231 lines
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231 lines
27 KiB
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<document id="3F10C05F48D36CCE7E19A9A01E501DBD" ID-DOI="10.11646/zootaxa.3630.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261763" ID-ZooBank="C7529E9C-73F7-4E52-92E7-FCACA8F99A7D" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629846447618" checkinUser="felipe" docAuthor="Liu, Ye, Shi, Hongliang & Liang, Hongbin" docDate="2013" docId="03CF5A00720BFFBAFF17FA25F602FD96" docLanguage="en" docName="zt03630p518.pdf" docOrigin="Zootaxa 3630 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Chlaenioctenus) aodai Liu & Shi & Liang 2013, sp. n." docType="treatment" docVersion="1" lastPageNumber="512" masterDocId="FFF62278720EFFBDFF80FFC0F270FF81" masterDocTitle="Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus" masterLastPageNumber="518" masterPageNumber="505" pageNumber="510" updateTime="1730973406347" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods id="623A48322D8ABBC69CB307C17DD06187" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E3DEA69A9F5F4BAD25CB5B23C9EDEDB4">
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<mods:title id="871A3D076742CA679AE9B77FCECE5DA9">Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus</mods:title>
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</mods:titleInfo>
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<mods:name id="1F5FF423C7D599B8051F2B9CAB33A3EB" type="personal">
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<mods:role id="BD1BFB749C50FFC67A1D7C89766989DA">
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<mods:roleTerm id="E803248E68C211C09C871C9445480AC1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="62C6D77FB21D60EB4D625B387BF9AB0B">Liu, Ye</mods:namePart>
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</mods:name>
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<mods:name id="EA25F73A75048D6CDEC178769E647BAE" type="personal">
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<mods:role id="DFAE4E32DA55257B3A1A5B605D87B10D">
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<mods:roleTerm id="D6AA446D58C5A2FE96D728C3742793E1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87F3B1B84B2BEC919011DC33018019F2">Shi, Hongliang</mods:namePart>
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</mods:name>
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<mods:name id="C7B997C7A33E7A8BA29352EA74F8A3A4" type="personal">
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<mods:role id="B6787164C8DD23905E80ED40C9F632AA">
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<mods:roleTerm id="8C9D8D09E63D9FA4E24F08A7C760ED67">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0EAB9A7AD97903EB86EA197A3260E9C5">Liang, Hongbin</mods:namePart>
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<mods:nameIdentifier id="EDC62C33C471D478FF2618FC6871098A" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="17066ED9BBEFCCDE8EE55160A0DF2979">text</mods:typeOfResource>
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<mods:relatedItem id="39F6C1EC677A2B6668924EBC2A2B03CE" type="host">
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<mods:titleInfo id="9D202E707B5E1E18B1C9E12F5409EDCD">
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<mods:title id="AD88F14CD6B8EC022A66A9AD5AEA0788">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="94717091B7A79860F33AA02E96398C07">
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<mods:date id="84E3FBEFE7E208967C305F395ED4AF7D">2013</mods:date>
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<mods:detail id="60B2AA07FC4C2927339DC6099F5AE03D" type="pubDate">
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<mods:number id="CB0F7D5EC263290B2AFFAFBAC44E4E7B">2013-03-21</mods:number>
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</mods:detail>
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<mods:detail id="BCE1BD18BE85A61587701348CE41562E" type="volume">
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<mods:number id="356916D42E7790523E8ADD509C70C599">3630</mods:number>
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</mods:detail>
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<mods:detail id="AFEFB0425BA79D85528339CE2288023B" type="issue">
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<mods:number id="9D83C84A290F61367ABED1552086BD3D">3</mods:number>
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</mods:detail>
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<mods:extent id="400BE8E1B0C380F2E81AB2781FE63FA3" unit="page">
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<mods:start id="EA23F3E3757507AC9EB13D61ED8B55C6">505</mods:start>
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<mods:end id="D596869824938E64B0EB6CCA841888EB">518</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="786648A325B9CB8E81F444DE5EAEA4F3">journal article</mods:classification>
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<mods:identifier id="7EB7BC9C3CA31DC4180D05AD783E7F7D" type="DOI">10.11646/zootaxa.3630.3.6</mods:identifier>
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<mods:identifier id="5608118152DCD1D3714EFC39CFDC94F9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="849745CA16C6372BBFBB724B26CD577A" type="Zenodo-Dep">5261763</mods:identifier>
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<mods:identifier id="DDD3A5C87DC911326BD64A9DD146FA51" type="ZooBank">C7529E9C-73F7-4E52-92E7-FCACA8F99A7D</mods:identifier>
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</mods:mods>
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<treatment id="03CF5A00720BFFBAFF17FA25F602FD96" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFBAFF17FA25F602FD96" httpUri="http://treatment.plazi.org/id/03CF5A00720BFFBAFF17FA25F602FD96" lastPageId="7" lastPageNumber="512" pageId="5" pageNumber="510">
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<subSubSection id="C37CB89D720BFFB8FF17FA25F175FA7C" box="[151,773,1509,1534]" pageId="5" pageNumber="510" type="nomenclature">
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<paragraph id="8BD9EB16720BFFB8FF17FA25F175FA7C" blockId="5.[151,773,1509,1569]" box="[151,773,1509,1534]" pageId="5" pageNumber="510">
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<taxonomicName id="4C669095720BFFB8FF17FA25F0CEFA7F" ID-CoL="5XW7J" authority="Liu & Liang" authorityName="Liu & Shi & Liang" authorityYear="2013" box="[151,702,1509,1534]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="aodai" status="sp. nov." subGenus="Chlaenioctenus">
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<emphasis id="B9123704720BFFB8FF17FA25F37DFA7F" bold="true" box="[151,269,1509,1534]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
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(
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<emphasis id="B9123704720BFFB8FE9DFA25F3A1FA7F" bold="true" box="[285,465,1509,1534]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
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)
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<emphasis id="B9123704720BFFB8FE62FA25F0CEFA7F" bold="true" box="[482,702,1509,1534]" pageId="5" pageNumber="510">
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<emphasis id="B9123704720BFFB8FE62FA25F051FA7F" bold="true" box="[482,545,1509,1534]" italics="true" pageId="5" pageNumber="510">aodai</emphasis>
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Liu & Liang
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</emphasis>
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</taxonomicName>
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<emphasis id="B9123704720BFFB8FD45FA25F175FA7C" bold="true" box="[709,773,1509,1533]" pageId="5" pageNumber="510">
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<taxonomicNameLabel id="A2218A7F720BFFB8FD45FA25F175FA7C" box="[709,773,1509,1533]" pageId="5" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37CB89D720BFFB8FF17F9C9F070F9A0" box="[151,512,1545,1569]" pageId="5" pageNumber="510" type="description">
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<paragraph id="8BD9EB16720BFFB8FF17F9C9F070F9A0" blockId="5.[151,773,1509,1569]" box="[151,512,1545,1569]" pageId="5" pageNumber="510">
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(
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<figureCitation id="135DF793720BFFB8FF1FF9C9F356F9A0" box="[159,294,1545,1569]" captionStart="FIGURES 17–18" captionStartId="9.[151,264,1143,1166]" captionTargetBox="[164,1388,193,1122]" captionTargetId="figure-349@9.[151,1436,193,1122]" captionTargetPageId="9" captionText="FIGURES 17–18. Chlaenius aodai sp. n. 17, Holotype, male, dorsal view. 18, Holotype, male, ventral view. FIGURE 19. Chlaenius langsonensis Kirschenhofer, 2008. Holotype, male. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261779" httpUri="https://zenodo.org/record/5261779/files/figure.png" pageId="5" pageNumber="510">Figs. 17, 18</figureCitation>
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,
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<figureCitation id="135DF793720BFFB8FEB2F9C9F320F9A0" box="[306,336,1545,1569]" captionStart="FIGURES 20–25" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 20–25. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="5" pageNumber="510">23</figureCitation>
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,
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<figureCitation id="135DF793720BFFB8FEDCF9C9F30AF9A0" box="[348,378,1545,1569]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 26–31. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="5" pageNumber="510">29</figureCitation>
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,
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<figureCitation id="135DF793720BFFB8FE06F9C9F3D4F9A0" box="[390,420,1545,1569]" captionStart="FIGURES 32–37" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 32–37. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="5" pageNumber="510">35</figureCitation>
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,
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<figureCitation id="135DF793720BFFB8FE30F9C9F387F9A0" box="[432,503,1545,1569]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="5" pageNumber="510">44, 45</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37CB89D720BFFB8FF17F991F1DDF930" pageId="5" pageNumber="510" type="materials_examined">
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<paragraph id="8BD9EB16720BFFB8FF17F991F1DDF930" blockId="5.[151,1436,1617,2037]" pageId="5" pageNumber="510">
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<emphasis id="B9123704720BFFB8FF17F991F0D7F9E8" box="[151,679,1617,1642]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
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<emphasis id="B9123704720BFFB8FF17F991F3E2F9EB" bold="true" box="[151,402,1617,1642]" pageId="5" pageNumber="510">Specimen examined</emphasis>
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(
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<emphasis id="B9123704720BFFB8FE30F991F0E6F9EB" bold="true" box="[432,662,1617,1642]" pageId="5" pageNumber="510">
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total
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<specimenCount id="9D60209F720BFFB8FE7BF991F0E6F9EB" box="[507,662,1617,1642]" pageId="5" pageNumber="510" type="generic">1 specimens</specimenCount>
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</emphasis>
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).
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</emphasis>
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<materialsCitation id="3B0EE14B720BFFB8FD3CF991F635F90C" collectingDate="1989-05-24" collectorName="Pacholatko" country="Vietnam" location="Tamdao" pageId="5" pageNumber="510" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54DD55B4720BFFB8FD3CF991F158F9E8" box="[700,808,1617,1641]" pageId="5" pageNumber="510" type="holotype">Holotype</typeStatus>
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: male (NHMB), “N.
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<collectingCountry id="F371AB86720BFFB8FBCAF991F6D9F9E8" box="[1098,1193,1617,1641]" name="Vietnam" pageId="5" pageNumber="510">Vietnam</collectingCountry>
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(Tonkin),
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<location id="8EB9BDCD720BFFB8FABBF991F7E8F9E8" LSID="urn:lsid:plazi:treatment:03CF5A00720BFFBAFF17FA25F602FD96:8EB9BDCD720BFFB8FABBF991F7E8F9E8" box="[1339,1432,1617,1641]" country="Vietnam" name="Tamdao" pageId="5" pageNumber="510">Tamdao</location>
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, 12−
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<date id="FFD8CDD6720BFFB8FF40F9B5F346F90C" box="[192,310,1653,1677]" pageId="5" pageNumber="510" value="1989-05-24">
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<collectingDate id="EF9C343E720BFFB8FF40F9B5F346F90C" box="[192,310,1653,1677]" pageId="5" pageNumber="510" value="1989-05-24">24.5.1989</collectingDate>
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</date>
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,
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<collectorName id="26938EC0720BFFB8FECCF9B5F3B8F90C" box="[332,456,1653,1677]" pageId="5" pageNumber="510">Pacholatko</collectorName>
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leg.” / “Freiwlliger, Museumsverein, Basel, 1989”
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</materialsCitation>
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/ “
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<typeStatus id="54DD55B4720BFFB8FBFBF9B5F764F90C" box="[1147,1300,1653,1677]" pageId="5" pageNumber="510" type="holotype">HOLOTYPE</typeStatus>
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,
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<taxonomicName id="4C669095720BFFB8FAAAF9B5F3EFF930" authority="Liu & Shi & Liang, 2013" authorityName="Liu & Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="aodai" status="sp. nov." subGenus="Chlaenioctenus">
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<emphasis id="B9123704720BFFB8FAAAF9B5F7EBF90C" box="[1322,1435,1653,1677]" italics="true" pageId="5" pageNumber="510">Chlaenius</emphasis>
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(
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<emphasis id="B9123704720BFFB8FF20F959F33EF930" box="[160,334,1689,1713]" italics="true" pageId="5" pageNumber="510">Chlaenioctenus</emphasis>
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)
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<emphasis id="B9123704720BFFB8FEDFF959F3EFF930" box="[351,415,1689,1713]" italics="true" pageId="5" pageNumber="510">aodai</emphasis>
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</taxonomicName>
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<emphasis id="B9123704720BFFB8FE25F959F394F930" bold="true" box="[421,484,1689,1713]" pageId="5" pageNumber="510">
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<taxonomicNameLabel id="A2218A7F720BFFB8FE25F959F394F930" box="[421,484,1689,1713]" pageId="5" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, des. by Liu & Liang, 2012’’ [red label].
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</paragraph>
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</subSubSection>
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<subSubSection id="C37CB89D720BFFBAFF49F97DF7FCFEBE" lastPageId="7" lastPageNumber="512" pageId="5" pageNumber="510" type="description">
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<paragraph id="8BD9EB16720BFFB8FF49F97DF1F5F978" blockId="5.[151,1436,1617,2037]" pageId="5" pageNumber="510">
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<emphasis id="B9123704720BFFB8FF49F97DF30FF954" box="[201,383,1725,1749]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
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<emphasis id="B9123704720BFFB8FF49F97DF308F954" bold="true" box="[201,376,1725,1749]" pageId="5" pageNumber="510">Measurements</emphasis>
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.
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</emphasis>
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BL =
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<quantity id="4C9E46F3720BFFB8FE4AF97DF040F954" box="[458,560,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="5" pageNumber="510" unit="mm" value="16.6">16.6 mm</quantity>
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, HW =
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<quantity id="4C9E46F3720BFFB8FD0BF97DF093F954" box="[651,739,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="510" unit="mm" value="2.6">2.6 mm</quantity>
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, EYL = 1.0 mm, PL =
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<quantity id="4C9E46F3720BFFB8FC6AF97DF633F954" box="[1002,1091,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="510" unit="mm" value="3.5">3.5 mm</quantity>
|
|||
|
|
, PW =
|
|||
|
|
<quantity id="4C9E46F3720BFFB8FB18F97DF681F954" box="[1176,1265,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="510" unit="mm" value="3.5">3.5 mm</quantity>
|
|||
|
|
, EL =
|
|||
|
|
<quantity id="4C9E46F3720BFFB8FABFF97DF7E7F954" box="[1343,1431,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="510" unit="mm" value="9.5">9.5 mm</quantity>
|
|||
|
|
, EW =
|
|||
|
|
<quantity id="4C9E46F3720BFFB8FF60F921F348F978" box="[224,312,1761,1785]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="510" unit="mm" value="5.2">5.2 mm</quantity>
|
|||
|
|
, ratio PL/PW = 1.0, EL/EW = 1.83, EW/PW = 1.49.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD9EB16720BFFB8FF49F8C5F3A9F808" blockId="5.[151,1436,1617,2037]" pageId="5" pageNumber="510">
|
|||
|
|
<emphasis id="B9123704720BFFB8FF49F8C5F32AF89C" box="[201,346,1797,1822]" inLineHeading="true" pageId="5" pageNumber="510" reason="1">
|
|||
|
|
<emphasis id="B9123704720BFFB8FF49F8C5F323F89F" bold="true" box="[201,339,1797,1822]" pageId="5" pageNumber="510">Description</emphasis>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Head and pronotum black, with distinct metallic shine; elytra black, with weak shine; coxae, trochanters, tibiae, antennomeres 2 and 3 black or dark brown; antennomeres 1 and 4−11, mandibles, palpi and tarsi reddish brown; femora yellow in the middle portion, black at base and apex, basal black areas on hind femora longer than hind trochanters.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD9EB16720BFFBAFF49F855F7FCFEBE" blockId="5.[151,1436,1617,2037]" lastBlockId="7.[151,1436,151,536]" lastPageId="7" lastPageNumber="512" pageId="5" pageNumber="510">
|
|||
|
|
Head convex, vertex and occiput punctate; genae glabrous, oblique, as long as one-fourth of eyes in dorsal view; lateral margins of mentum rounded. Pronotum (
|
|||
|
|
<figureCitation id="135DF793720BFFB8FC84F879F128F850" box="[772,856,1977,2001]" captionStart="FIGURES 20–25" captionStartId="10.[151,264,1268,1291]" captionTargetBox="[194,1395,220,1229]" captionTargetId="figure-108@10.[172,1416,193,1252]" captionTargetPageId="10" captionText="FIGURES 20–25. Pronotum of Chlaenius (Chlaenioctenus) species. 20, C. cheni, holotype. 21, C. bodhidharma, holotype. 22, C. li, holotype. 23, C. aodai, holotype. 24, C. eneides, lectotype. 25, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261781" httpUri="https://zenodo.org/record/5261781/files/figure.png" pageId="5" pageNumber="510">Fig. 23</figureCitation>
|
|||
|
|
) long-ovoid, widest in the middle; lateral margins indistinctly bordered, sinuate before posterior angles; disc finely punctate, glabrous, median line distinct; posterior angles nearly rectangular, rounded at apex; base punctate, basal foveae deep, sparsely punctate. Elytra moderately convex, widest near posterior third; intervals (
|
|||
|
|
<figureCitation id="135DF7937209FFBAFD2DFF7CF172FF55" box="[685,770,188,212]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1942,1965]" captionTargetBox="[151,1436,1371,1910]" captionTargetId="figure-112@10.[151,1436,1371,1921]" captionTargetPageId="10" captionText="FIGURES 26–31. Elytral intervals of Chlaenius (Chlaenioctenus) species. 26, C. cheni, holotype. 27, C. bodhidharma, holotype. 28, C. li, holotype. 29, C. aodai, holotype. 30, C. eneides, lectotype. 31, C. langsonensis, holotype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5261785" httpUri="https://zenodo.org/record/5261785/files/figure.png" pageId="7" pageNumber="512">Fig. 29</figureCitation>
|
|||
|
|
) slightly convex, with regular punctures and pubescence; yellow spot at posterior third of each elytron, small, covering half interval 4 to interval 8. Proepisterna very sparsely punctate. Hind femora with basal black areas slightly longer than hind trochanters, claws (
|
|||
|
|
<figureCitation id="135DF7937209FFBAFB69FEC4F749FE9D" box="[1257,1337,260,284]" captionStart="FIGURES 32–37" captionStartId="11.[151,264,927,950]" captionTargetBox="[157,1404,190,873]" captionTargetId="figure-162@11.[151,1436,173,906]" captionTargetPageId="11" captionText="FIGURES 32–37. Hind claws of Chlaenius (Chlaenioctenus) species. 32, C. cheni, holotype. 33, C. bodhidharma, holotype. 34, C. li, holotype. 35, C. aodai, Holotype. 36, C. eneides, lectotype. 37, C. langsonensis, holotype. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5261787" httpUri="https://zenodo.org/record/5261787/files/figure.png" pageId="7" pageNumber="512">Fig. 35</figureCitation>
|
|||
|
|
) curved, with 5−6 short teeth. Aedeagus (
|
|||
|
|
<figureCitation id="135DF7937209FFBAFD83FEE7F0FAFEBE" box="[515,650,295,319]" captionStart="FIGURES 38–47" captionStartId="11.[151,264,1880,1903]" captionTargetBox="[179,1409,1044,1854]" captionTargetId="figure-165@11.[167,1411,1030,1861]" captionTargetPageId="11" captionText="FIGURES 38–47. Male genitalia of Chlaenius (Chlaenioctenus) species. 38, C. cheni, holotype (right lateral view). 39, C. cheni holotype (dorsal view). 40, C. bodhidharma, holotype (right lateral view). 41, C. bodhidharma, holotype (dorsal view). 42, C. li, holotype (right lateral view). 43, C. li, holotype (dorsal view). 44, C. aodai, holotype (right lateral view). 45, C. aodai, holotype (dorsal view). 46, C. eneides, lectotype (right lateral view). 47, C. eneides, lectotype (dorsal view). Scale bars: A = 1.0 mm (Figs. 38, 40, 42, 44, 46); B = 0.5 mm (Figs. 39, 41, 43, 45, 47)." figureDoi="http://doi.org/10.5281/zenodo.5261791" httpUri="https://zenodo.org/record/5261791/files/figure.png" pageId="7" pageNumber="512">Figs. 44, 45</figureCitation>
|
|||
|
|
) moderately robust in lateral view, apical lamella rounded and short.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF19BB9E7208FFBBFF17F8B5F132F82B" ID-DOI="http://doi.org/10.5281/zenodo.5261773" ID-Zenodo-Dep="5261773" httpUri="https://zenodo.org/record/5261773/files/figure.png" pageId="6" pageNumber="511" startId="6.[151,264,1909,1932]" targetBox="[282,1273,195,1883]" targetPageId="6">
|
|||
|
|
<paragraph id="8BD9EB167208FFBBFF17F8B5F132F82B" blockId="6.[151,1436,1909,1962]" pageId="6" pageNumber="511">
|
|||
|
|
<emphasis id="B91237047208FFBBFF17F8B5F339F80D" bold="true" box="[151,329,1909,1932]" pageId="6" pageNumber="511">FIGURES 9–12.</emphasis>
|
|||
|
|
<emphasis id="B91237047208FFBBFED0F8B6F3BDF80D" box="[336,461,1910,1932]" italics="true" pageId="6" pageNumber="511">
|
|||
|
|
<taxonomicName id="4C6690957208FFBBFED0F8B6F3C7F80D" authority="Liu & Shi & Liang, 2013" authorityName="Liu & Shi & Liang" authorityYear="2013" box="[336,439,1910,1932]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="genus" status="sp. nov.">Chlaenius</taxonomicName>
|
|||
|
|
li
|
|||
|
|
</emphasis>
|
|||
|
|
<emphasis id="B91237047208FFBBFE55F8B6F060F80A" bold="true" box="[469,528,1910,1931]" pageId="6" pageNumber="511">
|
|||
|
|
<taxonomicNameLabel id="A2218A7F7208FFBBFE55F8B6F060F80A" box="[469,528,1910,1931]" pageId="6" pageNumber="511" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
9, Holotype, male, dorsal view. 10, Holotype, male, ventral view. 11, Paratype, female, dorsal view. 12, Paratype, female, ventral view. Scale bar = 5.0 mm
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C37CB89D7209FFBAFF49FE8CF0FBFE06" pageId="7" pageNumber="512" type="etymology">
|
|||
|
|
<paragraph id="8BD9EB167209FFBAFF49FE8CF0FBFE06" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="512">
|
|||
|
|
<emphasis id="B91237047209FFBAFF49FE8CF33FFEE2" box="[201,335,332,356]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
|
|||
|
|
<emphasis id="B91237047209FFBAFF49FE8CF338FEE5" bold="true" box="[201,328,332,356]" pageId="7" pageNumber="512">Etymology</emphasis>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Latinized name
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFD89FE8CF039FEE5" authorityName="Liu & Shi & Liang" authorityYear="2013" box="[521,585,332,356]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="aodai">
|
|||
|
|
<emphasis id="B91237047209FFBAFD89FE8CF039FEE5" box="[521,585,332,356]" italics="true" pageId="7" pageNumber="512">aodai</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
refers to Ao Dai, a traditional costume of Vietnamese women, implying that the new species is discovered from
|
|||
|
|
<collectingCountry id="F371AB867209FFBAFDA5FEAFF0F7FE06" box="[549,647,367,391]" name="Vietnam" pageId="7" pageNumber="512">Vietnam</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37CB89D7209FFBAFF49FE54F602FD96" pageId="7" pageNumber="512" type="discussion">
|
|||
|
|
<paragraph id="8BD9EB167209FFBAFF49FE54F602FD96" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="512">
|
|||
|
|
<emphasis id="B91237047209FFBAFF49FE54F34CFE2A" box="[201,316,404,428]" inLineHeading="true" pageId="7" pageNumber="512" reason="1">
|
|||
|
|
<emphasis id="B91237047209FFBAFF49FE54F346FE2D" bold="true" box="[201,310,404,428]" pageId="7" pageNumber="512">Remarks</emphasis>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFEC3FE54F38BFE2D" authorityName="Liu & Shi & Liang" authorityYear="2013" box="[323,507,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="aodai">
|
|||
|
|
<emphasis id="B91237047209FFBAFEC3FE54F38BFE2D" box="[323,507,404,428]" italics="true" pageId="7" pageNumber="512">Chlaenius aodai</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFD0BFE54F097FE2D" authorityName="Liu & Shi & Liang" authorityYear="2013" box="[651,743,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="cheni">
|
|||
|
|
<emphasis id="B91237047209FFBAFD0BFE54F097FE2D" box="[651,743,404,428]" italics="true" pageId="7" pageNumber="512">C. cheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFC9DFE54F1FEFE2D" authorityName="Kirschenhofer" authorityYear="2008" box="[797,910,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="sehnali">
|
|||
|
|
<emphasis id="B91237047209FFBAFC9DFE54F1FEFE2D" box="[797,910,404,428]" italics="true" pageId="7" pageNumber="512">C. sehnali</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having yellow spot at posterior third of each elytron, but differs from
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFE30FE78F07FFE51" authorityName="Liu & Shi & Liang" authorityYear="2013" box="[432,527,440,464]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="cheni">
|
|||
|
|
<emphasis id="B91237047209FFBAFE30FE78F07FFE51" box="[432,527,440,464]" italics="true" pageId="7" pageNumber="512">C. cheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having pronotum wider than long contrast to pronotum wider than long in
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFA02FE78F2A4FE75" authorityName="Liu & Shi & Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="cheni">
|
|||
|
|
<emphasis id="B91237047209FFBAFA02FE78F2A4FE75" italics="true" pageId="7" pageNumber="512">C. cheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and from
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFED4FE1CF3B7FE75" authorityName="Kirschenhofer" authorityYear="2008" box="[340,455,476,500]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="sehnali">
|
|||
|
|
<emphasis id="B91237047209FFBAFED4FE1CF3B7FE75" box="[340,455,476,500]" italics="true" pageId="7" pageNumber="512">C. sehnali</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having all elytral intervals regularly pubescent and pronotal disc finely punctate in contrast to intervals glabrous medially and pronotal disc coarsely punctate in
|
|||
|
|
<taxonomicName id="4C6690957209FFBAFC7AFDC0F61BFD99" authorityName="Kirschenhofer" authorityYear="2008" box="[1018,1131,512,536]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="sehnali">
|
|||
|
|
<emphasis id="B91237047209FFBAFC7AFDC0F61BFD99" box="[1018,1131,512,536]" italics="true" pageId="7" pageNumber="512">C. sehnali</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|