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<document id="B5FB18C8C14DD3707EF84A05F9839D1D" ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7746966DB33FA361E13FA67" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Bettongia Gray 1837" docType="treatment" docVersion="8" lastPageNumber="335" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="335" updateTime="1699350301078" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5FDC1C3B72D8144EA504E6FBFE49C2F7">Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:namePart id="9EB35A6FF0C6654C234A17BD4453E7F5">Beck, Robin M. D.</mods:namePart>
<mods:affiliation id="18F8AB8F10D0A242F0DEE04C05872005">School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="76AC7DE5B04B4EDA810E00DD0770A5F2">Voss, Robert S.</mods:namePart>
<mods:affiliation id="1D14A0420B7CB705B42F9460BD310A5C">Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="CC12FB3B59AEBA39E7CD9F31ADC3364D">Jansa, Sharon A.</mods:namePart>
<mods:affiliation id="F7542A50D1DB1CD600A7FE35A75BFFC1">Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<paragraph id="8BF96C4BF7746966DB33FA361BD2FA77" blockId="335.[312,432,1432,1456]" box="[312,432,1432,1456]" pageId="335" pageNumber="335">
<heading id="D0B1DB27F7746966DB33FA361BD2FA77" box="[312,432,1432,1456]" centered="true" fontSize="10" level="2" pageId="335" pageNumber="335" reason="4">
<emphasis id="B932B059F7746966DB33FA361BD2FA77" bold="true" box="[312,432,1432,1456]" pageId="335" pageNumber="335">
?
<taxonomicName id="4C4617C8F7746966DB48FA361BD2FA77" box="[323,432,1432,1456]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B932B059F7746966DB48FA361BD2FA77" bold="true" box="[323,432,1432,1456]" italics="true" pageId="335" pageNumber="335">Bettongia</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BF96C4BF7746966DA87FA641843FA25" blockId="335.[108,637,1482,1738]" box="[140,545,1482,1506]" pageId="335" pageNumber="335">
SPECIES SCORED:?
<taxonomicName id="4C4617C8F7746966DB53FA651BDCFA25" ID-CoL="39X8" authorityName="Gray" authorityYear="1837" box="[344,446,1483,1506]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis id="B932B059F7746966DB53FA651BDCFA25" box="[344,446,1483,1506]" italics="true" pageId="335" pageNumber="335">Bettongia</emphasis>
</taxonomicName>
<emphasis id="B932B059F7746966DBD9FA651879FA25" box="[466,539,1483,1506]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
.
</paragraph>
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<subSubSection id="C35C3FC0F7746966DA87FA45186EF9A1" pageId="335" pageNumber="335" type="materials_examined">
<paragraph id="8BF96C4BF7746966DA87FA45186EF9A1" blockId="335.[108,637,1482,1738]" pageId="335" pageNumber="335">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode id="ED57F48EF7746966D83DFA451AC5F9E3" pageId="335" pageNumber="335">SPECIMENS</collectionCode>
: Henks Hollow and Two Trees sites (Riversleigh Faunal Zone C), Riversleigh World Heritage Area,
<collectingRegion id="4982A2A9F7746966DB19F9E01BF4F9A1" box="[274,406,1614,1638]" country="Australia" name="Queensland" pageId="335" pageNumber="335">Queensland</collectingRegion>
,
<collectingCountry id="F3512CDBF7746966DBA9F9E0186AF9A1" box="[418,520,1614,1638]" name="Australia" pageId="335" pageNumber="335">Australia</collectingCountry>
.
</paragraph>
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<subSubSection id="C35C3FC0F7746966DA87F9DE1ECFFEB9" pageId="335" pageNumber="335" type="description">
<paragraph id="8BF96C4BF7746966DA87F9DE1922FE9A" blockId="335.[108,637,1482,1738]" lastBlockId="335.[684,1214,226,1440]" pageId="335" pageNumber="335">AGE OF SCORED SPECIMENS: Riversleigh Faunal Zone C is interpreted to be middle Miocene based on biostratigraphy (see above). In the absence of radiometric dates, we have assumed the entire span of the middle Miocene (Langhian to Serravallian; Cohen et al., 2013 [updated]) for this terminal.</paragraph>
<paragraph id="8BF96C4BF7746966D8C7FEC81ECFFEB9" blockId="335.[684,1214,226,1440]" box="[716,1197,358,382]" pageId="335" pageNumber="335">
ASSIGNED AGE RANGE:
<geoCoordinate id="EE720A8CF7746966D9D9FEC81E78FEB9" box="[978,1050,358,382]" degrees="15.970" direction="north" orientation="latitude" pageId="335" pageNumber="335" precision="55" value="15.97">15.970</geoCoordinate>
<geoCoordinate id="EE720A8CF7746966DE12FEC81E0DFEB9" box="[1049,1135,358,382]" degrees="11.630" direction="west" orientation="longitude" pageId="335" pageNumber="335" precision="55" value="-11.63">11.630</geoCoordinate>
Mya.
</paragraph>
</subSubSection>
<subSubSection id="C35C3FC0F7746966D8C7FE291E13FA67" pageId="335" pageNumber="335" type="discussion">
<paragraph id="8BF96C4BF7746966D8C7FE291E13FA67" blockId="335.[684,1214,226,1440]" pageId="335" pageNumber="335">
REMARKS: Flannery and Archer (1987b) referred this fossil taxon, which is represented by a partial skull and associated mandibles plus an additional left mandibular fragment, to the Recent potoroid genus
<taxonomicName id="4C4617C8F7746966D9B3FDA21E42FDE4" authorityName="Gray" authorityYear="1837" box="[952,1056,524,547]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis id="B932B059F7746966D9B3FDA21E42FDE4" box="[952,1056,524,547]" italics="true" pageId="335" pageNumber="335">Bettongia</emphasis>
</taxonomicName>
, but we consider it only questionably a member of this genus based on available evidence, which we summarize briefly here. Flannery and Archer (1987b) noted particular similarities between?
<emphasis id="B932B059F7746966DE41FD3E1E03FD60" box="[1098,1121,656,679]" italics="true" pageId="335" pageNumber="335">B.</emphasis>
<emphasis id="B932B059F7746966DE7FFD3E1EDEFD60" box="[1140,1212,656,679]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
and the living
<taxonomicName id="4C4617C8F7746966D94DFD1F19CFFD0F" baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1824" box="[838,941,688,712]" class="Mammalia" family="Potoroidae" genus="Bettongia" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="lesueur">
<emphasis id="B932B059F7746966D94DFD1F19CFFD0F" box="[838,941,688,712]" italics="true" pageId="335" pageNumber="335">B. lesueur</emphasis>
</taxonomicName>
. The phylogenetic analysis of Kear et al. (2007) recovered a sister-group relationship between?
<emphasis id="B932B059F7746966D9A9FD5D19DAFCCD" box="[930,952,755,778]" italics="true" pageId="335" pageNumber="335">B.</emphasis>
<emphasis id="B932B059F7746966D9C6FD5D1E74FCCD" box="[973,1046,755,778]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
and the extant potoroid
<taxonomicName id="4C4617C8F7746966D919FCBA198AFCEB" baseAuthorityName="Kerr" baseAuthorityYear="1792" box="[786,1000,788,812]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="tridactylus">
<emphasis id="B932B059F7746966D919FCBA198AFCEB" box="[786,1000,788,812]" italics="true" pageId="335" pageNumber="335">Potorous tridactylus</emphasis>
</taxonomicName>
to the exclusion of other macropodiforms in their taxon sample, supporting the potoroid affinities of?
<emphasis id="B932B059F7746966DE48FCF91E38FCA9" box="[1091,1114,855,878]" italics="true" pageId="335" pageNumber="335">B.</emphasis>
<emphasis id="B932B059F7746966DE66FCF91ED4FCA9" box="[1133,1206,855,878]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
. However,
<taxonomicName id="4C4617C8F7746966D913FCD619C3FC48" baseAuthorityName="Kerr" baseAuthorityYear="1792" box="[792,929,887,911]" class="Mammalia" family="Potoroidae" genus="Potorous" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="tridactylus">
<emphasis id="B932B059F7746966D913FCD619C3FC48" box="[792,929,887,911]" italics="true" pageId="335" pageNumber="335">P. tridactylus</emphasis>
</taxonomicName>
was the sole Recent potoroid included by Kear et al. (2007), so the precise position of?
<emphasis id="B932B059F7746966D93CFC14192CFC16" box="[823,846,954,977]" italics="true" pageId="335" pageNumber="335">B.</emphasis>
<emphasis id="B932B059F7746966D968FC1419CEFC16" box="[867,940,954,977]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
within
<taxonomicName id="4C4617C8F7746966DE0EFC171E19FC16" authorityName="Gray" authorityYear="1821" box="[1029,1147,953,977]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
(and, particularly, its relationship to recent
<taxonomicName id="4C4617C8F7746966DE5EFC751899FBD4" pageId="335" pageNumber="335">
<emphasis id="B932B059F7746966DE5EFC751EDEFC35" box="[1109,1212,987,1010]" italics="true" pageId="335" pageNumber="335">Bettongia</emphasis>
species
</taxonomicName>
) remained uncertain in this analysis. The phylogenetic analysis of Travouillon et al. (2016), meanwhile, recovered?
<emphasis id="B932B059F7746966D9BAFB9019AAFB92" box="[945,968,1086,1109]" italics="true" pageId="335" pageNumber="335">B.</emphasis>
<emphasis id="B932B059F7746966D9D6FB901E44FB92" box="[989,1062,1086,1109]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
within
<taxonomicName id="4C4617C8F7746966DE8BFB901893FBB1" authorityName="Gray" authorityYear="1821" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
but outside the crown clade. Some other published analyses have failed to place?
<emphasis id="B932B059F7746966DEADFB2E1EDEFB50" box="[1190,1212,1152,1175]" italics="true" pageId="335" pageNumber="335">B.</emphasis>
<emphasis id="B932B059F7746966D8B1FB0F1966FB7F" box="[698,772,1185,1208]" italics="true" pageId="335" pageNumber="335">moyesi</emphasis>
within
<taxonomicName id="4C4617C8F7746966D968FB0E19B9FB7F" authorityName="Gray" authorityYear="1821" box="[867,987,1184,1208]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
(Travouillon et al., 2015a: fig. 5A; Butler et al., 2016, 2018; den Boer and Kear, 2018: supplemental data), or have found its relationship relative to
<taxonomicName id="4C4617C8F7746966DE16FAAA1EDEFADB" baseAuthorityName="sensu Kear and Cooke" baseAuthorityYear="2001" box="[1053,1212,1284,1308]" class="Mammalia" family="Macropodidae" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="family">Macropodidae</taxonomicName>
and
<taxonomicName id="4C4617C8F7746966D8D0FA8B1933FAFA" authorityName="Gray" authorityYear="1821" box="[731,849,1317,1341]" class="Mammalia" family="Potoroidae" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="family">Potoroidae</taxonomicName>
to be largely unresolved (Bates et al., 2014; Black et al., 2014c; Travouillon et al., 2014b, 2015a: fig. 5B; Cooke et al., 2015; den Boer and Kear, 2018: supplemental data).
</paragraph>
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