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<document id="EA25DB6789B03EAB70C56BD2CB264A93" ID-DOI="10.1080/00222930701695629" ID-ISSN="1464-5262" ID-Zenodo-Dep="5233167" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629510120364" checkinUser="felipe" docAuthor="Gheerardyn, Hendrik, Fiers, Frank, Vincx, Magda &amp; Troch, Marleen De" docDate="2007" docId="D34587E1FFB9FFCEFE9AFE02572AFE15" docLanguage="en" docName="JNatHist.41.37-40.2479-2510.pdf" docOrigin="Journal of Natural History 41 (37 - 40)" docSource="http://dx.doi.org/10.1080/00222930701695629" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Tapholeon arenicolus Gheerardyn &amp; Fiers &amp; Vincx &amp; Troch 2007, comb. nov." docType="treatment" docVersion="3" lastPageNumber="2493" masterDocId="2F7CFF99FFB5FFDEFFCDFFCF547EFFEF" masterDocTitle="Revision of the genus Tapholeon Wells, 1967 (Copepoda, Harpacticoida, Laophontidae)" masterLastPageNumber="2510" masterPageNumber="2479" pageNumber="2489" updateTime="1699075389853" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="EDE3B50E1558461A2729A81BBBB529BD">Revision of the genus Tapholeon Wells, 1967 (Copepoda, Harpacticoida, Laophontidae)</mods:title>
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<mods:namePart id="3702A61FB512D90B078B65AA21F67469">Gheerardyn, Hendrik</mods:namePart>
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<mods:namePart id="5C78179344762DFC4D3CB3E4217AD92C">Fiers, Frank</mods:namePart>
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<mods:namePart id="3DC218BAB98D263729B225B8E3678B4A">Vincx, Magda</mods:namePart>
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<mods:namePart id="E598AE948375B77AEF3F85C32A6DFBE5">Troch, Marleen De</mods:namePart>
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<mods:typeOfResource id="B34840F0DF2542410077397F5E8F9070">text</mods:typeOfResource>
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<mods:title id="07B4DECF064483C0617B4F797246DCEA">Journal of Natural History</mods:title>
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<mods:date id="A22A7A4DDCB223D0C22080BBF50442F1">2007</mods:date>
<mods:detail id="59F9D39703C2992FC6B3C89B55719719" type="pubDate">
<mods:number id="2BA063CD0BD9C21B82AF613E542E8505">2010-12-02</mods:number>
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<mods:detail id="DED0E8A70BE6EC1382661CD0EDC0E39A" type="volume">
<mods:number id="C2A716B907B4FFE9309047779B77DBA7">41</mods:number>
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<mods:detail id="7FB05944EF72684FADE343469A5C9653" type="issue">
<mods:number id="047B89A5A9BAF87E5E3E678EB05C3494">37 - 40</mods:number>
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<mods:classification id="A86EE136B2E076F20AA78C073CBF69F5">journal article</mods:classification>
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<mods:identifier id="C52F0DCC14D6019649009C04173000A8" type="ISSN">1464-5262</mods:identifier>
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<treatment id="D34587E1FFB9FFCEFE9AFE02572AFE15" LSID="urn:lsid:plazi:treatment:D34587E1FFB9FFCEFE9AFE02572AFE15" httpUri="http://treatment.plazi.org/id/D34587E1FFB9FFCEFE9AFE02572AFE15" lastPageId="16" lastPageNumber="2493" pageId="12" pageNumber="2489">
<subSubSection id="13F6657CFFB9FFD2FE9AFE0257AAFE0A" box="[343,980,461,485]" pageId="12" pageNumber="2489" type="nomenclature">
<paragraph id="5B5336F7FFB9FFD2FE9AFE0257AAFE0A" blockId="12.[141,980,461,551]" box="[343,980,461,485]" pageId="12" pageNumber="2489">
<heading id="001B819BFFB9FFD2FE9AFE0257AAFE0A" box="[343,980,461,485]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="2489" reason="2">
<taxonomicName id="9CEC4D74FFB9FFD2FE9AFE025739FE0A" authority="(Chappuis, 1954)" authorityName="Gheerardyn &amp; Fiers &amp; Vincx &amp; Troch" authorityYear="2007" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[343,839,461,485]" class="Hexanauplia" family="Laophontidae" genus="Tapholeon" kingdom="Animalia" order="Harpacticoida" pageId="12" pageNumber="2489" phylum="Arthropoda" rank="species" species="arenicolus" status="comb. nov.">
<emphasis id="6998EAE5FFB9FFD2FE9AFE025611FE0A" bold="true" box="[343,623,461,485]" italics="true" pageId="12" pageNumber="2489">Tapholeon arenicolus</emphasis>
(
<bibRefCitation id="3F7D4B06FFB9FFD2FD4DFE02573EFE0A" author="Chappuis PA" box="[640,832,461,485]" pageId="12" pageNumber="2489" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72AB579EFFB9FFD2FC9CFE0257AAFE0A" box="[849,980,461,485]" pageId="12" pageNumber="2489" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13F6657CFFB9FFCEFD8AFE21572AFE15" lastPageId="16" lastPageNumber="2493" pageId="12" pageNumber="2489" type="description">
<paragraph id="5B5336F7FFB9FFD2FD8AFE21569AFDE9" blockId="12.[141,980,461,551]" box="[583,740,494,518]" pageId="12" pageNumber="2489">
(
<figureCitation id="C3D72A72FFB9FFD2FD82FE2156C3FDE9" box="[591,701,494,518]" captionStart="Figure 5" captionStartId="13.[125,185,1448,1467]" captionTargetBox="[134,1161,134,1410]" captionTargetId="figure-1@13.[133,1164,133,1412]" captionTargetPageId="13" captionText="Figure 5. (A–F) Tapholeon arenicolus (Chappuis, 1954) comb. nov.: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G) Tapholeon ornatus Wells, 1967: female anal somite and caudal rami, dorsal. Scale bars in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="12" pageNumber="2489">Figures 5</figureCitation>
,
<figureCitation id="C3D72A72FFB9FFD2FD06FE2156A5FDE9" box="[715,731,494,518]" captionStart="Figure 6" captionStartId="14.[138,198,1590,1609]" captionTargetBox="[164,1158,135,1552]" captionTargetId="figure-0@14.[163,1159,133,1554]" captionTargetPageId="14" captionText="Figure 6. Tapholeon arenicolus (Chappuis, 1954) comb. nov. (A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="12" pageNumber="2489">6</figureCitation>
)
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF40FDC056E1FDC8" blockId="12.[141,980,461,551]" box="[141,671,527,551]" pageId="12" pageNumber="2489">
<emphasis id="6998EAE5FFB9FFD2FF40FDC05482FDC8" box="[141,252,527,551]" italics="true" pageId="12" pageNumber="2489">Synonym.</emphasis>
<taxonomicName id="9CEC4D74FFB9FFD2FEC9FDC056E5FDC8" authority="Chappuis, 1954" authorityName="Chappuis" authorityYear="1954" box="[260,667,527,551]" class="Hexanauplia" family="Laophontidae" genus="Asellopsis" kingdom="Animalia" order="Harpacticoida" pageId="12" pageNumber="2489" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="6998EAE5FFB9FFD2FEC9FDC055ADFDC8" box="[260,467,527,551]" italics="true" pageId="12" pageNumber="2489">Asellopsis arenicola</emphasis>
<bibRefCitation id="3F7D4B06FFB9FFD2FE11FDC056E5FDC8" author="Chappuis PA" box="[476,667,527,551]" pageId="12" pageNumber="2489" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FDAF5568FD97" blockId="12.[138,278,608,632]" box="[138,278,608,632]" pageId="12" pageNumber="2489">
<emphasis id="6998EAE5FFB9FFD2FF47FDAF5568FD97" box="[138,278,608,632]" italics="true" pageId="12" pageNumber="2489">
<typeStatus id="84578855FFB9FFD2FF47FDAF54BEFD98" box="[138,192,608,631]" pageId="12" pageNumber="2489">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FD5F5695FD26" blockId="12.[138,1184,656,713]" pageId="12" pageNumber="2489">
<materialsCitation id="EB843CAAFFB9FFD2FF47FD5F5699FD26" country="Comoros" location="Grande Comore" pageId="12" pageNumber="2489" specimenCount="1" stateProvince="Grande Comore" typeStatus="holotype">
<collectingCountry id="23FB7667FFB9FFD2FF47FD5F5536FD47" box="[138,328,656,680]" name="Comoros" pageId="12" pageNumber="2489">Comoro Islands</collectingCountry>
,
<collectingRegion id="9928F815FFB9FFD2FE98FD5F566AFD47" box="[341,532,656,680]" country="Comoros" name="Grande Comore" pageId="12" pageNumber="2489">Grande Comore</collectingRegion>
(beach of Mitsamiouli), very fine coral sand, and gravel with coral rubble and shell debris (
<bibRefCitation id="3F7D4B06FFB9FFD2FDEEFD7E56A0FD26" author="Chappuis PA" box="[547,734,689,713]" pageId="12" pageNumber="2489" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis 1954</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FCCD555CFCF5" blockId="12.[138,290,770,794]" box="[138,290,770,794]" pageId="12" pageNumber="2489">
<emphasis id="6998EAE5FFB9FFD2FF47FCCD555CFCF5" box="[138,290,770,794]" italics="true" pageId="12" pageNumber="2489">
<typeStatus id="84578855FFB9FFD2FF47FCCD54BEFCF6" box="[138,192,770,793]" pageId="12" pageNumber="2489">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FCFD5578FCA5" blockId="12.[138,262,818,842]" box="[138,262,818,842]" pageId="12" pageNumber="2489">Unknown.</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FC4C5521FC74" blockId="12.[138,351,899,923]" box="[138,351,899,923]" pageId="12" pageNumber="2489">
<emphasis id="6998EAE5FFB9FFD2FF47FC4C5521FC74" box="[138,351,899,923]" italics="true" pageId="12" pageNumber="2489">Additional material</emphasis>
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FC7B5489FBA2" blockId="12.[138,1185,948,1101]" pageId="12" pageNumber="2489">
<materialsCitation id="EB843CAAFFB9FFD2FF47FC7B548DFBA2" collectingDate="1984-08-03" collectionCode="COP" country="Comoros" location="Groupe Plongee de l'Expedition Karthala" municipality="Ouroveni" pageId="12" pageNumber="2489" specimenCode="COP 1984, COP 1985, COP 1986, COP 1987" specimenCount="50" stateProvince="Grande Comore">
<collectingCountry id="23FB7667FFB9FFD2FF47FC7B5487FC23" box="[138,249,948,972]" name="Comoros" pageId="12" pageNumber="2489">Comoros</collectingCountry>
, southeastern coast of
<collectingRegion id="9928F815FFB9FFD2FDDBFC7B56A3FC23" box="[534,733,948,972]" country="Comoros" name="Grande Comore" pageId="12" pageNumber="2489">Grande Comore</collectingRegion>
,
<collectingMunicipality id="BB37AC8DFFB9FFD2FD23FC7B5725FC23" box="[750,859,948,972]" pageId="12" pageNumber="2489">Ouroveni</collectingMunicipality>
(11
<emphasis id="6998EAE5FFB9FFD2FC5DFC7B57E4FC23" bold="true" box="[912,922,948,972]" italics="true" pageId="12" pageNumber="2489">°</emphasis>
549S, 43
<emphasis id="6998EAE5FFB9FFD2FBCEFC7B5073FC23" bold="true" box="[1027,1037,948,972]" italics="true" pageId="12" pageNumber="2489">°</emphasis>
299E), small protected creek with mangrove, sample of fine sand, two dissected females (
<specimenCode id="0B4A9E8CFFB9FFD2FBDAFC1B50E2FC03" box="[1047,1180,980,1004]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="12" pageNumber="2489">COP 1984</specimenCode>
,
<specimenCode id="0B4A9E8CFFB9FFD2FF47FC3A5570FBE2" box="[138,270,1013,1037]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="12" pageNumber="2489">COP 1985</specimenCode>
), one dissected male (
<specimenCode id="0B4A9E8CFFB9FFD2FDD8FC3A56E5FBE2" box="[533,667,1013,1037]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="12" pageNumber="2489">COP 1986</specimenCode>
), and more than
<specimenCount id="4DEAFD7EFFB9FFD2FCA7FC3A5076FBE2" box="[874,1032,1013,1037]" pageId="12" pageNumber="2489" type="generic">50 specimens</specimenCount>
preserved in alcohol (
<specimenCode id="0B4A9E8CFFB9FFD2FF3BFBDA55FDFBC2" box="[246,387,1045,1069]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="12" pageNumber="2489">COP 1987</specimenCode>
); collected
<date id="2F521037FFB9FFD2FDD5FBDA56ABFBC2" box="[536,725,1045,1069]" pageId="12" pageNumber="2489" value="1984-08-03">
<collectingDate id="3F16E9DFFFB9FFD2FDD5FBDA56ABFBC2" box="[536,725,1045,1069]" pageId="12" pageNumber="2489" value="1984-08-03">3 August 1984</collectingDate>
</date>
by
<location id="5E33602CFFB9FFD2FCDCFBDA548DFBA2" LSID="urn:lsid:plazi:treatment:D34587E1FFB9FFCEFE9AFE02572AFE15:5E33602CFFB9FFD2FCDCFBDA548DFBA2" country="Comoros" municipality="Ouroveni" name="Groupe Plongee de l'Expedition Karthala" pageId="12" pageNumber="2489" stateProvince="Grande Comore">Groupe Plongée de l’Expedition Karthala</location>
</materialsCitation>
.
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF47FB4954B5FADF" blockId="12.[138,386,1158,1180]" lastBlockId="12.[138,1185,1206,1685]" pageId="12" pageNumber="2489">
<emphasis id="6998EAE5FFB9FFD2FF47FB4955FCFB71" box="[138,386,1158,1182]" italics="true" pageId="12" pageNumber="2489">Redescription of female</emphasis>
Total body length 354–398 Mm (
<emphasis id="6998EAE5FFB9FFD2FDC7FB775666FB20" box="[522,536,1208,1231]" italics="true" pageId="12" pageNumber="2489">n</emphasis>
510; average5373 Mm; measured from anterior margin of rostrum to posterior margin of caudal rami). Largest width measured at posterior margin of cephalothorax: 97 Mm. Measurements by
<bibRefCitation id="3F7D4B06FFB9FFD2FD6AFB385709FAE0" author="Chappuis PA" box="[679,887,1271,1295]" pageId="12" pageNumber="2489" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
:
<quantity id="9C149B12FFB9FFD2FC44FB38579FFAE0" box="[905,993,1271,1295]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="2489" unit="mm" value="0.4">0.4 mm</quantity>
without caudal setae.
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF68FAF65723FABE" blockId="12.[138,1185,1206,1685]" box="[165,861,1337,1361]" pageId="12" pageNumber="2489">Rostrum triangular and prominent; fused to cephalothorax.</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF68FA955588F9DC" blockId="12.[138,1185,1206,1685]" pageId="12" pageNumber="2489">
Habitus (
<figureCitation id="C3D72A72FFB9FFD2FEDCFA9555F4FA9D" box="[273,394,1370,1394]" captionStart="Figure 5" captionStartId="13.[125,185,1448,1467]" captionTargetBox="[134,1161,134,1410]" captionTargetId="figure-1@13.[133,1164,133,1412]" captionTargetPageId="13" captionText="Figure 5. (A–F) Tapholeon arenicolus (Chappuis, 1954) comb. nov.: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G) Tapholeon ornatus Wells, 1967: female anal somite and caudal rami, dorsal. Scale bars in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="12" pageNumber="2489">Figure 5A</figureCitation>
): body elongate and slender; urosome weakly tapering towards anal somite. Genital double-somite and following urosomite slightly extended laterally. Integument of cephalothorax and pleurotergites with irregular pattern of small denticles. Posterodorsal margin of cephalothorax smooth, of free prosomites smooth along median part but serrate laterally, of all urosomites serrate. Posterodorsal margins of all free somites (except second urosomite) clothed with slender hairs. Anal operculum convex with slightly serrate margin, not protruding.
</paragraph>
<paragraph id="5B5336F7FFB9FFD2FF68F9F35558F97A" blockId="12.[138,1185,1206,1685]" pageId="12" pageNumber="2489">Ventral surface of genital double-somite and following urosomites smooth. Posteroventral margins of genital double-somite and of next two urosomites with long, slender hairs.</paragraph>
<caption id="0F93667FFFB8FFD3FFB0FA675602F9E5" ID-DOI="http://doi.org/10.5281/zenodo.5233177" ID-Zenodo-Dep="5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="13" pageNumber="2490" startId="13.[125,185,1448,1467]" targetBox="[134,1161,134,1410]" targetPageId="13">
<paragraph id="5B5336F7FFB8FFD3FFB0FA675602F9E5" blockId="13.[125,1172,1448,1547]" pageId="13" pageNumber="2490">
Figure 5. (A–F)
<taxonomicName id="9CEC4D74FFB8FFD3FED7FA675604FA54" authority="(Chappuis, 1954)" authorityName="Gheerardyn &amp; Fiers &amp; Vincx &amp; Troch" authorityYear="2007" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[282,634,1448,1467]" class="Hexanauplia" family="Laophontidae" genus="Tapholeon" kingdom="Animalia" order="Harpacticoida" pageId="13" pageNumber="2490" phylum="Arthropoda" rank="species" species="arenicolus" status="comb. nov.">
<emphasis id="6998EAE5FFB8FFD3FED7FA6755AEFA54" box="[282,464,1448,1467]" italics="true" pageId="13" pageNumber="2490">Tapholeon arenicolus</emphasis>
(
<bibRefCitation id="3F7D4B06FFB8FFD3FE11FA67560CFA54" author="Chappuis PA" box="[476,626,1448,1467]" pageId="13" pageNumber="2490" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72AB579EFFB8FFD3FDB2FA675695FA54" box="[639,747,1448,1467]" pageId="13" pageNumber="2490" rank="species">comb. nov.</taxonomicNameLabel>
: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G)
<taxonomicName id="9CEC4D74FFB8FFD3FCFEFA125033FA1F" authority="Wells, 1967" authorityName="Wells" authorityYear="1967" box="[819,1101,1501,1520]" class="Hexanauplia" family="Laophontidae" genus="Tapholeon" kingdom="Animalia" order="Harpacticoida" pageId="13" pageNumber="2490" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="6998EAE5FFB8FFD3FCFEFA1257AAFA1F" box="[819,980,1501,1520]" italics="true" pageId="13" pageNumber="2490">Tapholeon ornatus</emphasis>
<bibRefCitation id="3F7D4B06FFB8FFD3FC16FA125033FA1F" author="Wells JBJ" box="[987,1101,1501,1520]" pageId="13" pageNumber="2490" pagination="189 - 358" refId="ref11471" refString="Wells JBJ. 1967. The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh 67 (7): 189 - 358." type="journal article" year="1967">Wells, 1967</bibRefCitation>
</taxonomicName>
: female anal somite and caudal rami, dorsal. Scale bars in Mm.
</paragraph>
</caption>
<caption id="0F93667FFFBBFFD0FF47F9F95505F991" ID-DOI="http://doi.org/10.5281/zenodo.5233179" ID-Zenodo-Dep="5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="14" pageNumber="2491" startId="14.[138,198,1590,1609]" targetBox="[164,1158,135,1552]" targetPageId="14">
<paragraph id="5B5336F7FFBBFFD0FF47F9F95505F991" blockId="14.[138,1186,1590,1663]" pageId="14" pageNumber="2491">
Figure 6.
<taxonomicName id="9CEC4D74FFBBFFD0FF27F9F95631F9A6" authority="(Chappuis, 1954)" authorityName="Gheerardyn &amp; Fiers &amp; Vincx &amp; Troch" authorityYear="2007" baseAuthorityName="Chappuis" baseAuthorityYear="1954" box="[234,591,1590,1609]" class="Hexanauplia" family="Laophontidae" genus="Tapholeon" kingdom="Animalia" order="Harpacticoida" pageId="14" pageNumber="2491" phylum="Arthropoda" rank="species" species="arenicolus" status="comb. nov.">
<emphasis id="6998EAE5FFBBFFD0FF27F9F955DEF9A6" box="[234,416,1590,1609]" italics="true" pageId="14" pageNumber="2491">Tapholeon arenicolus</emphasis>
(
<bibRefCitation id="3F7D4B06FFBBFFD0FE62F9F95639F9A6" author="Chappuis PA" box="[431,583,1590,1609]" pageId="14" pageNumber="2491" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis, 1954</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="72AB579EFFBBFFD0FD9BF9F956BEF9A6" box="[598,704,1590,1609]" pageId="14" pageNumber="2491" rank="species">comb. nov.</taxonomicNameLabel>
(A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm.
</paragraph>
</caption>
<paragraph id="5B5336F7FFBAFFD1FF55FF4657C4FECE" blockId="15.[125,1173,137,681]" pageId="15" pageNumber="2492">
Caudal rami (
<figureCitation id="C3D72A72FFBAFFD1FEF7FF4655ABFF4E" box="[314,469,137,161]" captionStart="Figure 5" captionStartId="13.[125,185,1448,1467]" captionTargetBox="[134,1161,134,1410]" captionTargetId="figure-1@13.[133,1164,133,1412]" captionTargetPageId="13" captionText="Figure 5. (A–F) Tapholeon arenicolus (Chappuis, 1954) comb. nov.: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G) Tapholeon ornatus Wells, 1967: female anal somite and caudal rami, dorsal. Scale bars in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="15" pageNumber="2492">Figure 5B, C</figureCitation>
) flattened; as long as wide, with subquadrate appearance in dorsal view; tapering strongly towards apical margin in lateral view. Surface strongly concave beyond implantation of dorsal seta. Distal inner corner rounded, strongly serrate. Lateral margin spinulose. Seta IV short; seta V about two times length of caudal ramus and pinnate; all other setae naked. Seta VII inserted near middle of ramus.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FEE5577CFE6C" blockId="15.[125,1173,137,681]" pageId="15" pageNumber="2492">
Antennule (
<figureCitation id="C3D72A72FFBAFFD1FEEDFEE555EBFEAD" box="[288,405,298,322]" captionStart="Figure 5" captionStartId="13.[125,185,1448,1467]" captionTargetBox="[134,1161,134,1410]" captionTargetId="figure-1@13.[133,1164,133,1412]" captionTargetPageId="13" captionText="Figure 5. (A–F) Tapholeon arenicolus (Chappuis, 1954) comb. nov.: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G) Tapholeon ornatus Wells, 1967: female anal somite and caudal rami, dorsal. Scale bars in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="15" pageNumber="2492">Figure 5E</figureCitation>
) six-segmented. First, second, and third segment each with row of spinules. Second segment with strong, armed spine along outer margin. Ultimate segment with rounded tip. Armature formula as in
<typeStatus id="84578855FFBAFFD1FDA2FEA456DEFE6C" box="[623,672,363,387]" pageId="15" pageNumber="2492">type</typeStatus>
species.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FE435759FE4B" blockId="15.[125,1173,137,681]" box="[152,807,396,420]" pageId="15" pageNumber="2492">
Antenna (
<figureCitation id="C3D72A72FFBAFFD1FEC1FE4355FEFE4B" box="[268,384,396,420]" captionStart="Figure 5" captionStartId="13.[125,185,1448,1467]" captionTargetBox="[134,1161,134,1410]" captionTargetId="figure-1@13.[133,1164,133,1412]" captionTargetPageId="13" captionText="Figure 5. (A–F) Tapholeon arenicolus (Chappuis, 1954) comb. nov.: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G) Tapholeon ornatus Wells, 1967: female anal somite and caudal rami, dorsal. Scale bars in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="15" pageNumber="2492">Figure 5F</figureCitation>
) and mouthparts as in
<typeStatus id="84578855FFBAFFD1FD59FE4356BBFE4B" box="[660,709,396,420]" pageId="15" pageNumber="2492">type</typeStatus>
species.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FE6256A2FE0A" blockId="15.[125,1173,137,681]" pageId="15" pageNumber="2492">
P1 (
<figureCitation id="C3D72A72FFBAFFD1FF02FE62553BFE2A" box="[207,325,429,453]" captionStart="Figure 6" captionStartId="14.[138,198,1590,1609]" captionTargetBox="[164,1158,135,1552]" captionTargetId="figure-0@14.[163,1159,133,1554]" captionTargetPageId="14" captionText="Figure 6. Tapholeon arenicolus (Chappuis, 1954) comb. nov. (A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="15" pageNumber="2492">Figure 6A</figureCitation>
) with pattern of spinules as figured. Exopod two-segmented; exp-2 bearing four setae. Enp-1 four times as long as exp.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FE215742FDC9" blockId="15.[125,1173,137,681]" pageId="15" pageNumber="2492">
P2–P4 (
<figureCitation id="C3D72A72FFBAFFD1FF37FE2155EAFDE9" box="[250,404,494,518]" captionStart="Figure 6" captionStartId="14.[138,198,1590,1609]" captionTargetBox="[164,1158,135,1552]" captionTargetId="figure-0@14.[163,1159,133,1554]" captionTargetPageId="14" captionText="Figure 6. Tapholeon arenicolus (Chappuis, 1954) comb. nov. (A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="15" pageNumber="2492">Figure 6B–D</figureCitation>
) with three-segmented exopods and two-segmented endopods, with pattern of spinules as figured. Setal formula in
<tableCitation id="166E034CFFBAFFD1FD2FFDC15742FDC9" box="[738,828,526,550]" captionStart="Table I" captionStartId="4.[138,192,1324,1343]" captionText="Table I. Species of Tapholeon Wells, 1967: number of exopodal segments in P1 and swimming leg setal formulae of P2–P4." pageId="15" pageNumber="2492">Table I.</tableCitation>
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FDE050EAFD67" blockId="15.[125,1173,137,681]" pageId="15" pageNumber="2492">
P5 (
<figureCitation id="C3D72A72FFBAFFD1FF0BFDE05541FDA8" box="[198,319,559,583]" captionStart="Figure 6" captionStartId="14.[138,198,1590,1609]" captionTargetBox="[164,1158,135,1552]" captionTargetId="figure-0@14.[163,1159,133,1554]" captionTargetPageId="14" captionText="Figure 6. Tapholeon arenicolus (Chappuis, 1954) comb. nov. (A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="15" pageNumber="2492">Figure 6D</figureCitation>
) with separate exopod and baseoendopod. Margins of rami furnished with spinules, surface smooth. Baseoendopod reaching beyond middle of exopod; bearing one apical and three lateral setae. Exopod ovate; two times as long as wide; bearing five setae.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FD5E55A3FD46" blockId="15.[125,1173,137,681]" box="[152,477,657,681]" pageId="15" pageNumber="2492">P6 vestige bearing one seta.</paragraph>
<paragraph id="5B5336F7FFBAFFD1FFB0FD2D568BFC43" blockId="15.[125,355,738,760]" lastBlockId="15.[125,1173,786,1168]" pageId="15" pageNumber="2492">
<emphasis id="6998EAE5FFBAFFD1FFB0FD2D551DFD15" box="[125,355,738,762]" italics="true" pageId="15" pageNumber="2492">Redescription of male</emphasis>
Total body length 346–384 Mm (
<emphasis id="6998EAE5FFBAFFD1FE33FCDC5672FCC5" box="[510,524,787,810]" italics="true" pageId="15" pageNumber="2492">n</emphasis>
510; average5362 Mm; measured from anterior margin of rostrum to posterior margin of caudal rami). Largest width measured at posterior margin of cephalothorax: 88 Mm. Habitus as in female; except for fully separated second and third urosomites and lack of lateral extensions in second to fourth urosomites. Ventral surface of third urosomite with transversal row of large spinules.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FC7A5652FC02" blockId="15.[125,1173,786,1168]" pageId="15" pageNumber="2492">
Antennule (
<figureCitation id="C3D72A72FFBAFFD1FEE9FC7A55E3FC22" box="[292,413,949,973]" captionStart="Figure 5" captionStartId="13.[125,185,1448,1467]" captionTargetBox="[134,1161,134,1410]" captionTargetId="figure-1@13.[133,1164,133,1412]" captionTargetPageId="13" captionText="Figure 5. (A–F) Tapholeon arenicolus (Chappuis, 1954) comb. nov.: (A) female habitus, dorsal; (B) female caudal rami, dorsal; (C) female caudal rami, lateral; (D) male antennule (armature omitted), ventral; (E) female antennule (armature omitted), ventral; (F) female antennary exopod. (G) Tapholeon ornatus Wells, 1967: female anal somite and caudal rami, dorsal. Scale bars in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233177" httpUri="https://zenodo.org/record/5233177/files/figure.png" pageId="15" pageNumber="2492">Figure 5D</figureCitation>
) sub-chirocer. Shape of first and second segment as in female. Armature formula as in
<typeStatus id="84578855FFBAFFD1FE54FC1A55B4FC02" box="[409,458,981,1005]" pageId="15" pageNumber="2492">type</typeStatus>
species.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FC3956C1FBE1" blockId="15.[125,1173,786,1168]" box="[152,703,1014,1038]" pageId="15" pageNumber="2492">Antenna, mouthparts, and P1–P4 as in female.</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FBD85508FBA0" blockId="15.[125,1173,786,1168]" pageId="15" pageNumber="2492">
P5 (
<figureCitation id="C3D72A72FFBAFFD1FF01FBD85536FBC0" box="[204,328,1047,1071]" captionStart="Figure 6" captionStartId="14.[138,198,1590,1609]" captionTargetBox="[164,1158,135,1552]" captionTargetId="figure-0@14.[163,1159,133,1554]" captionTargetPageId="14" captionText="Figure 6. Tapholeon arenicolus (Chappuis, 1954) comb. nov. (A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="15" pageNumber="2492">Figure 6G</figureCitation>
): endopodal lobe obsolete; without seta. Exopod small; bearing three strongly armed setae.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FF55FB9754CCFB7F" blockId="15.[125,1173,786,1168]" pageId="15" pageNumber="2492">
P6 vestiges (
<figureCitation id="C3D72A72FFBAFFD1FEE7FB9755E0FB9F" box="[298,414,1112,1136]" captionStart="Figure 6" captionStartId="14.[138,198,1590,1609]" captionTargetBox="[164,1158,135,1552]" captionTargetId="figure-0@14.[163,1159,133,1554]" captionTargetPageId="14" captionText="Figure 6. Tapholeon arenicolus (Chappuis, 1954) comb. nov. (A) Female P1, posterior; (B) female P2, anterior; (C) female P3, anterior; (D) female P5, anterior; (E) female P4, posterior; (F) male P6, anterior; (G) male P5, anterior. Scale bar in Mm." figureDoi="http://doi.org/10.5281/zenodo.5233179" httpUri="https://zenodo.org/record/5233179/files/figure.png" pageId="15" pageNumber="2492">Figure 6F</figureCitation>
) asymmetrical; each bearing a plumose inner and a naked outer seta.
</paragraph>
<paragraph id="5B5336F7FFBAFFD1FFB0FB0654A2FB0E" blockId="15.[125,220,1225,1249]" box="[125,220,1225,1249]" pageId="15" pageNumber="2492">
<emphasis id="6998EAE5FFBAFFD1FFB0FB0654A2FB0E" box="[125,220,1225,1249]" italics="true" pageId="15" pageNumber="2492">Remarks</emphasis>
</paragraph>
<paragraph id="5B5336F7FFBAFFCEFFB0FB355720FF2E" blockId="15.[125,1173,1274,1685]" lastBlockId="16.[138,1185,137,193]" lastPageId="16" lastPageNumber="2493" pageId="15" pageNumber="2492">
<bibRefCitation id="3F7D4B06FFBAFFD1FFB0FB355538FAFD" author="Chappuis PA" box="[125,326,1274,1298]" pageId="15" pageNumber="2492" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
assigned this species to the genus
<taxonomicName id="9CEC4D74FFBAFFD1FD12FB3550EAFAFD" authority="Brady and Robertson, 1873" authorityName="Brady and Robertson" authorityYear="1873" box="[735,1172,1274,1298]" class="Hexanauplia" family="Laophontidae" genus="Asellopsis" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="2492" phylum="Arthropoda" rank="genus">
<emphasis id="6998EAE5FFBAFFD1FD12FB35573AFAFD" box="[735,836,1274,1298]" italics="true" pageId="15" pageNumber="2492">Asellopsis</emphasis>
Brady and Robertson, 1873
</taxonomicName>
because of the remarkably flattened (asellopsiform) caudal rami. Also, at the time of description the genus
<taxonomicName id="9CEC4D74FFBAFFD1FE46FAF55582FABD" authorityName="Wells" authorityYear="1967" box="[395,508,1338,1362]" class="Hexanauplia" family="Laophontidae" genus="Tapholeon" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="2492" phylum="Arthropoda" rank="genus">
<emphasis id="6998EAE5FFBAFFD1FE46FAF55582FABD" box="[395,508,1338,1362]" italics="true" pageId="15" pageNumber="2492">Tapholeon</emphasis>
</taxonomicName>
had not been established. However,
<bibRefCitation id="3F7D4B06FFBAFFD1FC04FAF550EAFABD" author="Chappuis PA" box="[969,1172,1338,1362]" pageId="15" pageNumber="2492" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
made no reference to a three-segmented endopod P
<quantity id="9C149B12FFBAFFD1FD2BFA95576DFA9D" box="[742,787,1370,1394]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="15" pageNumber="2492" unit="in" value="3.0">3 in</quantity>
the male, but only mentioned a slightly shorter inner seta on the second endopodal segment of this leg. In the present material, neither a three-segmented male endopod nor a differentiated seta was found. Therefore,
<taxonomicName id="9CEC4D74FFBAFFD1FECAFA7455ECFA3C" authorityName="Chappuis" authorityYear="1954" box="[263,402,1467,1491]" class="Hexanauplia" family="Laophontidae" genus="Asellopsis" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="2492" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="6998EAE5FFBAFFD1FECAFA7455ECFA3C" box="[263,402,1467,1491]" italics="true" pageId="15" pageNumber="2492">A. arenicola</emphasis>
</taxonomicName>
should be excluded from the genus
<taxonomicName id="9CEC4D74FFBAFFD1FCA8FA7457B4FA3C" authorityName="Brady and Robertson" authorityYear="1873" box="[869,970,1467,1491]" class="Hexanauplia" family="Laophontidae" genus="Asellopsis" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="2492" phylum="Arthropoda" rank="genus">
<emphasis id="6998EAE5FFBAFFD1FCA8FA7457B4FA3C" box="[869,970,1467,1491]" italics="true" pageId="15" pageNumber="2492">Asellopsis</emphasis>
</taxonomicName>
, which typically displays pronounced sexual dimorphism in the third leg. Fortunately,
<bibRefCitation id="3F7D4B06FFBAFFD1FC07FA1350EAFA1B" author="Chappuis PA" box="[970,1172,1500,1524]" pageId="15" pageNumber="2492" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
illustrates the peculiar seta on the second antennular segment. The thickened antennular seta, the lack of sexually dimorphic structures in the legs, and the strongly reduced shape of the P
<quantity id="9C149B12FFBAFFD1FF0DF9F3548FF9BB" box="[192,241,1596,1620]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="15" pageNumber="2492" unit="in" value="5.0">5 in</quantity>
the male clearly indicate that
<taxonomicName id="9CEC4D74FFBAFFD1FDA0F9F35688F9BB" authorityName="Chappuis" authorityYear="1954" box="[621,758,1596,1620]" class="Hexanauplia" family="Laophontidae" genus="Asellopsis" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="2492" phylum="Arthropoda" rank="species" species="arenicola">
<emphasis id="6998EAE5FFBAFFD1FDA0F9F35688F9BB" box="[621,758,1596,1620]" italics="true" pageId="15" pageNumber="2492">A. arenicola</emphasis>
</taxonomicName>
must be transferred to the genus
<taxonomicName id="9CEC4D74FFBAFFD1FFB0F9925490F99A" authorityName="Wells" authorityYear="1967" box="[125,238,1629,1653]" class="Hexanauplia" family="Laophontidae" genus="Tapholeon" kingdom="Animalia" order="Harpacticoida" pageId="15" pageNumber="2492" phylum="Arthropoda" rank="genus">
<emphasis id="6998EAE5FFBAFFD1FFB0F9925490F99A" box="[125,238,1629,1653]" italics="true" pageId="15" pageNumber="2492">Tapholeon</emphasis>
</taxonomicName>
. Other differences with the original decription are: P1 exopod with four setae on the second segment, instead of three as shown by
<bibRefCitation id="3F7D4B06FFBAFFD1FD0DF9B257F7F97A" author="Chappuis PA" box="[704,905,1661,1685]" pageId="15" pageNumber="2492" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis (1954)</bibRefCitation>
; the much longer setae (reaching far beyond the exopod) of the endopods of P2–P4; the presence of an inner seta on exp-2 of P3, and the plumose setae on the rami of the P5.
</paragraph>
<paragraph id="5B5336F7FFA5FFCEFF47FF3B5510FEE3" blockId="16.[138,366,244,268]" box="[138,366,244,268]" pageId="16" pageNumber="2493">
<emphasis id="6998EAE5FFA5FFCEFF47FF3B5510FEE3" box="[138,366,244,268]" italics="true" pageId="16" pageNumber="2493">Differential diagnosis</emphasis>
</paragraph>
<paragraph id="5B5336F7FFA5FFCEFF47FEEA569BFE91" blockId="16.[138,1185,293,382]" pageId="16" pageNumber="2493">The present species bears three outer spines on the ultimate segments of the exopods of P2–P4, lacks an inner seta on exp-2 of P4, and bears two, three and three setae on the second endopodal segments of P2–P4, respectively.</paragraph>
<paragraph id="5B5336F7FFA5FFCEFF47FE7E5575FE26" blockId="16.[138,267,433,457]" box="[138,267,433,457]" pageId="16" pageNumber="2493">
<emphasis id="6998EAE5FFA5FFCEFF47FE7E5575FE26" box="[138,267,433,457]" italics="true" pageId="16" pageNumber="2493">Distribution</emphasis>
</paragraph>
<paragraph id="5B5336F7FFA5FFCEFF47FE2D572AFE15" blockId="16.[138,852,482,506]" box="[138,852,482,506]" pageId="16" pageNumber="2493">
<collectingRegion id="9928F815FFA5FFCEFF47FE2D5533FE15" box="[138,333,482,506]" country="Comoros" name="Grande Comore" pageId="16" pageNumber="2493">Grande Comore</collectingRegion>
,
<collectingCountry id="23FB7667FFA5FFCEFE96FE2D55B9FE15" box="[347,455,482,506]" name="Comoros" pageId="16" pageNumber="2493">Comoros</collectingCountry>
(
<bibRefCitation id="3F7D4B06FFA5FFCEFE1AFE2D56EEFE15" author="Chappuis PA" box="[471,656,482,506]" pageId="16" pageNumber="2493" pagination="45 - 73" refId="ref10823" refString="Chappuis PA. 1954. Recherches sur la faune interstitielle des sediments marins et d'eau douce a Madagascar: IV. Copepodes Harpacticoides psammiques de Madagascar. Memoires de l'Institut Scientifique de Madagascar 9: 45 - 73." type="journal article" year="1954">Chappuis 1954</bibRefCitation>
; present report).
</paragraph>
</subSubSection>
</treatment>
</document>
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