treatments-xml/data/03/9B/55/039B5575FF9DDF7A3956FD12FE1AFD82.xml

<document id="CD5C1E09BEBAD98034A696559F3FDE37" ID-DOI="10.25674/SO94iSS3iD303" ID-ISSN="2509-9523" ID-Zenodo-Dep="13193962" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631614709" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Mainda, Tobias" docDate="2022" docId="039B5575FF9DDF7A3956FD12FE1AFD82" docLanguage="en" docName="SoilOrg.94.3.139-147.pdf" docOrigin="Soil Organisms 94 (3)" docSource="http://dx.doi.org/10.25674/so94iss3id303" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus arfakmontium Frisch &amp; Mainda, 2022, spec. nov." docType="treatment" docVersion="3" lastPageNumber="143" masterDocId="FFA22D0DFF9EDF7E3A19FFECFFA6FFF1" masterDocTitle="The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species" masterLastPageNumber="147" masterPageNumber="139" pageNumber="142" updateTime="1722679412379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6EDD1A80D9D21A9BF0411D15AB222BB6">The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species</mods:title>
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<mods:namePart id="AC3811B871C429E338B1573A5BEB62D8">Frisch, Johannes</mods:namePart>
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<mods:namePart id="72EFEBCD8BD8FB2B0B1615277109A3D4">Mainda, Tobias</mods:namePart>
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<mods:title id="218437296C7737B08E3BE4A49DF9317D">Soil Organisms</mods:title>
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<mods:date id="7A41F6DA6750F9F61CB8BFC75E64A1F1">2022</mods:date>
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<paragraph id="8B8DE463FF9DDF7D3956FD12FB53FCE9" blockId="3.[847,1269,765,825]" box="[847,1269,765,792]" pageId="3" pageNumber="142">
<heading id="D0C5530FFF9DDF7D3956FD12FB53FCE9" box="[847,1269,765,792]" fontSize="11" level="2" pageId="3" pageNumber="142" reason="5">
<emphasis id="B9463871FF9DDF7D3956FD12FB53FCE9" bold="true" box="[847,1269,765,792]" pageId="3" pageNumber="142">
<taxonomicName id="4C329FE0FF9DDF7D3956FD12FBD0FCE9" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[847,1142,765,792]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="arfakmontium" status="sp. nov.">
<emphasis id="B9463871FF9DDF7D3956FD12FBD0FCE9" bold="true" box="[847,1142,765,792]" italics="true" pageId="3" pageNumber="142">Scopaeus arfakmontium</emphasis>
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<taxonomicNameLabel id="A275850AFF9DDF7D3E67FD12FB53FCE9" box="[1150,1269,766,792]" pageId="3" pageNumber="142" rank="species">spec. nov.</taxonomicNameLabel>
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<subSubSection id="C328B7E8FF9DDF7D3956FCCCFBE1FCC8" box="[847,1095,800,825]" pageId="3" pageNumber="142" type="description">
<paragraph id="8B8DE463FF9DDF7D3956FCCCFBE1FCC8" blockId="3.[847,1269,765,825]" box="[847,1095,800,825]" pageId="3" pageNumber="142">
(
<figureCitation id="1309F8E6FF9DDF7D394FFCCCFC3AFCC8" box="[854,924,800,825]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="3" pageNumber="142">Figs 2</figureCitation>
, 
<figureCitation id="1309F8E6FF9DDF7D39BEFCCCFC13FCC9" box="[935,949,800,824]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="3" pageNumber="142">5</figureCitation>
, 
<figureCitation id="1309F8E6FF9DDF7D39D8FCCDFC51FCC8" box="[961,1015,801,825]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="3" pageNumber="142">8–10</figureCitation>
, 
<figureCitation id="1309F8E6FF9DDF7D3E1BFCCDFB99FCC8" box="[1026,1087,801,825]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="3" pageNumber="142">17, 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9DDF7D3956FC8EFA31FC4D" pageId="3" pageNumber="142" type="materials_examined">
<paragraph id="8B8DE463FF9DDF7D3956FC8EFA31FC4D" blockId="3.[824,1437,866,1975]" pageId="3" pageNumber="142">
<emphasis id="B9463871FF9DDF7D3956FC8EFBA5FC8B" bold="true" box="[847,1027,866,890]" pageId="3" pageNumber="142">Type specimens</emphasis>
: 
<materialsCitation id="3B5AEE3EFF9DDF7D3E0BFC8EFA3FFC6A" collectingDate="2011-08-02" collectionCode="TWCF" collectorName="Watanabe" country="Indonesia" elevation="500" location="Arfak Mts" pageId="3" pageNumber="142" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="54895AC1FF9DDF7D3E0BFC8EFBD0FC8B" box="[1042,1142,866,890]" pageId="3" pageNumber="142" type="holotype">Holotype</typeStatus>
<specimenCount id="9D342FEAFF9DDF7D3E67FC8FFB34FC8B" box="[1150,1170,867,890]" count="1" pageId="3" pageNumber="142" type="male">♂</specimenCount>
, 
<collectingCountry id="F325A4F3FF9DDF7D3E86FC8EFAADFC8B" box="[1183,1291,866,890]" name="Indonesia" pageId="3" pageNumber="142">Indonesia</collectingCountry>
, 
<collectingRegion id="49F62A81FF9DDF7D3F0FFC8FFA3FFC8B" box="[1302,1433,867,890]" country="Indonesia" name="Papua Barat" pageId="3" pageNumber="142">West Papua</collectingRegion>
: 
<location id="8EEDB2B8FF9DDF7D3921FC6FFC08FC6A" LSID="urn:lsid:plazi:treatment:039B5575FF9DDF7A3956FD12FE1AFD82:8EEDB2B8FF9DDF7D3921FC6FFC08FC6A" box="[824,942,899,923]" country="Indonesia" name="Arfak Mts" pageId="3" pageNumber="142" stateProvince="Papua Barat">Arfak Mts</location>
, 
<quantity id="4CCA4986FF9DDF7D39A5FC6FFBA2FC6A" box="[956,1028,899,923]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="142" unit="m" value="500.0">
<elevation id="001F0350FF9DDF7D39A5FC6FFBA2FC6A" box="[956,1028,899,923]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="142" unit="m" value="500.0">500 m</elevation>
</quantity>
, 
<date id="FF8CC2A3FF9DDF7D3E0AFC6FFB2FFC6A" box="[1043,1161,899,923]" pageId="3" pageNumber="142" value="2011-08-02">
<collectingDate id="EFC83B4BFF9DDF7D3E0AFC6FFB2FFC6A" box="[1043,1161,899,923]" pageId="3" pageNumber="142" value="2011-08-02">08.02.2011</collectingDate>
</date>
, leg. 
<collectorName id="26C781B5FF9DDF7D3EDFFC6FFA97FC6A" box="[1222,1329,899,923]" pageId="3" pageNumber="142">Watanabe</collectorName>
(
<collectionCode id="ED237CA6FF9DDF7D3F5BFC6FFA34FC6A" box="[1346,1426,899,923]" pageId="3" pageNumber="142">TWCF</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B5AEE3EFF9DDF7D3921FC48FA34FC4D" box="[824,1426,932,956]" collectingDate="2011-08-02" collectionCode="MFNB, TWCF" collectorName="Watanabe" country="Indonesia" elevation="500" location="Arfak Mts" pageId="3" pageNumber="142" specimenCount="4" specimenCount-male="4" stateProvince="Papua Barat" typeStatus="paratype">
<typeStatus id="54895AC1FF9DDF7D3921FC48FC00FC4D" box="[824,934,932,956]" pageId="3" pageNumber="142" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D342FEAFF9DDF7D39A9FC48FC7EFC4A" box="[944,984,932,956]" count="4" pageId="3" pageNumber="142" type="male">4 ♂</specimenCount>
, same data as holotype (
<collectionCode id="ED237CA6FF9DDF7D3EF9FC48FA94FC4D" box="[1248,1330,932,956]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="3" pageNumber="142">MFNB</collectionCode>
, 
<collectionCode id="ED237CA6FF9DDF7D3F24FC48FA2DFC4D" box="[1341,1419,932,956]" pageId="3" pageNumber="142">TWCF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9DDF7D3956FC29FC1EF920" pageId="3" pageNumber="142" type="diagnosis">
<paragraph id="8B8DE463FF9DDF7D3956FC29FA93FA78" blockId="3.[824,1437,866,1975]" pageId="3" pageNumber="142">
<emphasis id="B9463871FF9DDF7D3956FC29FC18FC2C" bold="true" box="[847,958,965,989]" pageId="3" pageNumber="142">Diagnosis</emphasis>
: Body shape and coloring as in 
<figureCitation id="1309F8E6FF9DDF7D3F03FC29FAFCFC2C" box="[1306,1370,965,989]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="3" pageNumber="142">Fig. 2</figureCitation>
. Total body length 4.7 – 5.0 mm; forebody length 
<quantity id="4CCA4986FF9DDF7D3F14FC0AFA3FFC0C" box="[1293,1433,998,1022]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="3" pageNumber="142" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7 – 2.8 mm</quantity>
. Head pyriform, 1.13 – 1.15 (
<geoCoordinate id="EE0682A4FF9DDF7D3E6FFBE9FB65FBEE" box="[1142,1219,1029,1055]" degrees="1.14" direction="west" orientation="longitude" pageId="3" pageNumber="142" precision="555" value="-1.14">Ø 1.14</geoCoordinate>
) times as long as wide, across eyes 1.34 – 1.42 (
<geoCoordinate id="EE0682A4FF9DDF7D3E9DFBCAFB76FBCE" box="[1156,1232,1062,1087]" degrees="1.36" direction="west" orientation="longitude" pageId="3" pageNumber="142" precision="555" value="-1.36">Ø 1.36</geoCoordinate>
) times as wide as across tempora at half of their length from eyes to neck constriction. Eyes 0.5 – 0.55 (
<geoCoordinate id="EE0682A4FF9DDF7D3E99FB84FB68FB70" box="[1152,1230,1128,1153]" degrees="0.52" direction="west" orientation="longitude" pageId="3" pageNumber="142" precision="555" value="-0.52">Ø 0.52</geoCoordinate>
) times as long as tempora. Elytra about 1.14 times as long as pronotum. Penultimate segment of antenna about 1.3 times as long as wide. Tibia of mesothoracic leg 9.2 – 11.2 (
<geoCoordinate id="EE0682A4FF9DDF7D3F4AFB26FA33FB15" box="[1363,1429,1226,1252]" degrees="9.7" direction="west" orientation="longitude" pageId="3" pageNumber="142" precision="5555" value="-9.7">Ø 9.7</geoCoordinate>
) times as long as wide. Body black except for dark brown maxillary palpi with light brown apex of last segment, dark brown legs with medium brown tarsi, and antennae becoming evenly lighter from dark brown proximal segments towards light brown apical segments.
</paragraph>
<paragraph id="8B8DE463FF9DDF7D3956FA7DFC1EF920" blockId="3.[824,1437,866,1975]" pageId="3" pageNumber="142">
Male: Aedeagus (
<figureCitation id="1309F8E6FF9DDF7D3E0EFA7DFBFDFA58" box="[1047,1115,1425,1449]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="3" pageNumber="142">Fig. 5</figureCitation>
) about 
<quantity id="4CCA4986FF9DDF7D3EAEFA7DFAA8FA58" box="[1207,1294,1425,1449]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="142" unit="mm" value="0.9">0.9 mm</quantity>
long; apical lobes parallel in lateral view (
<figureCitation id="1309F8E6FF9DDF7D3E6BFA5EFB09FA3B" box="[1138,1199,1458,1482]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="3" pageNumber="142">Fig. 8</figureCitation>
), in ventral and dorsal view (
<figureCitation id="1309F8E6FF9DDF7D3964FA3FFC4AFA1A" box="[893,1004,1491,1515]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="3" pageNumber="142">Figs 9, 10</figureCitation>
) convex laterally; lateral lobes notably longer than wide (
<figureCitation id="1309F8E6FF9DDF7D3E1CFA18FBD1F9FD" box="[1029,1143,1524,1548]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="3" pageNumber="142">Figs 9, 10</figureCitation>
); dorsal lobe triangular in dorsal view with subacute apex (
<figureCitation id="1309F8E6FF9DDF7D3E8DF9F9FB46F9DC" box="[1172,1248,1557,1581]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="3" pageNumber="142">Fig. 10</figureCitation>
). Asetose median depression of abdominal sternite VII (
<figureCitation id="1309F8E6FF9DDF7D3F19F9DAFAF3F9BF" box="[1280,1365,1590,1614]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="3" pageNumber="142">Fig. 17</figureCitation>
) with parallel lateral margins. Abdominal sternite VIII (
<figureCitation id="1309F8E6FF9DDF7D3F6BF9BAFCF7F97E" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="3" pageNumber="142">Fig. 18</figureCitation>
) with median lobe of posterior margin notably longer than lateroposterior angles; end of median lobe notably emarginate.
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9DDF7A3956F936FD5AFDE0" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142" type="distribution">
<paragraph id="8B8DE463FF9DDF7D3956F936FAD2F8C2" blockId="3.[824,1437,866,1975]" pageId="3" pageNumber="142">
<emphasis id="B9463871FF9DDF7D3956F936FC7AF903" bold="true" box="[847,988,1754,1778]" pageId="3" pageNumber="142">Distribution</emphasis>
: 
<taxonomicName id="4C329FE0FF9DDF7D39E1F936FAAAF903" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1016,1292,1754,1778]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF9DDF7D39E1F936FAAAF903" box="[1016,1292,1754,1778]" italics="true" pageId="3" pageNumber="142">Scopaeus arfakmontium</emphasis>
</taxonomicName>
is hitherto known only from the 
<typeStatus id="54895AC1FF9DDF7D3E03F917FBECF8E2" box="[1050,1098,1787,1811]" pageId="3" pageNumber="142">type</typeStatus>
locality in the Arfak Mountains in the east of the Bird’s Head Peninsula, 
<collectingRegion id="49F62A81FF9DDF7D3EF7F8F0FAC9F8C2" box="[1262,1391,1820,1843]" country="Indonesia" name="Papua Barat" pageId="3" pageNumber="142">West Papua</collectingRegion>
.
</paragraph>
<paragraph id="8B8DE463FF9DDF7A3956F8D0FD5AFDE0" blockId="3.[824,1437,866,1975]" lastBlockId="4.[151,764,241,627]" lastPageId="4" lastPageNumber="143" pageId="3" pageNumber="142">
<emphasis id="B9463871FF9DDF7D3956F8D0FB91F8A4" bold="true" box="[847,1079,1852,1877]" pageId="3" pageNumber="142">Comparative notes</emphasis>
: Among the related species treated here, 
<taxonomicName id="4C329FE0FF9DDF7D39D0F8B2FB70F884" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[969,1238,1885,1909]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF9DDF7D39D0F8B2FB70F884" box="[969,1238,1885,1909]" italics="true" pageId="3" pageNumber="142">Scopaeus arfakmontium</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF9DDF7D3EFCF8B1FA80F887" box="[1253,1318,1885,1910]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="3" pageNumber="142">Fig. 2</figureCitation>
) is readily distinguishable from 
<taxonomicName id="4C329FE0FF9DDF7D3E07F893FB08F867" authorityName="Cameron" authorityYear="1938" box="[1054,1198,1918,1942]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="kokodanus">
<emphasis id="B9463871FF9DDF7D3E07F893FB08F867" box="[1054,1198,1918,1942]" italics="true" pageId="3" pageNumber="142">S. kokodanus</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF9DDF7D3EA3F892FB53F867" box="[1210,1269,1918,1942]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="3" pageNumber="142">Fig. 1</figureCitation>
) by the entirely black body without a light posterior margin of the elytra and the more elongate distal antennal segments. It differs from both 
<taxonomicName id="4C329FE0FF9ADF7A3B12FEFFFECCFEDB" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[267,362,274,298]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF9ADF7A3B12FEFFFECCFEDB" box="[267,362,274,298]" italics="true" pageId="4" pageNumber="143">S. balkei</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF9ADF7A3B60FEFEFE1FFEDB" box="[377,441,274,298]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="4" pageNumber="143">Fig. 3</figureCitation>
) and 
<taxonomicName id="4C329FE0FF9ADF7A3BEFFEFFFD25FEDB" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[502,643,274,298]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="potamides">
<emphasis id="B9463871FF9ADF7A3BEFFEFFFD25FEDB" box="[502,643,274,298]" italics="true" pageId="4" pageNumber="143">S. potamides</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF9ADF7A388BFEFEFD74FEDB" box="[658,722,274,298]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="4" pageNumber="143">Fig. 4</figureCitation>
) by larger body size, the pyriform head, the larger aedeagus (
<figureCitation id="1309F8E6FF9ADF7A3A86FEB8FF47FE9D" box="[159,225,340,364]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="4" pageNumber="143">Fig. 5</figureCitation>
) with elongate lateral lobes and the shape of the apical lobes, which are convexely widened laterally and parallel in lateral view (
<figureCitation id="1309F8E6FF9ADF7A3B87FE7AFDAAFE5F" box="[414,524,406,430]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="4" pageNumber="143">Figs 8–10</figureCitation>
), and by the posterior margin of abdominal sternite VIII with the median lobe longer than the lateroposterior angles (
<figureCitation id="1309F8E6FF9ADF7A3861FE34FD63FE01" box="[632,709,472,496]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="4" pageNumber="143">Fig. 18</figureCitation>
) and without four distinct, black macrosetae (see 
<figureCitation id="1309F8E6FF9ADF7A386EFE14FD56FDE0" box="[631,752,504,529]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="4" pageNumber="143">Figs 20, 22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9ADF7A3AB7FDF6FE1AFD82" pageId="4" pageNumber="143" type="etymology">
<paragraph id="8B8DE463FF9ADF7A3AB7FDF6FE1AFD82" blockId="4.[151,764,241,627]" pageId="4" pageNumber="143">
<emphasis id="B9463871FF9ADF7A3AB7FDF6FE8DFDC3" bold="true" box="[174,299,538,562]" pageId="4" pageNumber="143">Etymology</emphasis>
: The epithet 
<taxonomicName id="4C329FE0FF9ADF7A3BDBFDF6FDC7FDC0" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[450,609,537,562]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="arfakmontium">
‘ 
<emphasis id="B9463871FF9ADF7A3BD0FDF5FDC7FDC0" box="[457,609,537,561]" italics="true" pageId="4" pageNumber="143">arfakmontium</emphasis>
</taxonomicName>
’ (latin, noun, genitive: ‘of the Arfak Mountains’) refers to the 
<typeStatus id="54895AC1FF9ADF7A38D5FDD7FD5AFDA2" box="[716,764,571,595]" pageId="4" pageNumber="143">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>