<documentid="047F455496DE0DC1E3AB8988D6C5CEB7"ID-CLB-Dataset="304560"ID-DOI="10.11646/zootaxa.5519.2.3"ID-GBIF-Dataset="a3266865-054f-4cff-bc0c-faf89ec8074c"ID-ISSN="1175-5326"ID-Zenodo-Dep="13915771"ID-ZooBank="BE0CA3E4-3695-403A-AD98-3C279238518B"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="jonas"IM.materialsCitations_approvedBy="jonas"IM.metadata_approvedBy="jonas"IM.tables_approvedBy="jonas"IM.taxonomicNames_approvedBy="jonas"IM.treatments_approvedBy="jonas"checkinTime="1728585742752"checkinUser="plazi"docAuthor="Theischinger, Günther, Kalnins, Martins & Marinov, Milen"docDate="2024"docId="03C587C6CE6D9877FF24829B737EFA78"docLanguage="en"docName="zootaxa.5519.2.3.pdf"docOrigin="Zootaxa 5519 (2)"docSource="http://dx.doi.org/10.11646/zootaxa.5519.2.3"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Anax Leach 1815"docType="key"docVersion="3"lastPageNumber="241"masterDocId="FFFCFFBECE74986DFFB3832C7740FF90"masterDocTitle="Two new species of Anax Leach, 1815 from the Solomon Islands and Tonga (Odonata: Aeshnidae)"masterLastPageNumber="242"masterPageNumber="215"pageNumber="240"updateTime="1729087811061"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="5717C66597EA86F10A545EA9AE1C6FF4">Two new species of Anax Leach, 1815 from the Solomon Islands and Tonga (Odonata: Aeshnidae)</mods:title>
<mods:affiliationid="8EF51F9BA96F9671265C0AA064037EE7">Department of Biosystematics, Institute of Life Sciences and Technology, Daugavpils University, Parādes Street 1 A, Daugavpils, Latvia, LV- 5401</mods:affiliation>
<mods:affiliationid="D35A6285D514EAC283C75882611D2759">Biosecurity Surveillance & Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8544, New Zealand</mods:affiliation>
As characters such as e. g. ‘presence or absence and shape and definition of the postfrontal T-mark’, ‘colour of occipital triangle and vertex’ and ‘abdominal colour pattern’ have been found by other authors and ourselves as too variable to be used for specific distinction of
, the key is primarily based on the shape and proportions of the elements of the male terminalia (epiproct, cerci) that appear in a high degree consistent and thus to be most useful for the identification of taxa of the species group. An illustration of the abdominal patterns (
<figureCitationid="13572A55CE6D9874FABE819F7221FD5D"box="[1293,1377,690,717]"captionStart="FIGURE 53"captionStartId="21.[151,250,1856,1880]"captionTargetBox="[164,1436,181,1813]"captionTargetId="figure-17@21.[151,1436,181,1833]"captionTargetPageId="21"captionText="FIGURE 53. Anax spp., male terminalia: (a–e, h–p) dorsal; (f, g) base, lateral: (a, b, d) modified from Martin (1909), (c, e–j, l–m) modified from Lieftinck (1942); (k) modified from Kennedy (1934)."figureDoi="http://doi.org/10.5281/zenodo.13915810"httpUri="https://zenodo.org/record/13915810/files/figure.png"pageId="25"pageNumber="240">Fig. 53</figureCitation>
) that may also be useful is presented subsequent to the Key as a further aid to identification. However, it is important to point out inconsistencies and shortcomings in hitherto available keys or similar more or less comprehensive work. These are part of
<bibRefCitationid="EFFD4B21CE6D9874FED98033754EFCA9"author="Martin, R."box="[362,526,799,825]"pageId="25"pageNumber="240"pagination="157 - 223"refId="ref12817"refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223."type="journal article"year="1909">Martin (1909)</bibRefCitation>
,
<bibRefCitationid="EFFD4B21CE6D9874FDAE8033759AFCA9"author="Kennedy, C. H."box="[541,730,799,825]"pageId="25"pageNumber="240"pagination="346 - 356"refId="ref12221"refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346"type="journal article"year="1934">Kennedy (1934)</bibRefCitation>
,
<bibRefCitationid="EFFD4B21CE6D9874FD5A803374E8FCA9"author="Lieftinck, M. A."box="[745,936,799,825]"pageId="25"pageNumber="240"pagination="441 - 608"refId="ref12308"refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608."type="journal article"year="1942">Lieftinck (1942)</bibRefCitation>
,
<bibRefCitationid="EFFD4B21CE6D9874FC048033733FFCA9"author="Michalski, J."box="[951,1151,799,825]"pageId="25"pageNumber="240"refId="ref12846"refString="Michalski, J. (2012) A manual for the identification of the dragonflies and damselflies of New Guinea, Maluku, & the Solomon Islands. Kanduanum Books, Morristown, New Jersey, 561 pp."type="book"year="2012">Michalski (2012)</bibRefCitation>
,
<bibRefCitationid="EFFD4B21CE6D9874FB3D803372DCFCA9"author="Orr, A. G. & Kalkman, V. J."box="[1166,1436,798,825]"pageId="25"pageNumber="240"pagination="3 - 156"refId="ref12885"refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156."type="journal article"year="2015">Orr & Kalkman (2015)</bibRefCitation>
and
<bibRefCitationid="EFFD4B21CE6D9874FF7A806F76D7FCCD"author="Seehausen, M."box="[201,407,835,861]"pageId="25"pageNumber="240"pagination="1 - 34"refId="ref13074"refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34."type="journal article"year="2017">Seehausen (2017)</bibRefCitation>
.
<bibRefCitationid="EFFD4B21CE6D9874FE17806F7504FCCD"author="Martin, R."box="[420,580,835,861]"pageId="25"pageNumber="240"pagination="157 - 223"refId="ref12817"refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223."type="journal article"year="1909">Martin (1909)</bibRefCitation>
presents useful dorsal and lateral view illustrations of the male terminalia of
<bibRefCitationid="EFFD4B21CE6D9874FB8180A773AAFC35"author="Kennedy, C. H."box="[1074,1258,907,933]"pageId="25"pageNumber="240"pagination="346 - 356"refId="ref12221"refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346"type="journal article"year="1934">Kennedy (1934)</bibRefCitation>
includes dorsal and lateral illustrations of the terminalia of
<bibRefCitationid="EFFD4B21CE6D9874FB5180DB72DCFB81"author="Lieftinck, M. A."box="[1250,1436,1015,1041]"pageId="25"pageNumber="240"pagination="441 - 608"refId="ref12308"refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608."type="journal article"year="1942">Lieftinck (1942)</bibRefCitation>
presents excellent drawings of the available species including
<bibRefCitationid="EFFD4B21CE6D9874FB1187137225FBC9"author="Michalski, J."box="[1186,1381,1087,1113]"pageId="25"pageNumber="240"refId="ref12846"refString="Michalski, J. (2012) A manual for the identification of the dragonflies and damselflies of New Guinea, Maluku, & the Solomon Islands. Kanduanum Books, Morristown, New Jersey, 561 pp."type="book"year="2012">Michalski (2012)</bibRefCitation>
uses the illustrations of
<bibRefCitationid="EFFD4B21CE6D9874FEDF874F754BFBED"author="Martin, R."box="[364,523,1123,1149]"pageId="25"pageNumber="240"pagination="157 - 223"refId="ref12817"refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223."type="journal article"year="1909">Martin (1909)</bibRefCitation>
and
<bibRefCitationid="EFFD4B21CE6D9874FDF0874F75BCFBED"author="Lieftinck, M. A."box="[579,764,1123,1149]"pageId="25"pageNumber="240"pagination="441 - 608"refId="ref12308"refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608."type="journal article"year="1942">Lieftinck (1942)</bibRefCitation>
<bibRefCitationid="EFFD4B21CE6D9874FE0687AB75F5FB31"author="Orr, A. G. & Kalkman, V. J."box="[437,693,1158,1185]"pageId="25"pageNumber="240"pagination="3 - 156"refId="ref12885"refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156."type="journal article"year="2015">Orr & Kalkman (2015)</bibRefCitation>
give excellent colour illustrations including dorsal view terminalia illustrations of New
<bibRefCitationid="EFFD4B21CE6D9874FDF387E37448FB79"author="Seehausen, M."box="[576,776,1231,1257]"pageId="25"pageNumber="240"pagination="1 - 34"refId="ref13074"refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34."type="journal article"year="2017">Seehausen (2017)</bibRefCitation>
<bibRefCitationid="EFFD4B21CE6D9874FB5687DF72DCFA9D"author="Lieftinck, M. A."box="[1253,1436,1267,1293]"pageId="25"pageNumber="240"pagination="441 - 608"refId="ref12308"refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608."type="journal article"year="1942">Lieftinck (1942)</bibRefCitation>
<bibRefCitationid="EFFD4B21CE6D9874FD21863B74DDFAA1"author="Orr, A. G. & Kalkman, V. J."box="[658,925,1302,1329]"pageId="25"pageNumber="240"pagination="3 - 156"refId="ref12885"refString="Orr, A. G. & Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156."type="journal article"year="2015">Orr & Kalkman (2015)</bibRefCitation>
<bibRefCitationid="EFFD4B21CE6D9874FA84867377A4FA0D"author="Kennedy, C. H."pageId="25"pageNumber="240"pagination="346 - 356"refId="ref12221"refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346"type="journal article"year="1934">Kennedy (1934)</bibRefCitation>
, as well as a photo of this species by Yen Choong Chee and Hagen material of this species in Vienna. This raises doubt as to whether the abdomens figured by
<bibRefCitationid="EFFD4B21CE6D9874FD6A868B74E0FA51"author="Seehausen, M."box="[729,928,1447,1473]"pageId="25"pageNumber="240"pagination="1 - 34"refId="ref13074"refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34."type="journal article"year="2017">Seehausen (2017)</bibRefCitation>
<figureCitationid="13572A55CE6D9874FEB586C3761DF999"box="[262,349,1518,1545]"captionStart="FIGURE 52"captionStartId="19.[151,250,1810,1834]"captionTargetBox="[313,1275,633,1785]"captionTargetId="figure-268@19.[313,1275,633,1786]"captionTargetPageId="19"captionText="FIGURE 52a. Terms used in the descriptions and Key, presented on Anax gibbosulus, male terminalia, dorsal and lateral (modified from Lieftinck 1942): als = apico-lateral spike; bit = basal interior tooth; C = cercus; E = epiproct; S10 = abdominal segment 10; t = tubercles."figureDoi="http://doi.org/10.5281/zenodo.13915806"httpUri="https://zenodo.org/record/13915806/files/figure.png"pageId="25"pageNumber="240">Figs 52</figureCitation>
,
<figureCitationid="13572A55CE6D9874FEDA86C276C6F999"box="[361,390,1518,1545]"captionStart="FIGURE 53"captionStartId="21.[151,250,1856,1880]"captionTargetBox="[164,1436,181,1813]"captionTargetId="figure-17@21.[151,1436,181,1833]"captionTargetPageId="21"captionText="FIGURE 53. Anax spp., male terminalia: (a–e, h–p) dorsal; (f, g) base, lateral: (a, b, d) modified from Martin (1909), (c, e–j, l–m) modified from Lieftinck (1942); (k) modified from Kennedy (1934)."figureDoi="http://doi.org/10.5281/zenodo.13915810"httpUri="https://zenodo.org/record/13915810/files/figure.png"pageId="25"pageNumber="240">53</figureCitation>
<keyLeadid="309D95E5CE6D9874FF10854972D2F9ED"box="[163,1426,1637,1661]"pageId="25"pageNumber="240">- Epiproct not spiny and apically slightly to distinctly emarginate (b–p) 2</keyLead>
<keyLeadid="309D95E5CE6D9874FF1085FF7685F898"pageId="25"pageNumber="240">- Epiproct markedly shorter than one half of cerci, not much longer than wide, apically slightly 3 emarginate (c–p)</keyLead>
<keyLeadid="309D95E5CE6D9874FF1084727578F803"pageId="25"pageNumber="240">- Cerci distinctly increasing in width to at least 1/2 their length; with apico-lateral spike of 4 variable size and form (d–p)</keyLead>
<keyLeadid="309D95E5CE6E9877FF1083E7757EFE90"pageId="26"pageNumber="241">- Cerci widening from base to maximally 2/3 length, thence parallel sided or more or less 5 continuously narrowed (e–p)</keyLead>
<keyLeadid="309D95E5CE6E9877FF10827B7557FE1B"pageId="26"pageNumber="241">- Inner edge of cerci with some basal convexity or concavity; ventral edge of cerci with prominent 6 basal inferior tooth (g–p)</keyLead>
<keyLeadid="309D95E5CE6E9877FF1082E872D2FE4B"box="[163,1426,452,476]"pageId="26"pageNumber="241">- Inner edge of cerci undulate with basal concavity before larger convexity (i–p) 7</keyLead>
<keyLeadid="309D95E5CE6E9877FF1082C17605FDB1"pageId="26"pageNumber="241">7 (6) Cerci approximately three times as long as their greatest width, and apical 1/3 to 1/2 tapered 8 (i–j)</keyLead>
<keyLeadid="309D95E5CE6E9877FF10811E734BFDF7"pageId="26"pageNumber="241">- Cerci approximately three times as long as their greatest width and apical 1/4 parallel sided (m), 9 or cerci approximately four times as long as their greatest width (k–l, n–p)</keyLead>
<keyLeadid="309D95E5CE6E9877FF1081E475F9FD6D"pageId="26"pageNumber="241">9 (7) Cerci with apical 1/4 to 1/3 only slightly tapered and apically distinctly wider than at base, 10 giving them a truncate appearance (k–m)</keyLead>
<keyLeadid="309D95E5CE6E9877FF108022741AFCD2"pageId="26"pageNumber="241">- Cerci with apical 1/3 more distinctly tapered and apically barely wider than at base, giving them 12 a subtruncate to pointed (n) or rounded (o, p) appearance</keyLead>
<keyLeadid="309D95E5CE6E9877FF10873176B3FBC2"pageId="26"pageNumber="241">- Cerci with tiny apico-lateral spike pointing straight backward or slightly incurved, making them 13 appear rounded (o–p)</keyLead>
13 (12) Cerci inner margin with middle portion not distinctly bulging (o)
<taxonomicNameid="4C6C4D53CE6E9877FAA7874E72D1FBEA"authorityName="Theischinger & Kalnins & Marinov"authorityYear="2024"box="[1300,1425,1122,1146]"class="Insecta"family="Aeshnidae"genus="Anax"kingdom="Animalia"order="Odonata"pageId="26"pageNumber="241"phylum="Arthropoda"rank="species"species="insulanus">
- Cerci inner margin with middle portion distinctly bulging (p)
<taxonomicNameid="4C6C4D53CE6E9877FA8F87A672D2FB32"authorityName="Theischinger & Kalnins & Marinov"authorityYear="2024"box="[1340,1426,1162,1186]"class="Insecta"family="Aeshnidae"genus="Anax"kingdom="Animalia"order="Odonata"pageId="26"pageNumber="241"phylum="Arthropoda"rank="species"species="tonga">
<figureCitationid="13572A55CE6E9877FE2487D976B3FA9F"box="[407,499,1269,1295]"captionStart="FIGURE 54"captionStartId="22.[151,250,1874,1898]"captionTargetBox="[215,1368,193,1836]"captionTargetId="figure-22@22.[206,1381,181,1850]"captionTargetPageId="22"captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (m–o) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (b–g) modified from Orr & Kalkman (2015); (h–i) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, m–o) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened."figureDoi="http://doi.org/10.5281/zenodo.13915812"httpUri="https://zenodo.org/record/13915812/files/figure.png"pageId="26"pageNumber="241">Figs 54</figureCitation>
,
<figureCitationid="13572A55CE6E9877FDB387D9755FFA9F"box="[512,543,1269,1295]"captionStart="FIGURE 55"captionStartId="23.[151,250,1850,1874]"captionTargetBox="[161,1422,192,1813]"captionTargetId="figure-17@23.[151,1436,181,1826]"captionTargetPageId="23"captionText="FIGURE 55. Anax spp., males, dorsal, abdominal characters. (a–c) A. insulanus sp. nov.; (d–f) A. tonga sp. nov.: (a, d) abdominal pattern; (b, e) S2–3; (c, f) terminalia; (a, d) illustrated by Albert Orr."figureDoi="http://doi.org/10.5281/zenodo.13915814"httpUri="https://zenodo.org/record/13915814/files/figure.png"pageId="26"pageNumber="241">55</figureCitation>
Unfortunately it was not possible to present all species in colour, but
<figureCitationid="13572A55CE6E9877FC0086127361FAC8"box="[947,1057,1341,1368]"captionStart="FIGURE 54"captionStartId="22.[151,250,1874,1898]"captionTargetBox="[215,1368,193,1836]"captionTargetId="figure-22@22.[206,1381,181,1850]"captionTargetPageId="22"captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (m–o) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (b–g) modified from Orr & Kalkman (2015); (h–i) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, m–o) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened."figureDoi="http://doi.org/10.5281/zenodo.13915812"httpUri="https://zenodo.org/record/13915812/files/figure.png"pageId="26"pageNumber="241">Figure 54</figureCitation>
illustrated the dorsal view of the abdomen of all species, with colours where shown, as known or presumed to occur in life. Because of the small scale and the minute pale pattern elements (barely detectable in preserved specimens) in
<figureCitationid="13572A55CE6E9877FBD886AA73F9FA30"box="[1131,1209,1413,1440]"captionStart="FIGURE 54"captionStartId="22.[151,250,1874,1898]"captionTargetBox="[215,1368,193,1836]"captionTargetId="figure-22@22.[206,1381,181,1850]"captionTargetPageId="22"captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (m–o) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (b–g) modified from Orr & Kalkman (2015); (h–i) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, m–o) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened."figureDoi="http://doi.org/10.5281/zenodo.13915812"httpUri="https://zenodo.org/record/13915812/files/figure.png"pageId="26"pageNumber="241">Fig. 54</figureCitation>
the newly described species
<taxonomicNameid="4C6C4D53CE6E9877FF438686763BFA54"authorityName="Theischinger & Kalnins & Marinov"authorityYear="2024"box="[240,379,1450,1476]"class="Insecta"family="Aeshnidae"genus="Anax"kingdom="Animalia"order="Odonata"pageId="26"pageNumber="241"phylum="Arthropoda"rank="species"species="insulanus">
<taxonomicNameid="4C6C4D53CE6E9877FE0686867555FA53"authorityName="Theischinger & Kalnins & Marinov"authorityYear="2024"box="[437,533,1450,1476]"class="Insecta"family="Aeshnidae"genus="Anax"kingdom="Animalia"order="Odonata"pageId="26"pageNumber="241"phylum="Arthropoda"rank="species"species="tonga">
<taxonomicNameid="4C6C4D53CE6E9877FC6986867325FA54"authorityName="Theischinger & Kalnins & Marinov"authorityYear="2024"box="[986,1125,1450,1476]"class="Insecta"family="Aeshnidae"genus="Anax"kingdom="Animalia"order="Odonata"pageId="26"pageNumber="241"phylum="Arthropoda"rank="species"species="insulanus">
<taxonomicNameid="4C6C4D53CE6E9877FAA686867235FA53"authorityName="Theischinger & Kalnins & Marinov"authorityYear="2024"box="[1301,1397,1450,1476]"class="Insecta"family="Aeshnidae"genus="Anax"kingdom="Animalia"order="Odonata"pageId="26"pageNumber="241"phylum="Arthropoda"rank="species"species="tonga">
