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164 lines
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<document ID-DOI="http://doi.org/10.5281/zenodo.6103256" ID-GBIF-Dataset="f0d443f2-b8b9-46fa-a1aa-2d0f0d3c8748" ID-GBIF-Taxon="114923358" ID-ISSN="1175-5326" ID-Zenodo-Dep="6103256" ID-ZooBank="CEAB4AA5-C4F8-437E-A2F4-D82F383579C1" checkinTime="1447288886152" checkinUser="donat" docAuthor="Taylor, Robert W." docDate="2015" docId="03B3878D9B01FFF0FF534B7A9E79FBAF" docLanguage="en" docName="29627.pdf.imf" docOrigin="Zootaxa 3947 (2)" docTitle="Heteroponera pendergrasti Taylor, 2015, sp. n." docType="treatment" docVersion="5" lastPageId="28" lastPageNumber="179" masterDocId="FF8AFFF59B18FFECFFC44D279F4AF867" masterDocTitle="Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus" masterLastPageNumber="180" masterPageNumber="151" pageId="25" pageNumber="176" updateTime="1645000973839" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Taylor, Robert W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>3947</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>151</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3947.2.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="ZooBank">CEAB4AA5-C4F8-437E-A2F4-D82F383579C1</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6103256" ID-GBIF-Taxon="114923358" ID-Zenodo-Dep="6103256" LSID="urn:lsid:plazi:treatment:03B3878D9B01FFF0FF534B7A9E79FBAF" httpUri="http://treatment.plazi.org/id/03B3878D9B01FFF0FF534B7A9E79FBAF" lastPageId="28" lastPageNumber="179" pageId="25" pageNumber="176">
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<subSubSection pageId="25" pageNumber="176" type="nomenclature">
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<paragraph box="[151,594,1629,1655]" pageId="25" pageNumber="176">
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<emphasis bold="true" box="[151,594,1629,1655]" pageId="25" pageNumber="176">
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<heading bold="true" box="[151,594,1629,1655]" fontSize="11" level="1" pageId="25" pageNumber="176" reason="1">
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10.
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<emphasis bold="true" box="[195,521,1629,1655]" italics="true" pageId="25" pageNumber="176">
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<taxonomicName box="[195,521,1629,1655]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="176" phylum="Arthropoda" rank="species" species="pendergrasti" status="sp. n.">Heteroponera pendergrasti</taxonomicName>
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</emphasis>
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<taxonomicNameLabel box="[528,594,1629,1655]" pageId="25" pageNumber="176" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</emphasis>
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</paragraph>
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<paragraph box="[151,302,1663,1688]" pageId="25" pageNumber="176">
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(
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<figureCitation box="[159,290,1663,1688]" captionStart="FIGURES 50 – 53" captionStartId="27.[151,263,1954,1976]" captionTargetBox="[198,1387,193,1932]" captionTargetId="figure@27.[198,1388,193,1932]" captionTargetPageId="27" captionText="FIGURES 50 – 53. Heteroponera pendergrasti sp. n., holotype worker, standard views (HW = 1.23 mm, PW = 0.86 mm, WL = 1.54 mm, GW = 0.99 mm)." pageId="25" pageNumber="176">Figs 50–53</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="176" type="materials_examined">
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<paragraph pageId="25" pageNumber="176">
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<emphasis bold="true" box="[151,481,1735,1760]" pageId="25" pageNumber="176">Type locality. AUSTRALIA</emphasis>
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: QUEENSLAND:
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<emphasis box="[699,884,1735,1760]" italics="true" pageId="25" pageNumber="176">NE Lamb Range</emphasis>
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(
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<emphasis box="[900,1111,1735,1760]" italics="true" pageId="25" pageNumber="176">11 km SE Mareeba</emphasis>
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), -17 0 6, 145 34.
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<emphasis bold="true" box="[199,394,1772,1796]" pageId="25" pageNumber="176">Type deposition.</emphasis>
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Holotype ANIC. Paratypes in ANIC, QMBA.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="179" pageId="25" pageNumber="176" type="distribution">
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<paragraph pageId="25" pageNumber="176">
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<emphasis bold="true" box="[199,837,1807,1832]" pageId="25" pageNumber="176">Distribution, material examined. EA7 (Lamb Range):</emphasis>
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<emphasis box="[844,960,1808,1832]" italics="true" pageId="25" pageNumber="176">Mt Tiptree</emphasis>
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, -17 0 3, 145 37, (B. Halliday, 13.vi.1984), worker—900 m (RWT & JDF, 29.vi.1971), worker paratype -17 0 3, 145 38, 730 m (RWT & JDF, 29.vi.1971), worker paratype.
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<emphasis box="[354,620,1879,1904]" italics="true" pageId="25" pageNumber="176">Mt Edith, Lamb Range</emphasis>
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, -17 0 6, 145 37, QMBA berlesate 465, 1140 m (GBM, DKY & GIT, 11.x.1982) holotype, worker paratype.
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<emphasis box="[588,774,1916,1940]" italics="true" pageId="25" pageNumber="176">NE Lamb Range</emphasis>
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(
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<emphasis box="[791,1032,1916,1940]" italics="true" pageId="25" pageNumber="176">11 km SE of Mareeba</emphasis>
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), -17 0 6, 145 34, QMBA berlesates 804, 822, rainforest moss, 1200 m 11.xi.1988 (GBM, GIT), holotype, 4 worker paratypes.
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<emphasis box="[1177,1383,1952,1976]" italics="true" pageId="25" pageNumber="176">Upper Isley Creek</emphasis>
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, -17 0 3, 145 41, trees & logs, pyrethrum, 750 m (GBM & Janetzki, 29.xi.1993).
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</paragraph>
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<caption pageId="26" pageNumber="177" targetBox="[200,1386,193,1931]" targetPageId="26">
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<paragraph pageId="26" pageNumber="177">
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<emphasis bold="true" box="[151,342,1952,1974]" pageId="26" pageNumber="177">FIGURES 46–49.</emphasis>
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<taxonomicName box="[350,597,1953,1974]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="ecarinata" status="sp. n.">
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<emphasis box="[350,597,1953,1974]" italics="true" pageId="26" pageNumber="177">Heteroponera ecarinata</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[606,665,1953,1974]" pageId="26" pageNumber="177">
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<taxonomicNameLabel box="[606,665,1953,1974]" pageId="26" pageNumber="177" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, paratype worker, standard views (HW = 1.23mm, PW = 0.91mm, WL = 1.55mm, GW = 1.04mm).
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</paragraph>
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</caption>
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<caption pageId="27" pageNumber="178" targetBox="[198,1387,193,1932]" targetPageId="27">
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<paragraph pageId="27" pageNumber="178">
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<emphasis bold="true" box="[151,340,1954,1976]" pageId="27" pageNumber="178">FIGURES 50–53.</emphasis>
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<taxonomicName box="[347,623,1954,1976]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="178" phylum="Arthropoda" rank="species" species="pendergrasti" status="sp. n.">
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<emphasis box="[347,623,1954,1976]" italics="true" pageId="27" pageNumber="178">Heteroponera pendergrasti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[629,687,1955,1976]" pageId="27" pageNumber="178" rank="species">
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<emphasis bold="true" box="[629,687,1955,1976]" pageId="27" pageNumber="178">sp. n.</emphasis>
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</taxonomicNameLabel>
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, holotype worker, standard views (HW = 1.23mm, PW = 0.86mm, WL = 1.54mm, GW = 0.99mm).
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</paragraph>
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</caption>
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<paragraph pageId="28" pageNumber="179">
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<emphasis bold="true" box="[199,424,151,176]" pageId="28" pageNumber="179">Worker diagnosis.</emphasis>
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General features, conformation, color and major sculpturation as illustrated and in appropriate key couplets above. Characterized in the
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<taxonomicName box="[762,905,189,212]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="ecarinata">
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<emphasis box="[762,905,189,212]" italics="true" pageId="28" pageNumber="179">H. ecarinata</emphasis>
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</taxonomicName>
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species complex by heavy sculpturation, dark blackish color, with dark brown legs and antennae, and distinguished from the similar
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<emphasis box="[1126,1269,224,248]" italics="true" pageId="28" pageNumber="179">
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<taxonomicName box="[1126,1269,224,248]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="ecarinata">H. ecarinata</taxonomicName>
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</emphasis>
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by differences in sculpturation (particularly of the frons and second gastral tergite), coloration and structure of the petiolar node. Fore-coxae and posterior surface of node superficially microsculptured; propodeal declivity shining, with weak, superficial dorsally-directed radial striae; its sides distinctly sculpturally bordered, the posterolateral edge in profile micro-serrate as a result of minutely extended cuticular points along the sculptural border. Sculpturation of second gastral tergite more distinctly longitudinally deployed than in
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<emphasis box="[868,1013,405,428]" italics="true" pageId="28" pageNumber="179">
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<taxonomicName box="[868,1013,405,428]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="ecarinata">H. ecarinata</taxonomicName>
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</emphasis>
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(refer Figs). Dimensions: (holotype; smallest available specimen, largest available specimen (both Mt): TL (ca): 5.1, 4.8, 5.1; HW: 1.23, 1.18, 1.23; HL: 1.19, 1.17, 1.18; CI: 103, 100, 104; EL: 0.16, 0.16, 1.18; SL: 0.71, 0.69, 0.70; SI: 57, 58, 57; PW: 0.86, 0.86, 0.90;WL: 1.54, 1.49, 1.58; petH: 0.85, 0.79, 0.90; petW: 0.46, 0.47, 0.47; GW 0.99, 0.96, 1.03.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="179" type="description">
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<paragraph box="[199,703,548,572]" pageId="28" pageNumber="179">
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<emphasis bold="true" box="[199,356,549,572]" pageId="28" pageNumber="179">Related taxa.</emphasis>
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See above under
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<taxonomicName box="[555,696,549,572]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="ecarinata">
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<emphasis box="[555,696,549,572]" italics="true" pageId="28" pageNumber="179">H. ecarinata</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="28" pageNumber="179">
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<emphasis bold="true" box="[199,600,583,608]" pageId="28" pageNumber="179">In-group sympatric associations.</emphasis>
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<emphasis box="[612,853,583,608]" italics="true" pageId="28" pageNumber="179">
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<taxonomicName box="[612,853,583,608]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="relicta">Heteroponera relicta</taxonomicName>
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</emphasis>
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and
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<taxonomicName box="[919,1101,583,608]" class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="pendergrasti">
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<emphasis box="[919,1101,583,608]" italics="true" pageId="28" pageNumber="179">H. pendergrasti</emphasis>
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</taxonomicName>
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are both present in QMBA Berlesate 804.
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</paragraph>
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<paragraph pageId="28" pageNumber="179">
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<emphasis bold="true" box="[199,371,655,680]" pageId="28" pageNumber="179">Biogeography.</emphasis>
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known only from Lamb Range, the northern rampart of the Atherton Tableland.
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<emphasis italics="true" pageId="28" pageNumber="179">
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<taxonomicName class="Insecta" family="Formicidae" genus="Heteroponera" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="179" phylum="Arthropoda" rank="species" species="pendergrasti">H. pendergrasti</taxonomicName>
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</emphasis>
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is possibly endemic there. Note the frequency of accessions by various collectors, and the absence of records in the well-known, generally well collected and more frequently visited nearby EA9, Atherton Tableland and EA6 Black Mountain localities.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="179" type="discussion">
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<paragraph pageId="28" pageNumber="179">
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<emphasis bold="true" box="[199,315,800,824]" pageId="28" pageNumber="179">Remarks.</emphasis>
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The apically sub-dentate petiolar node is (probably homoplasiously and convergently) similar to those of some Neotropical species (Brown 1958; Feitosa 2011).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="179" type="etymology">
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<paragraph pageId="28" pageNumber="179">
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<emphasis bold="true" box="[199,333,872,896]" pageId="28" pageNumber="179">Etymology.</emphasis>
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Named for Professor J.G. (Jim) Pendergrast of Auckland, New Zealand, my first teacher in Zoology at both High School and (in a later position) at The University of Auckland, where he supervised my M.Sc. degree.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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