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<document id="44E04B31EE44B81DFB3F825A71A72504" ID-CLB-Dataset="44104" ID-DOI="10.5281/zenodo.211551" ID-GBIF-Dataset="1259ff0f-cb7c-4693-b809-e6a61e2c478c" ID-ISSN="1175-5326" ID-Zenodo-Dep="211551" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" checkinTime="1460557931563" checkinUser="plazi" docAuthor="Chani-Posse, Mariana &amp; Couturier, Guy" docDate="2012" docId="601CA77AFF95FF98BFD70B1B7BA5FE8F" docLanguage="en" docName="zt03568p088.pdf" docOrigin="Zootaxa 3568" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Delgadobius amazonensis Chani-Posse &amp; Couturier 2012, new species" docType="treatment" docVersion="12" lastPageNumber="87" masterDocId="9C25DF02FF96FF9FBF4008227815FFB0" masterDocTitle="Delgadobius amazonensis — a new genus and species of the subtribe Philonthina from Amazonia (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="88" masterPageNumber="81" pageNumber="84" updateTime="1729190770081" updateUser="ExternalLinkService">
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<mods:title id="ADC2F8968B130515B508E8F23070FD15">Delgadobius amazonensis — a new genus and species of the subtribe Philonthina from Amazonia (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="46F2FDF167C21D16B33384EA6458B026">Chani-Posse, Mariana</mods:namePart>
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<mods:namePart id="18148A690265CB3D1D3768B25F462495">Couturier, Guy</mods:namePart>
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<treatment id="601CA77AFF95FF98BFD70B1B7BA5FE8F" ID-DOI="http://doi.org/10.5281/zenodo.6172335" ID-GBIF-Taxon="119556166" ID-Zenodo-Dep="6172335" LSID="urn:lsid:plazi:treatment:601CA77AFF95FF98BFD70B1B7BA5FE8F" httpUri="http://treatment.plazi.org/id/601CA77AFF95FF98BFD70B1B7BA5FE8F" lastPageId="7" lastPageNumber="87" pageId="3" pageNumber="84">
<subSubSection id="A0AF45E7FF95FF9CBFD70B1B7935FCC4" pageId="3" pageNumber="84" type="nomenclature">
<paragraph id="E80A166CFF95FF9CBFD70B1B7BA7FCE2" blockId="3.[151,946,824,884]" box="[151,946,824,851]" pageId="3" pageNumber="84">
<heading id="B342A100FF95FF9CBFD70B1B7BA7FCE2" bold="true" box="[151,946,824,851]" fontSize="11" level="1" pageId="3" pageNumber="84" reason="1">
<emphasis id="DAC1CA7EFF95FF9CBFD70B1B7BA7FCE2" bold="true" box="[151,946,824,851]" pageId="3" pageNumber="84">
<taxonomicName id="2FB56DEFFF95FF9CBFD70B1B7B00FCE2" ID-CoL="34FTL" authority="Chani-Posse &amp; Couturier" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[151,789,824,851]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="amazonensis" status="sp. nov.">
<emphasis id="DAC1CA7EFF95FF9CBFD70B1B79C5FCE3" bold="true" box="[151,464,825,851]" italics="true" pageId="3" pageNumber="84">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier
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, 
<taxonomicNameLabel id="C1F27705FF95FF9CBC620B1A7BA7FCE2" box="[802,946,824,850]" pageId="3" pageNumber="84" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<paragraph id="E80A166CFF95FF9CBFD70B7E7935FCC4" blockId="3.[151,946,824,884]" box="[151,288,860,884]" pageId="3" pageNumber="84">
(
<figureCitation id="708E0AE9FF95FF9CBFDF0B7E7902FCC4" box="[159,279,860,884]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Figs. 1–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A0AF45E7FF95FF9CBFD70B817A4AFC0C" box="[151,607,931,957]" pageId="3" pageNumber="84" type="diagnosis">
<paragraph id="E80A166CFF95FF9CBFD70B817A4AFC0C" blockId="3.[151,1437,931,2036]" box="[151,607,931,957]" pageId="3" pageNumber="84">
<emphasis id="DAC1CA7EFF95FF9CBFD70B817904FC0C" bold="true" box="[151,273,931,956]" pageId="3" pageNumber="84">Diagnosis.</emphasis>
As for the genus (see above).
</paragraph>
</subSubSection>
<subSubSection id="A0AF45E7FF95FF9CBF870BE57935F844" pageId="3" pageNumber="84" type="description">
<paragraph id="E80A166CFF95FF9CBF870BE57D9DFC50" blockId="3.[151,1437,931,2036]" box="[199,1416,967,992]" pageId="3" pageNumber="84">
<emphasis id="DAC1CA7EFF95FF9CBF870BE5794CFC50" bold="true" box="[199,345,967,992]" pageId="3" pageNumber="84">Description.</emphasis>
Length of the body 
<quantity id="2F4DBB89FF95FF9CBD7C0BE57AD7FC6F" box="[572,706,967,991]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="7.9" metricValueMin="7.5" pageId="3" pageNumber="84" unit="mm" value="7.7" valueMax="7.9" valueMin="7.5">7.5–7.9 mm</quantity>
(
<quantity id="2F4DBB89FF95FF9CBD910BE57B4EFC6F" box="[721,859,967,991]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="3" pageNumber="84" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 mm</quantity>
, abdomen excluded). Coloration as for the genus.
</paragraph>
<paragraph id="E80A166CFF95FF9CBF870BCE78E7FADC" blockId="3.[151,1437,931,2036]" pageId="3" pageNumber="84">
Head of rounded-quadrangular shape, slightly broadened at distal third and with obtuse hind angles, as long as to slightly wider than long (HW/HL= 1.00–1.11) (
<figureCitation id="708E0AE9FF95FF9CBD950C2D7B0DFB97" box="[725,792,1039,1063]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 1</figureCitation>
), slightly narrower than pronotum at widest point (HW/ PW= 0.88–0.91); postmandibular ridge present; epicranium with medial interocular punctures separated by a distance less than 1.5 times as large as distance separating medial punctures from lateral punctures. Eyes distinctly longer than temples seen from above (EL/TL= 1.75) (
<figureCitation id="708E0AE9FF95FF9CBDAB0C5E7B39FB24" box="[747,812,1148,1172]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 1</figureCitation>
). Antennae with segment 1 shorter than segments 2 and 3 combined, segment 3 slightly longer than segment 2, segments 4 and 5 longer than wide, segment 6 about as long as wide, segments 7 to 10 transverse, last segment minutely emarginate. Mandibles (right and left) with a single tooth (
<figureCitation id="708E0AE9FF95FF9CBFA00CC57930FB4F" box="[224,293,1255,1279]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 2</figureCitation>
). Maxillary palpus with last segment gradually narrowed to subacute apex from apical half, about 1.5 times as long as preceding segment and not appreciably narrower (
<figureCitation id="708E0AE9FF95FF9CBCD70D2E7BCBFA94" box="[919,990,1292,1316]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 2</figureCitation>
). Mentum and submentum subequal in length. Labial palpus with segment 3 distinctly longer than segment 2 and gradually narrowed to subacute apex (
<figureCitation id="708E0AE9FF95FF9CBFDF0D7678F0FADC" box="[159,229,1364,1388]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="E80A166CFF95FF9CBF870D5B7B6FF990" blockId="3.[151,1437,931,2036]" pageId="3" pageNumber="84">
Pronotum slightly longer than wide (PW/PL= 0.89–0.95) (
<figureCitation id="708E0AE9FF95FF9CBC140D5A7B80FA20" box="[852,917,1400,1424]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 1</figureCitation>
), slightly narrowed anteriad; dorsal surface of pronotum with two rows of punctures, each with four punctures. Legs with front and middle tarsi as long as front and middle tibiae, hind tarsi shorter than hind tibiae; front tarsus with first segment shorter than segments 2 and 3 combined (
<figureCitation id="708E0AE9FF95FF9CBE560DC67948FA4C" box="[278,349,1508,1532]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 7</figureCitation>
); mid and hind tarsus with first segment longer than segments 2 and 3 combined (
<figureCitation id="708E0AE9FF95FF9CBA500DC67D43FA4C" box="[1296,1366,1508,1532]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 8</figureCitation>
); first segment of hind tarsus longer than last segment (S1/S5= 1.3–1.5).
</paragraph>
<paragraph id="E80A166CFF95FF9CBF870E0E7C08F9F4" blockId="3.[151,1437,931,2036]" box="[199,1053,1580,1604]" pageId="3" pageNumber="84">Elytra at sides distinctly longer than pronotum at midline (EtL/PL=1.2–1.3).</paragraph>
<paragraph id="E80A166CFF95FF9CBF870E727C1FF93C" blockId="3.[151,1437,931,2036]" pageId="3" pageNumber="84">Abdomen. Terga 3 to 5 with posterior transverse basal carina, acutely extended medially on terga 3 and 4, incomplete on tergum 5; terga 6 to 8 with only anterior transverse basal carina.</paragraph>
<paragraph id="E80A166CFF95FF9CBF870EBA7B3EF8D4" blockId="3.[151,1437,931,2036]" pageId="3" pageNumber="84">
Male genitalia. Sternum 7 slightly emarginate medio-apically; sternum 8 deeply emarginate medio-apically; tergum 10 subangulate at apex, with several apical setae and two to four long and strong subapical setae (
<figureCitation id="708E0AE9FF95FF9CBA080E9E7D98F964" box="[1352,1421,1724,1748]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 9</figureCitation>
); sternum 9 acutely emarginate apically, with several apical setae at each side of emargination and one to two long subapical macrosetae (
<figureCitation id="708E0AE9FF95FF9CBED60F2679FFF8AC" box="[406,490,1796,1820]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 10</figureCitation>
). Aedeagus with parameres fused as one short sclerite only attached to median lobe at base, without sensory peg setae; median lobe elongate, with apical part distinctly narrowed into a thin rod-like apex; internal sac with sclerotized structures (
<figureCitation id="708E0AE9FF95FF9CBDD70F6E7B0BF8D4" box="[663,798,1868,1892]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Figs. 11, 12</figureCitation>
).
</paragraph>
<paragraph id="E80A166CFF95FF9CBF870F527935F844" blockId="3.[151,1437,931,2036]" pageId="3" pageNumber="84">
Female genitalia. Sterna 7 and 8 straight to slightly sinuate apically. Genital segment with styli of tergum 9 similar to those of male; tergum 10 similar to that of male; gonocoxites strongly developed, second gonocoxites rather short, each with two to five strong setae along its outer margin, with a minute stylus (
<figureCitation id="708E0AE9FF95FF9CBBE60F9A7CEDF860" box="[1190,1272,1976,2000]" captionStart="FIGURES 1 – 13" captionStartId="5.[151,264,1891,1914]" captionTargetBox="[233,1358,196,1856]" captionTargetId="figure@5.[225,1360,187,1862]" captionTargetPageId="5" captionText="FIGURES 1 – 13. Delgadobius amazonensis: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR = infraorbital ridge); 5, prosternum and hypomeron (aL = additional line, Pr = pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S = segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars = 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="3" pageNumber="84">Fig. 13</figureCitation>
) and two long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="A0AF45E7FF92FF9BBF8708BA7B2DFF1F" box="[199,824,151,176]" pageId="4" pageNumber="85" type="etymology">
<paragraph id="E80A166CFF92FF9BBF8708BA7B2DFF1F" blockId="4.[151,1436,151,1977]" box="[199,824,151,176]" pageId="4" pageNumber="85">
<emphasis id="DAC1CA7EFF92FF9BBF8708BA795BFF00" bold="true" box="[199,334,152,176]" pageId="4" pageNumber="85">Etymology.</emphasis>
The name refers to the species distribution.
</paragraph>
</subSubSection>
<subSubSection id="A0AF45E7FF92FF9BBF87089E79D4FD5F" pageId="4" pageNumber="85" type="materials_examined">
<paragraph id="E80A166CFF92FF9BBF87089E79D4FD5F" blockId="4.[151,1436,151,1977]" pageId="4" pageNumber="85">
<emphasis id="DAC1CA7EFF92FF9BBF87089E79F2FF64" bold="true" box="[199,487,187,212]" pageId="4" pageNumber="85">
<typeStatus id="370EA8CEFF92FF9BBF87089E7917FF64" box="[199,258,188,212]" pageId="4" pageNumber="85">Type</typeStatus>
material. 
<typeStatus id="370EA8CEFF92FF9BBE3A089E79F2FF64" box="[378,487,188,212]" pageId="4" pageNumber="85" type="holotype">Holotype</typeStatus>
</emphasis>
3, with labels: “ 
<collectingCountry id="90A256FCFF92FF9BBDE1089E7AC2FF64" box="[673,727,188,212]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
– Loreto – 
<date id="9C0B30ACFF92FF9BBC17089E7BD9FF64" box="[855,972,188,212]" pageId="4" pageNumber="85" value="2010-10-30">30.X.2010</date>
/ Iquitos Nauta road, km 46/ 
<geoCoordinate id="8D8170ABFF92FF9BBA4C089E7D7AFF64" box="[1292,1391,188,212]" direction="south" orientation="latitude" pageId="4" pageNumber="85" precision="925" value="-4.0333333">04° 02´S</geoCoordinate>
<geoCoordinate id="8D8170ABFF92FF9BBA35089E78C3FF47" direction="west" orientation="longitude" pageId="4" pageNumber="85" precision="925" value="-73.4">73° 24´W</geoCoordinate>
/ G. Couturier, C. Delgado col.”, “on inflorescence 
<quantity id="2F4DBB89FF92FF9BBC5208C27B56FF47" box="[786,835,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="85" unit="in" value="3.0">3 in</quantity>
anthesis of 
<taxonomicName id="2FB56DEFFF92FF9BBC8A08C37C84FF48" box="[970,1169,224,248]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBC8A08C37C84FF48" box="[970,1169,224,248]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBC8A08C37C3BFF48" box="[970,1070,225,248]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
</taxonomicName>
<taxonomicName id="2FB56DEFFF92FF9BBBD808FD7D07FF47" box="[1176,1298,223,247]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
”, “ 
<typeStatus id="370EA8CEFF92FF9BBA7108FD7D8EFF47" box="[1329,1435,223,247]" pageId="4" pageNumber="85" type="holotype">Holotype</typeStatus>
<taxonomicName id="2FB56DEFFF92FF9BBFD709267B35FEAC" authority="Chani-Posse &amp; Couturier 2012" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[151,800,260,284]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBFD7092679A9FEAC" box="[151,444,260,284]" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier 2012
</taxonomicName>
” (red label) (
<collectionCode id="8EA48EA9FF92FF9BBCF509267C04FEAC" LSID="urn:lsid:biocol.org:col:34988" box="[949,1041,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
). Twenty-four 
<typeStatus id="370EA8CEFF92FF9BBBF709277D37FEAD" box="[1207,1314,261,285]" pageId="4" pageNumber="85" type="paratype">paratypes</typeStatus>
with same data, 1 3 3 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBE7D09057981FE8F" LSID="urn:lsid:biocol.org:col:34795" box="[317,404,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="85">FMNH</collectionCode>
), 2 3 3 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBD5109057A62FE8F" box="[529,631,295,319]" pageId="4" pageNumber="85">IADIZA</collectionCode>
), 1 3 5 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBDB409057B5AFE8F" LSID="urn:lsid:biocol.org:col:34988" box="[756,847,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
), 2 3 3 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBC8C09057C2CFE8F" box="[972,1081,295,319]" pageId="4" pageNumber="85">UNALM</collectionCode>
), 1 3 3 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBBF509057CE8FE8F" box="[1205,1277,295,319]" pageId="4" pageNumber="85">UTCI</collectionCode>
). Other thirtyseven 
<typeStatus id="370EA8CEFF92FF9BBF9F096F7958FED5" box="[223,333,333,357]" pageId="4" pageNumber="85" type="paratype">paratypes</typeStatus>
, with labels: “ 
<collectingCountry id="90A256FCFF92FF9BBEB0096E7A33FED4" box="[496,550,332,356]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
– Loreto – 
<date id="9C0B30ACFF92FF9BBDEE096E7B36FED4" box="[686,803,332,356]" pageId="4" pageNumber="85" value="2008-10-16">16.X.2008</date>
/ Pucaurquillo (Pevas)/ 
<geoCoordinate id="8D8170ABFF92FF9BBB6D096E7C93FED4" box="[1069,1158,332,356]" direction="south" orientation="latitude" pageId="4" pageNumber="85" precision="925" value="-3.3333333">03°20´S</geoCoordinate>
, 
<geoCoordinate id="8D8170ABFF92FF9BBBD3096E7CECFED4" box="[1171,1273,332,356]" direction="west" orientation="longitude" pageId="4" pageNumber="85" precision="925" value="-72.066666">72°04´W</geoCoordinate>
/ G. Couturier, W. Gonzáles col.,“on inflorescence 
<taxonomicName id="2FB56DEFFF92FF9BBD7209537B1CFE38" box="[562,777,369,392]" class="Insecta" family="Nymphalidae" genus="Euterpe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="precatoria">
<emphasis id="DAC1CA7EFF92FF9BBD7209537B1CFE38" box="[562,777,369,392]" italics="true" pageId="4" pageNumber="85">Euterpe precatoria</emphasis>
</taxonomicName>
<taxonomicName id="2FB56DEFFF92FF9BBC5309527B9DFE38" box="[787,904,368,392]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
(beginning anthesis)”, “ 
<typeStatus id="370EA8CEFF92FF9BBBDF094D7D11FE37" box="[1183,1284,367,391]" pageId="4" pageNumber="85" type="paratype">Paratype</typeStatus>
<taxonomicName id="2FB56DEFFF92FF9BBA4D09527A9EFE1C" authority="Chani-Posse &amp; Couturier 2012" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBA4D09527933FE1C" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier 2012
</taxonomicName>
”, 2 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBC4909B67B7AFE1C" box="[777,879,404,428]" pageId="4" pageNumber="85">IADIZA</collectionCode>
), 1 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBCAA09B67C50FE1C" LSID="urn:lsid:biocol.org:col:34988" box="[1002,1093,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
), 1 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBB8109B67D3AFE1C" box="[1217,1327,404,428]" pageId="4" pageNumber="85">MNHUB</collectionCode>
), 1 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBFDF099578F8FE7F" box="[159,237,439,463]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="85">NMW</collectionCode>
); “ 
<collectingCountry id="90A256FCFF92FF9BBE5009957953FE7F" box="[272,326,439,463]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
Loreto Iquitos/ Carretera Iquitos Nauta km/ 
<date id="9C0B30ACFF92FF9BBC2309957BC6FE7F" box="[867,979,439,463]" pageId="4" pageNumber="85" value="1991-10-31">31.X.1991</date>
/K. Mejia Col.”, “Host plant 
<taxonomicName id="2FB56DEFFF92FF9BBA78099B78F3FE44" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="carana">
<emphasis id="DAC1CA7EFF92FF9BBA78099B78F3FE44" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBA78099B7D89FE60" box="[1336,1436,441,464]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
carana
</emphasis>
</taxonomicName>
”, “ 
<typeStatus id="370EA8CEFF92FF9BBE4F09FE7966FE44" box="[271,371,476,500]" pageId="4" pageNumber="85" type="paratype">Paratype</typeStatus>
<taxonomicName id="2FB56DEFFF92FF9BBE3F09FE7C34FE44" authority="Chani-Posse &amp; Couturier 2012" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[383,1057,476,500]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBE3F09FE7ABCFE44" box="[383,681,476,500]" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier 2012
</taxonomicName>
”, 1 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBBF809FE7D01FE44" LSID="urn:lsid:biocol.org:col:34988" box="[1208,1300,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
); “ 
<collectingCountry id="90A256FCFF92FF9BBA7609FE7D81FE44" box="[1334,1428,476,500]" name="Peru" pageId="4" pageNumber="85">PEROU</collectingCountry>
/ Iquitos/ 
<date id="9C0B30ACFF92FF9BBFB809DD7966FDA7" box="[248,371,511,535]" pageId="4" pageNumber="85" value="1985-09">Sept. 1985</date>
/ F. Kahn Réc.”, “sur &quot;Huassai&quot;/ 
<taxonomicName id="2FB56DEFFF92FF9BBDBF0A237BCCFDA8" box="[767,985,513,536]" class="Insecta" family="Nymphalidae" genus="Euterpe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="precatoria">
<emphasis id="DAC1CA7EFF92FF9BBDBF0A237BCCFDA8" box="[767,985,513,536]" italics="true" pageId="4" pageNumber="85">Euterpe precatoria</emphasis>
</taxonomicName>
”, “ 
<typeStatus id="370EA8CEFF92FF9BBB4209DD7C73FDA7" box="[1026,1126,511,535]" pageId="4" pageNumber="85" type="paratype">Paratype</typeStatus>
<taxonomicName id="2FB56DEFFF92FF9BBB320A2279ECFD8C" authority="Chani-Posse &amp; Couturier 2012" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBB320A227D89FDA8" box="[1138,1436,512,536]" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier 2012
</taxonomicName>
”, 1 3 (
<collectionCode id="8EA48EA9FF92FF9BBD0B0A067AB2FD8C" LSID="urn:lsid:biocol.org:col:34988" box="[587,679,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
), 1 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBC6A0A067B94FD8C" box="[810,897,548,572]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="85">ZMUC</collectionCode>
); “ 
<collectingCountry id="90A256FCFF92FF9BBCDF0A067BE8FD8C" box="[927,1021,548,572]" name="Peru" pageId="4" pageNumber="85">PEROU</collectingCountry>
/ Iquitos/ 
<date id="9C0B30ACFF92FF9BBB2C0A067CF6FD8C" box="[1132,1251,548,572]" pageId="4" pageNumber="85" value="1985-09">Sept. 1985</date>
/ F. Kahn Réc.”, “sur &quot;Ungurahui&quot;&quot;/ 
<taxonomicName id="2FB56DEFFF92FF9BBE360A6B7A3FFDD0" box="[374,554,584,608]" class="Liliopsida" family="Arecaceae" genus="Jessenia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="bataua">
<emphasis id="DAC1CA7EFF92FF9BBE360A6B7A3FFDD0" box="[374,554,584,608]" italics="true" pageId="4" pageNumber="85">Jessenia bataua</emphasis>
</taxonomicName>
”, “ 
<typeStatus id="370EA8CEFF92FF9BBD0E0A657AA6FDEF" box="[590,691,583,607]" pageId="4" pageNumber="85" type="paratype">Paratype</typeStatus>
<taxonomicName id="2FB56DEFFF92FF9BBDFA0A6A7D52FDEF" authority="Chani-Posse &amp; Couturier 2012" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[698,1351,583,608]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBDFA0A6A7BCAFDD0" box="[698,991,584,608]" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier 2012
</taxonomicName>
”, 3 3 2 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBFFD0A4E7936FD34" box="[189,291,620,644]" pageId="4" pageNumber="85">IADIZA</collectionCode>
), 2 3 2 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBEED0A4E7A1CFD34" LSID="urn:lsid:biocol.org:col:34988" box="[429,521,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
), 2 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBDD20A4E7B15FD34" box="[658,768,620,644]" pageId="4" pageNumber="85">MNHUB</collectionCode>
), 2 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBCCA0A4E7BC2FD34" box="[906,983,620,644]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="85">NMW</collectionCode>
), 2 3 1 Ƥ (
<collectionCode id="8EA48EA9FF92FF9BBB200A4E7CDBFD34" box="[1120,1230,620,644]" pageId="4" pageNumber="85">UNALM</collectionCode>
);“ 
<collectingCountry id="90A256FCFF92FF9BBBA40A4E7D3CFD34" box="[1252,1321,620,644]" name="Brazil" pageId="4" pageNumber="85">Brésil</collectingCountry>
– Amaz./ Manaus/ 
<date id="9C0B30ACFF92FF9BBFBF0AAD7965FD17" box="[255,368,655,679]" pageId="4" pageNumber="85" value="1996-05-19">19.V.1996</date>
/ G. Couturier &amp; F. Kahn”, “Km. 130/ Br134/ Rodovia Boa Vista”, “ 
<taxonomicName id="2FB56DEFFF92FF9BBB390AB37D89FD17" authority="Palmae" authorityName="Palmae" box="[1145,1436,655,680]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBB390AB37D54FD18" box="[1145,1345,656,680]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBB390AB37CC8FD18" box="[1145,1245,657,680]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
Palmae
</taxonomicName>
sur inflorescence mâle”, “ 
<typeStatus id="370EA8CEFF92FF9BBEFD0A967A34FD7C" box="[445,545,692,716]" pageId="4" pageNumber="85" type="paratype">Paratype</typeStatus>
<taxonomicName id="2FB56DEFFF92FF9BBD6A0A967CA9FD7C" authority="Chani-Posse &amp; Couturier 2012" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[554,1212,692,716]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBD6A0A967B44FD7C" box="[554,849,692,716]" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
Chani-Posse &amp; Couturier 2012
</taxonomicName>
” 1 3, (
<collectionCode id="8EA48EA9FF92FF9BBA520A967D6CFD7C" box="[1298,1401,692,716]" pageId="4" pageNumber="85">IADIZA</collectionCode>
), 3 3, (
<collectionCode id="8EA48EA9FF92FF9BBF820AF57912FD5F" box="[194,263,727,751]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="4" pageNumber="85">INPA</collectionCode>
), 1 3, (
<collectionCode id="8EA48EA9FF92FF9BBE190AF579A0FD5F" LSID="urn:lsid:biocol.org:col:34988" box="[345,437,727,751]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="85">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A0AF45E7FF92FF98BF870AD97BA5FE8F" lastPageId="7" lastPageNumber="88" pageId="4" pageNumber="85" type="distribution">
<paragraph id="E80A166CFF92FF9BBF870AD97967FCCF" blockId="4.[151,1436,151,1977]" pageId="4" pageNumber="85">
<emphasis id="DAC1CA7EFF92FF9BBF870AD979EAFCA4" bold="true" box="[199,511,763,788]" pageId="4" pageNumber="85">Distribution and habitat.</emphasis>
<taxonomicName id="2FB56DEFFF92FF9BBD4C0ADE7B2DFCA4" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[524,824,764,788]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBD4C0ADE7B2DFCA4" box="[524,824,764,788]" italics="true" pageId="4" pageNumber="85">Delgadobius amazonensis</emphasis>
</taxonomicName>
has been recorded from 
<collectingCountry id="90A256FCFF92FF9BBB310ADF7CB1FCA4" box="[1137,1188,765,788]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
(Iquitos) and 
<collectingCountry id="90A256FCFF92FF9BBA170ADE7D8EFCA4" box="[1367,1435,764,788]" name="Brazil" pageId="4" pageNumber="85">Brazil</collectingCountry>
(Manaus). It was found in association with palm trees (
<taxonomicName id="2FB56DEFFF92FF9BBC650B3D7B88FC87" box="[805,925,799,823]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
) during anthesis, on male inflorescences of 
<taxonomicName id="2FB56DEFFF92FF9BBFD70B677974FCEC" box="[151,353,836,860]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBFD70B677974FCEC" box="[151,353,836,860]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBFD70B6778EEFCEC" box="[151,251,837,860]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
</taxonomicName>
L. f., 
<taxonomicName id="2FB56DEFFF92FF9BBEEF0B677AD8FCEC" authority="Wallace" authorityName="Wallace" box="[431,717,836,860]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="carana">
<emphasis id="DAC1CA7EFF92FF9BBEEF0B677A7EFCEC" box="[431,619,837,860]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBEEF0B677A06FCEC" box="[431,531,837,860]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
carana
</emphasis>
Wallace
</taxonomicName>
, 
<taxonomicName id="2FB56DEFFF92FF9BBDA00B677C34FCEC" authority="Martius" authorityName="Martius" box="[736,1057,836,860]" class="Liliopsida" family="Arecaceae" genus="Oenocarpus" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="bataua">
<emphasis id="DAC1CA7EFF92FF9BBDA00B677BD5FCEC" box="[736,960,836,860]" italics="true" pageId="4" pageNumber="85">Oenocarpus bataua</emphasis>
Martius
</taxonomicName>
<emphasis id="DAC1CA7EFF92FF9BBB6B0B667D12FCEC" box="[1067,1287,836,860]" italics="true" pageId="4" pageNumber="85">
(= 
<taxonomicName id="2FB56DEFFF92FF9BBB030B667CEAFCEC" box="[1091,1279,836,860]" class="Liliopsida" family="Arecaceae" genus="Jessenia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="bataua">Jessenia bataua</taxonomicName>
)
</emphasis>
and 
<taxonomicName id="2FB56DEFFF92FF9BBA030B677967FCCF" authority="Martius." authorityName="Martius." class="Insecta" family="Nymphalidae" genus="Euterpe" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="precatoria">
<emphasis id="DAC1CA7EFF92FF9BBA030B677919FC30" italics="true" pageId="4" pageNumber="85">Euterpe precatoria</emphasis>
Martius.
</taxonomicName>
</paragraph>
<paragraph id="E80A166CFF92FF9BBF870BAE7A95F808" blockId="4.[151,1436,151,1977]" pageId="4" pageNumber="85">
The occurrence of the new species in 
<taxonomicName id="2FB56DEFFF92FF9BBD300BAF7B2DFC14" box="[624,824,908,932]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBD300BAF7B2DFC14" box="[624,824,908,932]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBD300BAF7AC1FC14" box="[624,724,909,932]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
</taxonomicName>
was reported from two localities in 
<collectingCountry id="90A256FCFF92FF9BBB910BAF7D11FC14" box="[1233,1284,909,932]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
(Iquitos) and 
<collectingCountry id="90A256FCFF92FF9BBFD70B8D78CEFC77" box="[151,219,943,967]" name="Brazil" pageId="4" pageNumber="85">Brazil</collectingCountry>
(Manaus). Given this association, it seems probable that both the insect and the host plant may share a common distribution. 
<taxonomicName id="2FB56DEFFF92FF9BBED10BF77A4CFC5C" box="[401,601,980,1004]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBED10BF77A4CFC5C" box="[401,601,980,1004]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBED10BF779E0FC5C" box="[401,501,981,1004]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
</taxonomicName>
is an Amazonian native palm species (
<figureCitation id="708E0AE9FF92FF9BBB550BF67C8BFC5C" box="[1045,1182,980,1004]" captionStart="FIGURES 14 – 21" captionStartId="6.[151,264,1836,1859]" captionTargetBox="[304,1291,168,1816]" captionTargetId="figure@6.[304,1292,167,1817]" captionTargetPageId="6" captionText="FIGURES 14 – 21. Mauritia flexuosa: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits." figureDoi="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="4" pageNumber="85">Figs. 14–21</figureCitation>
), that is distributed all over the American tropics and the East Andes, mainly in the Amazonian basin, in areas such as 
<collectingCountry id="90A256FCFF92FF9BBBBE0BDB7D21FBA0" box="[1278,1332,1017,1040]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
, 
<collectingCountry id="90A256FCFF92FF9BBA020BD57D82FBBF" box="[1346,1431,1015,1039]" name="Bolivia" pageId="4" pageNumber="85">Bolivia</collectingCountry>
, 
<collectingCountry id="90A256FCFF92FF9BBFD70C3E791DFB84" box="[151,264,1052,1076]" name="Colombia" pageId="4" pageNumber="85">Colombia</collectingCountry>
, 
<collectingCountry id="90A256FCFF92FF9BBE540C3E7966FB84" box="[276,371,1052,1076]" name="Ecuador" pageId="4" pageNumber="85">Ecuador</collectingCountry>
, 
<collectingCountry id="90A256FCFF92FF9BBE3F0C3E79E0FB84" box="[383,501,1052,1076]" name="Venezuela" pageId="4" pageNumber="85">Venezuela</collectingCountry>
, 
<collectingCountry id="90A256FCFF92FF9BBD410C3E7A5DFB84" box="[513,584,1052,1076]" name="Brazil" pageId="4" pageNumber="85">Brazil</collectingCountry>
, the Guyanas, north of 
<collectingCountry id="90A256FCFF92FF9BBC100C3E7BA5FB84" box="[848,944,1052,1076]" name="Trinidad and Tobago" pageId="4" pageNumber="85">Trinidad</collectingCountry>
and 
<collectingCountry id="90A256FCFF92FF9BBCA80C3F7C54FB84" box="[1000,1089,1053,1076]" name="Panama" pageId="4" pageNumber="85">Panama</collectingCountry>
(
<bibRefCitation id="8C246B9DFF92FF9BBB100C3E7D43FB84" author="Henderson" box="[1104,1366,1052,1076]" pageId="4" pageNumber="85" refString="Henderson, A. H., Galeano, G. &amp; Bernal, R. (1995) Field guide to the palms of the Americas. Princeton University Press, Princeton, N. J." type="book" year="1995">
Henderson 
<emphasis id="DAC1CA7EFF92FF9BBB930C3F7D07FB84" box="[1235,1298,1052,1076]" italics="true" pageId="4" pageNumber="85">et al.,</emphasis>
1995
</bibRefCitation>
). This palm species plays a major role in the economic, social and ecological systems of the Peruvian Amazonia (
<bibRefCitation id="8C246B9DFF92FF9BBA7A0C1D790FFBCC" author="Vasquez" pageId="4" pageNumber="85" refString="Vasquez, J., Delgado, C., Couturier, G., Mejia, K., Freitas, L., Del Castillo, D. (2008) Pest insects of the palm tree Mauritia flexuosa L. f., dwarf form, in Peruvian Amazonia. Fruits, 63 (4), 227 - 238." type="journal article" year="2008">
Vásquez 
<emphasis id="DAC1CA7EFF92FF9BBFD70C4778CDFBCC" box="[151,216,1124,1148]" italics="true" pageId="4" pageNumber="85">et al.,</emphasis>
2008
</bibRefCitation>
, 
<bibRefCitation id="8C246B9DFF92FF9BBE680C467A6BFBCC" author="Trujillo-Gonzalez" box="[296,638,1124,1148]" pageId="4" pageNumber="85" refString="Trujillo-Gonzalez, J. M., Torres-Mora, M. A. &amp; Santana-Castaneda, E. (2011) La palma de Moriche (Mauritia flexuosa L. f) un ecosistema estrategico. Orinoquia, 15 (1), 62 - 70" type="journal article" year="2011">
Trujillo-González 
<emphasis id="DAC1CA7EFF92FF9BBEB80C477A3EFBCC" box="[504,555,1124,1148]" italics="true" pageId="4" pageNumber="85">et al</emphasis>
., 2011
</bibRefCitation>
). Because of its importance and in order to avoid its overexploitation, this plant is in process of domestication to be cultivated as a dwarf form (
<bibRefCitation id="8C246B9DFF92FF9BBCB80CA57CFBFB2F" author="Vasquez" box="[1016,1262,1159,1184]" pageId="4" pageNumber="85" refString="Vasquez, J., Delgado, C., Couturier, G., Mejia, K., Freitas, L., Del Castillo, D. (2008) Pest insects of the palm tree Mauritia flexuosa L. f., dwarf form, in Peruvian Amazonia. Fruits, 63 (4), 227 - 238." type="journal article" year="2008">
Vásquez 
<emphasis id="DAC1CA7EFF92FF9BBB240CAB7CB3FB10" box="[1124,1190,1160,1184]" italics="true" pageId="4" pageNumber="85">et al.,</emphasis>
2008
</bibRefCitation>
). However, the development of the crops involves the proliferation of harmful insects and knowledge about potential natural enemies is required. 
<taxonomicName id="2FB56DEFFF92FF9BBECA0CF37A40FB58" box="[394,597,1232,1256]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBECA0CF37A40FB58" box="[394,597,1232,1256]" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBECA0CF379FBFB58" box="[394,494,1233,1256]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
</taxonomicName>
grows naturally on flooded soils, on rivers and stream borders forming dense monospecific gatherings known in 
<collectingCountry id="90A256FCFF92FF9BBD2F0CD77AB7FABC" box="[623,674,1269,1292]" name="Peru" pageId="4" pageNumber="85">Peru</collectingCountry>
as “aguajales” (
<figureCitation id="708E0AE9FF92FF9BBC1C0CD67BFDFABC" box="[860,1000,1268,1292]" captionStart="FIGURES 14 – 21" captionStartId="6.[151,264,1836,1859]" captionTargetBox="[304,1291,168,1816]" captionTargetId="figure@6.[304,1292,167,1817]" captionTargetPageId="6" captionText="FIGURES 14 – 21. Mauritia flexuosa: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits." figureDoi="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="4" pageNumber="85">Figs. 14, 15</figureCitation>
). When they are associated with other palm species such as 
<taxonomicName id="2FB56DEFFF92FF9BBECC0D3A7A79FA80" box="[396,620,1304,1328]" class="Liliopsida" family="Arecaceae" genus="Oenocarpus" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="bataua">
<emphasis id="DAC1CA7EFF92FF9BBECC0D3A7A79FA80" box="[396,620,1304,1328]" italics="true" pageId="4" pageNumber="85">Oenocarpus bataua</emphasis>
</taxonomicName>
(as it is also the case here for the occurrence of 
<taxonomicName id="2FB56DEFFF92FF9BBBD90D3A7D3DFA80" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[1177,1320,1304,1328]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="DAC1CA7EFF92FF9BBBD90D3A7D3DFA80" box="[1177,1320,1304,1328]" italics="true" pageId="4" pageNumber="85">Delgadobius</emphasis>
</taxonomicName>
), they are called “sacha-aguajales” (
<bibRefCitation id="8C246B9DFF92FF9BBEF70D1E7A8AFAE4" author="Delgado" box="[439,671,1340,1364]" pageId="4" pageNumber="85" refString="Delgado C., Couturier G., Mejia K. (2007) Mauritia flexuosa (Arecaceae: Calamoideae), an Amazonian palm with cultivation purposes in Peru. Fruits, 62 (39), 157 - 169." type="journal article" year="2007">
Delgado 
<emphasis id="DAC1CA7EFF92FF9BBD5D0D1F7A49FAE4" box="[541,604,1340,1364]" italics="true" pageId="4" pageNumber="85">et al.,</emphasis>
2007
</bibRefCitation>
). In 
<taxonomicName id="2FB56DEFFF92FF9BBD8F0D1F7B44FAE4" box="[719,849,1340,1364]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBD8F0D1F7B44FAE4" box="[719,849,1340,1364]" italics="true" pageId="4" pageNumber="85">M. flexuosa</emphasis>
</taxonomicName>
, the new species has been observed actively running among the rachillae (
<figureCitation id="708E0AE9FF92FF9BBECF0D427A0FFAC8" box="[399,538,1376,1400]" captionStart="FIGURES 14 – 21" captionStartId="6.[151,264,1836,1859]" captionTargetBox="[304,1291,168,1816]" captionTargetId="figure@6.[304,1292,167,1817]" captionTargetPageId="6" captionText="FIGURES 14 – 21. Mauritia flexuosa: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits." figureDoi="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="4" pageNumber="85">Figs. 17–19</figureCitation>
) of the palm together with adults and larvae of 
<taxonomicName id="2FB56DEFFF92FF9BBB160D437D89FAC8" authority="Grouvelle" authorityName="Grouvelle" box="[1110,1436,1376,1400]" class="Insecta" family="Nitidulidae" genus="Mystrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="dalmasi">
<emphasis id="DAC1CA7EFF92FF9BBB160D437D35FAC8" box="[1110,1312,1376,1400]" italics="true" pageId="4" pageNumber="85">Mystrops dalmasi</emphasis>
Grouvelle
</taxonomicName>
(
<taxonomicName id="2FB56DEFFF92FF9BBFDE0DA6790AFA2C" box="[158,287,1412,1436]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
: 
<taxonomicName id="2FB56DEFFF92FF9BBE6C0DA679BAFA2C" box="[300,431,1412,1436]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
), which is quite abundant (Couturier, pers. obs.) and may be potential prey. 
<taxonomicName id="2FB56DEFFF92FF9BBA5E0DA67D89FA2C" box="[1310,1436,1412,1436]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
has also been reported as the most abundant family visiting male inflorescences of 
<taxonomicName id="2FB56DEFFF92FF9BBB7C0D8B7CAAFA70" box="[1084,1215,1448,1472]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBB7C0D8B7CAAFA70" box="[1084,1215,1448,1472]" italics="true" pageId="4" pageNumber="85">M. flexuosa</emphasis>
</taxonomicName>
among 
<taxonomicName id="2FB56DEFFF92FF9BBA5A0D8A7D8DFA70" box="[1306,1432,1448,1472]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
, which in turn has shown the highest representation among the insect orders associated with this palm species in Manaus, 
<collectingCountry id="90A256FCFF92FF9BBFBF0DD27956F9B8" box="[255,323,1520,1544]" name="Brazil" pageId="4" pageNumber="85">Brazil</collectingCountry>
(
<bibRefCitation id="8C246B9DFF92FF9BBE130DD279CBF9B8" author="Storti" box="[339,478,1520,1544]" pageId="4" pageNumber="85" refString="Storti E. (1993) Biologia floral de Mauritia flexuosa L. f. na Regiao de Manaus, AM, Brasil. Acta Amazonica, 23 (4), 371 - 381." type="journal article" year="1993">Storti, 1993</bibRefCitation>
). The presence of 
<taxonomicName id="2FB56DEFFF92FF9BBDF00DD37B76F9B8" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[688,867,1521,1544]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF92FF9BBDF00DD37B76F9B8" box="[688,867,1521,1544]" italics="true" pageId="4" pageNumber="85">D. amazonensis</emphasis>
</taxonomicName>
in 
<taxonomicName id="2FB56DEFFF92FF9BBCCA0DD37C18F9B8" box="[906,1037,1520,1544]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBCCA0DD37C18F9B8" box="[906,1037,1520,1544]" italics="true" pageId="4" pageNumber="85">M. flexuosa</emphasis>
</taxonomicName>
has only been observed during the flowering event, at the end of which the insect disappears (G. Couturier, pers. obs.). The host plant, 
<taxonomicName id="2FB56DEFFF92FF9BBA780E3778E1F9E0" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBA780E3778E1F9E0" italics="true" pageId="4" pageNumber="85">
<collectingCountry id="90A256FCFF92FF9BBA780E377D89F99C" box="[1336,1436,1557,1580]" name="Mauritius" pageId="4" pageNumber="85">Mauritia</collectingCountry>
flexuosa
</emphasis>
</taxonomicName>
or “aguaje” (
<figureCitation id="708E0AE9FF92FF9BBECE0E1A7A1EF9E0" box="[398,523,1592,1616]" captionStart="FIGURES 14 – 21" captionStartId="6.[151,264,1836,1859]" captionTargetBox="[304,1291,168,1816]" captionTargetId="figure@6.[304,1292,167,1817]" captionTargetPageId="6" captionText="FIGURES 14 – 21. Mauritia flexuosa: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits." figureDoi="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="4" pageNumber="85">Fig. 14, 15</figureCitation>
) as it is called in the Peruvian Amazonia (“buriti” or “miriti” in 
<collectingCountry id="90A256FCFF92FF9BBBA90E1A7D3AF9E0" box="[1257,1327,1592,1616]" name="Brazil" pageId="4" pageNumber="85">Brazil</collectingCountry>
) starts its commercial fructification at the age of 12–15 years when it naturally grows. Given its height (
<quantity id="2F4DBB89FF92FF9BBBEB0E7E7D07F9C4" box="[1195,1298,1628,1652]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="4" pageNumber="85" unit="m" value="37.5" valueMax="40.0" valueMin="35.0">35–40 m</quantity>
), it has been a common practice to cut the whole female individual in spite of the detrimental effects on the palm-tree populations (G. Couturier, pers. com.). The dwarf form or “aguaje enano” (
<figureCitation id="708E0AE9FF92FF9BBCAC0E867C2AF90C" box="[1004,1087,1700,1724]" captionStart="FIGURES 14 – 21" captionStartId="6.[151,264,1836,1859]" captionTargetBox="[304,1291,168,1816]" captionTargetId="figure@6.[304,1292,167,1817]" captionTargetPageId="6" captionText="FIGURES 14 – 21. Mauritia flexuosa: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits." figureDoi="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="4" pageNumber="85">Fig. 14</figureCitation>
) has not been found in natural conditions but only in culture. Its fructification happens at the age of 5 years, before the stem is less than 
<quantity id="2F4DBB89FF92FF9BBA710EEA7D49F950" box="[1329,1372,1736,1760]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="85" unit="m" value="1.0">1 m</quantity>
high. The precocity of its fructification and its short stem allows easy collection of both the fruit and the flower-visiting insects (
<figureCitation id="708E0AE9FF92FF9BBFB30F327967F898" box="[243,370,1808,1832]" captionStart="FIGURES 14 – 21" captionStartId="6.[151,264,1836,1859]" captionTargetBox="[304,1291,168,1816]" captionTargetId="figure@6.[304,1292,167,1817]" captionTargetPageId="6" captionText="FIGURES 14 – 21. Mauritia flexuosa: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits." figureDoi="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="4" pageNumber="85">Fig. 18, 21</figureCitation>
). Flowering can occur during the whole year. The anthesis of female and male flowers usually begins at 16 h and lasts an average of 24 h. When the flowers open, they produce a fragrance that may act as a mechanism for attracting insects (
<bibRefCitation id="8C246B9DFF92FF9BBD580F7A7B1CF8C0" author="Delgado" box="[536,777,1880,1904]" pageId="4" pageNumber="85" refString="Delgado C., Couturier G., Mejia K. (2007) Mauritia flexuosa (Arecaceae: Calamoideae), an Amazonian palm with cultivation purposes in Peru. Fruits, 62 (39), 157 - 169." type="journal article" year="2007">
Delgado 
<emphasis id="DAC1CA7EFF92FF9BBDC20F7B7AD6F8C0" box="[642,707,1880,1904]" italics="true" pageId="4" pageNumber="85">et al.,</emphasis>
2007
</bibRefCitation>
). A great amount of insects associated with this palm has been reported from male and female inflorescences during anthesis (
<bibRefCitation id="8C246B9DFF92FF9BBCD70F5E7C35F824" author="Storti" box="[919,1056,1916,1940]" pageId="4" pageNumber="85" refString="Storti E. (1993) Biologia floral de Mauritia flexuosa L. f. na Regiao de Manaus, AM, Brasil. Acta Amazonica, 23 (4), 371 - 381." type="journal article" year="1993">Storti, 1993</bibRefCitation>
). Since 
<taxonomicName id="2FB56DEFFF92FF9BBB390F5F7CEEF824" box="[1145,1275,1916,1940]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="85" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF92FF9BBB390F5F7CEEF824" box="[1145,1275,1916,1940]" italics="true" pageId="4" pageNumber="85">M. flexuosa</emphasis>
</taxonomicName>
is a dioecious palm, pollination by insects may be crucial.
</paragraph>
<caption id="BCCA46E4FF93FF9ABFD70F4179D7F844" ID-DOI="http://doi.org/10.5281/zenodo.211552" httpUri="https://zenodo.org/record/211552/files/figure.png" pageId="5" pageNumber="86" targetBox="[233,1358,196,1856]" targetPageId="5">
<paragraph id="E80A166CFF93FF9ABFD70F4179D7F844" blockId="5.[151,1436,1891,2036]" pageId="5" pageNumber="86">
<emphasis id="DAC1CA7EFF93FF9ABFD70F417958F8C8" bold="true" box="[151,333,1891,1914]" pageId="5" pageNumber="86">FIGURES 1–13.</emphasis>
<taxonomicName id="2FB56DEFFF93FF9ABE1A0F417A7FF8C9" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[346,618,1891,1913]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF93FF9ABE1A0F417A7FF8C9" box="[346,618,1891,1913]" italics="true" pageId="5" pageNumber="86">Delgadobius amazonensis</emphasis>
</taxonomicName>
: 1, habitus; 2, mouthparts; 3, mesonotum, metanotum and first abdominal segments; 4, head, ventral view (IoR= infraorbital ridge); 5, prosternum and hypomeron (aL= additional line, Pr= pronotal ridge); 6, pronotum and hypomeron, lateral view; 7, front tarsus, ventral view (S=segment); 8, mid tarsus, lateroventral view; 9, tergum 10 (male); 10, sternum 9; 111, aedeagus, dorsal view; 12, aedeagus, lateral view; 13, gonocoxites of female genital segment. Scale bars=0.4 mm.
</paragraph>
</caption>
<caption id="BCCA46E4FF90FF99BFD70F0E7D0FF8CF" ID-DOI="http://doi.org/10.5281/zenodo.211553" httpUri="https://zenodo.org/record/211553/files/figure.png" pageId="6" pageNumber="87" targetBox="[304,1291,168,1816]" targetPageId="6">
<paragraph id="E80A166CFF90FF99BFD70F0E7D0FF8CF" blockId="6.[151,1436,1836,1919]" pageId="6" pageNumber="87">
<emphasis id="DAC1CA7EFF90FF99BFD70F0E7946F8F1" bold="true" box="[151,339,1836,1859]" pageId="6" pageNumber="87">FIGURES 14–21.</emphasis>
<taxonomicName id="2FB56DEFFF90FF99BE190F0F7A18F8F2" box="[345,525,1836,1858]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="87" phylum="Tracheophyta" rank="species" species="flexuosa">
<emphasis id="DAC1CA7EFF90FF99BE190F0F7A18F8F2" box="[345,525,1836,1858]" italics="true" pageId="6" pageNumber="87">Mauritia flexuosa</emphasis>
</taxonomicName>
: 14, dwarf form; 15, palms in human-settled area; 16, rachillae before anthesis (male); 17, portion of rachilla at the beginning of anthesis (male); 18, Guy Couturier harvesting rachillae for insect sampling; 19, male inflorescence; 20, portion of rachilla during anthesis (female); 21, César Delgado next to a dwarf form with fruits.
</paragraph>
</caption>
<paragraph id="E80A166CFF90FF98BF870F8F7BA5FE8F" blockId="6.[151,1436,1964,2025]" lastBlockId="7.[151,1436,151,320]" lastPageId="7" lastPageNumber="88" pageId="6" pageNumber="87">
A common feature among the genera of palm trees from which 
<taxonomicName id="2FB56DEFFF90FF99BCDA0F8F7C59F874" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[922,1100,1965,1988]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF90FF99BCDA0F8F7C59F874" box="[922,1100,1965,1988]" italics="true" pageId="6" pageNumber="87">D. amazonensis</emphasis>
</taxonomicName>
has been recorded (
<emphasis id="DAC1CA7EFF90FF99BA710F8F78E5F858" italics="true" pageId="6" pageNumber="87">
<taxonomicName id="2FB56DEFFF90FF99BA710F8F7D82F874" box="[1329,1431,1965,1988]" class="Liliopsida" family="Arecaceae" genus="Mauritia" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="87" phylum="Tracheophyta" rank="genus">
<collectingCountry id="90A256FCFF90FF99BA710F8F7D82F874" box="[1329,1431,1965,1988]" name="Mauritius" pageId="6" pageNumber="87">Mauritia</collectingCountry>
</taxonomicName>
, 
<taxonomicName id="2FB56DEFFF90FF99BFD70FF378E5F858" box="[151,240,2001,2024]" class="Insecta" family="Nymphalidae" genus="Euterpe" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="genus">Euterpe</taxonomicName>
</emphasis>
and 
<taxonomicName id="2FB56DEFFF90FF99BE740FF379A8F858" box="[308,445,2001,2024]" class="Liliopsida" family="Arecaceae" genus="Oenocarpus" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="87" phylum="Tracheophyta" rank="genus">
<emphasis id="DAC1CA7EFF90FF99BE740FF379A8F858" box="[308,445,2001,2024]" italics="true" pageId="6" pageNumber="87">Oenocarpus</emphasis>
</taxonomicName>
) is the rather sparse structure of their inflorescence compared to other genera of 
<taxonomicName id="2FB56DEFFF91FF98BFD708BA791BFF00" box="[151,270,152,176]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="7" pageNumber="88" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
, such as 
<emphasis id="DAC1CA7EFF91FF98BE3008BA7A46FF00" box="[368,595,152,176]" italics="true" pageId="7" pageNumber="88">
<taxonomicName id="2FB56DEFFF91FF98BE3008BA7A16FF00" box="[368,515,152,176]" class="Liliopsida" family="Arecaceae" genus="Astrocaryum" kingdom="Plantae" order="Arecales" pageId="7" pageNumber="88" phylum="Tracheophyta" rank="genus">Astrocaryum</taxonomicName>
, 
<taxonomicName id="2FB56DEFFF91FF98BD4D08BA7A46FF00" box="[525,595,152,176]" class="Liliopsida" family="Arecaceae" genus="Elaeis" kingdom="Plantae" order="Arecales" pageId="7" pageNumber="88" phylum="Tracheophyta" rank="genus">Elaeis</taxonomicName>
</emphasis>
and 
<taxonomicName id="2FB56DEFFF91FF98BDC908BA7B07FF00" box="[649,786,152,176]" class="Liliopsida" family="Arecaceae" genus="Phytelephas" kingdom="Plantae" order="Arecales" pageId="7" pageNumber="88" phylum="Tracheophyta" rank="genus">
<emphasis id="DAC1CA7EFF91FF98BDC908BA7B07FF00" box="[649,786,152,176]" italics="true" pageId="7" pageNumber="88">Phytelephas</emphasis>
</taxonomicName>
. These genera have been also examined in the field but 
<taxonomicName id="2FB56DEFFF91FF98BAC108BB7933FF64" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF91FF98BAC108BB7933FF64" italics="true" pageId="7" pageNumber="88">D. amazonensis</emphasis>
</taxonomicName>
was not either observed or reported. We assume that the structure of the inflorescence may be a limiting factor in the predatory behavior of 
<taxonomicName id="2FB56DEFFF91FF98BDDB08C37B45FF48" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[667,848,225,248]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF91FF98BDDB08C37B45FF48" box="[667,848,225,248]" italics="true" pageId="7" pageNumber="88">D. amazonensis</emphasis>
</taxonomicName>
. Because of its association with a dominant plant species of economic and cultural importance in Amazonia, the ecological role of 
<taxonomicName id="2FB56DEFFF91FF98BB7009277CF7FEAC" authorityName="Chani-Posse &amp; Couturier" authorityYear="2012" box="[1072,1250,261,284]" class="Insecta" family="Staphylinidae" genus="Delgadobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="amazonensis">
<emphasis id="DAC1CA7EFF91FF98BB7009277CF7FEAC" box="[1072,1250,261,284]" italics="true" pageId="7" pageNumber="88">D. amazonensis</emphasis>
</taxonomicName>
in the “aguajal” ecosystem may deserve further studies of applied interest in the future.
</paragraph>
</subSubSection>
</treatment>
</document>