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<document id="07D8E8A1EEF5B5CB31269799CB7793BA" ID-CLB-Dataset="182443" ID-DOI="10.11646/phytotaxa.572.1.6" ID-GBIF-Dataset="0319c787-14ca-4528-8794-dee4f56c781a" ID-ISSN="1179-3163" ID-Zenodo-Dep="7305694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1667943407477" checkinUser="plazi" docAuthor="HOANG, THANH SON &amp; LIN, CHE-WEI" docDate="2022" docId="625287B9FF94A11FDAC569FD29EBC95B" docLanguage="en" docName="phytotaxa.572.1.6.pdf" docOrigin="Phytotaxa 572 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.572.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Begonia minhaniae T. S. Hoang &amp; C. W. Lin. A, B. Habit 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="88" masterDocId="9E6BFFC1FF94A11EDA4D6E6B2D0FCF26" masterDocTitle="Two new species, Begonia minhaniae and B. dieuanhiae (sect. Platycentrum, Begoniaceae), from northwestern Vietnam" masterLastPageNumber="96" masterPageNumber="87" pageNumber="87" updateTime="1699410986681" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="763DCD1107FA381F7B829BAE8FB4CBB8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F7A63A6792E3B9AD47B7C77B19DF150">
<mods:title id="3AF7B145DC5E05F78D6E57C662E7390E">Two new species, Begonia minhaniae and B. dieuanhiae (sect. Platycentrum, Begoniaceae), from northwestern Vietnam</mods:title>
</mods:titleInfo>
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<mods:namePart id="1C0411249FFECC48AE49C88F825B1776">HOANG, THANH SON</mods:namePart>
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<mods:namePart id="00D3A8C0E3708D627B3EC87C48B34AEA">LIN, CHE-WEI</mods:namePart>
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<mods:title id="F1A3E222FF1BAEB18077085393750CA8">Phytotaxa</mods:title>
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<mods:date id="D21F7DA8B36B633422007FCBB01974A9">2022</mods:date>
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<mods:number id="5DC9B3550319AC1230507888201E22F3">2022-11-08</mods:number>
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<treatment id="625287B9FF94A11FDAC569FD29EBC95B" ID-DOI="http://doi.org/10.5281/zenodo.7305704" ID-GBIF-Taxon="204325293" ID-Zenodo-Dep="7305704" LSID="urn:lsid:plazi:treatment:625287B9FF94A11FDAC569FD29EBC95B" httpUri="http://treatment.plazi.org/id/625287B9FF94A11FDAC569FD29EBC95B" lastPageId="1" lastPageNumber="88" pageId="0" pageNumber="87">
<subSubSection id="A2E16524FF94A11EDAC569FD2915C897" box="[136,1050,1942,1969]" pageId="0" pageNumber="87" type="nomenclature">
<paragraph id="EA4436AFFF94A11EDAC569FD2915C897" blockId="0.[136,1050,1942,1969]" box="[136,1050,1942,1969]" pageId="0" pageNumber="87">
<heading id="B10C81C3FF94A11EDAC569FD2915C897" box="[136,1050,1942,1969]" fontSize="11" level="2" pageId="0" pageNumber="87" reason="3">
<taxonomicName id="2DFB4D2CFF94A11EDAC569FD2F7FC896" ID-CoL="637c5a06-7207-4304-8f2c-52da372ec7ec" authority="T.S.Hoang &amp; C.W.Lin" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[136,624,1942,1969]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="0" pageNumber="87" phylum="Tracheophyta" rank="species" species="minhaniae" status="sp. nov.">
<emphasis id="D88FEABDFF94A11EDAC569FD2C63C897" bold="true" box="[136,364,1942,1969]" italics="true" pageId="0" pageNumber="87">Begonia minhaniae</emphasis>
T.S.Hoang &amp; C.W.Lin
</taxonomicName>
,
<emphasis id="D88FEABDFF94A11ED83669FC2FDFC896" box="[635,720,1943,1968]" italics="true" pageId="0" pageNumber="87">
<taxonomicNameLabel id="C3BC57C6FF94A11ED83669FC2FDFC896" box="[635,720,1943,1968]" pageId="0" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
§
<emphasis id="D88FEABDFF94A11ED8A169FD2E8CC896" box="[748,899,1942,1968]" italics="true" pageId="0" pageNumber="87">Platycentrum</emphasis>
(
<figureCitation id="72C02A2AFF94A11ED9DF69FD2ED0C897" box="[914,991,1942,1969]" captionStart="FIGURE 2" captionStartId="3.[136,234,1954,1976]" captionTargetBox="[210,1365,264,1886]" captionTargetId="figure-17@3.[189,1398,190,1930]" captionTargetPageId="3" captionText="FIGURE 2. Begonia minhaniae T.S.Hoang &amp; C.W.Lin. A. Habit; B, B’. Portion of leaf, upper and lower surfaces; C. Stipule; D, D’. Bracts; E, E’. Staminate flower, face and side views; F, F’, F’’. Stamens, dorsal, ventral and side views; G, G’. Immature pistillate flower, face and side views; H, H’. Style and stigmatic band, dorsal and ventral views; I. Cross section of ovary." figureDoi="http://doi.org/10.5281/zenodo.7305698" httpUri="https://zenodo.org/record/7305698/files/figure.png" pageId="0" pageNumber="87">Figs. 2</figureCitation>
&amp;
<figureCitation id="72C02A2AFF94A11EDE4E69FC291EC897" box="[1027,1041,1943,1969]" captionStart="FIGURE 3" captionStartId="4.[136,234,1935,1957]" captionTargetBox="[151,1436,190,1911]" captionTargetId="figure-21@4.[151,1436,190,1911]" captionTargetPageId="4" captionText="FIGURE 3. Begonia minhaniae T.S.Hoang &amp; C.W.Lin. A, B. Habit and habitat; C. Young leaf, also showing stipules, petiole and peduncle; D, E. Portion of leaf, upper and lower surfaces; F. Habit; G. Inflorescence; H. Immature staminate and pistillate flowers, also showing bracts and bracteoles; I. Immature staminate flowers; J. Cross section of an old capsule." figureDoi="http://doi.org/10.5281/zenodo.7305700" httpUri="https://zenodo.org/record/7305700/files/figure.png" pageId="0" pageNumber="87">3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2E16524FF94A11EDAC569D52E0BC8D1" pageId="0" pageNumber="87" type="materials_examined">
<paragraph id="EA4436AFFF94A11EDAC569D52E0BC8D1" blockId="0.[136,1452,1982,2040]" pageId="0" pageNumber="87">
<emphasis id="D88FEABDFF94A11EDAC569D52DD5C8F5" bold="true" box="[136,218,1982,2003]" pageId="0" pageNumber="87">Type:—</emphasis>
<materialsCitation id="5A933CF2FF94A11EDA9769D52E0FC8D1" ID-GBIF-Occurrence="3962336302" collectingDate="2021-07-22" collectionCode="VAFS, VNMN" country="Vietnam" elevation="512" latitude="21.707777" location="Van Tran District" longLatPrecision="1" longitude="104.34175" pageId="0" pageNumber="87" specimenCount="1" stateProvince="Yen Bai" typeStatus="holotype">
<collectingCountry id="92EC763FFF94A11EDA9769D52C42C8F5" box="[218,333,1982,2003]" name="Vietnam" pageId="0" pageNumber="87">VIETNAM</collectingCountry>
:
<collectingRegion id="283FF84DFF94A11EDB1869D42CF1C8F2" box="[341,510,1982,2004]" country="Vietnam" name="Yen Bai" pageId="0" pageNumber="87">Yen Bai province</collectingRegion>
,
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,
<geoCoordinate id="8FCF5068FF94A11ED8F669D52E3BC8F5" box="[699,820,1982,2003]" degrees="21.707777" direction="north" orientation="latitude" pageId="0" pageNumber="87" precision="1" value="21.707777">21.707777N</geoCoordinate>
,
<geoCoordinate id="8FCF5068FF94A11ED97369D52ECEC8F5" box="[830,961,1982,2003]" degrees="104.341748" direction="east" orientation="longitude" pageId="0" pageNumber="87" precision="1" value="104.34175">104.341748E</geoCoordinate>
,
<quantity id="2D039B4AFF94A11ED98669D52907C8F2" box="[971,1032,1982,2004]" metricMagnitude="2" metricUnit="m" metricValue="5.12" pageId="0" pageNumber="87" unit="m" value="512.0">
<elevation id="61D6D19CFF94A11ED98669D52907C8F2" box="[971,1032,1982,2004]" metricMagnitude="2" metricUnit="m" metricValue="5.12" pageId="0" pageNumber="87" unit="m" value="512.0">512 m</elevation>
</quantity>
elevation,
<date id="9E45106FFF94A11EDE3E69D529FEC8F5" box="[1139,1265,1982,2004]" pageId="0" pageNumber="87" value="2021-07-22">
<collectingDate id="8E01E987FF94A11EDE3E69D529FEC8F5" box="[1139,1265,1982,2004]" pageId="0" pageNumber="87" value="2021-07-22">22 July 2021</collectingDate>
</date>
, flowers and fruits,
<emphasis id="D88FEABDFF94A11EDAF069882CADC8D1" box="[189,418,2018,2040]" italics="true" pageId="0" pageNumber="87">Hoang Thanh Son 8754</emphasis>
(
<typeStatus id="3540880DFF94A11EDBE369892F06C8DE" box="[430,521,2018,2040]" pageId="0" pageNumber="87" type="holotype">Holotype</typeStatus>
<collectionCode id="8CEAAE6AFF94A11ED84269892F43C8D1" box="[527,588,2018,2039]" pageId="0" pageNumber="87">VAFS</collectionCode>
; Isotypes
<collectionCode id="8CEAAE6AFF94A11ED8E669892FF6C8D1" box="[683,761,2018,2039]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="0" pageNumber="87">VNMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A2E16524FF95A11FDAC56EF52E9DCCD2" pageId="1" pageNumber="88" type="description">
<paragraph id="EA4436AFFF95A11FDAC56EF52E9DCCD2" blockId="1.[136,1452,158,1661]" pageId="1" pageNumber="88">
Monoecious rhizomatous herb.
<emphasis id="D88FEABDFF95A11FDBBA6EF52F6ECF9E" bold="true" box="[503,609,158,184]" pageId="1" pageNumber="88">Rhizome</emphasis>
stout, reddish-brown,
<quantity id="2D039B4AFF95A11FD9276EF52ECECF9F" box="[874,961,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="88" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
or longer,
<quantity id="2D039B4AFF95A11FDE056EF529A4CF9F" box="[1096,1195,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="88" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5−7 mm</quantity>
thick, internodes
<quantity id="2D039B4AFF95A11FDFCD6EF52DBBCFFB" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="1" pageNumber="88" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2−7 mm</quantity>
long, sparsely pale brown tomentose.
<emphasis id="D88FEABDFF95A11FD8256EA92FC8CFFA" bold="true" box="[616,711,194,220]" pageId="1" pageNumber="88">Stipules</emphasis>
often caducous, yellow tinged crimson, triangular to very widely ovate-triangular, 4–6 ×
<quantity id="2D039B4AFF95A11FDBD86E8D2CF9CE27" box="[405,502,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="1" pageNumber="88" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
, herbaceous, keeled, glabrous or abaxially sparsely tomentose, margin ciliate, apex obtuse to rounded and mucronate.
<emphasis id="D88FEABDFF95A11FD8476F612F53CE02" bold="true" box="[522,604,266,292]" pageId="1" pageNumber="88">Leaves</emphasis>
alternate, petiole terete, brownish-red to crimson,
<quantity id="2D039B4AFF95A11FDEDD6F6129FDCE03" box="[1168,1266,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="1" pageNumber="88" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">4−10 cm</quantity>
long,
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diameter, densely subappressed yellowish-brown tomentose; leaf blade asymmetric, oblique, widely ovate, 8.5−12.5 ×
<quantity id="2D039B4AFF95A11FDAC56F392DFCCE4B" box="[136,243,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="9.5" metricValueMin="6.0" pageId="1" pageNumber="88" unit="cm" value="7.75" valueMax="9.5" valueMin="6.0">6–9.5 cm</quantity>
, broad side
<quantity id="2D039B4AFF95A11FDB346F392CF9CE4B" box="[377,502,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.5" metricValueMin="3.6" pageId="1" pageNumber="88" unit="cm" value="4.55" valueMax="5.5" valueMin="3.6">3.6–5.5 cm</quantity>
wide, basal lobes cordate,
<quantity id="2D039B4AFF95A11FD9696F392E83CE4B" box="[804,908,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="1" pageNumber="88" unit="cm" value="2.3" valueMax="3.0" valueMin="1.6">1.6–3 cm</quantity>
long, apex acute to acuminate, margin subentire to crenulate with a line of red puberulous hairs; leaf chartaceous, succulent, adaxially lime green, glabrous; abaxially pale green, glabrous except on veins; venation basally
<emphasis id="D88FEABDFF95A11FD9406FF02E28CE92" box="[781,807,411,436]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 7 palmate, veins green adaxially, yellowish-green and sparsely red puberulous abaxially; midrib distinct,
<emphasis id="D88FEABDFF95A11FD8896FD42FD1CEFE" box="[708,734,447,472]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 2 secondary veins on each side, other primary veins branching dichotomously or nearly so, tertiary veins reticulate.
<emphasis id="D88FEABDFF95A11FD8AF6F892E84CEDA" bold="true" box="[738,907,482,508]" pageId="1" pageNumber="88">Inflorescences</emphasis>
axillary, bisexual, cymose arising directly from rhizome,
<emphasis id="D88FEABDFF95A11FDAB86C6C2C00CD06" box="[245,271,519,544]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 2 orders of branching; peduncle reddish-green to crimson,
<emphasis id="D88FEABDFF95A11FD9856C6C2EEDCD06" box="[968,994,519,544]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FD9BE6C6D2934CD07" box="[1011,1083,518,545]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="88" unit="cm" value="10.0">10 cm</quantity>
long, sparsely yellowish-brown tomentose.
<emphasis id="D88FEABDFF95A11FDB456C412C59CD62" bold="true" box="[264,342,554,580]" pageId="1" pageNumber="88">Bracts</emphasis>
caducous, yellowish-red to pale brown, at basal node of inflorescence ovate,
<emphasis id="D88FEABDFF95A11FDEE46C4029CCCD62" box="[1193,1219,555,580]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 4 ×
<quantity id="2D039B4AFF95A11FDEB46C412834CD63" box="[1273,1339,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="88" unit="mm" value="2.0">2 mm</quantity>
, abaxially sparsely yellowish-brown tomentose or glabrous, margin ciliate; bracts of upper inflorescence similar but smaller.
<emphasis id="D88FEABDFF95A11FDAC56C192C6FCDAA" bold="true" box="[136,352,626,652]" pageId="1" pageNumber="88">Staminate flower:</emphasis>
pedicel yellowish-red,
<emphasis id="D88FEABDFF95A11FD8236C182F87CDAA" box="[622,648,627,652]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FD8D46C192FDCCDAB" box="[665,723,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="88" unit="cm" value="2.0">2 cm</quantity>
long, very sparsely yellowish-brown tomentose, tepals 4, pink, outer 2 orbicular to very widely ovate,
<emphasis id="D88FEABDFF95A11FD8716CFC2F59CD96" box="[572,598,663,688]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FD8286CFD2F94CD97" box="[613,667,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="88" unit="cm" value="1.0">1 cm</quantity>
across, adaxially very sparsely puberulous or glabrous, margin entire, apex rounded, inner 2 elliptic,
<emphasis id="D88FEABDFF95A11FDBAC6CD02CF4CDF2" box="[481,507,699,724]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 10 ×
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, glabrous, apex rounded; androecium zygomorphic,
<emphasis id="D88FEABDFF95A11FDE9A6CD029FECDF2" box="[1239,1265,699,724]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDF4D6CD12858CDF3" box="[1280,1367,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="88" unit="mm" value="3.5">3.5 mm</quantity>
across; stamens golden yellow,
<emphasis id="D88FEABDFF95A11FDBD56CB42CBDCDDE" box="[408,434,735,760]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 15; filaments
<emphasis id="D88FEABDFF95A11FD8136CB42F8FCDDE" box="[606,640,735,760]" italics="true" pageId="1" pageNumber="88">ca.</emphasis>
<quantity id="2D039B4AFF95A11FD8C56CB52FC5CDDF" box="[648,714,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="88" unit="mm" value="2.0">2 mm</quantity>
long, fused at base; anthers obovate,
<emphasis id="D88FEABDFF95A11FDE3A6CB4299ECDDE" box="[1143,1169,735,760]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDEED6CB529F7CDDF" box="[1184,1272,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="88" unit="mm" value="1.5">1.5 mm</quantity>
long, 2-locular, apex slightly retuse.
<emphasis id="D88FEABDFF95A11FDB3E6D692FC6CC3A" bold="true" box="[371,713,770,796]" pageId="1" pageNumber="88">Pistillate flower (immature):</emphasis>
pedicel
<emphasis id="D88FEABDFF95A11FD9676D682E4BCC3A" box="[810,836,771,796]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FD91E6D692E9ACC3B" box="[851,917,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="88" unit="mm" value="5.0">5 mm</quantity>
long with a pairs alternate bracteoles
<emphasis id="D88FEABDFF95A11FDF0B6D68286FCC3A" box="[1350,1376,771,796]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 2 ×
<quantity id="2D039B4AFF95A11FDFD06D692DB9CC67" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="88" unit="mm" value="1.0">1 mm</quantity>
, ciliate; sparsely yellowish-brown tomentose, tepals 3, pink, outer 2 orbicular,
<emphasis id="D88FEABDFF95A11FDE506D4C2938CC66" box="[1053,1079,807,832]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDE096D4D298BCC67" box="[1092,1156,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="88" unit="mm" value="8.0">8 mm</quantity>
across, subglabrous; inner 1 obovate,
<emphasis id="D88FEABDFF95A11FDB4F6D202C13CC42" box="[258,284,843,868]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 7 ×
<quantity id="2D039B4AFF95A11FDB196D212CA3CC43" box="[340,428,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="88" unit="mm" value="2.5">2.5 mm</quantity>
; ovary trigonous-ellipsoid,
<emphasis id="D88FEABDFF95A11FD8936D202FF7CC42" box="[734,760,843,868]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
. 5 ×
<quantity id="2D039B4AFF95A11FD97D6D212E87CC43" box="[816,904,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="88" unit="mm" value="2.5">2.5 mm</quantity>
; 3-winged, wings unequal,
<emphasis id="D88FEABDFF95A11FDEF46D2029DCCC42" box="[1209,1235,843,868]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDEAD6D21282ECC43" box="[1248,1313,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="88" unit="mm" value="6.0">6 mm</quantity>
long, lateral wings narrower, narrowly triangular, abaxial wing triangular, margin entire, glabrous or sparsely tomentose; ovary 3-locular, placenta bilamellate; styles 3, fused at base, yellow, U-shaped, stigma spirally twisted.
<emphasis id="D88FEABDFF95A11FDE9D6DF9283FCC8A" bold="true" box="[1232,1328,914,940]" pageId="1" pageNumber="88">Capsule</emphasis>
trigonousellipsoid,
<emphasis id="D88FEABDFF95A11FDAB56DDC2C1DCCF6" box="[248,274,951,976]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDB6F6DDD2C6ACCF7" box="[290,357,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="88" unit="mm" value="9.0">9 mm</quantity>
long,
<quantity id="2D039B4AFF95A11FDBE36DDD2CFECCF7" box="[430,497,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="88" unit="mm" value="6.0">6 mm</quantity>
thick (wings excluded), reddish-green when fresh; wings unequal,
<emphasis id="D88FEABDFF95A11FDEBC6DDC2804CCF6" box="[1265,1291,951,976]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDF566DDD2865CCF7" box="[1307,1386,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="88" unit="mm" value="11.0">11 mm</quantity>
long, lateral wings
<emphasis id="D88FEABDFF95A11FDB526DB02C36CCD2" box="[287,313,987,1012]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FDB0A6DB12C87CCD3" box="[327,392,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="88" unit="mm" value="2.0">2 mm</quantity>
wide, abaxial wing triangular,
<emphasis id="D88FEABDFF95A11FD8A96DB02FF1CCD2" box="[740,766,987,1012]" italics="true" pageId="1" pageNumber="88">ca</emphasis>
.
<quantity id="2D039B4AFF95A11FD9416DB12E42CCD3" box="[780,845,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="88" unit="mm" value="5.0">5 mm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="A2E16524FF95A11FDAF06D952807CB1B" pageId="1" pageNumber="88" type="distribution">
<paragraph id="EA4436AFFF95A11FDAF06D952807CB1B" blockId="1.[136,1452,158,1661]" pageId="1" pageNumber="88">
<emphasis id="D88FEABDFF95A11FDAF06D952F02CB3E" bold="true" box="[189,525,1022,1048]" pageId="1" pageNumber="88">Distribution and ecology:—</emphasis>
<taxonomicName id="2DFB4D2CFF95A11FD8436D942FE4CB3E" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[526,747,1022,1048]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="minhaniae">
<emphasis id="D88FEABDFF95A11FD8436D942FE4CB3E" box="[526,747,1022,1048]" italics="true" pageId="1" pageNumber="88">Begonia minhaniae</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="3540880DFF95A11FDE436D942931CB3F" box="[1038,1086,1023,1049]" pageId="1" pageNumber="88">type</typeStatus>
locality in Van Tran District. It grows on moist cliff faces in shady lower primary evergreen broad-leaved forests, elevation
<emphasis id="D88FEABDFF95A11FDEDC6A4829BCCB1A" box="[1169,1203,1059,1084]" italics="true" pageId="1" pageNumber="88">ca.</emphasis>
<specimenCount id="FCFDFD26FF95A11FDEF76A492807CB1B" box="[1210,1288,1058,1085]" pageId="1" pageNumber="88" type="generic">
<quantity id="2D039B4AFF95A11FDEF76A49280BCB1B" box="[1210,1284,1058,1085]" metricMagnitude="2" metricUnit="m" metricValue="5.12" pageId="1" pageNumber="88" unit="m" value="512.0">512 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="A2E16524FF95A11FDAF06A2D2EA6CBA2" pageId="1" pageNumber="88" type="etymology">
<paragraph id="EA4436AFFF95A11FDAF06A2D2EA6CBA2" blockId="1.[136,1452,158,1661]" pageId="1" pageNumber="88">
<emphasis id="D88FEABDFF95A11FDAF06A2D2C6BCB46" bold="true" box="[189,356,1094,1120]" pageId="1" pageNumber="88">Etymology:—</emphasis>
The species is named in honor of Ms. Minh An Hoang, the second daughter of the first author. She is always the first author’s motivation to research and find new species.
</paragraph>
</subSubSection>
<subSubSection id="A2E16524FF95A11FDAF06AE52961CA7B" pageId="1" pageNumber="88" type="conservation">
<paragraph id="EA4436AFFF95A11FDAF06AE52961CA7B" blockId="1.[136,1452,158,1661]" pageId="1" pageNumber="88">
<emphasis id="D88FEABDFF95A11FDAF06AE52CDCCB8E" bold="true" box="[189,467,1166,1192]" pageId="1" pageNumber="88">Conservation status:—</emphasis>
We assess
<taxonomicName id="2DFB4D2CFF95A11FD81C6AE42FE5CB8E" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[593,746,1166,1192]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="minhaniae">
<emphasis id="D88FEABDFF95A11FD81C6AE42FE5CB8E" box="[593,746,1166,1192]" italics="true" pageId="1" pageNumber="88">B. minhaniae</emphasis>
</taxonomicName>
to be Endangered based on criteria ENC2a(i) (IUCN 2022). This species has only four known populations in one single location with the number of mature individuals being less than 100, all of the populations were in areas which have no formal legal protection (its distribution overlapping the Mu Cang Chai Protection Forest, but it is not included on any national protected areas system). In the areas, the most prominent threat to the new species is deforestation, along with other human activities such as such as farming from around the habitat, which means it is prone to the risk of extinction due to chance events.
</paragraph>
</subSubSection>
<subSubSection id="A2E16524FF95A11FDAF06B0D29EBC95B" pageId="1" pageNumber="88" type="discussion">
<paragraph id="EA4436AFFF95A11FDAF06B0D29EBC95B" blockId="1.[136,1452,158,1661]" pageId="1" pageNumber="88">
<emphasis id="D88FEABDFF95A11FDAF06B0D2C29CAA6" bold="true" box="[189,294,1382,1408]" pageId="1" pageNumber="88">Notes:—</emphasis>
Like
<taxonomicName id="2DFB4D2CFF95A11FDB2E6B0C2EF1CAA6" authority="Lanors., Lamxay &amp; Souvann." authorityName="Lanors., Lamxay &amp; Souvann." box="[355,1022,1382,1409]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="phouchomvoyensis">
<emphasis id="D88FEABDFF95A11FDB2E6B0C2F90CAA6" box="[355,671,1382,1408]" italics="true" pageId="1" pageNumber="88">Begonia phouchomvoyensis</emphasis>
Lanors., Lamxay &amp; Souvann.
</taxonomicName>
(Lanorsavanh
<emphasis id="D88FEABDFF95A11FDEE66B0C29E8CAA6" box="[1195,1255,1382,1408]" italics="true" pageId="1" pageNumber="88">et al.</emphasis>
, 2020), this new species belongs to sect.
<emphasis id="D88FEABDFF95A11FDBDD6BE12F28CA82" box="[400,551,1418,1444]" italics="true" pageId="1" pageNumber="88">Platycentrum</emphasis>
. The two share several characters including rhizomatous habit, hairy petioles and abaxial lamina, 4-tepaled staminate flowers, pistillate flowers with 3 tepals and ovary 3 locules, but the new species can be distinguished by its stipule margin ciliate (vs. entire), the petiole densely subappressed yellowish-brown tomentose (vs. red villous), the leaf is glabrous on the adaxial surface (vs. strigose), the peduncle sparsely yellowish-brown tomentose (vs. red villous) and pedicel of pistillate flowers with two bracteoles (vs. bracteoles absent). A comparison of
<taxonomicName id="2DFB4D2CFF95A11FDAEA68542C32C97E" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[167,317,1598,1624]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="minhaniae">
<emphasis id="D88FEABDFF95A11FDAEA68542C32C97E" box="[167,317,1598,1624]" italics="true" pageId="1" pageNumber="88">B. minhaniae</emphasis>
</taxonomicName>
and
<taxonomicName id="2DFB4D2CFF95A11FDB3B68542F65C97E" box="[374,618,1598,1624]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="phouchomvoyensis">
<emphasis id="D88FEABDFF95A11FDB3B68542F65C97E" box="[374,618,1598,1624]" italics="true" pageId="1" pageNumber="88">B. phouchomvoyensis</emphasis>
</taxonomicName>
is presented in
<tableCitation id="A7790314FF95A11FD95168552E7EC97E" box="[796,881,1598,1624]" captionStart="TABLE 1" captionStartId="1.[136,229,1706,1732]" captionTargetPageId="1" captionText="TABLE 1. Comparison of Begonia minhaniae T.S.Hoang &amp; C.W.Lin and B. phouchomvoyensis Lanors., Lamxay &amp; Souvann." httpUri="http://table.plazi.org/id/BE846627FF95A11FDAC568C12DFDC9CE" pageId="1" pageNumber="88" tableUuid="BE846627FF95A11FDAC568C12DFDC9CE">Table 1</tableCitation>
. The character of pistillate flowers with unequal 3 tepals is uncommon in sect.
<emphasis id="D88FEABDFF95A11FDB8868092F53C95A" box="[453,604,1634,1660]" italics="true" pageId="1" pageNumber="88">Platycentrum</emphasis>
, shared between
<taxonomicName id="2DFB4D2CFF95A11FD9526808291DC95A" box="[799,1042,1634,1660]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="phouchomvoyensis">
<emphasis id="D88FEABDFF95A11FD9526808291DC95A" box="[799,1042,1634,1660]" italics="true" pageId="1" pageNumber="88">B. phouchomvoyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="2DFB4D2CFF95A11FDE05680829D2C95A" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[1096,1245,1634,1660]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="88" phylum="Tracheophyta" rank="species" species="minhaniae">
<emphasis id="D88FEABDFF95A11FDE05680829D2C95A" box="[1096,1245,1634,1660]" italics="true" pageId="1" pageNumber="88">B. minhaniae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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