<documentid="552CEAD9BD510E213FEE3F564B4BEF39"ID-DOI="10.25674/so95iss2id315"ID-ISSN="2509-9523"ID-Zenodo-Dep="13193924"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1722631509206"checkinUser="felipe"docAuthor="Seifert, Bernhard"docDate="2023"docId="03FAE936FFC5FA7E34E0F8E5FD88EF47"docLanguage="en"docName="SoilOrg.95.2.129-142.pdf"docOrigin="Soil Organisms 95 (2)"docSource="http://dx.doi.org/10.25674/so95iss2id315"docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article"docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2"docStyleName="SoilOrg.2018-.journal_article"docStyleVersion="2"docTitle="Formicoxenus sibiricus"docType="treatment"docVersion="2"lastPageNumber="137"masterDocId="FFC3914EFFC3FA763477FFD8FFBAE831"masterDocTitle="Two new species of Formicoxenus Mayr 1855 and Leptothorax Mayr 1855 from Tibet (Hymenoptera: Formicidae)"masterLastPageNumber="142"masterPageNumber="129"pageNumber="135"updateTime="1722679370904"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:titleid="E4A5139C05D5F8CA455892D5CE1488A5">Two new species of Formicoxenus Mayr 1855 and Leptothorax Mayr 1855 from Tibet (Hymenoptera: Formicidae)</mods:title>
<bibRefCitationid="EFC225D1FFC5FA7035DAF8E4FD92EF65"author="Forel, A."box="[429,552,1852,1876]"pageId="6"pageNumber="135"pagination="303 - 310"refId="ref7519"refString="Forel, A. (1899): Trois notices myrmecologiques. - Annales de la Societe Entomologique de Belgique 43: 303 - 310."type="journal article"year="1899">Forel 1899</bibRefCitation>
<bibRefCitationid="EFC225D1FFC5FA7035D4F8A6FD98EFA7"author="Forel, A."box="[419,546,1918,1942]"pageId="6"pageNumber="135"pagination="303 - 310"refId="ref7519"refString="Forel, A. (1899): Trois notices myrmecologiques. - Annales de la Societe Entomologique de Belgique 43: 303 - 310."type="journal article"year="1899">Forel 1899</bibRefCitation>
<bibRefCitationid="EFC225D1FFC5FA70375EFAD6FC15ED17"author="Forel, A."box="[809,943,1294,1318]"pageId="6"pageNumber="135"pagination="303 - 310"refId="ref7519"refString="Forel, A. (1899): Trois notices myrmecologiques. - Annales de la Societe Entomologique de Belgique 43: 303 - 310."type="journal article"year="1899">Forel (1899)</bibRefCitation>
described this taxon from East Siberia without giving a more precise type locality: ‘Sibérie orientale (recu de M. Nassonow).’ Investigated were the images of the
<bibRefCitationid="EFC225D1FFC5FA703093F9CDFC33EE7C"author="Francoeur, A. & Loiselle, R. & A. Buschinger"pageId="6"pageNumber="135"pagination="343 - 403"refId="ref7545"refString="Francoeur, A.; Loiselle, R. & A. Buschinger (1985): Biosystematique de la tribu Leptothoracini (Formicidae, Hymenoptera). 1. Le genre Formicoxenus dans la region Holarctique. - Le Naturaliste Canadien 112: 343 - 403."type="journal article"year="1985">Francoeur et al.(1985)</bibRefCitation>
. We have a very good idea of the morphological characters of this species as the images of the type correspond very well to the characters of my own sample from East Kazakhstan on the basis of which the species is re-described below.
<bibRefCitationid="EFC225D1FFC5FA703026F923FB46EF22"author="Dlussky, G. M."box="[1105,1276,1787,1811]"pageId="6"pageNumber="135"pagination="190 - 194"refId="ref7456"refString="Dlussky, G. M. (1963): Dva novykh vida murav'ev (Hymenoptera, Formicidae) iz vostochnovo Zabaikal'ya. - Entomologicheskoye Obozrenie 42 (1): 190 - 194."type="journal article"year="1963">Dlussky 1963</bibRefCitation>
<bibRefCitationid="EFC225D1FFC5FA70375EF8E5FC71EF64"author="Dlussky, G. M."box="[809,971,1853,1877]"pageId="6"pageNumber="135"pagination="190 - 194"refId="ref7456"refString="Dlussky, G. M. (1963): Dva novykh vida murav'ev (Hymenoptera, Formicidae) iz vostochnovo Zabaikal'ya. - Entomologicheskoye Obozrenie 42 (1): 190 - 194."type="journal article"year="1963">Dlussky (1963)</bibRefCitation>
<taxonomicNameid="4C5323A3FFC4FA7134CAFDC1FD14EA03"authority="Zhou et Chen 2011"authorityName="Zhou et Chen"authorityYear="2011"box="[189,686,537,562]"class="Insecta"family="Formicidae"genus="Leptothorax"kingdom="Animalia"order="Hymenoptera"pageId="7"pageNumber="136"phylum="Arthropoda"rank="species"species="zhengi">
<bibRefCitationid="EFC225D1FFC4FA7135C1FDC1FD14EA03"author="Zhou, S. Y. & Y. I. Peng & Z. L. Chen & X. G. Zhou & L. T. Liang"box="[438,686,537,562]"pageId="7"pageNumber="136"pagination="592 - 594"refId="ref7808"refString="Zhou, S. Y., Y. I. Peng, Z. L. Chen, X. G. Zhou & L. T. Liang (2011): A new species and a new record species of the genus Leptothorax Mayr (Hymenoptera, Formicidae) from China. - Acta Zootaxonomica Sinica 36: 592 - 594."type="journal article"year="2011">Zhou et Chen 2011</bibRefCitation>
<bibRefCitationid="EFC225D1FFC4FA7134A3FD83FE2DEA42"author="Zhou, S. Y. & Y. I. Peng & Z. L. Chen & X. G. Zhou & L. T. Liang"box="[212,407,603,627]"pageId="7"pageNumber="136"pagination="592 - 594"refId="ref7808"refString="Zhou, S. Y., Y. I. Peng, Z. L. Chen, X. G. Zhou & L. T. Liang (2011): A new species and a new record species of the genus Leptothorax Mayr (Hymenoptera, Formicidae) from China. - Acta Zootaxonomica Sinica 36: 592 - 594."type="journal article"year="2011">Zhou et al. (2011)</bibRefCitation>
<bibRefCitationid="EFC225D1FFC4FA7136D3FC99FEAFEB4B"author="Zhou, S. Y. & Y. I. Peng & Z. L. Chen & X. G. Zhou & L. T. Liang"pageId="7"pageNumber="136"pagination="592 - 594"refId="ref7808"refString="Zhou, S. Y., Y. I. Peng, Z. L. Chen, X. G. Zhou & L. T. Liang (2011): A new species and a new record species of the genus Leptothorax Mayr (Hymenoptera, Formicidae) from China. - Acta Zootaxonomica Sinica 36: 592 - 594."type="journal article"year="2011">Zhou et al. (2011)</bibRefCitation>
CL = 0.790, CW 0.581 (adapted to the measuring mode used here), SL 0.460, MW 0.385, ML 0.845. This translates into CS 0.686, CL/CW 1.360, SL/CS 0.671, MW/CS 0.560, ML/CS 1.230 (compare with
<tableCitationid="C6D16D9BFFC4FA71313EFECAFA34E91B"box="[1353,1422,274,298]"captionStart="Table 1"captionStartId="5.[189,242,246,267]"captionText="Table 1. Measurements of workers of the genera Leptothorax and Formicoxenus. Data are given in the sequence arithmetic mean ± standard deviation [minimum, maximum]."pageId="7"pageNumber="136">Tab. 1</tableCitation>
). Even if considering some measuring inaccuracy, these measurements as well as images given in the original description indicate a junior synonymy with
. This refers in particular to the much elongated head, the small eyes, the mesosoma and waist profile with a very broad subpetiolar process and an acute, well-developed spine of postpetiolar sternite, the characteristic sculpture on head, mesosoma and waist, and the short blunt setae on whole body. Hence there is in no character a disagreement with the re-description of
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the
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(for standard deviation, minimum and maximum values see
<tableCitationid="C6D16D9BFFCBFA7E35F1FDE2FE72EA63"box="[390,456,570,594]"captionStart="Table 1"captionStartId="5.[189,242,246,267]"captionText="Table 1. Measurements of workers of the genera Leptothorax and Formicoxenus. Data are given in the sequence arithmetic mean ± standard deviation [minimum, maximum]."pageId="8"pageNumber="137">Tab. 1</tableCitation>
). Rather small size,
<specimenCodeid="DBF5F05BFFCBFA7E36DBFDE2FD41EA63"box="[684,763,570,594]"collectionCode="CS"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS 676</specimenCode>
µm. Head much elongated, CL/CW 1.333. Genae in dorsal view parallel or even slightly diverging frontad. Anterior margin of clypeus in dorsal view semicircular. Occipital margin straight to weakly concave. Frons very broad (
<collectionCodeid="ED42C0E5FFCBFA7E3514FD07FE3EEAC6"box="[355,388,735,759]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
0.478). Scape comparably long, SL/
<collectionCodeid="ED42C0E5FFCBFA7E34C8FCD8FF65EB29"box="[191,223,768,792]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
<collectionCodeid="ED42C0E5FFCBFA7E363CFCD8FDD1EB29"box="[587,619,768,792]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
0.173, with notable microsetae. Dorsal profile of promesonotum largely linear, propodeum convex, metanotal depression moderately deep (MGr/
<specimenCodeid="DBF5F05BFFCBFA7E35E8FCBAFE4DEB4B"box="[415,503,866,890]"collectionCode="CS"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS 2.69</specimenCode>
%). Spines rather short and acute (SP/
<collectionCodeid="ED42C0E5FFCBFA7E3541FC5BFEECEBAA"box="[310,342,899,923]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
<collectionCodeid="ED42C0E5FFCBFA7E349CFC7CFEB1EB8D"box="[235,267,932,956]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
0.294). Petiole with strong anterolateral corners, in profile without peduncle and with a broad subpetiolar lobe. Postpetiole narrower than in other species (PpW/
<collectionCodeid="ED42C0E5FFCBFA7E3537FBDFFEE5EC2E"box="[320,351,1031,1055]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
0.344), its sternite with a well-developed spine. All surfaces of head, mesosoma, waist and appendages matt due to a reticulate microsculpture. Dorsal vertex strongly longitudinally carinulate-rugulose. Gaster glabrous and with very dilute pubescence. Head, mesosoma, waist, scape, femora and tibiae with very short (PnHL/
<collectionCodeid="ED42C0E5FFCBFA7E3585FB14FDA8ECD5"box="[498,530,1228,1252]"country="France"name="Musee des Dinosaures d'Esperaza (Aude)"pageId="8"pageNumber="137">CS</collectionCode>
0.062) setae, which widen from base to apex, are apically fringed (
<bibRefCitationid="EFC225D1FFCBFA7E35C5F9AFFDF7EEBE"author="Dlussky, G. M."box="[434,589,1655,1679]"pageId="8"pageNumber="137"pagination="1026 - 104"refId="ref7491"refString="Dlussky, G. M. (1964): Murav'i podroda Coptoformica roda Formica. - Zoologichesky Zhurnal' 43 (7): 1026 - 104."type="journal article"year="1964">Dlussky 1964</bibRefCitation>
</taxonomicName>
. The latter host had originally been determined by
<bibRefCitationid="EFC225D1FFCBFA7E366CF940FD7AEE81"author="Dlussky, G. M."box="[539,704,1688,1712]"pageId="8"pageNumber="137"pagination="190 - 194"refId="ref7456"refString="Dlussky, G. M. (1963): Dva novykh vida murav'ev (Hymenoptera, Formicidae) iz vostochnovo Zabaikal'ya. - Entomologicheskoye Obozrenie 42 (1): 190 - 194."type="journal article"year="1963">Dlussky (1963)</bibRefCitation>
<bibRefCitationid="EFC225D1FFCBFA7E355FF885FD9CEF47"author="Seifert, B. & R. Schultz"box="[296,550,1885,1910]"pageId="8"pageNumber="137"pagination="177 - 220"refId="ref7691"refString="Seifert, B. & R. Schultz (2021): A taxonomic revision of the Palaearctic ant subgenus Coptoformica Muller, 1923 (Hymenoptera, Formicidae). - Contributions to Entomology 71 (2): 177 - 220 [https: // doi. org / 10.21248 / contrib. entomol. 71.2.177 - 220]"type="journal article"year="2021">Seifert & Schultz 2021</bibRefCitation>
