304 lines
39 KiB
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304 lines
39 KiB
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<document id="160259A5F68DC2B42F92FACFAD746980" ID-CLB-Dataset="31499" ID-DOI="10.11646/zootaxa.4079.5.4" ID-GBIF-Dataset="2dec43b9-8411-461a-86a4-2031bd607a92" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050881" ID-ZooBank="0CD47C4D-95D3-4615-BB05-E319E040250D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510811756265" checkinUser="plazi" docAuthor="Niitsuma, Hiromi" docDate="2016" docId="C877FC1F5520FFCD7EDEF94FFC3B81BC" docLanguage="en" docName="zootaxa.4079.5.4.pdf" docOrigin="Zootaxa 4079 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nilothauma hibaratertium Sasa 1993" docType="treatment" docVersion="9" lastPageNumber="578" masterDocId="344E84675524FFC87E49FFCBFFB48620" masterDocTitle="Two species of Nilothauma Kieffer (Diptera, Chironomidae) from Japan, with description of a new species" masterLastPageNumber="581" masterPageNumber="573" pageNumber="577" updateTime="1698496518923" updateUser="ExternalLinkService">
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<mods:title id="6914D8167591C6A4B8C48F4D57C5AE0A">Two species of Nilothauma Kieffer (Diptera, Chironomidae) from Japan, with description of a new species</mods:title>
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<mods:namePart id="1C85DE29A14E68495DC9713D01C5E7DF">Niitsuma, Hiromi</mods:namePart>
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<mods:title id="DD846AFE47D98E6D2688D7B9FE8D5F27">Zootaxa</mods:title>
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<mods:date id="007D2BCF27DA5AAE149F46A31C266870">2016</mods:date>
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<mods:number id="5EFE76AE704F2CE4B71B8874EFDC9420">2016-02-16</mods:number>
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<mods:number id="5A8502D37069754036E8486BB6238DC3">4079</mods:number>
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<mods:number id="CFA115B7FA7F6FA1ECCA01665B43DBCD">5</mods:number>
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<mods:identifier id="AA8E2FB866C03EACE9FE587F9EC153B0" type="DOI">10.11646/zootaxa.4079.5.4</mods:identifier>
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<treatment id="C877FC1F5520FFCD7EDEF94FFC3B81BC" ID-DOI="http://doi.org/10.5281/zenodo.6088558" ID-GBIF-Taxon="137307472" ID-Zenodo-Dep="6088558" LSID="urn:lsid:plazi:treatment:C877FC1F5520FFCD7EDEF94FFC3B81BC" httpUri="http://treatment.plazi.org/id/C877FC1F5520FFCD7EDEF94FFC3B81BC" lastPageId="5" lastPageNumber="578" pageId="4" pageNumber="577">
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<subSubSection id="08C41E825520FFCC7EDEF94FFEFE809F" pageId="4" pageNumber="577" type="nomenclature">
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<paragraph id="40614D095520FFCC7EDEF94FFDA380BE" blockId="4.[151,535,1668,1727]" box="[151,535,1668,1694]" pageId="4" pageNumber="577">
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<heading id="1B29FA655520FFCC7EDEF94FFDA380BE" bold="true" box="[151,535,1668,1694]" fontSize="11" level="1" pageId="4" pageNumber="577" reason="1">
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<taxonomicName id="87DE368A5520FFCC7EDEF94FFDA380BE" ID-CoL="9HML8" authority="Sasa" authorityName="Sasa" authorityYear="1993" box="[151,535,1668,1694]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="hibaratertium">
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<emphasis id="72AA911B5520FFCC7EDEF94FFDA380BE" bold="true" box="[151,535,1668,1694]" pageId="4" pageNumber="577">
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<emphasis id="72AA911B5520FFCC7EDEF94FFE6380BE" bold="true" box="[151,471,1668,1694]" italics="true" pageId="4" pageNumber="577">Nilothauma hibaratertium</emphasis>
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Sasa
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="40614D095520FFCC7EDEF96DFEFE809F" blockId="4.[151,535,1668,1727]" box="[151,330,1702,1727]" pageId="4" pageNumber="577">
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(
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<figureCitation id="D8E5518C5520FFCC7ED7F96DFEF6809F" box="[158,322,1702,1727]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="4" pageNumber="577">Figures 10–13</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="08C41E825520FFCC7EDEF925FD1B8161" pageId="4" pageNumber="577" type="reference_group">
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<paragraph id="40614D095520FFCC7EDEF925FD858124" blockId="4.[151,736,1774,1858]" box="[151,561,1774,1796]" pageId="4" pageNumber="577">
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<treatmentCitationGroup id="60CE6A275520FFCC7EDEF925FD858124" box="[151,561,1774,1796]" pageId="4" pageNumber="577">
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<taxonomicName id="87DE368A5520FFCC7EDEF925FD988124" ID-CoL="9HML9" authority="Sasa, 1993: 73" authorityName="Sasa" authorityPageNumber="73" authorityYear="1993" box="[151,556,1774,1796]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="hibaratertia">
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<emphasis id="72AA911B5520FFCC7EDEF925FE3B8124" box="[151,399,1774,1796]" italics="true" pageId="4" pageNumber="577">Nilothauma hibaratertia</emphasis>
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<treatmentCitation id="C17F6B185520FFCC7FDCF925FD988124" author="Sasa" box="[405,556,1774,1796]" page="73" pageId="4" pageNumber="577" year="1993">
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<bibRefCitation id="244F30F85520FFCC7FDCF925FDB28124" author="Sasa" box="[405,518,1774,1796]" pageId="4" pageNumber="577" refString="Sasa, M. (1993) Studies on the chironomid midges (yusurika) collected in Toyama and other areas of Japan, 1993. Part 5. The chironomids collected from lakes in the Aizu district (Fukushima). Research Report from Toyama Prefectural environmental Pollution Research Centre, 1993, 69 - 95." type="journal article" year="1993">Sasa, 1993</bibRefCitation>
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: 73
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="40614D095520FFCC7EDEF8C6FD1B8161" blockId="4.[151,736,1774,1858]" pageId="4" pageNumber="577">
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<treatmentCitationGroup id="60CE6A275520FFCC7EDEF8C6FD1B8161" pageId="4" pageNumber="577">
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<taxonomicName id="87DE368A5520FFCC7EDEF8C6FD688103" authority="Sasa, Suzuki & Sakai, 1998: 52" authorityName="Sasa, Suzuki & Sakai" authorityPageNumber="52" authorityYear="1998" box="[151,732,1805,1827]" class="Insecta" family="Chironomidae" genus="Tosayusurika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="simantofea">
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<emphasis id="72AA911B5520FFCC7EDEF8C6FE268103" box="[151,402,1805,1827]" italics="true" pageId="4" pageNumber="577">Tosayusurika simantofea</emphasis>
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<treatmentCitation id="C17F6B185520FFCC7FD0F8C6FD688103" author="Sasa" box="[409,732,1805,1827]" page="52" pageId="4" pageNumber="577" year="1998">
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<bibRefCitation id="244F30F85520FFCC7FD0F8C6FD028103" author="Sasa" box="[409,694,1805,1827]" pageId="4" pageNumber="577" refString="Sasa, M., Suzuki, H. & Sakai, T. (1998) Studies on the chironomid midges collected on the shore of Shimanto River in April, 1998. Part 1. Description of species of the subfamily Chironominae. Tropical Medicine, 40, 47 - 89." type="journal article" year="1998">Sasa, Suzuki & Sakai, 1998</bibRefCitation>
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: 52
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</treatmentCitation>
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</taxonomicName>
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.
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<taxonomicName id="87DE368A5520FFCC7EDEF8E7FDEF8162" ID-CoL="92V5Y" authority="Adam & Saether, 1999: 61" authorityName="Adam & Saether. Recently I" authorityPageNumber="61" authorityYear="1999" box="[151,603,1836,1858]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="sasai">
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<emphasis id="72AA911B5520FFCC7EDEF8E7FEFD8162" box="[151,329,1836,1858]" italics="true" pageId="4" pageNumber="577">Nilothauma sasai</emphasis>
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<treatmentCitation id="C17F6B185520FFCC7F06F8E7FDEF8162" author="Adam" box="[335,603,1836,1858]" page="61" pageId="4" pageNumber="577" year="1999">
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<bibRefCitation id="244F30F85520FFCC7F06F8E7FD828162" author="Adam" box="[335,566,1836,1858]" pageId="4" pageNumber="577" refString="Adam, J. I. & Saether, O. A. (1999) Revision of the genus Nilothauma Kieffer, 1921 (Diptera: Chironomidae). Entomologica scandinavica, 56 (Supplement), 1 - 107." type="journal article" year="1999">Adam & Saether, 1999</bibRefCitation>
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: 61
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</treatmentCitation>
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</taxonomicName>
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.
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<emphasis id="72AA911B5520FFCC7C2CF8E0FD1B8161" bold="true" box="[613,687,1835,1858]" pageId="4" pageNumber="577">Syn. n.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="08C41E825520FFCD7EDEF8A5FEAC87D5" lastPageId="5" lastPageNumber="578" pageId="4" pageNumber="577" type="materials_examined">
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<paragraph id="40614D095520FFCD7EDEF8A5FEAC87D5" blockId="4.[151,1436,1902,2035]" lastBlockId="5.[151,1437,151,1949]" lastPageId="5" lastPageNumber="578" pageId="4" pageNumber="577">
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<emphasis id="72AA911B5520FFCC7EDEF8A5FE3181A7" bold="true" box="[151,389,1902,1927]" pageId="4" pageNumber="577">Material examined.</emphasis>
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Holotype of
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<taxonomicName id="87DE368A5520FFCC7C6EF8A5FCE781A7" authorityName="Sasa" authorityYear="1993" box="[551,851,1902,1927]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="hibaratertium">
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<emphasis id="72AA911B5520FFCC7C6EF8A5FCE781A7" box="[551,851,1902,1927]" italics="true" pageId="4" pageNumber="577">Nilothauma hibaratertium</emphasis>
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</taxonomicName>
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: male (NSMT-I-Dip-4842), labelled, “
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<taxonomicName id="87DE368A5520FFCC7B51F8A5FEE1818A" authority="No." authorityName="No." class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="hibaratertia">
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<emphasis id="72AA911B5520FFCC7B51F8A5FEAA818B" italics="true" pageId="4" pageNumber="577">Nilothauma hibaratertia</emphasis>
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No.
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</taxonomicName>
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223: 28”, light trap, JAPAN: Fukushima, Aizu, Bandai-Asahi National Park, Lake Hibara,
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<date id="34606BC95520FFCC7EDEF87DFEA781EF" box="[151,275,1974,1999]" pageId="4" pageNumber="577" value="1991-08-06">6.viii.1991</date>
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. Paratypes of
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<taxonomicName id="87DE368A5520FFCC7FFDF87DFD6881EF" authorityName="Sasa" authorityYear="1993" box="[436,732,1974,1999]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="hibaratertium">
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<emphasis id="72AA911B5520FFCC7FFDF87DFD6881EF" box="[436,732,1974,1999]" italics="true" pageId="4" pageNumber="577">Nilothauma hibaratertium</emphasis>
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</taxonomicName>
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: 2 males (NSMT), labelled respectively, “No.223: 29, 30”, as holotype. Holotype of
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<taxonomicName id="87DE368A5520FFCC7FE8F811FD0881D3" authorityName="Sasa, Suzuki & Sakai" authorityYear="1998" box="[417,700,2010,2035]" class="Insecta" family="Chironomidae" genus="Tosayusurika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="simantofea">
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<emphasis id="72AA911B5520FFCC7FE8F811FD0881D3" box="[417,700,2010,2035]" italics="true" pageId="4" pageNumber="577">Tosayusurika simantofea</emphasis>
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</taxonomicName>
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: male (NSMT-I-Dip-5198), labelled, “No. 358: 45”, light trap, JAPAN: Kochi, Nakamura, Shimanto River,
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<date id="34606BC95521FFCD7CC4FF5CFCB38690" box="[653,775,151,176]" pageId="5" pageNumber="578" value="1998-04-26">26.iv.1998</date>
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. Paratypes of
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<taxonomicName id="87DE368A5521FFCD7DEDFF5CFB0E8690" authorityName="Sasa, Suzuki & Sakai" authorityYear="1998" box="[932,1210,151,176]" class="Insecta" family="Chironomidae" genus="Tosayusurika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="simantofea">
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<emphasis id="72AA911B5521FFCD7DEDFF5CFB0E8690" box="[932,1210,151,176]" italics="true" pageId="5" pageNumber="578">Tosayusurika simantofea</emphasis>
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</taxonomicName>
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: 12 males (NSMT), labelled respectively, “No. 359: 51–54, 360: 43, 361: 1–7”, as holotype. Holotype of
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<taxonomicName id="87DE368A5521FFCD7A1EFF77FAA886F4" authorityName="Adam & Saether. Recently I" authorityYear="1999" box="[1111,1308,188,213]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="sasai">
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<emphasis id="72AA911B5521FFCD7A1EFF77FAA886F4" box="[1111,1308,188,213]" italics="true" pageId="5" pageNumber="578">Nilothauma sasai</emphasis>
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</taxonomicName>
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: male with pupal and larval exuviae (SUM-IC-C0028), reared from larva, JAPAN: Fukushima, Iwaki, Obisa River,
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<date id="34606BC95521FFCD7EDEFECFFEAA873D" box="[151,286,260,285]" pageId="5" pageNumber="578" value="1988-08-15">15.viii.1988</date>
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(emerged
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<date id="34606BC95521FFCD7FE9FECFFDA8873D" box="[416,540,260,285]" pageId="5" pageNumber="578" value="1988-09-12">12.ix.1988</date>
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). Non-types: male with pupal exuviae (SUM-IC-C0031), reared from pupa, JAPAN: Fukushima, Hirono, Asami River,
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<date id="34606BC95521FFCD7CCEFEECFCBA8760" box="[647,782,295,320]" pageId="5" pageNumber="578" value="1996-08-15">15.viii.1996</date>
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(emerged
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<date id="34606BC95521FFCD7DC3FEECFBA28760" box="[906,1046,295,320]" pageId="5" pageNumber="578" value="1996-08-29">29.viii.1996</date>
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); male with pupal exuviae (SUM- IC-C0032), as previous except
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<date id="34606BC95521FFCD7C44FE87FD208745" box="[525,660,332,357]" pageId="5" pageNumber="578" value="2001-08-15">15.viii.2001</date>
|
|||
|
|
(emerged
|
|||
|
|
<date id="34606BC95521FFCD7D51FE87FC328745" box="[792,902,332,357]" pageId="5" pageNumber="578" value="2001-09-03">3.ix.2001</date>
|
|||
|
|
); male (SUM-IC-C0033), as previous except
|
|||
|
|
<date id="34606BC95521FFCD7EDEFEA4FEAA87A8" box="[151,286,367,392]" pageId="5" pageNumber="578" value="2002-08-15">15.viii.2002</date>
|
|||
|
|
(emerged
|
|||
|
|
<date id="34606BC95521FFCD7FD0FEA4FD9187A8" box="[409,549,367,392]" pageId="5" pageNumber="578" value="2002-08-24">24.viii.2002</date>
|
|||
|
|
); 2 males with 1 pupal exuviae (SUM-IC-C0 0 34, C0035), as previous except
|
|||
|
|
<date id="34606BC95521FFCD7EDEFE5FFF43878D" box="[151,247,404,429]" pageId="5" pageNumber="578" value="2005-05-04">4.v.2005</date>
|
|||
|
|
(emerged
|
|||
|
|
<date id="34606BC95521FFCD7F27FE5EFDA1878D" box="[366,533,404,429]" pageId="5" pageNumber="578" value="2005-05-07" valueMax="2005-05-09" valueMin="2005-05-07">7 and 9.v.2005</date>
|
|||
|
|
); male (SUM-IC-C0036), reared from pupa, JAPAN: Fukushima, Iwaki, Yaguki,
|
|||
|
|
<date id="34606BC95521FFCD7EDEFE7CFF4687F0" box="[151,242,439,464]" pageId="5" pageNumber="578" value="2013-01-05">5.i.2013</date>
|
|||
|
|
(emerged
|
|||
|
|
<date id="34606BC95521FFCD7F3AFE7CFE5587F0" box="[371,481,439,464]" pageId="5" pageNumber="578" value="2013-01-26">26.i.2013</date>
|
|||
|
|
); male (MSU-IC-C0037), sweep net, JAPAN: Shizuoka, Shimizu, Yanbara River,
|
|||
|
|
<date id="34606BC95521FFCD7EDEFE17FEA087D5" box="[151,276,476,501]" pageId="5" pageNumber="578" value="2002-08-07">7.viii.2002</date>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="08C41E825521FFCD7E8EFE34FAA68335" pageId="5" pageNumber="578" type="description">
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFE34FCCB8437" blockId="5.[151,1437,151,1949]" box="[199,895,511,536]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFE34FE288438" bold="true" box="[199,412,511,536]" pageId="5" pageNumber="578">Description. Male</emphasis>
|
|||
|
|
(n = 24). Total length 2.2–4.2, 2.9 (8) mm.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFDEFFC768440" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFDEFFEFD841D" box="[199,329,548,573]" italics="true" pageId="5" pageNumber="578">Coloration.</emphasis>
|
|||
|
|
Generally green. Wing without any marking. Foreleg pale yellow with base and sub-apex of femora, apices of tibia and tarsomere 1, and tarsomeres 2–5 dark brown.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFDA7FAA98488" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFDA7FEB084A5" box="[199,260,620,645]" italics="true" pageId="5" pageNumber="578">Head</emphasis>
|
|||
|
|
. Temporals 8–14, 11. Frontal tubercles absent. AR 0.24–0.31, 0.26 (22). Palpomere 3 apically with 2 sensilla clavata, longest 23–28, 26 (18) µm long. Pm4/Pm3 1.3–1.6, 1.5 (21); Pm5/Pm4 1.2–1.4, 1.3 (21).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFD7DFA8084EF" blockId="5.[151,1437,151,1949]" box="[199,1332,694,719]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFD7DFEAC84EF" box="[199,280,694,719]" italics="true" pageId="5" pageNumber="578">Thorax</emphasis>
|
|||
|
|
. Acrostichals 14–23, 17; dorsocentrals 8–16, 12 (23); prealars 3–5, 4. Scutellum with 2 setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFD17FCFE8537" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFD17FEBC84D3" box="[199,264,732,755]" italics="true" pageId="5" pageNumber="578">Wing.</emphasis>
|
|||
|
|
Length 1.3–2.2, 1.9 (23) mm. VR 1.3–1.5, 1.4 (23). Brachiolum with 1 (8) seta; R with 9–20, 14 (8) setae; R1 with 7–16, 12 (8) setae; R4+5 with 2–19, 10 (8) setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFCEDFBB58542" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFCEDFEB7851D" box="[199,259,806,829]" italics="true" pageId="5" pageNumber="578">Legs.</emphasis>
|
|||
|
|
Mid tarsomere 1 with 1–5, 3 (20) sensilla chaetica, distalmost located 0.14–0.31, 0.21 (20) from apex. LR1 1.37–1.54, 1.46 (16); LR2 0.54–0.62, 0.58 (23); LR3 0.52–0.58, 0.56 (23).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFCBAFAA68335" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFCBAFEE285A8" box="[199,342,881,904]" italics="true" pageId="5" pageNumber="578">Hypopygium</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="D8E5518C5521FFCD7F2CFCBBFDBE85A9" box="[357,522,880,905]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="5" pageNumber="578">Figures 10, 11</figureCitation>
|
|||
|
|
). Tergite IX with 2 dorsal projections and 9–17, 13 (10) posterior setae. Anterior projection (
|
|||
|
|
<figureCitation id="D8E5518C5521FFCD7F52FC5FFE74858D" box="[283,448,916,941]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="5" pageNumber="578">Figures 12a–c</figureCitation>
|
|||
|
|
) 18–38, 28 (22) µm long, 0.75–1.2, 0.95 (22) times as long as broad, varying from simple lobe to two distinctly divided lobes, usually notched posteriorly, with about 20–50 plumose setae 40–59, 50 (22) µm long. Posterior projection 22–38, 30 (13) µm long, 0.73–1.2, 0.89 (11) times as long as basal width, basally microtrichiose, apically with 6–15, 9 (15) setae 12–20, 16 (17) µm long. Anal point 55–63, 59 (6) µm long, 1.8–1.9, 1.9 (6) times as long as broad, without microtrichia. Laterosternite IX with 2–7, 4 setae. Phallapodeme 50– 75, 64 (20) µm long. Sternapodeme bluntly triangular. Gonocoxite 88–125, 114 (22) µm long. Superior volsella (
|
|||
|
|
<figureCitation id="D8E5518C5521FFCD7ED7FBA7FEA482A5" box="[158,272,1132,1157]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="5" pageNumber="578">Figure 13</figureCitation>
|
|||
|
|
) pad-like, 37–50, 44 (22) µm long, microtrichiose with 1 long lateral and 5–8, 6 (21) short marginal setae on small tubercles; lateral seta 25–42, 32 (19) µm long. Median volsella 5–8, 7 (23) µm long, with 2–4, 3 (23) apical setae on small tubercles. Inferior volsella microtrichiose, with 7–10, 8 apically split setae. Gonostylus 105– 168, 149 (22) µm long, 5.3–5.9, 5.6 (6) times as long as broad at middle, parallel-sided for most of its length, with pointed apex; distal-median setae 6–10, 8 (22), uniserial, some split apically. HR 0.71–0.83, 0.77 (22).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="08C41E825521FFCD7E8EFAEBFC3B81BC" pageId="5" pageNumber="578" type="discussion">
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EFAEBFCF18078" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7E8EFAEBFE8F8319" bold="true" box="[199,315,1312,1337]" pageId="5" pageNumber="578">Remarks.</emphasis>
|
|||
|
|
The male of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7F9DFAEBFC158319" authority="Adam & Saether, 1999" authorityName="Adam & Saether. Recently I" authorityYear="1999" box="[468,929,1312,1337]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="sasai">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7F9DFAEBFD2E8318" box="[468,666,1312,1337]" italics="true" pageId="5" pageNumber="578">Nilothauma sasai</emphasis>
|
|||
|
|
<bibRefCitation id="244F30F85521FFCD7CEAFAEBFC158319" author="Adam" box="[675,929,1312,1337]" pageId="5" pageNumber="578" refString="Adam, J. I. & Saether, O. A. (1999) Revision of the genus Nilothauma Kieffer, 1921 (Diptera: Chironomidae). Entomologica scandinavica, 56 (Supplement), 1 - 107." type="journal article" year="1999">Adam & Saether, 1999</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was separated from that of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7A97FAEAFEA0837D" authority="Sasa, 1993" authorityName="Sasa" authorityYear="1993" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7A97FAEAFA288319" box="[1246,1436,1312,1337]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
<bibRefCitation id="244F30F85521FFCD7EDEFA8EFEA0837D" author="Sasa" box="[151,276,1349,1373]" pageId="5" pageNumber="578" refString="Sasa, M. (1993) Studies on the chironomid midges (yusurika) collected in Toyama and other areas of Japan, 1993. Part 5. The chironomids collected from lakes in the Aizu district (Fukushima). Research Report from Toyama Prefectural environmental Pollution Research Centre, 1993, 69 - 95." type="journal article" year="1993">Sasa, 1993</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the simple anterior projection on tergite IX (see the key to male imagines, fig. 23F in
|
|||
|
|
<bibRefCitation id="244F30F85521FFCD7B71FA8FFE9D83A1" author="Adam" pageId="5" pageNumber="578" refString="Adam, J. I. & Saether, O. A. (1999) Revision of the genus Nilothauma Kieffer, 1921 (Diptera: Chironomidae). Entomologica scandinavica, 56 (Supplement), 1 - 107." type="journal article" year="1999">Adam & Saether 1999</bibRefCitation>
|
|||
|
|
). However, re-examination of the
|
|||
|
|
<typeStatus id="9F65F3AB5521FFCD7CF8FAA3FCA783A1" box="[689,787,1384,1409]" pageId="5" pageNumber="578" type="holotype">holotype</typeStatus>
|
|||
|
|
male of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7D32FAA2FC6C83A0" authorityName="Adam & Saether. Recently I" authorityYear="1999" box="[891,984,1385,1408]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="sasai">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7D32FAA2FC6C83A0" box="[891,984,1385,1408]" italics="true" pageId="5" pageNumber="578">N. sasai</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
disclosed that the anterior projection is completely split into two lobes in conflict with the original description. I re-examined the following
|
|||
|
|
<typeStatus id="9F65F3AB5521FFCD7B4CFA46FA818385" box="[1285,1333,1421,1445]" pageId="5" pageNumber="578">type</typeStatus>
|
|||
|
|
series of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7EDEFA7AFEEC83E9" authorityName="Sasa" authorityYear="1993" box="[151,344,1456,1481]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7EDEFA7AFEEC83E9" box="[151,344,1456,1481]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7FDFFA7BFBB383E9" authority="Sasa, Suzuki & Sakai, 1998" authorityName="Sasa, Suzuki & Sakai" authorityYear="1998" box="[406,1031,1456,1481]" class="Insecta" family="Chironomidae" genus="Tosayusurika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="simantofea">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7FDFFA7BFD0483E9" box="[406,688,1456,1481]" italics="true" pageId="5" pageNumber="578">Tosayusurika simantofea</emphasis>
|
|||
|
|
<bibRefCitation id="244F30F85521FFCD7CF5FA7AFBB383E9" author="Sasa" box="[700,1031,1456,1481]" pageId="5" pageNumber="578" refString="Sasa, M., Suzuki, H. & Sakai, T. (1998) Studies on the chironomid midges collected on the shore of Shimanto River in April, 1998. Part 1. Description of species of the subfamily Chironominae. Tropical Medicine, 40, 47 - 89." type="journal article" year="1998">Sasa, Suzuki & Sakai, 1998</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, synonymized by
|
|||
|
|
<bibRefCitation id="244F30F85521FFCD7A97FA7BFF5283CD" author="Adam" pageId="5" pageNumber="578" refString="Adam, J. I. & Saether, O. A. (1999) Revision of the genus Nilothauma Kieffer, 1921 (Diptera: Chironomidae). Entomologica scandinavica, 56 (Supplement), 1 - 107." type="journal article" year="1999">Adam & Saether (1999)</bibRefCitation>
|
|||
|
|
: three male specimens of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7C5AFA1EFD6583CD" authorityName="Sasa" authorityYear="1993" box="[531,721,1492,1517]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7C5AFA1EFD6583CD" box="[531,721,1492,1517]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="56D886805521FFCD7D45FA1EFCDC83CD" box="[780,872,1492,1517]" pageId="5" pageNumber="578" type="male">13 male</specimenCount>
|
|||
|
|
specimens of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7A59FA1FFA9C83CD" authorityName="Sasa, Suzuki & Sakai" authorityYear="1998" box="[1040,1320,1492,1517]" class="Insecta" family="Chironomidae" genus="Tosayusurika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="simantofea">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7A59FA1FFA9C83CD" box="[1040,1320,1492,1517]" italics="true" pageId="5" pageNumber="578">Tosayusurika simantofea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
including each
|
|||
|
|
<typeStatus id="9F65F3AB5521FFCD7E99FA33FE828031" box="[208,310,1528,1553]" pageId="5" pageNumber="578" type="holotype">holotype</typeStatus>
|
|||
|
|
. Consequently, it was revealed that the anterior projection in the
|
|||
|
|
<typeStatus id="9F65F3AB5521FFCD7A4FFA32FBD28031" box="[1030,1126,1529,1553]" pageId="5" pageNumber="578" type="paratype">paratype</typeStatus>
|
|||
|
|
No. 359: 3 of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7B4FFA32FA288031" authorityName="Sasa, Suzuki & Sakai" authorityYear="1998" box="[1286,1436,1529,1553]" class="Insecta" family="Chironomidae" genus="Tosayusurika" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="simantofea">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7B4FFA32FA288031" box="[1286,1436,1529,1553]" italics="true" pageId="5" pageNumber="578">T. simantofea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is simple, not divided into two lobes. These show that the simple anterior projection is an individual variation in the species and
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7F57F98AFECD8078" authorityName="Adam & Saether. Recently I" authorityYear="1999" box="[286,377,1601,1624]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="sasai">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7F57F98AFECD8078" box="[286,377,1601,1624]" italics="true" pageId="5" pageNumber="578">N. sasai</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a junior synonym of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7CCBF98AFC8B8079" authorityName="Sasa" authorityYear="1993" box="[642,831,1600,1625]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7CCBF98AFC8B8079" box="[642,831,1600,1625]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="40614D095521FFCD7E8EF9AEFCA8812C" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
|||
|
|
I examined
|
|||
|
|
<specimenCount id="56D886805521FFCD7F02F9AEFE1B805D" box="[331,431,1636,1661]" pageId="5" pageNumber="578" type="male">23 males</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7F9DF9AEFD24805D" authorityName="Sasa" authorityYear="1993" box="[468,656,1636,1661]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
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<emphasis id="72AA911B5521FFCD7F9DF9AEFD24805D" box="[468,656,1636,1661]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
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</taxonomicName>
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, including seven newly collected males. They showed great variation in the morphology of the anterior anal tergal projection: in four specimens (17%) the projection is completely divided into two lobes (
|
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<figureCitation id="D8E5518C5521FFCD7FEDF967FD9780E5" box="[420,547,1708,1733]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="5" pageNumber="578">Figure 12a</figureCitation>
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), in
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<specimenCount id="56D886805521FFCD7C1EF967FD4480E5" box="[599,752,1708,1733]" pageId="5" pageNumber="578" type="generic">16 specimens</specimenCount>
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(70%) the projection is notched to a varying degree (
|
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<figureCitation id="D8E5518C5521FFCD7B18F967FF7080C9" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="5" pageNumber="578">Figure 12b</figureCitation>
|
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), and in three specimens (13%) the projection is simple (
|
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<figureCitation id="D8E5518C5521FFCD7D0DF91BFC7780C9" box="[836,963,1744,1769]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1286,1309]" captionTargetBox="[276,1303,210,1241]" captionTargetId="figure@6.[266,1321,195,1265]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Nilothauma hibaratertium Sasa, male. 10, hypopygium, ventral view (right) and dorsal view (left); 11, anal tergite with anal point, lateral view; 12 a – c, anterior projection on anal tergite, dorsal view, showing variation; 13, superior volsella, dorsal view." httpUri="https://zenodo.org/record/1051816/files/figure.png" pageId="5" pageNumber="578">Figure 12c</figureCitation>
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). The original description of
|
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<taxonomicName id="87DE368A5521FFCD7B43F91AFAD080C8" authorityName="Adam & Saether. Recently I" authorityYear="1999" box="[1290,1380,1745,1768]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="sasai">
|
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<emphasis id="72AA911B5521FFCD7B43F91AFAD080C8" box="[1290,1380,1745,1768]" italics="true" pageId="5" pageNumber="578">N. sasai</emphasis>
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</taxonomicName>
|
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may be based on another loaned specimen, not returned to me.
|
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</paragraph>
|
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<paragraph id="40614D095521FFCD7E8EF8D3FC3B81BC" blockId="5.[151,1437,151,1949]" pageId="5" pageNumber="578">
|
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<bibRefCitation id="244F30F85521FFCD7E8EF8D3FDA98111" author="Adam" box="[199,541,1816,1841]" pageId="5" pageNumber="578" refString="Adam, J. I. & Saether, O. A. (1999) Revision of the genus Nilothauma Kieffer, 1921 (Diptera: Chironomidae). Entomologica scandinavica, 56 (Supplement), 1 - 107." type="journal article" year="1999">Adam & Saether (1999: 71)</bibRefCitation>
|
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|
regarded the Portuguese species illustrated under the name of
|
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<taxonomicName id="87DE368A5521FFCD7B7EF8D2FE8D8175" authority="(Goetghebuer)" authorityName="Goetghebuer" baseAuthorityName="Goetghebuer" baseAuthorityYear="1921" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="brayi">
|
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|
|
<emphasis id="72AA911B5521FFCD7B7EF8D2FA2F8111" box="[1335,1435,1816,1841]" italics="true" pageId="5" pageNumber="578">N. brayi</emphasis>
|
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|
(Goetghebuer)
|
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|
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</taxonomicName>
|
|||
|
|
by
|
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|
|
<bibRefCitation id="244F30F85521FFCD7F2AF8F6FDF98175" author="Cranston" box="[355,589,1852,1877]" pageId="5" pageNumber="578" refString="Cranston, P. S., Dillon, M. E., Pinder, L. C. V. & Reiss, F. (1989) 10. The adult males of Chironominae (Diptera: Chironomidae) of the Holarctic region - Keys and diagnoses. In: Wiederholm, T. (Ed.), Chironomidae of the Holarctic region. Keys and diagnoses. Part 3. Adult males. Entomologica scandinavica, 34 (Supplement), 353 - 502." type="journal article" year="1989">
|
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|
|
Cranston
|
|||
|
|
<emphasis id="72AA911B5521FFCD7F87F8F6FE4B8175" box="[462,511,1852,1877]" italics="true" pageId="5" pageNumber="578">et al</emphasis>
|
|||
|
|
. (1989
|
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|
|
</bibRefCitation>
|
|||
|
|
fig. 10.41) as
|
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|
|
<taxonomicName id="87DE368A5521FFCD7CA5F8F6FC1C8175" authorityName="Sasa" authorityYear="1993" box="[748,936,1852,1877]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7CA5F8F6FC1C8175" box="[748,936,1852,1877]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. However, the species may be different from
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7EDEF8AAFEE28159" authorityName="Sasa" authorityYear="1993" box="[151,342,1888,1913]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7EDEF8AAFEE28159" box="[151,342,1888,1913]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
because of the stout and triangular posterior projection on the male anal tergite. The posterior projection is relatively slender and club-shaped in
|
|||
|
|
<taxonomicName id="87DE368A5521FFCD7C85F84EFC3C81BD" authorityName="Sasa" authorityYear="1993" box="[716,904,1924,1949]" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="hibaratertium">
|
|||
|
|
<emphasis id="72AA911B5521FFCD7C85F84EFC3C81BD" box="[716,904,1924,1949]" italics="true" pageId="5" pageNumber="578">N. hibaratertium</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
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</subSubSection>
|
|||
|
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</treatment>
|
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</document>
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