId="13" pageNumber="84">M. minax</emphasis>
</taxonomicName>
; b:
<taxonomicName id="D4D8E9D7292F6A7CFBFBE5F2FB2DFC30" authorityName="D. Sassi" authorityYear="2025" baseAuthorityName="Suffrian" baseAuthorityYear="1866" box="[1046,1176,948,972]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="abstersus">
<emphasis id="21AC4E46292F6A7CFBFBE5F2FB2DFC30" box="[1046,1176,948,972]" italics="true" pageId="13" pageNumber="84">M. abstersus</emphasis>
</taxonomicName>
; c:
<emphasis id="21AC4E46292F6A7CFB50E5F2FAFDFC30" box="[1213,1352,949,972]" italics="true" pageId="13" pageNumber="84">
<taxonomicName id="D4D8E9D7292F6A7CFB50E5F2FAF0FC30" authorityName="Sassi" authorityYear="2025" box="[1213,1349,949,972]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="84" phylum="Arthropoda" rank="species" species="tommasoi">M. tommasoi</taxonomicName>
.
</emphasis>
Circles: data from specimens analysed during the study. Star in fig. b: the available information allows to point out the presence in the state but without more precise data on the location. Star in fig. c: Identification uncertain based on females only, therefore excluded from the type series. In fig. a: record of the species for the state of Pará, indicated on the label of one of the paratypes (see text), is not shown on the map because this data is considered to require further confirmation.
</paragraph>
</caption>
<paragraph id="13679254292F6A7CFF2AE2E9FC75FB11" blockId="13.[151,1437,1198,2016]" pageId="13" pageNumber="84">
<taxonomicName id="D4D8E9D7292F6A7CFF2AE2E9FE80FB34" box="[199,309,1198,1224]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="13" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="21AC4E46292F6A7CFF2AE2E9FE80FB34" box="[199,309,1198,1224]" italics="true" pageId="13" pageNumber="84">Scutellum</emphasis>
</taxonomicName>
: completely black, moderately raised. Apex truncated in a straight line or mildly rounded. Surface smooth, minutely and sparsely punctured, with few, short, whitish setae.
</paragraph>
<paragraph id="13679254292F6A7CFF2AE2B1FE61F918" blockId="13.[151,1437,1198,2016]" pageId="13" pageNumber="84">
<emphasis id="21AC4E46292F6A7CFF2AE2B1FEB8FAEC" box="[199,269,1270,1296]" italics="true" pageId="13" pageNumber="84">Elytra</emphasis>
: elytral colour pattern remarkably variable as well. Often, elytra yellow with common, large, black stripe along suture, extending from anterior margin to elytral clivus. Black stripe, when complete, usually shortly broadened at its anterior third. Occasionally, black stripe may diminish progressively to almost disappearing towards elytral base, however, the black broadening on anterior third tends to persist in form of common rounded spot or two distinct, sometimes blurred markings. More rarely, black stripe reduced to very narrow black line along suture. Specimens with more extended dark elytral pattern, exhibit nearly rectangular or vaguely C-shaped, longitudinal black spot at middle of elytron. In some specimens, longitudinal spot merging with sutural stripe to give a pattern where dark colouration predominates over yellow (
<figureCitation id="8BE38ED1292F6A7CFD04E3B5FCF5F9F0" box="[745,832,1522,1548]" captionStart="FIGURE 3" captionStartId="10.[151,250,1600,1624]" captionTargetBox="[167,1420,181,1566]" captionTargetId="figure-33@10.[151,1436,181,1577]" captionTargetPageId="10" captionText="FIGURE 3. Metallactus tommasoi. ab: habitus, dorsal and lateral view (HT); c: frontal view, male (HT); d: frontal view, female (PT, Felipe Solá, Buenos Aires, Argentina); eh: colour variations (e: female, PT, Felipe Solá, Buenos Aires, Argentina; f: male, PT, Felipe Solá, Buenos Aires, Argentina; g: male, PT, Tandil, Buenos Aires, Argentina; h: male, PT, Felipe Solá, Buenos Aires, Argentina). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.14798008" httpUri="https://zenodo.org/record/14798008/files/figure.png" pageId="13" pageNumber="84">Fig. 3h</figureCitation>
). Humeral callus always spotted with black even in lighter specimens. In a single specimen (
<figureCitation id="8BE38ED1292F6A7CFDB8E051FD13F9CC" box="[597,678,1558,1584]" captionStart="FIGURE 3" captionStartId="10.[151,250,1600,1624]" captionTargetBox="[167,1420,181,1566]" captionTargetId="figure-33@10.[151,1436,181,1577]" captionTargetPageId="10" captionText="FIGURE 3. Metallactus tommasoi. ab: habitus, dorsal and lateral view (HT); c: frontal view, male (HT); d: frontal view, female (PT, Felipe Solá, Buenos Aires, Argentina); eh: colour variations (e: female, PT, Felipe Solá, Buenos Aires, Argentina; f: male, PT, Felipe Solá, Buenos Aires, Argentina; g: male, PT, Tandil, Buenos Aires, Argentina; h: male, PT, Felipe Solá, Buenos Aires, Argentina). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.14798008" httpUri="https://zenodo.org/record/14798008/files/figure.png" pageId="13" pageNumber="84">
Fig.
<quantity id="D4203FB1292F6A7CFD65E051FD13F9CC" box="[648,678,1558,1584]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="13" pageNumber="84" unit="g" value="3.0">3g</quantity>
</figureCitation>
), elytral dark pattern retained but faded into faint light brown design, almost not distinguished from yellow background. As a result, elytra might initially appear completely yellow. Elytral outline cylindrical with sides almost parallel. Lateral margins narrow, scarcely visible from above. Scutellar area not raised. Humeral callus scarcely prominent, not punctured. Elytral punctation moderately impressed, mostly sparse over anterior half, posteriorly arranged in almost regular rows. Intervals flat. Epipleura slightly convex, with smooth, impunctate surface.
</paragraph>
<paragraph id="13679254292F6A7CFF2AE0A9FCFAF8D1" blockId="13.[151,1437,1198,2016]" pageId="13" pageNumber="84">
<taxonomicName id="D4D8E9D7292F6A7CFF2AE0A9FE87F8F4" authorityName="Meyen" authorityYear="1834" box="[199,306,1774,1800]" family="Trichomycteridae" genus="Pygidium" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="84" phylum="Chordata" rank="genus">
<emphasis id="21AC4E46292F6A7CFF2AE0A9FE87F8F4" box="[199,306,1774,1800]" italics="true" pageId="13" pageNumber="84">Pygidium</emphasis>
</taxonomicName>
: usually black (but almost completely yellowish in some specimens), smooth, covered by thick, adpressed, whitish setae and sparse well-impressed punctures.
</paragraph>
<paragraph id="13679254292F6A7CFF2AE171FBC2F81C" blockId="13.[151,1437,1198,2016]" pageId="13" pageNumber="84">
<emphasis id="21AC4E46292F6A7CFF2AE171FE89F8AC" box="[199,316,1846,1872]" italics="true" pageId="13" pageNumber="84">Underside</emphasis>
: ventral surface black (sometimes part of the rear margin of the fifth abdominal ventrite yellowish), matt, covered with long, thick, adpressed, whitish setae, except shiny, scarcely setose, sparsely punctured posterior part of hypomera. Prosternal process narrow, fairly grooved at middle, with apex short triangular. Surface of prosternal process covered with long, thick, semi-erect whitish setae. Fifth abdominal ventrite with deep, heart-shaped, bald, shiny, impunctate median depression and deeply notched posterior margin.
</paragraph>
<paragraph id="13679254292C6A7FFF2AE6D0FEF5FF29" blockId="14.[151,1437,151,753]" pageId="14" pageNumber="85">
<emphasis id="21AC4E46292C6A7FFF2AE6D0FF49FF4C" box="[199,252,151,176]" italics="true" pageId="14" pageNumber="85">Legs</emphasis>
: black with yellow stripe along anterior margin of anterior femora. Median and posterior femora partly yellow as well.
</paragraph>
<paragraph id="13679254292C6A7FFF2AE698FECAFE75" blockId="14.[151,1437,151,753]" pageId="14" pageNumber="85">
<emphasis id="21AC4E46292C6A7FFF2AE698FEC5FF05" box="[199,368,223,249]" italics="true" pageId="14" pageNumber="85">Male genitalia</emphasis>
: Median lobe of aedeagus (
<figureCitation id="8BE38ED1292C6A7FFD56E698FC99FF05" box="[699,812,223,249]" captionStart="FIGURE 4" captionStartId="11.[151,250,1594,1618]" captionTargetBox="[171,1416,309,1554]" captionTargetId="figure-80@11.[171,1416,309,1570]" captionTargetPageId="11" captionText="FIGURE 4. ad: Metallactus minax, median lobe of aedeagus (LT) in ventral (a), dorsal (b) and lateral (c) view [1.5 mm]; d: variation in aedeagal morphology (specimen from “São Paulo”, Brazil); eg: M. tommasoi, median lobe of aedeagus (HT) in ventral (e), dorsal (f) and lateral (g) view [1.5 mm]; sd: indentation at sides of median carina (see text for explanations). Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798010" httpUri="https://zenodo.org/record/14798010/files/figure.png" pageId="14" pageNumber="85">Fig. 4eg</figureCitation>
) subcylindrical with apex rounded, flattened, paddleshaped in ventral and dorsal view, almost straight in lateral view. Setose depressions shifted to very lateral border of ventral surface, very sender, barely visible, marked with very few, tiny setae. Ventral surface just below apex raised in moderate, squat, roughly angulate median bulge. Endophallus very similar to that of
<taxonomicName id="D4D8E9D7292C6A7FFBB5E70CFB75FE98" authorityName="Suffrian" authorityYear="1866" box="[1112,1216,331,356]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="85" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="21AC4E46292C6A7FFBB5E70CFB75FE98" box="[1112,1216,331,356]" italics="true" pageId="14" pageNumber="85">M. minax</emphasis>
</taxonomicName>
, with no significant differences detected.
</paragraph>
<paragraph id="13679254292C6A7FFF2AE7D4FEC3FE2D" blockId="14.[151,1437,151,753]" pageId="14" pageNumber="85">
<emphasis id="21AC4E46292C6A7FFF2AE7D4FEAEFE51" box="[199,283,403,429]" italics="true" pageId="14" pageNumber="85">Female</emphasis>
. BL =
<quantity id="D4203FB1292C6A7FFE9FE7D5FE4BFE51" box="[370,510,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.4" metricValueMin="5.6" pageId="14" pageNumber="85" unit="mm" value="6.0" valueMax="6.4" valueMin="5.6">5.66.4 mm</quantity>
, BW =
<quantity id="D4203FB1292C6A7FFDB0E7D4FD5DFE51" box="[605,744,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="14" pageNumber="85" unit="mm" value="3.55" valueMax="3.9" valueMin="3.2">3.23.9 mm</quantity>
, PL =
<quantity id="D4203FB1292C6A7FFCD7E7D4FC73FE51" box="[826,966,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="14" pageNumber="85" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.82.1 mm</quantity>
, PW =
<quantity id="D4203FB1292C6A7FFBCCE7D4FB1BFE51" box="[1057,1198,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.3" metricValueMin="2.8" pageId="14" pageNumber="85" unit="mm" value="3.05" valueMax="3.3" valueMin="2.8">2.83.3 mm</quantity>
. Interocular distance 15.617.9 % of BL.
</paragraph>
<paragraph id="13679254292C6A7FFF2AE79CFD1DFDE5" blockId="14.[151,1437,151,753]" pageId="14" pageNumber="85">Female differs in the larger interocular distance and in the larger and stouter body. No significant differences have been noticed regarding the colour pattern.</paragraph>
<paragraph id="13679254292C6A7FFF2AE464FC20FD0D" blockId="14.[151,1437,151,753]" pageId="14" pageNumber="85">
The pit on the fifth abdominal ventrite in females is large and deep. The bottom of the pit is glabrous, matt, impunctate but covered by sparse, tiny wrinkles.The vasculum of the spermatheca (
<figureCitation id="8BE38ED1292C6A7FFBC7E400FBC2FD9D" box="[1066,1143,582,609]" captionStart="FIGURE 5" captionStartId="12.[151,250,1750,1774]" captionTargetBox="[159,1425,313,1713]" captionTargetId="figure-83@12.[151,1436,309,1727]" captionTargetPageId="12" captionText="FIGURE 5. a: Metallactus minax, endophallus [0.3 mm]; bd: spermatheca. b: M. minax [0.6 mm]; c: M. tommasoi [0.6 mm]; d: M. abstersus [0.6 mm]; scIscIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." figureDoi="http://doi.org/10.5281/zenodo.14798012" httpUri="https://zenodo.org/record/14798012/files/figure.png" pageId="14" pageNumber="85">Fig. 5c</figureCitation>
) is moderately pigmented, sickle-shaped with straight, not swollen proximal lobe. The distal lobe is long, not swollen, tapered toward the apex. The ampulla is not pigmented, small, sitting just at basal apex of vasculum. The duct and the sperm gland insertions are perceptibly distinct. The duct is very long, uniform in size, slender, floppy, strictly coiled, with its insertion on the bursa copulatrix forming a rigid, twisted, well pigmented sleeve.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,218 @@
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<mods:title id="FF1C5A7B74939278CEAD1E2C45BF1F4B">A new species of Neosilba (Diptera, Lonchaeidae) associated with fig orchards in Mexico</mods:title>
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<mods:affiliation id="38B118028DC7A9CB1C56DE5065541B3B">Instituto de Ecología AC (INECOL), Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:namePart id="EE2B094369890679BAAF891DC90B2FC0">Lasa, Rodrigo</mods:namePart>
<mods:affiliation id="00666FBBB16072DBF367771E54F3578A">Instituto de Ecología AC (INECOL), Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<subSubSection id="61D96578B737FFDA958EF8A5FD52363E" box="[151,736,1863,1890]" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="297C36F3B737FFDA958EF8A5FD52363E" blockId="1.[151,736,1863,1890]" box="[151,736,1863,1890]" pageId="1" pageNumber="196">
<heading id="7234819FB737FFDA958EF8A5FD52363E" bold="true" box="[151,736,1863,1890]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
<emphasis id="1BB7EAE1B737FFDA958EF8A5FD52363E" bold="true" box="[151,736,1863,1890]" pageId="1" pageNumber="196">
<taxonomicName id="EEC34D70B737FFDA958EF8A5FDC9363E" authority="MacGowan &amp; Lasa" authorityName="Macgowan &amp; Navarro-De-La-Fuente &amp; Williams &amp; Lasa" authorityYear="2025" box="[151,635,1863,1890]" class="Insecta" family="Lonchaeidae" genus="Neosilba" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="flavitarsis" status="sp. nov.">
<emphasis id="1BB7EAE1B737FFDA958EF8A5FECD363E" bold="true" box="[151,383,1863,1890]" italics="true" pageId="1" pageNumber="196">Neosilba flavitarsis</emphasis>
MacGowan &amp; Lasa
</taxonomicName>
<taxonomicNameLabel id="0084579AB737FFDA979AF8AAFD52363E" box="[643,736,1864,1890]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="61D96578B737FFD9958EF873FD5434C1" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196" type="description">
<paragraph id="297C36F3B737FFD9958EF873FB5D3189" blockId="1.[151,1436,1936,2034]" lastBlockId="2.[151,1437,151,1437]" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196">
<emphasis id="1BB7EAE1B737FFDA958EF873FD9036F6" box="[151,546,1936,1962]" italics="true" pageId="1" pageNumber="196">
Description.
<typeStatus id="F6788851B737FFDA9432F872FE2036F6" box="[299,402,1936,1962]" pageId="1" pageNumber="196" type="holotype">Holotype</typeStatus>
Male: Head
</emphasis>
: Eye bare. Frons matt black, ratio of frons width at narrowest point above the lunule to eye width 1:2.1. Frontal and interfrontal setulae short, approximately 0.2x length of orbital seta. Orbital plate shining black, bare apart from the orbital seta. Lunule matt black, parafacial and face slightly silver-grey pollinose. Antenna black, postpedicel length to depth ratio 4.1:1. Arista short plumose, plumosity at maximum extent 0.9x depth of postpedicel. Anterior genal setulae in a single row of 45 along mouth margin.
</paragraph>
<paragraph id="297C36F3B734FFD995DEFF3DFBBB333D" blockId="2.[151,1437,151,1437]" pageId="2" pageNumber="197">
<emphasis id="1BB7EAE1B734FFD995DEFF3DFEAA31A5" box="[199,280,223,249]" italics="true" pageId="2" pageNumber="197">Thorax</emphasis>
: Scutum sub-shining blue-black, covered in rather dense black setula approximately 0.4x as long as the orbital seta. Anepisternum anteriorly with a vertical row of three setae posteriorly with a vertical row of six, remainder of sclerite covered in long setulae, approximately 0.5x as long as the posterior setae. Katepisternum with two setae located near dorsal margin, anterior slightly weaker than the posterior, a scattering of setulae located anterior to these near the dorsal margin. Proepisternum and proepimeron each with a single long, recurved seta. Prosternum bare. Scutellum; scutum shining blue-black as thorax, margin between lateral and apical setae with six long setulae on right side, five on left, two setulae between apical setae. Calypter white with a white fringe within which, centrally placed, are nine long black setulae 2.5x longer than the other setulae in the fringe. Wing lightly fumose, less so along posterior margin, veins brownish-black, wing length 5.0 mm. Legs black, basal and second tarsomere of all legs brownish-yellow, covered with small black setulae, fore and hind basitarsi with a ventral fringe of stiff golden setulae. Halter greyish apically, contrasting with the black stem.
</paragraph>
<paragraph id="297C36F3B734FFD995DEFD89FEB333F5" blockId="2.[151,1437,151,1437]" pageId="2" pageNumber="197">
<emphasis id="1BB7EAE1B734FFD995DEFD89FE8333D9" box="[199,305,619,645]" italics="true" pageId="2" pageNumber="197">Abdomen</emphasis>
: sternite 1 bare, tergite 5 with a group of stronger spine-like setae apically. Yellow cerci often visible ventrally.
</paragraph>
<paragraph id="297C36F3B734FFD995DEFD51FAA832B1" blockId="2.[151,1437,151,1437]" pageId="2" pageNumber="197">
<emphasis id="1BB7EAE1B734FFD995DEFD51FE333391" box="[199,385,691,717]" italics="true" pageId="2" pageNumber="197">Male terminalia</emphasis>
: (
<figureCitation id="B1F82A76B734FFD99482FD51FDB83391" box="[411,522,691,717]" captionStart="FIGURES 16" captionStartId="3.[152,270,1936,1960]" captionTargetBox="[391,1196,306,1909]" captionTargetId="figure-83@3.[391,1196,306,1909]" captionTargetPageId="3" captionText="FIGURES 16. Neosilba flavitarsis sp. nov. 1. Holotype male, with the yellow cerci clearly visible at the tip of the abdomen. 2. Allotype female. 3. Male terminalia, lateral view. 4. Male terminalia, ventral view. 5. Postgonites, ventral view. 6. Phallus, lateral view. Ce=cerci, Ep = epandrium, Ph = phallus, Pg = postgonite." figureDoi="http://doi.org/10.5281/zenodo.14797957" httpUri="https://zenodo.org/record/14797957/files/figure.png" pageId="2" pageNumber="197">Figs. 36</figureCitation>
) In lateral view epandrium slightly higher than wide, extended into a narrow, recurved anterior process, with numerous strong setae along ventral margin and posteroventraly. Cerci fused, yellow in colour, 0.8x height of epandrium, with short black setulae on apical half. Surstylus visible on either side of anterior process of epandrium, outer margin with a regular row of strong setae. Inner surface of surstylus with a row of 6 black prensisetae along posterior margin near the base of the cerci, one additional prensisetae anterior to the most ventral prensisetae in this row. Inner surface of surstylus with approximately 15 faint lateral ridges. Postgonite Ushaped, not heavily sclerotised, densely long setulose along medial margin. Phallus, a simple J-shaped tube without processes, basiphallus dark brown with a translucent inner margin, distiphallus pale yellow and laterally flattened. there are tiny spicules present on the translucent part of the basiphallus and the base of the distiphallus.
</paragraph>
<paragraph id="297C36F3B734FFD995DEFC15FCA63521" blockId="2.[151,1437,151,1437]" pageId="2" pageNumber="197">
<emphasis id="1BB7EAE1B734FFD995DEFC15FEA9354D" box="[199,283,1015,1041]" italics="true" pageId="2" pageNumber="197">Female</emphasis>
: resembling the male but generally with shorter pilosity. Ratio of frons width at narrowest point above the lunule to eye width 1: 1.3. Apical segment of aculeus with a pair of dorso-basal setulae which are slightly longer than the apical segment, a pair of ventro-apical setulae the same length as the segment and a pair of dorso-apical setulae just shorter than the length of the apical segment.
</paragraph>
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<collectingCountry id="51D47663B734FFD9947EFB64FE6F35FC" box="[359,477,1158,1184]" name="Mexico" pageId="2" pageNumber="197">MEXICO</collectingCountry>
</emphasis>
:
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,
<collectingMunicipality id="C918AC89B734FFD99770FB65FD7235FD" box="[617,704,1159,1185]" pageId="2" pageNumber="197">Tatatila</collectingMunicipality>
,
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, emerged from
<taxonomicName id="EEC34D70B734FFD9919CFB65FAAB35FC" box="[1157,1305,1159,1184]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="197" phylum="Mollusca" rank="species" species="carica">
<emphasis id="1BB7EAE1B734FFD9919CFB65FAAB35FC" box="[1157,1305,1159,1184]" italics="true" pageId="2" pageNumber="197">Ficus carica</emphasis>
</taxonomicName>
(L.), fruit,
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<collectingDate id="4D39E9DBB734FFD9958EFB49FE8A3599" box="[151,312,1195,1221]" pageId="2" pageNumber="197" value="2023-10">October 2023</collectingDate>
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, leg.
<collectorName id="84365325B734FFD99462FB49FD2A3599" box="[379,664,1195,1221]" pageId="2" pageNumber="197">Rodrigo Lasa. Specimen</collectorName>
deposited in
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,
<collectorName id="84365325B734FFD991B2FB49FA963599" box="[1195,1316,1195,1221]" pageId="2" pageNumber="197">Edinburgh</collectorName>
, specimen NMS-10022229.
</materialsCitation>
</paragraph>
<paragraph id="297C36F3B734FFD995DEFB11FD5434C1" blockId="2.[151,1437,151,1437]" pageId="2" pageNumber="197">
<materialsCitation id="99AB3CAEB734FFD995DEFB11FABB3409" collectingDate="2112-10-02" collectingDateMax="2220-10-02" collectingDateMin="2112-10-02" collectorName="Rodrigo Lasa. Specimen &amp; Edinburgh" country="Mexico" latitude="19.6908" location="National Museum of Scotland" longLatPrecision="7" longitude="-97.1105" municipality="Edinburgh" pageId="2" pageNumber="197" specimenCount="19" specimenCount-female="10" specimenCount-male="9" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<typeStatus id="F6788851B734FFD995DEFB11FE8A3451" box="[199,312,1267,1293]" pageId="2" pageNumber="197" type="paratype">Paratypes</typeStatus>
:
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with the same data as the holotype in
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,
<collectingMunicipality id="C918AC89B734FFD991C8FB11FAF83451" box="[1233,1354,1267,1293]" pageId="2" pageNumber="197">Edinburgh</collectingMunicipality>
,
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<collectingDate id="4D39E9DBB734FFD9904FFB11FEBB346D" pageId="2" pageNumber="197" value="2112-10-02">NMS-10022112</collectingDate>
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, NMS-
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*. * Indicates
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on the same pin.
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with the same data as the holotype deposited in
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, Entomological collection of the
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.
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View file

@ -1,55 +1,56 @@
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,
<figureCitation id="A9FC2A4DFF904726FECAFD1DFEFBA2E1" box="[333,361,721,747]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">13</figureCitation>
<figureCitation id="A9FC2A4DFF904726FECAFD1DFEFBA2E1" box="[333,361,721,747]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="79DD6543FF904726FF10FCDEFA88A422" pageId="3" pageNumber="174" type="materials_examined">
@ -207,13 +208,13 @@ The new species,
<emphasis id="03B3EADAFF904726FB0BFBFEFB7FA446" box="[1164,1261,1074,1100]" italics="true" pageId="3" pageNumber="174">A. nitida</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF904726FB7AFBFEFAD9A447" box="[1277,1355,1074,1101]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="3" pageNumber="174">Figs. 5</figureCitation>
<figureCitation id="A9FC2A4DFF904726FB7AFBFEFAD9A447" box="[1277,1355,1074,1101]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="3" pageNumber="174">Figs. 5</figureCitation>
) in the general coloration and pattern of the forewing, particularly on the more contrasting black elements of the forewing, better developed shine on the surface of the forewing. The new species can be differentiated from the other species based on the features of male genitalia. Uncus is slightly thinner and shorter than in
<taxonomicName id="F6C74D4BFF904726FBAAFB52FB19A4B2" box="[1069,1163,1182,1208]" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="03B3EADAFF904726FBAAFB52FB19A4B2" box="[1069,1163,1182,1208]" italics="true" pageId="3" pageNumber="174">A. nitida</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF904726FB1EFB52FB79A4B2" box="[1177,1259,1182,1208]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">Fig. 14</figureCitation>
<figureCitation id="A9FC2A4DFF904726FB1EFB52FB79A4B2" box="[1177,1259,1182,1208]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">Fig. 14</figureCitation>
). Tegumen with two short, pointed lateral penicular extensions is rounded and not sclerotized in
<taxonomicName id="F6C74D4BFF904726FBA4FB0EFB16A4D6" box="[1059,1156,1218,1244]" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="03B3EADAFF904726FBA4FB0EFB16A4D6" box="[1059,1156,1218,1244]" italics="true" pageId="3" pageNumber="174">A. nitida</emphasis>
@ -227,13 +228,13 @@ The new species,
<emphasis id="03B3EADAFF904726FABBFAC6FA0EA52E" box="[1340,1436,1290,1316]" italics="true" pageId="3" pageNumber="174">A. nitida</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF904726FF18FAE2FF62A542" box="[159,240,1326,1352]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">Fig. 14</figureCitation>
<figureCitation id="A9FC2A4DFF904726FF18FAE2FF62A542" box="[159,240,1326,1352]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">Fig. 14</figureCitation>
),
<taxonomicName id="F6C74D4BFF904726FE83FAE2FE18A542" authorityName="Hreblay, Katona &amp; Toth" authorityYear="2023" box="[260,394,1326,1352]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="himalaya">
<emphasis id="03B3EADAFF904726FE83FAE2FE18A542" box="[260,394,1326,1352]" italics="true" pageId="3" pageNumber="174">A. himalaya</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF904726FE1EFAE2FE7BA543" box="[409,489,1326,1353]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">Fig. 15</figureCitation>
<figureCitation id="A9FC2A4DFF904726FE1EFAE2FE7BA543" box="[409,489,1326,1353]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">Fig. 15</figureCitation>
),
<taxonomicName id="F6C74D4BFF904726FE79FAE2FDE9A542" authorityName="Irungbam &amp; Fric" authorityYear="2025" box="[510,635,1326,1352]" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="albicans" status="sp. nov.">
<emphasis id="03B3EADAFF904726FE79FAE2FDE9A542" box="[510,635,1326,1352]" italics="true" pageId="3" pageNumber="174">A. albicans</emphasis>
@ -242,19 +243,19 @@ The new species,
<taxonomicNameLabel id="188057A1FF904726FD07FAE2FD4BA542" box="[640,729,1326,1352]" pageId="3" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A9FC2A4DFF904726FD60FAE2FCAAA542" box="[743,824,1326,1352]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">Fig. 16</figureCitation>
<figureCitation id="A9FC2A4DFF904726FD60FAE2FCAAA542" box="[743,824,1326,1352]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">Fig. 16</figureCitation>
), and
<taxonomicName id="F6C74D4BFF904726FCFCFAE2FC61A542" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[891,1011,1326,1352]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="infausta">
<emphasis id="03B3EADAFF904726FCFCFAE2FC61A542" box="[891,1011,1326,1352]" italics="true" pageId="3" pageNumber="174">A. infausta</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF904726FB86FAE2FBC1A542" box="[1025,1107,1326,1352]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." pageId="3" pageNumber="174">Fig. 18</figureCitation>
<figureCitation id="A9FC2A4DFF904726FB86FAE2FBC1A542" box="[1025,1107,1326,1352]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="3" pageNumber="174">Fig. 18</figureCitation>
). Saccus is broader and wider than in
<taxonomicName id="F6C74D4BFF904726FF6CFA9EFED9A566" box="[235,331,1362,1388]" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="03B3EADAFF904726FF6CFA9EFED9A566" box="[235,331,1362,1388]" italics="true" pageId="3" pageNumber="174">A. nitida</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF904726FEDCFA9EFE3FA566" box="[347,429,1362,1388]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">Fig. 14</figureCitation>
<figureCitation id="A9FC2A4DFF904726FEDCFA9EFE3FA566" box="[347,429,1362,1388]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">Fig. 14</figureCitation>
), which is narrow.
</paragraph>
</subSubSection>
@ -262,19 +263,19 @@ The new species,
<paragraph id="317836C8FF904726FF40FAB9FD69A74A" blockId="3.[151,1437,785,2036]" pageId="3" pageNumber="174">
<emphasis id="03B3EADAFF904726FF40FAB9FEC3A585" bold="true" box="[199,337,1397,1423]" pageId="3" pageNumber="174">Description</emphasis>
. Adult (
<figureCitation id="A9FC2A4DFF904726FE33FABAFDD7A59A" box="[436,581,1398,1424]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="3" pageNumber="174">Figs. 1, 2, 3</figureCitation>
<figureCitation id="A9FC2A4DFF904726FE33FABAFDD7A59A" box="[436,581,1398,1424]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="3" pageNumber="174">Figs. 1, 2, 3</figureCitation>
) Forewing length:
<quantity id="F63F9B2DFF904726FCA0FABAFCE8A59A" box="[807,890,1398,1424]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="174" unit="mm" value="23.0">23 mm</quantity>
in male. Head, thorax white with dark brown; antennae brown (base white); tegulae with two black-brown lines; patagia with two black-brown bars; thorax black-brown scales; pectus brown; tarsi black-brown bands; abdomen brown, blackish dorsal line mixed with white. Forewing whitish, dark brown suffusion; wide, slightly curved outer edge. Postmedial and antemedial lines wavy, black. Reniform stigmata reddish brown, outward curved, white-bordered. Orbicular stigmata misty gray, ovate, black-lined. Region between spots black. Subterminal line dentate, lighter shade; blackish marks, excurved middle. Terminal line yellowish; cilia mottled. Hindwing dark brown, faint veins, indistinct cell spot, underside shaded, postmedial lines faint; cilia light near veins.
<emphasis id="03B3EADAFF904726FD14F9BEFCADA681" bold="true" box="[659,831,1649,1676]" pageId="3" pageNumber="174">Male genitalia</emphasis>
(
<figureCitation id="A9FC2A4DFF904726FCC8F9BEFC36A686" box="[847,932,1650,1676]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="3" pageNumber="174">Fig. 13</figureCitation>
<figureCitation id="A9FC2A4DFF904726FCC8F9BEFC36A686" box="[847,932,1650,1676]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="3" pageNumber="174">Fig. 13</figureCitation>
). Uncus thin, short, wide at base, narrowing at apex. Tegumen with two short, pointed, hairy lateral extensions. Juxta pentagonal, plate-like, wider at base. Valva rounded, narrows at apex. Sacculus wide, robust, thinly haired. Harpe arises in distal valva, well-sclerotized, curved, pointed, shorter than valva. Saccus broad, wide. Aedeagus large, massive, widens from middle to apex. Carina bilateral, small, pointed plates, highly sclerotized. Vesica ventrally everted, large, sack-like, with diverticula covered in medium pin-like cornuti. Female unknown.
</paragraph>
<paragraph id="317836C8FF904726FF40F885FADAA7A7" blockId="3.[151,1437,785,2036]" pageId="3" pageNumber="174">
<emphasis id="03B3EADAFF904726FF40F885FE2CA769" bold="true" box="[199,446,1865,1891]" pageId="3" pageNumber="174">Genetic information</emphasis>
(
<figureCitation id="A9FC2A4DFF904726FE55F886FDB8A76E" box="[466,554,1866,1892]" captionStart="FIGURE 21" captionStartId="11.[151,250,1435,1459]" captionTargetBox="[156,1425,698,1408]" captionTargetId="figure-236@11.[151,1436,669,1412]" captionTargetPageId="11" captionText="FIGURE 21. Maximum Likelihood tree of the genus Anacronicta based on sequences of mitochondrial cytochrome c oxidase subunit I (COI)." pageId="3" pageNumber="174">Fig. 21</figureCitation>
<figureCitation id="A9FC2A4DFF904726FE55F886FDB8A76E" box="[466,554,1866,1892]" captionStart="FIGURE 21" captionStartId="11.[151,250,1435,1459]" captionTargetBox="[156,1425,698,1408]" captionTargetId="figure-236@11.[151,1436,669,1412]" captionTargetPageId="11" captionText="FIGURE 21. Maximum Likelihood tree of the genus Anacronicta based on sequences of mitochondrial cytochrome c oxidase subunit I (COI)." figureDoi="http://doi.org/10.5281/zenodo.14797507" httpUri="https://zenodo.org/record/14797507/files/figure.png" pageId="3" pageNumber="174">Fig. 21</figureCitation>
).
<specimenCount id="27C1FD41FF904726FDC4F886FD66A76E" box="[579,756,1866,1892]" count="2" pageId="3" pageNumber="174" type="generic">Two specimens</specimenCount>
including

View file

@ -1,53 +1,54 @@
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<mods:title id="D9BBA54B3D238C4E823AD7EE873099C6">Three new species of the genus Anacronicta Warren in Seitz, 1909 (Noctuidae: Pantheinae) from northeast India</mods:title>
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<paragraph id="317836C8FF964720FF10FF13FEB2A0F0" blockId="5.[151,957,151,250]" box="[151,288,223,250]" pageId="5" pageNumber="176">
(
<figureCitation id="A9FC2A4DFF964720FF18FF13FF7DA0F3" box="[159,239,223,249]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="5" pageNumber="176">Figs. 4</figureCitation>
<figureCitation id="A9FC2A4DFF964720FF18FF13FF7DA0F3" box="[159,239,223,249]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="5" pageNumber="176">Figs. 4</figureCitation>
,
<figureCitation id="A9FC2A4DFF964720FF7CFF13FE8AA0F0" box="[251,280,223,250]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="5" pageNumber="176">15</figureCitation>
<figureCitation id="A9FC2A4DFF964720FF7CFF13FE8AA0F0" box="[251,280,223,250]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="5" pageNumber="176">15</figureCitation>
)
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@ -145,19 +146,19 @@ None.
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</emphasis>
(
<figureCitation id="A9FC2A4DFF964720FC80FE7BFCD9A1DB" box="[775,843,439,465]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="5" pageNumber="176">Fig. 4</figureCitation>
<figureCitation id="A9FC2A4DFF964720FC80FE7BFCD9A1DB" box="[775,843,439,465]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="5" pageNumber="176">Fig. 4</figureCitation>
), is very similar to
<taxonomicName id="F6C74D4BFF964720FBA5FE7BFB3BA1DB" authorityName="Hreblay, Katona &amp; Toth" authorityYear="2023" box="[1058,1193,439,465]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="himalaya">
<emphasis id="03B3EADAFF964720FBA5FE7BFB3BA1DB" box="[1058,1193,439,465]" italics="true" pageId="5" pageNumber="176">A. himalaya</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FB3FFE7BFA98A1DB" box="[1208,1290,439,465]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 10</figureCitation>
<figureCitation id="A9FC2A4DFF964720FB3FFE7BFA98A1DB" box="[1208,1290,439,465]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Fig. 10</figureCitation>
), in the wing pattern but differs in the ground colour of the forewing, which is darker than the lighter grey base colour, and the reniform stigmata is filled with golden flaw in
<taxonomicName id="F6C74D4BFF964720FD21FE33FCBCA213" authorityName="Hreblay, Katona &amp; Toth" authorityYear="2023" box="[678,814,511,537]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="himalaya">
<emphasis id="03B3EADAFF964720FD21FE33FCBCA213" box="[678,814,511,537]" italics="true" pageId="5" pageNumber="176">A. himalaya</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FCB9FE33FC00A213" box="[830,914,511,537]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 10</figureCitation>
<figureCitation id="A9FC2A4DFF964720FCB9FE33FC00A213" box="[830,914,511,537]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Fig. 10</figureCitation>
). Based on the male genitalia, the new species belongs to the
<taxonomicName id="F6C74D4BFF964720FEBAFDEFFE25A237" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[317,439,547,573]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="infausta">
<emphasis id="03B3EADAFF964720FEBAFDEFFE25A237" box="[317,439,547,573]" italics="true" pageId="5" pageNumber="176">A. infausta</emphasis>
@ -167,7 +168,7 @@ group due to the structure of the basal diverticulum of the vesica. Compared to
<emphasis id="03B3EADAFF964720FF10FD8BFE8DA26B" box="[151,287,583,609]" italics="true" pageId="5" pageNumber="176">A. himalaya</emphasis>
</taxonomicName>
(
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<figureCitation id="A9FC2A4DFF964720FEA8FD8BFE10A26B" box="[303,386,583,609]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="5" pageNumber="176">Fig. 18</figureCitation>
), the male genitalia of
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<emphasis id="03B3EADAFF964720FD01FD8BFC81A26B" box="[646,787,583,609]" italics="true" pageId="5" pageNumber="176">A. manipura</emphasis>
@ -176,7 +177,7 @@ group due to the structure of the basal diverticulum of the vesica. Compared to
<taxonomicNameLabel id="188057A1FF964720FC9DFD8BFCE6A26B" box="[794,884,583,609]" pageId="5" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A9FC2A4DFF964720FC03FD8BFC44A268" box="[900,982,583,610]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="5" pageNumber="176">Fig. 15</figureCitation>
<figureCitation id="A9FC2A4DFF964720FC03FD8BFC44A268" box="[900,982,583,610]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="5" pageNumber="176">Fig. 15</figureCitation>
) has the much narrower and less curved uncus, the slightly more elongated tegumen, the broader juxta, harpe much shorter than valva, carina with strong toothed field, vesica recurved at the base, and bearing a smaller diverticulum with three further, smaller diverticuli, which are covered with dense cornuti.
</paragraph>
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@ -184,19 +185,19 @@ group due to the structure of the basal diverticulum of the vesica. Compared to
<paragraph id="317836C8FF964720FF40FD1BFAD5A41B" blockId="5.[151,1437,295,1294]" pageId="5" pageNumber="176">
<emphasis id="03B3EADAFF964720FF40FD1BFEC3A2FB" bold="true" box="[199,337,727,753]" pageId="5" pageNumber="176">Description</emphasis>
. Adult (
<figureCitation id="A9FC2A4DFF964720FE2AFD1BFE60A2FB" box="[429,498,727,753]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="5" pageNumber="176">Fig. 4</figureCitation>
<figureCitation id="A9FC2A4DFF964720FE2AFD1BFE60A2FB" box="[429,498,727,753]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="5" pageNumber="176">Fig. 4</figureCitation>
). Forewing length:
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in male. Antennae threadlike. Palps dark grey, few white scales at ends. Forehead dark brown center, white hair on edges and dorsal side. Collar dark brown, white dorsal edge. Tegulae, thorax mix of white, brown, black scales. Forewing dark grey, black markings scattered white scales. Postmedial/antemedial lines black, wide, double, blurred, zigzag. Reniform/orbicular stigmata unclear. Hind wing dark brown, faint veins, no cell spot. Cilia near veins lighter.
<emphasis id="03B3EADAFF964720FCCEFCABFC66A38B" bold="true" box="[841,1012,871,897]" pageId="5" pageNumber="176">Male genitalia</emphasis>
(
<figureCitation id="A9FC2A4DFF964720FB84FCABFBC7A388" box="[1027,1109,871,898]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="5" pageNumber="176">Fig. 15</figureCitation>
<figureCitation id="A9FC2A4DFF964720FB84FCABFBC7A388" box="[1027,1109,871,898]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="5" pageNumber="176">Fig. 15</figureCitation>
). Uncus long, thin. Tegumen short, broad, small, rounded lobes. Vinculum V-shaped, twice as long. Proximal valva wider, sclerotized, tapers at harpe. Cucullus round, sparsely haired. Harpe broad at base, sclerotized, curved, shorter than cucullus. Sacculus flat, almost straight. Clavus simple. Aedeagus short, broad. Carina crescent-shaped, toothed lateral fields. Vesica recurved, small diverticulum at base. Three smaller diverticula with dense cornuti fields. Female unknown.
</paragraph>
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<emphasis id="03B3EADAFF964720FF40FBD7FE28A43F" bold="true" box="[199,442,1051,1077]" pageId="5" pageNumber="176">Genetic information</emphasis>
(
<figureCitation id="A9FC2A4DFF964720FE4CFBD7FDB2A43F" box="[459,544,1051,1077]" captionStart="FIGURE 21" captionStartId="11.[151,250,1435,1459]" captionTargetBox="[156,1425,698,1408]" captionTargetId="figure-236@11.[151,1436,669,1412]" captionTargetPageId="11" captionText="FIGURE 21. Maximum Likelihood tree of the genus Anacronicta based on sequences of mitochondrial cytochrome c oxidase subunit I (COI)." pageId="5" pageNumber="176">Fig. 21</figureCitation>
<figureCitation id="A9FC2A4DFF964720FE4CFBD7FDB2A43F" box="[459,544,1051,1077]" captionStart="FIGURE 21" captionStartId="11.[151,250,1435,1459]" captionTargetBox="[156,1425,698,1408]" captionTargetId="figure-236@11.[151,1436,669,1412]" captionTargetPageId="11" captionText="FIGURE 21. Maximum Likelihood tree of the genus Anacronicta based on sequences of mitochondrial cytochrome c oxidase subunit I (COI)." figureDoi="http://doi.org/10.5281/zenodo.14797507" httpUri="https://zenodo.org/record/14797507/files/figure.png" pageId="5" pageNumber="176">Fig. 21</figureCitation>
). Only the
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of

View file

@ -1,55 +1,56 @@
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@ -72,9 +73,9 @@ urn:lsid:zoobank.org:act:
</paragraph>
<paragraph id="317836C8FF964720FF10FA6BFEAEA5CB" blockId="5.[151,951,1375,1474]" box="[151,316,1447,1474]" pageId="5" pageNumber="176">
(
<figureCitation id="A9FC2A4DFF964720FF18FA6BFE98A5CB" box="[159,266,1447,1474]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Figs. 7, 8</figureCitation>
<figureCitation id="A9FC2A4DFF964720FF18FA6BFE98A5CB" box="[159,266,1447,1474]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Figs. 7, 8</figureCitation>
,
<figureCitation id="A9FC2A4DFF964720FE90FA6BFEA6A5CB" box="[279,308,1447,1473]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="5" pageNumber="176">16</figureCitation>
<figureCitation id="A9FC2A4DFF964720FE90FA6BFEA6A5CB" box="[279,308,1447,1473]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="5" pageNumber="176">16</figureCitation>
)
</paragraph>
</subSubSection>
@ -168,31 +169,31 @@ were deposited in the collections of NZC, ZSI.
<taxonomicNameLabel id="188057A1FF964720FD12F8C5FD7CA729" box="[661,750,1801,1827]" pageId="5" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A9FC2A4DFF964720FD79F8C4FCFAA729" box="[766,872,1800,1827]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Figs. 7, 8</figureCitation>
<figureCitation id="A9FC2A4DFF964720FD79F8C4FCFAA729" box="[766,872,1800,1827]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Figs. 7, 8</figureCitation>
), is similar to
<taxonomicName id="F6C74D4BFF964720FB8DF8C5FB16A729" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[1034,1156,1801,1827]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="infausta">
<emphasis id="03B3EADAFF964720FB8DF8C5FB16A729" box="[1034,1156,1801,1827]" italics="true" pageId="5" pageNumber="176">A. infausta</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FB14F8C4FB71A729" box="[1171,1251,1800,1827]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 11</figureCitation>
<figureCitation id="A9FC2A4DFF964720FB14F8C4FB71A729" box="[1171,1251,1800,1827]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Fig. 11</figureCitation>
) and
<taxonomicName id="F6C74D4BFF964720FAA4F8C5FA0EA728" baseAuthorityName="Leech" baseAuthorityYear="1900" box="[1315,1436,1800,1827]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="03B3EADAFF964720FAA4F8C5FA0EA728" box="[1315,1436,1800,1827]" italics="true" pageId="5" pageNumber="176">A. obscura</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FF18F8E0FF61A74D" box="[159,243,1836,1863]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 12</figureCitation>
<figureCitation id="A9FC2A4DFF964720FF18F8E0FF61A74D" box="[159,243,1836,1863]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Fig. 12</figureCitation>
) in the general coloration and pattern of the forewing, particularly in the presence of dark grey ground coloration, a darker area near the reniform, a slightly curved, darker, and zigzag postmedial line. Externally, it differs clearly from
<taxonomicName id="F6C74D4BFF964720FEF2F8B9FE7FA785" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[373,493,1909,1935]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="infausta">
<emphasis id="03B3EADAFF964720FEF2F8B9FE7FA785" box="[373,493,1909,1935]" italics="true" pageId="5" pageNumber="176">A. infausta</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FE7BF8B8FDD9A785" box="[508,587,1908,1935]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 11</figureCitation>
<figureCitation id="A9FC2A4DFF964720FE7BF8B8FDD9A785" box="[508,587,1908,1935]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Fig. 11</figureCitation>
) and
<taxonomicName id="F6C74D4BFF964720FD0EF8B9FC93A784" baseAuthorityName="Leech" baseAuthorityYear="1900" box="[649,769,1908,1935]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="03B3EADAFF964720FD0EF8B9FC93A784" box="[649,769,1908,1935]" italics="true" pageId="5" pageNumber="176">A. obscura</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FC97F8B8FCF2A785" box="[784,864,1908,1935]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 12</figureCitation>
<figureCitation id="A9FC2A4DFF964720FC97F8B8FCF2A785" box="[784,864,1908,1935]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="5" pageNumber="176">Fig. 12</figureCitation>
) by the more contrasting dark elements of the wing pattern and the ochre-yellowish coloration of the inner part of the hindwings. In the structure of male genitalia,
<taxonomicName id="F6C74D4BFF964720FA03F855FF64A7DC" authorityName="Irungbam &amp; Fric" authorityYear="2025" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="albicans" status="sp. nov.">
<emphasis id="03B3EADAFF964720FA03F855FF64A7DC" italics="true" pageId="5" pageNumber="176">A. albicans</emphasis>
@ -201,23 +202,23 @@ were deposited in the collections of NZC, ZSI.
<taxonomicNameLabel id="188057A1FF964720FF7AF871FEC5A7DD" box="[253,343,1981,2007]" pageId="5" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="A9FC2A4DFF964720FEE1F870FE25A7DC" box="[358,439,1980,2006]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="5" pageNumber="176">Fig. 16</figureCitation>
<figureCitation id="A9FC2A4DFF964720FEE1F870FE25A7DC" box="[358,439,1980,2006]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="5" pageNumber="176">Fig. 16</figureCitation>
) differs from
<taxonomicName id="F6C74D4BFF964720FDD4F871FD5EA7DD" baseAuthorityName="Walker" baseAuthorityYear="1856" box="[595,716,1981,2007]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="infausta">
<emphasis id="03B3EADAFF964720FDD4F871FD5EA7DD" box="[595,716,1981,2007]" italics="true" pageId="5" pageNumber="176">A. infausta</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FD5BF870FCBFA7DD" box="[732,813,1980,2007]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 19</figureCitation>
<figureCitation id="A9FC2A4DFF964720FD5BF870FCBFA7DD" box="[732,813,1980,2007]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="5" pageNumber="176">Fig. 19</figureCitation>
) and
<taxonomicName id="F6C74D4BFF964720FCEBF871FC77A7DC" baseAuthorityName="Leech" baseAuthorityYear="1900" box="[876,997,1980,2007]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="03B3EADAFF964720FCEBF871FC77A7DC" box="[876,997,1980,2007]" italics="true" pageId="5" pageNumber="176">A. obscura</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF964720FC73F870FBD4A7DD" box="[1012,1094,1980,2007]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." pageId="5" pageNumber="176">Fig. 20</figureCitation>
<figureCitation id="A9FC2A4DFF964720FC73F870FBD4A7DD" box="[1012,1094,1980,2007]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="5" pageNumber="176">Fig. 20</figureCitation>
) by the shape of the valva, the shape of the aedeagus, and the position and size of the basal cornuti field of the vesica.
</paragraph>
</subSubSection>
<caption id="65B86640FF954723FF10F85DFD1BA7FA" pageId="6" pageNumber="177" startId="6.[151,264,1937,1961]" targetBox="[195,1376,196,1898]" targetPageId="6" targetType="figure">
<caption id="65B86640FF954723FF10F85DFD1BA7FA" ID-DOI="http://doi.org/10.5281/zenodo.14797499" ID-Zenodo-Dep="14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="6" pageNumber="177" startId="6.[151,264,1937,1961]" targetBox="[195,1376,196,1898]" targetPageId="6" targetType="figure">
<paragraph id="317836C8FF954723FF10F85DFD1BA7FA" blockId="6.[151,1437,1937,2033]" pageId="6" pageNumber="177">
<emphasis id="03B3EADAFF954723FF10F85DFD3BA7A3" bold="true" box="[151,681,1937,1961]" pageId="6" pageNumber="177">
FIGURES 712. Adults of genus
@ -273,7 +274,7 @@ male, Laos;
male, Laos.
</paragraph>
</caption>
<caption id="65B86640FF944722FF10F8BBFAF5A7B9" pageId="7" pageNumber="178" startId="7.[151,264,1911,1935]" targetBox="[177,1416,203,1862]" targetPageId="7" targetType="figure">
<caption id="65B86640FF944722FF10F8BBFAF5A7B9" ID-DOI="http://doi.org/10.5281/zenodo.14797501" ID-Zenodo-Dep="14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="7" pageNumber="178" startId="7.[151,264,1911,1935]" targetBox="[177,1416,203,1862]" targetPageId="7" targetType="figure">
<paragraph id="317836C8FF944722FF10F8BBFAF5A7B9" blockId="7.[151,1436,1911,1971]" pageId="7" pageNumber="178">
<emphasis id="03B3EADAFF944722FF10F8BBFD67A785" bold="true" box="[151,757,1911,1935]" pageId="7" pageNumber="178">
FIGURES 1316. Male genitalia genus
@ -317,7 +318,7 @@ male, HT., Japan;
<paragraph id="317836C8FF9B472DFF40FF5AFDD5A26B" blockId="8.[151,1437,150,933]" pageId="8" pageNumber="179">
<emphasis id="03B3EADAFF9B472DFF40FF5AFEC3A0BA" bold="true" box="[199,337,150,176]" pageId="8" pageNumber="179">Description</emphasis>
. Adult (
<figureCitation id="A9FC2A4DFF9B472DFE29FF5BFD89A0BB" box="[430,539,151,177]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="8" pageNumber="179">Figs. 7, 8</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFE29FF5BFD89A0BB" box="[430,539,151,177]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="8" pageNumber="179">Figs. 7, 8</figureCitation>
). Forewing length—
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in males.
@ -327,14 +328,14 @@ in males.
. Head, thorax black scales, black tegulae. Palpi short, blackish gray, gray irrorated.Antennae long.Abdomen black, brown, gray irrorated scales. Forewing warm gray with brown tints. Subbasal line black. Antemedial line double, black, straight, costa to dorsum. Postmedial line double, black, incurved below vein 4, continuous to dorsum. Reniform stigmata kidney-shaped, lighter, encircled with black. Orbicular stigmata brown-gray, black outline, connected by black streak. Subterminal area lighter, defined by black dentate marks, angled at veins 7, 6, excurved at middle. Terminal line thin yellowish outline. Cilia mottled. Hindwing cream in basal 2/3, greyish-brown terminal band, cilia greyish-brown, black opposite veins.
<emphasis id="03B3EADAFF9B472DFE0AFE5FFDAAA1A6" bold="true" box="[397,568,402,429]" pageId="8" pageNumber="179">Male genitalia</emphasis>
(
<figureCitation id="A9FC2A4DFF9B472DFDCFFE5FFD0EA1A7" box="[584,668,403,429]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." pageId="8" pageNumber="179">Fig. 16</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFDCFFE5FFD0EA1A7" box="[584,668,403,429]" captionStart="FIGURES 1316" captionStartId="7.[151,264,1911,1935]" captionTargetBox="[177,1416,203,1862]" captionTargetId="figure-21@7.[151,1436,181,1887]" captionTargetPageId="7" captionText="FIGURES 1316. Male genitalia genus Anacronicta. 13. A. pseudonitida sp. nov., male, PT, Manipur, India; 14. A. nitida, male, HT., Japan; 15. A. manipura sp. nov., male, HT, Manipur, India; 16. A. albicans sp. nov., male, PT, Manipur, India." figureDoi="http://doi.org/10.5281/zenodo.14797501" httpUri="https://zenodo.org/record/14797501/files/figure.png" pageId="8" pageNumber="179">Fig. 16</figureCitation>
). Uncus massive, straight, wide at base, narrower at apex. Tegumen with two short sclerotized penicular extensions, shorter than vinculum. Vinculum V-shaped. Juxta hexagonal, shieldlike. Valva large, lobe-like, constricted distally, narrower at apex. Sacculus robust, thin hairs. Saccular process sclerotized, thin, curved, distal half of valva, extends slightly beyond apex. Aedeagus large, massive, thicker at apex, straight. Carina crescent-shaped, highly sclerotized. Vesica large, ventrally everted, with basal, medial, apical diverticula covered in pin-like cornuti.
</paragraph>
<paragraph id="317836C8FF9B472DFF40FDA7FE1BA28F" blockId="8.[151,1437,150,933]" box="[199,393,619,645]" pageId="8" pageNumber="179">Female unknown</paragraph>
<paragraph id="317836C8FF9B472DFF40FD42FBA4A333" blockId="8.[151,1437,150,933]" pageId="8" pageNumber="179">
<emphasis id="03B3EADAFF9B472DFF40FD42FE2AA2A2" bold="true" box="[199,440,654,680]" pageId="8" pageNumber="179">Genetic information</emphasis>
(
<figureCitation id="A9FC2A4DFF9B472DFE41FD43FD85A2A3" box="[454,535,655,681]" captionStart="FIGURE 21" captionStartId="11.[151,250,1435,1459]" captionTargetBox="[156,1425,698,1408]" captionTargetId="figure-236@11.[151,1436,669,1412]" captionTargetPageId="11" captionText="FIGURE 21. Maximum Likelihood tree of the genus Anacronicta based on sequences of mitochondrial cytochrome c oxidase subunit I (COI)." pageId="8" pageNumber="179">Fig. 21</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFE41FD43FD85A2A3" box="[454,535,655,681]" captionStart="FIGURE 21" captionStartId="11.[151,250,1435,1459]" captionTargetBox="[156,1425,698,1408]" captionTargetId="figure-236@11.[151,1436,669,1412]" captionTargetPageId="11" captionText="FIGURE 21. Maximum Likelihood tree of the genus Anacronicta based on sequences of mitochondrial cytochrome c oxidase subunit I (COI)." figureDoi="http://doi.org/10.5281/zenodo.14797507" httpUri="https://zenodo.org/record/14797507/files/figure.png" pageId="8" pageNumber="179">Fig. 21</figureCitation>
).
<specimenCount id="27C1FD41FF9B472DFDADFD43FD7BA2A3" box="[554,745,655,681]" count="7" pageId="8" pageNumber="179" type="generic">Seven specimens</specimenCount>
including

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@ -67,9 +68,9 @@ Hreblay, Katona &amp; Tóth, 2023: 115
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(
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<figureCitation id="A9FC2A4DFF9A472CFF18FE5FFF6FA1A7" box="[159,253,403,429]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="9" pageNumber="180">Figs. 10</figureCitation>
,
<figureCitation id="A9FC2A4DFF9A472CFE8EFE5FFEB4A1A7" box="[265,294,403,429]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." pageId="9" pageNumber="180">18</figureCitation>
<figureCitation id="A9FC2A4DFF9A472CFE8EFE5FFEB4A1A7" box="[265,294,403,429]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="9" pageNumber="180">18</figureCitation>
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<paragraph id="317836C8FF9B472DFF10FA6BFEB2A5CB" blockId="8.[151,1033,1410,1473]" box="[151,288,1447,1473]" pageId="8" pageNumber="179">
(
<figureCitation id="A9FC2A4DFF9B472DFF18FA6BFF7DA5CB" box="[159,239,1447,1473]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." pageId="8" pageNumber="179">Figs. 9</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFF18FA6BFF7DA5CB" box="[159,239,1447,1473]" captionStart="FIGURES 712" captionStartId="6.[151,264,1937,1961]" captionTargetBox="[195,1376,196,1898]" captionTargetId="figure-26@6.[170,1416,178,1915]" captionTargetPageId="6" captionText="FIGURES 712. Adults of genus Anacronicta. 7. A. albicans sp. nov., male, HT, Manipur, India; 8. A. albicans sp. nov., male, PT, Manipur, India; 9. A. albergentea, male, Manipur, India. 10. A. himalaya, male, HT, Nepal (Balint et al. 2023); 11. A. infausta, male, Laos; 12. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797499" httpUri="https://zenodo.org/record/14797499/files/figure.png" pageId="8" pageNumber="179">Figs. 9</figureCitation>
,
<figureCitation id="A9FC2A4DFF9B472DFF7CFA6BFE8AA5CB" box="[251,280,1447,1473]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." pageId="8" pageNumber="179">17</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFF7CFA6BFE8AA5CB" box="[251,280,1447,1473]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="8" pageNumber="179">17</figureCitation>
)
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@ -186,14 +187,14 @@ The species resembles
<emphasis id="03B3EADAFF9B472DFDD6F8C8FD21A714" box="[593,691,1796,1822]" italics="true" pageId="8" pageNumber="179">A. nitida</emphasis>
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(
<figureCitation id="A9FC2A4DFF9B472DFD42F8C8FC9BA715" box="[709,777,1796,1823]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="8" pageNumber="179">Fig. 5</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFD42F8C8FC9BA715" box="[709,777,1796,1823]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="8" pageNumber="179">Fig. 5</figureCitation>
) and
<taxonomicName id="F6C74D4BFF9B472DFCCAF8C8FB22A714" authority="Matsumura, 1931" authorityName="Matsumura" authorityYear="1931" box="[845,1200,1796,1823]" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="horishana">
<emphasis id="03B3EADAFF9B472DFCCAF8C8FC72A714" box="[845,992,1796,1822]" italics="true" pageId="8" pageNumber="179">A. horishana</emphasis>
Matsumura, 1931
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(
<figureCitation id="A9FC2A4DFF9B472DFB45F8C8FA95A714" box="[1218,1287,1796,1822]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." pageId="8" pageNumber="179">Fig. 6</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFB45F8C8FA95A714" box="[1218,1287,1796,1822]" captionStart="FIGURES 16" captionStartId="4.[151,264,1922,1946]" captionTargetBox="[189,1401,195,1880]" captionTargetId="figure-26@4.[151,1436,178,1901]" captionTargetPageId="4" captionText="FIGURES 16. Adults of genus Anacronicta. 1. A. pseudonitida sp. nov., male, HT, Manipur, India; 2. A. pseudonitida sp. nov., male, PT, Myanmar; 3. A. pseudonitida sp. nov., male, PT, Manipur, India; 4. A. manipura sp. nov., male, HT, Manipur, India; 5. A. nitida, male, HT., Japan; 6. A. horishana, male, Types, Taiwan." figureDoi="http://doi.org/10.5281/zenodo.14797495" httpUri="https://zenodo.org/record/14797495/files/figure.png" pageId="8" pageNumber="179">Fig. 6</figureCitation>
) in forewing pattern, dark shadow near the reniform, and the dark brown streak from the lower reniform to the outer margin, as well as the light area between the reniform, dark streak, postmedian fascia, and costal margin.
<taxonomicName id="F6C74D4BFF9B472DFB2EF880FADFA76C" authorityName="Speidel &amp; Kononenko" authorityYear="1998" box="[1193,1357,1868,1894]" class="Insecta" family="Noctuidae" genus="Anacronicta" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="albargentea">
<emphasis id="03B3EADAFF9B472DFB2EF880FADFA76C" box="[1193,1357,1868,1894]" italics="true" pageId="8" pageNumber="179">A. albargentea</emphasis>
@ -203,7 +204,7 @@ differs by its whitish forewing ground color, more contrasting dark wing pattern
<emphasis id="03B3EADAFF9B472DFEE4F858FD9AA7A4" box="[355,520,1940,1966]" italics="true" pageId="8" pageNumber="179">A. albargentea</emphasis>
</taxonomicName>
(
<figureCitation id="A9FC2A4DFF9B472DFD9EF858FDF8A7A4" box="[537,618,1940,1966]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." pageId="8" pageNumber="179">Fig. 17</figureCitation>
<figureCitation id="A9FC2A4DFF9B472DFD9EF858FDF8A7A4" box="[537,618,1940,1966]" captionStart="FIGURES 1720" captionStartId="10.[151,264,1865,1889]" captionTargetBox="[172,1408,202,1827]" captionTargetId="figure-26@10.[151,1436,181,1841]" captionTargetPageId="10" captionText="FIGURES 1720. Adults of genus Anacronicta. 17. A. albergentea, male, Manipur, India; 18. A. himalaya, male, HT, Nepal (Balint et al. 2023); 19. A. infausta, male, Laos; 20. A. obscura, male, Laos." figureDoi="http://doi.org/10.5281/zenodo.14797503" httpUri="https://zenodo.org/record/14797503/files/figure.png" pageId="8" pageNumber="179">Fig. 17</figureCitation>
) has a more massive structure, longer sclerotized pointed peniculi of the tegumen, and a differently shaped and armed vesica.
</paragraph>
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