<documentid="4A3595CC73646A2BF3BEEF2F895F8B27"ID-CLB-Dataset="304902"ID-DOI="10.5852/ejt.2024.966.2697"ID-GBIF-Dataset="0bf71223-cd52-42bc-8bce-edad65412211"ID-ISSN="2118-9773"ID-Zenodo-Dep="14001790"ID-ZooBank="FFE41347-8B55-4CF1-B10B-53EBB4CF0E26"IM.bibliography_approvedBy="carolina"IM.illustrations_approvedBy="carolina"IM.materialsCitations_approvedBy="carolina"IM.metadata_approvedBy="carolina"IM.tables_approvedBy="carolina"IM.taxonomicNames_approvedBy="felipe"IM.treatmentCitations_approvedBy="carolina"IM.treatments_approvedBy="carolina"checkinTime="1730118416961"checkinUser="plazi"docAuthor="Henrard, Arnaud, Griswold, Charles & Jocqué, Rudy"docDate="2024"docId="3157E00AFFEFFFD6FDF60E9FFA8ED8E6"docLanguage="en"docName="EJT.2024.966.1-80.pdf"docOrigin="European Journal of Taxonomy 966 (1)"docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2697/12453"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Tabiboka milloti Henrard, Griswold & Jocqué, 2024, gen. et sp. nov."docType="treatment"docUuid="FF163E94-73C4-4D75-AC93-1BEB4FCA536F"docUuidSource="ZooBank"docVersion="8"lastPageNumber="29"masterDocId="CD6E9872FFF8FFCAFFAA0824FFD5DF0E"masterDocTitle="On new genera and species of crack-leg spiders (Araneae, Udubidae) from Madagascar"masterLastPageNumber="80"masterPageNumber="1"pageNumber="24"updateTime="1730317898278"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:affiliationid="B12CECB31F7CBB9C19AA05085D8F9FF2">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
<mods:affiliationid="0352BD36A3760EE9AA475E3B670E5DA5">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA.</mods:affiliation>
<mods:affiliationid="15660A7424936E2701D18AB3B57C9D8A">Royal Museum for Central Africa, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium.</mods:affiliation>
<taxonomicNameid="7EFE2A9FFFEFFFDDFDF60E9FFCF5D9D8"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[604,800,1723,1750]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="milloti"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFEFFFDDFC820E98FC0DD9D8"box="[808,984,1724,1750]"pageId="23"pageNumber="24"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="21C54D99FFEFFFDDFD7B0EDBFCECD817"box="[721,825,1791,1817]"captionStart-0="Fig"captionStart-1="Fig"captionStartId-0="25.[189,232,1717,1743]"captionStartId-1="27.[189,231,1595,1621]"captionTargetBox-1="[260,1379,284,1517]"captionTargetId-0="figure-19@26.[189,1398,304,1979]"captionTargetId-1="figure-11@27.[189,1398,265,1558]"captionTargetPageId-0="26"captionTargetPageId-1="27"captionText-0="Fig. 8(on next page). Tabiboka milloti gen. nov. et sp. nov. male holotype (CASENT_9016063; MNHN). A. Habitus, dorsal view. B. Carapace, dorsal view. C. SEM of right male palp (mirrored), prolateral view. D. Idem, ventral view. E. Idem, retrolateral view. F. Idem, detail of bulb, ventral view. G. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = 4.6 mm); C–E = 0.5 mm; F–G = 0.1 mm. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick."captionText-1="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold."figureDoi-0="http://doi.org/10.5281/zenodo.14014722"figureDoi-1="http://doi.org/10.5281/zenodo.14001810"httpUri-0="https://zenodo.org/record/14014722/files/figure.png"httpUri-1="https://zenodo.org/record/14001810/files/figure.png"pageId="23"pageNumber="24">Figs 8–9</figureCitation>
<treatmentCitationid="385F770DFFEFFFDDFE740F1BFD1CD857"author="Wheeler W. C. & Coddington J. A. & Crowley L. M. & Dimitrov D. & Goloboff P. A. & Griswold C. E. & Hormiga G. & Prendini L. & Ramirez M. J. & Sierwald P. & Almeida-Silva L. M. & Alvarez-Padilla F. & Arnedo M. A. & Benavides L. R. & Benjamin S. P. & Bond J. E. & Grismado C. J. & Hasan E. & Hedin M. & Izquierdo M. A. & Labarque F. M. & Ledford J. & Lopardo L. & Maddison W. P. & Miller J. A. & Piacentini L. N. & Platnick N. I. & Polotow D. & Silva-Davila D. & Scharff N. & Szuts T. & Ubick D. & Vink C. & Wood H. M. & Zhang J. X."box="[478,713,1855,1881]"pageId="23"pageNumber="24"year="2017">
<bibRefCitationid="DD6F2CEDFFEFFFDDFE740F1BFD1CD857"author="Wheeler W. C. & Coddington J. A. & Crowley L. M. & Dimitrov D. & Goloboff P. A. & Griswold C. E. & Hormiga G. & Prendini L. & Ramirez M. J. & Sierwald P. & Almeida-Silva L. M. & Alvarez-Padilla F. & Arnedo M. A. & Benavides L. R. & Benjamin S. P. & Bond J. E. & Grismado C. J. & Hasan E. & Hedin M. & Izquierdo M. A. & Labarque F. M. & Ledford J. & Lopardo L. & Maddison W. P. & Miller J. A. & Piacentini L. N. & Platnick N. I. & Polotow D. & Silva-Davila D. & Scharff N. & Szuts T. & Ubick D. & Vink C. & Wood H. M. & Zhang J. X."box="[478,713,1855,1881]"pageId="23"pageNumber="24"pagination="576 - 616"refId="ref39136"refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. & Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182"type="journal article"year="2017">
<taxonomicNameid="7EFE2A9FFFEFFFDDFEF30F88FDCDD8C8"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[345,536,1964,1990]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="milloti"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFEFFFDDFDB50F89FD19D8C9"box="[543,716,1965,1991]"pageId="23"pageNumber="24"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="21C54D99FFEFFFDDFB200FEAFAC4D8E6"box="[1162,1297,1998,2024]"captionStart="Fig"captionStartId="25.[189,232,1717,1743]"captionTargetId="figure-19@26.[189,1398,304,1979]"captionTargetPageId="26"captionText="Fig. 8(on next page). Tabiboka milloti gen. nov. et sp. nov. male holotype (CASENT_9016063; MNHN). A. Habitus, dorsal view. B. Carapace, dorsal view. C. SEM of right male palp (mirrored), prolateral view. D. Idem, ventral view. E. Idem, retrolateral view. F. Idem, detail of bulb, ventral view. G. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = 4.6 mm); C–E = 0.5 mm; F–G = 0.1 mm. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14014722"httpUri="https://zenodo.org/record/14014722/files/figure.png"pageId="23"pageNumber="24">Figs 8E, G</figureCitation>
<figureCitationid="21C54D99FFE0FFD2FED20909FE06DE46"box="[376,467,301,328]"captionStart="Fig"captionStartId="25.[189,232,1717,1743]"captionTargetId="figure-19@26.[189,1398,304,1979]"captionTargetPageId="26"captionText="Fig. 8(on next page). Tabiboka milloti gen. nov. et sp. nov. male holotype (CASENT_9016063; MNHN). A. Habitus, dorsal view. B. Carapace, dorsal view. C. SEM of right male palp (mirrored), prolateral view. D. Idem, ventral view. E. Idem, retrolateral view. F. Idem, detail of bulb, ventral view. G. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = 4.6 mm); C–E = 0.5 mm; F–G = 0.1 mm. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14014722"httpUri="https://zenodo.org/record/14014722/files/figure.png"pageId="24"pageNumber="25">Figs 8F</figureCitation>
<taxonomicNameid="7EFE2A9FFFE0FFD2FD28090AFD38DE49"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[642,749,301,327]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="24"pageNumber="25"phylum="Arthropoda"rank="species"species="milleri"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE0FFD2FD5C090AFC7FDE46"box="[758,938,302,328]"pageId="24"pageNumber="25"rank="species">gen. et sp. nov.</taxonomicNameLabel>
, the the VTA is smaller and the RTA is greatly elongated,
<figureCitationid="21C54D99FFE0FFD2FE320976FDC2DE63"box="[408,535,338,365]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="24"pageNumber="25">Fig. 5A–C</figureCitation>
, and the E and the MA are shorted and broader; and in
<taxonomicNameid="7EFE2A9FFFE0FFD2FB0C0977FAE8DE62"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[1190,1341,338,364]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="24"pageNumber="25"phylum="Arthropoda"rank="species"species="polotowae"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE0FFD2FAEF0976FEEFDE9F"pageId="24"pageNumber="25"rank="species">gen. et sp. nov.</taxonomicNameLabel>
, the VTA is finger-like and the MA is shorter,
<figureCitationid="21C54D99FFE0FFD2FCC10952FC2EDE9E"box="[875,1019,374,400]"captionStart="Fig"captionStartId="31.[189,231,1719,1745]"captionTargetBox="[211,1371,542,1665]"captionTargetId="figure-76@31.[189,1398,527,1680]"captionTargetPageId="31"captionText="Fig. 11. Tabiboka polotowae gen. et sp. nov., genitalia drawings; A–C= male holotype (CASENT_9016117; CAS); D–E = female paratype (CASENT_9016117; CAS). A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA= median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001819"httpUri="https://zenodo.org/record/14001819/files/figure.png"pageId="24"pageNumber="25">Fig. 11A–C</figureCitation>
). Females of
<taxonomicNameid="7EFE2A9FFFE0FFD2FB370953FADCDE9E"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[1181,1289,374,400]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="24"pageNumber="25"phylum="Arthropoda"rank="species"species="milloti">
have the epigyne with a heart-shaped median lobe (ML), broad anteriorly (anterior width greater than one-half total width,
<figureCitationid="21C54D99FFE0FFD2FEA1099BFEB1DED7"box="[267,356,447,473]"captionStart="Fig"captionStartId="27.[189,231,1595,1621]"captionTargetBox="[260,1379,284,1517]"captionTargetId="figure-11@27.[189,1398,265,1558]"captionTargetPageId="27"captionText="Fig. 9. Tabiboka milloti gen. nov. et sp. nov. genitalia drawings.A–C = male holotype (CASENT_9016063; MNHN). D–E = female paratype (CASENT_9018308; CAS).A. Left palp, prolateral view.B. Idem, ventral view. C. Idem, retrolateral view. D. Epigyne, ventral view.E. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold."figureDoi="http://doi.org/10.5281/zenodo.14001810"httpUri="https://zenodo.org/record/14001810/files/figure.png"pageId="24"pageNumber="25">Fig. 9D</figureCitation>
) (vs in
<taxonomicNameid="7EFE2A9FFFE0FFD2FE1309E4FDF7DED7"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[441,546,447,473]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="24"pageNumber="25"phylum="Arthropoda"rank="species"species="milleri">
, the ML is narrow anteriorly, anterior width less than one-half total width,
<figureCitationid="21C54D99FFE0FFD2FF1109C0FECDDEF1"box="[187,280,484,511]"captionStart="Fig"captionStartId="18.[189,233,1787,1813]"captionTargetBox="[219,1365,478,1733]"captionTargetId="figure-82@18.[189,1398,445,1749]"captionTargetPageId="18"captionText="Fig. 5. Tabiboka milleri gen. nov. et sp. nov. genitalia drawings of male and female paratypes (CASENT_9016086; CAS). A. Male left palp, prolateral view. B. Idem, ventral view.C. Idem, retrolateral view. D. Epigyne, ventral view. E. Idem, dorsal view. Abbreviations: C = conductor; CD = copulatory duct; CE = cymbial retrobasal extension; CO = copulatory opening; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; RTA = retrolateral tibial apophysis; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; VTA = ventral tibial apophysis. Scale bars = 0.5 mm. Illustrations by Jenny Speckels."figureDoi="http://doi.org/10.5281/zenodo.14001800"httpUri="https://zenodo.org/record/14001800/files/figure.png"pageId="24"pageNumber="25">Fig. 5D</figureCitation>
; and in
<taxonomicNameid="7EFE2A9FFFE0FFD2FEC409C1FDD6DEF0"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[366,515,484,510]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="24"pageNumber="25"phylum="Arthropoda"rank="species"species="polotowae">
<locationid="BC2107C7FFE0FFD2FB7D0B36FEE5DC5E"LSID="urn:lsid:plazi:treatment:3157E00AFFEFFFD6FDF60E9FFA8ED8E6:BC2107C7FFE0FFD2FB7D0B36FEE5DC5E"country="Madagascar"name="Parc National Andasibe"pageId="24"pageNumber="25"stateProvince="Toamasina Province">Parc National Andasibe</location>
<collectionCodeid="DFEFC9D9FFE0FFD2FCCB0B13FC6ADC5F"box="[865,959,823,849]"collectionName="France, Paris, Museum National d'Histoire Naturelle"country="France"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988"lsid="urn:lsid:biocol.org:col:34988"name="Museum National d'Histoire Naturelle"pageId="24"pageNumber="25">MNHN</collectionCode>
<materialsCitationid="09965B41FFE0FFD2FF170BB9FAF3DB0E"collectingDate="2003-01-16"collectingDateMax="2003-01-18"collectingDateMin="2003-01-16"collectingMethod="general collecting in rainforest"collectionCode="CAS"collectorName="C. Griswold & D. Silva & D. Andriamalala"country="Madagascar"elevation="960"latitude="-18.943888"location="Res. Analamazaotra"longLatPrecision="21"longitude="48.4175"pageId="24"pageNumber="25"specimenCode="CASENT_9018308"specimenCount="1"specimenCount-female="1"stateProvince="Toamasina Province"typeStatus="paratype">
<locationid="BC2107C7FFE0FFD2FC5C0BBAFAC5DCB9"LSID="urn:lsid:plazi:treatment:3157E00AFFEFFFD6FDF60E9FFA8ED8E6:BC2107C7FFE0FFD2FC5C0BBAFAC5DCB9"box="[1014,1296,925,952]"country="Madagascar"latitude="-18.943888"longLatPrecision="21"longitude="48.4175"name="Parc National Andasibe"pageId="24"pageNumber="25"stateProvince="Toamasina Province">Parc National Andasibe</location>
<collectingMethodid="60BF290BFFE0FFD2FE520BC2FCB3DB0E"box="[504,870,998,1024]"pageId="24"pageNumber="25">general collecting in rainforest</collectingMethod>
<collectionCodeid="DFEFC9D9FFE0FFD2FB470BC2FAF3DB0E"box="[1261,1318,998,1024]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="24"pageNumber="25"type="Museum">CAS</collectionCode>
<materialsCitationid="09965B41FFE0FFD2FAEB0BC2FDB8DB47"collectingDate="2003-01-16"collectingDateMax="2003-01-18"collectingDateMin="2003-01-16"collectionCode="CAS"collectorName="C. Griswold & D. Silva & D. Andriamalala"country="Madagascar"elevation="960"latitude="-18.943888"location="Parc National Andasibe"longLatPrecision="21"longitude="48.4175"pageId="24"pageNumber="25"specimenCode="CASENT_9005657"specimenCount="1"specimenCount-male="1"stateProvince="Toamasina"typeStatus="paratype">
<collectionCodeid="DFEFC9D9FFE0FFD2FD9E0C0BFDB8DB47"box="[564,621,1071,1097]"collectionName="USA, California, San Francisco, California Academy of Sciences"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="24"pageNumber="25"type="Museum">CAS</collectionCode>
<materialsCitationid="09965B41FFE0FFD2FD2C0C0BFAEBDB47"box="[646,1342,1071,1098]"collectingDate="2003-01-16"collectingDateMax="2003-01-18"collectingDateMin="2003-01-16"collectionCode="CAS"collectorName="C. Griswold & D. Silva & D. Andriamalala"country="Madagascar"elevation="960"latitude="-18.943888"location="Parc National Andasibe"longLatPrecision="21"longitude="48.4175"pageId="24"pageNumber="25"specimenCode="CASENT_9005261"specimenCount="1"specimenCount-male="1"stateProvince="Toamasina"typeStatus="paratype">
<collectionCodeid="DFEFC9D9FFE0FFD2FAAB0C0BFAEBDB47"box="[1281,1342,1071,1097]"collectionName="USA, California, San Francisco, California Academy of Sciences"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="24"pageNumber="25"type="Museum">CAS</collectionCode>
<collectionCodeid="DFEFC9D9FFE0FFD2FD480CCFFCCEDA0B"box="[738,795,1259,1285]"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="24"pageNumber="25"type="Museum">CAS</collectionCode>
<materialsCitationid="09965B41FFE0FFD2FC9D0CCFFE38DA27"collectingDate="2015-01-11"collectingDateMax="2015-01-14"collectingDateMin="2015-01-11"collectionCode="CAS"collectorName="B. L. Fisher"country="Madagascar"elevation="1385"latitude="-19.715721"location="Ankadivory"longLatPrecision="1"longitude="47.82084"municipality="Tsinjoarivo forest"pageId="24"pageNumber="25"specimenCode="CASENT_9065583"specimenCount="1"specimenCount-male="1"stateProvince="Antananarivo">
<collectionCodeid="DFEFC9D9FFE0FFD2FE1E0D2BFE38DA27"box="[436,493,1295,1321]"collectionName="USA, California, San Francisco, California Academy of Sciences"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="24"pageNumber="25"type="Museum">CAS</collectionCode>
<materialsCitationid="09965B41FFE0FFD2FDAC0D2BFB7ADA27"box="[518,1199,1295,1322]"collectingDate="2015-01-11"collectingDateMax="2015-01-14"collectingDateMin="2015-01-11"collectionCode="CAS"collectorName="B. L. Fisher"country="Madagascar"elevation="1385"latitude="-19.715721"location="Ankadivory"longLatPrecision="1"longitude="47.82084"municipality="Tsinjoarivo forest"pageId="24"pageNumber="25"specimenCode="CASENT_906558"specimenCount="1"specimenCount-male="1"stateProvince="Antananarivo">
<collectionCodeid="DFEFC9D9FFE0FFD2FBD80D2BFB7ADA27"box="[1138,1199,1295,1321]"collectionName="USA, California, San Francisco, California Academy of Sciences"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690"lsid="urn:lsid:biocol.org:col:34700"name="California Academy of Sciences"pageId="24"pageNumber="25"type="Museum">CAS</collectionCode>
<figureCitationid="21C54D99FFE0FFD2FEC70DCDFE23D90D"box="[365,502,1513,1539]"captionStart="Fig"captionStartId="25.[189,232,1717,1743]"captionTargetId="figure-19@26.[189,1398,304,1979]"captionTargetPageId="26"captionText="Fig. 8(on next page). Tabiboka milloti gen. nov. et sp. nov. male holotype (CASENT_9016063; MNHN). A. Habitus, dorsal view. B. Carapace, dorsal view. C. SEM of right male palp (mirrored), prolateral view. D. Idem, ventral view. E. Idem, retrolateral view. F. Idem, detail of bulb, ventral view. G. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = 4.6 mm); C–E = 0.5 mm; F–G = 0.1 mm. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14014722"httpUri="https://zenodo.org/record/14014722/files/figure.png"pageId="24"pageNumber="25">Fig. 8A–B</figureCitation>
). Carapace pale brownish yellow, darker along margin, with two faint, longitudinal dark bands along margin of pars cephalica; black surrounding all eyes; chelicerae brownish yellow; sternum yellow-white, unmarked; legs brownish yellow, femora, tibiae and metatarsi with faint median and subapical dusky annuli; abdomen pale yellow with darker cardiac mark and dorsolateral dark bands that break up into chevrons posteriorly, sides mottled with dark marks, venter yellow except with darker mottling around spinneret base.
<paragraphid="B941511CFFE0FFD2FF170F43FCCAD8A8"blockId="24.[188,1399,1895,1958]"pageId="24"pageNumber="25">EYES. ALE 0.33; AME 0.13; PLE 0.15; PME 0.14; ALE–AME 0.05; AME–AME 0.20; PLE–PME 0.13; PME–PME 0.06; MOQ: AW 0.26, PW 0.38, L 0.30.</paragraph>
<paragraphid="B941511CFFE0FFD2FF170FEAFD5AD8E7"blockId="24.[189,655,1998,2025]"box="[189,655,1998,2025]"pageId="24"pageNumber="25">CLYPEUS. 0.10 high, with six long setae.</paragraph>
<paragraphid="B941511CFFE1FFD3FF17092DFD49DE4B"blockId="25.[189,1398,265,325]"pageId="25"pageNumber="26">LEGS. Trochanters deeply notched. Unpaired claw smooth, rudimentary on legs III and IV. Metatarsi lacking subbasal crack. Scopulae absent.</paragraph>
<figureCitationid="21C54D99FFE1FFD3FEED0BB4FE11DCA5"box="[327,452,912,939]"captionStart="Fig"captionStartId="25.[189,232,1717,1743]"captionTargetId="figure-19@26.[189,1398,304,1979]"captionTargetPageId="26"captionText="Fig. 8(on next page). Tabiboka milloti gen. nov. et sp. nov. male holotype (CASENT_9016063; MNHN). A. Habitus, dorsal view. B. Carapace, dorsal view. C. SEM of right male palp (mirrored), prolateral view. D. Idem, ventral view. E. Idem, retrolateral view. F. Idem, detail of bulb, ventral view. G. Idem, detail of the RTA. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; MA = median apophysis; Pc = paraconductor; dRTA = retrodorsal tibial apophysis; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars: A–B = no scales provided (note: carapace length = 4.6 mm); C–E = 0.5 mm; F–G = 0.1 mm. Stacked microphotographs by Hannah Wood; SEM images by Charles Griswold and Darrell Ubick."figureDoi="http://doi.org/10.5281/zenodo.14014722"httpUri="https://zenodo.org/record/14014722/files/figure.png"pageId="25"pageNumber="26">Figs 8C–G</figureCitation>
, 9A–C). Tibia with VTA thumb-shaped, RTA broad, median with sharp, ventroapical point (vRTA), blunt retrodorsal projection (dRTA); cymbium oval with rounded apical extremity, without dorsal patch of chemosensitive setae, with small, sharp, conical retrobasal cymbial extension (CE) extending to tibial apex; subtegulum large, filling most of cymbial concavity, with transverse ridge (SL) that interacts with tegulum; tegulum oval with large, with rounded retrodistal protuberance and conical promedian lobe (TL) that locks with subtegulum; embolus (E) arises from retromargin of tegulum, short, simple, narrow; median apophysis (MA) strongly sclerotized elongate hook curved 270° prolaterad; conductor (C) hyaline, translucent, flattened, fan-shaped, widened at extremity; paraconductor (Pc) finger-like membranous, originating at base of MA and extending two-thirds length of E.
<paragraphid="B941511CFFE1FFD3FF170D6DFCB9DA88"blockId="25.[187,1399,1353,1414]"pageId="25"pageNumber="26">COLOUR.As in male except contrast between yellow-white background and dusky markings much bolder, especially annuli on posterior femora, tibiae and metatarsi.</paragraph>
<paragraphid="B941511CFFE1FFD3FF170D89FC6FDAC8"blockId="25.[189,954,1452,1479]"box="[189,954,1452,1479]"pageId="25"pageNumber="26">STERNUM. Oval with slightly sinuous sides; 2.20 wide, 2.40 long.</paragraph>
<taxonomicNameid="7EFE2A9FFFE1FFD3FE120E91FDA0D9C1"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[440,629,1717,1743]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="25"pageNumber="26"phylum="Arthropoda"rank="species"species="milloti"status="gen. nov. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE1FFD3FDD70E92FC8AD9DE"box="[637,863,1718,1744]"pageId="25"pageNumber="26"rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
<taxonomicNameid="7EFE2A9FFFE3FFD1FEA60E18FE13D958"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[268,454,1596,1622]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="27"pageNumber="28"phylum="Arthropoda"rank="species"species="milloti"status="gen. nov. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE3FFD1FE610E18FD72D958"box="[459,679,1596,1622]"pageId="27"pageNumber="28"rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
. Endogyne, dorsal view. Abbreviations: C = conductor; CE = cymbial retrobasal extension; dRTA = retrodorsal tibial apophysis; E = embolus; FD = fertilisation duct; LL = lateral lobe; MA = median apophysis; ML = median lobe; Pc = paraconductor; S = spermatheca; SL = subtegular locking lobe; TL = tegular locking lobe; vRTA = retroventral tibial apophysis; VTA = ventral tibial apophysis. Scale bars = 0.25 mm. Illustrations A–C by Jenny Speckels; D–E by Charles Griswold.
</paragraph>
</caption>
<paragraphid="B941511CFFE4FFD6FF17092DFD4CDE2D"blockId="28.[189,665,265,291]"box="[189,665,265,291]"pageId="28"pageNumber="29">CLYPEUS. 0.20 high, with five long setae.</paragraph>
<bibRefCitationid="DD6F2CEDFFE4FFD6FC680ECCFB14D80C"author="Wheeler W. C. & Coddington J. A. & Crowley L. M. & Dimitrov D. & Goloboff P. A. & Griswold C. E. & Hormiga G. & Prendini L. & Ramirez M. J. & Sierwald P. & Almeida-Silva L. M. & Alvarez-Padilla F. & Arnedo M. A. & Benavides L. R. & Benjamin S. P. & Bond J. E. & Grismado C. J. & Hasan E. & Hedin M. & Izquierdo M. A. & Labarque F. M. & Ledford J. & Lopardo L. & Maddison W. P. & Miller J. A. & Piacentini L. N. & Platnick N. I. & Polotow D. & Silva-Davila D. & Scharff N. & Szuts T. & Ubick D. & Vink C. & Wood H. M. & Zhang J. X."box="[962,1217,1768,1794]"pageId="28"pageNumber="29"pagination="576 - 616"refId="ref39136"refString="Wheeler W. C., Coddington J. A., Crowley L. M., Dimitrov D., Goloboff P. A., Griswold C. E., Hormiga G., Prendini L., Ramirez M. J., Sierwald P., Almeida-Silva L. M., Alvarez-Padilla F., Arnedo M. A., Benavides L. R., Benjamin S. P., Bond J. E., Grismado C. J., Hasan E., Hedin M., Izquierdo M. A., Labarque F. M., Ledford J., Lopardo L., Maddison W. P., Miller J. A., Piacentini L. N., Platnick N. I., Polotow D., Silva-Davila D., Scharff N., Szuts T., Ubick D., Vink C., Wood H. M. & Zhang J. X. 2017. The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics 33 (6): 576 - 616. https: // doi. org / 10.1111 / cla. 12182"type="journal article"year="2017">
<taxonomicNameid="7EFE2A9FFFE4FFD6FEE40F7FFDDBD87B"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[334,526,1883,1909]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="28"pageNumber="29"phylum="Arthropoda"rank="species"species="milloti"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE4FFD6FDBF0F78FD10D878"box="[533,709,1884,1910]"pageId="28"pageNumber="29"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicNameid="7EFE2A9FFFE4FFD6FF170FEAFEA8D8E6"authorityName="Henrard & Griswold & Jocqué"authorityYear="2024"box="[189,381,1998,2024]"class="Arachnida"family="Udubidae"genus="Tabiboka"kingdom="Animalia"order="Araneae"pageId="28"pageNumber="29"phylum="Arthropoda"rank="species"species="milloti"status="gen. et sp. nov.">
<taxonomicNameLabelid="90B93075FFE4FFD6FE2E0FEBFDE0D8E7"box="[388,565,1999,2025]"pageId="28"pageNumber="29"rank="species">gen. et sp. nov.</taxonomicNameLabel>
<figureCitationid="21C54D99FFE4FFD6FB5F0FEAFA98D8E6"box="[1269,1357,1998,2024]"captionStart="Fig"captionStartId="63.[189,231,1857,1883]"captionTargetBox="[189,1398,1005,1819]"captionTargetId="figure-208@63.[189,1398,1005,1819]"captionTargetPageId="63"captionText="Fig. 26. Distribution map of species, and samples localities in the northern and eastern part of Madagascar. Tabiboka milleri gen. et sp. nov. (orange), Tabiboka milloti gen. et sp. nov. (yellow), Tabiboka polotowae gen. et sp. nov. (purple), Zorascar pasunepipe gen. et sp. nov. (red), Zorascar pasunepomme gen. et sp. nov. (dark blue), Zorodictyna almae sp. nov. (pale blue), Zorodictyna silvadavilae sp. nov. (green), Zorodictyna oswaldi (Lenz, 1891) (pink star)."figureDoi="http://doi.org/10.5281/zenodo.14001851"httpUri="https://zenodo.org/record/14001851/files/figure.png"pageId="28"pageNumber="29">Fig. 26</figureCitation>