216 lines
26 KiB
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216 lines
26 KiB
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<document id="8BF6A582F1BC847227B0F9E16CB3083A" ID-ISSN="0024-4082" ID-ZooBank="7A6A530-3EBA-4AF2-94E2-E9B8A1D07459" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738713251128" checkinUser="plazi" docAuthor="Roggero, Angela, Dierkens, Michael, Barbero, Enrico & Palestrini, Claudia" docDate="2017" docId="03DC6B6BFFF1FFD6FC18FF03FED2FB81" docLanguage="en" docName="zoj.12498.pdf" docOrigin="Zoological Journal of the Linnean Society 180" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Morettius Roggero, Dierkens, Barbero & Palestrini, 2017, GEN. NOV." docType="treatment" docVersion="1" lastPageNumber="315" masterDocId="FFE51313FFE1FFC7FFA1FFD5FF82FFF0" masterDocTitle="Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)" masterLastPageNumber="320" masterPageNumber="298" pageNumber="314" updateTime="1738713343532" updateUser="GgImagineBatch">
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<mods:title id="1081C5D68D09B153596B7DD572C2214A">Combined phylogenetic analysis of two new Afrotropical genera of Onthophagini (Coleoptera, Scarabaeidae)</mods:title>
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<mods:namePart id="38116AC5FFEDBC3C2FA8BCE8321C1CB6">Roggero, Angela</mods:namePart>
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<mods:namePart id="932FA7FFAFD8910B0D2E06CE8CCC4EC3">Dierkens, Michael</mods:namePart>
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<mods:namePart id="FA5A11364F467559F6FAE4575B2AF39D">Barbero, Enrico</mods:namePart>
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<mods:namePart id="A35626DDFBB3DFC962DFD2967ED079C0">Palestrini, Claudia</mods:namePart>
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<mods:title id="3B54360D9CD61FB88A4BFEDFBB76B2CA">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="B7215CEB3A8D5474377FEDAC999D1B82">
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<mods:date id="DFD6F691DED98BCEFEBA9997AFD8CB9C">2017</mods:date>
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<mods:number id="5E1ED16368DBC5F1EA152BC804CAD54F">180</mods:number>
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<subSubSection id="C36F89F6FFF1FFD7FC18FF03FA9DFF1D" box="[953,1311,210,237]" pageId="16" pageNumber="314" type="nomenclature">
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<paragraph id="8BCADA7DFFF1FFD7FC18FF03FA9DFF1D" blockId="16.[953,1311,210,237]" box="[953,1311,210,237]" pageId="16" pageNumber="314">
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<heading id="D0826D11FFF1FFD7FC18FF03FA9DFF1D" box="[953,1311,210,237]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="314" reason="2">
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<smallCapsWord id="8D2C4CA1FFF1FFD7FC18FF03FB8AFF1B" baselines="230,231" box="[953,1032,214,235]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Genus" pageId="16" pageNumber="314">GENUS</smallCapsWord>
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<taxonomicName id="4C75A1FEFFF1FFD7FBB3FF01FB1EFF1A" ID-CoL="5V6K" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[1042,1180,212,236]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<smallCapsWord id="8D2C4CA1FFF1FFD7FBB3FF01FB1EFF1A" baselines="230,230" box="[1042,1180,212,236]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Morettius" pageId="16" pageNumber="314">MORETTIUS</smallCapsWord>
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</taxonomicName>
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<emphasis id="B901066FFFF1FFD7FB04FF0CFA9DFF1D" bold="true" box="[1189,1311,210,237]" pageId="16" pageNumber="314">
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<taxonomicNameLabel id="A232BB14FFF1FFD7FB04FF0CFA9DFF1D" box="[1189,1311,210,237]" pageId="16" pageNumber="314" rank="genus">
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<smallCapsWord id="8D2C4CA1FFF1FFD7FB04FF0CFB57FF1A" baselines="230" box="[1189,1237,217,234]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="gen" pageId="16" pageNumber="314">GEN</smallCapsWord>
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.
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<smallCapsWord id="8D2C4CA1FFF1FFD7FB46FF0CFA95FF1A" baselines="230" box="[1255,1303,217,234]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="16" pageNumber="314">NOV</smallCapsWord>
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.
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</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36F89F6FFF1FFD6FC99FECEFED2FB81" lastPageId="17" lastPageNumber="315" pageId="16" pageNumber="314" type="materials_examined">
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<paragraph id="8BCADA7DFFF1FFD7FC99FECEFA08FEC0" blockId="16.[824,1418,283,305]" box="[824,1418,283,305]" pageId="16" pageNumber="314">
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<emphasis id="B901066FFFF1FFD7FC99FECEFC49FEC0" box="[824,971,283,304]" italics="true" pageId="16" pageNumber="314">
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<typeStatus id="54CE64DFFFF1FFD7FC99FECEFCECFEC0" box="[824,878,283,304]" pageId="16" pageNumber="314">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C75A1FEFFF1FFD7FC7BFECEFA08FEC0" authority="d'Orbigny, 1908" authorityName="d'Orbigny" authorityYear="1908" box="[986,1418,283,305]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="pallens">
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<emphasis id="B901066FFFF1FFD7FC7BFECEFB4FFEC0" box="[986,1229,283,304]" italics="true" pageId="16" pageNumber="314">Onthophagus pallens</emphasis>
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<bibRefCitation id="EFE4A78CFFF1FFD7FB74FECEFA08FEC0" author="d'Orbigny H" box="[1237,1418,283,305]" pageId="16" pageNumber="314" pagination="65 - 208" refId="ref13608" refString="d'Orbigny H. 1908. Descriptions d'especes nouvelles d'Onthophagides africains et notes synonimiques. Annales de la Societe Entomologique de France 77: 65-208." type="journal article" year="1908">d’Orbigny, 1908</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BCADA7DFFF1FFD7FC99FE8DFC75FE7C" blockId="16.[824,1439,344,396]" pageId="16" pageNumber="314">
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<emphasis id="B901066FFFF1FFD7FC99FE8DFC7DFE9D" box="[824,1023,344,365]" italics="true" pageId="16" pageNumber="314">Included species:</emphasis>
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<taxonomicName id="4C75A1FEFFF1FFD7FBACFE8DFAE0FE9D" authority="(d'Orbigny, 1908)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[1037,1378,344,366]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="pallens">
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<emphasis id="B901066FFFF1FFD7FBACFE8DFB05FE9D" box="[1037,1159,344,365]" italics="true" pageId="16" pageNumber="314">M. pallens</emphasis>
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(
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<bibRefCitation id="EFE4A78CFFF1FFD7FB3DFE8DFAD8FE9D" author="d'Orbigny H" box="[1180,1370,344,366]" pageId="16" pageNumber="314" pagination="65 - 208" refId="ref13608" refString="d'Orbigny H. 1908. Descriptions d'especes nouvelles d'Onthophagides africains et notes synonimiques. Annales de la Societe Entomologique de France 77: 65-208." type="journal article" year="1908">d’Orbigny, 1908</bibRefCitation>
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)
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</taxonomicName>
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, and
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<taxonomicName id="4C75A1FEFFF1FFD7FC99FEA2FC14FE7C" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[824,918,375,396]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="utete" status="sp. nov.">
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<emphasis id="B901066FFFF1FFD7FC99FEA2FC14FE7C" box="[824,918,375,396]" italics="true" pageId="16" pageNumber="314">M. utete</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A232BB14FFF1FFD7FC3FFEA2FC75FE7C" box="[926,1015,375,396]" pageId="16" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BCADA7DFFF1FFD7FC99FE7AFB28FD49" blockId="16.[824,1439,430,697]" pageId="16" pageNumber="314">
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<emphasis id="B901066FFFF1FFD7FC99FE7AFC32FE34" box="[824,944,431,452]" italics="true" pageId="16" pageNumber="314">Diagnosis:</emphasis>
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Species of the genus
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<taxonomicName id="4C75A1FEFFF1FFD7FB17FE7AFAA1FE34" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[1206,1315,431,452]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus">
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<emphasis id="B901066FFFF1FFD7FB17FE7AFAA1FE34" box="[1206,1315,431,452]" italics="true" pageId="16" pageNumber="314">Morettius</emphasis>
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</taxonomicName>
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(
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<figureCitation id="134EC6F8FFF1FFD7FA97FE7BFA0AFE33" box="[1334,1416,430,452]" captionStart="Figure 3" captionStartId="5.[148,227,1580,1599]" captionTargetBox="[161,1409,195,1544]" captionTargetId="figure-178@5.[161,1409,195,1544]" captionTargetPageId="5" captionText="Figure 3. Scatterplots of the RW 1 and 2 of head, pronotum, right elytron and mentum (semilandmarks method). Only the specimens employed to build the matrix are shown here. The deformation grids corresponding to the minimum and maximum values of the axes are shown for each anatomical trait. See text for the codes." pageId="16" pageNumber="314">Fig. S3</figureCitation>
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<emphasis id="B901066FFFF1FFD7FA32FE64FA1DFE33" box="[1427,1439,433,451]" italics="true" pageId="16" pageNumber="314">–</emphasis>
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Appendix 4, Supporting Information) are characterized by the mostly rounded and only slightly notched anterior margin of the clypeus, and the pronotum covered by granules, sometimes mixed to points. The pygidium is always smooth, with some rade points. The species show a moderate sexual dimorphism in the fore tibiae, and the pygidium is larger in males than in females.
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</paragraph>
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<paragraph id="8BCADA7DFFF1FFD7FC99FD08FAAFFB6F" blockId="16.[824,1440,733,1183]" pageId="16" pageNumber="314">
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<emphasis id="B901066FFFF1FFD7FC99FD08FC47FD02" box="[824,965,733,754]" italics="true" pageId="16" pageNumber="314">Epipharynx:</emphasis>
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The epipharynx (
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<figureCitation id="134EC6F8FFF1FFD7FB01FD08FB66FD02" box="[1184,1252,733,755]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 2</figureCitation>
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) fore margin is arched, with a largely V-shaped notch in the middle. The corypha is reduced, the chaetopariae are arched with short, thick and almost equal-length setae. The pubescence of the haptomerum is dense. The proplegmatium is subequal along the whole length, with the posterior triangular sclerotization reaching at least as much or more than half the length of the anterior epitorma, which is rectilinear, well sclerotized and thin. The base of the triangular sclerotization reaches the small, thick and upwardturned apophyses. Laeotorma and dexiotorma are symmetrical, short and stout. Pternotormae are well sclerotized, and the crepis is short but evident, with a sharp apex. The plegmatic area is visible.
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</paragraph>
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<paragraph id="8BCADA7DFFF1FFD7FC99FB14FBE0FA3C" blockId="16.[824,1439,1217,1484]" pageId="16" pageNumber="314">
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<emphasis id="B901066FFFF1FFD7FC99FB14FC7CFB26" box="[824,1022,1217,1238]" italics="true" pageId="16" pageNumber="314">Male genitalia:</emphasis>
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Only the male genitalia of
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<taxonomicName id="4C75A1FEFFF1FFD7FC99FB34FC30FB05" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[824,946,1248,1270]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="pallens">
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<emphasis id="B901066FFFF1FFD7FC99FB34FC30FB05" box="[824,946,1248,1270]" italics="true" pageId="16" pageNumber="314">M. pallens</emphasis>
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</taxonomicName>
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could be examined. The phallobase (
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<figureCitation id="134EC6F8FFF1FFD7FCE0FB2AFC06FAE4" box="[833,900,1279,1301]" captionStart="Figure 7" captionStartId="8.[162,241,917,936]" captionTargetBox="[391,1215,201,883]" captionTargetId="figure-27@8.[388,1212,198,884]" captionTargetPageId="8" captionText="Figure 7. Aedeagus of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA); see text for the codes. Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 7</figureCitation>
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) is short, only slightly arched, slender and of equal size along the whole length. The parameres are symmetrical, and squared, with a small tip at the apex, and a small, rounded protrusion ventrally. The internal sac (
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<figureCitation id="134EC6F8FFF1FFD7FB8EFAACFBF0FA7E" box="[1071,1138,1401,1423]" captionStart="Figure 8" captionStartId="8.[162,241,1850,1869]" captionTargetBox="[391,1215,1064,1819]" captionTargetId="figure-59@8.[388,1212,1062,1817]" captionTargetPageId="8" captionText="Figure 8. Primary lamella of the species of the genera Hamonthophagus (AC, BI, DE, FA and LA) and Morettius (PA). Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 8</figureCitation>
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) is membranous, with various well-sclerotized parts differing greatly from
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<taxonomicName id="4C75A1FEFFF1FFD7FC99FA63FBE0FA3C" box="[824,1122,1462,1484]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B901066FFFF1FFD7FC99FA63FB80FA3B" box="[824,1026,1462,1483]" italics="true" pageId="16" pageNumber="314">Hamonthophagus</emphasis>
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species.
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</taxonomicName>
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</paragraph>
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<paragraph id="8BCADA7DFFF1FFD7FC99FA3BFAE2F907" blockId="16.[824,1439,1518,1784]" pageId="16" pageNumber="314">
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<emphasis id="B901066FFFF1FFD7FC99FA3BFB89F9F3" box="[824,1035,1518,1539]" italics="true" pageId="16" pageNumber="314">Female genitalia:</emphasis>
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The female genitalia are very peculiar (
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<figureCitation id="134EC6F8FFF1FFD7FC0AF9D9FC7EF9D1" box="[939,1020,1548,1570]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="16" pageNumber="314">Fig. 11</figureCitation>
|
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|
|
), as the vagina of both species is entirely membranous, and no sclerotization is present at all. Furthermore, the infundibular tube is lowered as in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF1FFD7FBACF9BDFB55F98D" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[1037,1239,1640,1661]" class="Insecta" family="Scarabaeidae" genus="Hamonthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="314" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B901066FFFF1FFD7FBACF9BDFB55F98D" box="[1037,1239,1640,1661]" italics="true" pageId="16" pageNumber="314">Hamonthophagus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but here an expanded portion is identifiable in the central part of the vagina, which is differently shaped in the two species. The receptaculum seminis is sickle-shaped, namely slim, arched and apically sharp (
|
|||
|
|
<figureCitation id="134EC6F8FFF1FFD7FAAFF937FAD0F907" box="[1294,1362,1762,1784]" captionStart="Figure 9" captionStartId="9.[148,227,955,974]" captionTargetBox="[376,1200,195,918]" captionTargetId="figure-117@9.[376,1200,195,918]" captionTargetPageId="9" captionText="Figure 9. Above, the receptaculum seminis; the features that characterize the two models are clearly represented by H. bituberculatus (BI) and M. pallens (PA). Scale bar = 0.2 mm. Below, a generic example of the vagina sclerotization (H. bituberculatus), after being cleared from membranes and cut off. The various parts were numbered (see text for further details)." pageId="16" pageNumber="314">Fig. 9</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF1FFD7FC99F8CFFB37F89D" blockId="16.[824,1439,1818,1901]" pageId="16" pageNumber="314">
|
|||
|
|
<emphasis id="B901066FFFF1FFD7FC99F8CFFB98F8DF" box="[824,1050,1818,1839]" italics="true" pageId="16" pageNumber="314">Specific diagnosis:</emphasis>
|
|||
|
|
These species can be easily distinguished on the basis of external morphology, epipharynx and female genitalia.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF0CFF12FE32FE21" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
|
|||
|
|
The pronotum in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FEDFFF12FE75FF2C" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[382,503,199,220]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FEDFFF12FE75FF2C" box="[382,503,199,220]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is covered by distinct rasping points mixed with smaller, yellow granules; the granules of the rasping points instead are large, darker than the background surface, and carry long, thick and light yellow setae. The pronotum of
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FF35FEB5FF70FE85" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[148,242,352,373]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FF35FEB5FF70FE85" box="[148,242,352,373]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is covered by only a few rasping setigerous points with thin, yellow setae and many small granules, which are very thick, evenly coloured as the base and without points.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF0CFE0FFF7BFDDC" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
|
|||
|
|
Elytral striae of
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FE20FE0FFE78FE1F" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[385,506,474,495]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FE20FE0FFE78FE1F" box="[385,506,474,495]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are as large as the points, being instead larger than the points in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FF35FDC2FF70FDDC" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[148,242,535,556]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FF35FDC2FF70FDDC" box="[148,242,535,556]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF0CFDE0FDFEFD56" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
|
|||
|
|
The smooth pygidium carries in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FDE4FDE3FD3CFDBA" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[581,702,565,587]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FDE4FDE3FD3CFDBA" box="[581,702,565,587]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
few, small, rade and deep setigerous points that are not granulated, and in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FE2DFDA6FE6BFD78" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[396,489,627,648]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FE2DFDA6FE6BFD78" box="[396,489,627,648]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
only some large and superficial vanishing points without setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF0CFD7AFDA7FCB0" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
|
|||
|
|
The epipharynx (see
|
|||
|
|
<figureCitation id="134EC6F8FFF0FFD6FE69FD7AFD89FD35" box="[456,523,687,709]" captionStart="Figure 2" captionStartId="4.[162,241,1141,1160]" captionTargetBox="[391,1215,203,1113]" captionTargetId="figure-431@4.[391,1215,203,1113]" captionTargetPageId="4" captionText="Figure 2. Epipharynx of the species of the genera Hamonthophagus (AC = acutus, BI = bitubeculatus, DE = depressus, FA = fallax, LA = laceratus) and Morettius (PA = pallens, UT = utete). Scale bar = 0.5 mm." pageId="17" pageNumber="315">Fig. 2</figureCitation>
|
|||
|
|
for a comparison among species) has the characteristic shape of
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FD11FD1AFE9CFCF3" pageId="17" pageNumber="315">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FD11FD1AFF3DFCF2" italics="true" pageId="17" pageNumber="315">Morettius</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but can be distinguished from
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FD3CFD38FD78FCF2" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[669,762,749,770]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FD3CFD38FD78FCF2" box="[669,762,749,770]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the more developed apotormae, and the more slender laeotorma and dexiotorma.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF0CFC9CFD79FC29" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
|
|||
|
|
In both species, the vagina is wholly desclerotized, but carries two globose symmetrical expansions that encircle the desclerotized and lowered infundibulum in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FF35FC70FE8EFC49" baseAuthorityName="d'Orbigny" baseAuthorityYear="1908" box="[148,268,932,954]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="pallens">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FF35FC70FE8EFC49" box="[148,268,932,954]" italics="true" pageId="17" pageNumber="315">M. pallens</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, while in
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FE29FC70FE67FC4A" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[392,485,933,954]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FE29FC70FE67FC4A" box="[392,485,933,954]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
there is a single, large expansion (see
|
|||
|
|
<figureCitation id="134EC6F8FFF0FFD6FE9AFC16FE09FC28" box="[315,395,963,985]" captionStart="Figure 11" captionStartId="10.[162,241,707,726]" captionTargetBox="[391,1215,198,674]" captionTargetId="figure-136@10.[391,1215,198,674]" captionTargetPageId="10" captionText="Figure 11. Vagina and receptaculum seminis of the species of the genus Morettius (PA and UT). Scale bar = 0.5 mm." pageId="17" pageNumber="315">Fig. 11</figureCitation>
|
|||
|
|
for a comparison among species).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF0CFC34FE68FBE6" blockId="17.[148,763,199,1046]" pageId="17" pageNumber="315">
|
|||
|
|
As the male of
|
|||
|
|
<taxonomicName id="4C75A1FEFFF0FFD6FEFAFC37FE3AFC07" authorityName="Roggero & Dierkens & Barbero & Palestrini" authorityYear="2017" box="[347,440,994,1015]" class="Insecta" family="Scarabaeidae" genus="Morettius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="315" phylum="Arthropoda" rank="species" species="utete">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FEFAFC37FE3AFC07" box="[347,440,994,1015]" italics="true" pageId="17" pageNumber="315">M. utete</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is unknown, no comparison can be made between species.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCADA7DFFF0FFD6FF35FBE8FED2FB81" blockId="17.[148,763,1085,1137]" pageId="17" pageNumber="315">
|
|||
|
|
<emphasis id="B901066FFFF0FFD6FF35FBE8FEDCFBA2" box="[148,350,1085,1106]" italics="true" pageId="17" pageNumber="315">General remarks:</emphasis>
|
|||
|
|
No preimaginal stages have been described so far.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|