<documentid="7E86CA7B82BCB9232D3A45A8346A63C5"ID-DOI="10.11646/phytotaxa.374.3.1"ID-ISSN="1179-3163"ID-Zenodo-Dep="13727297"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1725677943877"checkinUser="felipe"docAuthor="Michelangeli, Fabián A. & Goldenberg, Renato"docDate="2018"docId="03D7F370FFE2C86EFF5DFF6FA83BF8E7"docLanguage="en"docName="phytotaxa.374.3.1.pdf"docOrigin="Phytotaxa 374 (3)"docSource="http://dx.doi.org/10.11646/phytotaxa.374.3.1"docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article"docStyleId="96748F8F1B6C902996E134952A3A36B9"docStyleName="Phytotaxa.2014-.journal_article"docStyleVersion="13"docTitle="Miconia yanachagaensis Michelang. & R. Goldenb. 2018, sp. nov."docType="treatment"docVersion="2"lastPageNumber="199"masterDocId="FFEE8B08FFE9C860FFD5FFF0AD1AFFD6"masterDocTitle="New and noteworthy Melastomataceae from the Yanachaga-Chemillén National Park and surrounding areas in Oxapampa, Pasco, Peru"masterLastPageNumber="210"masterPageNumber="185"pageNumber="196"updateTime="1725690687117"updateUser="ExternalLinkService"zenodo-license-figures="UNSPECIFIED"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="8B2B4170206A0AA25697AA95960EB8AF">New and noteworthy Melastomataceae from the Yanachaga-Chemillén National Park and surrounding areas in Oxapampa, Pasco, Peru</mods:title>
<mods:affiliationid="4DD5DDE1A7A8A294A69D9310C3AFE6BC">Universidade Federal do Paraná, Caixa Postal 19031, CEP 81531 - 970, Curitiba, PR, Brazil.</mods:affiliation>
<taxonomicNameid="4C7E39E5FFE2C86BFF71FF6FAFF4FF6E"ID-CoL="42PT3"authority="Michelang. & R.Goldenb."authorityName="Michelang. & R. Goldenb."authorityYear="2018"box="[164,750,159,184]"class="Magnoliopsida"family="Melastomataceae"genus="Miconia"kingdom="Plantae"order="Myrtales"pageId="11"pageNumber="196"phylum="Tracheophyta"rank="species"species="yanachagaensis"status="sp. nov.">
<figureCitationid="13455EE3FFE2C86BFC8FFF6FAEF8FF6E"box="[858,994,159,184]"captionStart-0="FIGURE 7"captionStart-1="FIGURE 8"captionStartId-0="12.[136,229,1956,1977]"captionStartId-1="13.[136,229,1556,1577]"captionTargetBox-0="[194,1392,209,1912]"captionTargetBox-1="[197,1390,190,1531]"captionTargetId-0="figure-15@12.[178,1411,190,1932]"captionTargetId-1="figure-14@13.[197,1391,190,1531]"captionTargetPageId-0="12"captionTargetPageId-1="13"captionText-0="FIGURE 7. Miconia yanachagaensis. A. Fertile branch with flower buds. B. Fertile branch with flowers and young fruits. C. Flower bud. D. Flower, apical view, four stamens removed. E. Old flower, apical view, with persistent petals. F. Style and inflexed stamens in bud. G. Longitudinal section of the hypanthium plus ovary. H. Stamens, from left to right, ventral view, lateral view, ventral view of the filament and apical view of the stamen. A. Monteagudo 16057 (NY); B. Monteagudo 13759 (NY); C–H. Briceño 341 (NY)."captionText-1="FIGURE 8. Photos of Miconia yanachagaensis in the field. A. Fertile branch with flower buds. B. Leaf, abaxial surface. C. Young branch with annular interpetiolar ridge. D. Flower, lateral view, and buds. E. Flowers, top and lateral view. A–B. Michelangeli 2744. C. Michelangeli 2872. D–E. Michelangeli 2939."figureDoi-0="http://doi.org/10.5281/zenodo.13727315"figureDoi-1="http://doi.org/10.5281/zenodo.13727317"httpUri-0="https://zenodo.org/record/13727315/files/figure.png"httpUri-1="https://zenodo.org/record/13727317/files/figure.png"pageId="11"pageNumber="196">Figures 7–8</figureCitation>
<materialsCitationid="3B16483BFFE2C86BFF02FEC2AC9DFE59"collectingDate="2007-12-01"collectionCode="A, USM, HOXA, MO, NY"country="Peru"county="Province of Oxapampa"elevation="2650"latitude="-10.378333"location="Yanachaga-Palcazu"longLatPrecision="21"longitude="-75.45"municipality="Parque Nacional Yanachaga-Chemillen"pageId="11"pageNumber="196"specimenCount="2"stateProvince="Pasco"typeStatus="holotype">
<collectingCountyid="62A03AEAFFE2C86BFEA5FEC2AF43FE91"box="[368,601,306,327]"pageId="11"pageNumber="196">Province of Oxapampa</collectingCounty>
, Dist. Huancabamba,
<collectingMunicipalityid="6BA5D81CFFE2C86BFCECFEC2A9A7FE91"box="[825,1213,306,327]"pageId="11"pageNumber="196">Parque Nacional Yanachaga-Chemillén</collectingMunicipality>
, parte alta de la trocha
<locationid="8EA114BDFFE2C86BFF68FEA6AC60FEBD"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE2C86BFF68FEA6AC60FEBD"box="[189,378,342,363]"country="Peru"county="Province of Oxapampa"latitude="-10.378333"longLatPrecision="21"longitude="-75.45"municipality="Parque Nacional Yanachaga-Chemillen"name="Yanachaga-Palcazu"pageId="11"pageNumber="196"stateProvince="Pasco">Yanachaga-Palcazú</location>
<collectingDateid="EF849D4EFFE2C86BFD0BFEA6AE96FEBD"box="[734,908,342,363]"pageId="11"pageNumber="196"value="2007-12-01">1 December 2007</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFC15FEA6AEC8FEBD"box="[960,978,342,363]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="11"pageNumber="196"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFF68FE8AAC10FE59"box="[189,266,378,399]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFE84FE8AAC61FE59"box="[337,379,378,399]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
Shrubs or treelets 1.5–4(–7) m tall, occasionally decumbent. Young stems terete to slightly flattened and depressed at the internode apices, and terete with age, interpetiolar/subpetiolar ridge very thick and continuous, i.e. there is an annular structure around the stem node, the young stems densely covered with long dendritic-pedicellate trichomes,
long, erect, eglandular, the axis rough at the base then simple or with very short arms concentrated at its apex, ochraceous, the trichomes paler and somewhat caducous on older stems. Leaves opposite, isophyllous; petiole
, obovate to elliptic, apex obtuse to broadly acute, base acute or rounded, margin repand, hyaline and revolute, coriaceous, acrodromous nerves 5, suprabasal (the inner pair of secondaries joining the midvein 4.5–9(–13) mm above the base), the outer pair submarginal, distally fainter and disappearing towards the leaf apex, tertiary complete vein pairs 22–28, lacking intercalating, incomplete, percurrent veins, midvein and secondary veins strongly impressed, tertiary veins and reticulation slightly impressed on the adaxial surface, midvein, secondary and tertiary veins strongly prominent, and reticulation prominent on the abaxial surface, adaxial surface moderately to sparsely covered with dendritic trichomes in young leaves, and caducous or broken in older ones, sometimes persistent along the main nerves, abaxial surface moderately to densely covered (but never completely concealing the surface) with dendritic-penicellate trichomes
long, terminal, broadly pyramidal, multiflorous, with 3 main axes departing from the base, each one with 4–6 pairs of erect paraclades, nodes lacking accessory branches, the axis covered with the same indumentum as the branches; bracts and bracteoles not seen, and probably very early caducous. Flowers on short pedicels
, campanulate, green, outside densely covered with dendritic trichomes similar to the ones on young stems and inflorescences, inside glabrous; torus glabrous. Calyx tube
long, reddish, trapezoidal, the apex truncate, margins lacerate-ciliolate; sepals external projections a mere triangular hump, shorter than the laminae. Petals 6.3–7.1 ×
diam., slightly ventrally inclined on top of a very short beak, connective barely prolonged below the thecae, dorsally very shortly projected into a very broad, rounded or slightly emarginate appendage, lacking glands. Ovary 5-locular, inferior, apex glabrous; style ca.
<materialsCitationid="3B16483BFFE2C86BFF68FA07AC43F98E"collectingDate="2010-10-29"collectionCode="E, R, HOXA, MO, NY, USM"collectorName="Briceno & Rivera"country="Peru"county="Province of Oxapampa"elevation="2790"latitude="-10.53389"location="Parque Nacional Yanachaga-Chemillen"longLatPrecision="21"longitude="-75.34945"municipality="Oxapampa"pageId="11"pageNumber="196"specimenCount="1"stateProvince="Pasco"typeStatus="paratype">
<locationid="8EA114BDFFE2C86BFC32FA07A8B2F9C6"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE2C86BFC32FA07A8B2F9C6"box="[999,1448,1527,1552]"country="Peru"county="Province of Oxapampa"latitude="-10.53389"longLatPrecision="21"longitude="-75.34945"municipality="Oxapampa"name="Parque Nacional Yanachaga-Chemillen"pageId="11"pageNumber="196"stateProvince="Pasco">Parque Nacional Yanachaga-Chemillén</location>
,
<locationid="8EA114BDFFE2C86BFF5DF9EBAC5BF9E2"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE2C86BFF5DF9EBAC5BF9E2"box="[136,321,1563,1588]"country="Peru"county="Province of Oxapampa"latitude="-10.53389"longLatPrecision="21"longitude="-75.34945"municipality="Oxapampa"name="Abra Esperanza"pageId="11"pageNumber="196"stateProvince="Pasco">Abra Esperanza</location>
<collectingDateid="EF849D4EFFE2C86BFD3DF9EBAEB6F9E2"box="[744,940,1563,1588]"pageId="11"pageNumber="196"value="2010-10-29">29 October 2010</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFB41F9EBA9B2F9E2"box="[1172,1192,1563,1588]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFA9CF9EBA8BEF9E2"box="[1353,1444,1563,1588]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFF1BF9CFADE4F98E"box="[206,254,1599,1624]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<materialsCitationid="3B16483BFFE2C86BFEB3F9CFAE35F9AA"collectingDate="2006-10-13"collectionCode="L, R, CUZ, HOXA"collectorName="Cardenas & Francis"country="Peru"elevation="2877"latitude="-10.529166"location="Sector San Alberto"longLatPrecision="21"longitude="-75.35222"pageId="11"pageNumber="196"specimenCount="1"typeStatus="paratype">
<locationid="8EA114BDFFE2C86BFEB3F9CFAF25F98E"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE2C86BFEB3F9CFAF25F98E"box="[358,575,1599,1624]"country="Peru"latitude="-10.529166"longLatPrecision="21"longitude="-75.35222"name="Sector San Alberto"pageId="11"pageNumber="196">Sector San Alberto</location>
<collectionCodeid="ED6FDAA3FFE2C86BFE6EF993ACD5F9AA"box="[443,463,1635,1660]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFDA8F993AFA3F9AA"box="[637,697,1635,1660]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:14364"lsid="urn:lsid:biocol.org:col:14364"name="Universidad Nacional San Antonio Abad del Cusco"pageId="11"pageNumber="196"type="Herbarium">CUZ</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFD1CF993AE3EF9AA"box="[713,804,1635,1660]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
!)
</materialsCitation>
;
<materialsCitationid="3B16483BFFE2C86BFCEAF993AC3AF912"collectingDate="2009-04-18"collectingDateMax="2009-04-19"collectingDateMin="2009-04-18"collectionCode="M, R, HOXA, HUT, MO, USM"collectorName="Cueva & Rivera"country="Peru"elevation="2972"latitude="-10.541667"location="Refugio Abra Esperanza"longLatPrecision="21"longitude="-75.34889"pageId="11"pageNumber="196"specimenCount="1"typeStatus="paratype">
<collectingDateid="EF849D4EFFE2C86BFD5DF977AE4FF976"box="[648,853,1671,1696]"pageId="11"pageNumber="196"value="2009-04-18"valueMax="2009-04-19"valueMin="2009-04-18">18–19 April 2009</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFB81F977A972F976"box="[1108,1128,1671,1696]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFADCF977A844F976"box="[1289,1374,1671,1696]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFE0FF93FAF9DF93E"box="[474,647,1743,1768]"pageId="11"pageNumber="196"value="2016-03-21">21 March 2016</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFD2BF93FAE15F93E"box="[766,783,1743,1768]"country="USA"lsid="urn:lsid:biocol.org:col:15707"name="Field Museum of Natural History, Botany Department"pageId="11"pageNumber="196"type="Herbarium">F</collectionCode>
.
<collectionCodeid="ED6FDAA3FFE2C86BFCCFF93FAE34F93E"box="[794,814,1743,1768]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="11"pageNumber="196"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFB8AF93FA9A0F93E"box="[1119,1210,1743,1768]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
!,
<collectionCodeid="ED6FDAA3FFE2C86BFB1CF93FA9E3F93E"box="[1225,1273,1743,1768]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFAC0F903A8B1F8DA"box="[1301,1451,1779,1804]"pageId="11"pageNumber="196"value="2017-07-19">19 July 2017</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFF6FF8E7ADD1F8E6"box="[186,203,1815,1840]"country="USA"lsid="urn:lsid:biocol.org:col:15707"name="Field Museum of Natural History, Botany Department"pageId="11"pageNumber="196"type="Herbarium">F</collectionCode>
.
<collectionCodeid="ED6FDAA3FFE2C86BFF03F8E7ADF0F8E6"box="[214,234,1815,1840]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="11"pageNumber="196"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFDCDF8E7AF69F8E6"box="[536,627,1815,1840]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
!,
<collectionCodeid="ED6FDAA3FFE2C86BFD54F8E7AFABF8E6"box="[641,689,1815,1840]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFBA6F8E7A999F8E6"box="[1139,1155,1815,1840]"country="Sweden"lsid="urn:lsid:biocol.org:col:15668"name="Department of Botany, Swedish Museum of Natural History"pageId="11"pageNumber="196"type="Herbarium">S</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFE0CF8CBAFBFF882"box="[473,677,1851,1876]"pageId="11"pageNumber="196"value="2018-02-19">19 February 2018</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFCD3F8CBAE0DF882"box="[774,791,1851,1876]"country="USA"lsid="urn:lsid:biocol.org:col:15707"name="Field Museum of Natural History, Botany Department"pageId="11"pageNumber="196"type="Herbarium">F</collectionCode>
.
<collectionCodeid="ED6FDAA3FFE2C86BFCF1F8CBAE22F882"box="[804,824,1851,1876]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="11"pageNumber="196"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFBD7F8CBA909F882"box="[1026,1043,1851,1876]"country="Sweden"lsid="urn:lsid:biocol.org:col:15668"name="Department of Botany, Swedish Museum of Natural History"pageId="11"pageNumber="196"type="Herbarium">S</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFB73F8CBA81BF882"box="[1190,1281,1851,1876]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
!,
<collectionCodeid="ED6FDAA3FFE2C86BFAC7F8CBA858F882"box="[1298,1346,1851,1876]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFC9BF8AFA93CF8AE"box="[846,1062,1887,1912]"pageId="11"pageNumber="196"value="2010-11-12">12 November 2010</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFB8AF8AFA975F8AE"box="[1119,1135,1887,1912]"country="South Africa"lsid="urn:lsid:biocol.org:col:14647"name="University of the Witwatersrand"pageId="11"pageNumber="196"type="Herbarium">J</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFA9CF8AFA8BEF8AE"box="[1353,1444,1887,1912]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFCFCF873AED9F84A"box="[809,963,1923,1948]"pageId="11"pageNumber="196"value="2003-06-12">12 June 2003</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFC1AF873AEF9F84A"box="[975,995,1923,1948]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFB32F873A824F84A"box="[1255,1342,1923,1948]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15633"lsid="urn:lsid:biocol.org:col:15633"name="Universidad Nacional de la Amazónia Peruana"pageId="11"pageNumber="196"type="Herbarium">AMAZ</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFA9FF873A8B9F84A"box="[1354,1443,1923,1948]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFEC9F857AC47F816"box="[284,349,1959,1984]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530"lsid="urn:lsid:biocol.org:col:15530"name="Universidad Nacional Agraria La Molina"pageId="11"pageNumber="196"type="Herbarium">MOL</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFEBDF857AC82F816"box="[360,408,1959,1984]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFB91F857A9C2F816"box="[1092,1240,1959,1984]"pageId="11"pageNumber="196"value="2003-06-12">12 June 2003</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFAE5F857A85EF816"box="[1328,1348,1959,1984]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFEF3F83BAC67F832"box="[294,381,1995,2020]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15633"lsid="urn:lsid:biocol.org:col:15633"name="Universidad Nacional de la Amazónia Peruana"pageId="11"pageNumber="196"type="Herbarium">AMAZ</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFE5EF83BACFEF832"box="[395,484,1995,2020]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFD46F83BAFCEF832"box="[659,724,1995,2020]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530"lsid="urn:lsid:biocol.org:col:15530"name="Universidad Nacional Agraria La Molina"pageId="11"pageNumber="196"type="Herbarium">MOL</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFD36F83BAE09F832"box="[739,787,1995,2020]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFF29F81FAC0AF7DE"box="[252,272,2031,2056]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFDCEF81FAF68F7DE"box="[539,626,2031,2056]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15633"lsid="urn:lsid:biocol.org:col:15633"name="Universidad Nacional de la Amazónia Peruana"pageId="11"pageNumber="196"type="Herbarium">AMAZ</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFDAAF81FAFC2F7DE"box="[639,728,2031,2056]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFC54F81FAED8F7DE"box="[897,962,2031,2056]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530"lsid="urn:lsid:biocol.org:col:15530"name="Universidad Nacional Agraria La Molina"pageId="11"pageNumber="196"type="Herbarium">MOL</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFC1BF81FAEE4F7DE"box="[974,1022,2031,2056]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="11"pageNumber="196"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE2C86BFE31F7E3AF80F7FA"box="[484,666,2067,2092]"pageId="11"pageNumber="196"value="2003-08-01">01 August 2003</collectingDate>
<collectionCodeid="ED6FDAA3FFE2C86BFD0DF7E3AFF6F7FA"box="[728,748,2067,2092]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="11"pageNumber="196">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFC29F7E3A949F7FA"box="[1020,1107,2067,2092]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15633"lsid="urn:lsid:biocol.org:col:15633"name="Universidad Nacional de la Amazónia Peruana"pageId="11"pageNumber="196"type="Herbarium">AMAZ</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE2C86BFBB4F7E3A9A0F7FA"box="[1121,1210,2067,2092]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="11"pageNumber="196"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE2C86BFAB3F7E3A8BDF7FA"box="[1382,1447,2067,2092]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530"lsid="urn:lsid:biocol.org:col:15530"name="Universidad Nacional Agraria La Molina"pageId="11"pageNumber="196"type="Herbarium">MOL</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE4C86DFF5DF951ADA2F96C"box="[136,184,1697,1722]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE4C86DFC6DF951A970F96C"box="[952,1130,1697,1722]"pageId="13"pageNumber="198"value="2003-08-01">01 August 2003</collectingDate>
<collectionCodeid="ED6FDAA3FFE4C86DFB77F951A9ACF96C"box="[1186,1206,1697,1722]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="13"pageNumber="198">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFF45F935ADF1F908"box="[144,235,1733,1758]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFE96F935AC69F908"box="[323,371,1733,1758]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE4C86DFBEDF935A9F5F908"box="[1080,1263,1733,1758]"pageId="13"pageNumber="198"value="2004-08-20">20 August 2004</collectingDate>
<collectionCodeid="ED6FDAA3FFE4C86DFAF9F935A85AF908"box="[1324,1344,1733,1758]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="13"pageNumber="198">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFEFDF919AC99F8D4"box="[296,387,1769,1794]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFE08F919AF17F8D4"box="[477,525,1769,1794]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<materialsCitationid="3B16483BFFE4C86DFDACF919AE38F8B8"collectingDate="2011-04-18"collectingDateMax="2012-11-04"collectingDateMin="2011-04-18"collectionCode="R, HOXA, HUT, MO, USM, E, NY"collectorName="Vasquez & Briceno"country="Peru"county="Estacion Biologica San Alberto"elevation="2903"latitude="-10.380278"location="Distrito Huancabamba"longLatPrecision="21"longitude="-75.46166"municipality="Abra Esperanza"pageId="13"pageNumber="198"specimenCount="1"typeStatus="paratype">
<collectingCountyid="62A03AEAFFE4C86DFDACF919AEFDF8D4"box="[633,999,1769,1794]"pageId="13"pageNumber="198">Estacion Biológica San Alberto</collectingCounty>
<collectingDateid="EF849D4EFFE4C86DFEF1F8FDACEBF8F0"box="[292,497,1805,1830]"pageId="13"pageNumber="198"value="2012-11-04">4 November 2012</collectingDate>
<collectionCodeid="ED6FDAA3FFE4C86DFDF8F8FDAF5BF8F0"box="[557,577,1805,1830]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="13"pageNumber="198">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFC85F8FDAEB1F8F0"box="[848,939,1805,1830]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
<locationid="8EA114BDFFE4C86DFB73F8FDA8BDF8F0"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE4C86DFB73F8FDA8BDF8F0"box="[1190,1447,1805,1830]"country="Peru"county="Estacion Biologica San Alberto"latitude="-10.380278"longLatPrecision="21"longitude="-75.46166"municipality="Abra Esperanza"name="Distrito Huancabamba"pageId="13"pageNumber="198">Distrito Huancabamba</location>
,
<locationid="8EA114BDFFE4C86DFF5DF8C1AF5BF89C"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE4C86DFF5DF8C1AF5BF89C"box="[136,577,1841,1866]"country="Peru"county="Estacion Biologica San Alberto"latitude="-10.380278"longLatPrecision="21"longitude="-75.46166"municipality="Abra Esperanza"name="Parque Nacional Yanachaga-Chemillen"pageId="13"pageNumber="198">Parque Nacional Yanachaga-Chemillén</location>
,
<locationid="8EA114BDFFE4C86DFD9EF8C1AE4BF89C"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE4C86DFD9EF8C1AE4BF89C"box="[587,849,1841,1866]"country="Peru"county="Estacion Biologica San Alberto"latitude="-10.380278"longLatPrecision="21"longitude="-75.46166"municipality="Abra Esperanza"name="Sector Abra Yanachaga"pageId="13"pageNumber="198">Sector Abra Yanachaga</location>
<collectingDateid="EF849D4EFFE4C86DFB35F8C1A863F89C"box="[1248,1401,1841,1866]"pageId="13"pageNumber="198"value="2011-04-18">18 April 2011</collectingDate>
<collectionCodeid="ED6FDAA3FFE4C86DFE40F8A5ACEAF8B8"box="[405,496,1877,1902]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFD58F8A5AFA7F8B8"box="[653,701,1877,1902]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFBCFF889A958F844"box="[1050,1090,1913,1938]"country="Australia"lsid="urn:lsid:biocol.org:col:14575"name="University of New England"pageId="13"pageNumber="198"type="Herbarium">NE</collectionCode>
<collectingDateid="EF849D4EFFE4C86DFF78F86DAC5AF860"box="[173,320,1949,1974]"pageId="13"pageNumber="198"value="1988-06-25">25 June 1988</collectingDate>
<collectionCodeid="ED6FDAA3FFE4C86DFEA3F86DAC90F860"box="[374,394,1949,1974]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="13"pageNumber="198">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFE2AF86DAF09F860"box="[511,531,1949,1974]"country="Germany"lsid="urn:lsid:biocol.org:col:15534"name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet"pageId="13"pageNumber="198"type="Herbarium">B</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFD12F86DAFC6F860"box="[711,732,1949,1974]"country="USA"lsid="urn:lsid:biocol.org:col:15707"name="Field Museum of Natural History, Botany Department"pageId="13"pageNumber="198"type="Herbarium">F</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFEEEF831AC55F80C"box="[315,335,1985,2010]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="13"pageNumber="198"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFD55F831AFC1F80C"box="[640,731,1985,2010]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
!,
<collectionCodeid="ED6FDAA3FFE4C86DFD39F831AE07F80C"box="[748,797,1985,2010]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFF2AF815AC09F828"box="[255,275,2021,2046]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="13"pageNumber="198"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFD96F815AF84F828"box="[579,670,2021,2046]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFD39F815AE07F828"box="[748,797,2021,2046]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFF23F7F9AC10F7F4"box="[246,266,2057,2082]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="13"pageNumber="198"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFDFFF7F9AF9FF7F4"box="[554,645,2057,2082]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="13"pageNumber="198"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE4C86DFD19F7F9AFE7F7F4"box="[716,765,2057,2082]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="13"pageNumber="198"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE7C86EFDC7FF50AFB4FF6F"box="[530,686,160,185]"pageId="14"pageNumber="199"value="2007-04-22">22 April 2007</collectingDate>
<collectionCodeid="ED6FDAA3FFE7C86EFD3DFF50AFE6FF6F"box="[744,764,160,185]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="14"pageNumber="199"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFBE5FF50A991FF6F"box="[1072,1163,160,185]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFB0AFF50A815FF6F"box="[1247,1295,160,185]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="14"pageNumber="199"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE7C86EFE04FF34AFB3FF0B"box="[465,681,196,221]"pageId="14"pageNumber="199"value="2007-11-24">24 November 2007</collectingDate>
</date>
(fl),
<collectionCodeid="ED6FDAA3FFE7C86EFD0AFF34AFECFF0B"box="[735,758,196,221]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="14"pageNumber="199"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFBFCFF34A99EFF0B"box="[1065,1156,196,221]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFB03FF34A81CFF0B"box="[1238,1286,196,221]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="14"pageNumber="199"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE7C86EFC03FF18A9BBFED7"box="[982,1185,232,257]"pageId="14"pageNumber="199"value="2007-12-02">2 December 2007</collectingDate>
<collectionCodeid="ED6FDAA3FFE7C86EFB0AFF18A9E9FED7"box="[1247,1267,232,257]"country="USA"lsid="urn:lsid:biocol.org:col:15406"name="Harvard University - Arnold Arboretum"pageId="14"pageNumber="199"type="Herbarium">A</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFED7FEFCAC47FEF3"box="[258,349,268,293]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFDD6FEFCAF29FEF3"box="[515,563,268,293]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="14"pageNumber="199"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE7C86EFE57FEC0AF24FE9F"box="[386,574,304,329]"pageId="14"pageNumber="199"value="2003-10-10">10 October 2003</collectingDate>
<collectionCodeid="ED6FDAA3FFE7C86EFD4DFEC0AFB2FE9F"box="[664,680,304,329]"country="South Africa"lsid="urn:lsid:biocol.org:col:14647"name="University of the Witwatersrand"pageId="14"pageNumber="199"type="Herbarium">J</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFCA1FEC0AED5FE9F"box="[884,975,304,329]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFBBDFEC0A982FE9F"box="[1128,1176,304,329]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="14"pageNumber="199"type="Herbarium">NY</collectionCode>
<locationid="8EA114BDFFE7C86EFE01FEA4AF9FFEBB"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE7C86EFE01FEA4AF9FFEBB"box="[468,645,340,365]"country="Peru"latitude="-10.383333"longLatPrecision="1298"longitude="-75.45"municipality="Cordillera Yanachaga"name="Rio San Daniel"pageId="14"pageNumber="199">Rio San Daniel</location>
<collectingDateid="EF849D4EFFE7C86EFBC3FEA4A9B0FEBB"box="[1046,1194,340,365]"pageId="14"pageNumber="199"value="1984-07-13">13 July 1984</collectingDate>
<collectingDateid="EF849D4EFFE7C86EFC32FE88A981FE47"box="[999,1179,376,401]"pageId="14"pageNumber="199"value="2008-08-19">19 August 2008</collectingDate>
<collectionCodeid="ED6FDAA3FFE7C86EFECCFE6CAC6EFE63"box="[281,372,412,437]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFDDEFE6CAF21FE63"box="[523,571,412,437]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="14"pageNumber="199"type="Herbarium">NY</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFCE0FE30AE8AFE0F"box="[821,912,448,473]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<materialsCitationid="3B16483BFFE7C86EFB92FE30AC05FDF7"collectingDate="2008-03-01"collectionCode="R, AMAZ, HOXA, HUT, MO, USM"collectorName="Vasquez"country="Peru"elevation="3350"latitude="-10.429444"location="Sector San Daniel"longLatPrecision="21"longitude="-75.435"pageId="14"pageNumber="199"specimenCount="1"typeStatus="paratype">
<locationid="8EA114BDFFE7C86EFB92FE30A80DFE0F"LSID="urn:lsid:plazi:treatment:03D7F370FFE2C86EFF5DFF6FA83BF8E7:8EA114BDFFE7C86EFB92FE30A80DFE0F"box="[1095,1303,448,473]"country="Peru"latitude="-10.429444"longLatPrecision="21"longitude="-75.435"name="Sector San Daniel"pageId="14"pageNumber="199">Sector San Daniel</location>
<collectionCodeid="ED6FDAA3FFE7C86EFC54FE14AE8FFE2B"box="[897,917,484,509]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="14"pageNumber="199">R</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFB4AFE14A9ECFE2B"box="[1183,1270,484,509]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15633"lsid="urn:lsid:biocol.org:col:15633"name="Universidad Nacional de la Amazónia Peruana"pageId="14"pageNumber="199"type="Herbarium">AMAZ</collectionCode>
,
<collectionCodeid="ED6FDAA3FFE7C86EFAD6FE14A846FE2B"box="[1283,1372,484,509]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectingDateid="EF849D4EFFE7C86EFC91FDF8AEE5FDF7"box="[836,1023,520,545]"pageId="14"pageNumber="199"value="2009-10-31">31 October 2009</collectingDate>
<collectionCodeid="ED6FDAA3FFE7C86EFBE3FDF8A954FDF7"box="[1078,1102,520,545]"country="Finland"lsid="urn:lsid:biocol.org:col:15618"name="University of Helsinki"pageId="14"pageNumber="199"type="Herbarium">H</collectionCode>
.
<collectorNameid="268B27B0FFE7C86EFB8DFDF8A9F4FDF7"box="[1112,1262,520,545]"pageId="14"pageNumber="199">van der Werff</collectorName>
22901
</emphasis>
(
<collectionCodeid="ED6FDAA3FFE7C86EFA9CFDF8A8BEFDF7"box="[1353,1444,520,545]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFF05FDDCAC1AFD93"box="[208,256,556,581]"country="USA"lsid="urn:lsid:biocol.org:col:15556"name="William and Lynda Steere Herbarium of the New York Botanical Garden"pageId="14"pageNumber="199"type="Herbarium">NY</collectionCode>
<collectingDateid="EF849D4EFFE7C86EFC41FDDCA94DFD93"box="[916,1111,556,581]"pageId="14"pageNumber="199"value="2009-10-31">31 October 2009</collectingDate>
<collectionCodeid="ED6FDAA3FFE7C86EFB42FDDCA9B5FD93"box="[1175,1199,556,581]"country="Finland"lsid="urn:lsid:biocol.org:col:15618"name="University of Helsinki"pageId="14"pageNumber="199"type="Herbarium">H</collectionCode>
.
<collectorNameid="268B27B0FFE7C86EFB68FDDCA841FD93"box="[1213,1371,556,581]"pageId="14"pageNumber="199">van der Werff</collectorName>
22972
</emphasis>
(
<collectionCodeid="ED6FDAA3FFE7C86EFF45FDA0ADF1FDBF"box="[144,235,592,617]"country="Peru"httpUri="http://grbio.org/cool/yriw-6hkr"name="Estación biológica del Jardin Botanico de Missouri"pageId="14"pageNumber="199"type="Herbarium">HOXA</collectionCode>
<collectionCodeid="ED6FDAA3FFE7C86EFE56FDA0ACDEFDBF"box="[387,452,592,617]"country="Peru"httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530"lsid="urn:lsid:biocol.org:col:15530"name="Universidad Nacional Agraria La Molina"pageId="14"pageNumber="199"type="Herbarium">MOL</collectionCode>
. A. Fertile branch with flower buds. B. Fertile branch with flowers and young fruits. C. Flower bud. D. Flower, apical view, four stamens removed. E. Old flower, apical view, with persistent petals. F. Style and inflexed stamens in bud. G. Longitudinal section of the hypanthium plus ovary. H. Stamens, from left to right, ventral view, lateral view, ventral view of the filament and apical view of the stamen. A.
in the field. A. Fertile branch with flower buds. B. Leaf, abaxial surface. C. Young branch with annular interpetiolar ridge. D. Flower, lateral view, and buds. E. Flowers, top and lateral view. A–B.
<emphasisid="B90A9E74FFE7C86EFF68FD84AF9EFD5B"bold="true"box="[189,644,628,653]"pageId="14"pageNumber="199">Distribution, ecology, and phenology</emphasis>
is known from the dwarf high montane shrublands and exposed grasslands areas along the Cordillera Yanachaga and the North West portion of the Yanachaga-Chemillén National Park above
<figureCitationid="13455EE3FFE7C86EFBC2FD4CA963FD03"box="[1047,1145,700,725]"captionStart="FIGURE 6"captionStartId="10.[136,229,1969,1990]"captionTargetBox="[197,1392,571,1937]"captionTargetId="figure-217@10.[164,1424,567,1947]"captionTargetPageId="10"captionText="FIGURE 6. Distribution of Miconia palcazuana and Miconia yanachagaensis."figureDoi="http://doi.org/10.5281/zenodo.13727313"httpUri="https://zenodo.org/record/13727313/files/figure.png"pageId="14"pageNumber="199">Figure 6</figureCitation>
. In spite of its reduced EEO that would place it as Endangered, given the fact that the species seems to be locally abundant, most of the populations are inside a National Park in areas not frequented by the public and under no immediate threat, we recommend that this species is temporarily considered as Least Concern (
<bibRefCitationid="EFEF3F97FFE7C86EFC16FC80A951FC5F"author="IUCN"box="[963,1099,880,905]"pageId="14"pageNumber="199"refId="ref16074"refString="IUCN. (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK."type="book"year="2001">IUCN 2001</bibRefCitation>
;
<bibRefCitationid="EFEF3F97FFE7C86EFB8DFC80AC74FC7B"author="IUCN Standards & Petitions Subcommittee"pageId="14"pageNumber="199"refId="ref16104"refString="IUCN Standards and Petitions Subcommittee. (2017) Guidelines for using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommitee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 6 November 2018)"type="url"year="2017">IUCN Standards and Petitions Subcommittee 2017</bibRefCitation>
:—The specific epithet of this new species alludes to the name of the mountain range where most specimens have been collected and that also forms part of the name of the national park. Yanachaga is a Quechua word that means black bundle or black package (
<bibRefCitationid="EFEF3F97FFE7C86EFDBEFBF0AE03FBCF"author="INRENA"box="[619,793,1024,1049]"pageId="14"pageNumber="199"refId="ref16052"refString="INRENA. (2005) Plan Maestro del Parque Nacional Yanachaga-Chemillen 2005 - 2009, Lima, Peru. Editorial INRENA."type="book"year="2005">INRENA 2005</bibRefCitation>
<bibRefCitationid="EFEF3F97FFE7C86EFE94FBB8AF35FBB7"author="Naudin, C. V."box="[321,559,1096,1121]"pageId="14"pageNumber="199"pagination="83 - 246"refId="ref16879"refString="Naudin, C. V. (1850 a) Melastomacearum monographicae descriptiones. Annales des Sciences Naturelles; Botanique Series 3, 16: 83 - 246."type="journal article"year="1850">Naudin (1850a: 204)</bibRefCitation>
Triana ex Hook. f.
</taxonomicName>
in
<bibRefCitationid="EFEF3F97FFE7C86EFCE1FBB8A9B2FBB7"author="Bentham, G. & Hooker, J. D."box="[820,1192,1096,1121]"pageId="14"pageNumber="199"refId="ref14828"refString="Bentham, G. & Hooker, J. D. (1867) Genera planta rum. Vols. 1 - 2. Reeve, Williams & Norgate, London."type="book"year="1867">Bentham & Hooker (1867: 763)</bibRefCitation>
, particularly due to its ovate-oblong anther thecae that open by a minute terminal pore. While this section is clearly not monophyletic, most species in it are found in the Andes, concentrated in two clades (Mecranium + Anaectocalyx + Allies and
<taxonomicNameid="4C7E39E5FFE7C86EFAFEFB60ACCDFB1B"authority="III sensu Goldenberg et al. 2008"authorityName="Goldenberg"authorityYear="2008"class="Magnoliopsida"family="Melastomataceae"genus="Miconia"kingdom="Plantae"order="Myrtales"pageId="14"pageNumber="199"phylum="Tracheophyta"rank="genus">
Miconia III sensu
<bibRefCitationid="EFEF3F97FFE7C86EFF1BFB44ACCDFB1B"author="Goldenberg, R. & Penneys, D. S. & Almeda, F. & Judd, W. S. & Michelangeli, F. A."box="[206,471,1204,1229]"pageId="14"pageNumber="199"pagination="963 - 979"refId="ref15838"refString="Goldenberg, R., Penneys, D. S., Almeda, F., Judd, W. S. & Michelangeli, F. A. (2008) Phylogeny of Miconia (Melastomataceae): Patterns of stamen diversification in a megadiverse neotropical genus. International Journal Of Plant Sciences 169: 963 - 979. https: // doi. org / 10.1086 / 589697"type="journal article"year="2008">
<bibRefCitationid="EFEF3F97FFE7C86EFD12FB28AEA8FB27"author="Bonpland, A."box="[711,946,1240,1265]"pageId="14"pageNumber="199"refId="ref15021"refString="Bonpland, A. (1816) Monographie des Melastomatacees. Librairie Greque-Latine-Allemande, Paris."type="book"year="1816">Bonpland 1816: 123</bibRefCitation>
)
<bibRefCitationid="EFEF3F97FFE7C86EFC11FB28A9FEFB27"author="De Candolle, A. P."box="[964,1252,1240,1265]"pageId="14"pageNumber="199"refId="ref15698"refString="De Candolle, A. P. (1828) Memoires sur la famille des Melastomacees. Treuttel et Wurtz, Paris."type="book"year="1828">De Candolle (1828: 188)</bibRefCitation>
</taxonomicName>
, characterized by large flowers with yellow anthers and broad, capitate or funnelform stigmas. Moreover, the majority of these species also have glandular trichomes in the filament and/or style. This species group includes several species in the Andes and it is in dire need of a careful revision. In
<bibRefCitationid="EFEF3F97FFE7C86EFDF0FA98AE0BFA57"author="Gleason, H. A."box="[549,785,1384,1409]"pageId="14"pageNumber="199"pagination="215 - 262"refId="ref15752"refString="Gleason, H. A. (1931) Studies on the flora of northern South America-XV. Recent collections of Melastomataceae from Peru and Amazonian Brazil. Bulletin of the Torrey Botanical Club 58: 215 - 262. https: // doi. org / 10.2307 / 2480704"type="journal article"year="1931">Gleason (1931: 243)</bibRefCitation>
<bibRefCitationid="EFEF3F97FFE7C86EFA95FA98AC17FA73"author="Macbride, J. F."pageId="14"pageNumber="199"pagination="172 - 191"refId="ref16180"refString="Macbride, J. F. (1929) New and renamed Peruvian Melastomes. Publications of the Field Museum of Natural History Botanical Series, 4: 172 - 191."type="journal article"year="1929">Macbride (1929: 182)</bibRefCitation>
the long dendritic-pedicellate trichomes on the lower leaf surface; all other species are either glabrous or have stellate or stellate-pinnoid trichomes.
<bibRefCitationid="EFEF3F97FFE7C86EFF31F95CACF8F913"author="Macbride, J. F."box="[228,482,1708,1733]"pageId="14"pageNumber="199"pagination="172 - 191"refId="ref16180"refString="Macbride, J. F. (1929) New and renamed Peruvian Melastomes. Publications of the Field Museum of Natural History Botanical Series, 4: 172 - 191."type="journal article"year="1929">Macbride (1929: 188)</bibRefCitation>
also has ovate, coriaceous leaves with dendritic trichomes on the abaxial surface, and it is found in similar environments, this later species has four-merous flowers, with short and broad, white to cream-colored anthers that open by longitudinal slits, which would place it in
<taxonomicNameid="4C7E39E5FFE7C86EFB20F904AE95F8E7"authority="(Richard ex de Candolle 1828: 197) Triana ex Hook. f."authorityName="Hook. f."baseAuthorityName="de Candolle"baseAuthorityPageNumber="197"baseAuthorityYear="1828"class="Magnoliopsida"family="Melastomataceae"genus="Miconia"kingdom="Plantae"order="Myrtales"pageId="14"pageNumber="199"phylum="Tracheophyta"rank="section"section="Chaenopleura">
<bibRefCitationid="EFEF3F97FFE7C86EFE6FF8E8AFA3F8E7"author="De Candolle, A. P."box="[442,697,1816,1841]"pageId="14"pageNumber="199"refId="ref15698"refString="De Candolle, A. P. (1828) Memoires sur la famille des Melastomacees. Treuttel et Wurtz, Paris."type="book"year="1828">de Candolle 1828: 197</bibRefCitation>
) Triana ex Hook.f.
</taxonomicName>
in
<bibRefCitationid="EFEF3F97FFE7C86EFC67F8E8A806F8E7"author="Bentham, G. & Hooker, J. D."box="[946,1308,1816,1841]"pageId="14"pageNumber="199"refId="ref14828"refString="Bentham, G. & Hooker, J. D. (1867) Genera planta rum. Vols. 1 - 2. Reeve, Williams & Norgate, London."type="book"year="1867">Bentham & Hooker (1867: 764)</bibRefCitation>
