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New taxonomic status for
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is here elevated to species rank,
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<bibRefCitation id="EFFD4B21CE66987FFBD286367251FAA4" author="Lieftinck, M. A." box="[1121,1297,1306,1332]" pageId="18" pageNumber="233" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck, 1942</bibRefCitation>
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,
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<taxonomicNameLabel id="A22B57B9CE66987FFAAF8637723AFAA4" box="[1308,1402,1307,1332]" pageId="18" pageNumber="233">stat. rev.</taxonomicNameLabel>
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on the basis of its high S3 Index vs the low S3 Index of
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<taxonomicName id="4C6C4D53CE66987FFD5B8612748AFAC8" authorityName="Hagen" authorityYear="1867" box="[744,970,1342,1368]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="233" phylum="Arthropoda" rank="subSpecies" species="fumosus" subSpecies="fumosus">A. fumosus fumosus</taxonomicName>
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Calculated on the basis of measurements given by
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the S3 Index of
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is 5.00 vs
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Key for species identification (males only) of
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As characters such as e. g. presence or absence and shape and definition of the postfrontal T-mark, colour of occipital triangle and vertex and abdominal colour pattern have been found by other authors and ourselves as too variable to be used for specific distinction of
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</taxonomicName>
, the key is primarily based on the shape and proportions of the elements of the male terminalia (epiproct, cerci) that appear in a high degree consistent and thus to be most useful for the identification of taxa of the species group. An illustration of the abdominal patterns (
<figureCitation id="13572A55CE6D9874FABE819F7221FD5D" box="[1293,1377,690,717]" captionStart="FIGURE 53" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[164,1436,181,1813]" captionTargetId="figure-17@21.[151,1436,181,1833]" captionTargetPageId="21" captionText="FIGURE 53. Anax spp., male terminalia: (ae, hp) dorsal; (f, g) base, lateral: (a, b, d) modified from Martin (1909), (c, ej, lm) modified from Lieftinck (1942); (k) modified from Kennedy (1934)." figureDoi="http://doi.org/10.5281/zenodo.13915810" httpUri="https://zenodo.org/record/13915810/files/figure.png" pageId="25" pageNumber="240">Fig. 53</figureCitation>
) that may also be useful is presented subsequent to the Key as a further aid to identification. However, it is important to point out inconsistencies and shortcomings in hitherto available keys or similar more or less comprehensive work. These are part of
<bibRefCitation id="EFFD4B21CE6D9874FED98033754EFCA9" author="Martin, R." box="[362,526,799,825]" pageId="25" pageNumber="240" pagination="157 - 223" refId="ref12817" refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223." type="journal article" year="1909">Martin (1909)</bibRefCitation>
,
<bibRefCitation id="EFFD4B21CE6D9874FDAE8033759AFCA9" author="Kennedy, C. H." box="[541,730,799,825]" pageId="25" pageNumber="240" pagination="346 - 356" refId="ref12221" refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346" type="journal article" year="1934">Kennedy (1934)</bibRefCitation>
,
<bibRefCitation id="EFFD4B21CE6D9874FD5A803374E8FCA9" author="Lieftinck, M. A." box="[745,936,799,825]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
,
<bibRefCitation id="EFFD4B21CE6D9874FC048033733FFCA9" author="Michalski, J." box="[951,1151,799,825]" pageId="25" pageNumber="240" refId="ref12846" refString="Michalski, J. (2012) A manual for the identification of the dragonflies and damselflies of New Guinea, Maluku, &amp; the Solomon Islands. Kanduanum Books, Morristown, New Jersey, 561 pp." type="book" year="2012">Michalski (2012)</bibRefCitation>
,
<bibRefCitation id="EFFD4B21CE6D9874FB3D803372DCFCA9" author="Orr, A. G. &amp; Kalkman, V. J." box="[1166,1436,798,825]" pageId="25" pageNumber="240" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. &amp; Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr &amp; Kalkman (2015)</bibRefCitation>
and
<bibRefCitation id="EFFD4B21CE6D9874FF7A806F76D7FCCD" author="Seehausen, M." box="[201,407,835,861]" pageId="25" pageNumber="240" pagination="1 - 34" refId="ref13074" refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34." type="journal article" year="2017">Seehausen (2017)</bibRefCitation>
.
<bibRefCitation id="EFFD4B21CE6D9874FE17806F7504FCCD" author="Martin, R." box="[420,580,835,861]" pageId="25" pageNumber="240" pagination="157 - 223" refId="ref12817" refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223." type="journal article" year="1909">Martin (1909)</bibRefCitation>
presents useful dorsal and lateral view illustrations of the male terminalia of
<taxonomicName id="4C6C4D53CE6D9874FF0B804B7677FC10" authorityName="Hagen" authorityYear="1867" box="[184,311,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="strenuus">
<emphasis id="B918EAC2CE6D9874FF0B804B7677FC10" box="[184,311,871,897]" italics="true" pageId="25" pageNumber="240">A. strenuus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FEFB804B768AFC10" authorityName="Selys" authorityYear="1872" box="[328,458,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
<emphasis id="B918EAC2CE6D9874FEFB804B768AFC10" box="[328,458,871,897]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FE68804B7516FC10" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[475,598,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FE68804B7516FC10" box="[475,598,871,897]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FDD4804B7441FC11" authorityName="Rambur" authorityYear="1842" box="[615,769,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
<emphasis id="B918EAC2CE6D9874FDD4804B7441FC11" box="[615,769,871,897]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FCA1804B74FEFC11" authorityName="Forster" authorityYear="1898" box="[786,958,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="maclachlani">
<emphasis id="B918EAC2CE6D9874FCA1804B74FEFC11" box="[786,958,871,897]" italics="true" pageId="25" pageNumber="240">A. maclachlani</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FC7C804B736EFC11" authorityName="Forster" authorityYear="1900" box="[975,1070,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="selysi">
<emphasis id="B918EAC2CE6D9874FC7C804B736EFC11" box="[975,1070,871,897]" italics="true" pageId="25" pageNumber="240">A. selysi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE6D9874FBD9804B73BBFC10" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1130,1275,871,897]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B918EAC2CE6D9874FBD9804B73BBFC10" box="[1130,1275,871,897]" italics="true" pageId="25" pageNumber="240">A. papuensis</emphasis>
</taxonomicName>
. He mentions
<taxonomicName id="4C6C4D53CE6D9874FF2480A77662FC35" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[151,290,907,933]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FF2480A77662FC35" box="[151,290,907,933]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
only as a race of
<taxonomicName id="4C6C4D53CE6D9874FE5C80A77529FC34" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[495,617,907,933]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FE5C80A77529FC34" box="[495,617,907,933]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
and does not refer to
<taxonomicName id="4C6C4D53CE6D9874FCD680A774A2FC35" authorityName="Hagen" authorityYear="1867" box="[869,994,907,933]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">
<emphasis id="B918EAC2CE6D9874FCD680A774A2FC35" box="[869,994,907,933]" italics="true" pageId="25" pageNumber="240">A. fumosus</emphasis>
</taxonomicName>
at all.
<bibRefCitation id="EFFD4B21CE6D9874FB8180A773AAFC35" author="Kennedy, C. H." box="[1074,1258,907,933]" pageId="25" pageNumber="240" pagination="346 - 356" refId="ref12221" refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346" type="journal article" year="1934">Kennedy (1934)</bibRefCitation>
includes dorsal and lateral illustrations of the terminalia of
<taxonomicName id="4C6C4D53CE6D9874FD3E80837452FC58" box="[653,786,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="piraticus">
<emphasis id="B918EAC2CE6D9874FD3E80837452FC58" box="[653,786,943,969]" italics="true" pageId="25" pageNumber="240">A. piraticus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FC91808374EEFC59" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[802,942,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FC91808374EEFC59" box="[802,942,943,969]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FC0D80837318FC59" authorityName="Rambur" authorityYear="1842" box="[958,1112,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
<emphasis id="B918EAC2CE6D9874FC0D80837318FC59" box="[958,1112,943,969]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FBDA808373A4FC58" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1129,1252,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FBDA808373A4FC58" box="[1129,1252,943,969]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FB4680837234FC58" authorityName="Hagen" authorityYear="1867" box="[1269,1396,943,969]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="strenuus">
<emphasis id="B918EAC2CE6D9874FB4680837234FC58" box="[1269,1396,943,969]" italics="true" pageId="25" pageNumber="240">A. strenuus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FA3780837795FC7D" authorityName="Forster" authorityYear="1900" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="selysi">
<emphasis id="B918EAC2CE6D9874FA3780837795FC7D" italics="true" pageId="25" pageNumber="240">A. selysi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE6D9874FEBC80FF76F9FC7D" authorityName="Forster" authorityYear="1898" box="[271,441,979,1005]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="maclachlani">
<emphasis id="B918EAC2CE6D9874FEBC80FF76F9FC7D" box="[271,441,979,1005]" italics="true" pageId="25" pageNumber="240">A. maclachlani</emphasis>
</taxonomicName>
. However, what he presents as
<taxonomicName id="4C6C4D53CE6D9874FC9080FF74FDFC7D" authorityName="Rambur" authorityYear="1842" box="[803,957,979,1005]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
<emphasis id="B918EAC2CE6D9874FC9080FF74FDFC7D" box="[803,957,979,1005]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
</taxonomicName>
is misleading, because it is probably only a different version of what he presents correctly as
<emphasis id="B918EAC2CE6D9874FD5780DB7426FB80" box="[740,870,1015,1041]" italics="true" pageId="25" pageNumber="240">
<taxonomicName id="4C6C4D53CE6D9874FD5780DB7422FB80" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[740,866,1015,1041]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">A. guttatus</taxonomicName>
,
</emphasis>
and certainly not
<taxonomicName id="4C6C4D53CE6D9874FB8B80DB7392FB81" authorityName="Rambur" authorityYear="1842" box="[1080,1234,1015,1041]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
<emphasis id="B918EAC2CE6D9874FB8B80DB7392FB81" box="[1080,1234,1015,1041]" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFD4B21CE6D9874FB5180DB72DCFB81" author="Lieftinck, M. A." box="[1250,1436,1015,1041]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
presents excellent drawings of the available species including
<emphasis id="B918EAC2CE6D9874FCFD87377490FBA5" box="[846,976,1051,1077]" italics="true" pageId="25" pageNumber="240">
<taxonomicName id="4C6C4D53CE6D9874FCFD8737748CFBA5" authorityName="Hagen" authorityYear="1867" box="[846,972,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">A. fumosus</taxonomicName>
,
</emphasis>
but except
<taxonomicName id="4C6C4D53CE6D9874FBE287377390FBA4" authorityName="Selys" authorityYear="1872" box="[1105,1232,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
<emphasis id="B918EAC2CE6D9874FBE287377390FBA4" box="[1105,1232,1051,1077]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
</taxonomicName>
and
<emphasis id="B918EAC2CE6D9874FAB5873772DDFBA4" box="[1286,1437,1051,1077]" italics="true" pageId="25" pageNumber="240">
<taxonomicName id="4C6C4D53CE6D9874FAB5873772D9FBA4" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1286,1433,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">A. papuensis</taxonomicName>
.
</emphasis>
He adds two taxa of the species group he describes as
<taxonomicName id="4C6C4D53CE6D9874FD488713742DFBC8" authorityName="Lieftinck" authorityYear="1942" box="[763,877,1087,1113]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B918EAC2CE6D9874FD488713742DFBC8" box="[763,877,1087,1113]" italics="true" pageId="25" pageNumber="240">A. pugnax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE6D9874FC17871373D4FBC9" authorityName="Lieftinck" authorityYear="1942" box="[932,1172,1087,1113]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="subSpecies" species="fumosus" subSpecies="celebense">
<emphasis id="B918EAC2CE6D9874FC17871373D4FBC9" box="[932,1172,1087,1113]" italics="true" pageId="25" pageNumber="240">A. fumosus celebense</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFFD4B21CE6D9874FB1187137225FBC9" author="Michalski, J." box="[1186,1381,1087,1113]" pageId="25" pageNumber="240" refId="ref12846" refString="Michalski, J. (2012) A manual for the identification of the dragonflies and damselflies of New Guinea, Maluku, &amp; the Solomon Islands. Kanduanum Books, Morristown, New Jersey, 561 pp." type="book" year="2012">Michalski (2012)</bibRefCitation>
uses the illustrations of
<bibRefCitation id="EFFD4B21CE6D9874FEDF874F754BFBED" author="Martin, R." box="[364,523,1123,1149]" pageId="25" pageNumber="240" pagination="157 - 223" refId="ref12817" refString="Martin, R. (1909) Aeshnines. Collections zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et descriptif, 20, 157 - 223." type="journal article" year="1909">Martin (1909)</bibRefCitation>
and
<bibRefCitation id="EFFD4B21CE6D9874FDF0874F75BCFBED" author="Lieftinck, M. A." box="[579,764,1123,1149]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
for the presentation of the species of New
<collectingCountry id="F37B7640CE6D9874FB56874F7279FBED" box="[1253,1337,1123,1149]" name="Guinea-Bissau" pageId="25" pageNumber="240">Guinea</collectingCountry>
, Maluku and the
<collectingCountry id="F37B7640CE6D9874FF5D87AB76ECFB31" box="[238,428,1159,1185]" name="Solomon Islands" pageId="25" pageNumber="240">Solomon Islands</collectingCountry>
.
<bibRefCitation id="EFFD4B21CE6D9874FE0687AB75F5FB31" author="Orr, A. G. &amp; Kalkman, V. J." box="[437,693,1158,1185]" pageId="25" pageNumber="240" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. &amp; Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr &amp; Kalkman (2015)</bibRefCitation>
give excellent colour illustrations including dorsal view terminalia illustrations of New
<collectingCountry id="F37B7640CE6D9874FE3187877693FB55" box="[386,467,1195,1221]" name="Guinea-Bissau" pageId="25" pageNumber="240">Guinea</collectingCountry>
species, (but in the detailed line drawings of appendages
<taxonomicName id="4C6C4D53CE6D9874FBD8878773A7FB55" authorityName="Hagen" authorityYear="1867" box="[1131,1255,1195,1221]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">
<emphasis id="B918EAC2CE6D9874FBD8878773A7FB55" box="[1131,1255,1195,1221]" italics="true" pageId="25" pageNumber="240">A. fumosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE6D9874FA92878772DCFB54" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1313,1436,1195,1221]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FA92878772DCFB54" box="[1313,1436,1195,1221]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
are mistakenly interchanged). Finally
<bibRefCitation id="EFFD4B21CE6D9874FDF387E37448FB79" author="Seehausen, M." box="[576,776,1231,1257]" pageId="25" pageNumber="240" pagination="1 - 34" refId="ref13074" refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34." type="journal article" year="2017">Seehausen (2017)</bibRefCitation>
when illustrating
<taxonomicName id="4C6C4D53CE6D9874FC6187E37321FB78" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[978,1121,1231,1257]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B918EAC2CE6D9874FC6187E37321FB78" box="[978,1121,1231,1257]" italics="true" pageId="25" pageNumber="240">A. papuensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FBDC87E373A8FB78" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1135,1256,1231,1257]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FBDC87E373A8FB78" box="[1135,1256,1231,1257]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FB4587E37236FB78" authorityName="Selys" authorityYear="1872" box="[1270,1398,1231,1257]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
<emphasis id="B918EAC2CE6D9874FB4587E37236FB78" box="[1270,1398,1231,1257]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D53CE6D9874FA3787E37650FA9D" authorityName="Rambur" authorityYear="1842" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="gibbosulus">
<emphasis id="B918EAC2CE6D9874FA3787E37650FA9D" italics="true" pageId="25" pageNumber="240">A. gibbosulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE6D9874FEF587DF768EFA9D" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[326,462,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FEF587DF768EFA9D" box="[326,462,1267,1293]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
depicts
<taxonomicName id="4C6C4D53CE6D9874FD9987DF75E3FA9C" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[554,675,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FD9987DF75E3FA9C" box="[554,675,1267,1293]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
matching very closely the drawing
<emphasis id="B918EAC2CE6D9874FB8287DF7381FA9D" box="[1073,1217,1267,1293]" italics="true" pageId="25" pageNumber="240">
A. f.
<taxonomicName id="4C6C4D53CE6D9874FBD687DF7381FA9D" authorityName="Hagen" authorityYear="1867" box="[1125,1217,1267,1293]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">fumosus</taxonomicName>
</emphasis>
of
<bibRefCitation id="EFFD4B21CE6D9874FB5687DF72DCFA9D" author="Lieftinck, M. A." box="[1253,1436,1267,1293]" pageId="25" pageNumber="240" pagination="441 - 608" refId="ref12308" refString="Lieftinck, M. A. (1942) The dragonflies (Odonata) of New Guinea and neighbouring islands. Part VI, Results of the Third Archbold Expedition 1938 - 39 and of the Le Roux Expedition 1939 to Netherlands New Guinea (I. Anisoptera). Treubia, 18 (3), 441 - 608." type="journal article" year="1942">Lieftinck (1942)</bibRefCitation>
and
<taxonomicName id="4C6C4D53CE6D9874FF7B863B7603FAA0" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[200,323,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6D9874FF7B863B7603FAA0" box="[200,323,1303,1329]" italics="true" pageId="25" pageNumber="240">A. guttatus</emphasis>
</taxonomicName>
(but actually
<taxonomicName id="4C6C4D53CE6D9874FE50863B7520FAA1" authorityName="Hagen" authorityYear="1867" box="[483,608,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="fumosus">
<emphasis id="B918EAC2CE6D9874FE50863B7520FAA1" box="[483,608,1303,1329]" italics="true" pageId="25" pageNumber="240">A. fumosus</emphasis>
</taxonomicName>
) of
<bibRefCitation id="EFFD4B21CE6D9874FD21863B74DDFAA1" author="Orr, A. G. &amp; Kalkman, V. J." box="[658,925,1302,1329]" pageId="25" pageNumber="240" pagination="3 - 156" refId="ref12885" refString="Orr, A. G. &amp; Kalkman, V. J. (2015) Field guide to the dragonflies of New Guinea. Brachytron, 17 (Supplement), 3 - 156." type="journal article" year="2015">Orr &amp; Kalkman (2015)</bibRefCitation>
. Seehausens drawing of the terminalia of
<taxonomicName id="4C6C4D53CE6D9874FA37863B7641FAC5" baseAuthorityName="Seehausen" baseAuthorityYear="2017" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FA37863B7641FAC5" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
looks almost like a copy of Lieftincks illustration of
<emphasis id="B918EAC2CE6D9874FCD48617734BFAC5" box="[871,1035,1339,1365]" italics="true" pageId="25" pageNumber="240">
A. f.
<taxonomicName id="4C6C4D53CE6D9874FC2D8617734BFAC5" box="[926,1035,1339,1365]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="celebense">celebense</taxonomicName>
</emphasis>
and not at all similar to Lieftincks illustration of
<taxonomicName id="4C6C4D53CE6D9874FE8A86737684FAE9" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[313,452,1375,1401]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FE8A86737684FAE9" box="[313,452,1375,1401]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
that matches the illustration of the
<typeStatus id="54D78872CE6D9874FCEF867374CCFAE9" box="[860,908,1375,1401]" pageId="25" pageNumber="240">type</typeStatus>
of this species by F.M. Carpenter in
<bibRefCitation id="EFFD4B21CE6D9874FA84867377A4FA0D" author="Kennedy, C. H." pageId="25" pageNumber="240" pagination="346 - 356" refId="ref12221" refString="Kennedy, C. H. (1934) Anax piraticus a new species of dragonfly (Odonata) from Guam. Annals of the Entomological Society of America, 27, 346 - 356. https: // doi. org / 10.1093 / aesa / 27.2.346" type="journal article" year="1934">Kennedy (1934)</bibRefCitation>
, as well as a photo of this species by Yen Choong Chee and Hagen material of this species in Vienna. This raises doubt as to whether the abdomens figured by
<bibRefCitation id="EFFD4B21CE6D9874FD6A868B74E0FA51" author="Seehausen, M." box="[729,928,1447,1473]" pageId="25" pageNumber="240" pagination="1 - 34" refId="ref13074" refString="Seehausen, M. (2017) Survey of Odonata from Timor Island, with description of the female of Anax georgius (Odonata: Aeshnidae). Faunistic Studies in Southeast Asian and Pacific Island Odonata, 20, 1 - 34." type="journal article" year="2017">Seehausen (2017)</bibRefCitation>
as
<taxonomicName id="4C6C4D53CE6D9874FC77868B730DFA51" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[964,1101,1447,1473]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FC77868B730DFA51" box="[964,1101,1447,1473]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
(
<figureCitation id="13572A55CE6D9874FBE8868B7393FA51" box="[1115,1235,1447,1473]" captionStart="FIGURES 1825" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[151,1422,561,1845]" captionTargetId="figure-54@6.[151,1435,561,1845]" captionTargetPageId="6" captionText="FIGURES 1825. Anax spp., male, dorsal: (1821) S23; (2225) terminalia: (18, 22) A. tonga; (19, 23) A. celebensis; (20, 24) A. insulanus; (21, 25) A. piraticus." pageId="25" pageNumber="240">Fig. 21 jl</figureCitation>
), and in particular if the thick-waisted females (
<figureCitation id="13572A55CE6D9874FE6E86E7751CFA75" box="[477,604,1483,1509]" captionStart="FIGURES 1825" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[151,1422,561,1845]" captionTargetId="figure-54@6.[151,1435,561,1845]" captionTargetPageId="6" captionText="FIGURES 1825. Anax spp., male, dorsal: (1821) S23; (2225) terminalia: (18, 22) A. tonga; (19, 23) A. celebensis; (20, 24) A. insulanus; (21, 25) A. piraticus." pageId="25" pageNumber="240">Fig. 22 jk</figureCitation>
), really are
<taxonomicName id="4C6C4D53CE6D9874FD6C86E77428FA75" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[735,872,1483,1509]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6D9874FD6C86E77428FA75" box="[735,872,1483,1509]" italics="true" pageId="25" pageNumber="240">A. panybeus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C376655BCE6D9874FF7486C376CEF999" box="[199,398,1518,1545]" pageId="25" pageNumber="240" type="multiple">
<paragraph id="8BD336D0CE6D9874FF7486C376CEF999" blockId="25.[151,1437,547,1545]" box="[199,398,1518,1545]" pageId="25" pageNumber="240">
<emphasis id="B918EAC2CE6D9874FF7486C377B7F999" bold="true" box="[199,247,1519,1545]" pageId="25" pageNumber="240">Key</emphasis>
(
<figureCitation id="13572A55CE6D9874FEB586C3761DF999" box="[262,349,1518,1545]" captionStart="FIGURE 52" captionStartId="19.[151,250,1810,1834]" captionTargetBox="[313,1275,633,1785]" captionTargetId="figure-268@19.[313,1275,633,1786]" captionTargetPageId="19" captionText="FIGURE 52a. Terms used in the descriptions and Key, presented on Anax gibbosulus, male terminalia, dorsal and lateral (modified from Lieftinck 1942): als = apico-lateral spike; bit = basal interior tooth; C = cercus; E = epiproct; S10 = abdominal segment 10; t = tubercles." figureDoi="http://doi.org/10.5281/zenodo.13915806" httpUri="https://zenodo.org/record/13915806/files/figure.png" pageId="25" pageNumber="240">Figs 52</figureCitation>
,
<figureCitation id="13572A55CE6D9874FEDA86C276C6F999" box="[361,390,1518,1545]" captionStart="FIGURE 53" captionStartId="21.[151,250,1856,1880]" captionTargetBox="[164,1436,181,1813]" captionTargetId="figure-17@21.[151,1436,181,1833]" captionTargetPageId="21" captionText="FIGURE 53. Anax spp., male terminalia: (ae, hp) dorsal; (f, g) base, lateral: (a, b, d) modified from Martin (1909), (c, ej, lm) modified from Lieftinck (1942); (k) modified from Kennedy (1934)." figureDoi="http://doi.org/10.5281/zenodo.13915810" httpUri="https://zenodo.org/record/13915810/files/figure.png" pageId="25" pageNumber="240">53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C376655BCE6D9877FF10851172D2FB32" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240" type="key">
<key id="FFFD61E1CE6D9877FF10851172D2FB32" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240">
<keyStep id="30982E75CE6D9874FF10851172D2F9ED" pageId="25" pageNumber="240">
<paragraph id="8BD336D0CE6D9874FF10851172D1F9C4" blockId="25.[163,1426,1597,1980]" box="[163,1425,1597,1621]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF10851172D1F9C4" box="[163,1425,1597,1621]" pageId="25" pageNumber="240">
1 Epiproct dorsally spiny and apically not emarginate (a)
<taxonomicName id="4C6C4D53CE6D9874FAA3851172D1F9C4" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1296,1425,1597,1620]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B918EAC2CE6D9874FAA3851172D1F9C4" box="[1296,1425,1597,1620]" italics="true" pageId="25" pageNumber="240">A. papuensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6D9874FF10854972D2F9ED" blockId="25.[163,1426,1597,1980]" box="[163,1426,1637,1661]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF10854972D2F9ED" box="[163,1426,1637,1661]" pageId="25" pageNumber="240">- Epiproct not spiny and apically slightly to distinctly emarginate (bp) 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6D9874FF1085A17685F898" pageId="25" pageNumber="240">
<paragraph id="8BD336D0CE6D9874FF1085A176EEF952" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF1085A176EEF952" pageId="25" pageNumber="240">
2 (1) Epiproct much longer than one half of cerci, more than twice as long as wide, apically distinctly
<taxonomicName id="4C6C4D53CE6D9874FAAD85A272D2F935" authorityName="Selys" authorityYear="1872" box="[1310,1426,1678,1701]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="georgius">
<emphasis id="B918EAC2CE6D9874FAAD85A272D2F935" box="[1310,1426,1678,1701]" italics="true" pageId="25" pageNumber="240">A. georgius</emphasis>
</taxonomicName>
emarginate (b)
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6D9874FF1085FF7685F898" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF1085FF7685F898" pageId="25" pageNumber="240">- Epiproct markedly shorter than one half of cerci, not much longer than wide, apically slightly 3 emarginate (cp)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6D9874FF1084347578F803" pageId="25" pageNumber="240">
<paragraph id="8BD336D0CE6D9874FF10843472D1F8DE" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF10843472D1F8DE" pageId="25" pageNumber="240">
3 (2) Cerci distinctly increasing in width to markedly less than 1/2 their length; apically simply
<emphasis id="B918EAC2CE6D9874FACF843572D2F8A0" box="[1404,1426,1817,1840]" italics="true" pageId="25" pageNumber="240">A.</emphasis>
rounded without apico-lateral spike (c)
<taxonomicName id="4C6C4D53CE6D9874FAA7841A72D1F8DE" authorityName="Forster" authorityYear="1898" box="[1300,1425,1846,1870]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="maclachlani">
<emphasis id="B918EAC2CE6D9874FAA7841A72D1F8DE" box="[1300,1425,1846,1870]" italics="true" pageId="25" pageNumber="240">maclachlani</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6D9874FF1084727578F803" blockId="25.[163,1426,1597,1980]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF1084727578F803" pageId="25" pageNumber="240">- Cerci distinctly increasing in width to at least 1/2 their length; with apico-lateral spike of 4 variable size and form (dp)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6D9877FF108488757EFE90" lastPageId="26" lastPageNumber="241" pageId="25" pageNumber="240">
<paragraph id="8BD336D0CE6D9874FF10848872D2F82B" blockId="25.[163,1426,1597,1980]" box="[163,1426,1955,1980]" pageId="25" pageNumber="240">
<keyLead id="309D95E5CE6D9874FF10848872D2F82B" box="[163,1426,1955,1980]" pageId="25" pageNumber="240">
4 (3) Cerci widening from base to more than 3/4 length, thence abruptly narrowed (d)
<taxonomicName id="4C6C4D53CE6D9874FA93848872D2F82B" authorityName="Hagen" authorityYear="1867" box="[1312,1426,1956,1979]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="240" phylum="Arthropoda" rank="species" species="strenuus">
<emphasis id="B918EAC2CE6D9874FA93848872D2F82B" box="[1312,1426,1956,1979]" italics="true" pageId="25" pageNumber="240">A. strenuus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF1083E7757EFE90" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1083E7757EFE90" pageId="26" pageNumber="241">- Cerci widening from base to maximally 2/3 length, thence parallel sided or more or less 5 continuously narrowed (ep)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF10823C7557FE1B" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF10823C7697FED6" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10823C7697FED6" pageId="26" pageNumber="241">
5 (4) Inner edge of cerci straight beyond midlength; ventral edge of cerci without prominent basal
<taxonomicName id="4C6C4D53CE6E9877FA98823D72D2FEB8" authorityName="Lieftinck" authorityYear="1942" box="[1323,1426,273,296]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="pugnax">
<emphasis id="B918EAC2CE6E9877FA98823D72D2FEB8" box="[1323,1426,273,296]" italics="true" pageId="26" pageNumber="241">A. pugnax</emphasis>
</taxonomicName>
inferior tooth (ef)
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF10827B7557FE1B" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10827B7557FE1B" pageId="26" pageNumber="241">- Inner edge of cerci with some basal convexity or concavity; ventral edge of cerci with prominent 6 basal inferior tooth (gp)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF1082B772D2FE4B" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF1082B772D2FE24" blockId="26.[163,1426,203,1186]" box="[163,1426,411,436]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1082B772D2FE24" box="[163,1426,411,436]" pageId="26" pageNumber="241">
6 (5) Inner edge of cerci widely convex without basal concavity (h)
<taxonomicName id="4C6C4D53CE6E9877FABB82B072D2FE24" authorityName="Rambur" authorityYear="1842" box="[1288,1426,412,436]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="gibbosulus">
<emphasis id="B918EAC2CE6E9877FABB82B072D2FE24" box="[1288,1426,412,436]" italics="true" pageId="26" pageNumber="241">A. gibbosulus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF1082E872D2FE4B" blockId="26.[163,1426,203,1186]" box="[163,1426,452,476]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1082E872D2FE4B" box="[163,1426,452,476]" pageId="26" pageNumber="241">- Inner edge of cerci undulate with basal concavity before larger convexity (ip) 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF1082C1734BFDF7" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF1082C17605FDB1" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1082C17605FDB1" pageId="26" pageNumber="241">7 (6) Cerci approximately three times as long as their greatest width, and apical 1/3 to 1/2 tapered 8 (ij)</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF10811E734BFDF7" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10811E734BFDF7" pageId="26" pageNumber="241">- Cerci approximately three times as long as their greatest width and apical 1/4 parallel sided (m), 9 or cerci approximately four times as long as their greatest width (kl, np)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF10815B72D1FD27" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF10815B72D1FD1F" blockId="26.[163,1426,203,1186]" box="[163,1425,631,656]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10815B72D1FD1F" box="[163,1425,631,656]" pageId="26" pageNumber="241">
8 (7) Cerci greatest width at about 2/3 length and outer margin straight (i)
<taxonomicName id="4C6C4D53CE6E9877FAA8815472D1FD1F" box="[1307,1425,632,655]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="piraticus">
<emphasis id="B918EAC2CE6E9877FAA8815472D1FD1F" box="[1307,1425,632,655]" italics="true" pageId="26" pageNumber="241">A. piraticus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF10818C72D1FD27" blockId="26.[163,1426,203,1186]" box="[163,1425,671,696]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10818C72D1FD27" box="[163,1425,671,696]" pageId="26" pageNumber="241">
- Cerci greatest width at about 1/2 length and outer margin convex (j)
<taxonomicName id="4C6C4D53CE6E9877FA8E818C72D1FD27" authorityName="Forster" authorityYear="1900" box="[1341,1425,671,695]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="selysi">
<emphasis id="B918EAC2CE6E9877FA8E818C72D1FD27" box="[1341,1425,671,695]" italics="true" pageId="26" pageNumber="241">A. selysi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF1081E4741AFCD2" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF1081E475F9FD6D" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1081E475F9FD6D" pageId="26" pageNumber="241">9 (7) Cerci with apical 1/4 to 1/3 only slightly tapered and apically distinctly wider than at base, 10 giving them a truncate appearance (km)</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF108022741AFCD2" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF108022741AFCD2" pageId="26" pageNumber="241">- Cerci with apical 1/3 more distinctly tapered and apically barely wider than at base, giving them 12 a subtruncate to pointed (n) or rounded (o, p) appearance</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF10807F72D1FC03" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF10807F72D2FCFB" blockId="26.[163,1426,203,1186]" box="[163,1426,851,875]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10807F72D2FCFB" box="[163,1426,851,875]" pageId="26" pageNumber="241">
10 (9) Cerci inner margin with middle portion not distinctly bulging (k)
<taxonomicName id="4C6C4D53CE6E9877FAA5807F72D2FCFB" baseAuthorityName="Seehausen" baseAuthorityYear="2017" box="[1302,1426,851,875]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="panybeus">
<emphasis id="B918EAC2CE6E9877FAA5807F72D2FCFB" box="[1302,1426,851,875]" italics="true" pageId="26" pageNumber="241">A. panybeus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF10805072D1FC03" blockId="26.[163,1426,203,1186]" box="[163,1425,891,915]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10805072D1FC03" box="[163,1425,891,915]" pageId="26" pageNumber="241">- Cerci inner margin with middle portion distinctly bulging (lm) 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF10808872D1FC74" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF10808872D1FC2C" blockId="26.[163,1426,203,1186]" box="[163,1425,931,956]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10808872D1FC2C" box="[163,1425,931,956]" pageId="26" pageNumber="241">
11 (10) Cerci with middle bulge not set off from slightly tapered apical 1/4 (l)
<taxonomicName id="4C6C4D53CE6E9877FA91808872D1FC2C" authorityName="Hagen" authorityYear="1867" box="[1314,1425,932,956]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="fumosus">
<emphasis id="B918EAC2CE6E9877FA91808872D1FC2C" box="[1314,1425,932,956]" italics="true" pageId="26" pageNumber="241">A. fumosus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF1080E072D1FC74" blockId="26.[163,1426,203,1186]" box="[163,1425,972,996]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1080E072D1FC74" box="[163,1425,972,996]" pageId="26" pageNumber="241">
- Cerci with middle bulge set off from parallel sided apical 1/4 (m)
<taxonomicName id="4C6C4D53CE6E9877FABE80E072D1FC74" authorityName="Lieftinck" authorityYear="1942" box="[1293,1425,972,996]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="B918EAC2CE6E9877FABE80E072D1FC74" box="[1293,1425,972,996]" italics="true" pageId="26" pageNumber="241">A. celebensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF1080D876B3FBC2" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF1080D872D2FB9C" blockId="26.[163,1426,203,1186]" box="[163,1426,1012,1036]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1080D872D2FB9C" box="[163,1426,1012,1036]" pageId="26" pageNumber="241">
12 (9) Cerci with small apico-lateral spike pointing caudo-laterad, making them appear pointed (n)
<taxonomicName id="4C6C4D53CE6E9877FA9780D872D2FB9C" baseAuthorityName="Asahina" baseAuthorityYear="1940" box="[1316,1426,1012,1036]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE6E9877FA9780D872D2FB9C" box="[1316,1426,1012,1036]" italics="true" pageId="26" pageNumber="241">A. guttatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF10873176B3FBC2" blockId="26.[163,1426,203,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10873176B3FBC2" pageId="26" pageNumber="241">- Cerci with tiny apico-lateral spike pointing straight backward or slightly incurved, making them 13 appear rounded (op)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30982E75CE6E9877FF10874E72D2FB32" pageId="26" pageNumber="241">
<paragraph id="8BD336D0CE6E9877FF10874E72D1FBEA" blockId="26.[163,1426,203,1186]" box="[163,1425,1122,1146]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF10874E72D1FBEA" box="[163,1425,1122,1146]" pageId="26" pageNumber="241">
13 (12) Cerci inner margin with middle portion not distinctly bulging (o)
<taxonomicName id="4C6C4D53CE6E9877FAA7874E72D1FBEA" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[1300,1425,1122,1146]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="insulanus">
<emphasis id="B918EAC2CE6E9877FAA7874E72D1FBEA" box="[1300,1425,1122,1146]" italics="true" pageId="26" pageNumber="241">A. insulanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF1087A772D2FB32" blockId="26.[163,1426,203,1186]" box="[163,1426,1162,1186]" pageId="26" pageNumber="241">
<keyLead id="309D95E5CE6E9877FF1087A772D2FB32" box="[163,1426,1162,1186]" pageId="26" pageNumber="241">
- Cerci inner margin with middle portion distinctly bulging (p)
<taxonomicName id="4C6C4D53CE6E9877FA8F87A672D2FB32" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[1340,1426,1162,1186]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="tonga">
<emphasis id="B918EAC2CE6E9877FA8F87A672D2FB32" box="[1340,1426,1162,1186]" italics="true" pageId="26" pageNumber="241">A. tonga</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C376655BCE6E9877FF2487D9737EFA78" pageId="26" pageNumber="241" type="description">
<paragraph id="8BD336D0CE6E9877FF2487D9748FFA80" blockId="26.[151,975,1269,1296]" box="[151,975,1269,1296]" pageId="26" pageNumber="241">
<heading id="D09B81BCCE6E9877FF2487D9748FFA80" bold="true" box="[151,975,1269,1296]" fontSize="11" level="1" pageId="26" pageNumber="241" reason="1">
<emphasis id="B918EAC2CE6E9877FF2487D9748FFA80" bold="true" box="[151,975,1269,1296]" pageId="26" pageNumber="241">
Abdominal pattern (
<figureCitation id="13572A55CE6E9877FE2487D976B3FA9F" box="[407,499,1269,1295]" captionStart="FIGURE 54" captionStartId="22.[151,250,1874,1898]" captionTargetBox="[215,1368,193,1836]" captionTargetId="figure-22@22.[206,1381,181,1850]" captionTargetPageId="22" captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (mo) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (bg) modified from Orr &amp; Kalkman (2015); (hi) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, mo) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened." figureDoi="http://doi.org/10.5281/zenodo.13915812" httpUri="https://zenodo.org/record/13915812/files/figure.png" pageId="26" pageNumber="241">Figs 54</figureCitation>
,
<figureCitation id="13572A55CE6E9877FDB387D9755FFA9F" box="[512,543,1269,1295]" captionStart="FIGURE 55" captionStartId="23.[151,250,1850,1874]" captionTargetBox="[161,1422,192,1813]" captionTargetId="figure-17@23.[151,1436,181,1826]" captionTargetPageId="23" captionText="FIGURE 55. Anax spp., males, dorsal, abdominal characters. (ac) A. insulanus sp. nov.; (df) A. tonga sp. nov.: (a, d) abdominal pattern; (b, e) S23; (c, f) terminalia; (a, d) illustrated by Albert Orr." figureDoi="http://doi.org/10.5281/zenodo.13915814" httpUri="https://zenodo.org/record/13915814/files/figure.png" pageId="26" pageNumber="241">55</figureCitation>
) of the
<taxonomicName id="4C6C4D53CE6E9877FDCE87DA745BFA9F" box="[637,795,1269,1296]" pageId="26" pageNumber="241">
<emphasis id="B918EAC2CE6E9877FDCE87DA75FEFA80" bold="true" box="[637,702,1270,1296]" italics="true" pageId="26" pageNumber="241">Anax</emphasis>
species
</taxonomicName>
(males) keyed
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD336D0CE6E9877FF248612737EFA78" blockId="26.[151,1437,1341,1512]" pageId="26" pageNumber="241">
Unfortunately it was not possible to present all species in colour, but
<figureCitation id="13572A55CE6E9877FC0086127361FAC8" box="[947,1057,1341,1368]" captionStart="FIGURE 54" captionStartId="22.[151,250,1874,1898]" captionTargetBox="[215,1368,193,1836]" captionTargetId="figure-22@22.[206,1381,181,1850]" captionTargetPageId="22" captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (mo) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (bg) modified from Orr &amp; Kalkman (2015); (hi) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, mo) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened." figureDoi="http://doi.org/10.5281/zenodo.13915812" httpUri="https://zenodo.org/record/13915812/files/figure.png" pageId="26" pageNumber="241">Figure 54</figureCitation>
illustrated the dorsal view of the abdomen of all species, with colours where shown, as known or presumed to occur in life. Because of the small scale and the minute pale pattern elements (barely detectable in preserved specimens) in
<figureCitation id="13572A55CE6E9877FBD886AA73F9FA30" box="[1131,1209,1413,1440]" captionStart="FIGURE 54" captionStartId="22.[151,250,1874,1898]" captionTargetBox="[215,1368,193,1836]" captionTargetId="figure-22@22.[206,1381,181,1850]" captionTargetPageId="22" captionText="FIGURE 54. Anax spp., male abdominal pattern, dorsal: (a) A. papuensis; (b) A. guttatus; (c) A. fumosus; (d) A. selysi; (e) A. gibbosulus; (f) A. pugnax; (g) A. maclachlani; (h) A. insulanus; (i) A. tonga; (j) A. piraticus; (k) A. georgius; (l) A. celebensis; (mo) A. ??? (purportedly A. panybeus); (p, q) A. panybeus; (bg) modified from Orr &amp; Kalkman (2015); (hi) illustrated by Albert Orr; (j) modified from Kennedy (1934); (k, mo) modified from Seehausen (2017); (l) modified from Lieftinck (1942); (p) brightened." figureDoi="http://doi.org/10.5281/zenodo.13915812" httpUri="https://zenodo.org/record/13915812/files/figure.png" pageId="26" pageNumber="241">Fig. 54</figureCitation>
the newly described species
<taxonomicName id="4C6C4D53CE6E9877FF438686763BFA54" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[240,379,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="insulanus">
<emphasis id="B918EAC2CE6E9877FF438686763BFA54" box="[240,379,1450,1476]" italics="true" pageId="26" pageNumber="241">A. insulanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE6E9877FE0686867555FA53" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[437,533,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="tonga">
<emphasis id="B918EAC2CE6E9877FE0686867555FA53" box="[437,533,1450,1476]" italics="true" pageId="26" pageNumber="241">A. tonga</emphasis>
</taxonomicName>
are also illustrated enlarged as known (
<taxonomicName id="4C6C4D53CE6E9877FC6986867325FA54" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[986,1125,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="insulanus">
<emphasis id="B918EAC2CE6E9877FC6986867325FA54" box="[986,1125,1450,1476]" italics="true" pageId="26" pageNumber="241">A. insulanus</emphasis>
</taxonomicName>
) or presumed (
<taxonomicName id="4C6C4D53CE6E9877FAA686867235FA53" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[1301,1397,1450,1476]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="241" phylum="Arthropoda" rank="species" species="tonga">
<emphasis id="B918EAC2CE6E9877FAA686867235FA53" box="[1301,1397,1450,1476]" italics="true" pageId="26" pageNumber="241">A. tonga</emphasis>
</taxonomicName>
) in life, along with enlarged photos of critical characters S23 and terminalia (
<figureCitation id="13572A55CE6E9877FC6E86E27372FA78" box="[989,1074,1485,1512]" captionStart="FIGURE 55" captionStartId="23.[151,250,1850,1874]" captionTargetBox="[161,1422,192,1813]" captionTargetId="figure-17@23.[151,1436,181,1826]" captionTargetPageId="23" captionText="FIGURE 55. Anax spp., males, dorsal, abdominal characters. (ac) A. insulanus sp. nov.; (df) A. tonga sp. nov.: (a, d) abdominal pattern; (b, e) S23; (c, f) terminalia; (a, d) illustrated by Albert Orr." figureDoi="http://doi.org/10.5281/zenodo.13915814" httpUri="https://zenodo.org/record/13915814/files/figure.png" pageId="26" pageNumber="241">Fig. 55</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,66 +1,66 @@
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<mods:affiliation id="0483CE498E13CD0AFBBC98575B53ED91">Department of Biosystematics, Institute of Life Sciences and Technology, Daugavpils University, Parādes Street 1 A, Daugavpils, Latvia, LV- 5401</mods:affiliation>
<mods:namePart id="82F92B09D55CFAC332EDB60FA6E8FBFF">Kalnins, Martins</mods:namePart>
<mods:affiliation id="7567E239E6163EFF1300FD04460C7743">Department of Biosystematics, Institute of Life Sciences and Technology, Daugavpils University, Parādes Street 1 A, Daugavpils, Latvia, LV- 5401</mods:affiliation>
</mods:name>
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<mods:roleTerm id="640C9658AEE5B18A24C806A43A294E2A">Author</mods:roleTerm>
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<mods:namePart id="B4CAB2953765F0558D646473F39028A5">Marinov, Milen</mods:namePart>
<mods:affiliation id="D7AA51E906CFF277B0EB1237EE6425AB">Biosecurity Surveillance &amp; Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8544, New Zealand</mods:affiliation>
<mods:namePart id="60A5F615D6D4FD321C0CA44843E7FA45">Marinov, Milen</mods:namePart>
<mods:affiliation id="4D1D2B4442F39890921AAD7CDC23401E">Biosecurity Surveillance &amp; Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8544, New Zealand</mods:affiliation>
</mods:name>
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<mods:date id="12A81BF85335260D59A3F9B883FCC0A4">2024</mods:date>
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<treatment id="03C587C6CE779863FF24858F7362FB81" LSID="urn:lsid:plazi:treatment:03C587C6CE779863FF24858F7362FB81" httpUri="http://treatment.plazi.org/id/03C587C6CE779863FF24858F7362FB81" lastPageId="14" lastPageNumber="229" pageId="3" pageNumber="218">
<treatment id="03C587C6CE779862FF24858F74A0F9C1" LSID="urn:lsid:plazi:treatment:03C587C6CE779862FF24858F74A0F9C1" httpUri="http://treatment.plazi.org/id/03C587C6CE779862FF24858F74A0F9C1" lastPageId="15" lastPageNumber="230" pageId="3" pageNumber="218">
<subSubSection id="C376655BCE77986EFF24858F76FCF92D" box="[151,444,1698,1725]" pageId="3" pageNumber="218" type="nomenclature">
<paragraph id="8BD336D0CE77986EFF24858F76FCF92D" blockId="3.[151,785,1698,1761]" box="[151,444,1698,1725]" pageId="3" pageNumber="218">
<heading id="D09B81BCCE77986EFF24858F76FCF92D" bold="true" box="[151,444,1698,1725]" fontSize="11" level="1" pageId="3" pageNumber="218" reason="1">
<emphasis id="B918EAC2CE77986EFF24858F76FCF92D" bold="true" box="[151,444,1698,1725]" pageId="3" pageNumber="218">
<taxonomicName id="4C6C4D53CE77986EFF24858F7617F92D" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[151,343,1698,1725]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="insulanus" status="sp. nov.">
<taxonomicName id="4C6C4D53CE77986EFF24858F7617F92D" authority="Theischinger &amp; Kalnins &amp; Marinov, 2024" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[151,343,1698,1725]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="insulanus" status="sp. nov.">
<emphasis id="B918EAC2CE77986EFF24858F7617F92D" bold="true" box="[151,343,1698,1725]" italics="true" pageId="3" pageNumber="218">Anax insulanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57B9CE77986EFEED858F76FCF92D" box="[350,444,1699,1725]" pageId="3" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
@ -74,12 +74,16 @@
<figureCitation id="13572A55CE77986EFF2C85EB77A6F971" box="[159,230,1735,1761]" captionStart="FIGURES 16" captionStartId="4.[151,264,1672,1696]" captionTargetBox="[152,1415,181,1624]" captionTargetId="figure-22@4.[151,1435,181,1648]" captionTargetPageId="4" captionText="FIGURES 16. Anax spp., male terminalia, dorsal: (1) Anax sp. Q; (2) A. fumosus celebense; (3) A. fumosus (from Orr &amp; Kalkman 2015); (4) A. panybeus; (5) A. piraticus (from Kennedy 1934); (6) A. strenuus (from Martin 1909)." figureDoi="http://doi.org/10.5281/zenodo.13915773" httpUri="https://zenodo.org/record/13915773/files/figure.png" pageId="3" pageNumber="218">Figs 1</figureCitation>
,
<figureCitation id="13572A55CE77986EFF4785EB7652F971" box="[244,274,1735,1761]" captionStart="FIGURE 1317" captionStartId="6.[151,250,476,500]" captionTargetBox="[409,1172,181,452]" captionTargetId="figure-22@6.[408,1180,181,452]" captionTargetPageId="6" captionText="FIGURE 1317. Similarities: Anax spp., male terminalia, dorsal: (13) A. guttatus; (14) A. insulanus; (15) A. tonga; (16) A. maclachlani; (17) A. gibbosulus." figureDoi="http://doi.org/10.5281/zenodo.13915777" httpUri="https://zenodo.org/record/13915777/files/figure.png" pageId="3" pageNumber="218">14</figureCitation>
, 20, 24, 26, 29, 32, 33, 36, 39, 41, 43, 4547)
,
<figureCitation id="13572A55CE77986EFEAD85EB7626F971" box="[286,358,1735,1761]" captionStart="FIGURES 1825" captionStartId="6.[151,264,1868,1892]" captionTargetBox="[151,1422,561,1845]" captionTargetId="figure-54@6.[151,1435,561,1845]" captionTargetPageId="6" captionText="FIGURES 1825. Anax spp., male, dorsal: (1821) S23; (2225) terminalia: (18, 22) A. tonga; (19, 23) A. celebensis; (20, 24) A. insulanus; (21, 25) A. piraticus." pageId="3" pageNumber="218">20, 24</figureCitation>
,
<figureCitation id="13572A55CE77986EFEC185EB76FAF971" box="[370,442,1735,1761]" captionStart="FIGURES 2631" captionStartId="7.[151,264,1936,1960]" captionTargetBox="[177,1423,181,1894]" captionTargetId="figure-17@7.[151,1436,181,1913]" captionText="FIGURES 2631. Anax spp., female, dorsal: (2628) S23; (2931) terminalia: (26, 29) A. insulanus; (27, 30) A. guttatus; (28, 31) A. gibbosulus." pageId="3" pageNumber="218">26, 29</figureCitation>
, 32, 33, 36, 39, 41, 43, 4547)
</paragraph>
</subSubSection>
<subSubSection id="C376655BCE779865FF2484237577FC18" lastPageId="8" lastPageNumber="223" pageId="3" pageNumber="218" type="materials_examined">
<paragraph id="8BD336D0CE77986EFF248423766AF8E1" blockId="3.[151,1436,1806,1977]" pageId="3" pageNumber="218">
<materialsCitation id="3B043C8DCE77986EFF248423766AF8E1" collectingDate="2023-04-21" collectionCode="AM" collectorName="Martins Kalnins" country="Solomon Islands" elevation="400" latitude="-9.4827" location="Davigori" longLatPrecision="7" longitude="159.9828" municipality="Barana" pageId="3" pageNumber="218" specimenCount="1" specimenCount-male="1" stateProvince="Guadalcanal" typeStatus="holotype">
<materialsCitation id="3B043C8DCE77986EFF248423766AF8E1" collectingDate="2023-04-21" collectionCode="AM" collectorName="Martins Kalnins" country="Solomon Islands" elevation="400" latitude="-9.4827" location="Barana vill." longLatPrecision="7" longitude="159.9828" municipality="Honiara" pageId="3" pageNumber="218" specimenCount="1" specimenCount-male="1" stateProvince="Guadalcanal Is." typeStatus="holotype">
<emphasis id="B918EAC2CE77986EFF248423764DF8B9" bold="true" box="[151,269,1807,1833]" pageId="3" pageNumber="218">
<typeStatus id="54D78872CE77986EFF2484237648F8B9" box="[151,264,1807,1833]" pageId="3" pageNumber="218" type="holotype">Holotype</typeStatus>
:
@ -90,15 +94,18 @@
<collectingCountry id="F37B7640CE77986EFE048422753BF8B9" box="[439,635,1806,1833]" name="Solomon Islands" pageId="3" pageNumber="218">Solomon Islands</collectingCountry>
</emphasis>
,
<collectingRegion id="49A8F832CE77986EFD3B84237454F8B9" box="[648,788,1807,1833]" country="Solomon Islands" name="Guadalcanal" pageId="3" pageNumber="218">Guadalcanal</collectingRegion>
Is.,
<collectingRegion id="49A8F832CE77986EFCF1842374DCF8B9" box="[834,924,1807,1833]" country="Solomon Islands" name="Honiara" pageId="3" pageNumber="218">Honiara</collectingRegion>
<collectingRegion id="49A8F832CE77986EFD3B84237476F8B9" box="[648,822,1807,1833]" pageId="3" pageNumber="218">Guadalcanal Is.</collectingRegion>
,
<collectingMunicipality id="6BB7ACAACE77986EFCF1842374DCF8B9" box="[834,924,1807,1833]" pageId="3" pageNumber="218">Honiara</collectingMunicipality>
S env., ~
<locationDeviation id="9A612FF2CE77986EFBB284237321F8B9" bearing="135.0" box="[1025,1121,1807,1833]" distance="2000.0" pageId="3" pageNumber="218">
<quantity id="4C949B35CE77986EFBB28423737AF8B9" box="[1025,1082,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="218" unit="km" value="2.0">2 km</quantity>
SE from
<collectingMunicipality id="6BB7ACAACE77986EFB10842373B2F8B9" box="[1187,1266,1807,1833]" pageId="3" pageNumber="218">Barana</collectingMunicipality>
vill.,
<location id="8EB3600BCE77986EFA81842372D8F8B9" LSID="urn:lsid:plazi:treatment:03C587C6CE779863FF24858F7362FB81:8EB3600BCE77986EFA81842372D8F8B9" box="[1330,1432,1807,1833]" country="Solomon Islands" latitude="-9.4827" longLatPrecision="7" longitude="159.9828" municipality="Barana" name="Davigori" pageId="3" pageNumber="218" stateProvince="Guadalcanal">Davigori</location>
SE
</locationDeviation>
from
<location id="8EB3600BCE77986EFB1084237267F8B9" LSID="urn:lsid:plazi:treatment:03C587C6CE779862FF24858F74A0F9C1:8EB3600BCE77986EFB1084237267F8B9" box="[1187,1319,1807,1833]" country="Solomon Islands" latitude="-9.4827" longLatPrecision="7" longitude="159.9828" municipality="Honiara" name="Barana vill." pageId="3" pageNumber="218" stateProvince="Guadalcanal Is.">Barana vill.</location>
,
<location id="8EB3600BCE77986EFA81842372D8F8B9" LSID="urn:lsid:plazi:treatment:03C587C6CE779862FF24858F74A0F9C1:8EB3600BCE77986EFA81842372D8F8B9" box="[1330,1432,1807,1833]" country="Solomon Islands" latitude="-9.4827" longLatPrecision="7" longitude="159.9828" municipality="Honiara" name="Davigori" pageId="3" pageNumber="218" stateProvince="Guadalcanal Is.">Davigori</location>
,
<geoCoordinate id="EE585017CE77986EFF24841E77BFF8DD" box="[151,255,1842,1869]" degrees="9.4827" direction="south" orientation="latitude" pageId="3" pageNumber="218" precision="5" value="-9.4827">9.4827°S</geoCoordinate>
/
@ -119,7 +126,7 @@ leg. (in
</materialsCitation>
</paragraph>
<paragraph id="8BD336D0CE77986EFF748457768EF829" blockId="3.[151,1436,1806,1977]" pageId="3" pageNumber="218">
<materialsCitation id="3B043C8DCE77986EFF748457768AF829" collectingDate="2023-05-14" collectionCode="LINC" collectorName="Martins Kalnins" country="Solomon Islands" elevation="400" latitude="-9.4827" location="Davigori" longLatPrecision="7" longitude="159.9828" municipality="Barana" pageId="3" pageNumber="218" specimenCount="4" specimenCount-male="4" stateProvince="Guadalcanal" typeStatus="paratype">
<materialsCitation id="3B043C8DCE77986EFF748457768AF829" collectingDate="2023-05-14" collectionCode="AM, LINC" collectorName="Martins Kalnins" country="Solomon Islands" elevation="400" latitude="-9.4827" location="Solomon Islands" longLatPrecision="7" longitude="159.9828" municipality="Honiara" pageId="3" pageNumber="218" specimenCode="K.646375, Odo_2023_033" specimenCount="4" specimenCount-male="4" stateProvince="Guadalcanal Is." typeStatus="paratype">
<emphasis id="B918EAC2CE77986EFF748457755AF805" bold="true" box="[199,538,1914,1941]" pageId="3" pageNumber="218">
<typeStatus id="54D78872CE77986EFF7484577603F805" box="[199,323,1915,1941]" pageId="3" pageNumber="218" type="paratype">Paratypes</typeStatus>
:
@ -133,9 +140,15 @@ leg. (in
</date>
, other data as for holotype: (
<specimenCount id="9D6AFD59CE77986EFBD6845773C8F805" box="[1125,1160,1915,1941]" count="1" pageId="3" pageNumber="218" type="male">1♂</specimenCount>
(K.646375) in AM,
(
<specimenCode id="DBCA9EABCE77986EFB2A8456724BF805" box="[1177,1291,1914,1941]" pageId="3" pageNumber="218">K.646375</specimenCode>
) in
<collectionCode id="ED7DAE15CE77986EFA888457722BF805" box="[1339,1387,1915,1941]" pageId="3" pageNumber="218">AM</collectionCode>
,
<specimenCount id="9D6AFD59CE77986EFACA845772DCF805" box="[1401,1436,1915,1941]" count="1" pageId="3" pageNumber="218" type="male">1♂</specimenCount>
(Odo_2023_033) in
(
<specimenCode id="DBCA9EABCE77986EFF2C84B37610F829" box="[159,336,1951,1977]" pageId="3" pageNumber="218">Odo_2023_033</specimenCode>
) in
<collectionCode id="ED7DAE15CE77986EFECF84B37682F829" box="[380,450,1951,1977]" pageId="3" pageNumber="218">LINC</collectionCode>
)
</materialsCitation>
@ -257,6 +270,7 @@ spp.
.
</paragraph>
</caption>
<caption id="DF136658CE72986BFF248460755EF818" pageId="6" pageNumber="221" startId="6.[151,264,1868,1892]" targetBox="[151,1422,561,1845]" targetPageId="6">
<paragraph id="8BD336D0CE72986BFF248460755EF818" blockId="6.[151,1436,1868,1929]" pageId="6" pageNumber="221">
<emphasis id="B918EAC2CE72986BFF2484607616F8F4" bold="true" box="[151,342,1868,1892]" pageId="6" pageNumber="221">FIGURES 1825.</emphasis>
<taxonomicName id="4C6C4D53CE72986BFEEE84617685F8F4" box="[349,453,1869,1892]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="221" phylum="Arthropoda" rank="species" species="undetermined">
@ -282,6 +296,8 @@ spp.
.
</emphasis>
</paragraph>
</caption>
<caption id="DF136658CE73986AFF2484BC7610F85D" pageId="7" pageNumber="222" startId="7.[151,264,1936,1960]" targetBox="[177,1423,181,1894]" targetPageId="7">
<paragraph id="8BD336D0CE73986AFF2484BC7610F85D" blockId="7.[151,1436,1936,1997]" pageId="7" pageNumber="222">
<emphasis id="B918EAC2CE73986AFF2484BC7614F839" bold="true" box="[151,340,1936,1961]" pageId="7" pageNumber="222">FIGURES 2631.</emphasis>
<taxonomicName id="4C6C4D53CE73986AFEEA84BD76FFF838" box="[345,447,1937,1960]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="222" phylum="Arthropoda" rank="species" species="undetermined">
@ -302,6 +318,7 @@ spp.
.
</emphasis>
</paragraph>
</caption>
<caption id="DF136658CE7C9865FF2481077494FDF8" ID-DOI="http://doi.org/10.5281/zenodo.13915779" ID-Zenodo-Dep="13915779" httpUri="https://zenodo.org/record/13915779/files/figure.png" pageId="8" pageNumber="223" startId="8.[151,264,555,579]" targetBox="[436,1148,183,528]" targetPageId="8" targetType="figure">
<paragraph id="8BD336D0CE7C9865FF2481077494FDF8" blockId="8.[151,1436,555,616]" pageId="8" pageNumber="223">
<emphasis id="B918EAC2CE7C9865FF2481077617FDD4" bold="true" box="[151,343,555,580]" pageId="8" pageNumber="223">FIGURES 3235.</emphasis>
@ -321,11 +338,14 @@ spp.
</paragraph>
</caption>
<paragraph id="8BD336D0CE7C9865FF74819677B0FC8C" blockId="8.[151,1436,698,976]" pageId="8" pageNumber="223">
<materialsCitation id="3B043C8DCE7C9865FF748196730BFD44" box="[199,1099,698,724]" collectingDate="1999-06-30" country="Georgia" location="New" municipality="Munda" pageId="8" pageNumber="223" specimenCount="2" specimenCount-female="2">
<materialsCitation id="3B043C8DCE7C9865FF748196730BFD44" box="[199,1099,698,724]" collectingDate="1999-06-30" collectionCode="K" country="Solomon Islands" location="Munda" municipality="Munda" pageId="8" pageNumber="223" specimenCode="K.646377, K.646379" specimenCount="2" specimenCount-female="2" stateProvince="New Georgia">
<specimenCount id="9D6AFD59CE7C9865FF74819677BFFD44" box="[199,255,698,724]" count="2" pageId="8" pageNumber="223" type="female">2♀♀</specimenCount>
(K.646377, K.646379),
<location id="8EB3600BCE7C9865FDA081967508FD44" LSID="urn:lsid:plazi:treatment:03C587C6CE779863FF24858F7362FB81:8EB3600BCE7C9865FDA081967508FD44" box="[531,584,698,724]" country="Georgia" municipality="Munda" name="New" pageId="8" pageNumber="223">New</location>
<collectingCountry id="F37B7640CE7C9865FDFC819675ECFD44" box="[591,684,698,724]" name="Georgia" pageId="8" pageNumber="223">Georgia</collectingCountry>
(
<specimenCode id="DBCA9EABCE7C9865FEBD819676C1FD44" box="[270,385,698,724]" pageId="8" pageNumber="223">K.646377</specimenCode>
,
<specimenCode id="DBCA9EABCE7C9865FE3E81967540FD44" box="[397,512,698,724]" pageId="8" pageNumber="223">K.646379</specimenCode>
),
<collectingRegion id="49A8F832CE7C9865FDA0819675ECFD44" box="[531,684,698,724]" pageId="8" pageNumber="223">New Georgia</collectingRegion>
,
<collectingMunicipality id="6BB7ACAACE7C9865FD0581967448FD44" box="[694,776,698,724]" pageId="8" pageNumber="223">Munda</collectingMunicipality>
,
@ -334,9 +354,11 @@ spp.
</date>
, together with
</materialsCitation>
<materialsCitation id="3B043C8DCE7C9865FBE181967516FD68" collectionCode="AM" collectorName="D. M. Reeves &amp; J. H. Bryan" country="Georgia" pageId="8" pageNumber="223" specimenCount="1" specimenCount-male="1">
<materialsCitation id="3B043C8DCE7C9865FBE181967516FD68" collectionCode="AM" collectorName="D. M. Reeves &amp; J. H. Bryan" country="Georgia" pageId="8" pageNumber="223" specimenCode="K.646378" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D6AFD59CE7C9865FBE181967335FD44" box="[1106,1141,698,724]" count="1" pageId="8" pageNumber="223" type="male">1♂</specimenCount>
(K.646378) of
(
<specimenCode id="DBCA9EABCE7C9865FB37819673B6FD44" box="[1156,1270,698,724]" pageId="8" pageNumber="223">K.646378</specimenCode>
) of
<taxonomicName id="4C6C4D53CE7C9865FA90819672DCFD44" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[1315,1436,698,724]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="223" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE7C9865FA90819672DCFD44" box="[1315,1436,698,724]" italics="true" pageId="8" pageNumber="223">A. guttatus</emphasis>
</taxonomicName>
@ -349,9 +371,11 @@ spp.
leg.
</materialsCitation>
;
<materialsCitation id="3B043C8DCE7C9865FDD381F277ACFC8C" collectingDate="1999-07-01" collectionCode="AM" collectorName="D. M. Reeves" country="Georgia" pageId="8" pageNumber="223" specimenCount="1" specimenCount-male="1">
<materialsCitation id="3B043C8DCE7C9865FDD381F277ACFC8C" collectingDate="1999-07-01" collectionCode="AM" collectorName="D. M. Reeves" country="Georgia" location="Munda" municipality="Munda" pageId="8" pageNumber="223" specimenCode="K.646376" specimenCount="1" specimenCount-male="1" stateProvince="New Georgia">
<specimenCount id="9D6AFD59CE7C9865FDD381F275C3FD68" box="[608,643,734,760]" count="1" pageId="8" pageNumber="223" type="male">1♂</specimenCount>
(K.646376), same locality,
(
<specimenCode id="DBCA9EABCE7C9865FD2381F27445FD68" box="[656,773,734,760]" pageId="8" pageNumber="223">K.646376</specimenCode>
), same locality,
<date id="FFD21010CE7C9865FC0781F27371FD68" box="[948,1073,734,760]" pageId="8" pageNumber="223" value="1999-07-01">
<collectingDate id="EF96E9F8CE7C9865FC0781F27371FD68" box="[948,1073,734,760]" pageId="8" pageNumber="223" value="1999-07-01">1 July1999</collectingDate>
</date>
@ -406,7 +430,7 @@ The name refers to the presence of the species on several of the
; insulanus is Latin for island inhabitant and is used as a noun in apposition to the generic name.
</paragraph>
</subSubSection>
<subSubSection id="C376655BCE7C9863FF24870E7362FB81" lastPageId="14" lastPageNumber="229" pageId="8" pageNumber="223" type="description">
<subSubSection id="C376655BCE7C9862FF24870E7268FB82" lastPageId="15" lastPageNumber="230" pageId="8" pageNumber="223" type="description">
<paragraph id="8BD336D0CE7C9865FF24870E76FBFBAC" blockId="8.[151,443,1058,1084]" box="[151,443,1058,1084]" pageId="8" pageNumber="223">
<emphasis id="B918EAC2CE7C9865FF24870E76FBFBAC" bold="true" box="[151,443,1058,1084]" pageId="8" pageNumber="223">
Description of
@ -580,6 +604,161 @@ Cerci brownish black, markedly longer than width of S10, leaf-shaped with inner
<emphasis id="B918EAC2CE7A9863FF7480DB7690FB81" box="[199,464,1015,1041]" italics="true" pageId="14" pageNumber="229">Measurements (in mm).</emphasis>
Abdomen (including cerci) 55.6, cerci 4.2, Hw 52.2.
</paragraph>
<caption id="DF136658CE7A9863FF24843F758DF8DF" ID-DOI="http://doi.org/10.5281/zenodo.13915798" ID-Zenodo-Dep="13915798" httpUri="https://zenodo.org/record/13915798/files/figure.png" pageId="14" pageNumber="229" startId="14.[151,250,1811,1835]" targetBox="[151,1436,1065,1788]" targetPageId="14" targetType="figure">
<paragraph id="8BD336D0CE7A9863FF24843F758DF8DF" blockId="14.[151,1436,1811,1871]" pageId="14" pageNumber="229">
<emphasis id="B918EAC2CE7A9863FF24843F7660F8BB" bold="true" box="[151,288,1811,1835]" pageId="14" pageNumber="229">FIGURE 47.</emphasis>
Surrounding garden landscape ~400 m above sea level—
<taxonomicName id="4C6C4D53CE7A9863FCDB84387343F8BB" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[872,1027,1811,1835]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="229" phylum="Arthropoda" rank="species" species="insulanus" status="sp. nov.">
<emphasis id="B918EAC2CE7A9863FCDB84387343F8BB" box="[872,1027,1811,1835]" italics="true" pageId="14" pageNumber="229">Anax insulanus</emphasis>
</taxonomicName>
<emphasis id="B918EAC2CE7A9863FBBA843F731AF8BB" bold="true" box="[1033,1114,1811,1835]" pageId="14" pageNumber="229">
<taxonomicNameLabel id="A22B57B9CE7A9863FBBA843F731AF8BB" box="[1033,1114,1811,1835]" pageId="14" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
habitat, Honiara S environs, ~2 km SE from Barana vill., Davigori, Guadalcanal Island.
</paragraph>
</caption>
<paragraph id="8BD336D0CE7B9862FF2483BB769FFF21" blockId="15.[151,479,151,177]" box="[151,479,151,177]" pageId="15" pageNumber="230">
<heading id="D09B81BCCE7B9862FF2483BB769FFF21" bold="true" box="[151,479,151,177]" fontSize="11" level="1" pageId="15" pageNumber="230" reason="1">
<emphasis id="B918EAC2CE7B9862FF2483BB769FFF21" bold="true" box="[151,479,151,177]" pageId="15" pageNumber="230">
Variation in the
<typeStatus id="54D78872CE7B9862FED083BB769FFF21" box="[355,479,151,177]" pageId="15" pageNumber="230" type="paratype">paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD336D0CE7B9862FF2483F374F4FE8D" blockId="15.[151,1437,223,1618]" pageId="15" pageNumber="230">
<specimenCount id="9D6AFD59CE7B9862FF2483F37655FF69" box="[151,277,223,249]" count="3" pageId="15" pageNumber="230" type="male">Three male</specimenCount>
<typeStatus id="54D78872CE7B9862FEAF83CC76C7FF6A" box="[284,391,224,250]" pageId="15" pageNumber="230" type="paratype">paratypes</typeStatus>
and
<specimenCount id="9D6AFD59CE7B9862FE0F83CC7577FF69" box="[444,567,223,250]" count="1" pageId="15" pageNumber="230" type="female">one female</specimenCount>
<typeStatus id="54D78872CE7B9862FD8E83CC75DDFF6A" box="[573,669,224,250]" pageId="15" pageNumber="230" type="paratype">paratype</typeStatus>
are available. Their overall coloration ranges from distinctly paler to barely darker than that given in the description of
<typeStatus id="54D78872CE7B9862FD7B822F746AFE8D" box="[712,810,259,285]" pageId="15" pageNumber="230" type="holotype">holotype</typeStatus>
and female.
</paragraph>
<paragraph id="8BD336D0CE7B9862FF74820B74F4FCAA" blockId="15.[151,1437,223,1618]" pageId="15" pageNumber="230">
The heads of the male
<typeStatus id="54D78872CE7B9862FE7A82047574FED2" box="[457,564,296,322]" pageId="15" pageNumber="230" type="paratype">paratypes</typeStatus>
largely agree with the
<typeStatus id="54D78872CE7B9862FC8A820B74DEFED1" box="[825,926,295,321]" pageId="15" pageNumber="230" type="holotype">holotype</typeStatus>
, but the brownish black T-mark on top of the frons ranges from rather thin-stemmed to very thick-stemmed. The coloration of pterothorax and antealar sinus is pale green and greyish green as in the
<typeStatus id="54D78872CE7B9862FDE5824375F8FE19" box="[598,696,367,393]" pageId="15" pageNumber="230" type="holotype">holotype</typeStatus>
to pale orange brown or yellowish to brownish grey. The wings are much as in the
<typeStatus id="54D78872CE7B9862FEC182BF7694FE3D" box="[370,468,403,429]" pageId="15" pageNumber="230" type="holotype">holotype</typeStatus>
but humeral sclerites can also be greyish green and axillary sclerites greyish yellow. Venation greyish brown to black. The membrane ranges from Hw only slightly pale brownish tinged between the level of the anal loop and MA, added to which is slight brownish suffusion of almost all remaining Fw. Nodal index 89, 1922 / 2021,
<quantity id="4C949B35CE7B9862FE3182D37697FD89" box="[386,471,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="15" pageNumber="230" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">910 in</quantity>
Fw and 1012, 14 / 1214,
<quantity id="4C949B35CE7B9862FCA482D37439FD89" box="[791,889,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.556" metricValueMin="2.794" pageId="15" pageNumber="230" unit="in" value="12.5" valueMax="14.0" valueMin="11.0">1114 in</quantity>
Hw; Ax1 and Ax7 or Ax8 thickened in Fw, Ax1 and Ax6 or Ax
<quantity id="4C949B35CE7B9862FE8F810F7626FDAD" box="[316,358,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="15" pageNumber="230" unit="in" value="7.0">7 in</quantity>
Hw; wing triangles with 68 cells in Fw, with 46 cells in Hw; Arc from halfway between level Ax1 and Ax2 to very close to level Ax2; Pt braced, greyish brown to black, overlying 2 and 1/3 to 3 and 1/2 cells; 610 double cells, if 8 or 10, 3 triple, between MP and CuA of Hw; anal loop with 1215 cells. The abdomen closely agrees with the
<typeStatus id="54D78872CE7B9862FEF681A376EAFD39" box="[325,426,655,681]" pageId="15" pageNumber="230" type="holotype">holotype</typeStatus>
, but S3 Index 5.97.5. In
<specimenCount id="9D6AFD59CE7B9862FD7681BC742AFD39" box="[709,874,655,682]" count="2" pageId="15" pageNumber="230" type="generic">two specimens</specimenCount>
much of S2 and anterior half of S3 markedly more vividly whitish, blue and black. S48 black and barely detectably smudged with brown along lateral margin, S49 or S59 each with pair of small ill-defined dull whitish yellow to pale greenish subapical dorso-lateral spots; S9 latero-basally black to black smudged with dull yellowish. The terminalia closely agree with the
<typeStatus id="54D78872CE7B9862FB7281D77267FC85" box="[1217,1319,763,789]" pageId="15" pageNumber="230" type="holotype">holotype</typeStatus>
. Abdomen (including cerci) 63.267.0 mm, cerci
<quantity id="4C949B35CE7B9862FDF58033758FFCAA" box="[582,719,799,826]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="15" pageNumber="230" unit="mm" value="6.35" valueMax="6.5" valueMin="6.2">6.26.5 mm</quantity>
, Hw 52.956.0 mm.
</paragraph>
<paragraph id="8BD336D0CE7B9862FF74806F7268FB82" blockId="15.[151,1437,223,1618]" pageId="15" pageNumber="230">
The second and only other female
<typeStatus id="54D78872CE7B9862FDF1806875E2FCCE" box="[578,674,836,862]" pageId="15" pageNumber="230" type="paratype">paratype</typeStatus>
(K.646379) largely agrees well with
<typeStatus id="54D78872CE7B9862FB8E806873DDFCCE" box="[1085,1181,836,862]" pageId="15" pageNumber="230" type="paratype">paratype</typeStatus>
K. 646377 (see above) regarding head, thorax, abdomen and terminalia. The T-mark on top of the frons has a slightly more slender stem. However, the Fw membrane is slightly tinged pale brownish almost all over. Nodal index 918 / 19
<quantity id="4C949B35CE7B9862FABB80A77201FC35" box="[1288,1345,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="230" unit="in" value="10.0">10 in</quantity>
Fw and broken-14 / 14
<quantity id="4C949B35CE7B9862FEF8808376C3FC59" box="[331,387,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="15" pageNumber="230" unit="in" value="12.0">12 in</quantity>
Hw; Ax1 and Ax7 thickened in Fw, Ax1 and Ax6 or Ax
<quantity id="4C949B35CE7B9862FC4A80837363FC59" box="[1017,1059,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="15" pageNumber="230" unit="in" value="7.0">7 in</quantity>
Hw; Pt overlying 2 and 1/2 cells; 68 double cells between MP and CuA of Hw; anal loop with 1012 cells. The S3 Index is c. 4.5. There is no mid-dorsal yellow to orange patch on S10. Abdomen (including cerci)
<quantity id="4C949B35CE7B9862FCC880DB74A1FB82" box="[891,993,1015,1042]" metricMagnitude="-2" metricUnit="m" metricValue="5.93" pageId="15" pageNumber="230" unit="mm" value="59.3">59.3 mm</quantity>
, cerci
<quantity id="4C949B35CE7B9862FB9980DB73C2FB82" box="[1066,1154,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="15" pageNumber="230" unit="mm" value="4.6">4.6 mm</quantity>
, Hw
<quantity id="4C949B35CE7B9862FB0E80DB7264FB82" box="[1213,1316,1015,1042]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="15" pageNumber="230" unit="mm" value="55.6">55.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C376655BCE7B9862FF748737765BFAE9" pageId="15" pageNumber="230" type="diagnosis">
<paragraph id="8BD336D0CE7B9862FF748737765BFAE9" blockId="15.[151,1437,223,1618]" pageId="15" pageNumber="230">
<emphasis id="B918EAC2CE7B9862FF7487377602FBA5" bold="true" box="[199,322,1051,1077]" pageId="15" pageNumber="230">Diagnosis.</emphasis>
A rather large dark short-bodied
<taxonomicName id="4C6C4D53CE7B9862FD70873775BDFBA5" authorityName="Leach" authorityYear="1815" box="[707,765,1051,1077]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAC2CE7B9862FD70873775BDFBA5" box="[707,765,1051,1077]" italics="true" pageId="15" pageNumber="230">Anax</emphasis>
</taxonomicName>
(body length c.
<quantity id="4C949B35CE7B9862FC0E87377350FBA6" box="[957,1040,1051,1078]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="15" pageNumber="230" unit="mm" value="85.0">85 mm</quantity>
, wingspan c.
<quantity id="4C949B35CE7B9862FB0387377250FBA6" box="[1200,1296,1051,1078]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="230" unit="mm" value="110.0">110 mm</quantity>
, Abd
<quantity id="4C949B35CE7B9862FAE587377785FBCA" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="6.7" metricValueMin="6.3" pageId="15" pageNumber="230" unit="mm" value="65.0" valueMax="67.0" valueMin="63.0">6367 mm</quantity>
, Hw
<quantity id="4C949B35CE7B9862FEB38713763CFBCA" box="[256,380,1087,1114]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="15" pageNumber="230" unit="mm" value="54.5" valueMax="56.0" valueMin="53.0">5356 mm</quantity>
; Ratio &lt;1.3) with high S3 Index (male: 5.97.5, female 4.04.5). Mature male in life (
<figureCitation id="13572A55CE7B9862FAF1871372D4FBC9" box="[1346,1428,1087,1113]" captionStart="FIGURE 36" captionStartId="9.[151,250,1576,1600]" captionTargetBox="[151,1436,181,1551]" captionTargetId="figure-17@9.[151,1436,181,1552]" captionTargetPageId="9" captionText="FIGURE 36. Anax insulanus, male, live: (a) part of habitus, lateral; (b) head, frontal; (c) base of abdomen, dorsal; (d) abdomen, lateral. Freshly killed specimen photographed in natural light—a, b, d." figureDoi="http://doi.org/10.5281/zenodo.13915781" httpUri="https://zenodo.org/record/13915781/files/figure.png" pageId="15" pageNumber="230">Fig. 36</figureCitation>
) with head, eyes and thorax largely green, legs and most of abdomen largely black, abdomen base bright green, blue and black. Abdominal pattern of male generally made up by pair of small ill-defined dull greenish yellow to pale yellow subapical dorsolateral spots each on S48 or 9 or S58 or 9, none on S13 and
<quantity id="4C949B35CE7B9862FBEB878773D0FB55" box="[1112,1168,1195,1221]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="230" unit="in" value="10.0">10 in</quantity>
male, on S38, none on S1, 2, 9 and
<quantity id="4C949B35CE7B9862FE9587E3761FFB79" box="[294,351,1231,1257]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="230" unit="in" value="10.0">10 in</quantity>
female. Coloration of preserved material ranges from distinctly paler to darker (
<figureCitation id="13572A55CE7B9862FB5887E37202FB79" box="[1259,1346,1231,1257]" captionStart="FIGURES 3738. 37" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[274,1321,195,1900]" captionTargetId="figure-22@10.[250,1338,181,1920]" captionTargetPageId="10" captionText="FIGURES 3738. 37. Anax insulanus, male, holotype, dorsal: (a) habitus; (b) head; (c) S23; (d) terminalia, dorsal. 38. Anax guttatus, male, from Australia, dorsal: (a) habitus; (b) head; (c) S23; (d) terminalia." figureDoi="http://doi.org/10.5281/zenodo.13915783" httpUri="https://zenodo.org/record/13915783/files/figure.png" pageId="15" pageNumber="230">Figs 37</figureCitation>
,
<figureCitation id="13572A55CE7B9862FAFC87E3722DFB79" box="[1359,1389,1231,1257]" captionStart="FIGURES 3940. 39" captionStartId="11.[151,264,959,983]" captionTargetBox="[155,1435,181,936]" captionTargetId="figure-17@11.[151,1435,181,936]" captionTargetPageId="11" captionText="FIGURES 3940. 39. Anax insulanus, male, Paratype K.646376, lateral/dorsal: (a) habitus; (b) head; (c) S23. 40. Anax guttatus, male (K.646378), from New Georgia, lateral/dorsal: (a) habitus; (b) head; (c) S2S3." figureDoi="http://doi.org/10.5281/zenodo.13915787" httpUri="https://zenodo.org/record/13915787/files/figure.png" pageId="15" pageNumber="230">39</figureCitation>
,
<figureCitation id="13572A55CE7B9862FACA87E372D7FB79" box="[1401,1431,1231,1257]" captionStart="FIGURES 4142. 41" captionStartId="11.[151,264,1863,1887]" captionTargetBox="[151,1436,1044,1840]" captionTargetId="figure-52@11.[151,1436,1044,1840]" captionTargetPageId="11" captionText="FIGURES 4142. 41. Anax insulanus, female, Paratype K.646377, lateral/dorsal: (a) habitus; (b) head; (c) S23; (d) terminalia. 42. Anax guttatus, female, from Australia, lateral/dorsal: (a) habitus; (b) head; (c) S23; (d) terminalia." figureDoi="http://doi.org/10.5281/zenodo.13915789" httpUri="https://zenodo.org/record/13915789/files/figure.png" pageId="15" pageNumber="230">41</figureCitation>
,
<figureCitation id="13572A55CE7B9862FF2487DF77F4FA9D" box="[151,180,1267,1293]" captionStart="FIGURES 4344. 43" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[213,1371,186,1917]" captionTargetId="figure-22@12.[213,1374,181,1919]" captionTargetPageId="12" captionText="FIGURES 4344. 43. Anax insulanus, female, Paratype K.646379, dorsal: (a) habitus; (b) head; (c) S23; (d) terminalia. 44. Anax guttatus, female, from Australia, dorsal: (a) habitus; (b) head; (c) S23; (d) terminalia." figureDoi="http://doi.org/10.5281/zenodo.13915791" httpUri="https://zenodo.org/record/13915791/files/figure.png" pageId="15" pageNumber="230">43</figureCitation>
) than given above, depending on maturity of the individual when collected and on age and state of preservation of specimens.
<taxonomicName id="4C6C4D53CE7B9862FE8E863B76ABFAA1" authorityName="Theischinger &amp; Kalnins &amp; Marinov" authorityYear="2024" box="[317,491,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="230" phylum="Arthropoda" rank="species" species="insulanus">
<emphasis id="B918EAC2CE7B9862FE8E863B76ABFAA1" box="[317,491,1303,1329]" italics="true" pageId="15" pageNumber="230">Anax insulanus</emphasis>
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may be separated from the very similar
<taxonomicName id="4C6C4D53CE7B9862FC70863B7317FAA1" authorityName="Lieftinck" authorityYear="1942" box="[963,1111,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="230" phylum="Arthropoda" rank="species" species="celebensis">
<emphasis id="B918EAC2CE7B9862FC70863B7317FAA1" box="[963,1111,1303,1329]" italics="true" pageId="15" pageNumber="230">A. celebensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D53CE7B9862FB20863B7258FAA1" authorityName="Kennedy" authorityYear="1934" box="[1171,1304,1303,1329]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="230" phylum="Arthropoda" rank="species" species="piraticus">
<emphasis id="B918EAC2CE7B9862FB20863B7258FAA1" box="[1171,1304,1303,1329]" italics="true" pageId="15" pageNumber="230">A. piraticus</emphasis>
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by its very dark abdomen with only a few barely perceptible small paler spots and smudges vs much richer and larger pale maculation.
</paragraph>
</subSubSection>
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<emphasis id="B918EAC2CE7B9862FF7486AF766AFA0D" bold="true" box="[199,298,1411,1437]" pageId="15" pageNumber="230">Habitat.</emphasis>
The
<specimenCount id="9D6AFD59CE7B9862FED686AF76A5FA0D" box="[357,485,1411,1437]" count="3" pageId="15" pageNumber="230" type="male">three males</specimenCount>
of
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<emphasis id="B918EAC2CE7B9862FDBF86AF75D6FA0D" box="[524,662,1411,1437]" italics="true" pageId="15" pageNumber="230">A. insulanus</emphasis>
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were captured near a small artificial concrete pool serving to collect rainwater (
<figureCitation id="13572A55CE7B9862FEA2868B7625FA52" box="[273,357,1447,1474]" captionStart="FIGURE 45" captionStartId="13.[151,250,928,952]" captionTargetBox="[151,1436,181,904]" captionTargetId="figure-17@13.[151,1436,181,904]" captionTargetPageId="13" captionText="FIGURE 45. Anax insulanus sp. nov. habitat (caught place)—small concreted pool for collecting rainwater Honiara S env., ~2 km SE from Barana vill., Davigori, Guadalcanal Island." figureDoi="http://doi.org/10.5281/zenodo.13915793" httpUri="https://zenodo.org/record/13915793/files/figure.png" pageId="15" pageNumber="230">Fig. 45</figureCitation>
), located in a highly disturbed open garden landscape (
<figureCitation id="13572A55CE7B9862FC66868B736DFA51" box="[981,1069,1447,1473]" captionStart="FIGURE 46" captionStartId="13.[151,250,1758,1782]" captionTargetBox="[151,1436,1012,1735]" captionTargetId="figure-47@13.[151,1436,1012,1735]" captionTargetPageId="13" captionText="FIGURE 46. Surrounding garden landscape ~400 m above sea level—Anax insulanus sp. nov. habitat, Honiara S environs, ~2 km SE from Barana vill., Davigori, Guadalcanal Island." figureDoi="http://doi.org/10.5281/zenodo.13915795" httpUri="https://zenodo.org/record/13915795/files/figure.png" pageId="15" pageNumber="230">Figs 46</figureCitation>
,
<figureCitation id="13572A55CE7B9862FB8A868B7316FA51" box="[1081,1110,1447,1473]" captionStart="FIGURE 47" captionStartId="14.[151,250,1811,1835]" captionTargetBox="[151,1436,1065,1788]" captionTargetId="figure-403@14.[151,1436,1065,1788]" captionTargetPageId="14" captionText="FIGURE 47. Surrounding garden landscape ~400 m above sea level—Anax insulanus sp. nov. habitat, Honiara S environs, ~2 km SE from Barana vill., Davigori, Guadalcanal Island." figureDoi="http://doi.org/10.5281/zenodo.13915798" httpUri="https://zenodo.org/record/13915798/files/figure.png" pageId="15" pageNumber="230">47</figureCitation>
) in the vicinity of secondary forest. One patrolling individual was caught on
<date id="FFD21010CE7B9862FD1286E7747DFA75" box="[673,829,1483,1509]" pageId="15" pageNumber="230" value="2023-04-21">21 April 2023</date>
. On the following day,
<date id="FFD21010CE7B9862FB8B86E7738DFA75" box="[1080,1229,1483,1510]" pageId="15" pageNumber="230" value="2023-05-14">14 May 2023</date>
, another patrolling individual was caught there. Later that same day, the vacant territory was occupied by a third individual.
</paragraph>
</subSubSection>
<subSubSection id="C376655BCE7B9862FF74853F74A0F9C1" pageId="15" pageNumber="230" type="distribution">
<paragraph id="8BD336D0CE7B9862FF74853F74A0F9C1" blockId="15.[151,1437,223,1618]" pageId="15" pageNumber="230">
<emphasis id="B918EAC2CE7B9862FF74853F7621F9BD" bold="true" box="[199,353,1555,1581]" pageId="15" pageNumber="230">Distribution.</emphasis>
Known only from Guadalcanal and New
<collectingCountry id="F37B7640CE7B9862FCF6853F74DFF9BD" box="[837,927,1555,1581]" name="Georgia" pageId="15" pageNumber="230">Georgia</collectingCountry>
in the
<collectingCountry id="F37B7640CE7B9862FC41853F73F2F9BD" box="[1010,1202,1555,1581]" name="Solomon Islands" pageId="15" pageNumber="230">Solomon Islands</collectingCountry>
, where it was found coexisting with
<taxonomicName id="4C6C4D53CE7B9862FEF9851B7683F9C1" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[330,451,1591,1617]" class="Insecta" family="Aeshnidae" genus="Anax" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="230" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B918EAC2CE7B9862FEF9851B7683F9C1" box="[330,451,1591,1617]" italics="true" pageId="15" pageNumber="230">A. guttatus</emphasis>
</taxonomicName>
. Possibly endemic to the Solomon Archipelago.
</paragraph>
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</treatment>
</document>

View file

@ -1,60 +1,63 @@
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@ -212,9 +215,9 @@ Diagnosis.—
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has the following diagnostic characters: nasals oriented dorsally; a complete row of lorilabial scales between subocular and supralabial scales; temporal and head scales smooth; frontal usually divided transversely; dorsal color pattern consisting of a mid-dorsal white stripe and two dorsolateral stripes, with two series of paravertebral brown marks resembling triangles bordered by white bar (
<figureCitation id="EAFC2A2EFFC96A4D2CDBF90BFC49F94C" box="[927,1011,1672,1697]" captionStart="FIG" captionStartId="3.[698,713,895,914]" captionTargetBox="[677,1184,171,872]" captionTargetId="figure-12@3.[677,1184,171,875]" captionTargetPageId="3" captionText="FIG. 1.—Adult specimens of Liolaemus arambarensis from Arambaré, Rio Grande do Sul, Brazil (left female, UFRGS 2951, SVL = 55.85 mm; right male, UFRGS 2902, SVL = 59.84 mm = holotype)." pageId="3" pageNumber="107">Figs. 1</figureCitation>
<figureCitation id="EAFC2A2EFFC96A4D2CDBF90BFC49F94C" box="[927,1011,1672,1697]" captionStart="FIG" captionStartId="3.[698,713,895,914]" captionTargetBox="[677,1184,171,872]" captionTargetId="figure-12@3.[677,1184,171,875]" captionTargetPageId="3" captionText="FIG. 1.—Adult specimens of Liolaemus arambarensis from Arambaré, Rio Grande do Sul, Brazil (left female, UFRGS 2951, SVL = 55.85 mm; right male, UFRGS 2902, SVL = 59.84 mm = holotype)." figureDoi="http://doi.org/10.5281/zenodo.13940232" httpUri="https://zenodo.org/record/13940232/files/figure.png" pageId="3" pageNumber="107">Figs. 1</figureCitation>
,
<figureCitation id="EAFC2A2EFFC96A4D2B40F90AFBAFF94C" box="[1028,1045,1673,1697]" captionStart="FIG" captionStartId="4.[130,145,933,952]" captionTargetBox="[107,620,171,912]" captionTargetId="figure-13@4.[107,620,171,912]" captionTargetPageId="4" captionText="FIG. 2.—(A) Dorsal and (B) lateral view of the head of adult male Liolaemus arambarensis (UFRGS 2902; holotype). Scale = 5 mm." pageId="3" pageNumber="107">2</figureCitation>
<figureCitation id="EAFC2A2EFFC96A4D2B40F90AFBAFF94C" box="[1028,1045,1673,1697]" captionStart="FIG" captionStartId="4.[130,145,933,952]" captionTargetBox="[107,620,171,912]" captionTargetId="figure-13@4.[107,620,171,912]" captionTargetPageId="4" captionText="FIG. 2.—(A) Dorsal and (B) lateral view of the head of adult male Liolaemus arambarensis (UFRGS 2902; holotype). Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13940234" httpUri="https://zenodo.org/record/13940234/files/figure.png" pageId="3" pageNumber="107">2</figureCitation>
).
</paragraph>
<paragraph id="727836ABFFC96A4A2D84F925FDA6FB05" blockId="3.[675,1188,1056,1726]" lastBlockId="4.[107,620,1055,1726]" lastPageId="4" lastPageNumber="108" pageId="3" pageNumber="107">
@ -227,7 +230,7 @@ is referrable to the ̒̒
</taxonomicName>
̓̓ group based on the presence of lorilabial scales smaller than supralabial scales and narrow (longer than wide) supralabial scales, the posterior ones being wider; the mental scale is in contact with the sublabial scales and is posteriorly wide; infralabial scales are concave.
</paragraph>
<caption id="26B86623FFCE6A4A2FC6FC26FE19FC05" pageId="4" pageNumber="108" startId="4.[130,145,933,952]" targetBox="[107,620,171,912]" targetPageId="4" targetType="figure">
<caption id="26B86623FFCE6A4A2FC6FC26FE19FC05" ID-DOI="http://doi.org/10.5281/zenodo.13940234" ID-Zenodo-Dep="13940234" httpUri="https://zenodo.org/record/13940234/files/figure.png" pageId="4" pageNumber="108" startId="4.[130,145,933,952]" targetBox="[107,620,171,912]" targetPageId="4" targetType="figure">
<paragraph id="727836ABFFCE6A4A2FC6FC26FE19FC05" blockId="4.[107,619,932,1000]" pageId="4" pageNumber="108">
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. 2.—(A) Dorsal and (B) lateral view of the head of adult male
@ -368,7 +371,7 @@ and
;
<bibRefCitation id="16564B5AFFCE6A4A2B1EFC4BFD72FC13" author="CEI, J. M." firstAuthor="Cei" journalOrPublisher="Museo Regionale di Scienze Naturali, Torino, Monografie" pageId="4" pageNumber="108" pagination="1 - 527" part="4" refId="ref7632" refString=". 1986. Reptiles del centro, centro-oeste y sur de la Argentina. Museo Regionale di Scienze Naturali, Torino, Monografie 4: 1 - 527." title="Reptiles del centro, centro-oeste y sur de la Argentina" type="journal article" year="1986">Cei, 1986</bibRefCitation>
;
<figureCitation id="EAFC2A2EFFCE6A4A2D92FC65FCA4FC13" box="[726,798,998,1022]" captionStart="FIG" captionStartId="3.[698,713,895,914]" captionTargetBox="[677,1184,171,872]" captionTargetId="figure-12@3.[677,1184,171,875]" captionTargetPageId="3" captionText="FIG. 1.—Adult specimens of Liolaemus arambarensis from Arambaré, Rio Grande do Sul, Brazil (left female, UFRGS 2951, SVL = 55.85 mm; right male, UFRGS 2902, SVL = 59.84 mm = holotype)." pageId="4" pageNumber="108">Fig. 1</figureCitation>
<figureCitation id="EAFC2A2EFFCE6A4A2D92FC65FCA4FC13" box="[726,798,998,1022]" captionStart="FIG" captionStartId="3.[698,713,895,914]" captionTargetBox="[677,1184,171,872]" captionTargetId="figure-12@3.[677,1184,171,875]" captionTargetPageId="3" captionText="FIG. 1.—Adult specimens of Liolaemus arambarensis from Arambaré, Rio Grande do Sul, Brazil (left female, UFRGS 2951, SVL = 55.85 mm; right male, UFRGS 2902, SVL = 59.84 mm = holotype)." figureDoi="http://doi.org/10.5281/zenodo.13940232" httpUri="https://zenodo.org/record/13940232/files/figure.png" pageId="4" pageNumber="108">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
@ -1293,7 +1296,7 @@ and
<emphasis id="40B3EAB9FFC36A472E6EFC97FDB6FCC6" box="[298,524,787,811]" italics="true" pageId="9" pageNumber="113">Myrsine umbellata</emphasis>
</taxonomicName>
(
<figureCitation id="EAFC2A2EFFC36A472D5AFC90FDD2FCC6" box="[542,616,787,811]" captionStart="FIG" captionStartId="10.[130,145,858,877]" captionTargetBox="[107,1163,171,838]" captionTargetId="figure-14@10.[107,1163,171,838]" captionTargetPageId="10" captionText="FIG. 3.—Habitat at the type locality of Liolaemus arambarensis at Arambaré, RS, Brazil; note the sandy soil." pageId="9" pageNumber="113">Fig. 3</figureCitation>
<figureCitation id="EAFC2A2EFFC36A472D5AFC90FDD2FCC6" box="[542,616,787,811]" captionStart="FIG" captionStartId="10.[130,145,858,877]" captionTargetBox="[107,1163,171,838]" captionTargetId="figure-14@10.[107,1163,171,838]" captionTargetPageId="10" captionText="FIG. 3.—Habitat at the type locality of Liolaemus arambarensis at Arambaré, RS, Brazil; note the sandy soil." figureDoi="http://doi.org/10.5281/zenodo.13940236" httpUri="https://zenodo.org/record/13940236/files/figure.png" pageId="9" pageNumber="113">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
@ -1426,7 +1429,7 @@ values of Student̓s
<emphasis id="40B3EAB9FFC36A472D2DFCB2FE9EFC88" italics="true" pageId="9" pageNumber="113">L. arambarensis</emphasis>
</taxonomicName>
(
<figureCitation id="EAFC2A2EFFC36A472E7EFCCEFE32FC88" box="[314,392,845,869]" captionStart="FIG" captionStartId="11.[154,169,1007,1026]" captionTargetBox="[131,1187,174,987]" captionTargetId="figure-12@11.[131,1187,171,987]" captionTargetPageId="11" captionText="FIG. 4.—Karyotype and site of active NORs in Liolaemus arambarensis (UFRGS 2369, male) from Arambaré, Rio Grande do Sul, Brazil. 2N = 34 (12 M + 22 m). Scale = 10 µm." pageId="9" pageNumber="113">Fig. 4</figureCitation>
<figureCitation id="EAFC2A2EFFC36A472E7EFCCEFE32FC88" box="[314,392,845,869]" captionStart="FIG" captionStartId="11.[154,169,1007,1026]" captionTargetBox="[131,1187,174,987]" captionTargetId="figure-12@11.[131,1187,171,987]" captionTargetPageId="11" captionText="FIG. 4.—Karyotype and site of active NORs in Liolaemus arambarensis (UFRGS 2369, male) from Arambaré, Rio Grande do Sul, Brazil. 2N = 34 (12 M + 22 m). Scale = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.13940238" httpUri="https://zenodo.org/record/13940238/files/figure.png" pageId="9" pageNumber="113">Fig. 4</figureCitation>
) indicates a diploid number of 2N = 34. The karyotype has 12 metacentric macrochromosomes and 22 microchromosomes. The first pair of microchromosomes is distinctly larger than the others. Analysis of chromosomes in metaphase, stained with silver, indicated that NORs are located in the secondary constrictions of the long arm of Pair 2. The known diploid number for
<taxonomicName id="B5C74D28FFC36A472E89FBD3FF0AFB68" pageId="9" pageNumber="113">
<emphasis id="40B3EAB9FFC36A472E89FBD3FDF2FB85" box="[461,584,1104,1128]" italics="true" pageId="9" pageNumber="113">Liolaemus</emphasis>
@ -1574,7 +1577,7 @@ has a diet exclusively composed of arthropods, with no plant material (
).
</paragraph>
</subSubSection>
<caption id="26B86623FFC06A442FC6FCD9FF6EFC69" pageId="10" pageNumber="114" startId="10.[130,145,858,877]" targetBox="[107,1163,171,838]" targetPageId="10" targetType="figure">
<caption id="26B86623FFC06A442FC6FCD9FF6EFC69" ID-DOI="http://doi.org/10.5281/zenodo.13940236" ID-Zenodo-Dep="13940236" httpUri="https://zenodo.org/record/13940236/files/figure.png" pageId="10" pageNumber="114" startId="10.[130,145,858,877]" targetBox="[107,1163,171,838]" targetPageId="10" targetType="figure">
<paragraph id="727836ABFFC06A442FC6FCD9FF6EFC69" blockId="10.[107,1163,857,900]" pageId="10" pageNumber="114">
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. 3.—Habitat at the type locality of
@ -1618,7 +1621,7 @@ group, defined by
<bibRefCitation id="16564B5AFFC06A442DA4F9B3FC16F9A4" author="ETHERIDGE, R." box="[736,940,1584,1609]" editor="R. Estes &amp; G. Prigill" firstAuthor="Etheridge" journalOrPublisher="Stanford University Press, Stanford, California, U. S. A." pageId="10" pageNumber="114" pagination="283 - 367" refId="ref7845" refString=". 2000. A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs 14: 293 - 352." title="A review of lizards of the Liolaemus wiegmannii group (Squamata, Iguania, Tropiduridae), and a history of morphological change in the sand-dwelling species. Herpetological Monographs 14: 293 - 352. logeny of Iguanidae." type="book chapter" volumeTitle="Phylogenetic Relationships of the Lizard Families - Essays Commemorating Charles L. Camp" year="2000">Etheridge, 2000</bibRefCitation>
): Lorilabial scales small, more than seven between the posterior canthals (Character 4: State 1); supralabials narrow (Character 6: State 1); no contact between postmentals and the infralabials (Character 11: State 1); snout profile flat (Character 32: State 1); and diurnal escape behavior includes rapid burial under sand (Character 36: State 1).
</paragraph>
<caption id="26B86623FFC16A452FDEFC6CFC38FBF7" pageId="11" pageNumber="115" startId="11.[154,169,1007,1026]" targetBox="[131,1187,174,987]" targetPageId="11" targetType="figure">
<caption id="26B86623FFC16A452FDEFC6CFC38FBF7" ID-DOI="http://doi.org/10.5281/zenodo.13940238" ID-Zenodo-Dep="13940238" httpUri="https://zenodo.org/record/13940238/files/figure.png" pageId="11" pageNumber="115" startId="11.[154,169,1007,1026]" targetBox="[131,1187,174,987]" targetPageId="11" targetType="figure">
<paragraph id="727836ABFFC16A452FDEFC6CFC38FBF7" blockId="11.[131,1187,1007,1050]" pageId="11" pageNumber="115">
<smallCapsWord id="749EA077FFC16A452FDEFC6CFF04FBEC" baselines="1021,1021" box="[154,190,1007,1026]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Fig" pageId="11" pageNumber="115">FIG</smallCapsWord>
. 4.—Karyotype and site of active NORs in
@ -1676,7 +1679,7 @@ however, appears referable to Branch 23 by sharing the following synapomorphies:
</emphasis>
there are more than seven present; infratarsal scales surface smooth (Character 17: State 2); infratarsal scales sub-imbricate (Character 19: State 2); shape infratarsal scales of obovate (Character 20: State 1).
</paragraph>
<caption id="26B86623FFC66A422FC6FDC1FEE9FD59" pageId="12" pageNumber="116" startId="12.[130,145,578,597]" targetBox="[107,618,173,556]" targetPageId="12" targetType="figure">
<caption id="26B86623FFC66A422FC6FDC1FEE9FD59" ID-DOI="http://doi.org/10.5281/zenodo.13940240" ID-Zenodo-Dep="13940240" httpUri="https://zenodo.org/record/13940240/files/figure.png" pageId="12" pageNumber="116" startId="12.[130,145,578,597]" targetBox="[107,618,173,556]" targetPageId="12" targetType="figure">
<paragraph id="727836ABFFC66A422FC6FDC1FEE9FD59" blockId="12.[107,619,577,692]" pageId="12" pageNumber="116">
<smallCapsWord id="749EA077FFC66A422FC6FDC1FF1CFDBE" baselines="592,591" box="[130,166,578,597]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Fig" pageId="12" pageNumber="116">FIG</smallCapsWord>
. 5.—Phylogenetic hypothesis proposed by
@ -1735,7 +1738,7 @@ and
.
</emphasis>
Among the synapomorphies described for Branch 24 (
<figureCitation id="EAFC2A2EFFC66A422FDDF990FF5BF9C6" box="[153,225,1555,1579]" captionStart="FIG" captionStartId="12.[130,145,578,597]" captionTargetBox="[107,618,173,556]" captionTargetId="figure-13@12.[107,619,171,558]" captionTargetPageId="12" captionText="FIG. 5.—Phylogenetic hypothesis proposed by Etheridge (2000) for the ̒̒aeiegmannii̓̓ group, which suggests Liolaemus arambarensis as sister group for Liolaemus Branch 24; future studies will be required to confirm this hypothesis." pageId="12" pageNumber="116">Fig. 5</figureCitation>
<figureCitation id="EAFC2A2EFFC66A422FDDF990FF5BF9C6" box="[153,225,1555,1579]" captionStart="FIG" captionStartId="12.[130,145,578,597]" captionTargetBox="[107,618,173,556]" captionTargetId="figure-13@12.[107,619,171,558]" captionTargetPageId="12" captionText="FIG. 5.—Phylogenetic hypothesis proposed by Etheridge (2000) for the ̒̒aeiegmannii̓̓ group, which suggests Liolaemus arambarensis as sister group for Liolaemus Branch 24; future studies will be required to confirm this hypothesis." figureDoi="http://doi.org/10.5281/zenodo.13940240" httpUri="https://zenodo.org/record/13940240/files/figure.png" pageId="12" pageNumber="116">Fig. 5</figureCitation>
),
<taxonomicName id="B5C74D28FFC66A422FBCF997FE00F9C6" authorityName="Verrastro &amp; Veronese &amp; Bujes &amp; Filh" authorityYear="2003" box="[248,442,1555,1579]" class="Squamata" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" pageId="12" pageNumber="116" phylum="Chordata" rank="species" species="arambarensis">
<emphasis id="40B3EAB9FFC66A422FBCF997FE00F9C6" box="[248,442,1555,1579]" italics="true" pageId="12" pageNumber="116">L. arambarensis</emphasis>