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153 lines
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<document id="E097102F324EC27A8A790B92F26C10BF" ID-DOI="10.5281/zenodo.174106" ID-GBIF-Dataset="2239677a-be03-489b-a288-2112c6d383c8" ID-ISSN="11755326" ID-Zenodo-Dep="174106" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790819789" checkinUser="plazi" docAuthor="Tan, Jingjing, Ren, Dong & Shih, Chungkun" docDate="2006" docId="038D87F9FFDC98147120FBF6D341FE1C" docLanguage="en" docName="zt01326p068.pdf" docOrigin="Zootaxa 1326" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Priacma Leconte 1874" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="FFB4FF81FFDD98167028FFC0D71BFFCA" masterDocTitle="First record of fossil Priacma (Coleoptera: Archostemata: Cupedidae) from the Jehol Biota of western Liaoning, China" masterLastPageNumber="68" masterPageNumber="55" pageNumber="56" updateTime="1698219209044" updateUser="plazi">
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<mods:titleInfo id="498CCF5B79DD9E11FDB16153E611145E">
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<mods:title id="810505083C725F239CBDE3659346680A">First record of fossil Priacma (Coleoptera: Archostemata: Cupedidae) from the Jehol Biota of western Liaoning, China</mods:title>
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<mods:namePart id="2BA9540157478B6D826897054360880B">Tan, Jingjing</mods:namePart>
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<mods:namePart id="BCF5FBE4BC66F31AAA4B46AC1F474AB5">Ren, Dong</mods:namePart>
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<mods:namePart id="299593561DA81F85576A8D1CD93BBEB6">Shih, Chungkun</mods:namePart>
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<mods:typeOfResource id="08CAF6F01E263EFD59848D9939300466">text</mods:typeOfResource>
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<mods:title id="1254A2EA5D2F913216128AC88FD52FE6">Zootaxa</mods:title>
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<mods:date id="BC61ED4340E1BC3086967133CD315F45">2006</mods:date>
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<mods:detail id="9FA0C690AE0BA274A35422C2BC63CE05" type="volume">
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<mods:number id="87AF49AFCB94F3AFDF16018784D47745">1326</mods:number>
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<mods:identifier id="D7A9F3959E79688A4955658F0E99CD46" type="DOI">10.5281/zenodo.174106</mods:identifier>
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<treatment id="038D87F9FFDC98147120FBF6D341FE1C" ID-DOI="http://doi.org/10.5281/zenodo.6259429" ID-GBIF-Taxon="119350371" ID-Zenodo-Dep="6259429" LSID="urn:lsid:plazi:treatment:038D87F9FFDC98147120FBF6D341FE1C" httpUri="http://treatment.plazi.org/id/038D87F9FFDC98147120FBF6D341FE1C" lastPageId="2" lastPageNumber="57" pageId="1" pageNumber="56">
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<subSubSection id="C33E6564FFDC98177120FBF6D562FB9A" box="[264,633,1078,1104]" pageId="1" pageNumber="56" type="nomenclature">
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<paragraph id="8B9B36EFFFDC98177120FBF6D562FB9A" blockId="1.[264,633,1078,1104]" box="[264,633,1078,1104]" pageId="1" pageNumber="56">
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<heading id="D0D38183FFDC98177120FBF6D562FB9A" bold="true" box="[264,633,1078,1104]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
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<emphasis id="B950EAFDFFDC98177120FBF6D562FB9A" bold="true" box="[264,633,1078,1104]" pageId="1" pageNumber="56">
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Genus
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<taxonomicName id="4C244D6CFFDC98177177FBF6D562FB9A" ID-CoL="6422F" authority="Leconte, 1874" authorityName="Leconte" authorityYear="1874" box="[351,633,1078,1104]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B950EAFDFFDC98177177FBF6D6DEFB9A" bold="true" box="[351,453,1078,1104]" italics="true" pageId="1" pageNumber="56">Priacma</emphasis>
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<bibRefCitation id="EFB54B1EFFDC981771E5FBF6D562FB9A" author="Leconte" box="[461,633,1078,1104]" pageId="1" pageNumber="68" refString="LeConte, J. L. (1874) On Cupesidae of North America. Transactions of the American Entomological Society, 5, 87 - 88." type="journal article" year="1874">Leconte, 1874</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E6564FFDC98177120FB41D358FA41" pageId="1" pageNumber="56" type="materials_examined">
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<paragraph id="8B9B36EFFFDC98177120FB41D301FB51" blockId="1.[264,1323,1153,1899]" box="[264,1050,1153,1179]" pageId="1" pageNumber="56">
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<emphasis id="B950EAFDFFDC98177120FB41D6B1FB51" bold="true" box="[264,426,1153,1179]" pageId="1" pageNumber="56">
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<typeStatus id="549F884DFFDC98177120FB41D65EFB51" box="[264,325,1153,1179]" pageId="1" pageNumber="56">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C244D6CFFDC9817719AFB41D413FB51" authority="Leconte, 1861" authorityName="Leconte" authorityYear="1861" box="[434,776,1153,1179]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="serrata">
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<emphasis id="B950EAFDFFDC9817719AFB41D54CFB50" box="[434,599,1153,1178]" italics="true" pageId="1" pageNumber="56">Cupes serrata</emphasis>
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<bibRefCitation id="EFB54B1EFFDC98177276FB41D413FB51" author="Leconte" box="[606,776,1153,1179]" pageId="1" pageNumber="68" refString="LeConte, J. L. (1861) New species of Coleoptera inhabiting the Pacific district of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 66, 338 - 358." type="journal article" year="1861">Leconte, 1861</bibRefCitation>
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</taxonomicName>
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, by monotypy; Recent.
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</paragraph>
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<paragraph id="8B9B36EFFFDC98177110FB69D358FA41" blockId="1.[264,1323,1153,1899]" pageId="1" pageNumber="56">
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<emphasis id="B950EAFDFFDC98177110FB69D514FB09" bold="true" box="[312,527,1193,1219]" pageId="1" pageNumber="56">Included species.</emphasis>
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In addition to the new species described below, six species were known before.
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<taxonomicName id="4C244D6CFFDC981771E1FB11D419FB21" authority="Leconte, 1861" authorityName="Leconte" authorityYear="1861" box="[457,770,1233,1259]" class="Insecta" family="Cupedidae" genus="Cupes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="serrata">
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<emphasis id="B950EAFDFFDC981771E1FB11D559FB20" box="[457,578,1233,1258]" italics="true" pageId="1" pageNumber="56">P. serrata</emphasis>
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<bibRefCitation id="EFB54B1EFFDC98177279FB11D419FB21" author="Leconte" box="[593,770,1233,1259]" pageId="1" pageNumber="68" refString="LeConte, J. L. (1861) New species of Coleoptera inhabiting the Pacific district of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 66, 338 - 358." type="journal article" year="1861">Leconte, 1861</bibRefCitation>
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</taxonomicName>
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is the only one extant species from North
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<collectingCountry id="F333767FFFDC98177120FB39D66AFAD9" box="[264,369,1273,1299]" name="United States of America" pageId="1" pageNumber="56">America</collectingCountry>
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. Others are all fossil species from the lower Cretaceous:
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<taxonomicName id="4C244D6CFFDC981774B1FB39D6E0FAF1" authority="Ponomarenko, 1986" authorityName="Ponomarenko" authorityYear="1986" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="corrupta">
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<emphasis id="B950EAFDFFDC981774B1FB39D230FAD8" box="[1177,1323,1273,1298]" italics="true" pageId="1" pageNumber="56">P. corrupta</emphasis>
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<bibRefCitation id="EFB54B1EFFDC98177120FAE1D6E0FAF1" author="Ponomarenko" box="[264,507,1313,1339]" pageId="1" pageNumber="68" refString="Ponomarenko, A. G. (1986) Scarabaeida (= Coleoptera). In: Rasnitsyn, A. P. (Ed.), Insects in the Early Cretaceous ecosystems of the west Mongolia. Nauka, Moscow, 84 - 107. [in Russian]" type="book chapter" year="1986">Ponomarenko, 1986</bibRefCitation>
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</taxonomicName>
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from West
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<collectingCountry id="F333767FFFDC981772B1FAE1D415FAF1" box="[665,782,1313,1339]" name="Mongolia" pageId="1" pageNumber="56">Mongolia</collectingCountry>
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,
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<taxonomicName id="4C244D6CFFDC98177308FAE1D3D4FAF1" authority="Ponomarenko, 1997" authorityName="Ponomarenko" authorityYear="1997" box="[800,1231,1313,1339]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="longicapitis">
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<emphasis id="B950EAFDFFDC98177308FAE1D4D5FAF0" box="[800,974,1313,1338]" italics="true" pageId="1" pageNumber="56">P. longicapitis</emphasis>
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<bibRefCitation id="EFB54B1EFFDC981773F4FAE1D3D4FAF1" author="Ponomarenko" box="[988,1231,1313,1339]" pageId="1" pageNumber="68" refString="Ponomarenko, A. G. (1997) New beetles of the family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal, 31, 389 - 399." type="journal article" year="1997">Ponomarenko, 1997</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C244D6CFFDC9817753DFAE1D540FAA9" authority="Ponomarenko, 1997" authorityName="Ponomarenko" authorityYear="1997" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="oculata">
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<emphasis id="B950EAFDFFDC9817753DFAE1D67BFAA8" italics="true" pageId="1" pageNumber="56">P. oculata</emphasis>
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<bibRefCitation id="EFB54B1EFFDC98177143FA89D540FAA9" author="Ponomarenko" box="[363,603,1353,1379]" pageId="1" pageNumber="68" refString="Ponomarenko, A. G. (1997) New beetles of the family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal, 31, 389 - 399." type="journal article" year="1997">Ponomarenko, 1997</bibRefCitation>
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</taxonomicName>
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from South
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<collectingCountry id="F333767FFFDC981772D2FA89D474FAA9" box="[762,879,1353,1379]" name="Mongolia" pageId="1" pageNumber="56">Mongolia</collectingCountry>
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,
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<emphasis id="B950EAFDFFDC98177356FA89D4F3FAA8" box="[894,1000,1353,1378]" italics="true" pageId="1" pageNumber="56">P. s t r i a t a</emphasis>
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<bibRefCitation id="EFB54B1EFFDC981773DEFA89D3FCFAA9" author="Ponomarenko" box="[1014,1255,1353,1379]" pageId="1" pageNumber="68" refString="Ponomarenko, A. G. (2000) Beetles of the Family Cupedidae from the Lower Cretaceous locality of Semen, Transbaikalia. Paleontological Journal, 34, 317 - 322." type="journal article" year="2000">Ponomarenko, 2000</bibRefCitation>
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from East Transbaikalia, and
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<emphasis id="B950EAFDFFDC98177209FAB1D565FA40" box="[545,638,1393,1418]" italics="true" pageId="1" pageNumber="56">P. s a n z i i</emphasis>
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<bibRefCitation id="EFB54B1EFFDC981772A1FAB1D4A9FA41" author="Sorinao" box="[649,946,1393,1419]" pageId="1" pageNumber="68" refString="Sorinao, C. & Delcls, X. (2006) New cupedids from the lower Cretaceous of Spain and the palaeogeography of the family. Acta Palaeontologica Polonica, 51, 185 - 200." type="journal article" year="2006">Sorinao and Delcls, 2006</bibRefCitation>
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from
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<collectingCountry id="F333767FFFDC981773D1FAB1D35BFA41" box="[1017,1088,1393,1419]" name="Spain" pageId="1" pageNumber="56">Spain</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E6564FFDC98177110FA59D47CF901" pageId="1" pageNumber="56" type="diagnosis">
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<paragraph id="8B9B36EFFFDC98177110FA59D47CF901" blockId="1.[264,1323,1153,1899]" pageId="1" pageNumber="56">
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<emphasis id="B950EAFDFFDC98177110FA59D6AAFA79" bold="true" box="[312,433,1433,1459]" pageId="1" pageNumber="56">Diagnosis</emphasis>
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(based on the original description of the
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<typeStatus id="549F884DFFDC9817738CFA59D4CDFA79" box="[932,982,1433,1459]" pageId="1" pageNumber="56">type</typeStatus>
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species and on the study of four new species). Head with two pairs of subacute tubercles; antennae filiform, somewhat moniliform, less than half as long as entire insect; mandibles strongly extending, bidentate or tridentate at apex; gula wide, somewhat rectangular, widening posteriorly; genae separated for entire length by gula. Pronotum nearly rectangular, angles sharp; prosternum without tarsal groove; prosternal process only shallowly extending behind coxae. Elytra with nine or ten almost complete rows of punctures, dorsal surface convex, longitudinal ridges bearing small tubercles.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E6564FFDC98147110F919D341FE1C" lastPageId="2" lastPageNumber="57" pageId="1" pageNumber="56" type="discussion">
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<paragraph id="8B9B36EFFFDC98147110F919D414FE4C" blockId="1.[264,1323,1153,1899]" lastBlockId="2.[264,1324,284,470]" lastPageId="2" lastPageNumber="57" pageId="1" pageNumber="56">
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<emphasis id="B950EAFDFFDC98177110F919D6AAF939" bold="true" box="[312,433,1753,1779]" pageId="1" pageNumber="56">Remarks.</emphasis>
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The newly described cupedids from the Yixian Formation belong to the tribe Priacmini of the family
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<taxonomicName id="4C244D6CFFDC98177272F8C1D5CCF8D1" box="[602,727,1793,1819]" class="Insecta" family="Cupedidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="family">Cupedidae</taxonomicName>
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because of antennae less than half as long as entire insect, interantennal distance much greater than diameter of eyes and prosternal process only shallowly extending behind coxae. Priacmini includes two extinct genera (
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<bibRefCitation id="EFB54B1EFFDF98147138FEDCD511FEFC" author="Ponomarenko" box="[272,522,284,310]" pageId="2" pageNumber="68" refString="Ponomarenko, A. G. (1969) Historical development of the Coleoptera-Archostemata. Trudy Paleontologicheskogo Instituta, 125, 70 - 115. [in Russian]" type="journal article" year="1969">Ponomarenko 1969</bibRefCitation>
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):
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<taxonomicName id="4C244D6CFFDF98147204FEDCD4ABFEFC" authority="Ponomarenko, 1968" authorityName="Ponomarenko" authorityYear="1968" box="[556,944,284,310]" class="Insecta" family="Cupedidae" genus="Cupidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B950EAFDFFDF98147204FEDCD5BBFEFF" box="[556,672,284,309]" italics="true" pageId="2" pageNumber="57">Cupidium</emphasis>
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<bibRefCitation id="EFB54B1EFFDF9814729DFEDCD4ABFEFC" author="Ponomarenko" box="[693,944,284,310]" pageId="2" pageNumber="68" refString="Ponomarenko, A. G. (1968) Archostematan beetles from the Jurassic of Karatau (Coleoptera, Archostemata). In: Rohdendorf, B. B. (Ed.), Jurskie Nasekomye Karatau. Nauka, Moskva, 190 - 236. [in Russian]" type="book chapter" year="1968">Ponomarenko, 1968</bibRefCitation>
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</taxonomicName>
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(from the Late Jurassic of Kazakhastan),
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<taxonomicName id="4C244D6CFFDF98147190FE84D459FE94" authority="Ponomarenko, 1966" authorityName="Ponomarenko" authorityYear="1966" box="[440,834,324,350]" class="Insecta" family="Cupedidae" genus="Priacmopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
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<emphasis id="B950EAFDFFDF98147190FE84D550FE97" box="[440,587,324,349]" italics="true" pageId="2" pageNumber="57">Priacmopsis</emphasis>
|
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|
<bibRefCitation id="EFB54B1EFFDF9814727CFE84D459FE94" author="Ponomarenko" box="[596,834,324,350]" pageId="2" pageNumber="68" refString="Ponomarenko, A. G. (1966) New beetles of the family Cupedidae (Coleoptera) from Mesozoic Deposits of Transbaikalia. Revue d'Entomologie de l'URSS, 45, 138 - 143. [in Russian]" type="journal article" year="1966">Ponomarenko, 1966</bibRefCitation>
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</taxonomicName>
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(from the Early Cretaceous of
|
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<collectingCountry id="F333767FFFDF98147492FE84D237FE94" box="[1210,1324,324,350]" name="Mongolia" pageId="2" pageNumber="57">Mongolia</collectingCountry>
|
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|
and Siberia), and one extant genus
|
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<taxonomicName id="4C244D6CFFDF9814728BFEACD413FE4F" box="[675,776,364,389]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis id="B950EAFDFFDF9814728BFEACD413FE4F" box="[675,776,364,389]" italics="true" pageId="2" pageNumber="57">Priacma</emphasis>
|
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</taxonomicName>
|
|||
|
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.
|
|||
|
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</paragraph>
|
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<paragraph id="8B9B36EFFFDF98147110FE54D341FE1C" blockId="2.[264,1324,284,470]" pageId="2" pageNumber="57">
|
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|
|
The new fossils can be placed in
|
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|
|
<taxonomicName id="4C244D6CFFDF98147297FE54D43FFE67" box="[703,804,404,429]" class="Insecta" family="Cupedidae" genus="Priacma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B950EAFDFFDF98147297FE54D43FFE67" box="[703,804,404,429]" italics="true" pageId="2" pageNumber="57">Priacma</emphasis>
|
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|
</taxonomicName>
|
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|
|
based on the pedicel obviously shorter than third antennomere and anterior angles of pronotum sharp and extending.
|
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|
</paragraph>
|
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</subSubSection>
|
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|
</treatment>
|
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</document>
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